534 lines
59 KiB
XML
534 lines
59 KiB
XML
<document id="4DF06FFB601D81574354569D6B595DE9" ID-DOI="10.11646/zootaxa.3609.4.3" ID-GBIF-Dataset="efd85aa0-d2a5-4d72-8125-6d45ab6935fc" ID-ISSN="1175-5326" ID-Zenodo-Dep="224051" ID-ZooBank="4D73C473-2533-4E1B-A039-113DF838F42A" IM.bibliography_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460572518824" checkinUser="plazi" docAuthor="Nunes, Rafael V. & Vaz-De-Mello, Fernando Z." docDate="2013" docId="03C187996A47073EFF00F97CE4BE850D" docLanguage="en" docName="zt03609p420.pdf" docOrigin="Zootaxa 3609 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dichotomius (Luederwaldtinia)" docType="key" docVersion="5" lastPageNumber="418" masterDocId="FFF8FFE16A410739FF97FFF6E12B827D" masterDocTitle="New brachypterous species of Dichotomius Hope, with taxonomic notes in the subgenus Luederwaldtinia Martínez (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="420" masterPageNumber="411" pageNumber="417" updateTime="1698322432242" updateUser="plazi">
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<mods:mods id="A939549810F829015087A5B4F3233A56" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5074FD4E4767C46E4DD782DBCDE8A601">
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<mods:title id="75AE1BDC09FA5A05F3C292DF2A39DB66">New brachypterous species of Dichotomius Hope, with taxonomic notes in the subgenus Luederwaldtinia Martínez (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="79B0D57FC8A3EF578372AFA3D6A58830">Nunes, Rafael V.</mods:namePart>
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<mods:name id="A1EA466207AAE944EFC8F260DC39D879" type="personal">
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<mods:roleTerm id="6155A80F97DF802036597F843EED6856">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="65A6CD23B5E0F5B1986DA77F3B496D17">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:typeOfResource id="488E4C1E908378DF4E173009CB1E4373">text</mods:typeOfResource>
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<mods:title id="643699EF79F4D2B1D8FBB8A37F51899F">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="9E794737144EA20ACF44D001E80139E3">
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<mods:date id="C8FBAE630B7E51B8F47DD7A51E1038C3">2013</mods:date>
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<mods:detail id="050DD91E6129D04ECF80A15D3D6EBC7C" type="pubDate">
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<mods:number id="EE6AE7415C91E716420EF27482443660">2013-01-31</mods:number>
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<mods:detail id="03AB9CAB0BFE90A4F8F8CEE9337A2323" type="volume">
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<mods:number id="474FB9AC4D80FAAE3BAA0F1DAD61C3B6">3609</mods:number>
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<mods:detail id="EE449ACA3057995B239677DBCA7424F8" type="issue">
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<mods:number id="F20CA50083C79464193DD8FCDF87A2CD">4</mods:number>
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<mods:start id="3EB7564A625C4AF0AD7980A985560CE2">411</mods:start>
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<mods:classification id="2D01C8F2EFECBB8463BFA9607EF25BF4">journal article</mods:classification>
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<mods:identifier id="4C956F963718EEE9DD732516E9562E04" type="DOI">10.11646/zootaxa.3609.4.3</mods:identifier>
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<mods:identifier id="7EF83DBD10128B67CC1DCCF5753F787D" type="GBIF-Dataset">efd85aa0-d2a5-4d72-8125-6d45ab6935fc</mods:identifier>
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<mods:identifier id="08BAAC27BC2E999261172CD01056B9DC" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="3D9F2107B48334676262BF4B324F5C44" type="Zenodo-Dep">224051</mods:identifier>
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</mods:mods>
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<treatment id="03C187996A47073EFF00F97CE4BE850D" ID-DOI="http://doi.org/10.5281/zenodo.6148988" ID-GBIF-Taxon="159868436" ID-Zenodo-Dep="6148988" LSID="urn:lsid:plazi:treatment:03C187996A47073EFF00F97CE4BE850D" httpUri="http://treatment.plazi.org/id/03C187996A47073EFF00F97CE4BE850D" lastPageId="7" lastPageNumber="418" pageId="6" pageNumber="417">
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<subSubSection id="C37265046A47073FFF00F97CE2C784DF" box="[151,1004,1673,1698]" pageId="6" pageNumber="417" type="nomenclature">
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<paragraph id="8BD7368F6A47073FFF00F97CE2C784DF" blockId="6.[151,1004,1673,1698]" box="[151,1004,1673,1698]" pageId="6" pageNumber="417">
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<emphasis id="B91CEA9D6A47073FFF00F97CE2C784DF" bold="true" box="[151,1004,1673,1698]" pageId="6" pageNumber="417">
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Key for identification of species groups in the subgenus
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<taxonomicName id="4C684D0C6A47073FFCB8F97FE2C784DF" ID-CoL="63YGX" box="[815,1004,1673,1698]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="subGenus" subGenus="Luederwaldtinia">
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<emphasis id="B91CEA9D6A47073FFCB8F97FE2C784DF" bold="true" box="[815,1004,1673,1698]" italics="true" pageId="6" pageNumber="417">Luederwaldtinia</emphasis>
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</taxonomicName>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37265046A47073FFF00F924E06A8598" pageId="6" pageNumber="417" type="discussion">
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<paragraph id="8BD7368F6A47073FFF00F924E06A8598" blockId="6.[151,1436,1745,2021]" pageId="6" pageNumber="417">
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We define here thirteen species groups, named after the older (or only) included species:
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<taxonomicName id="4C684D0C6A47073FFB12F924E5DC8494" baseAuthorityName="Harold" baseAuthorityYear="1869" box="[1157,1271,1746,1769]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="agenor">
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<emphasis id="B91CEA9D6A47073FFB12F924E5B28494" box="[1157,1177,1746,1769]" italics="true" pageId="6" pageNumber="417">D</emphasis>
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.
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<emphasis id="B91CEA9D6A47073FFB3EF924E5DC8494" box="[1193,1271,1746,1769]" italics="true" pageId="6" pageNumber="417">agenor</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C684D0C6A47073FFA92F924E4598494" baseAuthorityName="Eschscholtz" baseAuthorityYear="1822" box="[1285,1394,1745,1769]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="assifer">
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<emphasis id="B91CEA9D6A47073FFA92F924E4328494" box="[1285,1305,1746,1769]" italics="true" pageId="6" pageNumber="417">D</emphasis>
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.
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<emphasis id="B91CEA9D6A47073FFABFF927E4598494" box="[1320,1394,1745,1769]" italics="true" pageId="6" pageNumber="417">assifer</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C684D0C6A47073FFA16F924E1F18570" baseAuthorityName="Harold" baseAuthorityYear="1869" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="batesi">
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<emphasis id="B91CEA9D6A47073FFA16F924E4BE8494" box="[1409,1429,1746,1769]" italics="true" pageId="6" pageNumber="417">D</emphasis>
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.
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<emphasis id="B91CEA9D6A47073FFF00F903E1F18570" box="[151,218,1781,1805]" italics="true" pageId="6" pageNumber="417">batesi</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C684D0C6A47073FFF70F900E0B98570" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[231,402,1781,1805]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="carbonarius">
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<emphasis id="B91CEA9D6A47073FFF70F900E1D08570" box="[231,251,1782,1805]" italics="true" pageId="6" pageNumber="417">D</emphasis>
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.
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<emphasis id="B91CEA9D6A47073FFE9EF903E0B98570" box="[265,402,1781,1805]" italics="true" pageId="6" pageNumber="417">carbonarius</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C684D0C6A47073FFE08F900E3198570" baseAuthorityName="Germar" baseAuthorityYear="1824" box="[415,562,1781,1805]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="crinicollis">
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<emphasis id="B91CEA9D6A47073FFE08F900E0988570" box="[415,435,1782,1805]" italics="true" pageId="6" pageNumber="417">D</emphasis>
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.
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<emphasis id="B91CEA9D6A47073FFE56F903E3198570" box="[449,562,1781,1805]" italics="true" pageId="6" pageNumber="417">crinicollis</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C684D0C6A47073FFDD7F900E3D88570" authorityName="Pereira & D'Andretta" authorityYear="1955" box="[576,755,1781,1805]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="eucranioides">
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<emphasis id="B91CEA9D6A47073FFDD7F900E37F8570" box="[576,596,1782,1805]" italics="true" pageId="6" pageNumber="417">D</emphasis>
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.
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<emphasis id="B91CEA9D6A47073FFDF5F903E3D88570" box="[610,755,1781,1805]" italics="true" pageId="6" pageNumber="417">eucranioides</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C684D0C6A47073FFC97F900E2BE8570" baseAuthorityName="Arrow" baseAuthorityYear="1913" box="[768,917,1782,1805]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="geminatus">
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<emphasis id="B91CEA9D6A47073FFC97F900E23F8570" box="[768,788,1782,1805]" italics="true" pageId="6" pageNumber="417">D</emphasis>
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.
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<emphasis id="B91CEA9D6A47073FFCB5F900E2BE8570" box="[802,917,1782,1805]" italics="true" pageId="6" pageNumber="417">geminatus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C684D0C6A47073FFC35F900E5308570" baseAuthorityName="Erichson" baseAuthorityYear="1847" box="[930,1051,1781,1805]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="inachus">
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<emphasis id="B91CEA9D6A47073FFC35F900E29D8570" box="[930,950,1782,1805]" italics="true" pageId="6" pageNumber="417">D</emphasis>
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.
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<emphasis id="B91CEA9D6A47073FFC53F903E5308570" box="[964,1051,1781,1805]" italics="true" pageId="6" pageNumber="417">inachus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C684D0C6A47073FFBBFF900E58A8570" baseAuthorityName="Luederwaldt" baseAuthorityYear="1922" box="[1064,1185,1782,1805]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="muticus">
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<emphasis id="B91CEA9D6A47073FFBBFF900E5178570" box="[1064,1084,1782,1805]" italics="true" pageId="6" pageNumber="417">D</emphasis>
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.
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<emphasis id="B91CEA9D6A47073FFBDDF900E58A8570" box="[1098,1185,1782,1805]" italics="true" pageId="6" pageNumber="417">muticus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C684D0C6A47073FFB38F900E4218570" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[1199,1290,1782,1805]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="nisus">
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<emphasis id="B91CEA9D6A47073FFB38F900E5E88570" box="[1199,1219,1782,1805]" italics="true" pageId="6" pageNumber="417">D</emphasis>
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.
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<emphasis id="B91CEA9D6A47073FFB47F900E4218570" box="[1232,1290,1782,1805]" italics="true" pageId="6" pageNumber="417">nisus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C684D0C6A47073FFA80F900E4BE8570" baseAuthorityName="Harold" baseAuthorityYear="1867" box="[1303,1429,1782,1805]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="sericeus">
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<emphasis id="B91CEA9D6A47073FFA80F900E4008570" box="[1303,1323,1782,1805]" italics="true" pageId="6" pageNumber="417">D</emphasis>
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.
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<emphasis id="B91CEA9D6A47073FFAAEF900E4BE8570" box="[1337,1429,1782,1805]" italics="true" pageId="6" pageNumber="417">sericeus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C684D0C6A47073FFF00F8ECE00F854C" baseAuthorityName="Waterhouse" baseAuthorityYear="1891" box="[151,292,1818,1841]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="speciosus">
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<emphasis id="B91CEA9D6A47073FFF00F8ECE180854C" box="[151,171,1818,1841]" italics="true" pageId="6" pageNumber="417">D</emphasis>
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.
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<emphasis id="B91CEA9D6A47073FFF2EF8ECE00F854C" box="[185,292,1818,1841]" italics="true" pageId="6" pageNumber="417">speciosus</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="4C684D0C6A47073FFEF6F8ECE0CC854C" baseAuthorityName="Felsche" baseAuthorityYear="1901" box="[353,487,1817,1841]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="superbus">
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<emphasis id="B91CEA9D6A47073FFEF6F8ECE05E854C" box="[353,373,1818,1841]" italics="true" pageId="6" pageNumber="417">D</emphasis>
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.
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<emphasis id="B91CEA9D6A47073FFE15F8EFE0CC854C" box="[386,487,1817,1841]" italics="true" pageId="6" pageNumber="417">superbus</emphasis>
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</taxonomicName>
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. The main changes in these groups from previous treatments are 1) the incorporation of
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<bibRefCitation id="EFF94B7E6A47073FFF21F8CBE0828528" author="Luederwaldt, H." box="[182,425,1853,1877]" journalOrPublisher="Revista do Museu Paulista" pageId="6" pageNumber="417" pagination="207 - 216" part="20" refId="ref7939" refString="Luederwaldt, H. (1936) Notas complementares " As especies Brasileiras do genero Pinotus ". Revista do Museu Paulista, 20, 207 - 216." title="Notas complementares " As especies Brasileiras do genero Pinotus" type="journal article" year="1936">Luederwaldt’s (1936)</bibRefCitation>
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<taxonomicName id="4C684D0C6A47073FFE26F8C8E3088528" baseAuthorityName="Luederwaldt" baseAuthorityYear="1923" box="[433,547,1854,1877]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="micans">
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<emphasis id="B91CEA9D6A47073FFE26F8C8E0EE8528" box="[433,453,1854,1877]" italics="true" pageId="6" pageNumber="417">D</emphasis>
|
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.
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<emphasis id="B91CEA9D6A47073FFE43F8C8E3088528" box="[468,547,1854,1877]" italics="true" pageId="6" pageNumber="417">micans</emphasis>
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</taxonomicName>
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species group into the
|
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<taxonomicName id="4C684D0C6A47073FFCBCF8C8E2EB8528" baseAuthorityName="Germar" baseAuthorityYear="1824" box="[811,960,1853,1877]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="crinicollis">
|
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<emphasis id="B91CEA9D6A47073FFCBCF8C8E2148528" box="[811,831,1854,1877]" italics="true" pageId="6" pageNumber="417">D</emphasis>
|
||
.
|
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<emphasis id="B91CEA9D6A47073FFCD9F8CBE2EB8528" box="[846,960,1853,1877]" italics="true" pageId="6" pageNumber="417">crinicollis</emphasis>
|
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</taxonomicName>
|
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species group; 2) the incorporation of the
|
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<taxonomicName id="4C684D0C6A47073FFF00F894E06D8504" baseAuthorityName="Luederwaldt" baseAuthorityYear="1931" box="[151,326,1889,1913]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="interstitialis">
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<emphasis id="B91CEA9D6A47073FFF00F894E1808504" box="[151,171,1890,1913]" italics="true" pageId="6" pageNumber="417">D</emphasis>
|
||
.
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<emphasis id="B91CEA9D6A47073FFF28F897E06D8504" box="[191,326,1889,1913]" italics="true" pageId="6" pageNumber="417">interstitialis</emphasis>
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</taxonomicName>
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species group into the
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<taxonomicName id="4C684D0C6A47073FFDFEF894E2318504" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[617,794,1889,1913]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="carbonarius">
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<emphasis id="B91CEA9D6A47073FFDFEF894E3568504" box="[617,637,1890,1913]" italics="true" pageId="6" pageNumber="417">D</emphasis>
|
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.
|
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<emphasis id="B91CEA9D6A47073FFD06F897E2318504" box="[657,794,1889,1913]" italics="true" pageId="6" pageNumber="417">carbonarius</emphasis>
|
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</taxonomicName>
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species group (see above); 3) the splitting of the
|
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<taxonomicName id="4C684D0C6A47073FFA16F894E02185E0" baseAuthorityName="Arrow" baseAuthorityYear="1913" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="geminatus">
|
||
<emphasis id="B91CEA9D6A47073FFA16F894E4BE8504" box="[1409,1429,1890,1913]" italics="true" pageId="6" pageNumber="417">D</emphasis>
|
||
.
|
||
<emphasis id="B91CEA9D6A47073FFF00F870E02185E0" box="[151,266,1926,1949]" italics="true" pageId="6" pageNumber="417">geminatus</emphasis>
|
||
</taxonomicName>
|
||
species group from the
|
||
<taxonomicName id="4C684D0C6A47073FFDB2F870E38985E0" baseAuthorityName="Erichson" baseAuthorityYear="1847" box="[549,674,1925,1949]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="inachus">
|
||
<emphasis id="B91CEA9D6A47073FFDB2F870E31285E0" box="[549,569,1926,1949]" italics="true" pageId="6" pageNumber="417">D</emphasis>
|
||
.
|
||
<emphasis id="B91CEA9D6A47073FFDDCF873E38985E0" box="[587,674,1925,1949]" italics="true" pageId="6" pageNumber="417">inachus</emphasis>
|
||
</taxonomicName>
|
||
species group; and 4) the addition of three monobasic groups,
|
||
<emphasis id="B91CEA9D6A47073FFA16F870E33885BC" italics="true" pageId="6" pageNumber="417">
|
||
<taxonomicName id="4C684D0C6A47073FFA16F870E1D785BC" baseAuthorityName="Felsche" baseAuthorityYear="1901" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="superbus">D. superbus</taxonomicName>
|
||
(see above),
|
||
<taxonomicName id="4C684D0C6A47073FFE00F85CE33885BC" baseAuthorityName="Luederwaldt" baseAuthorityYear="1922" box="[407,531,1962,1985]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="muticus">D. muticus</taxonomicName>
|
||
</emphasis>
|
||
(from the
|
||
<taxonomicName id="4C684D0C6A47073FFD07F85CE22085BC" baseAuthorityName="Erichson" baseAuthorityYear="1847" box="[656,779,1961,1985]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="inachus">
|
||
<emphasis id="B91CEA9D6A47073FFD07F85CE38F85BC" box="[656,676,1962,1985]" italics="true" pageId="6" pageNumber="417">D</emphasis>
|
||
.
|
||
<emphasis id="B91CEA9D6A47073FFD23F85FE22085BC" box="[692,779,1961,1985]" italics="true" pageId="6" pageNumber="417">inachus</emphasis>
|
||
</taxonomicName>
|
||
species group), and
|
||
<taxonomicName id="4C684D0C6A47073FFC6CF85CE59B85BC" authorityName="Pereira & D'Andretta" authorityYear="1955" box="[1019,1200,1961,1985]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="eucranioides">
|
||
<emphasis id="B91CEA9D6A47073FFC6CF85CE59B85BC" box="[1019,1200,1961,1985]" italics="true" pageId="6" pageNumber="417">D. eucranioides</emphasis>
|
||
</taxonomicName>
|
||
(from the
|
||
<taxonomicName id="4C684D0C6A47073FFABAF85CE4B085BC" baseAuthorityName="Eschscholtz" baseAuthorityYear="1822" box="[1325,1435,1961,1985]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="assifer">
|
||
<emphasis id="B91CEA9D6A47073FFABAF85CE46A85BC" box="[1325,1345,1962,1985]" italics="true" pageId="6" pageNumber="417">D</emphasis>
|
||
.
|
||
<emphasis id="B91CEA9D6A47073FFAC6F85FE4B085BC" box="[1361,1435,1961,1985]" italics="true" pageId="6" pageNumber="417">assifer</emphasis>
|
||
</taxonomicName>
|
||
species group).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37265046A46073EFF50FF6EE32383D1" pageId="7" pageNumber="418" type="multiple">
|
||
<paragraph id="8BD7368F6A46073EFF50FF6EE32383D1" blockId="7.[151,1436,151,428]" pageId="7" pageNumber="418">
|
||
We stress that we consider those groups as a starting point for further taxonomic reviews in
|
||
<taxonomicName id="4C684D0C6A46073EFB4BFF6EE4BE82CD" box="[1244,1429,152,176]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="417" phylum="Arthropoda" rank="subGenus" subGenus="Luederwaldtinia">
|
||
<emphasis id="B91CEA9D6A46073EFB4BFF6EE4BE82CD" box="[1244,1429,152,176]" italics="true" pageId="7" pageNumber="418">Luederwaldtinia</emphasis>
|
||
</taxonomicName>
|
||
. Currently, the
|
||
<taxonomicName id="4C684D0C6A46073EFEAAFF4BE0F882A9" baseAuthorityName="Germar" baseAuthorityYear="1824" box="[317,467,188,212]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="crinicollis">
|
||
<emphasis id="B91CEA9D6A46073EFEAAFF4BE07A82A9" box="[317,337,189,212]" italics="true" pageId="7" pageNumber="418">D</emphasis>
|
||
.
|
||
<emphasis id="B91CEA9D6A46073EFEF5FF4AE0F882A9" box="[354,467,188,212]" italics="true" pageId="7" pageNumber="418">crinicollis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C684D0C6A46073EFE73FF4BE35882A9" baseAuthorityName="Waterhouse" baseAuthorityYear="1891" box="[484,627,189,212]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="speciosus">
|
||
<emphasis id="B91CEA9D6A46073EFE73FF4BE0D382A9" box="[484,504,189,212]" italics="true" pageId="7" pageNumber="418">D</emphasis>
|
||
.
|
||
<emphasis id="B91CEA9D6A46073EFD9FFF4BE35882A9" box="[520,627,189,212]" italics="true" pageId="7" pageNumber="418">speciosus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C684D0C6A46073EFD21FF4BE20D82A9" baseAuthorityName="Eschscholtz" baseAuthorityYear="1822" box="[694,806,188,212]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="assifer">
|
||
<emphasis id="B91CEA9D6A46073EFD21FF4BE3E182A9" box="[694,714,189,212]" italics="true" pageId="7" pageNumber="418">D</emphasis>
|
||
.
|
||
<emphasis id="B91CEA9D6A46073EFD4CFF4AE20D82A9" box="[731,806,188,212]" italics="true" pageId="7" pageNumber="418">assifer</emphasis>
|
||
</taxonomicName>
|
||
species groups and the positioning of
|
||
<taxonomicName id="4C684D0C6A46073EFB71FF4BE45B82A9" baseAuthorityName="Felsche" baseAuthorityYear="1901" box="[1254,1392,188,212]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="superbus">
|
||
<emphasis id="B91CEA9D6A46073EFB71FF4BE45B82A9" box="[1254,1392,188,212]" italics="true" pageId="7" pageNumber="418">D. superbus</emphasis>
|
||
</taxonomicName>
|
||
are under review by other authors. We do not include a list of species of each group since there are many
|
||
<typeStatus id="54D3882D6A46073EFAB7FF16E4708285" box="[1312,1371,224,248]" pageId="7" pageNumber="418">types</typeStatus>
|
||
to be examined. However, we are confident on the names used here as we have either examined
|
||
<typeStatus id="54D3882D6A46073EFB5DFEF3E5D18360" box="[1226,1274,261,285]" pageId="7" pageNumber="418">type</typeStatus>
|
||
specimens or photographs of
|
||
<typeStatus id="54D3882D6A46073EFEDCFEDEE050833D" box="[331,379,296,320]" pageId="7" pageNumber="418">type</typeStatus>
|
||
specimens, or have conclusive information from the original descriptions. Following key is based on characteristics of both sexes but mainly on male characters. Females are distinguished from males by the last abdominal sternite, which is expanded or at least not shortened medially as in males. When not specified on the key, characters fit for both sexes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37265046A46073EFF00FE2DE4BE850D" pageId="7" pageNumber="418" type="key">
|
||
<key id="FFF961BE6A46073EFF00FE2DE4BE850D" pageId="7" pageNumber="418">
|
||
<keyStep id="309C2E2A6A46073EFF00FE2DE4B78073" pageId="7" pageNumber="418">
|
||
<paragraph id="8BD7368F6A46073EFF00FE2DE4B7838D" blockId="7.[151,1437,475,1905]" box="[151,1436,475,496]" pageId="7" pageNumber="418">
|
||
<keyLead id="309995BA6A46073EFF00FE2DE4B7838D" box="[151,1436,475,496]" pageId="7" pageNumber="418">
|
||
1 Tooth delimiting clypeogenal junction or positioned near it (
|
||
<figureCitation id="13532A0A6A46073EFCB5FE2DE24E838D" box="[802,869,475,496]" captionStart="FIGURE 2" captionStartId="6.[151,250,1417,1440]" captionTargetBox="[185,1391,209,1379]" captionTargetId="figure@6.[167,1403,195,1387]" captionTargetPageId="6" captionText="FIGURE 2. Dichotomius eucranioides (A) black arrow indicating crenulated clypeal margin; D. sexdentatus (B) black arrow indicating tooth or protuberance on clypeogenal junction (this is also seen on D. eucranioides); D. nisus (C) black arrow indicating the asymmetry of the parameres; D. crinicollis (D) black arrows indicating pilosity on the hypomeron; D. superbus (E) white arrow indicating gibbosities on the pronotum; D. schiffleri (D. sericeus species group) (F) white arrow indicating a fovea on the posterior angle of the pronotum; D. speciosus (G) black arrows indicating cephalic transversal carina; D. agenor (H) black arrow indicating male cephalic carina; D. glaucus (D. carbornarius species group) (I) elytra microsculpture." httpUri="https://zenodo.org/record/224053/files/figure.png" pageId="7" pageNumber="418">Figs. 2</figureCitation>
|
||
A, 2B)........................................ 2
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD7368F6A46073EFF00FE0FE4B78073" blockId="7.[151,1437,475,1905]" box="[151,1436,505,526]" pageId="7" pageNumber="418">
|
||
<keyLead id="309995BA6A46073EFF00FE0FE4B78073" box="[151,1436,505,526]" pageId="7" pageNumber="418">- Clypeogenal junction lacking tooth...................................................................... 3</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309C2E2A6A46073EFF00FDE0E4B78168" pageId="7" pageNumber="418">
|
||
<paragraph id="8BD7368F6A46073EFF00FDE0E4B780DD" blockId="7.[151,1437,475,1905]" pageId="7" pageNumber="418">
|
||
<keyLead id="309995BA6A46073EFF00FDE0E4B780DD" pageId="7" pageNumber="418">
|
||
2 Clypeal margin crenulate (
|
||
<figureCitation id="13532A0A6A46073EFE71FDE0E30A8056" box="[486,545,534,555]" captionStart="FIGURE 2" captionStartId="6.[151,250,1417,1440]" captionTargetBox="[185,1391,209,1379]" captionTargetId="figure@6.[167,1403,195,1387]" captionTargetPageId="6" captionText="FIGURE 2. Dichotomius eucranioides (A) black arrow indicating crenulated clypeal margin; D. sexdentatus (B) black arrow indicating tooth or protuberance on clypeogenal junction (this is also seen on D. eucranioides); D. nisus (C) black arrow indicating the asymmetry of the parameres; D. crinicollis (D) black arrows indicating pilosity on the hypomeron; D. superbus (E) white arrow indicating gibbosities on the pronotum; D. schiffleri (D. sericeus species group) (F) white arrow indicating a fovea on the posterior angle of the pronotum; D. speciosus (G) black arrows indicating cephalic transversal carina; D. agenor (H) black arrow indicating male cephalic carina; D. glaucus (D. carbornarius species group) (I) elytra microsculpture." httpUri="https://zenodo.org/record/224053/files/figure.png" pageId="7" pageNumber="418">Fig. 2</figureCitation>
|
||
A). Males with strong transversal carina on head with width subequal to interocular space. Male pronotal disc delimited anteriorly by four gibbosities. Female pronotal disc with two lateral groups of ocellate punctures. Pronotum of both sexes with longitudinal sulcus filled with ocellate punctures. Pronotum wider than elytra. Elytral suture fused.
|
||
<collectingCountry id="F37F761F6A46073EFEB6FD98E07A80FE" box="[289,337,622,643]" name="India" pageId="7" pageNumber="418">Hind</collectingCountry>
|
||
wing reduced, flightless species...................................................................................................
|
||
<emphasis id="B91CEA9D6A46073EFDEFFD7DE4B780DD" bold="true" box="[632,1436,651,672]" pageId="7" pageNumber="418">
|
||
<taxonomicName id="4C684D0C6A46073EFDEFFD7DE23880DD" ID-CoL="35NBK" authorityName="Pereira & D'Andretta" authorityYear="1955" box="[632,787,651,672]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="eucranioides">
|
||
<emphasis id="B91CEA9D6A46073EFDEFFD7DE3A280DD" bold="true" box="[632,649,651,672]" italics="true" pageId="7" pageNumber="418">D</emphasis>
|
||
.
|
||
<emphasis id="B91CEA9D6A46073EFD02FD7DE23880DD" bold="true" box="[661,787,651,672]" italics="true" pageId="7" pageNumber="418">eucranioides</emphasis>
|
||
</taxonomicName>
|
||
species group:
|
||
<taxonomicName id="4C684D0C6A46073EFC26FD7DE4B780DD" ID-CoL="35NBK" authority="Pereira & D'Andretta" authorityName="Pereira & D'Andretta" authorityYear="1955" box="[945,1436,651,672]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="eucranioides">
|
||
<emphasis id="B91CEA9D6A46073EFC26FD7DE59F80DD" bold="true" box="[945,1204,651,672]" italics="true" pageId="7" pageNumber="418">Dichotomius eucranioides</emphasis>
|
||
Pereira & D’Andretta
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD7368F6A46073EFF00FD5EE4B78168" blockId="7.[151,1437,475,1905]" pageId="7" pageNumber="418">
|
||
<keyLead id="309995BA6A46073EFF00FD5EE4B78168" pageId="7" pageNumber="418">
|
||
<emphasis id="B91CEA9D6A46073EFF00FD5EE1B480C0" bold="true" box="[151,159,680,701]" pageId="7" pageNumber="418">-</emphasis>
|
||
Clypeal margin smooth. Males with transversal carina on head higher than wide. Pronotal disc smooth, transversally roughened at most. Pronotum with simple longitudinal sulcus lacking ocellate punctures. Metasternal borders, mesepisternum, and hypomeron with abundant red pilosity (
|
||
<figureCitation id="13532A0A6A46073EFDCBFD12E3BD8084" box="[604,662,740,761]" captionStart="FIGURE 2" captionStartId="6.[151,250,1417,1440]" captionTargetBox="[185,1391,209,1379]" captionTargetId="figure@6.[167,1403,195,1387]" captionTargetPageId="6" captionText="FIGURE 2. Dichotomius eucranioides (A) black arrow indicating crenulated clypeal margin; D. sexdentatus (B) black arrow indicating tooth or protuberance on clypeogenal junction (this is also seen on D. eucranioides); D. nisus (C) black arrow indicating the asymmetry of the parameres; D. crinicollis (D) black arrows indicating pilosity on the hypomeron; D. superbus (E) white arrow indicating gibbosities on the pronotum; D. schiffleri (D. sericeus species group) (F) white arrow indicating a fovea on the posterior angle of the pronotum; D. speciosus (G) black arrows indicating cephalic transversal carina; D. agenor (H) black arrow indicating male cephalic carina; D. glaucus (D. carbornarius species group) (I) elytra microsculpture." httpUri="https://zenodo.org/record/224053/files/figure.png" pageId="7" pageNumber="418">Fig. 2</figureCitation>
|
||
D). Pilosity of hypomeron generally curved upwards....................................................................................................
|
||
<emphasis id="B91CEA9D6A46073EFB11FCF7E4B78168" bold="true" box="[1158,1436,768,790]" pageId="7" pageNumber="418">
|
||
<taxonomicName id="4C684D0C6A46073EFB11FCF7E427816B" box="[1158,1292,768,790]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="crinicolllis">
|
||
<emphasis id="B91CEA9D6A46073EFB11FCF7E5BC816B" bold="true" box="[1158,1175,769,790]" italics="true" pageId="7" pageNumber="418">D</emphasis>
|
||
.
|
||
<emphasis id="B91CEA9D6A46073EFB34FCF7E427816B" bold="true" box="[1187,1292,769,790]" italics="true" pageId="7" pageNumber="418">crinicolllis</emphasis>
|
||
</taxonomicName>
|
||
species group
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309C2E2A6A46073EFF00FCE8E4B78113" pageId="7" pageNumber="418">
|
||
<paragraph id="8BD7368F6A46073EFF00FCE8E4B7812D" blockId="7.[151,1437,475,1905]" pageId="7" pageNumber="418">
|
||
<keyLead id="309995BA6A46073EFF00FCE8E4B7812D" pageId="7" pageNumber="418">
|
||
3 Posterior pronotal margin with well-defined fovea with sharp edges at each posterior angle. Pronotum evenly rounded in both sexes (
|
||
<figureCitation id="13532A0A6A46073EFEB5FCCAE070812C" box="[290,347,828,849]" captionStart="FIGURE 2" captionStartId="6.[151,250,1417,1440]" captionTargetBox="[185,1391,209,1379]" captionTargetId="figure@6.[167,1403,195,1387]" captionTargetPageId="6" captionText="FIGURE 2. Dichotomius eucranioides (A) black arrow indicating crenulated clypeal margin; D. sexdentatus (B) black arrow indicating tooth or protuberance on clypeogenal junction (this is also seen on D. eucranioides); D. nisus (C) black arrow indicating the asymmetry of the parameres; D. crinicollis (D) black arrows indicating pilosity on the hypomeron; D. superbus (E) white arrow indicating gibbosities on the pronotum; D. schiffleri (D. sericeus species group) (F) white arrow indicating a fovea on the posterior angle of the pronotum; D. speciosus (G) black arrows indicating cephalic transversal carina; D. agenor (H) black arrow indicating male cephalic carina; D. glaucus (D. carbornarius species group) (I) elytra microsculpture." httpUri="https://zenodo.org/record/224053/files/figure.png" pageId="7" pageNumber="418">Fig. 2</figureCitation>
|
||
F)..................................................................
|
||
<emphasis id="B91CEA9D6A46073EFB08FCCDE4B7812D" bold="true" box="[1183,1436,827,848]" pageId="7" pageNumber="418">
|
||
<taxonomicName id="4C684D0C6A46073EFB08FCCDE427812D" ID-CoL="35NCX" baseAuthorityName="Harold" baseAuthorityYear="1867" box="[1183,1292,827,848]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="sericeus">
|
||
<emphasis id="B91CEA9D6A46073EFB08FCCDE59B812D" bold="true" box="[1183,1200,827,848]" italics="true" pageId="7" pageNumber="418">D</emphasis>
|
||
.
|
||
<emphasis id="B91CEA9D6A46073EFB2BFCCDE427812D" bold="true" box="[1212,1292,827,848]" italics="true" pageId="7" pageNumber="418">sericeus</emphasis>
|
||
</taxonomicName>
|
||
species group
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD7368F6A46073EFF00FCAFE4B78113" blockId="7.[151,1437,475,1905]" box="[151,1436,857,878]" pageId="7" pageNumber="418">
|
||
<keyLead id="309995BA6A46073EFF00FCAFE4B78113" box="[151,1436,857,878]" pageId="7" pageNumber="418">- Posterior pronotal margin lacking foveae.................................................................. 4</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309C2E2A6A46073EFF00FC80E4B781BB" pageId="7" pageNumber="418">
|
||
<paragraph id="8BD7368F6A46073EFF00FC80E4B781D5" blockId="7.[151,1437,475,1905]" pageId="7" pageNumber="418">
|
||
<keyLead id="309995BA6A46073EFF00FC80E4B781D5" pageId="7" pageNumber="418">
|
||
4 Hypomeron with a well-defined anterior excavation delimited by a dense brush of long, red setae. Male head with high transverse carina, pronotum with one or two median lobes delimiting anterior depression.............
|
||
<emphasis id="B91CEA9D6A46073EFB39FC65E4B781D5" bold="true" box="[1198,1436,915,936]" pageId="7" pageNumber="418">
|
||
<taxonomicName id="4C684D0C6A46073EFB39FC65E42781D5" ID-CoL="35NB5" baseAuthorityName="Eschscholtz" baseAuthorityYear="1822" box="[1198,1292,915,936]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="assifer">
|
||
<emphasis id="B91CEA9D6A46073EFB39FC65E59481D5" bold="true" box="[1198,1215,915,936]" italics="true" pageId="7" pageNumber="418">D</emphasis>
|
||
.
|
||
<emphasis id="B91CEA9D6A46073EFB5CFC65E42781D5" bold="true" box="[1227,1292,915,936]" italics="true" pageId="7" pageNumber="418">assifer</emphasis>
|
||
</taxonomicName>
|
||
species group
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD7368F6A46073EFF00FC46E4B781BB" blockId="7.[151,1437,475,1905]" box="[151,1436,944,966]" pageId="7" pageNumber="418">
|
||
<keyLead id="309995BA6A46073EFF00FC46E4B781BB" box="[151,1436,944,966]" pageId="7" pageNumber="418">
|
||
<emphasis id="B91CEA9D6A46073EFF00FC46E1B481B8" bold="true" box="[151,159,944,965]" pageId="7" pageNumber="418">-</emphasis>
|
||
Hypomeron lacking anteromedial excavation, or excavation very shallow and without medial setal brush............... 5
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309C2E2A6A46073EFF00FC38E4B78624" pageId="7" pageNumber="418">
|
||
<paragraph id="8BD7368F6A46073EFF00FC38E4B78660" blockId="7.[151,1437,475,1905]" pageId="7" pageNumber="418">
|
||
<keyLead id="309995BA6A46073EFF00FC38E4B78660" pageId="7" pageNumber="418">
|
||
5 Body shiny metallic red, purple, blue, green, and/or gold. Metasternal disc concave. Male head with transverse carina bearing single horn at each end (
|
||
<figureCitation id="13532A0A6A46073EFE5EFC1AE329867C" box="[457,514,1004,1025]" captionStart="FIGURE 2" captionStartId="6.[151,250,1417,1440]" captionTargetBox="[185,1391,209,1379]" captionTargetId="figure@6.[167,1403,195,1387]" captionTargetPageId="6" captionText="FIGURE 2. Dichotomius eucranioides (A) black arrow indicating crenulated clypeal margin; D. sexdentatus (B) black arrow indicating tooth or protuberance on clypeogenal junction (this is also seen on D. eucranioides); D. nisus (C) black arrow indicating the asymmetry of the parameres; D. crinicollis (D) black arrows indicating pilosity on the hypomeron; D. superbus (E) white arrow indicating gibbosities on the pronotum; D. schiffleri (D. sericeus species group) (F) white arrow indicating a fovea on the posterior angle of the pronotum; D. speciosus (G) black arrows indicating cephalic transversal carina; D. agenor (H) black arrow indicating male cephalic carina; D. glaucus (D. carbornarius species group) (I) elytra microsculpture." httpUri="https://zenodo.org/record/224053/files/figure.png" pageId="7" pageNumber="418">Fig. 2</figureCitation>
|
||
G); pronotum with a central, well-defined, usually bifurcate lobe. Pronotal disc finely punctured. Length less than
|
||
<quantity id="4C909B6A6A46073EFE13FBFFE0EC8663" box="[388,455,1033,1054]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="7" pageNumber="418" unit="mm" value="16.0">16 mm</quantity>
|
||
..........................................................
|
||
<emphasis id="B91CEA9D6A46073EFB05FBFFE4B78660" bold="true" box="[1170,1436,1032,1054]" pageId="7" pageNumber="418">
|
||
<taxonomicName id="4C684D0C6A46073EFB05FBFFE4278663" ID-CoL="35ND3" baseAuthorityName="Waterhouse" baseAuthorityYear="1891" box="[1170,1292,1032,1054]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="speciosus">
|
||
<emphasis id="B91CEA9D6A46073EFB05FBFFE5888663" bold="true" box="[1170,1187,1033,1054]" italics="true" pageId="7" pageNumber="418">D</emphasis>
|
||
.
|
||
<emphasis id="B91CEA9D6A46073EFB38FBFFE4278663" bold="true" box="[1199,1292,1033,1054]" italics="true" pageId="7" pageNumber="418">speciosus</emphasis>
|
||
</taxonomicName>
|
||
species group
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD7368F6A46073EFF00FBD0E4B78624" blockId="7.[151,1437,475,1905]" pageId="7" pageNumber="418">
|
||
<keyLead id="309995BA6A46073EFF00FBD0E4B78624" pageId="7" pageNumber="418">
|
||
<emphasis id="B91CEA9D6A46073EFF00FBD0E1B48646" bold="true" box="[151,159,1062,1083]" pageId="7" pageNumber="418">-</emphasis>
|
||
Variable color and shiny. Metasternal disc convex. If transversal carina is present on head, then it does not have single horns at each end. Pronotum with variable shape, never with a central bifurcate lobe. Variable length.......................... 6
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309C2E2A6A46073EFF00FB97E4B786B3" pageId="7" pageNumber="418">
|
||
<paragraph id="8BD7368F6A46073EFF00FB97E4B786EE" blockId="7.[151,1437,475,1905]" pageId="7" pageNumber="418">
|
||
<keyLead id="309995BA6A46073EFF00FB97E4B786EE" pageId="7" pageNumber="418">
|
||
6 Green to copper color. Pronotum with strong anterior depression with bilobed summit (
|
||
<figureCitation id="13532A0A6A46073EFBB0FB97E54A860B" box="[1063,1121,1121,1142]" captionStart="FIGURE 2" captionStartId="6.[151,250,1417,1440]" captionTargetBox="[185,1391,209,1379]" captionTargetId="figure@6.[167,1403,195,1387]" captionTargetPageId="6" captionText="FIGURE 2. Dichotomius eucranioides (A) black arrow indicating crenulated clypeal margin; D. sexdentatus (B) black arrow indicating tooth or protuberance on clypeogenal junction (this is also seen on D. eucranioides); D. nisus (C) black arrow indicating the asymmetry of the parameres; D. crinicollis (D) black arrows indicating pilosity on the hypomeron; D. superbus (E) white arrow indicating gibbosities on the pronotum; D. schiffleri (D. sericeus species group) (F) white arrow indicating a fovea on the posterior angle of the pronotum; D. speciosus (G) black arrows indicating cephalic transversal carina; D. agenor (H) black arrow indicating male cephalic carina; D. glaucus (D. carbornarius species group) (I) elytra microsculpture." httpUri="https://zenodo.org/record/224053/files/figure.png" pageId="7" pageNumber="418">Fig. 2</figureCitation>
|
||
E). Male head with conical horn, female with two closely set tubercles............................................
|
||
<taxonomicName id="4C684D0C6A46073EFBCBFB88E4B786EE" ID-CoL="35ND4" authority="(Felsche)" authorityName="Felsche" baseAuthorityName="Felsche" baseAuthorityYear="1901" box="[1116,1436,1150,1171]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="superbus">
|
||
<emphasis id="B91CEA9D6A46073EFBCBFB88E4B786EE" bold="true" box="[1116,1436,1150,1171]" pageId="7" pageNumber="418">
|
||
<emphasis id="B91CEA9D6A46073EFBCBFB88E41186EE" bold="true" box="[1116,1338,1150,1171]" italics="true" pageId="7" pageNumber="418">Dichotomius superbus</emphasis>
|
||
(Felsche)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD7368F6A46073EFF00FB6DE4B786B3" blockId="7.[151,1437,475,1905]" pageId="7" pageNumber="418">
|
||
<keyLead id="309995BA6A46073EFF00FB6DE4B786B3" pageId="7" pageNumber="418">
|
||
<emphasis id="B91CEA9D6A46073EFF00FB6DE1B486CD" bold="true" box="[151,159,1179,1200]" pageId="7" pageNumber="418">-</emphasis>
|
||
If body color is green to copper, than pronotum is simple, lacking lobe or anterior depression. If pronotal has anterior depressions or lobes, then body color is black or brown............................................................ 7
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309C2E2A6A46073EFF00FB20E4B7871C" pageId="7" pageNumber="418">
|
||
<paragraph id="8BD7368F6A46073EFF00FB20E4B78758" blockId="7.[151,1437,475,1905]" pageId="7" pageNumber="418">
|
||
<keyLead id="309995BA6A46073EFF00FB20E4B78758" pageId="7" pageNumber="418">
|
||
<specimenCount id="9D6EFD066A46073EFF00FB20E0398696" box="[151,274,1238,1259]" pageId="7" pageNumber="418" type="male">7 Male</specimenCount>
|
||
head with high transverse carina bearing strong medial tooth. Female head with transverse bituberculate carina, lacking medial tooth. Pronotum of both sexes simple, at most with a small knob on anterior portion and with fine punctures.........................................................................................
|
||
<emphasis id="B91CEA9D6A46073EFB3DFAE7E4B78758" bold="true" box="[1194,1436,1296,1318]" pageId="7" pageNumber="418">
|
||
<taxonomicName id="4C684D0C6A46073EFB3DFAE7E427875B" ID-CoL="6CS36" baseAuthorityName="Harold" baseAuthorityYear="1869" box="[1194,1292,1296,1318]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="agenor">
|
||
<emphasis id="B91CEA9D6A46073EFB3DFAE7E590875B" bold="true" box="[1194,1211,1297,1318]" italics="true" pageId="7" pageNumber="418">D</emphasis>
|
||
.
|
||
<emphasis id="B91CEA9D6A46073EFB50FAE7E427875B" bold="true" box="[1223,1292,1297,1318]" italics="true" pageId="7" pageNumber="418">agenor</emphasis>
|
||
</taxonomicName>
|
||
species group
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD7368F6A46073EFF00FAD8E4B7871C" blockId="7.[151,1437,475,1905]" pageId="7" pageNumber="418">
|
||
<keyLead id="309995BA6A46073EFF00FAD8E4B7871C" pageId="7" pageNumber="418">
|
||
<emphasis id="B91CEA9D6A46073EFF00FAD8E1B4873E" bold="true" box="[151,159,1326,1347]" pageId="7" pageNumber="418">-</emphasis>
|
||
Male and female head variable, but never with transverse carina bearing medial tooth. If the head has a head transverse carina, then the pronotum bears large punctures, strong knobs, or depressions or the elytra is chagreened...................... 8
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309C2E2A6A46073EFF00FA9FE4BE846C" pageId="7" pageNumber="418">
|
||
<paragraph id="8BD7368F6A46073EFF00FA9FE4B787C5" blockId="7.[151,1437,475,1905]" pageId="7" pageNumber="418">
|
||
<keyLead id="309995BA6A46073EFF00FA9FE4B787C5" pageId="7" pageNumber="418">
|
||
8 Metasternal sides, mesepisternum, and hypomeron with dense, long, red to tan pilosity. Pronotum of large males with welldefined anterior depression bearing large punctures anteriorly delimited posteriorly by bilobed knob. Males with left paramere of aedeagus with strong apical hook (
|
||
<figureCitation id="13532A0A6A46073EFDD0FA52E3AB87C4" box="[583,640,1444,1465]" captionStart="FIGURE 2" captionStartId="6.[151,250,1417,1440]" captionTargetBox="[185,1391,209,1379]" captionTargetId="figure@6.[167,1403,195,1387]" captionTargetPageId="6" captionText="FIGURE 2. Dichotomius eucranioides (A) black arrow indicating crenulated clypeal margin; D. sexdentatus (B) black arrow indicating tooth or protuberance on clypeogenal junction (this is also seen on D. eucranioides); D. nisus (C) black arrow indicating the asymmetry of the parameres; D. crinicollis (D) black arrows indicating pilosity on the hypomeron; D. superbus (E) white arrow indicating gibbosities on the pronotum; D. schiffleri (D. sericeus species group) (F) white arrow indicating a fovea on the posterior angle of the pronotum; D. speciosus (G) black arrows indicating cephalic transversal carina; D. agenor (H) black arrow indicating male cephalic carina; D. glaucus (D. carbornarius species group) (I) elytra microsculpture." httpUri="https://zenodo.org/record/224053/files/figure.png" pageId="7" pageNumber="418">Fig. 2</figureCitation>
|
||
C)....................
|
||
<emphasis id="B91CEA9D6A46073EFC00FA55E4B787C5" bold="true" box="[919,1436,1443,1464]" pageId="7" pageNumber="418">
|
||
<taxonomicName id="4C684D0C6A46073EFC00FA55E2C387C5" ID-CoL="35NCJ" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[919,1000,1443,1464]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="nisus">
|
||
<emphasis id="B91CEA9D6A46073EFC00FA55E28387C5" bold="true" box="[919,936,1443,1464]" italics="true" pageId="7" pageNumber="418">D</emphasis>
|
||
.
|
||
<emphasis id="B91CEA9D6A46073EFC23FA55E2C387C5" bold="true" box="[948,1000,1443,1464]" italics="true" pageId="7" pageNumber="418">nisus</emphasis>
|
||
</taxonomicName>
|
||
species group:
|
||
<taxonomicName id="4C684D0C6A46073EFB12FA55E4B787C5" ID-CoL="35NCJ" authority="(Olivier)" authorityName="Olivier" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[1157,1436,1443,1464]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="nisus">
|
||
<emphasis id="B91CEA9D6A46073EFB12FA55E41587C5" bold="true" box="[1157,1342,1443,1464]" italics="true" pageId="7" pageNumber="418">Dichotomius nisus</emphasis>
|
||
(Olivier)
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD7368F6A46073EFF00FA36E4BE846C" blockId="7.[151,1437,475,1905]" pageId="7" pageNumber="418">
|
||
<keyLead id="309995BA6A46073EFF00FA36E4BE846C" pageId="7" pageNumber="418">
|
||
<emphasis id="B91CEA9D6A46073EFF00FA36E1B487A8" bold="true" box="[151,159,1472,1493]" pageId="7" pageNumber="418">-</emphasis>
|
||
Dense and long red to tan pilosity lacking in one or more of the following areas: mestasternal sides, mesepisternum, or hypomeron. Pronotum of large males with at most weakly-defined anterior depression, lacking puncturing and delimited posteriorly by weakly-defined knob. Males with left paramere lacking apicolateral hook............................... 9
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309C2E2A6A46073EFF00F9EFE4B78436" pageId="7" pageNumber="418">
|
||
<paragraph id="8BD7368F6A46073EFF00F9EFE4B78450" blockId="7.[151,1437,475,1905]" box="[151,1436,1560,1582]" pageId="7" pageNumber="418">
|
||
<keyLead id="309995BA6A46073EFF00F9EFE4B78450" box="[151,1436,1560,1582]" pageId="7" pageNumber="418">
|
||
9 Pronotal disc with ocellate punctures (
|
||
<figureCitation id="13532A0A6A46073EFDC7F9EFE3AD8453" box="[592,646,1561,1582]" captionStart="FIGURE 1" captionStartId="3.[151,250,1932,1955]" captionTargetBox="[238,1366,207,1900]" captionTargetId="figure@3.[209,1377,193,1917]" captionTargetPageId="3" captionText="FIGURE 1. Dichotomius mysticus (A) dorsal habitus; (B) dorsal view of genitalia; (C) hind wing; and (D) tarsi and tarsal claw (indicated by black arrow). Dichotomius vidaurrei (E) dorsal habitus; (F) dorsal view of genitalia; and (G) hind wing. Scale bar = 1.5 mm." httpUri="https://zenodo.org/record/224052/files/figure.png" pageId="7" pageNumber="418">Fig. 1</figureCitation>
|
||
E, see pronotal disc)............................
|
||
<emphasis id="B91CEA9D6A46073EFB21F9EFE4B78450" bold="true" box="[1206,1436,1560,1582]" pageId="7" pageNumber="418">
|
||
<taxonomicName id="4C684D0C6A46073EFB21F9EFE4268453" ID-CoL="35NB6" baseAuthorityName="Harold" baseAuthorityYear="1869" box="[1206,1293,1560,1582]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="batesi">
|
||
<emphasis id="B91CEA9D6A46073EFB21F9EFE5EC8453" bold="true" box="[1206,1223,1561,1582]" italics="true" pageId="7" pageNumber="418">D</emphasis>
|
||
.
|
||
<emphasis id="B91CEA9D6A46073EFB44F9EFE4268453" bold="true" box="[1235,1293,1561,1582]" italics="true" pageId="7" pageNumber="418">batesi</emphasis>
|
||
</taxonomicName>
|
||
species group
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD7368F6A46073EFF00F9C0E4B78436" blockId="7.[151,1437,475,1905]" box="[151,1436,1590,1611]" pageId="7" pageNumber="418">
|
||
<keyLead id="309995BA6A46073EFF00F9C0E4B78436" box="[151,1436,1590,1611]" pageId="7" pageNumber="418">- Pronotal disc without ocellate punctures. Ocellate punctures might occur on anterior angles or pronotal margins......... 10</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309C2E2A6A46073EFF00F9A2E4B08486" pageId="7" pageNumber="418">
|
||
<paragraph id="8BD7368F6A46073EFF00F9A2E4B784DE" blockId="7.[151,1437,475,1905]" pageId="7" pageNumber="418">
|
||
<keyLead id="309995BA6A46073EFF00F9A2E4B784DE" pageId="7" pageNumber="418">
|
||
10 Dull black to green-copper metallic species. Pronotum (except sometimes on disc) and elytra evenly chagreened (
|
||
<date id="FFD6104F6A46073EFAA3F9A2E4748414" box="[1332,1375,1620,1641]" pageId="7" pageNumber="418" value="1920-10">20X</date>
|
||
) (
|
||
<figureCitation id="13532A0A6A46073EFAE2F9A2E1C484FB" captionStart="FIGURE 2" captionStartId="6.[151,250,1417,1440]" captionTargetBox="[185,1391,209,1379]" captionTargetId="figure@6.[167,1403,195,1387]" captionTargetPageId="6" captionText="FIGURE 2. Dichotomius eucranioides (A) black arrow indicating crenulated clypeal margin; D. sexdentatus (B) black arrow indicating tooth or protuberance on clypeogenal junction (this is also seen on D. eucranioides); D. nisus (C) black arrow indicating the asymmetry of the parameres; D. crinicollis (D) black arrows indicating pilosity on the hypomeron; D. superbus (E) white arrow indicating gibbosities on the pronotum; D. schiffleri (D. sericeus species group) (F) white arrow indicating a fovea on the posterior angle of the pronotum; D. speciosus (G) black arrows indicating cephalic transversal carina; D. agenor (H) black arrow indicating male cephalic carina; D. glaucus (D. carbornarius species group) (I) elytra microsculpture." httpUri="https://zenodo.org/record/224053/files/figure.png" pageId="7" pageNumber="418">
|
||
Fig.
|
||
<date id="FFD6104F6A46073EFF76F987E1C484FB" box="[225,239,1649,1670]" pageId="7" pageNumber="418">2</date>
|
||
</figureCitation>
|
||
I). Head with either a conical horn, a pair of closely-set tubercles, or a short transverse carina.....................................................................................................
|
||
<emphasis id="B91CEA9D6A46073EFBE0F978E4B784DE" bold="true" box="[1143,1436,1678,1699]" pageId="7" pageNumber="418">
|
||
<taxonomicName id="4C684D0C6A46073EFBE0F978E42784DE" ID-CoL="35NBC" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[1143,1292,1678,1699]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="carbonarius">
|
||
<emphasis id="B91CEA9D6A46073EFBE0F978E5A384DE" bold="true" box="[1143,1160,1678,1699]" italics="true" pageId="7" pageNumber="418">D</emphasis>
|
||
.
|
||
<emphasis id="B91CEA9D6A46073EFB03F978E42784DE" bold="true" box="[1172,1292,1678,1699]" italics="true" pageId="7" pageNumber="418">carbonarius</emphasis>
|
||
</taxonomicName>
|
||
species group
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD7368F6A46073EFF00F95DE4B08486" blockId="7.[151,1437,475,1905]" pageId="7" pageNumber="418">
|
||
<keyLead id="309995BA6A46073EFF00F95DE4B08486" pageId="7" pageNumber="418">
|
||
<emphasis id="B91CEA9D6A46073EFF00F95DE1B484BD" bold="true" box="[151,159,1707,1728]" pageId="7" pageNumber="418">-</emphasis>
|
||
Dorsal surface black, smooth and shiny (if somewhat opaque on elytra, opacity due to punctures or reticulation and not to chagreened). Head generally with a conical horn or tubercle, or simply flat; if with a pair of closely-set tubercles then dorsum strongly shiny........................................................................................ 11
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309C2E2A6A46073EFF00F8F2E4B7854B" pageId="7" pageNumber="418">
|
||
<paragraph id="8BD7368F6A46073EFF00F8F2E4B78565" blockId="7.[151,1437,475,1905]" box="[151,1436,1795,1817]" pageId="7" pageNumber="418">
|
||
<keyLead id="309995BA6A46073EFF00F8F2E4B78565" box="[151,1436,1795,1817]" pageId="7" pageNumber="418">
|
||
11 Hypomeron laterally with sparse, long setae. Hypomeral disc not covered by setae.............
|
||
<emphasis id="B91CEA9D6A46073EFB37F8F5E4B78565" bold="true" box="[1184,1436,1795,1816]" pageId="7" pageNumber="418">
|
||
<taxonomicName id="4C684D0C6A46073EFB37F8F5E4268565" ID-CoL="35NC4" baseAuthorityName="Erichson" baseAuthorityYear="1847" box="[1184,1293,1795,1816]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="inachus">
|
||
<emphasis id="B91CEA9D6A46073EFB37F8F5E59A8565" bold="true" box="[1184,1201,1795,1816]" italics="true" pageId="7" pageNumber="418">D</emphasis>
|
||
.
|
||
<emphasis id="B91CEA9D6A46073EFB29F8F5E4268565" bold="true" box="[1214,1293,1795,1816]" italics="true" pageId="7" pageNumber="418">inachus</emphasis>
|
||
</taxonomicName>
|
||
species group
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD7368F6A46073EFF00F8D6E4B7854B" blockId="7.[151,1437,475,1905]" box="[151,1436,1824,1846]" pageId="7" pageNumber="418">
|
||
<keyLead id="309995BA6A46073EFF00F8D6E4B7854B" box="[151,1436,1824,1846]" pageId="7" pageNumber="418">
|
||
<emphasis id="B91CEA9D6A46073EFF00F8D6E1B48548" bold="true" box="[151,159,1824,1845]" pageId="7" pageNumber="418">-</emphasis>
|
||
Hypomeron laterally with dense, long setae. Most of surface of the hypomeral disc covered with setae.................. 12
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309C2E2A6A46073EFF00F8C8E4BE850D" pageId="7" pageNumber="418">
|
||
<paragraph id="8BD7368F6A46073EFF00F8C8E4B7852E" blockId="7.[151,1437,475,1905]" box="[151,1436,1854,1875]" pageId="7" pageNumber="418">
|
||
<keyLead id="309995BA6A46073EFF00F8C8E4B7852E" box="[151,1436,1854,1875]" pageId="7" pageNumber="418">
|
||
12 Head with a central horn, tubercle, or knob..........................................
|
||
<emphasis id="B91CEA9D6A46073EFB1FF8C8E4B7852E" bold="true" box="[1160,1436,1854,1875]" pageId="7" pageNumber="418">
|
||
<taxonomicName id="4C684D0C6A46073EFB1FF8C8E427852E" ID-CoL="6CRR7" baseAuthorityName="Arrow" baseAuthorityYear="1913" box="[1160,1292,1854,1875]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="geminatus">
|
||
<emphasis id="B91CEA9D6A46073EFB1FF8C8E5B2852E" bold="true" box="[1160,1177,1854,1875]" italics="true" pageId="7" pageNumber="418">D</emphasis>
|
||
.
|
||
<emphasis id="B91CEA9D6A46073EFB31F8C8E427852E" bold="true" box="[1190,1292,1854,1875]" italics="true" pageId="7" pageNumber="418">geminatus</emphasis>
|
||
</taxonomicName>
|
||
species group
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD7368F6A46073EFF00F8ADE4BE850D" blockId="7.[151,1437,475,1905]" box="[151,1429,1883,1905]" pageId="7" pageNumber="418">
|
||
<keyLead id="309995BA6A46073EFF00F8ADE4BE850D" box="[151,1429,1883,1905]" pageId="7" pageNumber="418">
|
||
<emphasis id="B91CEA9D6A46073EFF00F8ADE1B4850D" bold="true" box="[151,159,1883,1904]" pageId="7" pageNumber="418">-</emphasis>
|
||
Head flat, without tubercle, horn, or knob............
|
||
<emphasis id="B91CEA9D6A46073EFC87F8ADE520850D" bold="true" box="[784,1035,1883,1904]" pageId="7" pageNumber="418">
|
||
<taxonomicName id="4C684D0C6A46073EFC87F8ADE257850D" ID-CoL="35NCG" baseAuthorityName="Luederwaldt" baseAuthorityYear="1922" box="[784,892,1883,1904]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="muticus">
|
||
<emphasis id="B91CEA9D6A46073EFC87F8ADE20A850D" bold="true" box="[784,801,1883,1904]" italics="true" pageId="7" pageNumber="418">D</emphasis>
|
||
.
|
||
<emphasis id="B91CEA9D6A46073EFCBAF8ADE257850D" bold="true" box="[813,892,1883,1904]" italics="true" pageId="7" pageNumber="418">muticus</emphasis>
|
||
</taxonomicName>
|
||
species group
|
||
</emphasis>
|
||
...
|
||
<taxonomicName id="4C684D0C6A46073EFBBEF8ADE4BE850D" ID-CoL="35NCG" authority="(Luederwaldt)" authorityName="Luederwaldt" baseAuthorityName="Luederwaldt" baseAuthorityYear="1922" box="[1065,1429,1883,1904]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="muticus">
|
||
<emphasis id="B91CEA9D6A46073EFBBEF8ADE4BE850D" bold="true" box="[1065,1429,1883,1904]" pageId="7" pageNumber="418">
|
||
<emphasis id="B91CEA9D6A46073EFBBEF8ADE5D0850D" bold="true" box="[1065,1275,1883,1904]" italics="true" pageId="7" pageNumber="418">Dichotomius muticus</emphasis>
|
||
(Luederwaldt)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |