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<document id="9594D128AFDF380AE2453759657063C0" ID-CLB-Dataset="24427" ID-DOI="10.11646/zootaxa.4718.2.8" ID-GBIF-Dataset="64a14054-6473-4319-96ca-bbb539975f57" ID-ISSN="1175-5326" ID-Zenodo-Dep="3602406" ID-ZooBank="824A0CEA-92F9-482C-B967-9050D946" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1578564737112" checkinUser="plazi" docAuthor="Pagola-Carte, Santiago &amp; Matocq, Armand" docDate="2020" docId="4A3887ECA71C297AFF5C2125FE91FB54" docLanguage="en" docName="zootaxa.4718.2.8.pdf" docOrigin="Zootaxa 4718 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Delphiphylus Pagola-Carte &amp; Matocq 2020, n. gen." docType="treatment" docVersion="6" lastPageNumber="272" masterDocId="B601FF94A71D2979FFCB224CFF86FFB8" masterDocTitle="A new genus to accommodate the misplaced Nearctic species Hoplomachus affiguratus (Uhler, 1895) (Heteroptera: Miridae: Phylinae)" masterLastPageNumber="276" masterPageNumber="269" pageNumber="270" updateTime="1698776555722" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C2F76362ED6E243ABD667C79C0811DB1">A new genus to accommodate the misplaced Nearctic species Hoplomachus affiguratus (Uhler, 1895) (Heteroptera: Miridae: Phylinae)</mods:title>
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<mods:namePart id="09430EF0A820022EBC2FE3CBC0DB9CD8">Pagola-Carte, Santiago</mods:namePart>
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<mods:namePart id="979332783B47503AAD150F43466DC88C">Matocq, Armand</mods:namePart>
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<mods:date id="C939AD7FFC90C2A5BE0F455E608A032D">2020</mods:date>
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<subSubSection id="8A8B6571A71C2978FF5C2125FC0EFC18" pageId="1" pageNumber="270" type="nomenclature">
<paragraph id="C22E36FAA71C2978FF5C2125FE14FC3B" blockId="1.[151,904,873,928]" box="[151,402,873,900]" pageId="1" pageNumber="270">
<heading id="99668196A71C2978FF5C2125FE14FC3B" bold="true" box="[151,402,873,900]" fontSize="11" level="1" pageId="1" pageNumber="270" reason="1">
<emphasis id="F0E5EAE8A71C2978FF5C2125FE14FC3B" bold="true" box="[151,402,873,900]" pageId="1" pageNumber="270">
<taxonomicName id="05914D79A71C2978FF5C2125FEBEFC3C" authority="Pagola-Carte &amp; Matocq, 2020" authorityName="Pagola-Carte &amp; Matocq" authorityYear="2020" box="[151,312,873,900]" class="Insecta" family="Miridae" genus="Delphiphylus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="270" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="F0E5EAE8A71C2978FF5C2125FEBEFC3C" bold="true" box="[151,312,873,900]" italics="true" pageId="1" pageNumber="270">Delphiphylus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EBD65793A71C2978FE8B2125FE14FC3B" box="[320,402,873,899]" pageId="1" pageNumber="270" rank="genus">n. gen.</taxonomicNameLabel>
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<typeStatus id="1D2A8858A71C2978FF5C2187FF4FFC5B" box="[151,201,971,995]" pageId="1" pageNumber="270">Type</typeStatus>
species:
<taxonomicName id="05914D79A71C2978FEED2186FD34FC5B" authority="(Uhler, 1895)" authorityName="Pagola-Carte &amp; Matocq" authorityYear="2020" baseAuthorityName="Uhler" baseAuthorityYear="1895" box="[294,690,970,995]" class="Insecta" family="Miridae" genus="Delphiphylus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="270" phylum="Arthropoda" rank="species" species="affiguratus" status="comb. nov.">
<emphasis id="F0E5EAE8A71C2978FEED2186FDA5FC5B" box="[294,547,970,995]" italics="true" pageId="1" pageNumber="270">Delphiphylus affiguratus</emphasis>
(
<bibRefCitation id="A6004B0BA71C2978FDFB2186FD2DFC5B" author="Uhler, P. R." box="[560,683,970,995]" pageId="1" pageNumber="270" pagination="53 - 55" refId="ref4585" refString="Uhler, P. R. (1895) [Descriptions of new genera and species]. In: Gillette, G. P. &amp; Baker, C. F. (Eds.), A preliminary list of Hemiptera of Colorado. Bulletin of the Colorado Agricultural Experiment Station, 31, pp. 53 - 55" type="journal article" year="1895">Uhler, 1895</bibRefCitation>
)
</taxonomicName>
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<taxonomicNameLabel id="EBD65793A71C2978FD732187FC93FC5B" box="[696,789,971,995]" pageId="1" pageNumber="270" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="C22E36FAA71C2978FF5C265CFF71FB20" blockId="1.[151,1437,1040,2039]" pageId="1" pageNumber="270">
<emphasis id="F0E5EAE8A71C2978FF5C265CFEDAFB93" bold="true" box="[151,348,1040,1067]" pageId="1" pageNumber="270">Material studied</emphasis>
:
<specimenCount id="D497FD73A71C2978FEA2265CFE70FB93" box="[361,502,1040,1067]" pageId="1" pageNumber="270" type="generic">5 specimens</specimenCount>
:
<specimenCount id="D497FD73A71C2978FE34265DFDB9FB93" box="[511,575,1041,1067]" pageId="1" pageNumber="270" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="D497FD73A71C2978FD82265DFDF4FB93" box="[585,626,1041,1067]" pageId="1" pageNumber="270" type="female">1 ♀</specimenCount>
,
<materialsCitation id="72F93CA7A71C2978FDB4265DFABDFB93" ID-GBIF-Occurrence="2545448295" box="[639,1339,1041,1067]" collectingDate="1964-07-16" collectorName="H. H. Knight" location="Steamboat Springs" pageId="1" pageNumber="270" specimenCount="1" stateProvince="Colorado">
<collectingRegion id="0055F818A71C2978FDB4265DFD6AFB93" box="[639,748,1041,1067]" country="United States of America" name="Colorado" pageId="1" pageNumber="270">Colorado</collectingRegion>
:
<location id="C74E6021A71C2978FD3F265DFC4EFB93" LSID="urn:lsid:plazi:treatment:4A3887ECA71C297AFF5C2125FE91FB54:C74E6021A71C2978FD3F265DFC4EFB93" box="[756,968,1041,1067]" name="Steamboat Springs" pageId="1" pageNumber="270" stateProvince="Colorado">Steamboat Springs</location>
,
<date id="B62F103AA71C2978FC19265DFBE6FB93" box="[978,1120,1041,1067]" pageId="1" pageNumber="270" value="1964-07-16">
<collectingDate id="A66BE9D2A71C2978FC19265DFBE6FB93" box="[978,1120,1041,1067]" pageId="1" pageNumber="270" value="1964-07-16">16 july 1964</collectingDate>
</date>
, 7000
<collectorName id="6F64532CA71C2978FB65265DFABDFB93" box="[1198,1339,1041,1067]" pageId="1" pageNumber="270">H.H. Knight</collectorName>
</materialsCitation>
;
<specimenCount id="D497FD73A71C2978FA8D265DFAF7FB93" box="[1350,1393,1041,1067]" pageId="1" pageNumber="270" type="male">1 ♂</specimenCount>
:
<materialsCitation id="72F93CA7A71C2978FAB7265DFA1EFBF7" ID-GBIF-Occurrence="2545448296" collectingDate="1992-07-28" collectionCode="T" collectorName="W. A. Jones. &amp; A. Matocq &amp; Smithsonian Institution &amp; Henry" location="Co Routt Co Yampa" pageId="1" pageNumber="270" specimenCount="1" stateProvince="Colorado">
<location id="C74E6021A71C2978FAB7265DFED5FBF7" LSID="urn:lsid:plazi:treatment:4A3887ECA71C297AFF5C2125FE91FB54:C74E6021A71C2978FAB7265DFED5FBF7" name="Co Routt Co Yampa" pageId="1" pageNumber="270" stateProvince="Colorado">Co Routt Co Yampa</location>
,
<date id="B62F103AA71C2978FE962679FE68FBF7" box="[349,494,1077,1103]" pageId="1" pageNumber="270" value="1992-07-28">
<collectingDate id="A66BE9D2A71C2978FE962679FE68FBF7" box="[349,494,1077,1103]" pageId="1" pageNumber="270" value="1992-07-28">28-VII-1992</collectingDate>
</date>
,
<collectorName id="6F64532CA71C2978FE3C2679FDF8FBF7" box="[503,638,1077,1103]" pageId="1" pageNumber="270">W.A. Jones.</collectorName>
(coll.
<collectorName id="6F64532CA71C2978FD092679FCBCFBF7" box="[706,826,1077,1103]" pageId="1" pageNumber="270">A. Matocq</collectorName>
by exchange with
<collectorName id="6F64532CA71C2978FBC12679FA96FBF7" box="[1034,1296,1077,1103]" pageId="1" pageNumber="270">Smithsonian Institution</collectorName>
(
<collectionCode id="A480AE3FA71C2978FAD42679FAB4FBF7" box="[1311,1330,1077,1103]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="270">T</collectionCode>
.
<collectorName id="6F64532CA71C2978FAF72679FA00FBF7" box="[1340,1414,1077,1103]" pageId="1" pageNumber="270">Henry</collectorName>
))
</materialsCitation>
;
<specimenCount id="D497FD73A71C2978FF5C2615FF47FBCB" box="[151,193,1113,1139]" pageId="1" pageNumber="270" type="female">1 ♀</specimenCount>
,
<materialsCitation id="72F93CA7A71C2978FF052615FF75FB20" ID-GBIF-Occurrence="2545448294" collectingDate="1979-06-05" collectionCode="AMNH, R, T" collectorName="Lm N Kamela &amp; Coll. Poman &amp; Schuh" location="Umatilla CO." pageId="1" pageNumber="270" specimenCode="IZC 00110670" specimenCount="1" stateProvince="Colorado">
OR
<location id="C74E6021A71C2978FF372615FE15FBCB" LSID="urn:lsid:plazi:treatment:4A3887ECA71C297AFF5C2125FE91FB54:C74E6021A71C2978FF372615FE15FBCB" box="[252,403,1113,1139]" name="Umatilla CO." pageId="1" pageNumber="270" stateProvince="Colorado">Umatilla CO.</location>
<collectorName id="6F64532CA71C2978FE522615FDBDFBCB" box="[409,571,1113,1139]" pageId="1" pageNumber="270">Lm N Kamela</collectorName>
,
<date id="B62F103AA71C2978FD8E2614FD44FBCB" box="[581,706,1112,1139]" pageId="1" pageNumber="270" value="1979-06-05">
<collectingDate id="A66BE9D2A71C2978FD8E2614FD44FBCB" box="[581,706,1112,1139]" pageId="1" pageNumber="270" value="1979-06-05">5 Jun 1979</collectingDate>
</date>
,
<collectorName id="6F64532CA71C2978FD062615FCDAFBCB" box="[717,860,1113,1139]" pageId="1" pageNumber="270">Coll. Poman</collectorName>
,
<collectionCode id="A480AE3FA71C2978FCAF2615FC3CFBCB" box="[868,954,1113,1139]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="1" pageNumber="270">AMNH</collectionCode>
_
<specimenCode id="92379E81A71C2978FC032615FBEDFBCB" box="[968,1131,1113,1139]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="1" pageNumber="270">IZC 00110670</specimenCode>
(loaned from
<collectionCode id="A480AE3FA71C2978FACC2615FADBFBCB" box="[1287,1373,1113,1139]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="1" pageNumber="270">AMNH</collectionCode>
(
<collectionCode id="A480AE3FA71C2978FAA72615FA04FBCB" box="[1388,1410,1113,1139]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="270">R</collectionCode>
.
<collectionCode id="A480AE3FA71C2978FA4E2615FA1EFBCB" box="[1413,1432,1113,1139]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="270">T</collectionCode>
.
<collectorName id="6F64532CA71C2978FF5C2631FF67FB20" box="[151,225,1149,1176]" pageId="1" pageNumber="270">Schuh</collectorName>
))
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8A8B6571A71C2978FF0C26ECFA9DF99B" pageId="1" pageNumber="270" type="diagnosis">
<paragraph id="C22E36FAA71C2978FF0C26ECFA9DF99B" blockId="1.[151,1437,1040,2039]" pageId="1" pageNumber="270">
<emphasis id="F0E5EAE8A71C2978FF0C26ECFEBCFB03" bold="true" box="[199,314,1184,1211]" pageId="1" pageNumber="270">Diagnosis</emphasis>
:
<taxonomicName id="05914D79A71C2978FE8226EDFE2DFB03" authorityName="Douglas &amp; Scott" authorityYear="1865" box="[329,427,1185,1211]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="270" phylum="Arthropoda" rank="subFamily" subFamily="Phylinae">Phylinae</taxonomicName>
elongate and rather flat dorsally. Tegument shiny. Vestiture consisting of long, erect, black setae, arising from dark spots. Antennae robust but not incrassate. Calli of pronotum strongly marked, both structurally and chromatically, and delimited by dark spots. Mesothoracic spiracle and metasternal scent gland as in
<figureCitation id="5AAA2A7FA71C2978FABB26A5FF21FA9F" captionStart="FIGURES 34" captionStartId="2.[151,264,1714,1739]" captionTargetBox="[157,1434,1140,1692]" captionTargetId="figure@2.[151,1436,1138,1694]" captionTargetPageId="2" captionText="FIGURES 34. Meso- and metapleura: 3. Delphiphylus n. gen. affiguratus (Uhler, 1895) n. comb., female; 4. Hoplomachus thunbergi (Fallén, 1807), female. (Only mesothoracic spiracle and metasternal scent-efferent system highlighted.)" figureDoi="http://doi.org/10.5281/zenodo.3602410" httpUri="https://zenodo.org/record/3602410/files/figure.png" pageId="1" pageNumber="270">Fig. 3</figureCitation>
. Hemelytral membrane greyish with whitish veins. Femora spotted. Tibial spines black, arising from dark spots. Claws (
<figureCitation id="5AAA2A7FA71C2978FF25277DFEA9FAF3" box="[238,303,1329,1355]" captionStart="FIGURES 12" captionStartId="2.[151,264,1063,1088]" captionTargetBox="[158,1426,559,1034]" captionTargetId="figure@2.[151,1435,553,1043]" captionTargetPageId="2" captionText="FIGURES 12. Delphiphylus n. gen. affiguratus (Uhler, 1895) n. comb. 1. Metatarsal claw (Scale bar = 0.05 mm); 2. Pygo- phore, dorsal and lateral views. (Scale bar = 0.5 mm)" figureDoi="http://doi.org/10.5281/zenodo.3602408" httpUri="https://zenodo.org/record/3602408/files/figure.png" pageId="1" pageNumber="270">Fig. 1</figureCitation>
) robust, strongly bent medially with the apical part nearly straight and the basal part conspicuously wide; pulvillus large but narrow, largely joined to the basal part of the claw. Male genitalia: pygophore (
<figureCitation id="5AAA2A7FA71C2978FAD22719FADAFAD7" box="[1305,1372,1365,1391]" captionStart="FIGURES 12" captionStartId="2.[151,264,1063,1088]" captionTargetBox="[158,1426,559,1034]" captionTargetId="figure@2.[151,1435,553,1043]" captionTargetPageId="2" captionText="FIGURES 12. Delphiphylus n. gen. affiguratus (Uhler, 1895) n. comb. 1. Metatarsal claw (Scale bar = 0.05 mm); 2. Pygo- phore, dorsal and lateral views. (Scale bar = 0.5 mm)" figureDoi="http://doi.org/10.5281/zenodo.3602408" httpUri="https://zenodo.org/record/3602408/files/figure.png" pageId="1" pageNumber="270">Fig. 2</figureCitation>
) very long; right paramere (
<figureCitation id="5AAA2A7FA71C2978FE442735FE55FA2B" box="[399,467,1400,1427]" captionStart="FIGURES 57" captionStartId="4.[151,264,1977,2002]" captionTargetBox="[249,1331,180,1940]" captionTargetId="figure@4.[239,1347,166,1958]" captionTargetPageId="4" captionText="FIGURES 57. Delphiphylus n. gen. affiguratus (Uhler, 1895) n. comb., male genitalia. 5. Right paramere, different views; 6. Left paramere, lateral and (simplified) caudal views; 7. Vesica, two views. (Scale bar = 0.4 mm)" figureDoi="http://doi.org/10.5281/zenodo.3602412" httpUri="https://zenodo.org/record/3602412/files/figure.png" pageId="1" pageNumber="270">Fig. 5</figureCitation>
) rectilinear, with a basal protuberance; left paramere (
<figureCitation id="5AAA2A7FA71C2978FBF52735FB05FA2C" box="[1086,1155,1401,1428]" captionStart="FIGURES 57" captionStartId="4.[151,264,1977,2002]" captionTargetBox="[249,1331,180,1940]" captionTargetId="figure@4.[239,1347,166,1958]" captionTargetPageId="4" captionText="FIGURES 57. Delphiphylus n. gen. affiguratus (Uhler, 1895) n. comb., male genitalia. 5. Right paramere, different views; 6. Left paramere, lateral and (simplified) caudal views; 7. Vesica, two views. (Scale bar = 0.4 mm)" figureDoi="http://doi.org/10.5281/zenodo.3602412" httpUri="https://zenodo.org/record/3602412/files/figure.png" pageId="1" pageNumber="270">Fig. 6</figureCitation>
) with sensory lobe short and blunt; vesica (
<figureCitation id="5AAA2A7FA71C2978FEA927D1FE23FA0F" box="[354,421,1437,1463]" captionStart="FIGURES 57" captionStartId="4.[151,264,1977,2002]" captionTargetBox="[249,1331,180,1940]" captionTargetId="figure@4.[239,1347,166,1958]" captionTargetPageId="4" captionText="FIGURES 57. Delphiphylus n. gen. affiguratus (Uhler, 1895) n. comb., male genitalia. 5. Right paramere, different views; 6. Left paramere, lateral and (simplified) caudal views; 7. Vesica, two views. (Scale bar = 0.4 mm)" figureDoi="http://doi.org/10.5281/zenodo.3602412" httpUri="https://zenodo.org/record/3602412/files/figure.png" pageId="1" pageNumber="270">Fig. 7</figureCitation>
) “J”-shaped with lateral torsion, composed of two thick strands ending in two apical points forming a sub-right angle, with the secondary gonopore in subapical position. Female genitalia: dorsal wall of the genital chamber (
<figureCitation id="5AAA2A7FA71C2978FE9227A9FE1DFA47" box="[345,411,1509,1535]" captionStart="FIGURES 810" captionStartId="5.[151,264,1971,1996]" captionTargetBox="[184,1402,488,1945]" captionTargetId="figure@5.[169,1419,481,1951]" captionTargetPageId="5" captionText="FIGURES 810. Delphiphylus n. gen. affiguratus (Uhler, 1895) n. comb., female genitalia. 8. Dorsal wall of the genital chamber; 9. Vestibulum (vulvar area), ventral view; 10. Posterior wall. (Scale bar = 0.4 mm)" figureDoi="http://doi.org/10.5281/zenodo.3602416" httpUri="https://zenodo.org/record/3602416/files/figure.png" pageId="1" pageNumber="270">Fig. 8</figureCitation>
) sub-oval, with sclerotized rings sub-triangular and dorsal sac protruding dorso-posteriorly; vestibulum greatly developed, particularly to the right side (
<figureCitation id="5AAA2A7FA71C2978FCFC2445FC22F99B" box="[823,932,1545,1571]" captionStart="FIGURES 810" captionStartId="5.[151,264,1971,1996]" captionTargetBox="[184,1402,488,1945]" captionTargetId="figure@5.[169,1419,481,1951]" captionTargetPageId="5" captionText="FIGURES 810. Delphiphylus n. gen. affiguratus (Uhler, 1895) n. comb., female genitalia. 8. Dorsal wall of the genital chamber; 9. Vestibulum (vulvar area), ventral view; 10. Posterior wall. (Scale bar = 0.4 mm)" figureDoi="http://doi.org/10.5281/zenodo.3602416" httpUri="https://zenodo.org/record/3602416/files/figure.png" pageId="1" pageNumber="270">Figs. 89</figureCitation>
); posterior wall present (
<figureCitation id="5AAA2A7FA71C2978FB712445FA88F99B" box="[1210,1294,1545,1571]" captionStart="FIGURES 810" captionStartId="5.[151,264,1971,1996]" captionTargetBox="[184,1402,488,1945]" captionTargetId="figure@5.[169,1419,481,1951]" captionTargetPageId="5" captionText="FIGURES 810. Delphiphylus n. gen. affiguratus (Uhler, 1895) n. comb., female genitalia. 8. Dorsal wall of the genital chamber; 9. Vestibulum (vulvar area), ventral view; 10. Posterior wall. (Scale bar = 0.4 mm)" figureDoi="http://doi.org/10.5281/zenodo.3602416" httpUri="https://zenodo.org/record/3602416/files/figure.png" pageId="1" pageNumber="270">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8A8B6571A71C2978FF0C2461FF5DF937" pageId="1" pageNumber="270" type="etymology">
<paragraph id="C22E36FAA71C2978FF0C2461FF5DF937" blockId="1.[151,1437,1040,2039]" pageId="1" pageNumber="270">
<emphasis id="F0E5EAE8A71C2978FF0C2461FECEF9FF" bold="true" box="[199,328,1581,1607]" pageId="1" pageNumber="270">Etymology</emphasis>
: the generic name
<taxonomicName id="05914D79A71C2978FDD12461FD29F9FF" authority="Pagola-Carte &amp; Matocq, 2020" authorityName="Pagola-Carte &amp; Matocq" authorityYear="2020" box="[538,687,1581,1607]" class="Insecta" family="Miridae" genus="Delphiphylus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="274" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="F0E5EAE8A71C2978FDD12461FD29F9FF" box="[538,687,1581,1607]" italics="true" pageId="1" pageNumber="270">Delphiphylus</emphasis>
</taxonomicName>
is formed by combination of
<taxonomicName id="05914D79A71C2978FC332461FBFAF9FF" box="[1016,1148,1581,1607]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="1" pageNumber="270" phylum="Tracheophyta" rank="genus">
<emphasis id="F0E5EAE8A71C2978FC332461FBFAF9FF" box="[1016,1148,1581,1607]" italics="true" pageId="1" pageNumber="270">Delphinium</emphasis>
</taxonomicName>
(generic name of the host plant) and
<taxonomicName id="05914D79A71C2978FED8241DFED9F9D3" authorityName="Hahn" authorityYear="1831" box="[275,351,1617,1643]" class="Insecta" family="Miridae" genus="Phylus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="270" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71C2978FED8241DFED9F9D3" box="[275,351,1617,1643]" italics="true" pageId="1" pageNumber="270">Phylus</emphasis>
</taxonomicName>
(name of a genus of
<taxonomicName id="05914D79A71C2978FD93241DFD3CF9D3" authorityName="Douglas &amp; Scott" authorityYear="1865" box="[600,698,1617,1643]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="270" phylum="Arthropoda" rank="subFamily" subFamily="Phylinae">Phylinae</taxonomicName>
abundantly used as a suffix in this subfamily). It is a masculine name.
