170 lines
21 KiB
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170 lines
21 KiB
XML
<document ID-CLB-Dataset="68943" ID-DOI="http://dx.doi.org/10.3897/zitteliana.96.83571" ID-GBIF-Dataset="f614d115-44b1-41ef-89ce-8651629bceb0" ID-Pensoft-Pub="2747-8106-96-103" ID-Pensoft-UUID="4E0188A4D3D7563C980CB7C5B9424F3C" ID-ZooBank="5B10FFF604074E09B986F931635E9BCC" ModsDocID="2747-8106-96-103" checkinTime="1654860115112" checkinUser="pensoft" docAuthor="Schwarzhans, Werner, Ohe, Fumio, Tsuchiya, Yuki & Ujihara, Atsushi" docDate="2022" docId="F2E750DBD50A568ABC186F0B71D6B8DB" docLanguage="en" docName="Zitteliana 96: 103-134" docOrigin="Zitteliana 96" docPubDate="2022-06-10" docSource="http://dx.doi.org/10.3897/zitteliana.96.83571" docTitle="Diaphus watatsumi Schwarzhans & Ohe & Tsuchiya & Ujihara 2022, sp. nov." docType="treatment" docUuid="211177B6-5D3E-4052-808F-C2DC9D55EB37" docUuidSource="ZooBank" docVersion="5" id="4E0188A4D3D7563C980CB7C5B9424F3C" lastPageNumber="103" masterDocId="4E0188A4D3D7563C980CB7C5B9424F3C" masterDocTitle="Lanternfish otoliths (Myctophidae, Teleostei) from the Miocene of Japan" masterLastPageNumber="134" masterPageNumber="103" pageNumber="103" updateTime="1732684795358" updateUser="ExternalLinkService">
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<mods:mods id="6661E65CC5F3860D516EC6496C8A46EF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="3FF58252B0DB77EB57534E5CED811EDF">
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<mods:title id="272F21863371B8ADB5E533361B61391B">Lanternfish otoliths (Myctophidae, Teleostei) from the Miocene of Japan</mods:title>
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</mods:titleInfo>
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<mods:name id="FAA5575EF2581F1B7E3D59EBAE7C334A" type="personal">
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<mods:role id="28E8CBCF72F18F3AEB03590B7D4F9546">
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<mods:roleTerm id="FD71DAB91488E80F8DE912E41AF538CF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4097C5FCD334D2A38D01A71904A52921">Schwarzhans, Werner</mods:namePart>
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<mods:affiliation id="7C91FDA437D1CFEB279EC43030503225">Zoological Museum, Natural History Museum of Denmark, Universitetsparken 15, 2100 Kobenhavn, Denmark & Ahrensburger Weg 103, 22359 Hamburg, Germany</mods:affiliation>
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<mods:nameIdentifier id="BEC43DE3000219B9538821DC29D5B17C" type="email">wwschwarz@aol.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="99CFC6D4D827A8D4C4AE751D6F799DB4" type="personal">
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<mods:roleTerm id="5B9EDF559704F088406FF3362C16A1BB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="35F0FA9FE2A1ABAC44E8C5592E016778">Ohe, Fumio</mods:namePart>
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<mods:affiliation id="BEB1687AB418C19F1207315141A4D961">Nara National Research Institute for Cultural Properties, Nara 630 - 8577, Japan</mods:affiliation>
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</mods:name>
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<mods:name id="EA7181F02D940A2D74411EB769697E21" type="personal">
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<mods:roleTerm id="721BBE12BD53D9A8D5A8E6FEE4EC0980">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F69435F1A192A485F0050AD0FD371549">Tsuchiya, Yuki</mods:namePart>
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<mods:affiliation id="32E0EC080F3CCAFE7D7B1F288AC5C23E">Graduate School of Environmental Studies, Nagoya University, Nagoya, Japan</mods:affiliation>
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</mods:name>
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<mods:name id="6AD50DC0383E7FE79F97FBDC96F36E61" type="personal">
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<mods:roleTerm id="EC8A381E41ECA4C97DBD7B427500351F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8BCF4C8694D5E9DD3A7DDB2AA00DEFCD">Ujihara, Atsushi</mods:namePart>
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<mods:affiliation id="0051AA10C801EB1067B0C6E245669641">Graduate School of Environmental Studies, Nagoya University, Nagoya, Japan</mods:affiliation>
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<mods:typeOfResource id="B88B4C32B3718EEDA418D849F86C81EE">text</mods:typeOfResource>
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<mods:titleInfo id="5EF59C4363178D5377BB257589F5EB1E">
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<mods:title id="A706BF5C558C9DD82F72DA17AA580FC2">Zitteliana</mods:title>
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<mods:part id="8DFBABAE267166FB9FCFB5C4D437DF2D">
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<mods:date id="2336D7784FE5D67158EDAA46FE534D7B">2022</mods:date>
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<mods:number id="C815BEC4DCB4B100F33A6894E7F80A8F">2022-06-10</mods:number>
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<mods:detail id="ECB70463A1C3CA26503020313F8357EB" type="volume">
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<mods:number id="2080A150E060FCDA4FD703C3F0115063">96</mods:number>
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<mods:start id="0FA075954D5060215287CF4DCE97C5AF">103</mods:start>
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<mods:end id="461589BAFDB25D2928575B56F96A8609">134</mods:end>
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<mods:url id="A75B97696215B944F4882332F9AA2C53">http://dx.doi.org/10.3897/zitteliana.96.83571</mods:url>
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<mods:classification id="A3023FFE0D00C8ADAFD200DD7115FCA1">journal article</mods:classification>
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<mods:identifier id="6AFEC7D662D2BBC2C39C8340FB35EF10" type="DOI">http://dx.doi.org/10.3897/zitteliana.96.83571</mods:identifier>
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<treatment id="F2E750DBD50A568ABC186F0B71D6B8DB" ID-GBIF-Taxon="195832369" LSID="urn:lsid:zoobank.org:act:211177B6-5D3E-4052-808F-C2DC9D55EB37" httpUri="http://treatment.plazi.org/id/F2E750DBD50A568ABC186F0B71D6B8DB" lastPageNumber="103" pageId="0" pageNumber="103" scope_family="Myctophidae" scope_order="Myctophiformes" scope_phylum="Teleostei">
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<subSubSection id="3F5233B76C36219F1C132111E209D292" pageId="0" pageNumber="103" type="nomenclature">
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<paragraph id="51F281D88267AEAA1065487B811D68F4" pageId="0" pageNumber="103">
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<taxonomicName id="9DD0AD61B492D355A6105079510D0476" LSID="http://zoobank.org/211177B6-5D3E-4052-808F-C2DC9D55EB37" authority="Schwarzhans & Ohe & Tsuchiya & Ujihara, 2022" authorityName="Schwarzhans & Ohe & Tsuchiya & Ujihara" authorityYear="2022" family="Myctophidae" genus="Diaphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diaphus watatsumi" order="Myctophiformes" pageId="0" pageNumber="103" phylum="Chordata" rank="species" species="watatsumi" status="sp. nov.">Diaphus watatsumi</taxonomicName>
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<taxonomicNameLabel id="9ABF1FC0768AF4ADDFEEC39A9B484F9C" pageId="0" pageNumber="103">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="5AA519B05F1C82EB6BD90D57D06F70DD" pageId="0" pageNumber="103" type="description">
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<paragraph id="57324D63629767C69AD6A33DB015B713" pageId="0" pageNumber="103">
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<figureCitation id="45EFF5E418D973A956CBAF147F2BD600" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-F: Diaphus angulatus Ohe & Araki, 1973; A-C: SMF PO 101.130, Inabacho, Tsu City, Katada Formation, level ML 2305; D-E: SMF PO 101.131, Fuchu machi, Toyama City, Higashibessho Formation, level H 20; G-N: Diaphus epipedus sp. nov.; G-I: holotype, SMF PO 101.132, Tsuyama City, Takakura Formation, level MS 15; K-J: paratype, SMF PO 101.133, Tsuyama City, Takakura Formation, level MS 15; L-N: paratype, SMF PO 101.134, Yatsuo, Toyama City, Higashibessho Formation, level H 21; O-T: Diaphus metopoclampoides Steurbaut, 1983, SMF PO 101.135, Tsuyama City, Takakura Formation level MS 06, f; U-AB: Diaphus watatsumi sp. nov., Tsuyama City, Takakura Formation; U-X: holotype, SMF PO 101.136, level MS 52; Y-AB: paratypes, SMF PO 101.137, levels gh, j; AC-AE: Diaphus sp. 1, col. Tsuchiya, Ichishi Group." figureDoi="10.3897/zitteliana.96.83571.figure8" httpUri="https://binary.pensoft.net/fig/694121" pageId="0" pageNumber="103">Fig. 8U</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="25BA5653F79221655C04FF87A3CB3457" pageId="0" pageNumber="103" type="etymology">
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<paragraph id="D1B9B879FC4370391F5BA177DAC7B9ED" pageId="0" pageNumber="103">Etymology.</paragraph>
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<paragraph id="9629E8779F1325ABE4D0C24CB04FC17D" pageId="0" pageNumber="103">Named after Watatsumi, a Shinto spirit of the sea in Japanese.</paragraph>
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</subSubSection>
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<subSubSection id="B2C4878CB3F3381E528758B5A584C9BC" pageId="0" pageNumber="103" type="holotype">
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<paragraph id="57E2DB8532990FEF7C1A023F36A90834" pageId="0" pageNumber="103">Holotype.</paragraph>
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<paragraph id="3458612139F7326EB74866F7710411ED" pageId="0" pageNumber="103">
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SMF PO 101.136 (Fig.
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<figureCitation id="BBB22798DE69D554E3611A56125D3EAC" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-F: Diaphus angulatus Ohe & Araki, 1973; A-C: SMF PO 101.130, Inabacho, Tsu City, Katada Formation, level ML 2305; D-E: SMF PO 101.131, Fuchu machi, Toyama City, Higashibessho Formation, level H 20; G-N: Diaphus epipedus sp. nov.; G-I: holotype, SMF PO 101.132, Tsuyama City, Takakura Formation, level MS 15; K-J: paratype, SMF PO 101.133, Tsuyama City, Takakura Formation, level MS 15; L-N: paratype, SMF PO 101.134, Yatsuo, Toyama City, Higashibessho Formation, level H 21; O-T: Diaphus metopoclampoides Steurbaut, 1983, SMF PO 101.135, Tsuyama City, Takakura Formation level MS 06, f; U-AB: Diaphus watatsumi sp. nov., Tsuyama City, Takakura Formation; U-X: holotype, SMF PO 101.136, level MS 52; Y-AB: paratypes, SMF PO 101.137, levels gh, j; AC-AE: Diaphus sp. 1, col. Tsuchiya, Ichishi Group." figureDoi="10.3897/zitteliana.96.83571.figure8" httpUri="https://binary.pensoft.net/fig/694121" pageId="0" pageNumber="103">8U-X</figureCitation>
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), Miya River at Numa, Tsuyama City, Okayama Prefecture, Takakura FM, level MS 52, late Burdigalian, diatom zone 3A-3B.
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</paragraph>
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</subSubSection>
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<subSubSection id="C9DECFC131246755DE15C91DBF399D85" pageId="0" pageNumber="103" type="paratypes">
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<paragraph id="2E6C257076BB30E9D96E23E698418C09" pageId="0" pageNumber="103">Paratypes.</paragraph>
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<paragraph id="F6EEAE1EB1B5CE86281C5E815A81434B" pageId="0" pageNumber="103">3 specimens (SMF PO 101.137), same data as holotype, levels gh, h, j.</paragraph>
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</subSubSection>
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<subSubSection id="052333F17D8C6BC8DE644C6D4164F87A" pageId="0" pageNumber="103" type="materials_examined">
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<paragraph id="9AF448ADB599555157561B0114A080EA" pageId="0" pageNumber="103">Additional material.</paragraph>
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<paragraph id="0B76F8298AADAD5D47176A19276EC3A8" pageId="0" pageNumber="103">
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<materialsCitation id="20F1ED0639ED77A5DCB8548F2E1ACAAD" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" specimenCount="7">
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<specimenCount id="5DD7C5ECA8242A025F7A3980EB1485C3" type="generic">7 specimens</specimenCount>
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, same data as holotype, levels MS 15, MS 22, MS 27, MS 36, MS 70, f9, j
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="AD67664F797ACD5384690C4C91C4A4FE" pageId="0" pageNumber="103" type="diagnosis">
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<paragraph id="80F0E0CE0AFB24CA1CBA081897768294" pageId="0" pageNumber="103">Diagnosis.</paragraph>
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<paragraph id="B79300F2677434D397618B720469C8F8" pageId="0" pageNumber="103">OL:OH = 1.35-1.45; OH:OT = 3.8-4.5. Dorsal rim with distinct postdorsal angle above central position of cauda, slightly concave postdorsal section and depressed predorsal section. Rostrum moderately long, 16-18% of OL. Ventral rim with 13-15 feeble denticles. Sulcus long, relatively narrow, slightly bent; OCL:CCL = 2.0.</paragraph>
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</subSubSection>
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<subSubSection id="7285D64F8BD69C7785E263AC4045A5D0" pageId="0" pageNumber="103" type="description">
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<paragraph id="98FCB741608109C1A04133B6FF425BF1" pageId="0" pageNumber="103">Description.</paragraph>
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<paragraph id="9D13405B07C548C0B5DF1300C1144923" pageId="0" pageNumber="103">Slender, thin and relatively large otoliths reaching about 4.1 mm in length (holotype 3.8 mm). Rostrum moderately long and pointed; excisura mostly shallow and narrow, antirostrum much shorter than rostrum. Dorsal rim anteriorly depressed, ascending to distinct postdorsal angle positioned relatively forward above center of cauda; postdorsal section inclined, slightly concave. Ventral rim rather shallow and very regularly bent, with 13-15 feeble denticles which erode easily and are rarely completely preserved like in holotype. Posterior rim gently and regularly curved, slightly crenulated.</paragraph>
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<paragraph id="6351405279989856AC5C77ABC2073F5B" pageId="0" pageNumber="103">Inner face slightly bent in horizontal direction, relatively smooth, with long, slightly supramedian and slightly bent, shallow sulcus. Ostium about as wide as cauda but twice as long. Ostial colliculum only slightly wider than caudal colliculum (OCH:CCH = 1.1-1.3). Caudal pseudocolliculum distinct, long, fine. Dorsal margin of ostium slightly oscillating. Ventral furrow very indistinct; dorsal depression large, well marked towards crista superior. Outer face nearly flat except slightly thickened central and postcentral region, with numerous rapidly fading radial furrows on dorsal and ventral fields.</paragraph>
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</subSubSection>
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<subSubSection id="4605C9F9E953E2C89F664EC09DBE6D8F" pageId="0" pageNumber="103" type="discussion">
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<paragraph id="5AF26A5B8185765BF591ED8FA645B699" pageId="0" pageNumber="103">Discussion.</paragraph>
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<paragraph id="B7058D16B91A6075E78D177EDD5CFEB3" pageId="0" pageNumber="103">
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<taxonomicName id="49BF8EA8AC08DA88D623AFCFD86344A4" authorityName="Schwarzhans & Ohe & Tsuchiya & Ujihara" authorityYear="2022" family="Myctophidae" genus="Diaphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diaphus watatsumi" order="Myctophiformes" pageId="0" pageNumber="103" phylum="Chordata" rank="species" species="watatsumi">
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<emphasis id="690F6D68E46E9C148852ABE99118AA7C" italics="true" pageId="0" pageNumber="103">Diaphus watatsumi</emphasis>
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</taxonomicName>
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is superficially an inconspicuous
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<taxonomicName id="6A519C3D8BA644D9AA99D9C081F6C6E1" family="Myctophidae" genus="Diaphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diaphus" order="Myctophiformes" pageId="0" pageNumber="103" phylum="Chordata" rank="genus">
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<emphasis id="152DB4594BB6390825148339FDD40EAD" italics="true" pageId="0" pageNumber="103">Diaphus</emphasis>
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</taxonomicName>
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otolith but can be relatively easily identified by its proportions and the shape and number of delicate denticles along the ventral rim. It resembles
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<taxonomicName id="D4C7EE82914C40EF9D0E584B926F283E" lsidName="D. marwicki" pageId="0" pageNumber="103" rank="species" species="marwicki">
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<emphasis id="8AA1015866FCC5DC1F255C927355A32B" italics="true" pageId="0" pageNumber="103">D. marwicki</emphasis>
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</taxonomicName>
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(Frost, 1933), a common species in the early and middle Miocene of the southern Pacific in New Zealand and Chile (see
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<bibRefCitation id="028E6A79D65EB6F6080D5A7B9FBC91FB" author="Schwarzhans, W" journalOrPublisher="Memorie Della Societa Italiana Di Scienze Naturali e Del Museon Di Storia Naturale Di Milano" pageId="0" pageNumber="103" pagination="3 - 326" refId="B60" refString="Schwarzhans, W, 2019. Reconstruction of the fossil marine bony fish fauna (Teleostei) from the Eocene to Pleistocene of New Zealand by means of otoliths. Memorie Della Societa Italiana Di Scienze Naturali e Del Museon Di Storia Naturale Di Milano 46: 3 - 326" title="Reconstruction of the fossil marine bony fish fauna (Teleostei) from the Eocene to Pleistocene of New Zealand by means of otoliths." volume="46" year="2019">Schwarzhans 2019</bibRefCitation>
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and
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<bibRefCitation id="F9F04B2E94F3834AB924C743898D9EAB" DOI="https://doi.org/10.1186/s13358-021-00228-w" author="Schwarzhans, WW" journalOrPublisher="Paleobiology" pageId="0" pageNumber="103" refId="B63" refString="Schwarzhans, WW, Nielsen, SN, 2021. Fish otoliths from the early Miocene of Chile: a window into the evolution of marine bony fishes in the Southeast Pacific. Swiss Journal of Palaeontology 140: e16. [62 pp] https://doi.org/10.1186/s13358-021-00228-w" title="Fish otoliths from the early Miocene of Chile: a window into the evolution of marine bony fishes in the Southeast Pacific. Swiss Journal of Palaeontology 140: e 16. [62 pp]" url="https://doi.org/10.1186/s13358-021-00228-w" year="2021">Schwarzhans and Nielsen 2021</bibRefCitation>
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), but differs in the more elongate shape (OL:OH = 1.35-1.45 vs. 1.25-1.35), the thin appearance (OH:OT = 3.8-4.5 vs. 3.5-3.6), the larger number of denticles on the ventral rim (13-15 vs. 10-13), and the anteriorly depressed dorsal rim.
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<taxonomicName id="BA0D0ED97791E04A14D2A2CF462715B8" authorityName="Schwarzhans & Ohe & Tsuchiya & Ujihara" authorityYear="2022" family="Myctophidae" genus="Diaphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diaphus watatsumi" order="Myctophiformes" pageId="0" pageNumber="103" phylum="Chordata" rank="species" species="watatsumi">
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<emphasis id="9098881403A31EC71657A37A380A79CC" italics="true" pageId="0" pageNumber="103">Diaphus watatsumi</emphasis>
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</taxonomicName>
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resembles even more closely
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<taxonomicName id="D535D2E844F65DF3E20DD4B637B186D9" lsidName="D. sulcatus" pageId="0" pageNumber="103" rank="species" species="sulcatus">
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<emphasis id="850DB46968C38C843890E7D8D52698F0" italics="true" pageId="0" pageNumber="103">D. sulcatus</emphasis>
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</taxonomicName>
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(Bassoli, 1906) from the middle and late Miocene of Europe and the extant
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<taxonomicName id="3DE29BB7C4D68D6269077098836874B3" lsidName="D. splendidus" pageId="0" pageNumber="103" rank="species" species="splendidus">
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<emphasis id="65D302AF49634A97AB97E56784BE35EE" italics="true" pageId="0" pageNumber="103">D. splendidus</emphasis>
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</taxonomicName>
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(Brauer, 1904), which is known since at least the early Pliocene, and is therefore associated with the
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<taxonomicName id="E5FA9B3AE6A414A70A86EA7AD6170A65" baseAuthorityName="Brauer" baseAuthorityYear="1904" family="Myctophidae" genus="Diaphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diaphus splendidus" order="Myctophiformes" pageId="0" pageNumber="103" phylum="Chordata" rank="species" species="splendidus">
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<emphasis id="0AE8FFC409D13A895178E342CCA784B6" italics="true" pageId="0" pageNumber="103">Diaphus splendidus</emphasis>
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</taxonomicName>
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Otolith Group.
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<taxonomicName id="E93D6CCECBA9A17F63FD2264F2AE7DA9" authorityName="Schwarzhans & Ohe & Tsuchiya & Ujihara" authorityYear="2022" family="Myctophidae" genus="Diaphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diaphus watatsumi" order="Myctophiformes" pageId="0" pageNumber="103" phylum="Chordata" rank="species" species="watatsumi">
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<emphasis id="4AFCAB930C63495C21DEEC5BB2EC0C97" italics="true" pageId="0" pageNumber="103">Diaphus watatsumi</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName id="4CFDD373AE4DDFD910E393BB8431AB7E" lsidName="D. sulcatus" pageId="0" pageNumber="103" rank="species" species="sulcatus">
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<emphasis id="A5EFAB0061835FC7C228EE9124781AA2" italics="true" pageId="0" pageNumber="103">D. sulcatus</emphasis>
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</taxonomicName>
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|
in the rostrum being much longer than the antirostrum (vs. slightly longer to equal length) and the more depressed and longer predorsal region. It differs from
|
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<taxonomicName id="24DD78FA365ED953211195892C9EADC6" lsidName="D. splendidus" pageId="0" pageNumber="103" rank="species" species="splendidus">
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<emphasis id="B4475ADC44BA9B793287CAC0F8BF0D0A" italics="true" pageId="0" pageNumber="103">D. splendidus</emphasis>
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</taxonomicName>
|
|
in the relatively higher ratio OCL:CCL (2.0 vs. 1.6-1.8) and the higher number of denticles on the ventral rim (13-15 vs. 9-13).
|
|
<taxonomicName id="9C872E36BA603DBEC4172BD850D9E9E7" authorityName="Schwarzhans & Ohe & Tsuchiya & Ujihara" authorityYear="2022" family="Myctophidae" genus="Diaphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diaphus watatsumi" order="Myctophiformes" pageId="0" pageNumber="103" phylum="Chordata" rank="species" species="watatsumi">
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|
<emphasis id="DBAE43D9E6B9515E1018D5D31269F788" italics="true" pageId="0" pageNumber="103">Diaphus watatsumi</emphasis>
|
|
</taxonomicName>
|
|
also resembles otoliths of the extant
|
|
<taxonomicName id="66672C34EDD014A3A99AE9FE21971871" baseAuthorityName="Cocco" baseAuthorityYear="1838" class="Actinopterygii" family="Myctophidae" genus="Lobianchia" kingdom="Animalia" lsidName="Lobianchia gemellarii" order="Myctophiformes" pageId="0" pageNumber="103" phylum="Chordata" rank="species" species="gemellarii">
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<emphasis id="C8686F78046258DEEBFC4AD81CF820BC" italics="true" pageId="0" pageNumber="103">Lobianchia gemellarii</emphasis>
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|
</taxonomicName>
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|
(Cocco, 1838) (see
|
|
<bibRefCitation id="2BCF1DFFCAD4201F37732E0F6779D88F" author="Schwarzhans, W" journalOrPublisher="Rivista Italiana di Paleontologia e Stratigrafia" pageId="0" pageNumber="103" pagination="355 - 400" refId="B64" refString="Schwarzhans, W, Ohe, F, 2019. Lanternfish otoliths (Teleostei, Myctophidae) from the Pliocene and Pleistocene of Japan. Rivista Italiana di Paleontologia e Stratigrafia 125: 355 - 400" title="Lanternfish otoliths (Teleostei, Myctophidae) from the Pliocene and Pleistocene of Japan." volume="125" year="2019">Schwarzhans and Ohe 2019</bibRefCitation>
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for figures) but differs in the higher number of denticles along the ventral rim (13-15 vs. 5-9), the more regularly rounded posterior rim and the many radial furrows on the outer face.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |