184 lines
21 KiB
XML
184 lines
21 KiB
XML
<document id="2447ECEFC3EEA4EA1421228AE5FE9425" ID-CLB-Dataset="7400" ID-DOI="10.11646/zootaxa.4951.1.5" ID-GBIF-Dataset="08feafa7-d493-476f-b92e-493ac9f31a3e" ID-ISSN="1175-5326" ID-Zenodo-Dep="4655795" ID-ZooBank="6BEAE3FC-629E-4BE4-88A5-E342C14272C3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617267136329" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2021" docId="DE5FC219820CFFC6FF0BCCD8FAE157CA" docLanguage="en" docName="zootaxa.4951.1.5.pdf" docOrigin="Zootaxa 4951 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heterotetramelus (Heterotetramelus) baloghi Jałoszyński 2021, comb. n." docType="treatment" docVersion="9" lastPageNumber="134" masterDocId="2266BA618208FFC2FF9CCC4FFFCC5162" masterDocTitle="New Caledonian species misplaced in Horaeomorphus Schaufuss belong in Heterotetramelus Franz (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="136" masterPageNumber="130" pageNumber="134" updateTime="1698935831422" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="4975FD76D36476B8F4221D7445ECBEFB">
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<mods:title id="35A2E783B3098CC83E791D9F732A76B0">New Caledonian species misplaced in Horaeomorphus Schaufuss belong in Heterotetramelus Franz (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:name id="0FEAD154861BA12D51D2F6BE7C942D01" type="personal">
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<mods:namePart id="CF3363DFDF41C99427CFEA1D586A89B9">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="67FD9DF95E5B610F55472596D3EBE460">Zootaxa</mods:title>
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<mods:part id="6DB8C149036854A1753A121C83E5B966">
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<mods:date id="C71FF0B056AFFBA2BEDE27164E92996C">2021</mods:date>
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<mods:number id="3A465C1674EC1F92F6A4812DAA06821E">2021-04-01</mods:number>
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<mods:number id="6BA30D8B505ACEFD67A40D07229ED377">4951</mods:number>
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<mods:number id="1A85835709F1768663EE969347E3B419">1</mods:number>
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<mods:start id="671021FB8E18B3E5518AF36BAF07D14D">130</mods:start>
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<mods:end id="63D04388761AF6DA90B4D94694BEA619">136</mods:end>
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<treatment id="DE5FC219820CFFC6FF0BCCD8FAE157CA" ID-DOI="http://doi.org/10.5281/zenodo.4683467" ID-GBIF-Taxon="181002200" ID-Zenodo-Dep="4683467" LSID="urn:lsid:plazi:treatment:DE5FC219820CFFC6FF0BCCD8FAE157CA" httpUri="http://treatment.plazi.org/id/DE5FC219820CFFC6FF0BCCD8FAE157CA" lastPageNumber="134" pageId="4" pageNumber="134">
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<subSubSection id="1EEC2084820CFFC6FF0BCCD8FC5451D3" box="[151,920,151,178]" pageId="4" pageNumber="134" type="nomenclature">
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<paragraph id="5649730F820CFFC6FF0BCCD8FC5451D3" blockId="4.[151,920,151,178]" box="[151,920,151,178]" pageId="4" pageNumber="134">
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<heading id="0D01C463820CFFC6FF0BCCD8FC5451D3" bold="true" box="[151,920,151,178]" fontSize="11" level="1" pageId="4" pageNumber="134" reason="1">
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<emphasis id="6482AF1D820CFFC6FF0BCCD8FC5451D3" bold="true" box="[151,920,151,178]" pageId="4" pageNumber="134">
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<taxonomicName id="91F6088C820CFFC6FF0BCCD8FCEC51D3" ID-CoL="8Y4HG" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2021" baseAuthorityName="Franz" box="[151,800,151,178]" class="Insecta" family="Staphylinidae" genus="Heterotetramelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="134" phylum="Arthropoda" rank="species" species="baloghi" status="comb. nov." subGenus="Heterotetramelus">
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<emphasis id="6482AF1D820CFFC6FF0BCCD8FEA551D0" bold="true" box="[151,361,151,178]" italics="true" pageId="4" pageNumber="134">Heterotetramelus</emphasis>
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(
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<emphasis id="6482AF1D820CFFC6FEE6CCD8FD8051D0" bold="true" box="[378,588,151,178]" italics="true" pageId="4" pageNumber="134">Heterotetramelus</emphasis>
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)
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<emphasis id="6482AF1D820CFFC6FDC2CCD8FD7551D0" bold="true" box="[606,697,151,178]" italics="true" pageId="4" pageNumber="134">baloghi</emphasis>
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(Franz)
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</taxonomicName>
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,
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<taxonomicNameLabel id="7FB11266820CFFC6FCB2CCD8FC5451D3" box="[814,920,151,177]" pageId="4" pageNumber="134" rank="species">comb. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="1EEC2084820CFFC6FF0BCC93FF20507A" pageId="4" pageNumber="134" type="reference_group">
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<paragraph id="5649730F820CFFC6FF0BCC93FDF25191" blockId="4.[151,574,219,280]" box="[151,574,219,244]" pageId="4" pageNumber="134">
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<treatmentCitationGroup id="76E65421820CFFC6FF0BCC93FDF25191" box="[151,574,219,244]" pageId="4" pageNumber="134">
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<treatmentCitation id="D757551E820CFFC6FF0BCC93FDF65191" author="Franz, H." box="[151,570,219,244]" page="66" pageId="4" pageNumber="134" year="1979">
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<taxonomicName id="91F6088C820CFFC6FF0BCC93FDF65191" authority="Franz, 1979: 66" authorityName="Franz" authorityPageNumber="66" authorityYear="1979" box="[151,570,219,244]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="134" phylum="Arthropoda" rank="species" species="baloghi">
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<emphasis id="6482AF1D820CFFC6FF0BCC93FE5D5196" box="[151,401,220,244]" italics="true" pageId="4" pageNumber="134">Horaeomorphus baloghi</emphasis>
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<bibRefCitation id="32670EFE820CFFC6FE0BCC93FDF65191" author="Franz, H." box="[407,570,219,244]" pageId="4" pageNumber="134" pagination="61 - 67" refId="ref4346" refString="Franz, H. (1979) Scydmaeniden aus Neukaledonien (Coleoptera). II. Acta Zoologica Academiae Scientarum Hungaricae, 25 (1 - 2), 61 - 67." type="journal article" year="1979">Franz, 1979: 66</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="5649730F820CFFC6FF0BCCB1FF20507A" blockId="4.[151,574,219,280]" box="[151,236,254,280]" pageId="4" pageNumber="134">
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(
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<figureCitation id="CECD6F8A820CFFC6FF3CCCB1FF2F507A" box="[160,227,254,280]" captionStart="FIGURES 1–3" captionStartId="3.[151,275,1233,1259]" captionTargetBox="[151,1436,179,1209]" captionTargetId="figure-19@3.[151,1436,179,1209]" captionTargetPageId="3" captionText="FIGURES 1–3. Dorsal habitus of type specimens. Heterotetramelus australis (Franz), holotype female (1); H. baloghi (Franz), paratype female (2); H. novaecaledoniae (Franz), holotype male (3)." figureDoi="http://doi.org/10.5281/zenodo.4655797" httpUri="https://zenodo.org/record/4655797/files/figure.png" pageId="4" pageNumber="134">Fig. 2</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="1EEC2084820CFFC6FF0BCD09FECB53E6" pageId="4" pageNumber="134" type="materials_examined">
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<paragraph id="5649730F820CFFC6FF0BCD09FB8D50CB" blockId="4.[151,1437,326,1705]" pageId="4" pageNumber="134">
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<emphasis id="6482AF1D820CFFC6FF0BCD09FE725002" bold="true" box="[151,446,326,352]" pageId="4" pageNumber="134">Type material examined.</emphasis>
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<materialsCitation id="E69E7952820CFFC6FE59CD09FBD050E7" ID-GBIF-Occurrence="3071254301" collectingDate="1977-02-12" collectingDateMax="1977-02-13" collectingDateMin="1977-02-12" collectorName="Dr. J." country="New Caledonia" location="Noumea" municipality="Mt. Koghi" pageId="4" pageNumber="134" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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<emphasis id="6482AF1D820CFFC6FE59CD09FC855002" bold="true" box="[453,841,326,352]" pageId="4" pageNumber="134">
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<typeStatus id="894DCDAD820CFFC6FE59CD09FDFE5002" box="[453,562,326,352]" pageId="4" pageNumber="134" type="paratype">Paratype</typeStatus>
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(
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<collectingCountry id="2EE1339F820CFFC6FDDCCD09FCF05002" box="[576,828,326,352]" name="New Caledonia" pageId="4" pageNumber="134">NEW CALEDONIA</collectingCountry>
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):
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</emphasis>
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<specimenCount id="40F0B886820CFFC6FCD3CD08FCA85003" box="[847,868,327,353]" pageId="4" pageNumber="134" type="female">♀</specimenCount>
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, four labels: „NEW-CALEDONIE /
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<location id="532925D4820CFFC6FA9DCD09FAAC5002" LSID="urn:lsid:plazi:treatment:DE5FC219820CFFC6FF0BCCD8FAE157CA:532925D4820CFFC6FA9DCD09FAAC5002" box="[1281,1376,326,352]" country="New Caledonia" municipality="Mt. Koghi" name="Noumea" pageId="4" pageNumber="134">Nouméa</location>
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/
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<collectingMunicipality id="B62DE975820CFFC6FAE9CD08FF2C50E7" pageId="4" pageNumber="134">Mt. Koghi</collectingMunicipality>
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” [white, printed], “
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<date id="224855CF820CFFC6FE2ACD24FD9E50E7" box="[438,594,363,389]" pageId="4" pageNumber="134" value="1977-02-12" valueMax="1977-02-13" valueMin="1977-02-12">
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<collectingDate id="320CAC27820CFFC6FE2ACD24FD9E50E7" box="[438,594,363,389]" pageId="4" pageNumber="134" value="1977-02-12" valueMax="1977-02-13" valueMin="1977-02-12">12-13.II.1977</collectingDate>
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</date>
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/ leg.
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<collectorName id="FB0316D9820CFFC6FD0CCD24FD0950E7" box="[656,709,363,389]" pageId="4" pageNumber="134">Dr.J.</collectorName>
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BALOGH” [white, printed], “
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</materialsCitation>
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<materialsCitation id="E69E7952820CFFC6FB80CD24FBF150CB" ID-GBIF-Occurrence="3071254302" collectionCode="NHMW" country="New Caledonia" pageId="4" pageNumber="134" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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<specimenCount id="40F0B886820CFFC6FB80CD24FBE250E7" box="[1052,1070,363,389]" pageId="4" pageNumber="134" type="female">♀</specimenCount>
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” [white, handwritten], „Horaeo- morphus / baloghi m. /
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<typeStatus id="894DCDAD820CFFC6FE01CDC0FDF250CB" box="[413,574,399,425]" pageId="4" pageNumber="134" type="paratype">PARATYPUS</typeStatus>
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” [yellow, handwritten and printed] (
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<collectionCode id="30E7EBCA820CFFC6FC4FCDC0FBF950CB" box="[979,1077,399,425]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="4" pageNumber="134" type="Museum">NHMW</collectionCode>
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)
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</materialsCitation>
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||
.
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</paragraph>
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<paragraph id="5649730F820CFFC6FF5BCDFBFECB53E6" blockId="4.[151,1437,326,1705]" pageId="4" pageNumber="134">
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<emphasis id="6482AF1D820CFFC6FF5BCDFBFE7B50AC" bold="true" box="[199,439,436,462]" pageId="4" pageNumber="134">Emended diagnosis.</emphasis>
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Female body over
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<quantity id="910EDEEA820CFFC6FD08CDFBFD1A50AC" box="[660,726,436,462]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="134" unit="mm" value="2.0">2 mm</quantity>
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in length, bicolored and with strongly contrasting parts: elytra and legs brown with umbra hue, head and prothorax nearly black, scapes nearly as dark as head, remaining antennomeres becoming slightly lighter toward antennal apices; eyes large (diameter much longer than apical width of scape), bean-shaped and strongly convex; antennomeres 7–8 each elongate (8 indistinctly so), 9–10 each as long as broad; pronotum lacking antebasal pits, with barely perceivable, superficial transverse impression. Male characters unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="1EEC2084820CFFC6FF5BCEC0FBA154CA" pageId="4" pageNumber="134" type="description">
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<paragraph id="5649730F820CFFC6FF5BCEC0FEE05275" blockId="4.[151,1437,326,1705]" pageId="4" pageNumber="134">
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<emphasis id="6482AF1D820CFFC6FF5BCEC0FEB953CB" bold="true" box="[199,373,655,681]" pageId="4" pageNumber="134">Redescription.</emphasis>
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Male unknown. Body of female (
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<figureCitation id="CECD6F8A820CFFC6FD60CEC0FCF253CB" box="[764,830,655,681]" captionStart="FIGURES 1–3" captionStartId="3.[151,275,1233,1259]" captionTargetBox="[151,1436,179,1209]" captionTargetId="figure-19@3.[151,1436,179,1209]" captionTargetPageId="3" captionText="FIGURES 1–3. Dorsal habitus of type specimens. Heterotetramelus australis (Franz), holotype female (1); H. baloghi (Franz), paratype female (2); H. novaecaledoniae (Franz), holotype male (3)." figureDoi="http://doi.org/10.5281/zenodo.4655797" httpUri="https://zenodo.org/record/4655797/files/figure.png" pageId="4" pageNumber="134">Fig. 1</figureCitation>
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) moderately slender and strongly convex, bicolored: elytra and legs brown with umbra hue, head and prothorax nearly black, scapes nearly as dark as head, remaining antennomeres becoming slightly lighter toward antennal apices, setae distinctly lighter than elytra, yellowish-brown; BL
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||
<quantity id="910EDEEA820CFFC6FF5DCEB3FEE45275" box="[193,296,764,791]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="134" unit="mm" value="2.5">2.50 mm</quantity>
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.
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</paragraph>
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<paragraph id="5649730F820CFFC6FF5BCF6EFD105559" blockId="4.[151,1437,326,1705]" pageId="4" pageNumber="134">
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Head sub-pentagonal, broadest at eyes, HL
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||
<quantity id="910EDEEA820CFFC6FD2FCF6EFCD5525E" box="[691,793,801,828]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="4" pageNumber="134" unit="mm" value="0.45">0.45 mm</quantity>
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, HW
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<quantity id="910EDEEA820CFFC6FCC7CF6EFC0F525E" box="[859,963,801,828]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="134" unit="mm" value="0.5">0.50 mm</quantity>
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. Vertex and frons confluent, together sub- trapezoidal and weakly convex; supraantennal tubercles weakly elevated; tempora in dorsal view 1.5 × as long as eyes; eyes large (eye diameter much longer than apical width of scape), in lateral view bean-shaped with distinct posterior emargination, strongly convex and finely faceted. Punctures on frons and vertex very small but distinct, sharply marked and dense, separated by spaces subequal to diameters of punctures; setae dense, moderately long, suberect or erect; tempora with distinct long and dense thick bristles. Antennae slender and weakly gradually thickened, AnL
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||
<quantity id="910EDEEA820CFFC6FE8DCFB2FEBA5575" box="[273,374,1021,1047]" metricMagnitude="-3" metricUnit="m" metricValue="1.38" pageId="4" pageNumber="134" unit="mm" value="1.38">1.38 mm</quantity>
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, antennomeres 1–8 each elongate (8 weakly so), 9 and 10 each about as long as broad, 11 much shorter than 9–10 combined, 1.8 × as long as broad.
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</paragraph>
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<paragraph id="5649730F820CFFC6FF5BC809FDCE5590" blockId="4.[151,1437,326,1705]" pageId="4" pageNumber="134">
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Pronotum in dorsal view bell-shaped, broadest near anterior third, PL
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<quantity id="910EDEEA820CFFC6FC45C809FBF35502" box="[985,1087,1094,1120]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="4" pageNumber="134" unit="mm" value="0.73">0.73 mm</quantity>
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||
, PW
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||
<quantity id="910EDEEA820CFFC6FBE0C80AFB2F5502" box="[1148,1251,1093,1120]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="4" pageNumber="134" unit="mm" value="0.58">0.58 mm</quantity>
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||
; anterior margin and sides in anterior third rounded together; lateral margins in posterior half distinctly sinuate, strongly constricted near posterior third; posterior corners indistinctly obtuse-angled and blunt; posterior margin weakly arcuate. Base of pronotum with barely discernible, superficial transverse impression, lacking pits. Punctures, setae and lateral bristles similar to those on head.
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||
</paragraph>
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<paragraph id="5649730F820CFFC6FF5BC8B3FD0354E6" blockId="4.[151,1437,326,1705]" pageId="4" pageNumber="134">
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||
Elytra together oval, strongly and evenly convex, broadest distinctly in front of middle; EL
|
||
<quantity id="910EDEEA820CFFC6FB54C8B3FAE25475" box="[1224,1326,1276,1303]" metricMagnitude="-3" metricUnit="m" metricValue="1.33" pageId="4" pageNumber="134" unit="mm" value="1.33">1.33 mm</quantity>
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||
, EW
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||
<quantity id="910EDEEA820CFFC6FAF7C8B3FF095459" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="4" pageNumber="134" unit="mm" value="1.05">1.05 mm</quantity>
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||
, EI 1.26; basal elytral impressions short but distinct; humeral calli weakly elevated, elongate; elytral apices separately rounded. Punctures on elytra dense but very shallow and inconspicuous; setae distinctly thicker and denser than those on head and pronotum, suberect.
