286 lines
77 KiB
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286 lines
77 KiB
XML
<document id="FE9F72E4A282A38FF9D213E2723C7824" ID-DOI="10.2478/aemnp-2018-0047" ID-GBIF-Dataset="3c9f44c6-59ff-4e7b-95e6-9c60bceb9ee5" ID-ISSN="1804-6487" ID-Zenodo-Dep="3550123" ID-ZooBank="661E0E32-1600-442D-95AC-4E2D427A9DBD" IM.bibliography_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1574365496940" checkinUser="tatiana" docAuthor="Ruschel, Tatiana Petersen" docDate="2018" docId="03C187F69F27FF93FEF5FD2BFDFE092C" docLanguage="en" docName="ActaEntMusNatPra.58.2.559-566.pdf.imf" docOrigin="Acta Entomologica Musei Nationalis Pragae 58 (2)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Gibbocicada Ruschel 2018, gen. nov." docType="treatment" docVersion="8" lastPageNumber="561" masterDocId="FFF8FF8E9F25FF96FFB8FF95FF850B39" masterDocTitle="Gibbocicada brasiliana, new genus and new species from Brazil and a key for the genera of Tibicinini (Hemiptera: Auchenorrhyncha: Cicadidae)" masterLastPageNumber="566" masterPageNumber="559" pageNumber="561" updateTime="1684257795048" updateUser="tatiana">
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<mods:titleInfo id="EF319CE77C09C53C579240C389A12F3D">
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<mods:title id="A8CFD05D76B9B190F96DB1FB54876454">Gibbocicada brasiliana, new genus and new species from Brazil and a key for the genera of Tibicinini (Hemiptera: Auchenorrhyncha: Cicadidae)</mods:title>
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<mods:name id="F2F08F29AC455BDE38FE8DB701C7E4B2" type="personal">
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<mods:namePart id="C4E26BED75B3E69F2703E38D1D59C683">Ruschel, Tatiana Petersen</mods:namePart>
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<mods:affiliation id="1BA216CDCC30833ADA6BC90A382334A4">Departamento de Zoologia, Universidade Federal do Rio Grande do Sul. Avenida Bento Gonçalves 9500, Agronomia, 91501 – 970 Porto Alegre, RS, Brazil</mods:affiliation>
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<mods:nameIdentifier id="60407C8F3B949F422771FA1ACAC628C1" type="email">tatiana.ruschel@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="15A64B2AA81B58AB9C30F9BD05C837DA">text</mods:typeOfResource>
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<mods:title id="45B340F1D2C9DFB56B2273748BB6287A">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:part id="D32B107B80B1B2CEE89DBBA541E18D89">
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<mods:date id="358E5F619816B0FB6A36B8BC528D4D98">2018</mods:date>
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<mods:number id="540D79CB808C0D41289CAAADD21FEC25">2018-11-26</mods:number>
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<mods:number id="406321B25F2BA50288F3E98C27642AF1">58</mods:number>
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<mods:classification id="8AF2985CD8BBE21CF30E93468AE4DD03">journal article</mods:classification>
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<mods:identifier id="03BB286321783DC9334407B013B9E7E4" type="DOI">10.2478/aemnp-2018-0047</mods:identifier>
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<treatment id="03C187F69F27FF93FEF5FD2BFDFE092C" ID-DOI="http://doi.org/10.5281/zenodo.6485692" ID-GBIF-Taxon="160511694" ID-Zenodo-Dep="6485692" LSID="urn:lsid:plazi:treatment:03C187F69F27FF93FEF5FD2BFDFE092C" httpUri="http://treatment.plazi.org/id/03C187F69F27FF93FEF5FD2BFDFE092C" lastPageId="5" lastPageNumber="561" pageId="2" pageNumber="561">
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<subSubSection id="C372656B9F27FF94FEF5FD2BFDC709EC" box="[333,578,702,725]" pageId="2" pageNumber="561" type="nomenclature">
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<paragraph id="8BD736E09F27FF94FEF5FD2BFDC709EC" blockId="2.[151,761,702,2071]" box="[333,578,702,725]" pageId="2" pageNumber="561">
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<heading id="D09F818C9F27FF94FEF5FD2BFDC709EC" bold="true" box="[333,578,702,725]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="561" reason="4">
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<emphasis id="B91CEAF29F27FF94FEF5FD2BFDC709EC" bold="true" box="[333,578,702,725]" pageId="2" pageNumber="561">
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<taxonomicName id="4C684D639F27FF94FEF5FD2BFE5209EC" ID-CoL="8JL7N" authority="Ruschel, 2018" authorityName="Ruschel" authorityYear="2018" box="[333,471,702,725]" class="Insecta" family="Cicadidae" genus="Gibbocicada" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B91CEAF29F27FF94FEF5FD2BFE5209EC" bold="true" box="[333,471,702,725]" italics="true" pageId="2" pageNumber="561">Gibbocicada</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22F57899F27FF94FE66FD2BFDC709EC" box="[478,578,702,725]" pageId="2" pageNumber="561" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C372656B9F27FF94FE2EFD49FE7C09D6" box="[406,505,732,751]" pageId="2" pageNumber="561" type="description">
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<paragraph id="8BD736E09F27FF94FE2EFD49FE7C09D6" blockId="2.[151,761,702,2071]" box="[406,505,732,751]" pageId="2" pageNumber="561">
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(
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<figureCitation id="13532A659F27FF94FE24FD49FE7709D6" box="[412,498,732,751]" captionStart-0="Figs 2–11" captionStart-1="Figs 12–14" captionStart-2="Figs 15–18" captionStartId-0="3.[151,188,1930,1949]" captionStartId-1="4.[151,188,908,927]" captionStartId-2="4.[151,188,2010,2029]" captionTargetId-0="figure@3.[151,1436,414,1916]" captionTargetId-1="figure@4.[151,1436,189,887]" captionTargetId-2="figure@4.[151,1436,1096,1992]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="4" captionText-0="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." captionText-1="Figs 12–14. Gibbocicada brasiliana gen. et sp. nov., paratype, male. 12 – Pygofer, uncus and aedeagus, in latero-ventral view; 13 – same, in lateral view; 14 – aedeagus in ventral view. Abbreviations: aed – aedeagus, as – anal styles, at – anal tube, bl – basal lobe, pyg – pygofer, un – uncus, vb – ventral branches, ve – vesica, vbp – ventrobasal pocket. Scale bars: 12, 13 – 1 mm; 14 – 0.5 mm." captionText-2="Figs 15–18. Gibbocicada brasiliana gen. et sp. nov., paratype, female. 15 – habitus in dorsal view; 16 – operculum in latero-ventral view; 17 – sternite VII, abdominal segment 9 and ovipositor sheath in ventral view; 18 – tergites 5–9 in lateral view.Abbreviations: ab9 – abdominal segment 9, db – dorsal beak, mc – meracanthus, op – operculum, os – ovipositor sheath, st – sternite. Scale bars: 15 – 2 cm; 16 – 1 mm; 17, 18 – 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3550127" figureDoi-1="http://doi.org/10.5281/zenodo.3550129" figureDoi-2="http://doi.org/10.5281/zenodo.3550131" httpUri-0="https://zenodo.org/record/3550127/files/figure.png" httpUri-1="https://zenodo.org/record/3550129/files/figure.png" httpUri-2="https://zenodo.org/record/3550131/files/figure.png" pageId="2" pageNumber="561">Figs 2–18</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C372656B9F27FF94FF2FFC95FF72080E" pageId="2" pageNumber="561" type="type_taxon">
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<paragraph id="8BD736E09F27FF94FF2FFC95FF72080E" blockId="2.[151,761,702,2071]" pageId="2" pageNumber="561">
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<emphasis id="B91CEAF29F27FF94FF2FFC95FEA8082E" bold="true" box="[151,301,768,791]" pageId="2" pageNumber="561">
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<typeStatus id="54D388429F27FF94FF2FFC95FF4A082E" box="[151,207,768,791]" pageId="2" pageNumber="561">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C684D639F27FF94FE8DFC95FDB0082E" authority="Ruschel, 2018" authorityName="Ruschel" authorityYear="2018" box="[309,565,768,791]" class="Insecta" family="Cicadidae" genus="Gibbocicada" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="brasiliana" status="sp. nov.">
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<emphasis id="B91CEAF29F27FF94FE8DFC95FDB0082E" box="[309,565,768,791]" italics="true" pageId="2" pageNumber="561">Gibbocicada brasiliana</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22F57899F27FF94FD85FC95FD16082E" box="[573,659,768,791]" pageId="2" pageNumber="561" rank="species">sp. nov.</taxonomicNameLabel>
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, here designated.
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</paragraph>
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</subSubSection>
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<subSubSection id="C372656B9F27FF94FF2FFCD5FDA60DCE" pageId="2" pageNumber="561" type="diagnosis">
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<paragraph id="8BD736E09F27FF94FF2FFCD5FDA60DCE" blockId="2.[151,761,702,2071]" pageId="2" pageNumber="561">
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<emphasis id="B91CEAF29F27FF94FF2FFCD5FE89086E" bold="true" box="[151,268,832,855]" pageId="2" pageNumber="561">Diagnosis.</emphasis>
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The new genus can be distinguished from all other genera in the
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<taxonomicName id="4C684D639F27FF94FEE2FCF5FE45084E" authorityName="Distant" authorityYear="1905" box="[346,448,864,887]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="tribe" tribe="Tibicinini">Tibicinini</taxonomicName>
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by the following combination of characters: head short, three times shorter than pronotoum in dorsal view (
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<figureCitation id="13532A659F27FF94FECCFC35FE30088E" box="[372,437,928,951]" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetBox="[151,1436,414,1915]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">Fig. 6</figureCitation>
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); lateral ocelli almost reaching the anterior margin of pronotum in dorsal view (
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<figureCitation id="13532A659F27FF94FD1FFC55FD6F08EE" box="[679,746,960,983]" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">Fig. 6</figureCitation>
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); anterior margin of lateral lobe of pronotum reaching the eyes (
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<figureCitation id="13532A659F27FF94FF6DFB95FE930F2E" box="[213,278,1024,1047]" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">Fig. 6</figureCitation>
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); pronotum and mesonotum pronounced dorsally in lateral view (
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<figureCitation id="13532A659F27FF94FE38FBB5FE410F0E" box="[384,452,1056,1079]" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetBox="[151,1436,414,1915]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">Fig. 5</figureCitation>
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); dorsal midline of pronotal collar very broad, as wide as the diameter of eye (
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<figureCitation id="13532A659F27FF94FD10FBD5FD6F0F6E" box="[680,746,1088,1111]" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">Fig. 6</figureCitation>
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); lateral angle of pronotal collar laterally pronounced (
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<figureCitation id="13532A659F27FF94FD75FBF5FF220FAE" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">Fig. 6</figureCitation>
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); meracanthus large, of the same length as operculum (
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<figureCitation id="13532A659F27FF94FF27FB35FF650F8E" box="[159,224,1184,1207]" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetBox="[151,1436,414,1915]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">Fig. 9</figureCitation>
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); fore wings with the median and cubitus anterior veins rising from the same point in the basal cell (
|
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<figureCitation id="13532A659F27FF94FD11FB55FD6C0FEE" box="[681,745,1216,1239]" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetBox="[151,1436,414,1915]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">Fig. 3</figureCitation>
|
||
); tergite 1 fully covered by cruciform elevation; uncus posteriorly developed (
|
||
<figureCitation id="13532A659F27FF94FEF6FA95FE1C0E2E" box="[334,409,1280,1303]" captionStart="Figs 12–14" captionStartId="4.[151,188,908,927]" captionTargetBox="[161,1410,215,870]" captionTargetId="figure@4.[151,1436,189,887]" captionTargetPageId="4" captionText="Figs 12–14. Gibbocicada brasiliana gen. et sp. nov., paratype, male. 12 – Pygofer, uncus and aedeagus, in latero-ventral view; 13 – same, in lateral view; 14 – aedeagus in ventral view. Abbreviations: aed – aedeagus, as – anal styles, at – anal tube, bl – basal lobe, pyg – pygofer, un – uncus, vb – ventral branches, ve – vesica, vbp – ventrobasal pocket. Scale bars: 12, 13 – 1 mm; 14 – 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3550129" httpUri="https://zenodo.org/record/3550129/files/figure.png" pageId="2" pageNumber="561">Fig. 13</figureCitation>
|
||
); basal lobe long, almostreaching the uncus (
|
||
<figureCitation id="13532A659F27FF94FEB5FAB5FED80E0E" box="[269,349,1312,1335]" captionStart="Figs 12–14" captionStartId="4.[151,188,908,927]" captionTargetId="figure@4.[151,1436,189,887]" captionTargetPageId="4" captionText="Figs 12–14. Gibbocicada brasiliana gen. et sp. nov., paratype, male. 12 – Pygofer, uncus and aedeagus, in latero-ventral view; 13 – same, in lateral view; 14 – aedeagus in ventral view. Abbreviations: aed – aedeagus, as – anal styles, at – anal tube, bl – basal lobe, pyg – pygofer, un – uncus, vb – ventral branches, ve – vesica, vbp – ventrobasal pocket. Scale bars: 12, 13 – 1 mm; 14 – 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3550129" httpUri="https://zenodo.org/record/3550129/files/figure.png" pageId="2" pageNumber="561">Fig. 13</figureCitation>
|
||
). The new genus can be distinguished from
|
||
<taxonomicName id="4C684D639F27FF94FF6FFAD5FEE50E6E" authorityName="Davis" authorityYear="1919" box="[215,352,1344,1367]" class="Insecta" family="Cicadidae" genus="Okanagodes" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91CEAF29F27FF94FF6FFAD5FEE50E6E" box="[215,352,1344,1367]" italics="true" pageId="2" pageNumber="561">Okanagodes</emphasis>
|
||
</taxonomicName>
|
||
by the distance between the lateral ocellus and eyes greater than diameter of ocellus, and by the postclypeus short, slightly prominent relative to the anterior margin of head in dorsal view; from
|
||
<taxonomicName id="4C684D639F27FF94FDCFFA35FD7D0E8E" authorityName="Van Duzee" authorityYear="1915" box="[631,760,1440,1463]" class="Insecta" family="Cicadidae" genus="Clidophleps" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91CEAF29F27FF94FDCFFA35FD7D0E8E" box="[631,760,1440,1463]" italics="true" pageId="2" pageNumber="561">Clidophleps</emphasis>
|
||
</taxonomicName>
|
||
by the medial cell notmuch narrower than radial cell; from
|
||
<taxonomicName id="4C684D639F27FF94FF2FFA75FE960ECE" authorityName="Distant" authorityYear="1905" box="[151,275,1504,1527]" class="Insecta" family="Cicadidae" genus="Okanagana" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91CEAF29F27FF94FF2FFA75FE960ECE" box="[151,275,1504,1527]" italics="true" pageId="2" pageNumber="561">Okanagana</emphasis>
|
||
</taxonomicName>
|
||
by the head (including eyes) broader than the pronotum; from
|
||
<taxonomicName id="4C684D639F27FF94FEFDF995FE190D2E" authorityName="Distant" authorityYear="1905" box="[325,412,1536,1559]" class="Insecta" family="Cicadidae" genus="Paharia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91CEAF29F27FF94FEFDF995FE190D2E" box="[325,412,1536,1559]" italics="true" pageId="2" pageNumber="561">Paharia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C684D639F27FF94FE10F995FDA60D2E" authorityName="Chen" authorityYear="1943" box="[424,547,1536,1559]" class="Insecta" family="Cicadidae" genus="Subpsaltria" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91CEAF29F27FF94FE10F995FDA60D2E" box="[424,547,1536,1559]" italics="true" pageId="2" pageNumber="561">Subpsaltria</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C684D639F27FF94FDE2F995FD340D2E" authorityName="Kolenati" authorityYear="1857" box="[602,689,1536,1559]" class="Insecta" family="Cicadidae" genus="Tibicina" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91CEAF29F27FF94FDE2F995FD340D2E" box="[602,689,1536,1559]" italics="true" pageId="2" pageNumber="561">Tibicina</emphasis>
|
||
</taxonomicName>
|
||
by the pronotum long, three timeslonger than head, the aedeagus not elongated nor curved anteriorly; from
|
||
<taxonomicName id="4C684D639F27FF94FDE4F9D5FD500D6E" authorityName="Lee" authorityYear="2012" box="[604,725,1600,1623]" class="Insecta" family="Cicadidae" genus="Subtibicina" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91CEAF29F27FF94FDE4F9D5FD500D6E" box="[604,725,1600,1623]" italics="true" pageId="2" pageNumber="561">Subtibicina</emphasis>
|
||
</taxonomicName>
|
||
by the fore wings being broad (less than three times the length), not tinged with yellowish brown, and by the uncus being arched and not flat; and from
|
||
<taxonomicName id="4C684D639F27FF94FD9FF935FD2C0D8E" authorityName="Distant" authorityYear="1914" box="[551,681,1696,1719]" class="Insecta" family="Cicadidae" genus="Tibicinoides" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91CEAF29F27FF94FD9FF935FD2C0D8E" box="[551,681,1696,1719]" italics="true" pageId="2" pageNumber="561">Tibicinoides</emphasis>
|
||
</taxonomicName>
|
||
by the base of wings colorless, and the medial cell of fore wings almost twice longer than ulnar cell 3.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C372656B9F27FF94FF2FF895FBD80F8C" pageId="2" pageNumber="561" type="description">
|
||
<paragraph id="8BD736E09F27FF94FF2FF895FBD80F8C" blockId="2.[151,761,702,2071]" lastBlockId="2.[828,1437,190,2069]" pageId="2" pageNumber="561">
|
||
<emphasis id="B91CEAF29F27FF94FF2FF895FEA70C2E" bold="true" box="[151,290,1792,1815]" pageId="2" pageNumber="561">Description.</emphasis>
|
||
Head (
|
||
<figureCitation id="13532A659F27FF94FED4F895FE2F0C2E" box="[364,426,1792,1815]" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">Fig. 6</figureCitation>
|
||
) (including eyes) wide, slightly broader than pronotum (excluding the pronotal collar), and narrower than mesonotum; head short, three times shorter than pronotoum in dorsal view. Lateral ocelli widelyseparated and almost reaching the anteriormargin of pronotum. Vertex wide, the distance between the lateral ocellus and eyes greater than diameter of ocellus. Supra-antennal plates not prominent and not meeting eye. Postclypeus short and narrow in dorsal view, its width equal to the distance between the lateral ocelli; slightly prominent relative to the anterior margin of head in dorsal view; rectangular in ventral view (
|
||
<figureCitation id="13532A659F27FF94FC75FF6BFB890A2C" box="[973,1036,254,277]" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">Fig. 7</figureCitation>
|
||
) and obtuse in lateral view. Pronotum and mesonotum pronounced dorsally in lateral view (
|
||
<figureCitation id="13532A659F27FF94FAC9FE8BFCCE0A6C" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">Fig. 5</figureCitation>
|
||
). Pronotum (
|
||
<figureCitation id="13532A659F27FF94FC60FEABFB9D0A6C" box="[984,1048,318,341]" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">Fig. 6</figureCitation>
|
||
) long, three times longer than head, not depressed laterally; anterior margin of lateral lobe of pronotum reaching the eyes; pronotal collar broad in dorsal view, as wide as diameter of eye; lateral angle of pronotal collar laterally pronounced, exceeding the lateral margin of mesonotum; lateralpart of pronotalcollar narrow relative to lateral angle. Mesonotal stridulatory apparatus absent. Operculum (
|
||
<figureCitation id="13532A659F27FF94FBA8FD8BFBCA090C" box="[1040,1103,542,565]" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">Fig. 9</figureCitation>
|
||
) broad, the lateral andposterior margins slightly convex, and distal margins not reaching thedistal margins of tympanal cavities. Meracanthus (
|
||
<figureCitation id="13532A659F27FF94FAC9FDCBFCCF09AC" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">Fig. 9</figureCitation>
|
||
) large, of the same length as operculum. Basisternum 3 flat, covered by the meso- and metacoxae. Cruciform elevation with the central and lateral areas flat. Profemora armed with two spines, the primary one the longest and parallel to the posterior margin of femur, and the secondary one sharp and straight. Tarsi three-segmented. Fore wings (
|
||
<figureCitation id="13532A659F27FF94FC35FCABFC4A086C" box="[909,975,830,853]" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">Fig. 2</figureCitation>
|
||
) hyaline; broad, width almost three times the length; with eight apical cells; the median and cubitus anterior veins rising from the same point in the basal cell; medial cell almost twice longer than ulnar cell 3 and not much narrower than radial cell; hind wings (
|
||
<figureCitation id="13532A659F27FF94FAABFC2BFAD708EC" box="[1299,1362,958,981]" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetBox="[151,1436,414,1915]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">Fig. 2</figureCitation>
|
||
) about half the length of fore wings, with six apical cells. Male timbal covers absent (
|
||
<figureCitation id="13532A659F27FF94FB8AFC6BFB030F2C" box="[1074,1158,1022,1045]" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">Fig. 10</figureCitation>
|
||
). Tergite 1 fully covered by cruciform elevation. Sternite II (
|
||
<figureCitation id="13532A659F27FF94FB7CFB8BFA8A0F0C" box="[1220,1295,1054,1077]" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetBox="[151,1436,414,1915]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">
|
||
<figureCitation id="13532A659F27FF94FB7CFB8BFA800F0C" box="[1220,1285,1054,1077]" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">Fig. 8</figureCitation>
|
||
)
|
||
</figureCitation>
|
||
distant from metacoxae. Uncus longer than sternite VIII, elevated and posteriorly developed. Basal lobe of pygofer long, almost reaching the uncus (
|
||
<figureCitation id="13532A659F27FF94FBAAFBEBFBE50FAC" box="[1042,1120,1150,1173]" captionStart="Figs 12–14" captionStartId="4.[151,188,908,927]" captionTargetBox="[161,1410,215,870]" captionTargetId="figure@4.[151,1436,189,887]" captionTargetPageId="4" captionText="Figs 12–14. Gibbocicada brasiliana gen. et sp. nov., paratype, male. 12 – Pygofer, uncus and aedeagus, in latero-ventral view; 13 – same, in lateral view; 14 – aedeagus in ventral view. Abbreviations: aed – aedeagus, as – anal styles, at – anal tube, bl – basal lobe, pyg – pygofer, un – uncus, vb – ventral branches, ve – vesica, vbp – ventrobasal pocket. Scale bars: 12, 13 – 1 mm; 14 – 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3550129" httpUri="https://zenodo.org/record/3550129/files/figure.png" pageId="2" pageNumber="561">Fig. 13</figureCitation>
|
||
). Aedeagus not elongated nor curved anteriorly (
|
||
<figureCitation id="13532A659F27FF94FBB9FB0BFBD40F8C" box="[1025,1105,1182,1205]" captionStart="Figs 12–14" captionStartId="4.[151,188,908,927]" captionTargetBox="[161,1410,215,870]" captionTargetId="figure@4.[151,1436,189,887]" captionTargetPageId="4" captionText="Figs 12–14. Gibbocicada brasiliana gen. et sp. nov., paratype, male. 12 – Pygofer, uncus and aedeagus, in latero-ventral view; 13 – same, in lateral view; 14 – aedeagus in ventral view. Abbreviations: aed – aedeagus, as – anal styles, at – anal tube, bl – basal lobe, pyg – pygofer, un – uncus, vb – ventral branches, ve – vesica, vbp – ventrobasal pocket. Scale bars: 12, 13 – 1 mm; 14 – 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3550129" httpUri="https://zenodo.org/record/3550129/files/figure.png" pageId="2" pageNumber="561">Fig. 12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C372656B9F27FF94FC84FB2BFAE30E0C" pageId="2" pageNumber="561" type="etymology">
|
||
<paragraph id="8BD736E09F27FF94FC84FB2BFAE30E0C" blockId="2.[828,1437,190,2069]" pageId="2" pageNumber="561">
|
||
<emphasis id="B91CEAF29F27FF94FC84FB2BFC390FEC" bold="true" box="[828,956,1214,1237]" pageId="2" pageNumber="561">Etymology.</emphasis>
|
||
The name is derived from the Latin adjective
|
||
<emphasis id="B91CEAF29F27FF94FC84FB4BFC070FCC" box="[828,898,1246,1269]" italics="true" pageId="2" pageNumber="561">gibbus</emphasis>
|
||
(= humped) and the noun
|
||
<emphasis id="B91CEAF29F27FF94FB36FB4BFB510FCC" box="[1166,1236,1246,1269]" italics="true" pageId="2" pageNumber="561">cicada</emphasis>
|
||
(= cicada). Itrefers to the pronotum and mesonotum which are both pronounced dorsally in lateral view. The gendre is feminine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C372656B9F27FF93FC84FAABFDFE092C" lastPageId="5" lastPageNumber="564" pageId="2" pageNumber="561" type="discussion">
|
||
<paragraph id="8BD736E09F27FF94FC84FAABFB740C6C" blockId="2.[828,1437,190,2069]" pageId="2" pageNumber="561">
|
||
<emphasis id="B91CEAF29F27FF94FC84FAABFC470E6C" bold="true" box="[828,962,1342,1365]" pageId="2" pageNumber="561">Comments.</emphasis>
|
||
Based on the diagnosis of the subfamily proposed by
|
||
<bibRefCitation id="EFF94B119F27FF94FC74FACBFB4B0E4C" author="MARSHALL D. C. & MOULDS M. & HILL K. B. R. & PRICE B. W. & WADE E. & OWEN C. L. & GOEMANS G. & MARATHE K. & SARKAR V. & COOLEY J. R. & SANBORN A. F. & KUNTE K. & VILLET M. H. & SIMON C." box="[972,1230,1374,1397]" journalOrPublisher="Zootaxa" pageId="2" pageNumber="561" pagination="1 - 64" part="4424" refId="ref5320" refString="MARSHALL D. C., MOULDS M., HILL K. B. R., PRICE B. W., WADE E., OWEN C. L., GOEMANS G., MARATHE K., SARKAR V., COOLEY J. R., SANBORN A. F., KUNTE K., VILLET M. H. & SIMON C. 2018: A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily classification. Zootaxa 4424: 1 - 64." title="A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily classification" type="journal article" year="2018">MARSHALL et al. (2018)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C684D639F27FF94FB63FACBFAE00E4C" authority="Ruschel, 2018" authorityName="Ruschel" authorityYear="2018" box="[1243,1381,1374,1397]" class="Insecta" family="Cicadidae" genus="Gibbocicada" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B91CEAF29F27FF94FB63FACBFAE00E4C" box="[1243,1381,1374,1397]" italics="true" pageId="2" pageNumber="561">Gibbocicada</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22F57899F27FF94FAD7FACBFCEC0EAC" pageId="2" pageNumber="561" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
clearly belongs to the
|
||
<taxonomicName id="4C684D639F27FF94FBE4FAEBFB500EAC" authorityName="Distant" authorityYear="1905" box="[1116,1237,1406,1429]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="subFamily" subFamily="Tibicininae">Tibicininae</taxonomicName>
|
||
by the metanotum concealed at dorsal midline (
|
||
<figureCitation id="13532A659F27FF94FBC9FA0BFB310E8C" box="[1137,1204,1438,1461]" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">Fig. 2</figureCitation>
|
||
); the hind wing veins RP and Munfused at their bases (
|
||
<figureCitation id="13532A659F27FF94FB01FA2BFAA50EEC" box="[1209,1312,1470,1493]" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">Figs 3, 4</figureCitation>
|
||
); the distal marginsof male operculum not reachingthe distal margins of tympanal cavities and basallynot extendingbeyond meracanthus, but notin S-shape (
|
||
<figureCitation id="13532A659F27FF94FBCBF98BFB340D0C" box="[1139,1201,1566,1589]" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetBox="[151,1436,414,1915]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">Fig. 9</figureCitation>
|
||
); males lackingtimbal covers (
|
||
<figureCitation id="13532A659F27FF94FC2CF9ABFC6D0D6C" box="[916,1000,1598,1621]" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetBox="[151,1436,414,1915]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">Fig. 10</figureCitation>
|
||
); pygofer distal shoulder undeveloped; pygofer upper lobe absent; uncus very long and not retractable within pygofer; claspers absent; and the presence of a ventrobasalpocket (
|
||
<figureCitation id="13532A659F27FF94FB85F90BFB3C0D8C" box="[1085,1209,1694,1717]" captionStart="Figs 12–14" captionStartId="4.[151,188,908,927]" captionTargetId="figure@4.[151,1436,189,887]" captionTargetPageId="4" captionText="Figs 12–14. Gibbocicada brasiliana gen. et sp. nov., paratype, male. 12 – Pygofer, uncus and aedeagus, in latero-ventral view; 13 – same, in lateral view; 14 – aedeagus in ventral view. Abbreviations: aed – aedeagus, as – anal styles, at – anal tube, bl – basal lobe, pyg – pygofer, un – uncus, vb – ventral branches, ve – vesica, vbp – ventrobasal pocket. Scale bars: 12, 13 – 1 mm; 14 – 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3550129" httpUri="https://zenodo.org/record/3550129/files/figure.png" pageId="2" pageNumber="561">Figs 12–14</figureCitation>
|
||
). The forewing veins CuP and 1A are unfused in most genera of
|
||
<taxonomicName id="4C684D639F27FF94FB4EF92BFAF40DEC" authorityName="Distant" authorityYear="1905" box="[1270,1393,1726,1749]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="subFamily" subFamily="Tibicininae">Tibicininae</taxonomicName>
|
||
, but fused in
|
||
<taxonomicName id="4C684D639F27FF94FC2CF94BFB020DCC" authority="Uhler, 1888" authorityName="Uhler" authorityYear="1888" box="[916,1159,1758,1781]" class="Insecta" family="Cicadidae" genus="Platypedia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91CEAF29F27FF94FC2CF94BFB820DCC" box="[916,1031,1758,1781]" italics="true" pageId="2" pageNumber="561">Platypedia</emphasis>
|
||
Uhler, 1888
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C684D639F27FF94FB00F94BFCF40C2C" authority="Davis, 1920" authorityName="Davis" authorityYear="1920" class="Insecta" family="Cicadidae" genus="Neoplatypedia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91CEAF29F27FF94FB00F94BFAD60DCC" box="[1208,1363,1758,1781]" italics="true" pageId="2" pageNumber="561">Neoplatypedia</emphasis>
|
||
Davis, 1920
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C684D639F27FF94FCC6F96BFB870C2C" authorityName="Kato" authorityYear="1932" box="[894,1026,1790,1813]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="tribe" tribe="Platypediini">Platypediini</taxonomicName>
|
||
) (
|
||
<bibRefCitation id="EFF94B119F27FF94FBA0F96BFA840C2C" author="MARSHALL D. C. & MOULDS M. & HILL K. B. R. & PRICE B. W. & WADE E. & OWEN C. L. & GOEMANS G. & MARATHE K. & SARKAR V. & COOLEY J. R. & SANBORN A. F. & KUNTE K. & VILLET M. H. & SIMON C." box="[1048,1281,1790,1813]" journalOrPublisher="Zootaxa" pageId="2" pageNumber="561" pagination="1 - 64" part="4424" refId="ref5320" refString="MARSHALL D. C., MOULDS M., HILL K. B. R., PRICE B. W., WADE E., OWEN C. L., GOEMANS G., MARATHE K., SARKAR V., COOLEY J. R., SANBORN A. F., KUNTE K., VILLET M. H. & SIMON C. 2018: A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily classification. Zootaxa 4424: 1 - 64." title="A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily classification" type="journal article" year="2018">MARSHALLet al. 2018</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C684D639F27FF94FAAAF96BFA190C2C" authority="Ruschel, 2018" authorityName="Ruschel" authorityYear="2018" box="[1298,1436,1790,1813]" class="Insecta" family="Cicadidae" genus="Gibbocicada" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B91CEAF29F27FF94FAAAF96BFA190C2C" box="[1298,1436,1790,1813]" italics="true" pageId="2" pageNumber="561">Gibbocicada</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22F57899F27FF94FC84F88BFC190C0C" box="[828,924,1822,1845]" pageId="2" pageNumber="561" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
presents these veins partially fused, i.e. thebasal third of CuP and 1A are unfused (
|
||
<figureCitation id="13532A659F27FF94FB1AF8ABFB660C6C" box="[1186,1251,1854,1877]" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD736E09F27FF93FCE2F8CBFDFE092C" blockId="2.[828,1437,190,2069]" lastBlockId="5.[151,760,190,533]" lastPageId="5" lastPageNumber="564" pageId="2" pageNumber="561">
|
||
Among all tribes of
|
||
<taxonomicName id="4C684D639F27FF94FB92F8CBFB200C4C" authorityName="Distant" authorityYear="1905" box="[1066,1189,1886,1909]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="subFamily" subFamily="Tibicininae">Tibicininae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C684D639F27FF94FB16F8CBFABD0C4C" authority="Ruschel, 2018" authorityName="Ruschel" authorityYear="2018" box="[1198,1336,1886,1909]" class="Insecta" family="Cicadidae" genus="Gibbocicada" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B91CEAF29F27FF94FB16F8CBFABD0C4C" box="[1198,1336,1886,1909]" italics="true" pageId="2" pageNumber="561">Gibbocicada</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22F57899F27FF94FA85F8CBFA180C4C" box="[1341,1437,1886,1909]" pageId="2" pageNumber="561" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
is more similar to the genera classified in
|
||
<taxonomicName id="4C684D639F27FF94FB4FF8EBFAE70CAC" authorityName="Distant" authorityYear="1905" box="[1271,1378,1918,1941]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="tribe" tribe="Tibicinini">Tibicinini</taxonomicName>
|
||
, with the pronotum more or less distinctly narrowed anteriorly (
|
||
<figureCitation id="13532A659F27FF94FCFCF82BFC010CEC" box="[836,900,1982,2005]" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">Fig. 6</figureCitation>
|
||
); fore wings talc-like and obscurely wrinkled, their greatest breadth more than one third of their length (
|
||
<figureCitation id="13532A659F27FF94FAC9F84BFCC9032C" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">Fig. 2</figureCitation>
|
||
); abdomen subcylindrical (
|
||
<figureCitation id="13532A659F27FF94FBC6F86BFB41032C" box="[1150,1220,2046,2069]" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="2" pageNumber="561">Fig. 2</figureCitation>
|
||
); and uncus longer than sternite VIII (
|
||
<bibRefCitation id="EFF94B119F26FF95FEE5FF2BFE770BEC" author="DISTANT W. L." box="[349,498,190,213]" journalOrPublisher="Annals and Magazine of Natural History, Series 7" pageId="3" pageNumber="562" pagination="22 - 35" part="16" refId="ref5150" refString="DISTANT W. L. 1905: Rhynchotal notes - XXXIII. Annals and Magazine of Natural History, Series 7 16: 22 - 35." title="Rhynchotal notes - XXXIII" type="journal article" year="1905">DISTANT 1905</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="EFF94B119F26FF95FDBDFF2BFDFC0BEC" author="LEE Y. J." box="[517,633,190,213]" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="3" pageNumber="562" pagination="225 - 31" part="59" refId="ref5283" refString="LEE Y. J. 2012: Descriptions of two new genera and species of Cicadidae (Hemiptera) from India with some notes on tribal classification. Deutsche Entomologische Zeitschrift 59: 225 - 31." title="Descriptions of two new genera and species of Cicadidae (Hemiptera) from India with some notes on tribal classification" type="journal article" year="2012">LEE (2012)</bibRefCitation>
|
||
listed other characteristics shared by the genera of
|
||
<taxonomicName id="4C684D639F26FF95FD9FFF4BFD080BCC" authorityName="Distant" authorityYear="1905" box="[551,653,222,245]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="562" phylum="Arthropoda" rank="tribe" tribe="Tibicinini">Tibicinini</taxonomicName>
|
||
that Ialso observed in
|
||
<taxonomicName id="4C684D639F26FF95FEAFFF6BFE1B0A2C" authority="Ruschel, 2018" authorityName="Ruschel" authorityYear="2018" box="[279,414,254,277]" class="Insecta" family="Cicadidae" genus="Gibbocicada" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="562" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B91CEAF29F26FF95FEAFFF6BFE1B0A2C" box="[279,414,254,277]" italics="true" pageId="3" pageNumber="562">Gibbocicada</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22F57899F26FF95FE1DFF6BFD8D0A2C" box="[421,520,254,277]" pageId="3" pageNumber="562" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
: hind wing first cubital cell width at distal end about equal to second cubital cell; male operculum not reaching posterior margins of sternite II (
|
||
<figureCitation id="13532A659F26FF95FF0EFECBFF730A4C" box="[182,246,350,373]" captionStart="Figs 2–11" captionStartId="3.[151,188,1930,1949]" captionTargetId="figure@3.[151,1436,414,1916]" captionTargetPageId="3" captionText="Figs 2–11. Gibbocicada brasiliana gen. et sp. nov., holotype, male. 2 – habitus in dorsal view; 3 – wings in dorsal view (arrow points to the basal third of CuP and 1A which are unfused); 4 – detail of hind wing veins in dorsal view (arrow points to veins RP and M which are unfused at their bases); 5 – habitus in lateral view (wings not extended); 6 – head and pronotum in dorsal view; 7 – postclypeus in ventral view; 8 – thorax in ventral view; 9 – operculum in latero-ventral view; 10 – timbal in latero-ventral view; 11 – sternites VII and VIII in ventral view.Abbreviations: 1A – anal vein, CuA – cubitus anterior vein, CuP – cubitus posterior vein, M – median vein, mc – meracanthus, op – operculum, RP – radius posterior vein, tim – timbal, tymc – tympanal cavity, st – sternite. Scale bars: 2 – 2 cm; 3, 5 – 5 mm; 6–8, 11 – 2 mm; 4, 9, 10 –1 mm." figureDoi="http://doi.org/10.5281/zenodo.3550127" httpUri="https://zenodo.org/record/3550127/files/figure.png" pageId="3" pageNumber="562">Fig. 9</figureCitation>
|
||
); pygofer upper lobes absent, claspers absent and uncus very long (
|
||
<figureCitation id="13532A659F26FF95FC4FFF2BFBF00BEC" box="[1015,1141,190,213]" captionStart="Figs 12–14" captionStartId="4.[151,188,908,927]" captionTargetId="figure@4.[151,1436,189,887]" captionTargetPageId="4" captionText="Figs 12–14. Gibbocicada brasiliana gen. et sp. nov., paratype, male. 12 – Pygofer, uncus and aedeagus, in latero-ventral view; 13 – same, in lateral view; 14 – aedeagus in ventral view. Abbreviations: aed – aedeagus, as – anal styles, at – anal tube, bl – basal lobe, pyg – pygofer, un – uncus, vb – ventral branches, ve – vesica, vbp – ventrobasal pocket. Scale bars: 12, 13 – 1 mm; 14 – 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3550129" httpUri="https://zenodo.org/record/3550129/files/figure.png" pageId="3" pageNumber="562">Figs 13, 14</figureCitation>
|
||
). The new genus cannot be placed in the other tribes of
|
||
<taxonomicName id="4C684D639F26FF95FBE3FF4BFB540BCC" authorityName="Distant" authorityYear="1905" box="[1115,1233,222,245]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="562" phylum="Arthropoda" rank="subFamily" subFamily="Tibicininae">Tibicininae</taxonomicName>
|
||
for the lack of their diagnostic characters, i.e.
|
||
<taxonomicName id="4C684D639F26FF95FBEAFF6BFB470A2C" authorityName="Distant" authorityYear="1905" box="[1106,1218,254,277]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="562" phylum="Arthropoda" rank="tribe" tribe="Tettigadini">Tettigadini</taxonomicName>
|
||
because the species in the tribe are characterized by the lateral margins of the pronotum dilated and usually medially angulate (
|
||
<bibRefCitation id="EFF94B119F26FF95FAF1FEABFCF60A4C" author="DISTANT W. L." journalOrPublisher="Annals and Magazine of Natural History, Series 7" pageId="3" pageNumber="562" pagination="22 - 35" part="16" refId="ref5150" refString="DISTANT W. L. 1905: Rhynchotal notes - XXXIII. Annals and Magazine of Natural History, Series 7 16: 22 - 35." title="Rhynchotal notes - XXXIII" type="journal article" year="1905">DISTANT 1905</bibRefCitation>
|
||
), and by a mesonotal stridulatory apparatus (
|
||
<bibRefCitation id="EFF94B119F26FF93FAE0FECBFF4B0BEC" author="JACOBI A." journalOrPublisher="Zoologischer Anzeiger" lastPageId="5" lastPageNumber="564" pageId="3" pageNumber="562" pagination="67 - 71" part="32" refId="ref5265" refString="JACOBI A. 1907: Ein Schrillapparat bei Singcicaden. Zoologischer Anzeiger 32: 67 - 71." title="Ein Schrillapparat bei Singcicaden" type="journal article" year="1907">JACOBI 1907</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFF94B119F20FF93FF62FF2BFEF30BEC" author="BOULARD M." box="[218,374,190,213]" journalOrPublisher="Journal of Natural History" pageId="5" pageNumber="564" pagination="399 - 407" part="10" refId="ref4982" refString="BOULARD M. 1976: Un type nouveau d'appareil stridulant accessoire pour les Cicadoidea. Revision de la classification superieure de la superfamille (Hom.). Journal of Natural History 10: 399 - 407." title="Un type nouveau d'appareil stridulant accessoire pour les Cicadoidea. Revision de la classification superieure de la superfamille (Hom.)" type="journal article" year="1976">BOULARD 1976</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFF94B119F20FF93FE39FF2BFD900BEC" author="MOULDS M. S." box="[385,533,190,213]" journalOrPublisher="Records of the Australian Museum" pageId="5" pageNumber="564" pagination="375 - 446" part="57" refId="ref5458" refString="MOULDS M. S. 2005: An appraisal of the higher classification of cicadas (Hemiptera: Cicadoidea) with special reference to the Australian fauna. Records of the Australian Museum 57: 375 - 446." title="An appraisal of the higher classification of cicadas (Hemiptera: Cicadoidea) with special reference to the Australian fauna" type="journal article" year="2005">MOULDS 2005</bibRefCitation>
|
||
); Selymbrini because they have supra-antennal plates almostreaching the eye, a rounded postclypeus, hind coxae without lateral protruberance and a very short theca laterally expanded and distally ornamented (
|
||
<bibRefCitation id="EFF94B119F20FF93FEA6FEABFE790A6C" author="MARSHALL D. C. & MOULDS M. & HILL K. B. R. & PRICE B. W. & WADE E. & OWEN C. L. & GOEMANS G. & MARATHE K. & SARKAR V. & COOLEY J. R. & SANBORN A. F. & KUNTE K. & VILLET M. H. & SIMON C." box="[286,508,318,341]" journalOrPublisher="Zootaxa" pageId="5" pageNumber="564" pagination="1 - 64" part="4424" refId="ref5320" refString="MARSHALL D. C., MOULDS M., HILL K. B. R., PRICE B. W., WADE E., OWEN C. L., GOEMANS G., MARATHE K., SARKAR V., COOLEY J. R., SANBORN A. F., KUNTE K., VILLET M. H. & SIMON C. 2018: A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily classification. Zootaxa 4424: 1 - 64." title="A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily classification" type="journal article" year="2018">MARSHALL et al. 2018</bibRefCitation>
|
||
);
|
||
<taxonomicName id="4C684D639F20FF93FDB4FEABFF4E0A4C" authority="Sanborn, 2014" authorityName="Sanborn" authorityYear="2014" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="564" phylum="Arthropoda" rank="tribe" tribe="Chilecicadini">
|
||
Chilecicadini
|
||
<bibRefCitation id="EFF94B119F20FF93FD22FEABFF4E0A4C" author="SANBORN A. F." journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="5" pageNumber="564" pagination="339 - 348" part="116" refId="ref5652" refString="SANBORN A. F. 2014: A new genus and new tribe of cicada from South America (Hemiptera: Cicadoidea: Cicadidae) with a note on the taxonomic position of Ahomana Distant, 1905. Proceedings of the Entomological Society of Washington 116: 339 - 348." title="A new genus and new tribe of cicada from South America (Hemiptera: Cicadoidea: Cicadidae) with a note on the taxonomic position of Ahomana Distant, 1905" type="journal article" year="2014">Sanborn, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
because they havethe anteriorpronotum about as wide as the posterior pronotum, the lateral angle of the pronotal collar anteriorto thehind margin, and the partiallyexcavated lateral margin of the pronotal collar (
|
||
<bibRefCitation id="EFF94B119F20FF93FD9AFE2BFD3A0AEC" author="SANBORN A. F." box="[546,703,446,469]" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="5" pageNumber="564" pagination="339 - 348" part="116" refId="ref5652" refString="SANBORN A. F. 2014: A new genus and new tribe of cicada from South America (Hemiptera: Cicadoidea: Cicadidae) with a note on the taxonomic position of Ahomana Distant, 1905. Proceedings of the Entomological Society of Washington 116: 339 - 348." title="A new genus and new tribe of cicada from South America (Hemiptera: Cicadoidea: Cicadidae) with a note on the taxonomic position of Ahomana Distant, 1905" type="journal article" year="2014">SANBORN 2014</bibRefCitation>
|
||
); and
|
||
<taxonomicName id="4C684D639F20FF93FF2FFE4BFE1F0ACC" authority="Kato, 1932" authorityName="Kato" authorityYear="1932" box="[151,410,478,501]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="564" phylum="Arthropoda" rank="tribe" tribe="Platypediini">Platypediini Kato, 1932</taxonomicName>
|
||
because the tribe includes only species without timbal organs (
|
||
<bibRefCitation id="EFF94B119F20FF93FE62FE6BFDEA092C" author="MOULDS M. S." box="[474,623,510,533]" journalOrPublisher="Records of the Australian Museum" pageId="5" pageNumber="564" pagination="375 - 446" part="57" refId="ref5458" refString="MOULDS M. S. 2005: An appraisal of the higher classification of cicadas (Hemiptera: Cicadoidea) with special reference to the Australian fauna. Records of the Australian Museum 57: 375 - 446." title="An appraisal of the higher classification of cicadas (Hemiptera: Cicadoidea) with special reference to the Australian fauna" type="journal article" year="2005">MOULDS 2005</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |