treatments-xml/data/9E/2F/87/9E2F87B8FFA1B8298BFDFC6E9F7AFE39.xml
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<document id="BFCD1AB6BBAB31A615161D77BFC03E8C" ID-CLB-Dataset="25485" ID-DOI="10.11646/zootaxa.4668.2.6" ID-GBIF-Dataset="bf522066-d756-4fe2-be4d-d2b6b170562e" ID-ISSN="1175-5326" ID-Zenodo-Dep="3449341" ID-ZooBank="C8C5FA5C-7704-4336-9E73-1BB94E89249C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1568897552708" checkinUser="plazi" docAuthor="Eagderi, Soheil, Mousavi-Sabet, Hamed &amp; Freyhof, Jörg" docDate="2019" docId="9E2F87B8FFA1B8298BFDFC6E9F7AFE39" docLanguage="en" docName="zootaxa.4668.2.6.pdf" docOrigin="Zootaxa 4668 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraschistura makranensis Eagderi &amp; Mousavi-Sabet &amp; Freyhof 2019, new species" docType="treatment" docVersion="9" lastPageNumber="263" masterDocId="6216FFC0FFA2B82F8B6AFFF39B58FFB2" masterDocTitle="Paraschistura makranensis, a new loach from the Jegin River drainage in southern Iran with comments on P. ilamensis and P. pasatigris (Teleostei: Nemacheilidae)" masterLastPageNumber="270" masterPageNumber="258" pageNumber="261" updateTime="1698755553086" updateUser="ExternalLinkService">
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<mods:title id="F38FADB97E3166DEABFF0EBCFFCA445F">Paraschistura makranensis, a new loach from the Jegin River drainage in southern Iran with comments on P. ilamensis and P. pasatigris (Teleostei: Nemacheilidae)</mods:title>
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<mods:namePart id="28E5CA7C01A0D87CD96C8DAE45EC8F61">Eagderi, Soheil</mods:namePart>
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<mods:namePart id="22D45851CF7041675206524A6E331ED4">Mousavi-Sabet, Hamed</mods:namePart>
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<mods:namePart id="2752F484FDF7A876CC098DEFCF172049">Freyhof, Jörg</mods:namePart>
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<mods:title id="1E6D05E0AF6E34F43DE8D56FC9719171">Zootaxa</mods:title>
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<mods:date id="6EE807907403F2447A23A6D37087FB5B">2019</mods:date>
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<mods:number id="416BA6C3E79DE40325095A430ECAA9A6">2019-09-10</mods:number>
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<mods:number id="B01E56665DE32B3CC3AADD948393BF6B">4668</mods:number>
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<treatment id="9E2F87B8FFA1B8298BFDFC6E9F7AFE39" ID-GBIF-Taxon="159406874" LSID="urn:lsid:plazi:treatment:9E2F87B8FFA1B8298BFDFC6E9F7AFE39" httpUri="http://treatment.plazi.org/id/9E2F87B8FFA1B8298BFDFC6E9F7AFE39" lastPageId="6" lastPageNumber="263" pageId="3" pageNumber="261">
<subSubSection id="5E9C6525FFA1B82C8BFDFC6E99D9FC0A" box="[151,641,925,952]" pageId="3" pageNumber="261" type="nomenclature">
<paragraph id="163936AEFFA1B82C8BFDFC6E99D9FC0A" blockId="3.[151,641,925,988]" box="[151,641,925,952]" pageId="3" pageNumber="261">
<heading id="4D7181C2FFA1B82C8BFDFC6E99D9FC0A" bold="true" box="[151,641,925,952]" fontSize="11" level="1" pageId="3" pageNumber="261" reason="1">
<emphasis id="24F2EABCFFA1B82C8BFDFC6E99D9FC0A" bold="true" box="[151,641,925,952]" pageId="3" pageNumber="261">
<taxonomicName id="D1864D2DFFA1B82C8BFDFC6E9ABBFC0A" authority="Eagderi &amp; Mousavi-Sabet &amp; Freyhof, 2019" authorityName="Eagderi &amp; Mousavi-Sabet &amp; Freyhof" authorityYear="2019" box="[151,483,925,952]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="261" phylum="Chordata" rank="species" species="makranensis" status="sp. nov.">
<emphasis id="24F2EABCFFA1B82C8BFDFC6E9ABBFC0A" bold="true" box="[151,483,925,952]" italics="true" pageId="3" pageNumber="261">Paraschistura makranensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3FC157C7FFA1B82C8A98FC6D99D9FC0A" box="[498,641,926,952]" pageId="3" pageNumber="261" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5E9C6525FFA1B82C8BFDFC319A5FFC6E" box="[151,263,962,988]" pageId="3" pageNumber="261" type="description">
<paragraph id="163936AEFFA1B82C8BFDFC319A5FFC6E" blockId="3.[151,641,925,988]" box="[151,263,962,988]" pageId="3" pageNumber="261">
(
<figureCitation id="8EBD2A2BFFA1B82C8BCAFC319BA7FC6E" box="[160,255,962,988]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStart-6="FIGURE 8" captionStartId-0="5.[151,250,1314,1339]" captionStartId-1="6.[151,250,1978,2003]" captionStartId-2="7.[151,250,1299,1324]" captionStartId-3="9.[151,250,1344,1369]" captionStartId-4="9.[151,250,1889,1914]" captionStartId-5="10.[151,250,1162,1187]" captionStartId-6="10.[151,250,1968,1993]" captionTargetBox-0="[151,1435,309,1289]" captionTargetBox-1="[196,1391,417,1953]" captionTargetBox-2="[177,1411,181,1273]" captionTargetBox-3="[177,1390,193,1302]" captionTargetBox-4="[151,1435,1463,1864]" captionTargetBox-5="[196,1391,181,1136]" captionTargetBox-6="[196,1391,1209,1943]" captionTargetId-0="figure@5.[151,1435,309,1289]" captionTargetId-1="figure@6.[196,1391,417,1953]" captionTargetId-2="figure@7.[177,1411,181,1273]" captionTargetId-3="figure@9.[151,1436,181,1319]" captionTargetId-4="figure@9.[151,1435,1463,1864]" captionTargetId-5="figure@10.[196,1391,181,1136]" captionTargetId-6="figure@10.[196,1391,1209,1943]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="9" captionTargetPageId-4="9" captionTargetPageId-5="10" captionTargetPageId-6="10" captionText-0="FIGURE 2. Paraschistura makranensis, IMNRF-UT-1093-16, holotype, 32.7 mm SL; Iran: Jegin River." captionText-1="FIGURE 3. Paraschistura makranensis, paratypes; a, IMNRF-UT-1093-15, 31.7 mm SL; b, IMNRF-UT-1093-17, 27.0 mm SL. Iran: Jegin River." captionText-2="FIGURE 4. Paraschistura makranensis, VMFC PM-P, paratypes; from above: 35 mm SL; 35 mm SL; 34 mm SL; 34 mm SL; Iran: Jegin River." captionText-3="FIGURE 5. Paraschistura makranensis, VMFC PM-P, paratypes; from above: 35 mm SL; 35 mm SL; 34 mm SL; 34 mm SL; Iran: Jegin River." captionText-4="FIGURE 6. Paraschistura makranensis, VMFC PM-P 35.0 mm SL; Iran: Jegin River." captionText-5="FIGURE 7. Paraschistura makranensis, IMNRF-UT-1093-15, paratype, 31.7 mm SL; Iran: Jegin River." captionText-6="FIGURE 8. Paraschistura makranensis, showing suborbital flap; IMNRF-UT- 1093-15, paratype, 31.7 mm SL; Iran: Jegin River." figureDoi-0="http://doi.org/10.5281/zenodo.3449347" figureDoi-1="http://doi.org/10.5281/zenodo.3449349" figureDoi-2="http://doi.org/10.5281/zenodo.3449353" figureDoi-3="http://doi.org/10.5281/zenodo.3449355" figureDoi-4="http://doi.org/10.5281/zenodo.3449359" figureDoi-5="http://doi.org/10.5281/zenodo.3449361" figureDoi-6="http://doi.org/10.5281/zenodo.3449365" httpUri-0="https://zenodo.org/record/3449347/files/figure.png" httpUri-1="https://zenodo.org/record/3449349/files/figure.png" httpUri-2="https://zenodo.org/record/3449353/files/figure.png" httpUri-3="https://zenodo.org/record/3449355/files/figure.png" httpUri-4="https://zenodo.org/record/3449359/files/figure.png" httpUri-5="https://zenodo.org/record/3449361/files/figure.png" httpUri-6="https://zenodo.org/record/3449365/files/figure.png" pageId="3" pageNumber="261">Fig. 28</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5E9C6525FFA1B82C8BFDFBF99848FB6A" pageId="3" pageNumber="261" type="materials_examined">
<paragraph id="163936AEFFA1B82C8BFDFBF99BF7FBFA" blockId="3.[151,1437,1033,2033]" pageId="3" pageNumber="261">
<materialsCitation id="A6EE3CF3FFA1B82C8BFDFBF99BF7FBFA" ID-GBIF-Occurrence="2416712046" country="Iran" latitude="26.161972" location="Jegin" longLatPrecision="1" longitude="57.89167" municipality="Jegin River" pageId="3" pageNumber="261" specimenCount="1" stateProvince="Hormozgan" typeStatus="holotype">
<emphasis id="24F2EABCFFA1B82C8BFDFBF99A53FB96" bold="true" box="[151,267,1034,1060]" pageId="3" pageNumber="261">
<typeStatus id="C93D880CFFA1B82C8BFDFBF99A5EFB96" box="[151,262,1034,1060]" pageId="3" pageNumber="261" type="holotype">Holotype</typeStatus>
.
</emphasis>
IMNRF-UT-1093-16, 33 mm SL;
<collectingCountry id="6E91763EFFA1B82C89E9FBF999ECFB96" box="[643,692,1034,1060]" name="Iran" pageId="3" pageNumber="261">Iran</collectingCountry>
:
<collectingRegion id="D442F84CFFA1B82C89D7FBF998DBFB96" box="[701,899,1034,1060]" country="Iran" name="Hormozgan" pageId="3" pageNumber="261">Hormuzgan prov.</collectingRegion>
:
<collectingMunicipality id="F65DACD4FFA1B82C88E0FBF99F50FB96" box="[906,1032,1034,1060]" pageId="3" pageNumber="261">Jegin River</collectingMunicipality>
at
<location id="13596075FFA1B82C8F4FFBF99F3BFB96" LSID="urn:lsid:plazi:treatment:9E2F87B8FFA1B8298BFDFC6E9F7AFE39:13596075FFA1B82C8F4FFBF99F3BFB96" box="[1061,1123,1034,1060]" country="Iran" latitude="26.161972" longLatPrecision="1" longitude="57.89167" municipality="Jegin River" name="Jegin" pageId="3" pageNumber="261" stateProvince="Hormozgan">Jegin</location>
,
<geoCoordinate id="73B25069FFA1B82C8F01FBFA9E56FB96" box="[1131,1294,1033,1060]" degrees="26" direction="north" minutes="09" orientation="latitude" pageId="3" pageNumber="261" precision="1" seconds="43.1" value="26.161972">26°0943.1” N</geoCoordinate>
<geoCoordinate id="73B25069FFA1B82C8E78FBFA9BF3FBFA" degrees="57" direction="east" minutes="53" orientation="longitude" pageId="3" pageNumber="261" precision="1" seconds="30.0" value="57.89167">57°5330.0” E</geoCoordinate>
.
</materialsCitation>
</paragraph>
<paragraph id="163936AEFFA1B82C8BADFBA1982BFB22" blockId="3.[151,1437,1033,2033]" pageId="3" pageNumber="261">
<emphasis id="24F2EABCFFA1B82C8BADFBA19A1EFBDE" bold="true" box="[199,326,1106,1132]" pageId="3" pageNumber="261">
<typeStatus id="C93D880CFFA1B82C8BADFBA19A19FBDE" box="[199,321,1106,1132]" pageId="3" pageNumber="261" type="paratype">Paratypes</typeStatus>
.
</emphasis>
IMNRF-UT-1093, 2,
<quantity id="D17E9B4BFFA1B82C892BFBA199E4FBDE" box="[577,700,1106,1132]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" metricValueMax="3.2" metricValueMin="2.7" pageId="3" pageNumber="261" unit="mm" value="29.5" valueMax="32.0" valueMin="27.0">2732 mm</quantity>
SL; IMNRF-UT-2093, 3,
<quantity id="D17E9B4BFFA1B82C888DFBA29F2FFBDE" box="[999,1143,1105,1132]" metricMagnitude="-2" metricUnit="m" metricValue="2.82" metricValueMax="3.04" metricValueMin="2.6" pageId="3" pageNumber="261" unit="mm" value="28.2" valueMax="30.4" valueMin="26.0">2630.4 mm</quantity>
SL;
<materialsCitation id="A6EE3CF3FFA1B82C8FC5FBA19BB6FB22" ID-GBIF-Occurrence="2416712047" collectionCode="FSJF" pageId="3" pageNumber="261" specimenCode="FSJF 3684, 2" specimenCount="1" typeStatus="paratype">
<specimenCode id="46209ED5FFA1B82C8FC5FBA19E74FBD9" box="[1199,1324,1105,1132]" collectionCode="FSJF" pageId="3" pageNumber="261">FSJF 3684</specimenCode>
,
<specimenCode id="46209ED5FFA1B82C8E53FBA19E11FBDE" box="[1337,1353,1106,1132]" collectionCode="FSJF" pageId="3" pageNumber="261">2</specimenCode>
,
<quantity id="D17E9B4BFFA1B82C8E3CFBA19B9BFB22" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="2.7" metricValueMin="2.3" pageId="3" pageNumber="261" unit="mm" value="25.0" valueMax="27.0" valueMin="23.0">2327 mm</quantity>
SL
</materialsCitation>
;
<materialsCitation id="A6EE3CF3FFA1B82C8B93FB859837FB22" ID-GBIF-Occurrence="2416712045" box="[249,879,1142,1168]" collectionCode="VMFC, PM-P" country="Iran" latitude="26.161972" location="Jegin" longLatPrecision="1" longitude="57.89167" municipality="Jegin River" pageId="3" pageNumber="261" specimenCount="1" stateProvince="Hormozgan" typeStatus="paratype">
<collectionCode id="7097AE6BFFA1B82C8B93FB859AC8FB22" box="[249,400,1142,1168]" pageId="3" pageNumber="261">VMFC PM-P</collectionCode>
, 14, 27
<quantity id="D17E9B4BFFA1B82C8A9AFB859966FB22" box="[496,574,1142,1168]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="3" pageNumber="261" unit="mm" value="39.0">39 mm</quantity>
SL; same data as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="163936AEFFA1B82C8BADFB6A9848FB6A" blockId="3.[151,1437,1033,2033]" pageId="3" pageNumber="261">
<emphasis id="24F2EABCFFA1B82C8BADFB6A99FCFB06" bold="true" box="[199,676,1177,1204]" pageId="3" pageNumber="261">Material for molecular genetic analysis.</emphasis>
IMNRF-UT-2093- CO1-3, 3, same data as
<typeStatus id="C93D880CFFA1B82C8FF8FB699FA0FB06" box="[1170,1272,1178,1204]" pageId="3" pageNumber="261" type="holotype">holotype</typeStatus>
; (Genbank accession numbers:
<accessionNumber id="09D5AB4DFFA1B82C8A0AFB4E9ABCFB6A" box="[352,484,1213,1240]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN258033" pageId="3" pageNumber="261">
<collectionCode id="7097AE6BFFA1B82C8A0AFB4E9AD6FB6A" box="[352,398,1213,1240]" country="Brazil" name="Museu Nacional, Universidade Federal do Rio de Janeiro" pageId="3" pageNumber="261">MN</collectionCode>
258033
</accessionNumber>
,
<accessionNumber id="09D5AB4DFFA1B82C8A85FB4E992BFB6A" box="[495,627,1213,1240]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN258034" pageId="3" pageNumber="261">
<collectionCode id="7097AE6BFFA1B82C8A85FB4E9945FB6A" box="[495,541,1213,1240]" country="Brazil" name="Museu Nacional, Universidade Federal do Rio de Janeiro" pageId="3" pageNumber="261">MN</collectionCode>
258034
</accessionNumber>
,
<accessionNumber id="09D5AB4DFFA1B82C8914FB4E985CFB6A" box="[638,772,1213,1240]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN258035" pageId="3" pageNumber="261">
<collectionCode id="7097AE6BFFA1B82C8914FB4E99F5FB6A" box="[638,685,1213,1240]" country="Brazil" name="Museu Nacional, Universidade Federal do Rio de Janeiro" pageId="3" pageNumber="261">MN</collectionCode>
258035
</accessionNumber>
).
</paragraph>
</subSubSection>
<subSubSection id="5E9C6525FFA1B82C8BADFB129852F842" pageId="3" pageNumber="261" type="diagnosis">
<paragraph id="163936AEFFA1B82C8BADFB129918FA3E" blockId="3.[151,1437,1033,2033]" pageId="3" pageNumber="261">
<emphasis id="24F2EABCFFA1B82C8BADFB129A1AFB4E" bold="true" box="[199,322,1249,1276]" pageId="3" pageNumber="261">Diagnosis.</emphasis>
<taxonomicName id="D1864D2DFFA1B82C8A23FB119925FB4E" authorityName="Eagderi &amp; Mousavi-Sabet &amp; Freyhof" authorityYear="2019" box="[329,637,1250,1276]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="261" phylum="Chordata" rank="species" species="makranensis">
<emphasis id="24F2EABCFFA1B82C8A23FB119925FB4E" box="[329,637,1250,1276]" italics="true" pageId="3" pageNumber="261">Paraschistura makranensis</emphasis>
</taxonomicName>
is distinguished from all other
<taxonomicName id="D1864D2DFFA1B82C88B6FB119F23FB4E" authorityName="Prokofiev" authorityYear="2009" box="[988,1147,1250,1276]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="261" phylum="Chordata" rank="genus">
<emphasis id="24F2EABCFFA1B82C88B6FB119F23FB4E" box="[988,1147,1250,1276]" italics="true" pageId="3" pageNumber="261">Paraschistura</emphasis>
</taxonomicName>
species in
<collectingCountry id="6E91763EFFA1B82C8F9DFB119E7CFB4E" box="[1271,1316,1250,1276]" name="Iran" pageId="3" pageNumber="261">Iran</collectingCountry>
by a combination of characters, none of them unique. It is distinguished from its congeners in
<collectingCountry id="6E91763EFFA1B82C8F1DFAF59FFCFA92" box="[1143,1188,1286,1312]" name="Iran" pageId="3" pageNumber="261">Iran</collectingCountry>
except
<emphasis id="24F2EABCFFA1B82C8E6BFAF59A4CFAF1" italics="true" pageId="3" pageNumber="261">
<taxonomicName id="D1864D2DFFA1B82C8E6BFAF59E22FA92" authorityName="Mousavi-Sabet &amp; Eagderi" authorityYear="2015" box="[1281,1402,1286,1312]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="261" phylum="Chordata" rank="species" species="delvarii">P. delvarii</taxonomicName>
,
<taxonomicName id="D1864D2DFFA1B82C8EE2FAF59A4CFAF1" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="261" phylum="Chordata" rank="species" species="turcmenica">P. turcmenica</taxonomicName>
</emphasis>
, and some populations of
<taxonomicName id="D1864D2DFFA1B82C892CFAD9998BFAF1" authorityName="Freyhof, Sayyadzadeh, Esmaeili &amp; Geiger" authorityYear="2015" box="[582,723,1322,1347]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="261" phylum="Chordata" rank="species" species="naumanni">
<emphasis id="24F2EABCFFA1B82C892CFAD9998BFAF1" box="[582,723,1322,1347]" italics="true" pageId="3" pageNumber="261">P. naumanni</emphasis>
</taxonomicName>
by having a plain brown or slightly mottled colour pattern on the flank (vs. flank with bars, at least on the caudal peduncle), and from all species except
<taxonomicName id="D1864D2DFFA1B82C8FF4FABD9E48FAD5" baseAuthorityName="Berg" baseAuthorityYear="1898" box="[1182,1296,1358,1383]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="261" phylum="Chordata" rank="species" species="cristata">
<emphasis id="24F2EABCFFA1B82C8FF4FABD9E48FAD5" box="[1182,1296,1358,1383]" italics="true" pageId="3" pageNumber="261">P. cristata</emphasis>
</taxonomicName>
by having a complete (vs. incomplete) lateral line.
</paragraph>
<paragraph id="163936AEFFA1B82C8BADFA659919F9D6" blockId="3.[151,1437,1033,2033]" pageId="3" pageNumber="261">
It is further distinguished from
<taxonomicName id="D1864D2DFFA1B82C8942FA65998AFA02" baseAuthorityName="Nikolskii" baseAuthorityYear="1900" box="[552,722,1430,1456]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="261" phylum="Chordata" rank="species" species="bampurensis">
<emphasis id="24F2EABCFFA1B82C8942FA65998AFA02" box="[552,722,1430,1456]" italics="true" pageId="3" pageNumber="261">P. bampurensis</emphasis>
</taxonomicName>
and
<taxonomicName id="D1864D2DFFA1B82C8863FA659823FA1D" baseAuthorityName="Berg" baseAuthorityYear="1898" box="[777,891,1430,1455]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="261" phylum="Chordata" rank="species" species="cristata">
<emphasis id="24F2EABCFFA1B82C8863FA659823FA1D" box="[777,891,1430,1455]" italics="true" pageId="3" pageNumber="261">P. cristata</emphasis>
</taxonomicName>
by having 7½ branched dorsal-fin rays (vs. usually 8½, rarely 7½ or 9½), and a shorter caudal fin (1416% SL vs. 1719% SL); from
<taxonomicName id="D1864D2DFFA1B82C8FFAFA499F81FA66" baseAuthorityName="Nalbant &amp; Bianco" baseAuthorityYear="1998" box="[1168,1241,1466,1492]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="261" phylum="Chordata" rank="species" species="alta">
<emphasis id="24F2EABCFFA1B82C8FFAFA499F81FA66" box="[1168,1241,1466,1492]" italics="true" pageId="3" pageNumber="261">P. alta</emphasis>
</taxonomicName>
by presence (vs. absence) of a small and pointed processus dentiformis in the upper jaw, and an emarginate (vs. deeply forked) caudal fin. It is further distinguished from
<taxonomicName id="D1864D2DFFA1B82C8939F9F1999DF9A9" baseAuthorityName="Berg" baseAuthorityYear="1898" box="[595,709,1538,1563]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="261" phylum="Chordata" rank="species" species="cristata">
<emphasis id="24F2EABCFFA1B82C8939F9F1999DF9A9" box="[595,709,1538,1563]" italics="true" pageId="3" pageNumber="261">P. cristata</emphasis>
</taxonomicName>
by having a shallow dorsal adipose keel on the caudal peduncle without procurrent rays (vs. a prominent dorsal adipose crest supported by 2225 procurrent rays) and presence (vs. absence) of a suborbital flap in males.
</paragraph>
<paragraph id="163936AEFFA1B82C8BADF99D9AA2F946" blockId="3.[151,1437,1033,2033]" pageId="3" pageNumber="261">
The new species is further distinguished from
<taxonomicName id="D1864D2DFFA1B82C89A7F99D9818F93A" authorityName="Mousavi-Sabet &amp; Eagderi" authorityYear="2015" box="[717,832,1646,1672]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="261" phylum="Chordata" rank="species" species="delvarii">
<emphasis id="24F2EABCFFA1B82C89A7F99D9818F93A" box="[717,832,1646,1672]" italics="true" pageId="3" pageNumber="261">P. delvarii</emphasis>
</taxonomicName>
by presence (vs. absence) of a shallow dorsal adipose keel on the caudal peduncle, and absence (vs. presence) of flank scales anterior to the dorsal-fin origin. It is further distinguished from
<emphasis id="24F2EABCFFA1B82C8A1BF9459936F97D" box="[369,622,1718,1744]" italics="true" pageId="3" pageNumber="261">
<taxonomicName id="D1864D2DFFA1B82C8A1BF9459AA0F962" baseAuthorityName="Vatandoust &amp; Eagderi" baseAuthorityYear="2015" box="[369,504,1718,1744]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="261" phylum="Chordata" rank="species" species="ilamensis">P. ilamensis</taxonomicName>
,
<taxonomicName id="D1864D2DFFA1B82C896EF9459936F97D" authorityName="Freyhof, Sayyadzadeh, Esmaeili &amp; Geiger" authorityYear="2015" box="[516,622,1718,1743]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="261" phylum="Chordata" rank="species" species="susiani">P. susiani</taxonomicName>
</emphasis>
and
<taxonomicName id="D1864D2DFFA1B82C89CFF9459840F962" baseAuthorityName="Nalbant &amp; Bianco" baseAuthorityYear="1998" box="[677,792,1718,1744]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="261" phylum="Chordata" rank="species" species="nielseni">
<emphasis id="24F2EABCFFA1B82C89CFF9459840F962" box="[677,792,1718,1744]" italics="true" pageId="3" pageNumber="261">P. nielseni</emphasis>
</taxonomicName>
by absence (vs. presence) of scales on the back and flank anterior to the dorsal-fin origin.
</paragraph>
<paragraph id="163936AEFFA1B82C8BADF90D9852F842" blockId="3.[151,1437,1033,2033]" pageId="3" pageNumber="261">
<taxonomicName id="D1864D2DFFA1B82C8BADF90D9AA4F8AA" authorityName="Eagderi &amp; Mousavi-Sabet &amp; Freyhof" authorityYear="2019" box="[199,508,1790,1816]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="261" phylum="Chordata" rank="species" species="makranensis">
<emphasis id="24F2EABCFFA1B82C8BADF90D9AA4F8AA" box="[199,508,1790,1816]" italics="true" pageId="3" pageNumber="261">Paraschistura makranensis</emphasis>
</taxonomicName>
is further distinguished from
<taxonomicName id="D1864D2DFFA1B82C883BF90D9886F8A5" authorityName="Freyhof, Sayyadzadeh, Esmaeili &amp; Geiger" authorityYear="2015" box="[849,990,1790,1815]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="261" phylum="Chordata" rank="species" species="naumanni">
<emphasis id="24F2EABCFFA1B82C883BF90D9886F8A5" box="[849,990,1790,1815]" italics="true" pageId="3" pageNumber="261">P. naumanni</emphasis>
</taxonomicName>
by having the pelvic-fin origin located posterior to the dorsal-fin origin (vs. pelvic-fin origin situated below or slightly anterior to the vertical through the dorsal-fin origin), and absence (vs. presence) of scales on the back and flank anterior to the dorsal-fin origin. It is further distinguished from
<taxonomicName id="D1864D2DFFA1B82C8AABF899993FF836" authorityName="Sayyadzadeh, Teimori &amp; Esmaeili" authorityYear="2019" box="[449,615,1898,1924]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="261" phylum="Chordata" rank="species" species="kermanensis">
<emphasis id="24F2EABCFFA1B82C8AABF899993FF836" box="[449,615,1898,1924]" italics="true" pageId="3" pageNumber="261">P. kermanensis</emphasis>
</taxonomicName>
by presence (vs. absence) of a suborbital flap in males and having the pelvic-fin origin situated posterior to the vertical through the dorsal-fin origin (vs. in anterior to or below). It is further distinguished from
<emphasis id="24F2EABCFFA1B82C8A1AF84199D9F879" box="[368,641,1970,1996]" italics="true" pageId="3" pageNumber="261">
<taxonomicName id="D1864D2DFFA1B82C8A1AF8419ABBF87E" baseAuthorityName="Günther" baseAuthorityYear="1889" box="[368,483,1970,1996]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="261" phylum="Chordata" rank="species" species="kessleri">P. kessleri</taxonomicName>
,
<taxonomicName id="D1864D2DFFA1B82C8A84F84199D9F879" baseAuthorityName="Nikolskii" baseAuthorityYear="1947" box="[494,641,1970,1995]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="261" phylum="Chordata" rank="species" species="turcomana">P. turcomana</taxonomicName>
</emphasis>
and
<taxonomicName id="D1864D2DFFA1B82C89DDF8419817F879" box="[695,847,1970,1995]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="261" phylum="Chordata" rank="species" species="turcmenica">
<emphasis id="24F2EABCFFA1B82C89DDF8419817F879" box="[695,847,1970,1995]" italics="true" pageId="3" pageNumber="261">P. turcmenica</emphasis>
</taxonomicName>
by presence (vs. absence) of caudal peduncle scales, and presence (vs. absence) of a suborbital flap in males.
</paragraph>
</subSubSection>
<caption id="42F96626FFA6B82B8BFDFA5D99D7F9BD" ID-DOI="http://doi.org/10.5281/zenodo.3449345" ID-Zenodo-Dep="3449345" httpUri="https://zenodo.org/record/3449345/files/figure.png" pageId="4" pageNumber="262" startId="4.[151,250,1454,1479]" targetBox="[275,1298,207,1420]" targetPageId="4">
<paragraph id="163936AEFFA6B82B8BFDFA5D99D7F9BD" blockId="4.[151,1437,1454,1551]" pageId="4" pageNumber="262">
<emphasis id="24F2EABCFFA6B82B8BFDFA5D9A55FA75" bold="true" box="[151,269,1454,1479]" pageId="4" pageNumber="262">FIGURE 1</emphasis>
. Maximum likelihood estimation of the phylogenetic relationships based on the mitochondrial COI barcode region. Nucleotide positions with less than 95% site coverage were eliminated, resulting in 652 analysed positions. Values at nodes correspond to BI posterior probability/ML bootstrap.
</paragraph>
</caption>
<subSubSection id="5E9C6525FFA6B82A8BADF9CE9F80F959" lastPageId="5" lastPageNumber="263" pageId="4" pageNumber="262" type="description">
<paragraph id="163936AEFFA6B82B8BADF9CE9ADEF82E" blockId="4.[151,1436,1597,2020]" pageId="4" pageNumber="262">
<emphasis id="24F2EABCFFA6B82B8BADF9CE9A01F9EA" bold="true" box="[199,345,1597,1624]" pageId="4" pageNumber="262">Description.</emphasis>
For general appearance, see
<figureCitation id="8EBD2A2BFFA6B82B89CBF9CE9872F9E5" box="[673,810,1597,1623]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStartId-0="5.[151,250,1314,1339]" captionStartId-1="6.[151,250,1978,2003]" captionStartId-2="7.[151,250,1299,1324]" captionStartId-3="9.[151,250,1344,1369]" captionStartId-4="9.[151,250,1889,1914]" captionTargetBox-0="[151,1435,309,1289]" captionTargetBox-1="[196,1391,417,1953]" captionTargetBox-2="[177,1411,181,1273]" captionTargetBox-3="[177,1390,193,1302]" captionTargetBox-4="[151,1435,1463,1864]" captionTargetId-0="figure@5.[151,1435,309,1289]" captionTargetId-1="figure@6.[196,1391,417,1953]" captionTargetId-2="figure@7.[177,1411,181,1273]" captionTargetId-3="figure@9.[151,1436,181,1319]" captionTargetId-4="figure@9.[151,1435,1463,1864]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="9" captionTargetPageId-4="9" captionText-0="FIGURE 2. Paraschistura makranensis, IMNRF-UT-1093-16, holotype, 32.7 mm SL; Iran: Jegin River." captionText-1="FIGURE 3. Paraschistura makranensis, paratypes; a, IMNRF-UT-1093-15, 31.7 mm SL; b, IMNRF-UT-1093-17, 27.0 mm SL. Iran: Jegin River." captionText-2="FIGURE 4. Paraschistura makranensis, VMFC PM-P, paratypes; from above: 35 mm SL; 35 mm SL; 34 mm SL; 34 mm SL; Iran: Jegin River." captionText-3="FIGURE 5. Paraschistura makranensis, VMFC PM-P, paratypes; from above: 35 mm SL; 35 mm SL; 34 mm SL; 34 mm SL; Iran: Jegin River." captionText-4="FIGURE 6. Paraschistura makranensis, VMFC PM-P 35.0 mm SL; Iran: Jegin River." figureDoi-0="http://doi.org/10.5281/zenodo.3449347" figureDoi-1="http://doi.org/10.5281/zenodo.3449349" figureDoi-2="http://doi.org/10.5281/zenodo.3449353" figureDoi-3="http://doi.org/10.5281/zenodo.3449355" figureDoi-4="http://doi.org/10.5281/zenodo.3449359" httpUri-0="https://zenodo.org/record/3449347/files/figure.png" httpUri-1="https://zenodo.org/record/3449349/files/figure.png" httpUri-2="https://zenodo.org/record/3449353/files/figure.png" httpUri-3="https://zenodo.org/record/3449355/files/figure.png" httpUri-4="https://zenodo.org/record/3449359/files/figure.png" pageId="4" pageNumber="262">Figures 26</figureCitation>
. Morphometric data are provided in
<tableCitation id="5B040315FFA6B82B8FA2F9CE9E46F9EA" box="[1224,1310,1597,1624]" captionStart="TABLE 3" captionStartId="8.[151,244,646,673]" captionTargetBox="[163,1364,740,2024]" captionText="TABLE 3. Morphometric data of Paraschistura makranensis: holotype (IMNRF-UT-1093-16) and paratypes (IM- NRF-UT-1093, n=3, IMNRF-UT-2093, n=4). Holotype is not included into calculations of ranges, means and SD." httpUri="http://table.plazi.org/id/42F96626FFAAB8278BFDFD759ECBFD77" pageId="4" pageNumber="262" tableUuid="42F96626FFAAB8278BFDFD759ECBFD77">Table 3</tableCitation>
. Small and slender species with relatively short head. Body deepest midway between nape and dorsal fin origin, depth decreasing slowly towards caudal-fin base. Greatest body width at centre of pectoral-fin base. Section of head roundish, flattened on ventral face. Caudal peduncle compressed laterally, 1.21.4 (mean 1.3) times longer than deep. Pectoral fin reaching approximately 5065% of distance from pectoral-fin origin to pelvic-fin origin. Pelvic axillary lobe absent, or if present fully attached to body. Pelvic-fin origin below second or third branched dorsal-fin ray. Pelvic fin not reaching to anus. Anal-fin origin about one eye diameter posterior to anus. Anal-fin origin at vertical through mid-distance between dorsal- and caudal-fin origins. A shallow dorsal adipose keel without procurrent rays on caudal peduncle. Margin of dorsal fin straight or slightly convex. Caudal fin deeply emarginated. Largest known specimen
<quantity id="D17E9B4BFFA6B82B8A62F8719A0FF82E" box="[264,343,1922,1948]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="262" unit="mm" value="40.0">40 mm</quantity>
SL.
</paragraph>
<paragraph id="163936AEFFA6B82A8BADF8569A19FEAD" blockId="4.[151,1436,1597,2020]" lastBlockId="5.[151,1437,153,287]" lastPageId="5" lastPageNumber="263" pageId="4" pageNumber="262">Dorsal fin with 7½ and anal fin with 5½ branched rays. Caudal fin with 8+8 branched rays. Pectoral fin with 910 (usually 9) and pelvic fin with 67 branched rays. Scales absent on back and flank anterior to dorsal-fin ori- gin. Only small, isolated, deeply-embedded scales on caudal peduncle. Lateral line complete, reaching to or almost touching caudal-fin base. Anterior nostril opening at tip of a pointed and flap-like tube. Nostrils separated by a narrow space, posterior tip of anterior nostril not overlapping posterior nostril when folded backwards. Male with a suborbital flap.</paragraph>
<caption id="42F96626FFA7B82A8BFDFAD19FE7FA89" ID-DOI="http://doi.org/10.5281/zenodo.3449347" ID-Zenodo-Dep="3449347" box="[151,1215,1314,1339]" httpUri="https://zenodo.org/record/3449347/files/figure.png" pageId="5" pageNumber="263" startId="5.[151,250,1314,1339]" targetBox="[151,1435,309,1289]" targetPageId="5">
<paragraph id="163936AEFFA7B82A8BFDFAD19FE7FA89" blockId="5.[151,1215,1314,1339]" box="[151,1215,1314,1339]" pageId="5" pageNumber="263">
<emphasis id="24F2EABCFFA7B82A8BFDFAD19A4BFA89" bold="true" box="[151,275,1314,1339]" pageId="5" pageNumber="263">FIGURE 2.</emphasis>
<taxonomicName id="D1864D2DFFA7B82A8A73FAD19977FA88" authorityName="Eagderi &amp; Mousavi-Sabet &amp; Freyhof" authorityYear="2019" box="[281,559,1314,1338]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="263" phylum="Chordata" rank="species" species="makranensis">
<emphasis id="24F2EABCFFA7B82A8A73FAD19977FA88" box="[281,559,1314,1338]" italics="true" pageId="5" pageNumber="263">Paraschistura makranensis</emphasis>
</taxonomicName>
, IMNRF-UT-1093-16, holotype, 32.7 mm SL; Iran: Jegin River.
</paragraph>
</caption>
<paragraph id="163936AEFFA7B82A8BADFA9A9AB8F9A1" blockId="5.[151,1437,1385,2023]" pageId="5" pageNumber="263">
Mouth small, strongly arched (
<figureCitation id="8EBD2A2BFFA7B82A8949FA9A9930FA31" box="[547,616,1385,1411]" captionStart="FIGURE 7" captionStartId="10.[151,250,1162,1187]" captionTargetBox="[196,1391,181,1136]" captionTargetId="figure@10.[196,1391,181,1136]" captionTargetPageId="10" captionText="FIGURE 7. Paraschistura makranensis, IMNRF-UT-1093-15, paratype, 31.7 mm SL; Iran: Jegin River." figureDoi="http://doi.org/10.5281/zenodo.3449361" httpUri="https://zenodo.org/record/3449361/files/figure.png" pageId="5" pageNumber="263">Fig. 7</figureCitation>
). Lips moderately thick, with many deep furrows. A median interruption in lower lip. Upper lip with median incision. Processus dentiformis small and pointed. A shallow median notch in lower jaw. Barbels short, inner rostral barbel not reaching base of maxillary barbell, outer one reaching to or slightly beyond base of maxillary barbel, reaching vertical through anterior nostril. Maxillary barbel reaching vertical through distal part of eye.
</paragraph>
<paragraph id="163936AEFFA7B82A8BADF9EF9F80F959" blockId="5.[151,1437,1385,2023]" pageId="5" pageNumber="263">
<emphasis id="24F2EABCFFA7B82A8BADF9EF9A17F985" bold="true" box="[199,335,1564,1591]" pageId="5" pageNumber="263">Coloration.</emphasis>
Body plain grey or pale brown, slightly mottled with yellow belly in life (
<figureCitation id="8EBD2A2BFFA7B82A8FDBF9EE9FA0F98A" box="[1201,1272,1565,1592]" captionStart="FIGURE 6" captionStartId="9.[151,250,1889,1914]" captionTargetBox="[151,1435,1463,1864]" captionTargetId="figure@9.[151,1435,1463,1864]" captionTargetPageId="9" captionText="FIGURE 6. Paraschistura makranensis, VMFC PM-P 35.0 mm SL; Iran: Jegin River." figureDoi="http://doi.org/10.5281/zenodo.3449359" httpUri="https://zenodo.org/record/3449359/files/figure.png" pageId="5" pageNumber="263">Fig. 6</figureCitation>
), yellowish in preserved individuals (
<figureCitation id="8EBD2A2BFFA7B82A8AF3F9B29950F9E9" box="[409,520,1600,1627]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="7.[151,250,1299,1324]" captionStartId-1="9.[151,250,1344,1369]" captionTargetBox-0="[177,1411,181,1273]" captionTargetBox-1="[177,1390,193,1302]" captionTargetId-0="figure@7.[177,1411,181,1273]" captionTargetId-1="figure@9.[151,1436,181,1319]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionText-0="FIGURE 4. Paraschistura makranensis, VMFC PM-P, paratypes; from above: 35 mm SL; 35 mm SL; 34 mm SL; 34 mm SL; Iran: Jegin River." captionText-1="FIGURE 5. Paraschistura makranensis, VMFC PM-P, paratypes; from above: 35 mm SL; 35 mm SL; 34 mm SL; 34 mm SL; Iran: Jegin River." figureDoi-0="http://doi.org/10.5281/zenodo.3449353" figureDoi-1="http://doi.org/10.5281/zenodo.3449355" httpUri-0="https://zenodo.org/record/3449353/files/figure.png" httpUri-1="https://zenodo.org/record/3449355/files/figure.png" pageId="5" pageNumber="263">Figs. 45</figureCitation>
). Head dark-grey on top and side, cheek silvery cream. A prominent, irregularlyshaped, dark brown blotch at posterior extremity of caudal peduncle, notched or interrupted by 12 white streaks shortly above mid-height in some individuals. Dorsal and caudal fins hyaline, with elongated spots on rays. A prominent, usually large black, spot at base of unbranched dorsal-fin rays. Dorsal and caudal fins hyaline with faded black spots and stripes on rays, absent in some individuals. Pectoral, pelvic and anal fins hyaline.
</paragraph>
</subSubSection>
<subSubSection id="5E9C6525FFA7B82A8BADF90699FCF881" pageId="5" pageNumber="263" type="etymology">
<paragraph id="163936AEFFA7B82A8BADF90699FCF881" blockId="5.[151,1437,1385,2023]" pageId="5" pageNumber="263">
<emphasis id="24F2EABCFFA7B82A8BADF9069A16F8BD" bold="true" box="[199,334,1781,1807]" pageId="5" pageNumber="263">Etymology.</emphasis>
The species name
<taxonomicName id="D1864D2DFFA7B82A8945F90699E5F8BD" authorityName="Eagderi &amp; Mousavi-Sabet &amp; Freyhof" authorityYear="2019" box="[559,701,1781,1807]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="263" phylum="Chordata" rank="species" species="makranensis">
<emphasis id="24F2EABCFFA7B82A8945F90699E5F8BD" box="[559,701,1781,1807]" italics="true" pageId="5" pageNumber="263">makranensis</emphasis>
</taxonomicName>
refers to Makran, an ancient Persian word referencing the area along the coast of the
<collectingCountry id="6E91763EFFA7B82A8AE4F8EA9A8AF881" box="[398,466,1817,1843]" name="Oman" pageId="5" pageNumber="263">Oman</collectingCountry>
Sea. An adjective.
</paragraph>
</subSubSection>
<subSubSection id="5E9C6525FFA7B82A8BADF8CF9E5DF8E5" box="[199,1285,1852,1879]" pageId="5" pageNumber="263" type="distribution">
<paragraph id="163936AEFFA7B82A8BADF8CF9E5DF8E5" blockId="5.[151,1437,1385,2023]" box="[199,1285,1852,1879]" pageId="5" pageNumber="263">
<emphasis id="24F2EABCFFA7B82A8BADF8CF9A39F8E5" bold="true" box="[199,353,1852,1879]" pageId="5" pageNumber="263">Distribution.</emphasis>
<taxonomicName id="D1864D2DFFA7B82A8A0DF8CE99C3F8E5" authorityName="Eagderi &amp; Mousavi-Sabet &amp; Freyhof" authorityYear="2019" box="[359,667,1853,1879]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="263" phylum="Chordata" rank="species" species="makranensis">
<emphasis id="24F2EABCFFA7B82A8A0DF8CE99C3F8E5" box="[359,667,1853,1879]" italics="true" pageId="5" pageNumber="263">Paraschistura makranensis</emphasis>
</taxonomicName>
is known from the upper Jegin River drainage (
<figureCitation id="8EBD2A2BFFA7B82A8FD9F8CE9FAFF8E5" box="[1203,1271,1853,1879]" captionStart="FIGURE 9" captionStartId="11.[151,250,779,804]" captionTargetBox="[151,1435,181,753]" captionTargetId="figure@11.[151,1435,181,753]" captionTargetPageId="11" captionText="FIGURE 9. Type locality of Paraschistura makranensis: Jegin River near Jegin, Iran." figureDoi="http://doi.org/10.5281/zenodo.3449369" httpUri="https://zenodo.org/record/3449369/files/figure.png" pageId="5" pageNumber="263">Fig. 9</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5E9C6525FFA7B8298BADF8929F7AFE39" lastPageId="6" lastPageNumber="264" pageId="5" pageNumber="263" type="discussion">
<paragraph id="163936AEFFA7B8298BADF8929F7AFE39" blockId="5.[151,1437,1385,2023]" lastBlockId="6.[151,1437,153,395]" lastPageId="6" lastPageNumber="264" pageId="5" pageNumber="263">
<emphasis id="24F2EABCFFA7B82A8BADF8929A63F8C9" bold="true" box="[199,315,1889,1915]" pageId="5" pageNumber="263">Remarks.</emphasis>
The descriptions of
<taxonomicName id="D1864D2DFFA7B82A8974F89198CEF8C9" authority="(Freyhof et al. 2015)" authorityName="Freyhof" authorityYear="2015" baseAuthorityName="Freyhof" baseAuthorityYear="2015" box="[542,918,1888,1915]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="263" phylum="Chordata" rank="species" species="pasatigris">
<emphasis id="24F2EABCFFA7B82A8974F89199F1F8C9" box="[542,681,1890,1915]" italics="true" pageId="5" pageNumber="263">P. pasatigris</emphasis>
(
<bibRefCitation id="72174B5FFFA7B82A89DCF89298D7F8C9" author="Freyhof, J. &amp; Sayyadzadeh, G. &amp; Esmaeili, H. R. &amp; Geiger, M." box="[694,911,1888,1915]" pageId="5" pageNumber="263" pagination="1 - 48" refId="ref5479" refString="Freyhof, J., Sayyadzadeh, G., Esmaeili, H. R. &amp; Geiger, M. (2015) Review of the genus Paraschistura from Iran with description of six new species (Teleostei: Nemacheilidae). Ichthyological Exploration of Freshwaters, 26 (1), 1 - 48." type="journal article" year="2015">
Freyhof
<emphasis id="24F2EABCFFA7B82A887DF8919810F8C9" box="[791,840,1889,1915]" italics="true" pageId="5" pageNumber="263">et al</emphasis>
. 2015
</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="D1864D2DFFA7B82A88A1F8919EC4F8C9" authority="(Vatandoust &amp; Eagderi 2015)" authorityName="Vatandoust &amp; Eagderi" authorityYear="2015" baseAuthorityName="Vatandoust &amp; Eagderi" baseAuthorityYear="2015" box="[971,1436,1888,1915]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="263" phylum="Chordata" rank="species" species="ilamensis">
<emphasis id="24F2EABCFFA7B82A88A1F8919F08F8C9" box="[971,1104,1889,1915]" italics="true" pageId="5" pageNumber="263">P. ilamensis</emphasis>
(
<bibRefCitation id="72174B5FFFA7B82A8F37F8929ECDF8C9" author="Vatandoust, S. &amp; Eagderi, S." box="[1117,1429,1888,1915]" pageId="5" pageNumber="263" pagination="177 - 182" refId="ref6521" refString="Vatandoust, S. &amp; Eagderi, S. (2015) Paraschistura ilamensis, a new species of loach from the Tigris River drainage (Teleostei: Nemacheilidae). International Journal of Aquatic Biology, 3 (3), 177 - 182." type="journal article" year="2015">Vatandoust &amp; Eagderi 2015</bibRefCitation>
)
</taxonomicName>
were published almost simultaneously and both species occur in the same watershed. We were able to compare materials from both
<typeStatus id="C93D880CFFA7B82A8A17F85A9AF5F871" box="[381,429,1961,1987]" pageId="5" pageNumber="263">type</typeStatus>
localities as well as molecular characters (COI) of both species and found no differences between them. We therefore conclude that these two names represent a single species. To clarify the synonymy, the Principle of Priority (ICZN Article 23) is applied (
<bibRefCitation id="72174B5FFFA4B82989BFFF6A9805FF01" author="ICZN &amp; International Code of Zoological Nomenclature" box="[725,861,153,179]" pageId="6" pageNumber="264" refId="ref5536" refString="ICZN (International Code of Zoological Nomenclature) (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomeclature, London, 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
).
<taxonomicName id="D1864D2DFFA4B8298818FF6A9FDBFF01" baseAuthorityName="Vatandoust &amp; Eagderi" baseAuthorityYear="2015" box="[882,1155,153,179]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="264" phylum="Chordata" rank="species" species="ilamensis">
<emphasis id="24F2EABCFFA4B8298818FF6A9FDBFF01" box="[882,1155,153,179]" italics="true" pageId="6" pageNumber="264">Paraschistura ilamensis</emphasis>
</taxonomicName>
became available online on
<date id="6238106EFFA4B8298BD7FF4F9A0FFF65" box="[189,343,188,215]" pageId="6" pageNumber="264" value="2015-06-25">25 June 2015</date>
with no mention of its online archiving in ZooBank, i.e., the online publication does not meet the criteria for archiving on ZooBank (despite it being archived on
<date id="6238106EFFA4B82988CBFF139F63FF49" box="[929,1083,224,251]" pageId="6" pageNumber="264" value="2015-06-25">
25 June
<quantity id="D17E9B4BFFA4B8298F69FF139F63FF49" box="[1027,1083,224,251]" metricMagnitude="1" metricUnit="m" metricValue="5.1181" pageId="6" pageNumber="264" unit="in" value="2015.0">2015</quantity>
</date>
in Zenodo; https://zenodo.org/ record/845474#.XJ4EzdgyXIU). The description itself was printed and distributed on
<date id="6238106EFFA4B8298F0DFEF69E40FEAD" box="[1127,1304,260,287]" pageId="6" pageNumber="264" value="2015-06-30">30th June 2015</date>
, thus
<emphasis id="24F2EABCFFA4B8298E31FEF59A36FEF1" italics="true" pageId="6" pageNumber="264">
Para- schistura
<taxonomicName id="D1864D2DFFA4B8298A6EFEDA9A36FEF1" baseAuthorityName="Vatandoust &amp; Eagderi" baseAuthorityYear="2015" box="[260,366,297,323]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="264" phylum="Chordata" rank="species" species="ilamensis">ilamensis</taxonomicName>
</emphasis>
is available only from this printed description (
<bibRefCitation id="72174B5FFFA4B8298812FEDA98A6FEF1" author="ICZN &amp; International Code of Zoological Nomenclature" box="[888,1022,297,323]" pageId="6" pageNumber="264" refId="ref5536" refString="ICZN (International Code of Zoological Nomenclature) (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomeclature, London, 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
). The description of
<taxonomicName id="D1864D2DFFA4B8298F8AFED99E33FEF1" box="[1248,1387,298,323]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="264" phylum="Chordata" rank="species" species="pasatigris">
<emphasis id="24F2EABCFFA4B8298F8AFED99E33FEF1" box="[1248,1387,298,323]" italics="true" pageId="6" pageNumber="264">P. pasatigris</emphasis>
</taxonomicName>
was printed and distributed on
<date id="6238106EFFA4B8298AABFEBE9939FED5" box="[449,609,332,359]" pageId="6" pageNumber="264" value="2015-07-02">2nd July 2015</date>
(F. Pfeil, pers. com), therefore
<taxonomicName id="D1864D2DFFA4B82988AEFEBD9F12FED5" baseAuthorityName="Vatandoust &amp; Eagderi" baseAuthorityYear="2015" box="[964,1098,333,359]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="264" phylum="Chordata" rank="species" species="ilamensis">
<emphasis id="24F2EABCFFA4B82988AEFEBD9F12FED5" box="[964,1098,333,359]" italics="true" pageId="6" pageNumber="264">P. ilamensis</emphasis>
</taxonomicName>
has two days priority over
<taxonomicName id="D1864D2DFFA4B8298EE2FEBD9A5FFE39" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="264" phylum="Chordata" rank="species" species="pasatigris">
<emphasis id="24F2EABCFFA4B8298EE2FEBD9A5FFE39" italics="true" pageId="6" pageNumber="264">P. pasatigris</emphasis>
</taxonomicName>
and
<taxonomicName id="D1864D2DFFA4B8298A54FE819A91FE39" box="[318,457,370,395]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="264" phylum="Chordata" rank="species" species="pasatigris">
<emphasis id="24F2EABCFFA4B8298A54FE819A91FE39" box="[318,457,370,395]" italics="true" pageId="6" pageNumber="264">P. pasatigris</emphasis>
</taxonomicName>
has to be treated as a junior synonym of
<taxonomicName id="D1864D2DFFA4B82988FFFE819F43FE39" baseAuthorityName="Vatandoust &amp; Eagderi" baseAuthorityYear="2015" box="[917,1051,369,395]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="264" phylum="Chordata" rank="species" species="ilamensis">
<emphasis id="24F2EABCFFA4B82988FFFE819F43FE39" box="[917,1051,369,395]" italics="true" pageId="6" pageNumber="264">P. ilamensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>