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<document id="E7BBE4F97C7E150D2D9D6EC46D91D4A5" ID-DOI="10.11646/zootaxa.4755.2.10" ID-GBIF-Dataset="0cb0c88f-3e8b-4552-9567-18b4178d846d" ID-ISSN="1175-5334" ID-Zenodo-Dep="3733685" ID-ZooBank="urn:lsid:zoobank.org:pub:4D1861B9-0582-4AE8-9A34-3F903B872D08" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1585597297038" checkinUser="carolina" docAuthor="Albornoz-Garzón, Juan G., Acosta-Santos, Astrid, Bogotá-Gregory, Juan D. &amp; Agudelo-Córdoba, Edwin" docDate="2020" docId="830C1D4EFFA53F41FF4CFBD0FE6C3088" docLanguage="en" docName="zootaxa.4755.2.10.pdf.imf" docOrigin="Zootaxa 4755 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Creagrutus andaki Albornoz-Garzón &amp; Acosta-Santos &amp; Bogotá-Gregory &amp; Agudelo-Córdoba 2020, new species" docType="treatment" docVersion="7" lastPageNumber="370" masterDocId="7F356536FFA43F44FFDBFFB5FFFB3652" masterDocTitle="A new species of Creagrutus Günther (Characidae: Stevardiinae) from the Amazonian Piedmont of Colombia" masterLastPageNumber="374" masterPageNumber="365" pageNumber="366" updateTime="1698733407910" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="CCE6F770EE691D6F44F8F673B619ADE1">A new species of Creagrutus Günther (Characidae: Stevardiinae) from the Amazonian Piedmont of Colombia</mods:title>
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<mods:namePart id="05C4C8CE4663C405F4487D3877057C9F">Albornoz-Garzón, Juan G.</mods:namePart>
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<mods:namePart id="959A6FB1B03FDCD0F2FA79C288240E54">Acosta-Santos, Astrid</mods:namePart>
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<mods:namePart id="BFAFA479267CB580B6E1B311F6E50F16">Bogotá-Gregory, Juan D.</mods:namePart>
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<treatment id="830C1D4EFFA53F41FF4CFBD0FE6C3088" ID-DOI="http://doi.org/10.5281/zenodo.3812491" ID-GBIF-Taxon="163392050" ID-Zenodo-Dep="3812491" LSID="urn:lsid:plazi:treatment:830C1D4EFFA53F41FF4CFBD0FE6C3088" httpUri="http://treatment.plazi.org/id/830C1D4EFFA53F41FF4CFBD0FE6C3088" lastPageId="5" lastPageNumber="370" pageId="1" pageNumber="366">
<subSubSection id="43BFFFD3FFA53F45FF4CFBD0FDE2322D" box="[151,537,1125,1152]" pageId="1" pageNumber="366" type="nomenclature">
<paragraph id="0B1AAC58FFA53F45FF4CFBD0FDE2322D" blockId="1.[151,537,1125,1187]" box="[151,537,1125,1152]" pageId="1" pageNumber="366">
<heading id="50521B34FFA53F45FF4CFBD0FDE2322D" bold="true" box="[151,537,1125,1152]" fontSize="10" level="2" pageId="1" pageNumber="366" reason="2">
<emphasis id="39D1704AFFA53F45FF4CFBD0FDE2322D" bold="true" box="[151,537,1125,1152]" pageId="1" pageNumber="366">
<emphasis id="39D1704AFFA53F45FF4CFBD0FE7932D2" bold="true" box="[151,386,1125,1152]" italics="true" pageId="1" pageNumber="366">
<taxonomicName id="CCA5D7DBFFA53F45FF4CFBD0FE8032D2" authority="Albornoz-Garzón &amp; Acosta-Santos &amp; Bogotá-Gregory &amp; Agudelo-Córdoba, 2020" authorityName="Albornoz-Garzón &amp; Acosta-Santos &amp; Bogotá-Gregory &amp; Agudelo-Córdoba" authorityYear="2020" box="[151,379,1125,1152]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="366" phylum="Chordata" rank="species" species="andaki" status="sp. nov.">Creagrutus andaki</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="22E2CD31FFA53F45FE52FBD3FDE2322D" box="[393,537,1125,1152]" pageId="1" pageNumber="366" rank="species">new species</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="43BFFFD3FFA53F45FF4CFB3CFE9732F1" box="[151,364,1161,1187]" pageId="1" pageNumber="366" type="description">
<paragraph id="0B1AAC58FFA53F45FF4CFB3CFE9732F1" blockId="1.[151,537,1125,1187]" box="[151,364,1161,1187]" pageId="1" pageNumber="366">
(
<figureCitation id="939EB0DDFFA53F45FF7BFB3CFEFF32F1" box="[160,260,1161,1187]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,250,1908,1933]" captionStartId-1="2.[151,250,1352,1377]" captionStartId-2="3.[151,250,1281,1306]" captionTargetBox-0="[183,1404,1418,1881]" captionTargetBox-1="[265,1323,181,1327]" captionTargetBox-2="[333,1255,181,1255]" captionTargetId-0="figure@1.[183,1404,1418,1881]" captionTargetId-1="figure@2.[265,1323,181,1327]" captionTargetId-2="figure@3.[333,1255,181,1255]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Creagrutus andaki, holotype, CIACOL 3462, 56.8 mm SL, Colombia, Caquetá, Florencia, Río Caquetá basin, Río Hacha drainage, Río Sucre." captionText-1="FIGURE 2. Creagrutus andaki, paratypes: (A) CIACOL 2910, 58.7 mm LS. (B) CZUT-IC 19973, 57.4 mm LS. (C) CIACOL 3463, 38.6 mm LS. All from Colombia, Caquetá, Florencia, Río Caquetá basin, Río Hacha drainage, Río Sucre." captionText-2="FIGURE 3. Creagrutus andaki, IAvH-P 20565, paratype, 50.6 mm SL, premaxillary teeth in ventral view, left side." figureDoi-0="http://doi.org/10.5281/zenodo.3733687" figureDoi-1="http://doi.org/10.5281/zenodo.3733689" figureDoi-2="http://doi.org/10.5281/zenodo.3733691" httpUri-0="https://zenodo.org/record/3733687/files/figure.png" httpUri-1="https://zenodo.org/record/3733689/files/figure.png" httpUri-2="https://zenodo.org/record/3733691/files/figure.png" pageId="1" pageNumber="366">Figs.1–3</figureCitation>
; Table 1)
</paragraph>
</subSubSection>
<subSubSection id="43BFFFD3FFA53F45FF4CFB78FC9332B4" box="[151,872,1229,1254]" pageId="1" pageNumber="366" type="description">
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urn:lsid:zoobank.org:act:
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<subSubSection id="43BFFFD3FFA53F46FF4CFAA1FED4300C" lastPageId="2" lastPageNumber="367" pageId="1" pageNumber="366" type="materials_examined">
<paragraph id="0B1AAC58FFA53F45FF4CFAA1FDD93324" blockId="1.[151,1436,1299,1398]" pageId="1" pageNumber="366">
<materialsCitation id="BBCDA605FFA53F45FF4CFAA1FDD93324" ID-GBIF-Occurrence="2592573804" collectingDate="2018-08-10" collectingDateMax="2017-08-12" collectingDateMin="2016-08-12" collectionCode="CIACOL" collectorName="J. D. Bogota-Gregory &amp; I. Gonzalez-Gomez" country="Colombia" county="Florencia" elevation="1015" latitude="1.7953333" location="Rio Hacha" longLatPrecision="1" longitude="-75.64691" municipality="Rio" pageId="1" pageNumber="366" specimenCode="CIACOL 2130, CZUT-IC 19973, IAvH-P 20565, CIACOL 2910" specimenCount="1" stateProvince="Caqueta" typeStatus="holotype">
<emphasis id="39D1704AFFA53F45FF4CFAA1FEF0337C" bold="true" box="[151,267,1300,1326]" pageId="1" pageNumber="366">
<typeStatus id="D41E12FAFFA53F45FF4CFAA1FEFD337C" box="[151,262,1300,1326]" pageId="1" pageNumber="366" type="holotype">Holotype</typeStatus>
.
</emphasis>
CIACOL 3462,
<quantity id="CC5D01BDFFA53F45FE16FAA6FDC8337D" box="[461,563,1299,1327]" metricMagnitude="-2" metricUnit="m" metricValue="5.68" pageId="1" pageNumber="366" unit="mm" value="56.8">56.8 mm</quantity>
SL,
<collectingCountry id="73B2ECC8FFA53F45FDB7FAA1FD26337C" box="[620,733,1300,1326]" name="Colombia" pageId="1" pageNumber="366">Colombia</collectingCountry>
,
<collectingRegion id="C96162BAFFA53F45FD30FAA1FCB1337C" box="[747,842,1300,1326]" country="Colombia" name="Caqueta" pageId="1" pageNumber="366">Caquetá</collectingRegion>
,
<collectingCounty id="E27BD4D4FFA53F45FC8CFAA1FC3F337C" box="[855,964,1300,1326]" pageId="1" pageNumber="366">Florencia</collectingCounty>
,
<collectingMunicipality id="EB7E3622FFA53F45FC0AFAA1FC02337C" box="[977,1017,1300,1326]" pageId="1" pageNumber="366">Río</collectingMunicipality>
<collectingRegion id="C96162BAFFA53F45FBD9FAA1FBA5337C" box="[1026,1118,1300,1326]" country="Colombia" name="Caqueta" pageId="1" pageNumber="366">Caquetá</collectingRegion>
basin,
<location id="0E7AFA83FFA53F45FB68FAA1FAD7337C" LSID="urn:lsid:plazi:treatment:830C1D4EFFA53F41FF4CFBD0FE6C3088:0E7AFA83FFA53F45FB68FAA1FAD7337C" box="[1203,1324,1300,1326]" country="Colombia" county="Florencia" latitude="1.7953333" longLatPrecision="1" longitude="-75.64691" municipality="Rio" name="Rio Hacha" pageId="1" pageNumber="366" stateProvince="Caqueta">Río Hacha</location>
drainage,
<location id="0E7AFA83FFA53F45FF4CFA8DFF443300" LSID="urn:lsid:plazi:treatment:830C1D4EFFA53F41FF4CFBD0FE6C3088:0E7AFA83FFA53F45FF4CFA8DFF443300" box="[151,191,1336,1362]" country="Colombia" county="Florencia" latitude="1.7953333" longLatPrecision="1" longitude="-75.64691" municipality="Rio" name="Rio" pageId="1" pageNumber="366" stateProvince="Caqueta">Río</location>
<collectingRegion id="C96162BAFFA53F45FF1DFA8DFEF23300" box="[198,265,1336,1362]" country="Colombia" name="Sucre" pageId="1" pageNumber="366">Sucre</collectingRegion>
, km 61, bridge at old road to
<location id="0E7AFA83FFA53F45FD94FA8DFD6E3300" LSID="urn:lsid:plazi:treatment:830C1D4EFFA53F41FF4CFBD0FE6C3088:0E7AFA83FFA53F45FD94FA8DFD6E3300" box="[591,661,1336,1362]" country="Colombia" county="Florencia" latitude="1.7953333" longLatPrecision="1" longitude="-75.64691" municipality="Rio" name="Neiva" pageId="1" pageNumber="366" stateProvince="Caqueta">Neiva</location>
,
<geoCoordinate id="6E91CA9FFFA53F45FD7BFA8DFCC53300" box="[672,830,1336,1362]" degrees="01" direction="north" minutes="47" orientation="latitude" pageId="1" pageNumber="366" precision="1" seconds="43.2" value="1.7953333">01º47’43.2”N</geoCoordinate>
,
<geoCoordinate id="6E91CA9FFFA53F45FC92FA82FC103300" box="[841,1003,1335,1362]" degrees="75" direction="west" minutes="38" orientation="longitude" pageId="1" pageNumber="366" precision="1" seconds="48.9" value="-75.64691">75º38’48.9”W</geoCoordinate>
,
<quantity id="CC5D01BDFFA53F45FC2DFA82FB973300" box="[1014,1132,1335,1362]" metricMagnitude="3" metricUnit="m" metricValue="1.015" pageId="1" pageNumber="366" unit="m" value="1015.0">
<elevation id="80884B6BFFA53F45FC2DFA82FB973300" box="[1014,1132,1335,1362]" metricMagnitude="3" metricUnit="m" metricValue="1.015" pageId="1" pageNumber="366" unit="m" value="1015.0">1015 masl</elevation>
</quantity>
;
<collectorName id="A650C98EFFA53F45FBACFA8CFA913300" box="[1143,1386,1336,1363]" pageId="1" pageNumber="366">J. D. Bogotá-Gregory</collectorName>
&amp;
<collectorName id="A650C98EFFA53F45FA57FA8CFEA53324" pageId="1" pageNumber="366">I. González-Gómez</collectorName>
,
<date id="7F1B8A98FFA53F45FEB2FAE9FDE53324" box="[361,542,1372,1398]" pageId="1" pageNumber="366" value="2018-08-10">
<collectingDate id="6F5F7370FFA53F45FEB2FAE9FDE53324" box="[361,542,1372,1398]" pageId="1" pageNumber="366" value="2018-08-10">10 August 2018</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<caption id="5FDAFCD0FFA53F45FF4CF8C1FE5531E3" ID-DOI="http://doi.org/10.5281/zenodo.3733687" ID-Zenodo-Dep="3733687" httpUri="https://zenodo.org/record/3733687/files/figure.png" pageId="1" pageNumber="366" startId="1.[151,250,1908,1933]" targetBox="[183,1404,1418,1881]" targetPageId="1">
<paragraph id="0B1AAC58FFA53F45FF4CF8C1FE5531E3" blockId="1.[151,1437,1908,1969]" pageId="1" pageNumber="366">
<emphasis id="39D1704AFFA53F45FF4CF8C1FEE831DF" bold="true" box="[151,275,1908,1933]" pageId="1" pageNumber="366">FIGURE 1.</emphasis>
<taxonomicName id="CCA5D7DBFFA53F45FEC2F8C0FE2C31DF" authorityName="Albornoz-Garzón &amp; Acosta-Santos &amp; Bogotá-Gregory &amp; Agudelo-Córdoba" authorityYear="2020" box="[281,471,1909,1933]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="366" phylum="Chordata" rank="species" species="andaki">
<emphasis id="39D1704AFFA53F45FEC2F8C0FE2C31DF" box="[281,471,1909,1933]" italics="true" pageId="1" pageNumber="366">Creagrutus andaki</emphasis>
</taxonomicName>
, holotype, CIACOL 3462, 56.8 mm SL, Colombia, Caquetá, Florencia, Río Caquetá basin, Río Hacha drainage, Río Sucre.
</paragraph>
</caption>
<caption id="5FDAFCD0FFA63F46FF4CFAFDFAF833D7" ID-DOI="http://doi.org/10.5281/zenodo.3733689" ID-Zenodo-Dep="3733689" httpUri="https://zenodo.org/record/3733689/files/figure.png" pageId="2" pageNumber="367" startId="2.[151,250,1352,1377]" targetBox="[265,1323,181,1327]" targetPageId="2">
<paragraph id="0B1AAC58FFA63F46FF4CFAFDFAF833D7" blockId="2.[151,1436,1352,1413]" pageId="2" pageNumber="367">
<emphasis id="39D1704AFFA63F46FF4CFAFDFEE83333" bold="true" box="[151,275,1352,1377]" pageId="2" pageNumber="367">FIGURE 2.</emphasis>
<taxonomicName id="CCA5D7DBFFA63F46FEC1FAFCFE233333" authorityName="Albornoz-Garzón &amp; Acosta-Santos &amp; Bogotá-Gregory &amp; Agudelo-Córdoba" authorityYear="2020" box="[282,472,1353,1377]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="367" phylum="Chordata" rank="species" species="andaki">
<emphasis id="39D1704AFFA63F46FEC1FAFCFE233333" box="[282,472,1353,1377]" italics="true" pageId="2" pageNumber="367">Creagrutus andaki</emphasis>
</taxonomicName>
, paratypes: (A) CIACOL 2910, 58.7 mm LS. (B) CZUT-IC 19973, 57.4 mm LS. (C) CIACOL 3463, 38.6 mm LS. All from Colombia, Caquetá, Florencia, Río Caquetá basin, Río Hacha drainage, Río Sucre.
</paragraph>
</caption>
<paragraph id="0B1AAC58FFA63F46FF1CFA01FED4300C" blockId="2.[151,1437,1459,2026]" pageId="2" pageNumber="367">
<materialsCitation id="BBCDA605FFA63F46FF1CFA01FF1E33A3" ID-GBIF-Occurrence="2592573801" collectingDate="2018-08-10" collectingDateMax="2017-08-12" collectingDateMin="2016-08-12" collectionCode="CIACOL" collectorName="J. D. Bogota-Gregory &amp; I. Gonzalez-Gomez" country="Colombia" county="Florencia" elevation="1015" latitude="1.7953333" location="Rio Hacha" longLatPrecision="1" longitude="-75.64691" municipality="Rio" pageId="2" pageNumber="367" specimenCode="CIACOL 2130, CZUT-IC 19973, IAvH-P 20565, CIACOL 2910" specimenCount="1" stateProvince="Caqueta" typeStatus="paratype">
<emphasis id="39D1704AFFA63F46FF1CFA01FEBD339C" bold="true" box="[199,326,1460,1486]" pageId="2" pageNumber="367">
<typeStatus id="D41E12FAFFA63F46FF1CFA01FEBA339C" box="[199,321,1460,1486]" pageId="2" pageNumber="367" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectorName id="A650C98EFFA63F46FE95FA06FE89339F" box="[334,370,1459,1485]" pageId="2" pageNumber="367">All</collectorName>
from the same locality of the
<typeStatus id="D41E12FAFFA63F46FD12FA06FCD4339F" box="[713,815,1459,1485]" pageId="2" pageNumber="367" type="holotype">holotype</typeStatus>
: CIACOL 3463 (4,
<quantity id="CC5D01BDFFA63F46FBCBFA06FB64339C" box="[1040,1183,1459,1486]" metricMagnitude="-2" metricUnit="m" metricValue="4.835" metricValueMax="5.8" metricValueMin="3.87" pageId="2" pageNumber="367" unit="mm" value="48.35" valueMax="58.0" valueMin="38.7">38.7–58 mm</quantity>
SL), same data as
<typeStatus id="D41E12FAFFA63F46FAACFA06FF1A33A3" pageId="2" pageNumber="367" type="holotype">holotype</typeStatus>
;
</materialsCitation>
<materialsCitation id="BBCDA605FFA63F46FF36FA6DFD0633A0" ID-GBIF-Occurrence="2592573805" box="[237,765,1495,1522]" collectionCode="C, S, SL" country="Colombia" county="Florencia" elevation="1015" latitude="1.7953333" location="Rio Hacha" longLatPrecision="1" longitude="-75.64691" municipality="Rio" pageId="2" pageNumber="367" specimenCode="CIACOL 2130" specimenCount="1" stateProvince="Caqueta" typeStatus="paratype">
<specimenCode id="5B030423FFA63F46FF36FA6DFE6F33A0" box="[237,404,1496,1522]" collectionCode="CIACOL" pageId="2" pageNumber="367">CIACOL 2130</specimenCode>
(7, 1
<collectionCode id="6DB4349DFFA63F46FE03FA6DFE1733A0" box="[472,492,1496,1522]" pageId="2" pageNumber="367">C</collectionCode>
&amp;
<collectionCode id="6DB4349DFFA63F46FDDFFA6DFDE833A0" box="[516,531,1496,1522]" pageId="2" pageNumber="367">S</collectionCode>
,
<quantity id="CC5D01BDFFA63F46FDC4FA62FD3833A0" box="[543,707,1495,1522]" metricMagnitude="-2" metricUnit="m" metricValue="5.02" metricValueMax="6.459999999999999" metricValueMin="3.5799999999999996" pageId="2" pageNumber="367" unit="mm" value="50.199999999999996" valueMax="64.6" valueMin="35.8">35.8–64.6 mm</quantity>
<collectionCode id="6DB4349DFFA63F46FD10FA6DFD0B33A0" box="[715,752,1496,1522]" pageId="2" pageNumber="367">SL</collectionCode>
);
</materialsCitation>
<materialsCitation id="BBCDA605FFA63F46FCDFFA6DFECB306B" ID-GBIF-Occurrence="2592573803" collectionCode="SL, C, S" country="Colombia" county="Florencia" elevation="1015" latitude="1.7953333" location="Rio Hacha" longLatPrecision="1" longitude="-75.64691" municipality="Rio" pageId="2" pageNumber="367" specimenCode="CZUT-IC 19973, IAvH-P 20565" specimenCount="1" stateProvince="Caqueta" typeStatus="paratype">
<specimenCode id="5B030423FFA63F46FCDFFA6DFC4533A0" box="[772,958,1496,1522]" collectionCode="CZUT-IC" pageId="2" pageNumber="367">CZUT-IC 19973</specimenCode>
(2,
<quantity id="CC5D01BDFFA63F46FC37FA62FB8033A0" box="[1004,1147,1495,1522]" metricMagnitude="-2" metricUnit="m" metricValue="4.645" metricValueMax="5.7" metricValueMin="3.59" pageId="2" pageNumber="367" unit="mm" value="46.45" valueMax="57.0" valueMin="35.9">35.9–57 mm</quantity>
<collectionCode id="6DB4349DFFA63F46FB58FA6DFB5333A0" box="[1155,1192,1496,1522]" pageId="2" pageNumber="367">SL</collectionCode>
);
<specimenCode id="5B030423FFA63F46FB66FA6DFA9B33A0" box="[1213,1376,1495,1522]" collectionCode="IAvH-P" pageId="2" pageNumber="367">IAvH-P 20565</specimenCode>
(2, 1
<collectionCode id="6DB4349DFFA63F46FF4CFA49FF503044" box="[151,171,1532,1558]" pageId="2" pageNumber="367">C</collectionCode>
&amp;
<collectionCode id="6DB4349DFFA63F46FF18FA49FF293044" box="[195,210,1532,1558]" pageId="2" pageNumber="367">S</collectionCode>
,
<quantity id="CC5D01BDFFA63F46FF05FA4EFE7A3044" box="[222,385,1531,1558]" metricMagnitude="-2" metricUnit="m" metricValue="5.265000000000001" metricValueMax="5.470000000000001" metricValueMin="5.0600000000000005" pageId="2" pageNumber="367" unit="mm" value="52.65" valueMax="54.7" valueMin="50.6">50.6–54.7 mm</quantity>
<collectionCode id="6DB4349DFFA63F46FE53FA49FE573044" box="[392,428,1532,1558]" pageId="2" pageNumber="367">SL</collectionCode>
),
<collectorName id="A650C98EFFA63F46FE1BFA49FD503047" box="[448,683,1531,1558]" pageId="2" pageNumber="367">E. Agudelo-Córdoba</collectorName>
,
<collectorName id="A650C98EFFA63F46FD6CFA49FC3D3044" box="[695,966,1532,1558]" pageId="2" pageNumber="367">J D. Bogotá-Gregory, A</collectorName>
. Acosta-Santos &amp;
<collectorName id="A650C98EFFA63F46FB4DFA49FA8E3047" box="[1174,1397,1531,1558]" pageId="2" pageNumber="367">I. González-Gómez</collectorName>
,
<date id="7F1B8A98FFA63F46FA5BFA49FED6306B" pageId="2" pageNumber="367" value="2016-08-12">
<collectingDate id="6F5F7370FFA63F46FA5BFA49FED6306B" pageId="2" pageNumber="367" value="2016-08-12">12 August 2016</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="BBCDA605FFA63F46FEE2F995FED0300C" ID-GBIF-Occurrence="2592573802" collectorName="E. Agudelo-Cordoba &amp; J D. Bogota-Gregory, A" country="Colombia" county="Florencia" elevation="1015" latitude="1.7953333" location="Rio Hacha" longLatPrecision="1" longitude="-75.64691" municipality="Rio" pageId="2" pageNumber="367" specimenCode="CIACOL 2910" specimenCount="1" stateProvince="Caqueta" typeStatus="paratype">
<specimenCode id="5B030423FFA63F46FEE2F995FE1A3068" box="[313,481,1568,1594]" collectionCode="CIACOL" pageId="2" pageNumber="367">
<collectionCode id="6DB4349DFFA63F46FEE2F995FE5B3068" box="[313,416,1568,1594]" pageId="2" pageNumber="367">CIACOL</collectionCode>
2910
</specimenCode>
(1,
<quantity id="CC5D01BDFFA63F46FDCBF9AAFD8D3068" box="[528,630,1567,1594]" metricMagnitude="-2" metricUnit="m" metricValue="5.87" pageId="2" pageNumber="367" unit="mm" value="58.7">58.7 mm</quantity>
<collectionCode id="6DB4349DFFA63F46FDA5F995FD583068" box="[638,675,1568,1594]" pageId="2" pageNumber="367">SL</collectionCode>
);
<collectorName id="A650C98EFFA63F46FD63F995FC5E306B" box="[696,933,1567,1594]" pageId="2" pageNumber="367">E. Agudelo-Córdoba</collectorName>
,
<collectorName id="A650C98EFFA63F46FC6AF995FB3F3068" box="[945,1220,1568,1594]" pageId="2" pageNumber="367">J D. Bogotá-Gregory, A</collectorName>
. Acosta-Santos,
<date id="7F1B8A98FFA63F46FA5BF995FED0300C" pageId="2" pageNumber="367" value="2017-08-12">
<collectingDate id="6F5F7370FFA63F46FA5BF995FED0300C" pageId="2" pageNumber="367" value="2017-08-12">12 August 2017</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="43BFFFD3FFA63F46FF1CF9D2FE8531B8" pageId="2" pageNumber="367" type="diagnosis">
<paragraph id="0B1AAC58FFA63F46FF1CF9D2FE8531B8" blockId="2.[151,1437,1459,2026]" pageId="2" pageNumber="367">
<emphasis id="39D1704AFFA63F46FF1CF9D2FEC130D0" bold="true" box="[199,314,1639,1666]" pageId="2" pageNumber="367">Diagnosis</emphasis>
.
<taxonomicName id="CCA5D7DBFFA63F46FE93F9DDFDE130D3" authorityName="Albornoz-Garzón &amp; Acosta-Santos &amp; Bogotá-Gregory &amp; Agudelo-Córdoba" authorityYear="2020" box="[328,538,1639,1666]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="367" phylum="Chordata" rank="species" species="andaki">
<emphasis id="39D1704AFFA63F46FE93F9DDFDE130D3" box="[328,538,1639,1666]" italics="true" pageId="2" pageNumber="367">Creagrutus andaki</emphasis>
</taxonomicName>
differs from all its congeners, except for
<emphasis id="39D1704AFFA63F46FC32F9DDFAF830D3" box="[1001,1283,1639,1666]" italics="true" pageId="2" pageNumber="367">
<taxonomicName id="CCA5D7DBFFA63F46FC32F9DDFB9F30D3" box="[1001,1124,1640,1666]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="367" phylum="Chordata" rank="species" species="cacique">C. cacique</taxonomicName>
,
<taxonomicName id="CCA5D7DBFFA63F46FBB4F9DDFAF830D3" authorityName="Vari &amp; Harold" authorityYear="2001" box="[1135,1283,1639,1666]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="367" phylum="Chordata" rank="species" species="gyrospilus">C. gyrospilus</taxonomicName>
</emphasis>
,
<taxonomicName id="CCA5D7DBFFA63F46FACAF9DDFA6D30D3" authorityName="Vari, Harold &amp; Taphorn" authorityYear="1994" box="[1297,1430,1639,1666]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="367" phylum="Chordata" rank="species" species="melasma">
<emphasis id="39D1704AFFA63F46FACAF9DDFA6D30D3" box="[1297,1430,1639,1666]" italics="true" pageId="2" pageNumber="367">C. melasma</emphasis>
</taxonomicName>
,
<taxonomicName id="CCA5D7DBFFA63F46FF4CF939FEA030F7" authorityName="Dagosta &amp; Pastana" authorityYear="2014" box="[151,347,1676,1702]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="367" phylum="Chordata" rank="species" species="nigrotaeniatus">
<emphasis id="39D1704AFFA63F46FF4CF939FEA030F7" box="[151,347,1676,1702]" italics="true" pageId="2" pageNumber="367">C. nigrotaeniatus</emphasis>
</taxonomicName>
,
<taxonomicName id="CCA5D7DBFFA63F46FEB1F939FDD930F7" authorityName="Vari &amp; Harold" authorityYear="2001" box="[362,546,1676,1702]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="367" phylum="Chordata" rank="species" species="ouranonastes">
<emphasis id="39D1704AFFA63F46FEB1F939FDD930F7" box="[362,546,1676,1702]" italics="true" pageId="2" pageNumber="367">C. ouranonastes</emphasis>
</taxonomicName>
,
<taxonomicName id="CCA5D7DBFFA63F46FDEAF939FD2B30F7" box="[561,720,1675,1702]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="367" phylum="Chordata" rank="species" species="planquettei">
<emphasis id="39D1704AFFA63F46FDEAF939FD2B30F7" box="[561,720,1675,1702]" italics="true" pageId="2" pageNumber="367">C. planquettei</emphasis>
</taxonomicName>
,
<taxonomicName id="CCA5D7DBFFA63F46FD04F939FCB030F7" authorityName="Vari &amp; Lima" authorityYear="2003" box="[735,843,1675,1702]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="367" phylum="Chordata" rank="species" species="tuyuka">
<emphasis id="39D1704AFFA63F46FD04F939FCB030F7" box="[735,843,1675,1702]" italics="true" pageId="2" pageNumber="367">C. tuyuka</emphasis>
</taxonomicName>
,
<taxonomicName id="CCA5D7DBFFA63F46FC82F939FC4B30F7" authorityName="Ribeiro, Benine &amp; Figueiredo" authorityYear="2004" box="[857,944,1676,1702]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="367" phylum="Chordata" rank="species" species="varii">
<emphasis id="39D1704AFFA63F46FC82F939FC4B30F7" box="[857,944,1676,1702]" italics="true" pageId="2" pageNumber="367">C. varii</emphasis>
</taxonomicName>
and
<taxonomicName id="CCA5D7DBFFA63F46FC33F939FBA430F7" authorityName="Harold &amp; Salcedo" authorityYear="2010" box="[1000,1119,1675,1702]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="367" phylum="Chordata" rank="species" species="yanatili">
<emphasis id="39D1704AFFA63F46FC33F939FBA430F7" box="[1000,1119,1675,1702]" italics="true" pageId="2" pageNumber="367">C. yanatili</emphasis>
</taxonomicName>
by having a dark, relatively broad midlateral stripe starting at anteriormost scale of lateral line (vs. midlateral stripe absent or starting posterior to humeral blotch). The new species is differentiated from
<taxonomicName id="CCA5D7DBFFA63F46FCF6F961FC3930BF" authorityName="Vari &amp; Harold" authorityYear="2001" box="[813,962,1747,1774]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="367" phylum="Chordata" rank="species" species="gyrospilus">
<emphasis id="39D1704AFFA63F46FCF6F961FC3930BF" box="[813,962,1747,1774]" italics="true" pageId="2" pageNumber="367">C. gyrospilus</emphasis>
</taxonomicName>
,
<taxonomicName id="CCA5D7DBFFA63F46FC0BF961FB8B30BF" box="[976,1136,1747,1774]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="367" phylum="Chordata" rank="species" species="planquettei">
<emphasis id="39D1704AFFA63F46FC0BF961FB8B30BF" box="[976,1136,1747,1774]" italics="true" pageId="2" pageNumber="367">C. planquettei</emphasis>
</taxonomicName>
, and
<taxonomicName id="CCA5D7DBFFA63F46FB75F961FADE30BF" authorityName="Harold &amp; Salcedo" authorityYear="2010" box="[1198,1317,1747,1774]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="367" phylum="Chordata" rank="species" species="yanatili">
<emphasis id="39D1704AFFA63F46FB75F961FADE30BF" box="[1198,1317,1747,1774]" italics="true" pageId="2" pageNumber="367">C. yanatili</emphasis>
</taxonomicName>
by having humeral blotch vertically elongated (vs. humeral blotch absent or oval shaped).
<taxonomicName id="CCA5D7DBFFA63F46FBC0F94DFB163143" authorityName="Albornoz-Garzón &amp; Acosta-Santos &amp; Bogotá-Gregory &amp; Agudelo-Córdoba" authorityYear="2020" box="[1051,1261,1783,1810]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="367" phylum="Chordata" rank="species" species="andaki">
<emphasis id="39D1704AFFA63F46FBC0F94DFB163143" box="[1051,1261,1783,1810]" italics="true" pageId="2" pageNumber="367">Creagrutus andaki</emphasis>
</taxonomicName>
differs from
<taxonomicName id="CCA5D7DBFFA63F46FA58F94DFF153167" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="367" phylum="Chordata" rank="species" species="cacique">
<emphasis id="39D1704AFFA63F46FA58F94DFF153167" italics="true" pageId="2" pageNumber="367">C. cacique</emphasis>
</taxonomicName>
,
<taxonomicName id="CCA5D7DBFFA63F46FF27F8A9FE7A3167" authorityName="Vari, Harold &amp; Taphorn" authorityYear="1994" box="[252,385,1819,1846]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="367" phylum="Chordata" rank="species" species="melasma">
<emphasis id="39D1704AFFA63F46FF27F8A9FE7A3167" box="[252,385,1819,1846]" italics="true" pageId="2" pageNumber="367">C. melasma</emphasis>
</taxonomicName>
and
<taxonomicName id="CCA5D7DBFFA63F46FE63F8A9FDDF3167" authorityName="Vari &amp; Lima" authorityYear="2003" box="[440,548,1819,1846]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="367" phylum="Chordata" rank="species" species="tuyuka">
<emphasis id="39D1704AFFA63F46FE63F8A9FDDF3167" box="[440,548,1819,1846]" italics="true" pageId="2" pageNumber="367">C. tuyuka</emphasis>
</taxonomicName>
by lacking a dark blotch on dorsal fin (vs. dorsal fin with a dark blotch).
<taxonomicName id="CCA5D7DBFFA63F46FA87F8A9FEC8310B" authorityName="Albornoz-Garzón &amp; Acosta-Santos &amp; Bogotá-Gregory &amp; Agudelo-Córdoba" authorityYear="2020" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="367" phylum="Chordata" rank="species" species="andaki">
<emphasis id="39D1704AFFA63F46FA87F8A9FEC8310B" italics="true" pageId="2" pageNumber="367">Creagrutus andaki</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="CCA5D7DBFFA63F46FDB1F8F5FCD8310B" authorityName="Vari &amp; Harold" authorityYear="2001" box="[618,803,1856,1882]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="367" phylum="Chordata" rank="species" species="ouranonastes">
<emphasis id="39D1704AFFA63F46FDB1F8F5FCD8310B" box="[618,803,1856,1882]" italics="true" pageId="2" pageNumber="367">C. ouranonastes</emphasis>
</taxonomicName>
and
<taxonomicName id="CCA5D7DBFFA63F46FC80F8F5FC49310B" authorityName="Ribeiro, Benine &amp; Figueiredo" authorityYear="2004" box="[859,946,1856,1882]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="367" phylum="Chordata" rank="species" species="varii">
<emphasis id="39D1704AFFA63F46FC80F8F5FC49310B" box="[859,946,1856,1882]" italics="true" pageId="2" pageNumber="367">C. varii</emphasis>
</taxonomicName>
by the absence of a dark blotch on base of middle rays of caudal fin (vs. presence of a dark blotch on base of middle rays of caudal fin).
<taxonomicName id="CCA5D7DBFFA63F46FB6AF8D1FA79312F" authorityName="Albornoz-Garzón &amp; Acosta-Santos &amp; Bogotá-Gregory &amp; Agudelo-Córdoba" authorityYear="2020" box="[1201,1410,1891,1918]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="367" phylum="Chordata" rank="species" species="andaki">
<emphasis id="39D1704AFFA63F46FB6AF8D1FA79312F" box="[1201,1410,1891,1918]" italics="true" pageId="2" pageNumber="367">Creagrutus andaki</emphasis>
</taxonomicName>
is differentiated from
<taxonomicName id="CCA5D7DBFFA63F46FEB4F83DFDCA31F3" authorityName="Dagosta &amp; Pastana" authorityYear="2014" box="[367,561,1928,1954]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="367" phylum="Chordata" rank="species" species="nigrotaeniatus">
<emphasis id="39D1704AFFA63F46FEB4F83DFDCA31F3" box="[367,561,1928,1954]" italics="true" pageId="2" pageNumber="367">C. nigrotaeniatus</emphasis>
</taxonomicName>
by having a larger number of predorsal scales (11–12 vs. 8–9), by the morphology of the dentigerous surface in ventral view (triangular vs. longitudinally elongate), and by the number of dentary teeth (5–6 vs. 8–10).
</paragraph>
</subSubSection>
<caption id="5FDAFCD0FFA73F47FF4CFAB4FAD73348" ID-DOI="http://doi.org/10.5281/zenodo.3733691" ID-Zenodo-Dep="3733691" box="[151,1324,1281,1306]" httpUri="https://zenodo.org/record/3733691/files/figure.png" pageId="3" pageNumber="368" startId="3.[151,250,1281,1306]" targetBox="[333,1255,181,1255]" targetPageId="3">
<paragraph id="0B1AAC58FFA73F47FF4CFAB4FAD73348" blockId="3.[151,1324,1281,1306]" box="[151,1324,1281,1306]" pageId="3" pageNumber="368">
<emphasis id="39D1704AFFA73F47FF4CFAB4FEE83348" bold="true" box="[151,275,1281,1306]" pageId="3" pageNumber="368">FIGURE 3.</emphasis>
<taxonomicName id="CCA5D7DBFFA73F47FEC2FAB7FE2C3348" authorityName="Albornoz-Garzón &amp; Acosta-Santos &amp; Bogotá-Gregory &amp; Agudelo-Córdoba" authorityYear="2020" box="[281,471,1282,1306]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="3" pageNumber="368" phylum="Chordata" rank="species" species="andaki">
<emphasis id="39D1704AFFA73F47FEC2FAB7FE2C3348" box="[281,471,1282,1306]" italics="true" pageId="3" pageNumber="368">Creagrutus andaki</emphasis>
</taxonomicName>
, IAvH-P 20565, paratype, 50.6 mm SL, premaxillary teeth in ventral view, left side.
</paragraph>
</caption>
<subSubSection id="43BFFFD3FFA73F41FF1CFAF2FC7F3711" lastPageId="5" lastPageNumber="370" pageId="3" pageNumber="368" type="description">
<paragraph id="0B1AAC58FFA73F47FF1CFAF2FDBE30BC" blockId="3.[151,1437,1351,2027]" pageId="3" pageNumber="368">
<emphasis id="39D1704AFFA73F47FF1CFAF2FEA23330" bold="true" box="[199,345,1351,1378]" pageId="3" pageNumber="368">Description.</emphasis>
Morphometric data are given in Table 1. Body moderately elongate. Greatest body depth between half distance between pectoral- and pelvic-fin origins. Head slightly pointed, its dorsal profile convex from the margin of the upper lip to vertical through posterior margin of posterior nostrils, then straight to slightly convex from latter point to end of supraoccipital spine; interorbital region transversely convex. Dorsal profile slightly convex from tip of supraoccipital spine to dorsal-fin origin; nearly straight and posteroventrally slanted from latter point to adipose fin, and concave from later point to anteriormost procurrent caudal-fin rays. Ventral profile of head slightly convex to antero-ventral corner of dentary, then straight to isthmus. Ventral profile of body slightly convex from isthmus to anal-fin origin; anal-fin base straight; straight to slightly concave from the base of last anal-fin ray to procurrent caudal-fin rays. Some specimens with tubercles with keratinized caps distributed on dorsal region of head. Infraorbital series moderately developed. Central portion of ventral margin of third infraorbital horizontally straight, in contact with horizontal limb of preopercle. Posterior margins of third through fifth infraorbitals distinctly separated from vertical limb of preopercle.
</paragraph>
<paragraph id="0B1AAC58FFA73F40FF1CF94DFCE13685" blockId="3.[151,1437,1351,2027]" lastBlockId="4.[151,1437,153,755]" lastPageId="4" lastPageNumber="369" pageId="3" pageNumber="368">
Mouth subterminal, upper jaw conspicuously protruding beyond lower jaw; outer row of premaxillary teeth slightly exposed in ventral view. Corner of mouth located at horizontal line that crosses below ventral margin of eye. Premaxilla with three series of tricuspid teeth: first row irregular, with 5(1) or 6(1) teeth of equal size forming arc without distinct gap between first and second teeth, 5 from 17 examined specimens presenting gap (
<figureCitation id="939EB0DDFFA73F47FA91F8D1FA75312C" box="[1354,1422,1892,1918]" captionStart="FIGURE 3" captionStartId="3.[151,250,1281,1306]" captionTargetBox="[333,1255,181,1255]" captionTargetId="figure@3.[333,1255,181,1255]" captionTargetPageId="3" captionText="FIGURE 3. Creagrutus andaki, IAvH-P 20565, paratype, 50.6 mm SL, premaxillary teeth in ventral view, left side." figureDoi="http://doi.org/10.5281/zenodo.3733691" httpUri="https://zenodo.org/record/3733691/files/figure.png" pageId="3" pageNumber="368">Fig. 3</figureCitation>
). Second row in cluster of 3(2) teeth, larger than teeth from first row, third row with 1(1) or 2(1) teeth, slightly thinner than remaining teeth, located next to third and fourth teeth from first row. Maxilla with 2(1) or 3(1) tricuspid teeth, of equal size. Dentary with 5(1) or 6(1) tricuspid teeth, anteriormost three teeth larger than remaining teeth. First branchial arch with 1(1) or 2(1) gill rakers on hypobranchial, 7(1) or 8(1) gill rakers on ceratobranchial, and 6(1) or 7(1) gill rakers on epibranchial. Branchiostegal rays 4(2).
</paragraph>
<paragraph id="0B1AAC58FFA03F40FF1CFF54FD6E37D9" blockId="4.[151,1437,153,755]" pageId="4" pageNumber="369">
Scales cycloid,
<emphasis id="39D1704AFFA03F40FEAFFF54FE4736A9" box="[372,444,225,251]" italics="true" pageId="4" pageNumber="369">circuli</emphasis>
restricted to anterior part of scales, with 10–16 wavy
<emphasis id="39D1704AFFA03F40FBD0FF54FBBA36A9" box="[1035,1089,225,251]" italics="true" pageId="4" pageNumber="369">radii</emphasis>
, that extend parallel from scale center to its posterior margin. Lateral line slightly curved ventrally, with 40*(9) or 41(6) pored scales. Four(2) or 5*(12) scales between dorsal-fin origin and lateral line; 3*(15) scales between lateral line and pelvic-fin insertion. Predorsal scales 11*(14) or 12(1). Postanal scales 2*(7), 3(7) or 4(1). Caudal fin with scales on first or second third of each lobe, similar in size to scales of body.
</paragraph>
<paragraph id="0B1AAC58FFA03F40FF1CFE20FBA434A1" blockId="4.[151,1437,153,755]" pageId="4" pageNumber="369">Dorsal-fin rays ii,8*(15), dorsal-fin origin anterior to pelvic-fin insertion, distal dorsal-fin margin slightly concave, with third and last two rays longer than remaining rays, fleshy interradial folds present. Anteriormost dorsal-fin pterygiophore inserted posterior to neural spine of 10th(2) vertebra. Anal-fin rays iii(2), 9(3) or 10*(12), distal profile concave, last unbranched and first two branched rays slightly longer. Fleshy interradial folds present. Anteriormost anal-fin pterygiophore inserted posterior to neural spine of 19th (2) vertebra. Pectoral-fin rays i,9(3) or 10*(12), its posterior tip not reaching vertical through pelvic-fin insertion. Fleshy interradial folds present. Pelvic-fin rays i,7*(15), distal tip not reaching anal-fin origin. Fleshy interradial folds present. Adipose-fin origin at vertical posterior through last anal-fin ray insertion. Caudal fin forked, lobes approximately similar in size. Principal caudal-fin rays i,10+9,i(2). Dorsal caudal fin procurrent rays 13(1) or 14(1), ventral caudal-fin procurrent rays 12(2). One pair of uroneurals. Supraneurals 5(2). Ribs 12(1) or 13(1). Vertebrae 38(2).</paragraph>
<caption id="5FDAFCD0FFA03F40FF4CFC97FA983569" ID-Table-UUID="5FDAFCD0FFA03F40FF4CFC97FA983569" box="[151,1379,802,827]" httpUri="http://table.plazi.org/id/5FDAFCD0FFA03F40FF4CFC97FA983569" pageId="4" pageNumber="369" startId="4.[151,239,802,826]" targetBox="[163,1363,842,1791]" targetIsTable="true" targetPageId="4">
<paragraph id="0B1AAC58FFA03F40FF4CFC97FA983569" blockId="4.[151,1379,802,827]" box="[151,1379,802,827]" pageId="4" pageNumber="369">
<tableNote id="F643ADD6FFA03F40FF4CFC97FA983569" box="[151,1379,802,827]" pageId="4" pageNumber="369" targetBox="[163,1363,842,1791]" targetPageId="4">
<emphasis id="39D1704AFFA03F40FF4CFC97FEF33569" bold="true" box="[151,264,802,827]" pageId="4" pageNumber="369">TABLE I.</emphasis>
Morphometric data for holotype and paratypes (n=14) of
<emphasis id="39D1704AFFA03F40FCA9FC96FBBA3569" box="[882,1089,803,827]" italics="true" pageId="4" pageNumber="369">
<taxonomicName id="CCA5D7DBFFA03F40FCA9FC96FBC63569" authorityName="Albornoz-Garzón &amp; Acosta-Santos &amp; Bogotá-Gregory &amp; Agudelo-Córdoba" authorityYear="2020" box="[882,1085,803,827]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="369" phylum="Chordata" rank="species" species="andaki">Creagrutus andaki</taxonomicName>
.
</emphasis>
S.D. = Standard deviation.
</tableNote>
</paragraph>
</caption>
<paragraph id="0B1AAC58FFA03F40FD2BFCFFFAB730AD" pageId="4" pageNumber="369">
<table id="79A55EF8FFA0C0BBFF78FCFFFAA830AD" box="[163,1363,842,1791]" gridcols="5" gridrows="23" pageId="4" pageNumber="369">
<tr id="B595AE1AFFA0C0BBFF78FCFFFAA83531" box="[163,1363,842,867]" gridrow="0" pageId="4" pageNumber="369" rowspan-0="1" rowspan-2="1">
<th id="F644C766FFA0C0BBFD2BFCFFFCB53531" box="[752,846,842,867]" gridcol="1" gridrow="0" pageId="4" pageNumber="369">Holotype</th>
<th id="F644C766FFA0C0BBFB8DFCFFFAA83531" box="[1110,1363,842,867]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="4" pageNumber="369">Paratypes</th>
</tr>
<tr id="B595AE1AFFA0C0BBFF78FCC0FAA835DF" box="[163,1363,885,909]" gridrow="1" pageId="4" pageNumber="369" rowspan-0="1" rowspan-1="1">
<td id="F644C766FFA0C0BBFC67FCC0FC0D35DF" box="[956,1014,885,909]" gridcol="2" gridrow="1" pageId="4" pageNumber="369">Mean</td>
<td id="F644C766FFA0C0BBFB8DFCC0FB4735DF" box="[1110,1212,885,909]" gridcol="3" gridrow="1" pageId="4" pageNumber="369">Range</td>
<td id="F644C766FFA0C0BBFAFDFCC0FAA835DF" box="[1318,1363,885,909]" gridcol="4" gridrow="1" pageId="4" pageNumber="369">S.D.</td>
</tr>
<tr id="B595AE1AFFA0C0BBFF78FC2BFAA835E5" box="[163,1363,926,951]" gridrow="2" pageId="4" pageNumber="369">
<th id="F644C766FFA0C0BBFF78FC2BFDDC35E5" box="[163,551,926,951]" gridcol="0" gridrow="2" pageId="4" pageNumber="369">Standard length (mm)</th>
<td id="F644C766FFA0C0BBFD2BFC2BFCB535E5" box="[752,846,926,951]" gridcol="1" gridrow="2" pageId="4" pageNumber="369">56.8</td>
<td id="F644C766FFA0C0BBFC67FC2BFC0D35E5" box="[956,1014,926,951]" gridcol="2" gridrow="2" pageId="4" pageNumber="369">-</td>
<td id="F644C766FFA0C0BBFB8DFC2BFB4735E5" box="[1110,1212,926,951]" gridcol="3" gridrow="2" pageId="4" pageNumber="369">35.9–65.2</td>
<td id="F644C766FFA0C0BBFAFDFC2BFAA835E5" box="[1318,1363,926,951]" gridcol="4" gridrow="2" pageId="4" pageNumber="369">-</td>
</tr>
<tr id="B595AE1AFFA0C0BBFF78FC7DFAA835B3" box="[163,1363,968,993]" gridrow="3" pageId="4" pageNumber="369">
<th id="F644C766FFA0C0BBFF78FC7DFAA835B3" box="[163,1363,968,993]" colspan="5" colspanRight="4" gridcol="0" gridrow="3" pageId="4" pageNumber="369">
<emphasis id="39D1704AFFA03F40FF78FC7CFE9F35B3" bold="true" box="[163,356,968,993]" pageId="4" pageNumber="369">Percentages of SL</emphasis>
</th>
</tr>
<tr id="B595AE1AFFA0C0BBFF78FC47FAA83259" box="[163,1363,1010,1035]" gridrow="4" pageId="4" pageNumber="369">
<th id="F644C766FFA0C0BBFF78FC47FDDC3259" box="[163,551,1010,1035]" gridcol="0" gridrow="4" pageId="4" pageNumber="369">Snout to dorsal-fin origin</th>
<td id="F644C766FFA0C0BBFD2BFC47FCB53259" box="[752,846,1010,1035]" gridcol="1" gridrow="4" pageId="4" pageNumber="369">46.8</td>
<td id="F644C766FFA0C0BBFC67FC47FC0D3259" box="[956,1014,1010,1035]" gridcol="2" gridrow="4" pageId="4" pageNumber="369">46.5</td>
<td id="F644C766FFA0C0BBFB8DFC47FB473259" box="[1110,1212,1010,1035]" gridcol="3" gridrow="4" pageId="4" pageNumber="369">43.3–48.9</td>
<td id="F644C766FFA0C0BBFAFDFC47FAA83259" box="[1318,1363,1010,1035]" gridcol="4" gridrow="4" pageId="4" pageNumber="369">1.3</td>
</tr>
<tr id="B595AE1AFFA0C0BBFF78FBA9FAA83267" box="[163,1363,1052,1077]" gridrow="5" pageId="4" pageNumber="369">
<th id="F644C766FFA0C0BBFF78FBA9FDDC3267" box="[163,551,1052,1077]" gridcol="0" gridrow="5" pageId="4" pageNumber="369">Snout to pectoral-fin origin</th>
<td id="F644C766FFA0C0BBFD2BFBA9FCB53267" box="[752,846,1052,1077]" gridcol="1" gridrow="5" pageId="4" pageNumber="369">24.2</td>
<td id="F644C766FFA0C0BBFC67FBA9FC0D3267" box="[956,1014,1052,1077]" gridcol="2" gridrow="5" pageId="4" pageNumber="369">24.0</td>
<td id="F644C766FFA0C0BBFB8DFBA9FB473267" box="[1110,1212,1052,1077]" gridcol="3" gridrow="5" pageId="4" pageNumber="369">21.8–27.0</td>
<td id="F644C766FFA0C0BBFAFDFBA9FAA83267" box="[1318,1363,1052,1077]" gridcol="4" gridrow="5" pageId="4" pageNumber="369">1.4</td>
</tr>
<tr id="B595AE1AFFA0C0BBFF78FBF3FAA8320D" box="[163,1363,1094,1119]" gridrow="6" pageId="4" pageNumber="369">
<th id="F644C766FFA0C0BBFF78FBF3FDDC320D" box="[163,551,1094,1119]" gridcol="0" gridrow="6" pageId="4" pageNumber="369">Snout to pelvic-fin origin</th>
<td id="F644C766FFA0C0BBFD2BFBF3FCB5320D" box="[752,846,1094,1119]" gridcol="1" gridrow="6" pageId="4" pageNumber="369">44.6</td>
<td id="F644C766FFA0C0BBFC67FBF3FC0D320D" box="[956,1014,1094,1119]" gridcol="2" gridrow="6" pageId="4" pageNumber="369">46.2</td>
<td id="F644C766FFA0C0BBFB8DFBF3FB47320D" box="[1110,1212,1094,1119]" gridcol="3" gridrow="6" pageId="4" pageNumber="369">42.8–48.5</td>
<td id="F644C766FFA0C0BBFAFDFBF3FAA8320D" box="[1318,1363,1094,1119]" gridcol="4" gridrow="6" pageId="4" pageNumber="369">1.6</td>
</tr>
<tr id="B595AE1AFFA0C0BBFF78FBC5FAA832DB" box="[163,1363,1136,1161]" gridrow="7" pageId="4" pageNumber="369">
<th id="F644C766FFA0C0BBFF78FBC5FDDC32DB" box="[163,551,1136,1161]" gridcol="0" gridrow="7" pageId="4" pageNumber="369">Snout to anal-fin origin</th>
<td id="F644C766FFA0C0BBFD2BFBC5FCB532DB" box="[752,846,1136,1161]" gridcol="1" gridrow="7" pageId="4" pageNumber="369">60.5</td>
<td id="F644C766FFA0C0BBFC67FBC5FC0D32DB" box="[956,1014,1136,1161]" gridcol="2" gridrow="7" pageId="4" pageNumber="369">61.7</td>
<td id="F644C766FFA0C0BBFB8DFBC5FB4732DB" box="[1110,1212,1136,1161]" gridcol="3" gridrow="7" pageId="4" pageNumber="369">57.8–65.2</td>
<td id="F644C766FFA0C0BBFAFDFBC5FAA832DB" box="[1318,1363,1136,1161]" gridcol="4" gridrow="7" pageId="4" pageNumber="369">1.9</td>
</tr>
<tr id="B595AE1AFFA0C0BBFF78FB2FFAA832E1" box="[163,1363,1178,1203]" gridrow="8" pageId="4" pageNumber="369">
<th id="F644C766FFA0C0BBFF78FB2FFDDC32E1" box="[163,551,1178,1203]" gridcol="0" gridrow="8" pageId="4" pageNumber="369">Dorsal-fin origin to caudal-fin base</th>
<td id="F644C766FFA0C0BBFD2BFB2FFCB532E1" box="[752,846,1178,1203]" gridcol="1" gridrow="8" pageId="4" pageNumber="369">57.7</td>
<td id="F644C766FFA0C0BBFC67FB2FFC0D32E1" box="[956,1014,1178,1203]" gridcol="2" gridrow="8" pageId="4" pageNumber="369">56.5</td>
<td id="F644C766FFA0C0BBFB8DFB2FFB4732E1" box="[1110,1212,1178,1203]" gridcol="3" gridrow="8" pageId="4" pageNumber="369">52.9–58.5</td>
<td id="F644C766FFA0C0BBFAFDFB2FFAA832E1" box="[1318,1363,1178,1203]" gridcol="4" gridrow="8" pageId="4" pageNumber="369">1.5</td>
</tr>
<tr id="B595AE1AFFA0C0BBFF78FB71FAA8328F" box="[163,1363,1220,1245]" gridrow="9" pageId="4" pageNumber="369">
<th id="F644C766FFA0C0BBFF78FB71FDDC328F" box="[163,551,1220,1245]" gridcol="0" gridrow="9" pageId="4" pageNumber="369">Dorsal-fin origin to anal-fin origin</th>
<td id="F644C766FFA0C0BBFD2BFB71FCB5328F" box="[752,846,1220,1245]" gridcol="1" gridrow="9" pageId="4" pageNumber="369">28.2</td>
<td id="F644C766FFA0C0BBFC67FB71FC0D328F" box="[956,1014,1220,1245]" gridcol="2" gridrow="9" pageId="4" pageNumber="369">27.9</td>
<td id="F644C766FFA0C0BBFB8DFB71FB47328F" box="[1110,1212,1220,1245]" gridcol="3" gridrow="9" pageId="4" pageNumber="369">26.4–28.8</td>
<td id="F644C766FFA0C0BBFAFDFB71FAA8328F" box="[1318,1363,1220,1245]" gridcol="4" gridrow="9" pageId="4" pageNumber="369">0.7</td>
</tr>
<tr id="B595AE1AFFA0C0BBFF78FB5BFAA83355" box="[163,1363,1262,1287]" gridrow="10" pageId="4" pageNumber="369">
<th id="F644C766FFA0C0BBFF78FB5BFDDC3355" box="[163,551,1262,1287]" gridcol="0" gridrow="10" pageId="4" pageNumber="369">Dorsal-fin origin to pelvic-fin origin</th>
<td id="F644C766FFA0C0BBFD2BFB5BFCB53355" box="[752,846,1262,1287]" gridcol="1" gridrow="10" pageId="4" pageNumber="369">24.6</td>
<td id="F644C766FFA0C0BBFC67FB5BFC0D3355" box="[956,1014,1262,1287]" gridcol="2" gridrow="10" pageId="4" pageNumber="369">23.3</td>
<td id="F644C766FFA0C0BBFB8DFB5BFB473355" box="[1110,1212,1262,1287]" gridcol="3" gridrow="10" pageId="4" pageNumber="369">21.6–25.0</td>
<td id="F644C766FFA0C0BBFAFDFB5BFAA83355" box="[1318,1363,1262,1287]" gridcol="4" gridrow="10" pageId="4" pageNumber="369">1.0</td>
</tr>
<tr id="B595AE1AFFA0C0BBFF78FAADFAA83363" box="[163,1363,1304,1329]" gridrow="11" pageId="4" pageNumber="369">
<th id="F644C766FFA0C0BBFF78FAADFDDC3363" box="[163,551,1304,1329]" gridcol="0" gridrow="11" pageId="4" pageNumber="369">Dorsal-fin origin to pectoral-fin origin</th>
<td id="F644C766FFA0C0BBFD2BFAADFCB53363" box="[752,846,1304,1329]" gridcol="1" gridrow="11" pageId="4" pageNumber="369">31.4</td>
<td id="F644C766FFA0C0BBFC67FAADFC0D3363" box="[956,1014,1304,1329]" gridcol="2" gridrow="11" pageId="4" pageNumber="369">30.9</td>
<td id="F644C766FFA0C0BBFB8DFAADFB473363" box="[1110,1212,1304,1329]" gridcol="3" gridrow="11" pageId="4" pageNumber="369">28.6–33.5</td>
<td id="F644C766FFA0C0BBFAFDFAADFAA83363" box="[1318,1363,1304,1329]" gridcol="4" gridrow="11" pageId="4" pageNumber="369">1.2</td>
</tr>
<tr id="B595AE1AFFA0C0BBFF78FAF7FAA83309" box="[163,1363,1346,1371]" gridrow="12" pageId="4" pageNumber="369">
<th id="F644C766FFA0C0BBFF78FAF7FDDC3309" box="[163,551,1346,1371]" gridcol="0" gridrow="12" pageId="4" pageNumber="369">Caudal-peduncle depth</th>
<td id="F644C766FFA0C0BBFD2BFAF7FCB53309" box="[752,846,1346,1371]" gridcol="1" gridrow="12" pageId="4" pageNumber="369">12.1</td>
<td id="F644C766FFA0C0BBFC67FAF7FC0D3309" box="[956,1014,1346,1371]" gridcol="2" gridrow="12" pageId="4" pageNumber="369">11.3</td>
<td id="F644C766FFA0C0BBFB8DFAF7FB473309" box="[1110,1212,1346,1371]" gridcol="3" gridrow="12" pageId="4" pageNumber="369">10.5–12.0</td>
<td id="F644C766FFA0C0BBFAFDFAF7FAA83309" box="[1318,1363,1346,1371]" gridcol="4" gridrow="12" pageId="4" pageNumber="369">0.5</td>
</tr>
<tr id="B595AE1AFFA0C0BBFF78FAD9FAA833D7" box="[163,1363,1388,1413]" gridrow="13" pageId="4" pageNumber="369">
<th id="F644C766FFA0C0BBFF78FAD9FDDC33D7" box="[163,551,1388,1413]" gridcol="0" gridrow="13" pageId="4" pageNumber="369">Pectoral-fin length</th>
<td id="F644C766FFA0C0BBFD2BFAD9FCB533D7" box="[752,846,1388,1413]" gridcol="1" gridrow="13" pageId="4" pageNumber="369">18.2</td>
<td id="F644C766FFA0C0BBFC67FAD9FC0D33D7" box="[956,1014,1388,1413]" gridcol="2" gridrow="13" pageId="4" pageNumber="369">16.8</td>
<td id="F644C766FFA0C0BBFB8DFAD9FB4733D7" box="[1110,1212,1388,1413]" gridcol="3" gridrow="13" pageId="4" pageNumber="369">14.2–18.4</td>
<td id="F644C766FFA0C0BBFAFDFAD9FAA833D7" box="[1318,1363,1388,1413]" gridcol="4" gridrow="13" pageId="4" pageNumber="369">1.4</td>
</tr>
<tr id="B595AE1AFFA0C0BBFF78FA23FAA833FD" box="[163,1363,1430,1455]" gridrow="14" pageId="4" pageNumber="369">
<th id="F644C766FFA0C0BBFF78FA23FDDC33FD" box="[163,551,1430,1455]" gridcol="0" gridrow="14" pageId="4" pageNumber="369">Pelvic-fin length</th>
<td id="F644C766FFA0C0BBFD2BFA23FCB533FD" box="[752,846,1430,1455]" gridcol="1" gridrow="14" pageId="4" pageNumber="369">14.1</td>
<td id="F644C766FFA0C0BBFC67FA23FC0D33FD" box="[956,1014,1430,1455]" gridcol="2" gridrow="14" pageId="4" pageNumber="369">13.4</td>
<td id="F644C766FFA0C0BBFB8DFA23FB4733FD" box="[1110,1212,1430,1455]" gridcol="3" gridrow="14" pageId="4" pageNumber="369">11.8–14.5</td>
<td id="F644C766FFA0C0BBFAFDFA23FAA833FD" box="[1318,1363,1430,1455]" gridcol="4" gridrow="14" pageId="4" pageNumber="369">0.8</td>
</tr>
<tr id="B595AE1AFFA0C0BBFF78FA75FAA8338B" box="[163,1363,1472,1497]" gridrow="15" pageId="4" pageNumber="369">
<th id="F644C766FFA0C0BBFF78FA75FDDC338B" box="[163,551,1472,1497]" gridcol="0" gridrow="15" pageId="4" pageNumber="369">Dorsal-fin length</th>
<td id="F644C766FFA0C0BBFD2BFA75FCB5338B" box="[752,846,1472,1497]" gridcol="1" gridrow="15" pageId="4" pageNumber="369">22.2</td>
<td id="F644C766FFA0C0BBFC67FA75FC0D338B" box="[956,1014,1472,1497]" gridcol="2" gridrow="15" pageId="4" pageNumber="369">20.5</td>
<td id="F644C766FFA0C0BBFB8DFA75FB47338B" box="[1110,1212,1472,1497]" gridcol="3" gridrow="15" pageId="4" pageNumber="369">18.8–23.6</td>
<td id="F644C766FFA0C0BBFAFDFA75FAA8338B" box="[1318,1363,1472,1497]" gridcol="4" gridrow="15" pageId="4" pageNumber="369">1.4</td>
</tr>
<tr id="B595AE1AFFA0C0BBFF78FA5FFAA83051" box="[163,1363,1514,1539]" gridrow="16" pageId="4" pageNumber="369">
<th id="F644C766FFA0C0BBFF78FA5FFDDC3051" box="[163,551,1514,1539]" gridcol="0" gridrow="16" pageId="4" pageNumber="369">Anal-fin length</th>
<td id="F644C766FFA0C0BBFD2BFA5FFCB53051" box="[752,846,1514,1539]" gridcol="1" gridrow="16" pageId="4" pageNumber="369">16.6</td>
<td id="F644C766FFA0C0BBFC67FA5FFC0D3051" box="[956,1014,1514,1539]" gridcol="2" gridrow="16" pageId="4" pageNumber="369">16.7</td>
<td id="F644C766FFA0C0BBFB8DFA5FFB473051" box="[1110,1212,1514,1539]" gridcol="3" gridrow="16" pageId="4" pageNumber="369">15.1–19.6</td>
<td id="F644C766FFA0C0BBFAFDFA5FFAA83051" box="[1318,1363,1514,1539]" gridcol="4" gridrow="16" pageId="4" pageNumber="369">1.1</td>
</tr>
<tr id="B595AE1AFFA0C0BBFF78F9A1FAA8307F" box="[163,1363,1556,1581]" gridrow="17" pageId="4" pageNumber="369">
<th id="F644C766FFA0C0BBFF78F9A1FDDC307F" box="[163,551,1556,1581]" gridcol="0" gridrow="17" pageId="4" pageNumber="369">Head length</th>
<td id="F644C766FFA0C0BBFD2BF9A1FCB5307F" box="[752,846,1556,1581]" gridcol="1" gridrow="17" pageId="4" pageNumber="369">24.4</td>
<td id="F644C766FFA0C0BBFC67F9A1FC0D307F" box="[956,1014,1556,1581]" gridcol="2" gridrow="17" pageId="4" pageNumber="369">24.5</td>
<td id="F644C766FFA0C0BBFB8DF9A1FB47307F" box="[1110,1212,1556,1581]" gridcol="3" gridrow="17" pageId="4" pageNumber="369">22.1–26.7</td>
<td id="F644C766FFA0C0BBFAFDF9A1FAA8307F" box="[1318,1363,1556,1581]" gridcol="4" gridrow="17" pageId="4" pageNumber="369">1.2</td>
</tr>
<tr id="B595AE1AFFA0C0BBFF78F98BFAA83005" box="[163,1363,1598,1623]" gridrow="18" pageId="4" pageNumber="369">
<th id="F644C766FFA0C0BBFF78F98BFAA83005" box="[163,1363,1598,1623]" colspan="5" colspanRight="4" gridcol="0" gridrow="18" pageId="4" pageNumber="369">
<emphasis id="39D1704AFFA03F40FF78F98AFE913005" bold="true" box="[163,362,1598,1623]" pageId="4" pageNumber="369">Percentages of HL</emphasis>
</th>
</tr>
<tr id="B595AE1AFFA0C0BBFF78F9DDFAA830D3" box="[163,1363,1640,1665]" gridrow="19" pageId="4" pageNumber="369">
<th id="F644C766FFA0C0BBFF78F9DDFDDC30D3" box="[163,551,1640,1665]" gridcol="0" gridrow="19" pageId="4" pageNumber="369">Postorbital length</th>
<td id="F644C766FFA0C0BBFD2BF9DDFCB530D3" box="[752,846,1640,1665]" gridcol="1" gridrow="19" pageId="4" pageNumber="369">42.9</td>
<td id="F644C766FFA0C0BBFC67F9DDFC0D30D3" box="[956,1014,1640,1665]" gridcol="2" gridrow="19" pageId="4" pageNumber="369">42.5</td>
<td id="F644C766FFA0C0BBFB8DF9DDFB4730D3" box="[1110,1212,1640,1665]" gridcol="3" gridrow="19" pageId="4" pageNumber="369">40.8–47.3</td>
<td id="F644C766FFA0C0BBFAFDF9DDFAA830D3" box="[1318,1363,1640,1665]" gridcol="4" gridrow="19" pageId="4" pageNumber="369">1.7</td>
</tr>
<tr id="B595AE1AFFA0C0BBFF78F927FAA830F9" box="[163,1363,1682,1707]" gridrow="20" pageId="4" pageNumber="369">
<th id="F644C766FFA0C0BBFF78F927FDDC30F9" box="[163,551,1682,1707]" gridcol="0" gridrow="20" pageId="4" pageNumber="369">Snout length</th>
<td id="F644C766FFA0C0BBFD2BF927FCB530F9" box="[752,846,1682,1707]" gridcol="1" gridrow="20" pageId="4" pageNumber="369">26.4</td>
<td id="F644C766FFA0C0BBFC67F927FC0D30F9" box="[956,1014,1682,1707]" gridcol="2" gridrow="20" pageId="4" pageNumber="369">27.0</td>
<td id="F644C766FFA0C0BBFB8DF927FB4730F9" box="[1110,1212,1682,1707]" gridcol="3" gridrow="20" pageId="4" pageNumber="369">25.4–29.3</td>
<td id="F644C766FFA0C0BBFAFDF927FAA830F9" box="[1318,1363,1682,1707]" gridcol="4" gridrow="20" pageId="4" pageNumber="369">1.1</td>
</tr>
<tr id="B595AE1AFFA0C0BBFF78F909FAA83087" box="[163,1363,1724,1749]" gridrow="21" pageId="4" pageNumber="369">
<th id="F644C766FFA0C0BBFF78F909FDDC3087" box="[163,551,1724,1749]" gridcol="0" gridrow="21" pageId="4" pageNumber="369">Horizontal eye diameter</th>
<td id="F644C766FFA0C0BBFD2BF909FCB53087" box="[752,846,1724,1749]" gridcol="1" gridrow="21" pageId="4" pageNumber="369">30.0</td>
<td id="F644C766FFA0C0BBFC67F909FC0D3087" box="[956,1014,1724,1749]" gridcol="2" gridrow="21" pageId="4" pageNumber="369">28.8</td>
<td id="F644C766FFA0C0BBFB8DF909FB473087" box="[1110,1212,1724,1749]" gridcol="3" gridrow="21" pageId="4" pageNumber="369">26.5–32.2</td>
<td id="F644C766FFA0C0BBFAFDF909FAA83087" box="[1318,1363,1724,1749]" gridcol="4" gridrow="21" pageId="4" pageNumber="369">1.4</td>
</tr>
<tr id="B595AE1AFFA0C0BBFF78F953FAA830AD" box="[163,1363,1766,1791]" gridrow="22" pageId="4" pageNumber="369">
<th id="F644C766FFA0C0BBFF78F953FDDC30AD" box="[163,551,1766,1791]" gridcol="0" gridrow="22" pageId="4" pageNumber="369">Interorbital width</th>
<td id="F644C766FFA0C0BBFD2BF953FCB530AD" box="[752,846,1766,1791]" gridcol="1" gridrow="22" pageId="4" pageNumber="369">33.3</td>
<td id="F644C766FFA0C0BBFC67F953FC0D30AD" box="[956,1014,1766,1791]" gridcol="2" gridrow="22" pageId="4" pageNumber="369">32.5</td>
<td id="F644C766FFA0C0BBFB8DF953FB4730AD" box="[1110,1212,1766,1791]" gridcol="3" gridrow="22" pageId="4" pageNumber="369">31.6–34.2</td>
<td id="F644C766FFA0C0BBFAFDF953FAA830AD" box="[1318,1363,1766,1791]" gridcol="4" gridrow="22" pageId="4" pageNumber="369">0.7</td>
</tr>
</table>
</paragraph>
<paragraph id="0B1AAC58FFA03F41FF1CF89BFC6036A9" blockId="4.[151,1437,1838,2044]" lastBlockId="5.[151,1437,153,395]" lastPageId="5" lastPageNumber="370" pageId="4" pageNumber="369">
<emphasis id="39D1704AFFA03F40FF1CF89BFE69311B" bold="true" box="[199,402,1838,1865]" pageId="4" pageNumber="369">Color in alcohol.</emphasis>
Overall ground coloration beige. Dorsal region of body dark. Humeral blotch dark, vertically elongated, in some specimens anteriorly concave, encompassing longitudinally from second to fourth scale of lateral line and five scales vertically. Dark, relatively broad midlateral stripe starting at anteriormost scale of lateral line, extending to base of caudal peduncle. Dark chromatophores aligned with margins of myomeres, starting slightly posterior to anal-fin origin. Dorsal region of the head, upper lip, and anterior third of maxilla lightly dark; ventral and dorsal margins of orbit dark; preopercle and opercle silvery. Dorsal fin with melanophores concentrated on interradial membranes; pectoral fins with some melanophores along border of rays; pelvic, anal and caudal fins with scattered melanophores. Melanophores on caudal fin somewhat concentrated in the basal portion of the lobes. Dorsal and ventral procurrent rays dusky. Adipose fin dusky (
<figureCitation id="939EB0DDFFA13F41FC93FF54FC7136A9" box="[840,906,225,251]" captionStart="FIGURE 1" captionStartId="1.[151,250,1908,1933]" captionTargetBox="[183,1404,1418,1881]" captionTargetId="figure@1.[183,1404,1418,1881]" captionTargetPageId="1" captionText="FIGURE 1. Creagrutus andaki, holotype, CIACOL 3462, 56.8 mm SL, Colombia, Caquetá, Florencia, Río Caquetá basin, Río Hacha drainage, Río Sucre." figureDoi="http://doi.org/10.5281/zenodo.3733687" httpUri="https://zenodo.org/record/3733687/files/figure.png" pageId="5" pageNumber="370">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="0B1AAC58FFA13F41FF1CFEB1FC7F3711" blockId="5.[151,1437,153,395]" pageId="5" pageNumber="370">
<emphasis id="39D1704AFFA13F41FF1CFEB1FE4D374D" bold="true" box="[199,438,260,287]" pageId="5" pageNumber="370">Sexual dimorphism.</emphasis>
No secondary sexually dimorphic characters were observed. However, it should be noted that all specimens were collected during the same month (August).
</paragraph>
</subSubSection>
<subSubSection id="43BFFFD3FFA13F41FF1CFEF9FD8637D9" pageId="5" pageNumber="370" type="distribution">
<paragraph id="0B1AAC58FFA13F41FF1CFEF9FD8637D9" blockId="5.[151,1437,153,395]" pageId="5" pageNumber="370">
<emphasis id="39D1704AFFA13F41FF1CFEF9FE9A3735" bold="true" box="[199,353,332,359]" pageId="5" pageNumber="370">Distribution.</emphasis>
<taxonomicName id="CCA5D7DBFFA13F41FEBDFEF8FDCC3735" authorityName="Albornoz-Garzón &amp; Acosta-Santos &amp; Bogotá-Gregory &amp; Agudelo-Córdoba" authorityYear="2020" box="[358,567,333,359]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="370" phylum="Chordata" rank="species" species="andaki">
<emphasis id="39D1704AFFA13F41FEBDFEF8FDCC3735" box="[358,567,333,359]" italics="true" pageId="5" pageNumber="370">Creagrutus andaki</emphasis>
</taxonomicName>
is known only from its
<typeStatus id="D41E12FAFFA13F41FCE7FEF8FC973735" box="[828,876,333,359]" pageId="5" pageNumber="370">type</typeStatus>
locality in the Río
<collectingRegion id="C96162BAFFA13F41FBE5FEF8FB7A3735" box="[1086,1153,333,359]" country="Colombia" name="Sucre" pageId="5" pageNumber="370">Sucre</collectingRegion>
, Río Hacha drainage, Río
<collectingRegion id="C96162BAFFA13F41FF4CFEC4FF0837D9" box="[151,243,369,395]" country="Colombia" name="Caqueta" pageId="5" pageNumber="370">Caquetá</collectingRegion>
basin,
<collectingRegion id="C96162BAFFA13F41FE98FEC4FE5937D9" box="[323,418,369,395]" country="Colombia" name="Caqueta" pageId="5" pageNumber="370">Caquetá</collectingRegion>
,
<collectingCountry id="73B2ECC8FFA13F41FE76FEC4FDE037D9" box="[429,539,369,395]" name="Colombia" pageId="5" pageNumber="370">Colombia</collectingCountry>
(
<figureCitation id="939EB0DDFFA13F41FDF0FEC4FD8B37D9" box="[555,624,369,395]" captionStart="FIGURE 4" captionStartId="5.[151,250,1297,1322]" captionTargetBox="[189,1403,421,1271]" captionTargetId="figure@5.[183,1404,417,1272]" captionTargetPageId="5" captionText="FIGURE 4. Map of southern portion of Colombia showing type locality (yellow star) of Creagrutus andaki." figureDoi="http://doi.org/10.5281/zenodo.3733693" httpUri="https://zenodo.org/record/3733693/files/figure.png" pageId="5" pageNumber="370">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="5FDAFCD0FFA13F41FF4CFAA4FB1F3378" ID-DOI="http://doi.org/10.5281/zenodo.3733693" ID-Zenodo-Dep="3733693" box="[151,1252,1297,1322]" httpUri="https://zenodo.org/record/3733693/files/figure.png" pageId="5" pageNumber="370" startId="5.[151,250,1297,1322]" targetBox="[189,1403,421,1271]" targetPageId="5">
<paragraph id="0B1AAC58FFA13F41FF4CFAA4FB1F3378" blockId="5.[151,1252,1297,1322]" box="[151,1252,1297,1322]" pageId="5" pageNumber="370">
<emphasis id="39D1704AFFA13F41FF4CFAA4FEE83378" bold="true" box="[151,275,1297,1322]" pageId="5" pageNumber="370">FIGURE 4.</emphasis>
Map of southern portion of Colombia showing type locality (yellow star) of
<taxonomicName id="CCA5D7DBFFA13F41FBC4FAA7FB1B3378" authorityName="Albornoz-Garzón &amp; Acosta-Santos &amp; Bogotá-Gregory &amp; Agudelo-Córdoba" authorityYear="2020" box="[1055,1248,1298,1322]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="370" phylum="Chordata" rank="species" species="andaki">Creagrutus andaki</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="43BFFFD3FFA13F41FF1CFAEDFEEF33E8" pageId="5" pageNumber="370" type="etymology">
<paragraph id="0B1AAC58FFA13F41FF1CFAEDFEEF33E8" blockId="5.[151,1437,1368,1754]" pageId="5" pageNumber="370">
<emphasis id="39D1704AFFA13F41FF1CFAEDFEB53320" bold="true" box="[199,334,1368,1394]" pageId="5" pageNumber="370">Etymology.</emphasis>
The specific epithet “andaki” honor the native indigenous people Andakí or Andaquí, who occupied several sites of the Upper Río
<collectingRegion id="C96162BAFFA13F41FD97FAC9FD5333C4" box="[588,680,1404,1430]" country="Colombia" name="Caqueta" pageId="5" pageNumber="370">Caquetá</collectingRegion>
basin and fought bravely during the Spanish conquest. A noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="43BFFFD3FFA13F41FF1CFA76FE6C3088" pageId="5" pageNumber="370" type="discussion">
<paragraph id="0B1AAC58FFA13F41FF1CFA76FE6C3088" blockId="5.[151,1437,1368,1754]" pageId="5" pageNumber="370">
<emphasis id="39D1704AFFA13F41FF1CFA76FDDA338C" bold="true" box="[199,545,1475,1502]" pageId="5" pageNumber="370">Habitat and ecological notes.</emphasis>
The Río
<collectingRegion id="C96162BAFFA13F41FD50FA71FD30338C" box="[651,715,1476,1502]" country="Colombia" name="Sucre" pageId="5" pageNumber="370">Sucre</collectingRegion>
is a piedmont stream with transparent waters, with many rapids alternated with relatively few pools, substrate composed mainly by boulders, with some areas with cobbles, and margins with a well-developed riparian vegetation (
<figureCitation id="939EB0DDFFA13F41FD00F9B9FCE43074" box="[731,799,1547,1574]" captionStart="FIGURE 5" captionStartId="6.[151,250,1286,1311]" captionTargetBox="[231,1357,417,1262]" captionTargetId="figure@6.[231,1357,417,1262]" captionTargetPageId="6" captionText="FIGURE 5. Type locality of Creagrutus andaki, Colombia, Caquetá, Río Caquetá basin, Florencia, Río Hacha drainage, Río Sucre." figureDoi="http://doi.org/10.5281/zenodo.3733695" httpUri="https://zenodo.org/record/3733695/files/figure.png" pageId="5" pageNumber="370">Fig. 5</figureCitation>
). Some physical and chemical parameters of the Río
<collectingRegion id="C96162BAFFA13F41FAAEF9B9FF423019" country="Colombia" name="Sucre" pageId="5" pageNumber="370">Sucre</collectingRegion>
are: temperature 18.4–20.2°C, dissolved oxygen
<quantity id="CC5D01BDFFA13F41FD39F985FC9A3019" box="[738,865,1584,1611]" metricMagnitude="-6" metricUnit="kg" metricValue="7.95" metricValueMax="8.3" metricValueMin="7.6" pageId="5" pageNumber="370" unit="mg" value="7.95" valueMax="8.3" valueMin="7.6">7.6–8.3 mg</quantity>
O/l, pH 5.5–8, conductivity 19–23 µS/cm, average flow
<quantity id="CC5D01BDFFA13F41FF08F9E1FEC8303C" box="[211,307,1620,1646]" metricMagnitude="-1" metricUnit="m" metricValue="3.7" pageId="5" pageNumber="370" unit="m" value="0.37">0.370 m</quantity>
/s, total width
<quantity id="CC5D01BDFFA13F41FE0CF9E6FDD2303D" box="[471,553,1619,1647]" metricMagnitude="0" metricUnit="m" metricValue="7.5" pageId="5" pageNumber="370" unit="m" value="7.5">7.50 m</quantity>
, and discharge
<quantity id="CC5D01BDFFA13F41FD00F9E1FCF3303D" box="[731,776,1620,1647]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="370" unit="m" value="2.0">2 m</quantity>
<superScript id="FCD00110FFA13F41FCD3F9E6FCEB3033" attach="left" box="[776,784,1619,1633]" fontSize="6" pageId="5" pageNumber="370">3</superScript>
/s and 1997 L/s.
<taxonomicName id="CCA5D7DBFFA13F41FC0AF9E0FB5E303C" authorityName="Albornoz-Garzón &amp; Acosta-Santos &amp; Bogotá-Gregory &amp; Agudelo-Córdoba" authorityYear="2020" box="[977,1189,1620,1647]" class="Actinopterygii" family="Characidae" genus="Creagrutus" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="370" phylum="Chordata" rank="species" species="andaki">
<emphasis id="39D1704AFFA13F41FC0AF9E0FB5E303C" box="[977,1189,1620,1647]" italics="true" pageId="5" pageNumber="370">Creagrutus andaki</emphasis>
</taxonomicName>
is found syntopically with
<taxonomicName id="CCA5D7DBFFA13F41FF0BF9CDFE6A30C0" authorityName="Myers" authorityYear="1927" box="[208,401,1656,1682]" class="Actinopterygii" family="Characidae" genus="Othonocheirodus" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="370" phylum="Chordata" rank="genus">
<emphasis id="39D1704AFFA13F41FF0BF9CDFE6A30C0" box="[208,401,1656,1682]" italics="true" pageId="5" pageNumber="370">Othonocheirodus</emphasis>
</taxonomicName>
sp.,
<taxonomicName id="CCA5D7DBFFA13F41FE1DF9CDFD0530C0" authority="Gery" authorityName="Gery" authorityYear="1972" box="[454,766,1656,1682]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="370" phylum="Chordata" rank="species" species="boehlkei">
<emphasis id="39D1704AFFA13F41FE1DF9CDFD4430C0" box="[454,703,1656,1682]" italics="true" pageId="5" pageNumber="370">Characidium boehlkei</emphasis>
Géry
</taxonomicName>
and
<taxonomicName id="CCA5D7DBFFA13F41FCEFF9CDFB1230C0" authority="Fowler. Stomach" authorityName="Fowler. Stomach" authorityYear="1943" box="[820,1257,1656,1682]" class="Actinopterygii" family="Astroblepidae" genus="Astroblepus" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="370" phylum="Chordata" rank="species" species="caquetae">
<emphasis id="39D1704AFFA13F41FCEFF9CDFBDE30C0" box="[820,1061,1656,1682]" italics="true" pageId="5" pageNumber="370">Astroblepus caquetae</emphasis>
Fowler. Stomach
</taxonomicName>
contents of two cleared and stained specimens (CIACOL 2130 and IAvH-P 20565) included insects, fragments of seeds and unidentified organic material.
</paragraph>
</subSubSection>
</treatment>
</document>
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