</paragraph>
</subSubSection>
<subSubSection id="8A8B6571A71C2978FF0C24D4FB43F90B" box="[199,1221,1688,1715]" pageId="1" pageNumber="270" type="distribution">
<paragraph id="C22E36FAA71C2978FF0C24D4FB43F90B" blockId="1.[151,1437,1040,2039]" box="[199,1221,1688,1715]" pageId="1" pageNumber="270">
<emphasis id="F0E5EAE8A71C2978FF0C24D4FEDFF90B" bold="true" box="[199,345,1688,1715]" pageId="1" pageNumber="270">Distribution</emphasis>
: Nearctic, with a wide distribution in western
<collectingCountry id="BA86766AA71C2978FCA924D5FC1EF90B" box="[866,920,1689,1715]" name="United States of America" pageId="1" pageNumber="270">USA</collectingCountry>
and southwestern
<collectingCountry id="BA86766AA71C2978FBA224D5FB47F90B" box="[1129,1217,1689,1715]" name="Canada" pageId="1" pageNumber="270">Canada</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="8A8B6571A71C2978FF0C24F1FB9DF96F" box="[199,1051,1725,1751]" pageId="1" pageNumber="270" type="biology_ecology">
<paragraph id="C22E36FAA71C2978FF0C24F1FB9DF96F" blockId="1.[151,1437,1040,2039]" box="[199,1051,1725,1751]" pageId="1" pageNumber="270">
<emphasis id="F0E5EAE8A71C2978FF0C24F1FEC9F96F" bold="true" box="[199,335,1725,1751]" pageId="1" pageNumber="270">Host plants</emphasis>
:
<taxonomicName id="05914D79A71C2978FE9524F1FD8DF96F" box="[350,523,1725,1751]" class="Magnoliopsida" family="Ranunculaceae" kingdom="Plantae" order="Ranunculales" pageId="1" pageNumber="270" phylum="Tracheophyta" rank="family">Ranunculaceae</taxonomicName>
. In the only known species, genus
<taxonomicName id="05914D79A71C2978FC5B24F1FB92F96F" box="[912,1044,1725,1751]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="1" pageNumber="270" phylum="Tracheophyta" rank="genus">
<emphasis id="F0E5EAE8A71C2978FC5B24F1FB92F96F" box="[912,1044,1725,1751]" italics="true" pageId="1" pageNumber="270">Delphinium</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="8A8B6571A71C297AFF0C24ACFE91FB54" lastPageId="3" lastPageNumber="272" pageId="1" pageNumber="270" type="discussion">
<paragraph id="C22E36FAA71C2978FF0C24ACFCBFF808" blockId="1.[151,1437,1040,2039]" pageId="1" pageNumber="270">
<emphasis id="F0E5EAE8A71C2978FF0C24ACFEC3F943" bold="true" box="[199,325,1760,1787]" pageId="1" pageNumber="270">Discussion</emphasis>
: Even though a shallow examination of the habitus reveals similarities between
<taxonomicName id="05914D79A71C2978FB0424AEFAEAF943" authorityName="Pagola-Carte &amp; Matocq" authorityYear="2020" baseAuthorityName="Uhler" baseAuthorityYear="1895" box="[1231,1388,1761,1787]" class="Insecta" family="Miridae" genus="Delphiphylus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="270" phylum="Arthropoda" rank="species" species="affiguratus">
<emphasis id="F0E5EAE8A71C2978FB0424AEFAEAF943" box="[1231,1388,1761,1787]" italics="true" pageId="1" pageNumber="270">H. affiguratus</emphasis>
</taxonomicName>
and the other members of
<taxonomicName id="05914D79A71C2978FE462549FDAFF8A7" authorityName="Fieber" authorityYear="1858" box="[397,553,1797,1823]" class="Insecta" family="Miridae" genus="Hoplomachus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="270" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71C2978FE462549FDAFF8A7" box="[397,553,1797,1823]" italics="true" pageId="1" pageNumber="270">Hoplomachus</emphasis>
</taxonomicName>
such as the structure and patterns on head, scutellum and mesoscutum, as well as similarly dark and well-defined pronotal calli, a number of characters distinguish it. Needless to say, the obvious differences of colouration (in
<taxonomicName id="05914D79A71C2978FE2C2502FD03F8DF" authorityName="Pagola-Carte &amp; Matocq" authorityYear="2020" baseAuthorityName="Uhler" baseAuthorityYear="1895" box="[487,645,1869,1895]" class="Insecta" family="Miridae" genus="Delphiphylus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="270" phylum="Arthropoda" rank="species" species="affiguratus">
<emphasis id="F0E5EAE8A71C2978FE2C2502FD03F8DF" box="[487,645,1869,1895]" italics="true" pageId="1" pageNumber="270">H. affiguratus</emphasis>
</taxonomicName>
both sexes are greenish brown to grey) are put aside, in a similar way as they are ignored concerning the Palaearctic members of the genus:
<taxonomicName id="05914D79A71C2978FC7A253EFBDDF833" authorityName="Reuter" authorityYear="1879" box="[945,1115,1905,1931]" class="Insecta" family="Miridae" genus="Tinicephalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="270" phylum="Arthropoda" rank="species" species="flavopilosus">
<emphasis id="F0E5EAE8A71C2978FC7A253EFBDDF833" box="[945,1115,1905,1931]" italics="true" pageId="1" pageNumber="270">H. flavopilosus</emphasis>
</taxonomicName>
(yellow-ochre-greenish),
<taxonomicName id="05914D79A71C2978FA4A253EFF59F817" authorityName="Reuter" authorityYear="1876" class="Insecta" family="Miridae" genus="Phoenicocapsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="270" phylum="Arthropoda" rank="species" species="regina">
<emphasis id="F0E5EAE8A71C2978FA4A253EFF59F817" italics="true" pageId="1" pageNumber="270">H. regina</emphasis>
</taxonomicName>
(orange-red) and
<taxonomicName id="05914D79A71C2978FE6225DAFDB1F817" baseAuthorityName="Fallen" baseAuthorityYear="1807" box="[425,567,1941,1967]" class="Insecta" family="Miridae" genus="Hoplomachus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="270" phylum="Arthropoda" rank="species" species="thunbergi">
<emphasis id="F0E5EAE8A71C2978FE6225DAFDB1F817" box="[425,567,1941,1967]" italics="true" pageId="1" pageNumber="270">H. thunbergi</emphasis>
</taxonomicName>
(grey-ochre-greenish).
</paragraph>
<paragraph id="C22E36FAA71C297BFF0C25F5FEBCFEFB" blockId="1.[151,1437,1040,2039]" lastBlockId="2.[151,1437,153,539]" lastPageId="2" lastPageNumber="271" pageId="1" pageNumber="270">
Beyond the superficial appearance, the following distinguishing characters are considered as relevant: tegument shiny, setae arising from dark spots, segment II of antennae not incrassate in males, rostrum only reaching meso- coxae, tibial spines arising from dark spots, claws with basal part wide and pulvillus narrow (
<figureCitation id="5AAA2A7FA71F297BFB7A22D5FB75FF0B" box="[1201,1267,153,179]" captionStart="FIGURES 12" captionStartId="2.[151,264,1063,1088]" captionTargetBox="[158,1426,559,1034]" captionTargetId="figure@2.[151,1435,553,1043]" captionTargetPageId="2" captionText="FIGURES 12. Delphiphylus n. gen. affiguratus (Uhler, 1895) n. comb. 1. Metatarsal claw (Scale bar = 0.05 mm); 2. Pygo- phore, dorsal and lateral views. (Scale bar = 0.5 mm)" figureDoi="http://doi.org/10.5281/zenodo.3602408" httpUri="https://zenodo.org/record/3602408/files/figure.png" pageId="2" pageNumber="271">Fig. 1</figureCitation>
), metathoracic scent-efferent system with a large and protruding auricle (
<figureCitation id="5AAA2A7FA71F297BFCD522F1FCE4FF6F" box="[798,866,189,215]" captionStart="FIGURES 34" captionStartId="2.[151,264,1714,1739]" captionTargetBox="[157,1434,1140,1692]" captionTargetId="figure@2.[151,1436,1138,1694]" captionTargetPageId="2" captionText="FIGURES 34. Meso- and metapleura: 3. Delphiphylus n. gen. affiguratus (Uhler, 1895) n. comb., female; 4. Hoplomachus thunbergi (Fallén, 1807), female. (Only mesothoracic spiracle and metasternal scent-efferent system highlighted.)" figureDoi="http://doi.org/10.5281/zenodo.3602410" httpUri="https://zenodo.org/record/3602410/files/figure.png" pageId="2" pageNumber="271">Fig. 3</figureCitation>
). More interestingly, the male and female genitalia are quite different in
<taxonomicName id="05914D79A71F297BFE4922AEFDA6FF43" authorityName="Pagola-Carte &amp; Matocq" authorityYear="2020" baseAuthorityName="Uhler" baseAuthorityYear="1895" box="[386,544,225,251]" class="Insecta" family="Miridae" genus="Delphiphylus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="271" phylum="Arthropoda" rank="species" species="affiguratus">
<emphasis id="F0E5EAE8A71F297BFE4922AEFDA6FF43" box="[386,544,225,251]" italics="true" pageId="2" pageNumber="271">H. affiguratus</emphasis>
</taxonomicName>
(
<figureCitation id="5AAA2A7FA71F297BFDE422ADFD21FF43" box="[559,679,224,251]" captionStart-0="FIGURES 57" captionStart-1="FIGURES 810" captionStartId-0="4.[151,264,1977,2002]" captionStartId-1="5.[151,264,1971,1996]" captionTargetBox-0="[249,1331,180,1940]" captionTargetBox-1="[184,1402,488,1945]" captionTargetId-0="figure@4.[239,1347,166,1958]" captionTargetId-1="figure@5.[169,1419,481,1951]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 57. Delphiphylus n. gen. affiguratus (Uhler, 1895) n. comb., male genitalia. 5. Right paramere, different views; 6. Left paramere, lateral and (simplified) caudal views; 7. Vesica, two views. (Scale bar = 0.4 mm)" captionText-1="FIGURES 810. Delphiphylus n. gen. affiguratus (Uhler, 1895) n. comb., female genitalia. 8. Dorsal wall of the genital chamber; 9. Vestibulum (vulvar area), ventral view; 10. Posterior wall. (Scale bar = 0.4 mm)" figureDoi-0="http://doi.org/10.5281/zenodo.3602412" figureDoi-1="http://doi.org/10.5281/zenodo.3602416" httpUri-0="https://zenodo.org/record/3602412/files/figure.png" httpUri-1="https://zenodo.org/record/3602416/files/figure.png" pageId="2" pageNumber="271">Figs. 510</figureCitation>
) with respect to the Palaearctic
<taxonomicName id="05914D79A71F297BFBC722ADFB2EFF43" authorityName="Fieber" authorityYear="1858" box="[1036,1192,225,251]" class="Insecta" family="Miridae" genus="Hoplomachus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="271" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71F297BFBC722ADFB2EFF43" box="[1036,1192,225,251]" italics="true" pageId="2" pageNumber="271">Hoplomachus</emphasis>
</taxonomicName>
especies (see
<bibRefCitation id="A6004B0BA71F297BFA8C22ADFE08FEA7" author="Matocq, A. &amp; Pagola-Carte, S." pageId="2" pageNumber="271" pagination="435 - 440" refId="ref4038" refString="Matocq, A. &amp; Pagola-Carte, S. (2008) Transfert de deux especes des genres Phoenicocapsus et Tinicephalus dans le genre Hoplomachus (Heteroptera, Miridae, Phylinae). Bulletin de la Societe Entomologique de France, 113 (4), 435 - 440." type="journal article" year="2008">Matocq &amp; Pagola-Carte 2008</bibRefCitation>
: figs. 48, 913, 1418). At this point, we can assume that
<taxonomicName id="05914D79A71F297BFB892349FB3BFEA7" authorityName="Pagola-Carte &amp; Matocq" authorityYear="2020" baseAuthorityName="Uhler" baseAuthorityYear="1895" box="[1090,1213,261,287]" class="Insecta" family="Miridae" genus="Delphiphylus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="271" phylum="Arthropoda" rank="species" species="affiguratus">
<emphasis id="F0E5EAE8A71F297BFB892349FB3BFEA7" box="[1090,1213,261,287]" italics="true" pageId="2" pageNumber="271">affiguratus</emphasis>
</taxonomicName>
does not belong in
<taxonomicName id="05914D79A71F297BFF5C2365FEB5FEFB" authorityName="Fieber" authorityYear="1858" box="[151,307,297,323]" class="Insecta" family="Miridae" genus="Hoplomachus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="271" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71F297BFF5C2365FEB5FEFB" box="[151,307,297,323]" italics="true" pageId="2" pageNumber="271">Hoplomachus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C22E36FAA71F297BFF0C2301FB15FDA3" blockId="2.[151,1437,153,539]" pageId="2" pageNumber="271">
The vesica is extensively used as a primary character to define the genera of
<taxonomicName id="05914D79A71F297BFB802301FB37FEDF" authorityName="Douglas &amp; Scott" authorityYear="1865" box="[1099,1201,333,359]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="271" phylum="Arthropoda" rank="subFamily" subFamily="Phylinae">Phylinae</taxonomicName>
. Accordingly, in our search for a genus into which
<taxonomicName id="05914D79A71F297BFE38233DFDE8FE33" authorityName="Pagola-Carte &amp; Matocq" authorityYear="2020" baseAuthorityName="Uhler" baseAuthorityYear="1895" box="[499,622,369,395]" class="Insecta" family="Miridae" genus="Delphiphylus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="271" phylum="Arthropoda" rank="species" species="affiguratus">
<emphasis id="F0E5EAE8A71F297BFE38233DFDE8FE33" box="[499,622,369,395]" italics="true" pageId="2" pageNumber="271">affiguratus</emphasis>
</taxonomicName>
could be accommodated, we checked the available information on this male genitalic structure. The whole literature available to us was thoroughly studied, with emphasis on Nearctic+Palaearctic+Holarctic
<taxonomicName id="05914D79A71F297BFEBF23F5FE5CFE6B" authorityName="Douglas &amp; Scott" authorityYear="1865" box="[372,474,441,467]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="271" phylum="Arthropoda" rank="subFamily" subFamily="Phylinae">Phylinae</taxonomicName>
. Only a slight resemblance to the Holarctic
<taxonomicName id="05914D79A71F297BFC7123F5FB43FE6B" authority="Reuter, 1909" authorityName="Reuter" authorityYear="1909" box="[954,1221,441,467]" class="Insecta" family="Miridae" genus="Europiella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="271" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71F297BFC7123F5FBB7FE6B" box="[954,1073,441,467]" italics="true" pageId="2" pageNumber="271">Europiella</emphasis>
<bibRefCitation id="A6004B0BA71F297BFBFC23F5FB43FE6B" author="Reuter, O. M." box="[1079,1221,441,467]" pageId="2" pageNumber="271" pagination="1 - 86" refId="ref4146" refString="Reuter, O. M. (1909) Bemerkungen uber nearktische Capsiden nebst Beschreibung neuer Arten. Acta Societatis Scientiarum Fennicae, 36 (2), 1 - 86, 3 pls. https: // doi. org / 10.5962 / bhl. title. 9445" type="book chapter" year="1909">Reuter, 1909</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="05914D79A71F297BFB3223F5FEA1FE4F" authority="Fieber, 1858" authorityName="Fieber" authorityYear="1858" class="Insecta" family="Miridae" genus="Plagiognathus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="271" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71F297BFB3223F5FA1AFE6B" box="[1273,1436,441,467]" italics="true" pageId="2" pageNumber="271">Plagiognathus</emphasis>
Fieber, 1858
</taxonomicName>
, and the Palaearctic
<taxonomicName id="05914D79A71F297BFDC52391FC98FE4F" authority="Fieber, 1858" authorityName="Fieber" authorityYear="1858" box="[526,798,476,503]" class="Insecta" family="Miridae" genus="Amblytylus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="271" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71F297BFDC52391FD0CFE4F" box="[526,650,477,503]" italics="true" pageId="2" pageNumber="271">Amblytylus</emphasis>
Fieber, 1858
</taxonomicName>
and
<taxonomicName id="05914D79A71F297BFC9D2391FBE8FE4F" authority="Fieber, 1858" authorityName="Fieber" authorityYear="1858" box="[854,1134,476,503]" class="Insecta" family="Miridae" genus="Placochilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="271" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71F297BFC9D2391FC5CFE4F" box="[854,986,477,503]" italics="true" pageId="2" pageNumber="271">Placochilus</emphasis>
Fieber, 1858
</taxonomicName>
was noticed, each of these genera showing a disparate combination of characters which precludes further comparison.
</paragraph>
<caption id="96EE6672A71F297BFF5C266BFD37FBDC" ID-DOI="http://doi.org/10.5281/zenodo.3602408" ID-Zenodo-Dep="3602408" httpUri="https://zenodo.org/record/3602408/files/figure.png" pageId="2" pageNumber="271" startId="2.[151,264,1063,1088]" targetBox="[158,1426,559,1034]" targetPageId="2">
<paragraph id="C22E36FAA71F297BFF5C266BFD37FBDC" blockId="2.[151,1436,1063,1124]" pageId="2" pageNumber="271">
<emphasis id="F0E5EAE8A71F297BFF5C266BFEBDFBF8" bold="true" box="[151,315,1063,1088]" pageId="2" pageNumber="271">FIGURES 12.</emphasis>
<taxonomicName id="05914D79A71F297BFE882664FE4CFBF8" authority="Pagola-Carte &amp; Matocq, 2020" authorityName="Pagola-Carte &amp; Matocq" authorityYear="2020" box="[323,458,1064,1088]" class="Insecta" family="Miridae" genus="Delphiphylus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="271" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="F0E5EAE8A71F297BFE882664FE4CFBF8" box="[323,458,1064,1088]" italics="true" pageId="2" pageNumber="271">Delphiphylus</emphasis>
</taxonomicName>
<emphasis id="F0E5EAE8A71F297BFE1A2664FD9FFBF8" bold="true" box="[465,537,1064,1088]" pageId="2" pageNumber="271">
<taxonomicNameLabel id="EBD65793A71F297BFE1A2664FD9FFBF8" box="[465,537,1064,1088]" pageId="2" pageNumber="271" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
<taxonomicName id="05914D79A71F297BFDEB2664FCA6FBF8" authority="(Uhler, 1895)" authorityName="Pagola-Carte &amp; Matocq" authorityYear="2020" baseAuthorityName="Uhler" baseAuthorityYear="1895" box="[544,800,1063,1088]" class="Insecta" family="Miridae" genus="Delphiphylus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="271" phylum="Arthropoda" rank="species" species="affiguratus" status="comb. nov.">
<emphasis id="F0E5EAE8A71F297BFDEB2664FD09FBF8" box="[544,655,1064,1088]" italics="true" pageId="2" pageNumber="271">affiguratus</emphasis>
(
<bibRefCitation id="A6004B0BA71F297BFD55266BFC9CFBF8" author="Uhler, P. R." box="[670,794,1063,1088]" pageId="2" pageNumber="271" pagination="53 - 55" refId="ref4585" refString="Uhler, P. R. (1895) [Descriptions of new genera and species]. In: Gillette, G. P. &amp; Baker, C. F. (Eds.), A preliminary list of Hemiptera of Colorado. Bulletin of the Colorado Agricultural Experiment Station, 31, pp. 53 - 55" type="journal article" year="1895">Uhler, 1895</bibRefCitation>
)
</taxonomicName>
<emphasis id="F0E5EAE8A71F297BFCE32664FC00FBF8" bold="true" box="[808,902,1064,1088]" pageId="2" pageNumber="271">
<taxonomicNameLabel id="EBD65793A71F297BFCE32664FC00FBF8" box="[808,902,1064,1088]" pageId="2" pageNumber="271" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
1. Metatarsal claw (Scale bar = 0.05 mm); 2. Pygo- phore, dorsal and lateral views. (Scale bar = 0.5 mm)
</paragraph>
</caption>
<caption id="96EE6672A71F297BFF5C24FEFA91F957" ID-DOI="http://doi.org/10.5281/zenodo.3602410" ID-Zenodo-Dep="3602410" httpUri="https://zenodo.org/record/3602410/files/figure.png" pageId="2" pageNumber="271" startId="2.[151,264,1714,1739]" targetBox="[157,1434,1140,1692]" targetPageId="2">
<paragraph id="C22E36FAA71F297BFF5C24FEFA91F957" blockId="2.[151,1436,1714,1775]" pageId="2" pageNumber="271">
<emphasis id="F0E5EAE8A71F297BFF5C24FEFEBAF973" bold="true" box="[151,316,1714,1739]" pageId="2" pageNumber="271">FIGURES 34.</emphasis>
Meso- and metapleura: 3.
<taxonomicName id="05914D79A71F297BFD9A24FFFD5EF973" authority="Pagola-Carte &amp; Matocq, 2020" authorityName="Pagola-Carte &amp; Matocq" authorityYear="2020" box="[593,728,1715,1739]" class="Insecta" family="Miridae" genus="Delphiphylus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="271" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="F0E5EAE8A71F297BFD9A24FFFD5EF973" box="[593,728,1715,1739]" italics="true" pageId="2" pageNumber="271">Delphiphylus</emphasis>
</taxonomicName>
<emphasis id="F0E5EAE8A71F297BFD1424FFFCAEF973" bold="true" box="[735,808,1715,1739]" pageId="2" pageNumber="271">
<taxonomicNameLabel id="EBD65793A71F297BFD1424FFFCAEF973" box="[735,808,1715,1739]" pageId="2" pageNumber="271" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
<taxonomicName id="05914D79A71F297BFCE424FFFBA9F973" authority="(Uhler, 1895)" authorityName="Pagola-Carte &amp; Matocq" authorityYear="2020" baseAuthorityName="Uhler" baseAuthorityYear="1895" box="[815,1071,1714,1739]" class="Insecta" family="Miridae" genus="Delphiphylus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="271" phylum="Arthropoda" rank="species" species="affiguratus" status="comb. nov.">
<emphasis id="F0E5EAE8A71F297BFCE424FFFC18F973" box="[815,926,1715,1739]" italics="true" pageId="2" pageNumber="271">affiguratus</emphasis>
(
<bibRefCitation id="A6004B0BA71F297BFC6724FEFBAEF973" author="Uhler, P. R." box="[940,1064,1714,1739]" pageId="2" pageNumber="271" pagination="53 - 55" refId="ref4585" refString="Uhler, P. R. (1895) [Descriptions of new genera and species]. In: Gillette, G. P. &amp; Baker, C. F. (Eds.), A preliminary list of Hemiptera of Colorado. Bulletin of the Colorado Agricultural Experiment Station, 31, pp. 53 - 55" type="journal article" year="1895">Uhler, 1895</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="EBD65793A71F297BFBFC24FFFB12F972" box="[1079,1172,1715,1739]" pageId="2" pageNumber="271" rank="species">
<emphasis id="F0E5EAE8A71F297BFBFC24FFFB08F973" bold="true" box="[1079,1166,1715,1739]" pageId="2" pageNumber="271">n. comb</emphasis>
.
</taxonomicNameLabel>
, female; 4.
<taxonomicName id="05914D79A71F297BFAC424FFFE17F957" authority="(Fallen, 1807)" authorityName="Fallen" authorityYear="1807" baseAuthorityName="Fallen" baseAuthorityYear="1807" class="Insecta" family="Miridae" genus="Hoplomachus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="271" phylum="Arthropoda" rank="species" species="thunbergi">
<emphasis id="F0E5EAE8A71F297BFAC424FFFF7FF957" italics="true" pageId="2" pageNumber="271">Hoplomachus thunbergi</emphasis>
(Fallén, 1807)
</taxonomicName>
, female. (Only mesothoracic spiracle and metasternal scent-efferent system highlighted.)
</paragraph>
</caption>
<paragraph id="C22E36FAA71F297AFF0C2551FD58FE33" blockId="2.[151,1436,1821,2028]" lastBlockId="3.[151,1437,153,1260]" lastPageId="3" lastPageNumber="272" pageId="2" pageNumber="271">
Another approach consisted in a thorough study of the work by
<bibRefCitation id="A6004B0BA71F297BFC6A2551FB3CF88F" author="Schuh, R. T. &amp; Menard, K. L." box="[929,1210,1821,1847]" pageId="2" pageNumber="271" pagination="1 - 72" refId="ref4392" refString="Schuh, R. T. &amp; Menard, K. L. (2013) A revised classification of the Phylinae (Insecta: Heteroptera: Miridae): arguments for the placement of genera. American Museum Novitates, 3785, 1 - 72. https: // doi. org / 10.1206 / 3785.2" type="journal article" year="2013">Schuh &amp; Menard (2013)</bibRefCitation>
, since it is the most comprehensive and recent classification of the
<taxonomicName id="05914D79A71F297BFD7A250DFC95F8E3" authorityName="Douglas &amp; Scott" authorityYear="1865" box="[689,787,1857,1883]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="271" phylum="Arthropoda" rank="subFamily" subFamily="Phylinae">Phylinae</taxonomicName>
at a generic level. These authors place the Holarctic genus
<taxonomicName id="05914D79A71F297BFF0C2529FEE5F8C7" authorityName="Fieber" authorityYear="1858" box="[199,355,1893,1919]" class="Insecta" family="Miridae" genus="Hoplomachus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="271" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71F297BFF0C2529FEE5F8C7" box="[199,355,1893,1919]" italics="true" pageId="2" pageNumber="271">Hoplomachus</emphasis>
</taxonomicName>
in the subtribe Oncotylina (within the tribe
<taxonomicName id="05914D79A71F297BFCAB2529FC35F838" authorityName="Douglas &amp; Scott" authorityYear="1865" box="[864,947,1893,1920]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="271" phylum="Arthropoda" rank="tribe" tribe="Phylini">Phylini</taxonomicName>
) because “the male genitalia and pretarsus in
<taxonomicName id="05914D79A71F297BFF7E25C5FED7F81B" authorityName="Fieber" authorityYear="1858" box="[181,337,1929,1955]" class="Insecta" family="Miridae" genus="Hoplomachus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="271" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71F297BFF7E25C5FED7F81B" box="[181,337,1929,1955]" italics="true" pageId="2" pageNumber="271">Hoplomachus</emphasis>
</taxonomicName>
are very similar to what is seen in
<taxonomicName id="05914D79A71F297BFD1425C5FCE8F81B" authorityName="Fieber" authorityYear="1858" box="[735,878,1929,1955]" class="Insecta" family="Miridae" genus="Tinicephalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="271" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71F297BFD1425C5FCE8F81B" box="[735,878,1929,1955]" italics="true" pageId="2" pageNumber="271">Tinicephalus</emphasis>
</taxonomicName>
and the habitus is similar to some
<taxonomicName id="05914D79A71F297BFB3225C5FA1AF81B" authorityName="Fieber" authorityYear="1858" box="[1273,1436,1929,1955]" class="Insecta" family="Miridae" genus="Plagiognathus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="271" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71F297BFB3225C5FA1AF81B" box="[1273,1436,1929,1955]" italics="true" pageId="2" pageNumber="271">Plagiognathus</emphasis>
</taxonomicName>
spp.”, in spite of several characters of the external morphology that suggest placement in the subtribe Cremnorrhina (within the tribe Cremnorrhini). Being aware of the difficult task faced by these authors, we are obliged to be cau- tious in making no more than a little contribution to the tribal placement discussion. For instance, we partly agree with the above arguments for
<taxonomicName id="05914D79A71E297AFE2B22F1FDFAFF6F" authorityName="Fieber" authorityYear="1858" box="[480,636,189,215]" class="Insecta" family="Miridae" genus="Hoplomachus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71E297AFE2B22F1FDFAFF6F" box="[480,636,189,215]" italics="true" pageId="3" pageNumber="272">Hoplomachus</emphasis>
</taxonomicName>
, but only concerning the three Palaearctic species. The male genitalia of
<taxonomicName id="05914D79A71E297AFF5C22AEFEB2FF43" authorityName="Pagola-Carte &amp; Matocq" authorityYear="2020" baseAuthorityName="Uhler" baseAuthorityYear="1895" box="[151,308,225,251]" class="Insecta" family="Miridae" genus="Delphiphylus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="species" species="affiguratus">
<emphasis id="F0E5EAE8A71E297AFF5C22AEFEB2FF43" box="[151,308,225,251]" italics="true" pageId="3" pageNumber="272">H. affiguratus</emphasis>
</taxonomicName>
has nothing to do with that of
<taxonomicName id="05914D79A71E297AFD4722ADFC9DFF43" authorityName="Fieber" authorityYear="1858" box="[652,795,225,251]" class="Insecta" family="Miridae" genus="Tinicephalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71E297AFD4722ADFC9DFF43" box="[652,795,225,251]" italics="true" pageId="3" pageNumber="272">Tinicephalus</emphasis>
</taxonomicName>
. On the other hand,
<taxonomicName id="05914D79A71E297AFBCB22ADFB1AFF43" authorityName="Fieber" authorityYear="1858" box="[1024,1180,225,251]" class="Insecta" family="Miridae" genus="Hoplomachus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71E297AFBCB22ADFB1AFF43" box="[1024,1180,225,251]" italics="true" pageId="3" pageNumber="272">Hoplomachus</emphasis>
</taxonomicName>
is clearly separated by external morphology from the speciose genus
<taxonomicName id="05914D79A71E297AFD552349FCC7FEA7" authorityName="Fieber" authorityYear="1858" box="[670,833,261,287]" class="Insecta" family="Miridae" genus="Plagiognathus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71E297AFD552349FCC7FEA7" box="[670,833,261,287]" italics="true" pageId="3" pageNumber="272">Plagiognathus</emphasis>
</taxonomicName>
(see, for example,
<bibRefCitation id="A6004B0BA71E297AFBD32349FB27FEA7" author="Schuh, R. T." box="[1048,1185,261,287]" pageId="3" pageNumber="272" pagination="1 - 267" refId="ref4267" refString="Schuh, R. T. (2001) Revision of New World Plagiognathus Fieber, with comments on the Palearctic fauna and the description of a new genus (Heteroptera: Miridae: Phylinae). Bulletin of the American Museum of Natural History, 266, 1 - 267. https: // doi. org / 10.1206 / 0003 - 0090 (2001) 266 &lt;0001: RONWPF&gt; 2.0. CO; 2" type="journal article" year="2001">Schuh 2001</bibRefCitation>
: figs. 514). Concerning the recently erected genus
<taxonomicName id="05914D79A71E297AFE272365FC0BFEFB" authority="Schuh &amp; Salas, 2018" authorityName="Schuh &amp; Salas" authorityYear="2018" box="[492,909,297,323]" class="Insecta" family="Miridae" genus="Henryognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71E297AFE272365FD1FFEFB" box="[492,665,297,323]" italics="true" pageId="3" pageNumber="272">Henryognathus</emphasis>
<bibRefCitation id="A6004B0BA71E297AFD6B2365FC0BFEFB" author="Schuh, R. T. &amp; Salas, R." box="[672,909,297,323]" pageId="3" pageNumber="272" pagination="281 - 289" refId="ref4449" refString="Schuh, R. T. &amp; Salas, R. (2018) Henryognathus thomasi, a new genus and new species of Arctostaphylos - feeding plant bug from western North America (Miridae, Phylinae, Phylini). Zookeys, 796, 281 - 289. https: // doi. org / 10.3897 / zookeys. 796.21432" type="journal article" year="2018">Schuh &amp; Salas, 2018</bibRefCitation>
</taxonomicName>
, it “falls within the diagnosis of
<taxonomicName id="05914D79A71E297AFB322365FA1AFEFB" authorityName="Fieber" authorityYear="1858" box="[1273,1436,297,323]" class="Insecta" family="Miridae" genus="Plagiognathus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71E297AFB322365FA1AFEFB" box="[1273,1436,297,323]" italics="true" pageId="3" pageNumber="272">Plagiognathus</emphasis>
</taxonomicName>
[...], with the exception of the structure of the male genitalia”, as stated by the authors (
<bibRefCitation id="A6004B0BA71E297AFB962301FAC0FEDF" author="Schuh, R. T. &amp; Salas, R." box="[1117,1350,333,359]" pageId="3" pageNumber="272" pagination="281 - 289" refId="ref4449" refString="Schuh, R. T. &amp; Salas, R. (2018) Henryognathus thomasi, a new genus and new species of Arctostaphylos - feeding plant bug from western North America (Miridae, Phylinae, Phylini). Zookeys, 796, 281 - 289. https: // doi. org / 10.3897 / zookeys. 796.21432" type="journal article" year="2018">Schuh &amp; Salas 2018</bibRefCitation>
); again, its vesica is very dissimilar to that of
<taxonomicName id="05914D79A71E297AFDF1233EFD51FE33" authorityName="Pagola-Carte &amp; Matocq" authorityYear="2020" baseAuthorityName="Uhler" baseAuthorityYear="1895" box="[570,727,369,395]" class="Insecta" family="Miridae" genus="Delphiphylus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="species" species="affiguratus">
<emphasis id="F0E5EAE8A71E297AFDF1233EFD51FE33" box="[570,727,369,395]" italics="true" pageId="3" pageNumber="272">H. affiguratus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C22E36FAA71E297AFF0C23D9FCF7FD14" blockId="3.[151,1437,153,1260]" pageId="3" pageNumber="272">
Among the vast diversity of phyline genera, we logically focused on tribes other than
<taxonomicName id="05914D79A71E297AFB5B23D9FA91FE17" authorityName="Van Duzee" authorityYear="1916" box="[1168,1303,405,431]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="tribe" tribe="Hallodapini">Hallodapini</taxonomicName>
,
<taxonomicName id="05914D79A71E297AFAE823D9FF71FE6B" authorityName="Schuh" authorityYear="1974" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="tribe" tribe="Leucophoropterini">Leucophoropterini</taxonomicName>
and
<taxonomicName id="05914D79A71E297AFEF923F5FE35FE6B" authorityName="Douglas &amp; Scott" authorityYear="1876" box="[306,435,441,467]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="tribe" tribe="Pilophorini">Pilophorini</taxonomicName>
, devoting special attention to all explicit mentions to
<taxonomicName id="05914D79A71E297AFBD523F5FB3CFE6B" authorityName="Fieber" authorityYear="1858" box="[1054,1210,441,467]" class="Insecta" family="Miridae" genus="Hoplomachus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71E297AFBD523F5FB3CFE6B" box="[1054,1210,441,467]" italics="true" pageId="3" pageNumber="272">Hoplomachus</emphasis>
</taxonomicName>
throughout the text by
<bibRefCitation id="A6004B0BA71E297AFF712391FE56FE4F" author="Schuh, R. T. &amp; Menard, K. L." box="[186,464,477,503]" pageId="3" pageNumber="272" pagination="1 - 72" refId="ref4392" refString="Schuh, R. T. &amp; Menard, K. L. (2013) A revised classification of the Phylinae (Insecta: Heteroptera: Miridae): arguments for the placement of genera. American Museum Novitates, 3785, 1 - 72. https: // doi. org / 10.1206 / 3785.2" type="journal article" year="2013">Schuh &amp; Menard (2013)</bibRefCitation>
. The Palaearctic genus
<taxonomicName id="05914D79A71E297AFD1C2391FB8AFE4F" authority="Reuter, 1901" authorityName="Reuter" authorityYear="1901" box="[727,1036,477,503]" class="Insecta" family="Miridae" genus="Opisthotaenia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71E297AFD1C2391FCF0FE4F" box="[727,886,477,503]" italics="true" pageId="3" pageNumber="272">Opisthotaenia</emphasis>
Reuter, 1901
</taxonomicName>
is also classified within Oncotylina “because of the similarity of appearance with species of
<taxonomicName id="05914D79A71E297AFCD0204DFC24FDA3" authorityName="Knight" authorityYear="1919" box="[795,930,513,539]" class="Insecta" family="Miridae" genus="Phyllopidea" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71E297AFCD0204DFC24FDA3" box="[795,930,513,539]" italics="true" pageId="3" pageNumber="272">Phyllopidea</emphasis>
</taxonomicName>
[Knight, 1919] from the Nearctic, including similarity of structure of the endosoma, and its resemblance to
<taxonomicName id="05914D79A71E297AFCAB2069FC7AFD87" authorityName="Fieber" authorityYear="1858" box="[864,1020,549,575]" class="Insecta" family="Miridae" genus="Hoplomachus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71E297AFCAB2069FC7AFD87" box="[864,1020,549,575]" italics="true" pageId="3" pageNumber="272">Hoplomachus</emphasis>
</taxonomicName>
and
<taxonomicName id="05914D79A71E297AFBFF2069FB45FD87" authorityName="Puton" authorityYear="1875" box="[1076,1219,549,575]" class="Insecta" family="Miridae" genus="Thermocoris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71E297AFBFF2069FB45FD87" box="[1076,1219,549,575]" italics="true" pageId="3" pageNumber="272">Thermocoris</emphasis>
</taxonomicName>
[Puton, 1875] (
<bibRefCitation id="A6004B0BA71E297AFABD2069FEDFFDDC" author="Seidenstucker, G." pageId="3" pageNumber="272" pagination="31 - 38" refId="ref4510" refString="Seidenstucker, G. (1968) Noch zwei Opisthotaenia - Arten (Heteroptera, Miridae). Reichenbachia, 10 (6), 31 - 38." type="journal article" year="1968">Seidenstücker 1968</bibRefCitation>
) in the Palearctic.” We verified that the male genitalia of
<taxonomicName id="05914D79A71E297AFC232006FB00FDDB" authorityName="Pagola-Carte &amp; Matocq" authorityYear="2020" baseAuthorityName="Uhler" baseAuthorityYear="1895" box="[1000,1158,585,611]" class="Insecta" family="Miridae" genus="Delphiphylus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="species" species="affiguratus">
<emphasis id="F0E5EAE8A71E297AFC232006FB00FDDB" box="[1000,1158,585,611]" italics="true" pageId="3" pageNumber="272">H. affiguratus</emphasis>
</taxonomicName>
are completely different from those of
<taxonomicName id="05914D79A71E297AFEF82021FE54FD3F" authorityName="Reuter" authorityYear="1901" box="[307,466,621,647]" class="Insecta" family="Miridae" genus="Opisthotaenia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71E297AFEF82021FE54FD3F" box="[307,466,621,647]" italics="true" pageId="3" pageNumber="272">Opisthotaenia</emphasis>
</taxonomicName>
and
<taxonomicName id="05914D79A71E297AFDCD2021FD13FD3F" authorityName="Puton" authorityYear="1875" box="[518,661,621,647]" class="Insecta" family="Miridae" genus="Thermocoris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71E297AFDCD2021FD13FD3F" box="[518,661,621,647]" italics="true" pageId="3" pageNumber="272">Thermocoris</emphasis>
</taxonomicName>
(see
<bibRefCitation id="A6004B0BA71E297AFD052021FCE0FD3F" author="Wagner, E." box="[718,870,621,647]" pageId="3" pageNumber="272" pagination="78 - 81" refId="ref4646" refString="Wagner, E. (1966) Ribesophylus nov. gen., eine neue Gattung aus Spanien (Hem. Het. Miridae). Mitteilungen der Deutschen Entomologischen Gesellschaft, 25 (5), 78 - 81." type="journal article" year="1966">Wagner 1966</bibRefCitation>
,
<bibRefCitation id="A6004B0BA71E297AFCBB2020FC2DFD3F" author="Wagner, E." box="[880,939,620,647]" pageId="3" pageNumber="272" pagination="1 - 483" refId="ref4687" refString="Wagner, E. (1975) Die Miridae Hahn, 1831, des Mittelmeerraumes und der Makaronesischen Inseln (Hemiptera, Heteroptera). Teil 3. Entomologische Abhandlungen herausgegeben vom Staatlichen Museum fur Tierkunde in Dresden, 40 (Supplement), 1 - 483." type="journal article" year="1975">1975</bibRefCitation>
,
<bibRefCitation id="A6004B0BA71E297AFC7E2021FB15FD3F" author="Seidenstucker, G." box="[949,1171,621,647]" pageId="3" pageNumber="272" pagination="31 - 38" refId="ref4510" refString="Seidenstucker, G. (1968) Noch zwei Opisthotaenia - Arten (Heteroptera, Miridae). Reichenbachia, 10 (6), 31 - 38." type="journal article" year="1968">Seidenstucker 1968</bibRefCitation>
,
<bibRefCitation id="A6004B0BA71E297AFB562021FAB4FD3F" author="Matocq, A." box="[1181,1330,620,647]" pageId="3" pageNumber="272" pagination="273 - 277" refId="ref4005" refString="Matocq, A. (1985) Un genre et une espece de Miridae nouveaux pour la France (Heteroptera). L'Entomologiste, 41 (6), 273 - 277." type="journal article" year="1985">Matocq 1985</bibRefCitation>
) and also different from those of
<taxonomicName id="05914D79A71E297AFE5720DDFDA5FD13" authorityName="Knight" authorityYear="1919" box="[412,547,657,683]" class="Insecta" family="Miridae" genus="Phyllopidea" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71E297AFE5720DDFDA5FD13" box="[412,547,657,683]" italics="true" pageId="3" pageNumber="272">Phyllopidea</emphasis>
</taxonomicName>
(see
<bibRefCitation id="A6004B0BA71E297AFD9520DDFD61FD13" author="Schuh, R. T." box="[606,743,657,683]" pageId="3" pageNumber="272" pagination="1 - 267" refId="ref4267" refString="Schuh, R. T. (2001) Revision of New World Plagiognathus Fieber, with comments on the Palearctic fauna and the description of a new genus (Heteroptera: Miridae: Phylinae). Bulletin of the American Museum of Natural History, 266, 1 - 267. https: // doi. org / 10.1206 / 0003 - 0090 (2001) 266 &lt;0001: RONWPF&gt; 2.0. CO; 2" type="journal article" year="2001">Schuh 2001</bibRefCitation>
: figs. 1cd).
</paragraph>
<paragraph id="C22E36FAA71E297AFF0C20F9FE24FC57" blockId="3.[151,1437,153,1260]" pageId="3" pageNumber="272">
The western Nearctic species
<taxonomicName id="05914D79A71E297AFDD820F9FC46FD77" authority="(Uhler, 1893)" authorityName="Uhler" authorityYear="1893" baseAuthorityName="Uhler" baseAuthorityYear="1893" box="[531,960,693,719]" class="Insecta" family="Miridae" genus="Hoplomachidea" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="species" species="consors">
<emphasis id="F0E5EAE8A71E297AFDD820F9FCA5FD77" box="[531,803,693,719]" italics="true" pageId="3" pageNumber="272">Hoplomachidea consors</emphasis>
(
<bibRefCitation id="A6004B0BA71E297AFCFA20F9FC3EFD77" author="Uhler, P. R." box="[817,952,693,719]" pageId="3" pageNumber="272" pagination="260 - 265" refId="ref4539" refString="Uhler, P. R. (1893) Hemiptera in report on small collection of insects made during the Death Valley expedition. North American Fauna, 7, 260 - 265. https: // doi. org / 10.3996 / nafa. 7.0005" type="journal article" year="1893">Uhler, 1893</bibRefCitation>
)
</taxonomicName>
was originally described in
<taxonomicName id="05914D79A71E297AFB3120F9FA10FD77" authorityName="Fieber" authorityYear="1858" box="[1274,1430,693,719]" class="Insecta" family="Miridae" genus="Hoplomachus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71E297AFB3120F9FA10FD77" box="[1274,1430,693,719]" italics="true" pageId="3" pageNumber="272">Hoplomachus</emphasis>
</taxonomicName>
.
<bibRefCitation id="A6004B0BA71E297AFF5C2095FEB4FD4B" author="Reuter, O. M." box="[151,306,729,755]" pageId="3" pageNumber="272" pagination="1 - 86" refId="ref4146" refString="Reuter, O. M. (1909) Bemerkungen uber nearktische Capsiden nebst Beschreibung neuer Arten. Acta Societatis Scientiarum Fennicae, 36 (2), 1 - 86, 3 pls. https: // doi. org / 10.5962 / bhl. title. 9445" type="book chapter" year="1909">Reuter (1909)</bibRefCitation>
created the genus
<taxonomicName id="05914D79A71E297AFE352095FD37FD4B" authorityName="Reuter" authorityYear="1909" box="[510,689,729,755]" class="Insecta" family="Miridae" genus="Hoplomachidea" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71E297AFE352095FD37FD4B" box="[510,689,729,755]" italics="true" pageId="3" pageNumber="272">Hoplomachidea</emphasis>
</taxonomicName>
for it, highlighting the main differences, among them the conspicuous pale squamiform setae in
<taxonomicName id="05914D79A71E297AFE2F20B2FDDBFCAF" baseAuthorityName="Uhler" baseAuthorityYear="1893" box="[484,605,766,791]" class="Insecta" family="Miridae" genus="Hoplomachidea" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="species" species="consors">
<emphasis id="F0E5EAE8A71E297AFE2F20B2FDDBFCAF" box="[484,605,766,791]" italics="true" pageId="3" pageNumber="272">H. consors</emphasis>
</taxonomicName>
.
<bibRefCitation id="A6004B0BA71E297AFDA020B1FCFBFCAF" author="Schuh, R. T. &amp; Menard, K. L." box="[619,893,765,791]" pageId="3" pageNumber="272" pagination="1 - 72" refId="ref4392" refString="Schuh, R. T. &amp; Menard, K. L. (2013) A revised classification of the Phylinae (Insecta: Heteroptera: Miridae): arguments for the placement of genera. American Museum Novitates, 3785, 1 - 72. https: // doi. org / 10.1206 / 3785.2" type="journal article" year="2013">Schuh &amp; Menard (2013)</bibRefCitation>
place the genus
<taxonomicName id="05914D79A71E297AFBFC20B1FA06FCAF" authority="Reuter, 1909" authorityName="Reuter" authorityYear="1909" box="[1079,1408,765,791]" class="Insecta" family="Miridae" genus="Hoplomachidea" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71E297AFBFC20B1FB6CFCAF" box="[1079,1258,765,791]" italics="true" pageId="3" pageNumber="272">Hoplomachidea</emphasis>
<bibRefCitation id="A6004B0BA71E297AFB3B20B1FA06FCAF" author="Reuter, O. M." box="[1264,1408,765,791]" pageId="3" pageNumber="272" pagination="1 - 86" refId="ref4146" refString="Reuter, O. M. (1909) Bemerkungen uber nearktische Capsiden nebst Beschreibung neuer Arten. Acta Societatis Scientiarum Fennicae, 36 (2), 1 - 86, 3 pls. https: // doi. org / 10.5962 / bhl. title. 9445" type="book chapter" year="1909">
Reuter,
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</bibRefCitation>
</taxonomicName>
in the subtribe Semiina (tribe Semiini), separating it from
<taxonomicName id="05914D79A71E297AFD36216DFC1FFC83" authorityName="Fieber" authorityYear="1858" box="[765,921,801,827]" class="Insecta" family="Miridae" genus="Hoplomachus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71E297AFD36216DFC1FFC83" box="[765,921,801,827]" italics="true" pageId="3" pageNumber="272">Hoplomachus</emphasis>
</taxonomicName>
and also from the subtribe Cremnorhina (tribe Cremnorhini), with which it shares several characters of external morphology and male genitalia. They conclude that “even though the name and some morphological attributes suggest an affinity with
<taxonomicName id="05914D79A71E297AFB4F2125FAA6FC3B" authorityName="Fieber" authorityYear="1858" box="[1156,1312,873,899]" class="Insecta" family="Miridae" genus="Hoplomachus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71E297AFB4F2125FAA6FC3B" box="[1156,1312,873,899]" italics="true" pageId="3" pageNumber="272">Hoplomachus</emphasis>
</taxonomicName>
, these two taxa are not members of the same clade with the
<taxonomicName id="05914D79A71E297AFD7621C1FCA3FC1F" authorityName="Douglas &amp; Scott" authorityYear="1865" box="[701,805,909,935]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="subFamily" subFamily="Phylinae">Phylinae</taxonomicName>
”. Although we did not find any publication dealing with its genitalia, we also treat
<taxonomicName id="05914D79A71E297AFE0221FDFDFAFC73" authorityName="Reuter" authorityYear="1909" box="[457,636,945,971]" class="Insecta" family="Miridae" genus="Hoplomachidea" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71E297AFE0221FDFDFAFC73" box="[457,636,945,971]" italics="true" pageId="3" pageNumber="272">Hoplomachidea</emphasis>
</taxonomicName>
as a taxon clearly separated from both Palaearctic
<taxonomicName id="05914D79A71E297AFB0421FDFAEDFC73" authorityName="Fieber" authorityYear="1858" box="[1231,1387,945,971]" class="Insecta" family="Miridae" genus="Hoplomachus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71E297AFB0421FDFAEDFC73" box="[1231,1387,945,971]" italics="true" pageId="3" pageNumber="272">Hoplomachus</emphasis>
</taxonomicName>
and Nearctic
<emphasis id="F0E5EAE8A71E297AFF36219AFE24FC57" box="[253,418,981,1007]" italics="true" pageId="3" pageNumber="272">
<taxonomicName id="05914D79A71E297AFF36219AFE18FC57" authorityName="Pagola-Carte &amp; Matocq" authorityYear="2020" baseAuthorityName="Uhler" baseAuthorityYear="1895" box="[253,414,981,1007]" class="Insecta" family="Miridae" genus="Delphiphylus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="species" species="affiguratus">H. affiguratus</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="C22E36FAA71E297AFF0C21B5FE91FB54" blockId="3.[151,1437,153,1260]" pageId="3" pageNumber="272">
Thirdly, we also took into account the host plants of the phyline genera/species. As far as it is known (see
<bibRefCitation id="A6004B0BA71E297AFF5C2651FEA6FB8F" author="Schuh, R. T." box="[151,288,1053,1079]" pageId="3" pageNumber="272" refId="ref4345" refString="Schuh, R. T. (2002 - 2013) On-line systematic catalog of Plant Bugs (Insecta: Heteroptera: Miridae). Available from: http: // research. amnh. org / pbi / catalog / (accessed 13 October 2019)" type="url" year="2002">
Schuh
<date id="B62F103AA71E297AFF2D2651FEA6FB8F" box="[230,288,1053,1079]" pageId="3" pageNumber="272" value="2002" valueMax="2013">2002</date>
</bibRefCitation>
<bibRefCitation id="A6004B0BA71E297AFEE42651FEEEFB8F" author="Schuh, R. T. &amp; Menard, K. L." box="[303,360,1053,1079]" pageId="3" pageNumber="272" pagination="1 - 72" refId="ref4392" refString="Schuh, R. T. &amp; Menard, K. L. (2013) A revised classification of the Phylinae (Insecta: Heteroptera: Miridae): arguments for the placement of genera. American Museum Novitates, 3785, 1 - 72. https: // doi. org / 10.1206 / 3785.2" type="journal article" year="2013">2013</bibRefCitation>
), very few species develop on
<taxonomicName id="05914D79A71E297AFD012651FCF5FB8F" box="[714,883,1053,1079]" class="Magnoliopsida" family="Ranunculaceae" kingdom="Plantae" order="Ranunculales" pageId="3" pageNumber="272" phylum="Tracheophyta" rank="family">Ranunculaceae</taxonomicName>
or even
<taxonomicName id="05914D79A71E297AFC122651FBF6FB8F" box="[985,1136,1053,1079]" class="Magnoliopsida" kingdom="Plantae" order="Ranunculales" pageId="3" pageNumber="272" phylum="Tracheophyta" rank="order">Ranunculales</taxonomicName>
(including, among others, families
<taxonomicName id="05914D79A71E297AFF32260DFE1FFBE3" authorityName="de Jussieu" authorityYear="1789" box="[249,409,1089,1115]" class="Magnoliopsida" family="Berberidaceae" kingdom="Plantae" order="Ranunculales" pageId="3" pageNumber="272" phylum="Tracheophyta" rank="family">Berberidaceae</taxonomicName>
or
<taxonomicName id="05914D79A71E297AFE0B260DFDD8FBE3" box="[448,606,1089,1115]" class="Magnoliopsida" family="Papaveraceae" kingdom="Plantae" order="Ranunculales" pageId="3" pageNumber="272" phylum="Tracheophyta" rank="family">Papaveraceae</taxonomicName>
). Except for
<taxonomicName id="05914D79A71E297AFD25260DFB30FBE3" authority="Knight, 1923" authorityName="Knight" authorityYear="1923" box="[750,1206,1089,1115]" class="Insecta" family="Miridae" genus="Plagiognathus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="F0E5EAE8A71E297AFD25260DFB9CFBE3" box="[750,1050,1089,1115]" italics="true" pageId="3" pageNumber="272">Plagiognathus brevirostris</emphasis>
Knight, 1923
</taxonomicName>
(on
<taxonomicName id="05914D79A71E297AFB21260DFAE0FBE3" box="[1258,1382,1089,1115]" class="Magnoliopsida" family="Ranunculaceae" genus="Thalictrum" kingdom="Plantae" order="Ranunculales" pageId="3" pageNumber="272" phylum="Tracheophyta" rank="genus">
<emphasis id="F0E5EAE8A71E297AFB21260DFAE0FBE3" box="[1258,1382,1089,1115]" italics="true" pageId="3" pageNumber="272">Thalictrum</emphasis>
</taxonomicName>
spp. (
<bibRefCitation id="A6004B0BA71E297AFF542629FEA0FBC7" author="Schuh, R. T." box="[159,294,1125,1151]" pageId="3" pageNumber="272" pagination="1 - 267" refId="ref4267" refString="Schuh, R. T. (2001) Revision of New World Plagiognathus Fieber, with comments on the Palearctic fauna and the description of a new genus (Heteroptera: Miridae: Phylinae). Bulletin of the American Museum of Natural History, 266, 1 - 267. https: // doi. org / 10.1206 / 0003 - 0090 (2001) 266 &lt;0001: RONWPF&gt; 2.0. CO; 2" type="journal article" year="2001">Schuh 2001</bibRefCitation>
)) and
<taxonomicName id="05914D79A71E297AFEA52629FD70FBC7" authority="Knight, 1929" authorityName="Knight" authorityYear="1929" box="[366,758,1125,1151]" class="Insecta" family="Miridae" genus="Plagiognathus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="species" species="luteus">
<emphasis id="F0E5EAE8A71E297AFEA52629FDDAFBC7" box="[366,604,1125,1151]" italics="true" pageId="3" pageNumber="272">Plagiognathus luteus</emphasis>
Knight, 1929
</taxonomicName>
(on
<taxonomicName id="05914D79A71E297AFCE22629FC71FBC7" box="[809,1015,1125,1151]" class="Magnoliopsida" family="Berberidaceae" genus="Berberis" kingdom="Plantae" order="Ranunculales" pageId="3" pageNumber="272" phylum="Tracheophyta" rank="species" species="fremontii">
<emphasis id="F0E5EAE8A71E297AFCE22629FC71FBC7" box="[809,1015,1125,1151]" italics="true" pageId="3" pageNumber="272">Berberis fremontii</emphasis>
</taxonomicName>
and
<taxonomicName id="05914D79A71E297AFBE6262AFA1EFBC7" authority="(Schuh 2001))" authorityName="Schuh" authorityYear="2001" baseAuthorityName="Schuh" baseAuthorityYear="2001" box="[1069,1432,1125,1151]" class="Magnoliopsida" family="Berberidaceae" genus="Berberis" kingdom="Plantae" order="Ranunculales" pageId="3" pageNumber="272" phylum="Tracheophyta" rank="species" species="haematocarpa">
<emphasis id="F0E5EAE8A71E297AFBE6262AFB68FBC7" box="[1069,1262,1125,1151]" italics="true" pageId="3" pageNumber="272">B. haematocarpa</emphasis>
(
<bibRefCitation id="A6004B0BA71E297AFB372629FA00FBC7" author="Schuh, R. T." box="[1276,1414,1125,1151]" pageId="3" pageNumber="272" pagination="1 - 267" refId="ref4267" refString="Schuh, R. T. (2001) Revision of New World Plagiognathus Fieber, with comments on the Palearctic fauna and the description of a new genus (Heteroptera: Miridae: Phylinae). Bulletin of the American Museum of Natural History, 266, 1 - 267. https: // doi. org / 10.1206 / 0003 - 0090 (2001) 266 &lt;0001: RONWPF&gt; 2.0. CO; 2" type="journal article" year="2001">Schuh 2001</bibRefCitation>
))
</taxonomicName>
, the remaining ones are obviously not related with
<taxonomicName id="05914D79A71E297AFD1326C6FCF1FB1B" authorityName="Pagola-Carte &amp; Matocq" authorityYear="2020" baseAuthorityName="Uhler" baseAuthorityYear="1895" box="[728,887,1161,1187]" class="Insecta" family="Miridae" genus="Delphiphylus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="species" species="affiguratus">
<emphasis id="F0E5EAE8A71E297AFD1326C6FCF1FB1B" box="[728,887,1161,1187]" italics="true" pageId="3" pageNumber="272">H. affiguratus</emphasis>
</taxonomicName>
. On the other hand, the Palaearctic members of
<taxonomicName id="05914D79A71E297AFF5C26E1FEB5FB7F" authorityName="Fieber" authorityYear="1858" box="[151,307,1197,1223]" class="Insecta" family="Miridae" genus="Hoplomachus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="F0E5EAE8A71E297AFF5C26E1FEB5FB7F" box="[151,307,1197,1223]" italics="true" pageId="3" pageNumber="272">Hoplomachus</emphasis>
</taxonomicName>
are found on
<taxonomicName id="05914D79A71E297AFE1926E1FDD7FB7F" box="[466,593,1197,1223]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="272" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
. Concerning
<taxonomicName id="05914D79A71E297AFD2C26E1FC7FFB7F" baseAuthorityName="Uhler" baseAuthorityYear="1893" box="[743,1017,1197,1223]" class="Insecta" family="Miridae" genus="Hoplomachidea" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="272" phylum="Arthropoda" rank="species" species="consors">
<emphasis id="F0E5EAE8A71E297AFD2C26E1FC7FFB7F" box="[743,1017,1197,1223]" italics="true" pageId="3" pageNumber="272">Hoplomachidea consors</emphasis>
</taxonomicName>
, it lives on
<taxonomicName id="05914D79A71E297AFB4926E1FE95FB54" class="Magnoliopsida" family="Boraginaceae" genus="Phacelia" kingdom="Plantae" order="Boraginales" pageId="3" pageNumber="272" phylum="Tracheophyta" rank="species" species="crenulata">
<emphasis id="F0E5EAE8A71E297AFB4926E1FADEFB7F" box="[1154,1368,1197,1223]" italics="true" pageId="3" pageNumber="272">Phacelia crenulata</emphasis>
(Boraginaceae)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>