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||
</paragraph>
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||
<paragraph id="5649730F820CFFC6FF5BC9C0FBA154CA" blockId="4.[151,1437,326,1705]" box="[199,1133,1422,1449]" pageId="4" pageNumber="134">Legs moderately long and slender, pro- and mesotibiae indistinctly curved inwards.</paragraph>
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</subSubSection>
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<subSubSection id="1EEC2084820CFFC6FF5BC9FCFAE157CA" pageId="4" pageNumber="134" type="discussion">
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<paragraph id="5649730F820CFFC6FF5BC9FCFAE157CA" blockId="4.[151,1437,326,1705]" pageId="4" pageNumber="134">
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||
<emphasis id="6482AF1D820CFFC6FF5BC9FCFEF754AF" bold="true" box="[199,315,1459,1485]" pageId="4" pageNumber="134">Remarks.</emphasis>
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During this study it was not possible to examine the
|
||
<typeStatus id="894DCDAD820CFFC6FC16C9FCFC2054AF" box="[906,1004,1459,1485]" pageId="4" pageNumber="134" type="holotype">holotype</typeStatus>
|
||
of this species. However, the
|
||
<typeStatus id="894DCDAD820CFFC6FAA6C9FCFA5054AF" box="[1338,1436,1459,1485]" pageId="4" pageNumber="134" type="holotype">holotype</typeStatus>
|
||
is a female, and therefore the identification of
|
||
<taxonomicName id="91F6088C820CFFC6FD3FC997FCD05493" authorityName="Franz" authorityYear="1979" box="[675,796,1495,1521]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="134" phylum="Arthropoda" rank="species" species="baloghi">
|
||
<emphasis id="6482AF1D820CFFC6FD3FC997FCD05493" box="[675,796,1495,1521]" italics="true" pageId="4" pageNumber="134">H. baloghi</emphasis>
|
||
</taxonomicName>
|
||
may be still problematic, as species of
|
||
<taxonomicName id="91F6088C820CFFC6FB45C998FA505493" authorityName="Franz" authorityYear="1971" box="[1241,1436,1495,1521]" class="Insecta" family="Staphylinidae" genus="Heterotetramelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="134" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6482AF1D820CFFC6FB45C998FA505493" box="[1241,1436,1495,1521]" italics="true" pageId="4" pageNumber="134">Heterotetramelus</emphasis>
|
||
</taxonomicName>
|
||
may be similar one to another. Three specimens (one male and two females) collected on Mt. Koghi, the type local- ity of
|
||
<taxonomicName id="91F6088C820CFFC6FF40CA6EFE995758" authorityName="Franz" authorityYear="1979" box="[220,341,1568,1594]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="134" phylum="Arthropoda" rank="species" species="baloghi">
|
||
<emphasis id="6482AF1D820CFFC6FF40CA6EFE995758" box="[220,341,1568,1594]" italics="true" pageId="4" pageNumber="134">H. baloghi</emphasis>
|
||
</taxonomicName>
|
||
, were seen, which very much resemble the
|
||
<typeStatus id="894DCDAD820CFFC6FCDBCA6EFC6B5759" box="[839,935,1569,1595]" pageId="4" pageNumber="134" type="paratype">paratype</typeStatus>
|
||
of this species. The only differences are the eye of females conspicuously smaller and less convex than those in
|
||
<taxonomicName id="91F6088C820CFFC6FC36CA09FBE8573D" authorityName="Franz" authorityYear="1979" box="[938,1060,1605,1631]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="134" phylum="Arthropoda" rank="species" species="baloghi">
|
||
<emphasis id="6482AF1D820CFFC6FC36CA09FBE8573D" box="[938,1060,1605,1631]" italics="true" pageId="4" pageNumber="134">H. baloghi</emphasis>
|
||
</taxonomicName>
|
||
, slightly different proportions of antennomeres, and distinct (but still relatively shallow) antebasal pronotal pits. It is possible that pairs of extremely similar species occur sympatrically in
|
||
<collectingCountry id="2EE1339F820CFFC6FDD9CAC0FD3957CA" box="[581,757,1678,1705]" name="New Caledonia" pageId="4" pageNumber="134">New Caledonia</collectingCountry>
|
||
, and care must be taken when identifying females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |