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<mods:title id="21C5E7C71DAC49DCE67581EC4BA70891">Pamphagidae (Orthoptera: Caelifera) of North Africa: key to genera and the annotated check-list of species</mods:title>
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<mods:namePart id="A4FA05B17C40C8AB0CFC5EB4EBFD1089">Massa, Bruno</mods:namePart>
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<treatment id="03C187FAFFB2B354FF64D2E45FACFA42" ID-DOI="http://doi.org/10.5281/zenodo.5697544" ID-GBIF-Taxon="119574800" ID-Zenodo-Dep="5697544" LSID="urn:lsid:plazi:treatment:03C187FAFFB2B354FF64D2E45FACFA42" httpUri="http://treatment.plazi.org/id/03C187FAFFB2B354FF64D2E45FACFA42" lastPageNumber="451" pageId="16" pageNumber="451">
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<paragraph id="8BD736ECFFB2B354FF64D2E45EDDFE29" blockId="16.[151,1022,473,499]" box="[151,1022,473,499]" pageId="16" pageNumber="451">
<heading id="D09F8180FFB2B354FF64D2E45EDDFE29" bold="true" box="[151,1022,473,499]" fontSize="11" level="1" pageId="16" pageNumber="451" reason="1">
<emphasis id="B91CEAFEFFB2B354FF64D2E45EDDFE29" bold="true" box="[151,1022,473,499]" pageId="16" pageNumber="451">
<taxonomicName id="4C684D6FFFB2B354FF64D2E45CC8FE29" ID-CoL="8NRP5" authority="Fischer, 1853" authorityName="Fischer" authorityYear="1853" box="[151,491,473,499]" class="Insecta" family="Pamphagidae" genus="Euryparyphes" kingdom="Animalia" order="Orthoptera" pageId="16" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEAFEFFB2B354FF64D2E45C63FE29" bold="true" box="[151,320,473,499]" italics="true" pageId="16" pageNumber="451">Euryparyphes</emphasis>
Fischer, 1853
</taxonomicName>
(type-species:
<taxonomicName id="4C684D6FFFB2B354FD50D2E45ED5FE29" authority="Serville" authorityName="Serville" box="[675,1014,473,499]" class="Insecta" family="Pamphagidae" genus="Porthetis" kingdom="Animalia" order="Orthoptera" pageId="16" pageNumber="451" phylum="Arthropoda" rank="species" species="terrulenta">
<emphasis id="B91CEAFEFFB2B354FD50D2E45EACFE29" bold="true" box="[675,911,473,499]" italics="true" pageId="16" pageNumber="451">Porthetis terrulenta</emphasis>
Serville
</taxonomicName>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD736ECFFB2B354FF64D11D5FACFA42" blockId="16.[151,1436,543,1432]" pageId="16" pageNumber="451">
A revision of the genus has been carried out by La Greca (1993), and a key to species by Massa (2012b). Characters of the genus are: prosternal process straight or slightly sloping, bi-pointed, vertex not protruding; median pronotal keel longitudinally furrowed (in some taxa only visible at high magnification), more or less raised, generally interrupted by the transverse sulcus (with some exceptions); pronotal lateral keels present or just evident (more evident when the median keel is straight). Mesosternal space is as wide as long or just wider than long in males and wider than long in females, metasternal space two times wider than long in males, three times in females.
<collectingCountry id="F37F767CFFB2B354FF0DD1CA5C15FCCA" box="[254,310,759,784]" name="India" pageId="16" pageNumber="451">Hind</collectingCountry>
border of pronotum is undulated or festooned; tegmina are oval, ca. two times longer than broad, brown with a whitish dorsal side, marked with a narrow brown line (absent in some species). First abdominal tergites are more or less toothed. Dorsal keel of hind femora is undulated or with very small spines, inner ventral keel with more or less evident spines (La Greca 1993, Llorente del Moral &amp; Presa Asensio 1997, Massa 2012b). It is closely related to
<taxonomicName id="4C684D6FFFB2B354FE74D0B45FEBFC7A" authority="Bolivar, 1914" authorityName="Bolivar" authorityYear="1914" box="[391,712,903,928]" class="Insecta" family="Pamphagidae" genus="Paraeumigus" kingdom="Animalia" order="Orthoptera" pageId="16" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEAFEFFB2B354FE74D0B45F3FFC7A" box="[391,540,905,928]" italics="true" pageId="16" pageNumber="451">Paraeumigus</emphasis>
Bolivar, 1914
</taxonomicName>
. According to Chopard (1943) the latter is characterized by a prosternal gorget (
<figureCitation id="13532A69FFB2B354FE95D0915CF0FC1E" box="[358,467,940,965]" captionStart="FIGURES 123 148. 123 145" captionStartId="36.[151,264,1951,1973]" captionTargetBox="[267,1317,179,1917]" captionTargetId="figure@36.[267,1317,179,1935]" captionTargetPageId="36" captionText="FIGURES 123 148. 123 145. Characters of species of the Moroccan genus Paraeumigus, included P. lopezi n. sp. (141 145, holotype and allotype, BMCP). 146 148. Lateral view of Amigus nigroadspersus and its phallic complex. Figs. 124, 135 137, 139 140 after La Greca (1993)." httpUri="https://zenodo.org/record/249387/files/figure.png" pageId="16" pageNumber="451">Figs. 142</figureCitation>
,
<figureCitation id="13532A69FFB2B354FE13D0915F65FC1F" box="[480,582,940,965]" captionStart="FIGURES 149 160" captionStartId="37.[151,264,1953,1975]" captionTargetBox="[268,1321,179,1934]" captionTargetId="figure@37.[268,1322,179,1935]" captionTargetPageId="37" captionText="FIGURES 149 160. Females of some species of the Morocco genus Paraeumigus: P. fortius syntype (149 150), P. sobrinus syntypes (151 153), P. escalerai syntype (154), P. l o pe zi n. sp. allotype (155), P. bolivari paratype (156), P. montanus (157, after Chopard 1943), P. tricoloripes (158), P. parvulus (159 160)." httpUri="https://zenodo.org/record/249388/files/figure.png" pageId="16" pageNumber="451">149, 153</figureCitation>
), but La Greca (1993) considers that only the shape of the aedeagus valves permits the separation of the two genera. In
<taxonomicName id="4C684D6FFFB2B354FD74D0F25E07FC32" box="[647,804,975,1000]" class="Insecta" family="Pamphagidae" genus="Euryparyphes" kingdom="Animalia" order="Orthoptera" pageId="16" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEAFEFFB2B354FD74D0F25E07FC32" box="[647,804,975,1000]" italics="true" pageId="16" pageNumber="451">Euryparyphes</emphasis>
</taxonomicName>
, valves in ventral view are more or less rhomboidal and may be of two
<typeStatus id="54D3884EFFB2B354FEBCD0C85CADFBD7" box="[335,398,1013,1037]" pageId="16" pageNumber="451">types</typeStatus>
: 1) present in the species characterized by raised pronotum, in which the central part of aedeagus is concave and consists of two triangular median plates, each paired with a main keel; lateral secondary keels are present, which produce a lateral concavity; 2) present in the species with low or straight pronotum keel, in which the apical borders are much longer than basal ones, the base of apical valves shows short columnar keels, separated by similar furrows; main keels do not differ from secondary keels, and the lateral concavity is absent. In contrast to
<taxonomicName id="4C684D6FFFB2B354FEEBD79A5C96FB1A" box="[280,437,1191,1216]" class="Insecta" family="Pamphagidae" genus="Euryparyphes" kingdom="Animalia" order="Orthoptera" pageId="16" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEAFEFFB2B354FEEBD79A5C96FB1A" box="[280,437,1191,1216]" italics="true" pageId="16" pageNumber="451">Euryparyphes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C684D6FFFB2B354FE35D7945F78FB1A" box="[454,603,1193,1216]" class="Insecta" family="Pamphagidae" genus="Paraeumigus" kingdom="Animalia" order="Orthoptera" pageId="16" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEAFEFFB2B354FE35D7945F78FB1A" box="[454,603,1193,1216]" italics="true" pageId="16" pageNumber="451">Paraeumigus</emphasis>
</taxonomicName>
shows a swollen base of valves, provided with columnar keels, without distinction between main and secondary keels; the apical part over the base is narrow, develops parallel to one other, and enlarges apically in two pairs of membranous lobes, one pair per valve; the base of apical valves is characterized by 34 columnar keels per side. Additionally,
<taxonomicName id="4C684D6FFFB2B354FCCBD6295EF6FAF7" box="[824,981,1300,1325]" class="Insecta" family="Pamphagidae" genus="Euryparyphes" kingdom="Animalia" order="Orthoptera" pageId="16" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEAFEFFB2B354FCCBD6295EF6FAF7" box="[824,981,1300,1325]" italics="true" pageId="16" pageNumber="451">Euryparyphes</emphasis>
</taxonomicName>
species have clearly oval tegmina (
<figureCitation id="13532A69FFB2B354FA90D6295DD9FA8A" captionStart="FIGURES 110 122" captionStartId="35.[151,264,1503,1525]" captionTargetBox="[152,1436,182,1472]" captionTargetId="figure@35.[151,1436,177,1481]" captionTargetPageId="35" captionText="FIGURES 110 122. Characters of Euryparyphes rungsi n. sp. from Middle Atlas; holotype (110, 111, 114, 115, 116) and allotype (112, 113, 117, 118); epiphallus (119) and aedeagus valves of holotype, in lateral (120), front (121) and back view (122) (MNCN)." httpUri="https://zenodo.org/record/249386/files/figure.png" pageId="16" pageNumber="451">Figs. 110113</figureCitation>
), while
<taxonomicName id="4C684D6FFFB2B354FEA0D6045CCBFA8A" box="[339,488,1337,1360]" class="Insecta" family="Pamphagidae" genus="Paraeumigus" kingdom="Animalia" order="Orthoptera" pageId="16" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEAFEFFB2B354FEA0D6045CCBFA8A" box="[339,488,1337,1360]" italics="true" pageId="16" pageNumber="451">Paraeumigus</emphasis>
</taxonomicName>
ones have less oval tegmina, with upper border more or less straight (
<figureCitation id="13532A69FFB2B354FB07D60A58BBFA8A" box="[1268,1432,1335,1360]" captionStart="FIGURES 123 148. 123 145" captionStartId="36.[151,264,1951,1973]" captionTargetBox="[267,1317,179,1917]" captionTargetId="figure@36.[267,1317,179,1935]" captionTargetPageId="36" captionText="FIGURES 123 148. 123 145. Characters of species of the Moroccan genus Paraeumigus, included P. lopezi n. sp. (141 145, holotype and allotype, BMCP). 146 148. Lateral view of Amigus nigroadspersus and its phallic complex. Figs. 124, 135 137, 139 140 after La Greca (1993)." httpUri="https://zenodo.org/record/249387/files/figure.png" pageId="16" pageNumber="451">Figs. 123145</figureCitation>
,
<figureCitation id="13532A69FFB2B354FF64D6615DDEFAAF" box="[151,253,1372,1397]" captionStart="FIGURES 149 160" captionStartId="37.[151,264,1953,1975]" captionTargetBox="[268,1321,179,1934]" captionTargetId="figure@37.[268,1322,179,1935]" captionTargetPageId="37" captionText="FIGURES 149 160. Females of some species of the Morocco genus Paraeumigus: P. fortius syntype (149 150), P. sobrinus syntypes (151 153), P. escalerai syntype (154), P. l o pe zi n. sp. allotype (155), P. bolivari paratype (156), P. montanus (157, after Chopard 1943), P. tricoloripes (158), P. parvulus (159 160)." httpUri="https://zenodo.org/record/249388/files/figure.png" pageId="16" pageNumber="451">150160</figureCitation>
). Massa (2012b) recognized 14 species of
<taxonomicName id="4C684D6FFFB2B354FD04D6615EB7FAAF" box="[759,916,1372,1397]" class="Insecta" family="Pamphagidae" genus="Euryparyphes" kingdom="Animalia" order="Orthoptera" pageId="16" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEAFEFFB2B354FD04D6615EB7FAAF" box="[759,916,1372,1397]" italics="true" pageId="16" pageNumber="451">Euryparyphes</emphasis>
</taxonomicName>
; another one is here described from Middle Atlas, amounting to 15 species on the whole.
</paragraph>
</subSubSection>
</treatment>
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