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<mods:title id="879A1435879451B04F37749841B0537A">Integrative revision of the Iberian species of Coscinia Hübner, [1819] sensu lato and Spiris Hübner, [1819], (Lepidoptera: Erebidae, Arctiinae)</mods:title>
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<paragraph id="684EBA299F2D1C36FF5061C07FCA3F8A" blockId="3.[151,642,477,504]" box="[151,642,477,504]" pageId="3" pageNumber="404">
<heading id="33060D459F2D1C36FF5061C07FCA3F8A" bold="true" box="[151,642,477,504]" fontSize="11" level="1" pageId="3" pageNumber="404" reason="1">
<emphasis id="5A85663B9F2D1C36FF5061C07FCA3F8A" bold="true" box="[151,642,477,504]" pageId="3" pageNumber="404">
Subtribe
<taxonomicName id="AFF1C1AA9F2D1C36FECB61C07F6F3F85" authority="Walker" authorityName="Walker" authorityYear="1865" box="[268,551,477,504]" class="Insecta" family="Erebidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="subTribe" subTribe="Callimorphina">Callimorphina Walker</taxonomicName>
, [1865]
</emphasis>
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<paragraph id="684EBA299F2D1C36FF50623B7F103D6A" blockId="3.[151,1437,549,1872]" pageId="3" pageNumber="404">
Recent authors, such as
<bibRefCitation id="0C60C7D89F2D1C36FE62623B7F893C3D" author="Dacosta, M. A. &amp; Weller, S. J." box="[421,705,549,576]" pageId="3" pageNumber="404" pagination="1 - 94" refId="ref23178" refString="Dacosta, M. A. &amp; Weller, S. J. (2005) Phylogeny and classification of Callimorphini (Lepidoptera: Arctiidae: Arctiinae). Zootaxa, 1025 (1), 1 - 94. https: // doi. org / 10.11646 / zootaxa. 1025.1.1" type="journal article" year="2005">Dacosta &amp; Weller (2005)</bibRefCitation>
,
<bibRefCitation id="0C60C7D89F2D1C36FD0B62387EDF3C3D" author="Dubatolov, V. V." box="[716,919,549,576]" pageId="3" pageNumber="404" pagination="95 - 104" refId="ref23392" refString="Dubatolov, V. V. (2006) Cladogenesis of tiger-moths of the subfamily Arctiinae: development of a cladogenetic model of the tribe Callimorphini (Lepidoptera, Arctiidae) by the SYNAP method. Euroasian Entomological Journal, 5 (2), 95 - 104. [in Russian]" type="journal article" year="2006">Dubatolov (2006)</bibRefCitation>
,
<bibRefCitation id="0C60C7D89F2D1C36FC65623879E83C42" author="Witt, T. J. &amp; Ronkay, L." box="[930,1184,549,575]" pageId="3" pageNumber="404" refId="ref27927" refString="Witt, T. J. &amp; Ronkay, L. (2011) Noctuidae Europaea. Lymantrinae and Arctiinae including Phylogeny and Check List of the Quadrifid Noctuoidea of Europe, 13. Entomological Press, Soro, 448 pp." type="book" year="2011">Witt &amp; Ronkay (2011)</bibRefCitation>
,
<bibRefCitation id="0C60C7D89F2D1C36FB6C623878D43C42" author="Vives Moreno, A." box="[1195,1436,549,576]" pageId="3" pageNumber="404" refId="ref27675" refString="Vives Moreno, A. (2014) Catalogo sistematico y sinonimico de los Lepidoptera de la Peninsula Iberica, de Ceuta, de Melilla y de las Islas Azores, Baleares, Canarias, Madeira y Salvajes (Insecta: Lepidoptera). Suplemento de SHILAP Revista de Lepidopterologia, Madrid, 1184 pp." type="book" year="2014">Vives Moreno (2014)</bibRefCitation>
and
<bibRefCitation id="0C60C7D89F2D1C36FF0162547CE03C19" author="Zenker, M. M. &amp; Wahlberg, N. &amp; Brehm, G. &amp; Teston, J. A. &amp; Przybylowicz, L. &amp; Pie, M. R. &amp; Freitas, A. V. L." box="[198,424,585,612]" pageId="3" pageNumber="404" pagination="348 - 362" refId="ref28558" refString="Zenker, M. M., Wahlberg, N., Brehm, G., Teston, J. A., Przybylowicz, L., Pie, M. R. &amp; Freitas, A. V. L. (2016) Systematics and origin of moths in the subfamily Arctiinae (Lepidoptera, Erebidae) in the Neotropical region. Zoologica Scripta, 46 (3), 348 - 362." type="journal article" year="2016">
Zenker
<emphasis id="5A85663B9F2D1C36FEDB62577C1C3C1E" box="[284,340,585,611]" italics="true" pageId="3" pageNumber="404">et al.</emphasis>
(2016)
</bibRefCitation>
, include
<taxonomicName id="AFF1C1AA9F2D1C36FDCB62577F383C19" baseAuthorityName="Marmet &amp; Schmid" baseAuthorityYear="2000" box="[524,624,586,612]" class="Insecta" family="Arctiidae" genus="Coscinia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="5A85663B9F2D1C36FDCB62577F383C19" box="[524,624,586,612]" italics="true" pageId="3" pageNumber="404">Coscinia</emphasis>
</taxonomicName>
and
<taxonomicName id="AFF1C1AA9F2D1C36FD6162577FAF3C19" authorityName="Hubner" authorityYear="1819" box="[678,743,586,612]" class="Insecta" family="Erebidae" genus="Spiris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="5A85663B9F2D1C36FD6162577FAF3C19" box="[678,743,586,612]" italics="true" pageId="3" pageNumber="404">Spiris</emphasis>
</taxonomicName>
in the subtribe
<taxonomicName id="AFF1C1AA9F2D1C36FC53625479C63C1E" authority="Walker" authorityName="Walker" authorityYear="1865" box="[916,1166,585,611]" class="Insecta" family="Erebidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="subTribe" subTribe="Callimorphina">Callimorphina Walker</taxonomicName>
, [1865]. However, there are distinct morphological traits, explained below, which perhaps indicate that these genera do not belong in
<taxonomicName id="AFF1C1AA9F2D1C36FAAB62707C5A3CD6" authorityName="Walker" authorityYear="1865" class="Insecta" family="Erebidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="subTribe" subTribe="Callimorphina">Callimorphina</taxonomicName>
but in another subtribe. This is something that requires a more extensive and detailed study, and is not discussed here. The authors are fully aware that the morphological features described here only apply to a subset of
<taxonomicName id="AFF1C1AA9F2D1C36FF5062C47C743C8E" authorityName="Walker" authorityYear="1865" box="[151,316,729,755]" class="Insecta" family="Erebidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="subTribe" subTribe="Callimorphina">Callimorphina</taxonomicName>
, and that they do not represent a sound diagnosis for the whole subtribe; however, they are valid for the Iberian and Western Europe species.
</paragraph>
</subSubSection>
<subSubSection id="20EBE9A29F2D1C36FF00633C792A3DB6" pageId="3" pageNumber="404" type="description">
<paragraph id="684EBA299F2D1C36FF00633C792A3DB6" blockId="3.[151,1437,549,1872]" pageId="3" pageNumber="404">
Morphological differences within the genus
<taxonomicName id="AFF1C1AA9F2D1C36FD0A633F7E793D41" baseAuthorityName="Marmet &amp; Schmid" baseAuthorityYear="2000" box="[717,817,802,828]" class="Insecta" family="Arctiidae" genus="Coscinia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="5A85663B9F2D1C36FD0A633F7E793D41" box="[717,817,802,828]" italics="true" pageId="3" pageNumber="404">Coscinia</emphasis>
</taxonomicName>
have led the authors to create the genus
<taxonomicName id="AFF1C1AA9F2D1C36FADA633C7F773D1D" authority="Macia, Mally, Ylla, Gaston &amp; Huertas" authorityName="Macià &amp; Mally &amp; Ylla &amp; Gastón &amp; Huertas" authorityYear="2019" class="Insecta" family="Erebidae" genus="Sagarriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5A85663B9F2D1C36FADA633C78D53D46" box="[1309,1437,801,827]" italics="true" pageId="3" pageNumber="404">Sagarriella</emphasis>
Macià, Mally, Ylla, Gastón &amp; Huertas
</taxonomicName>
<emphasis id="5A85663B9F2D1C36FD82635B7FE63D1D" bold="true" box="[581,686,838,864]" pageId="3" pageNumber="404">
<taxonomicNameLabel id="41B6DB409F2D1C36FD82635B7FE63D1D" box="[581,686,838,864]" pageId="3" pageNumber="404" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
and to restore the genus
<taxonomicName id="AFF1C1AA9F2D1C36FC05635B796A3D22" authorityName="Kemal &amp; Kocak" authorityYear="2006" box="[962,1058,838,863]" class="Insecta" family="Arctiidae" genus="Lerautia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="5A85663B9F2D1C36FC05635B796A3D22" box="[962,1058,838,863]" italics="true" pageId="3" pageNumber="404">Lerautia</emphasis>
</taxonomicName>
<emphasis id="5A85663B9F2D1C36FBEE635B79C63D1D" bold="true" box="[1065,1166,838,864]" pageId="3" pageNumber="404">stat. rev.</emphasis>
Based on their morphological characteristics,
<taxonomicName id="AFF1C1AA9F2D1C36FE5063747F5F3DFE" authority="Macià &amp; Mally &amp; Ylla &amp; Gastón &amp; Huertas, 2019" authorityName="Macià &amp; Mally &amp; Ylla &amp; Gastón &amp; Huertas" authorityYear="2019" box="[407,535,873,899]" class="Insecta" family="Erebidae" genus="Sagarriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5A85663B9F2D1C36FE5063747F5F3DFE" box="[407,535,873,899]" italics="true" pageId="3" pageNumber="404">Sagarriella</emphasis>
</taxonomicName>
and
<taxonomicName id="AFF1C1AA9F2D1C36FD8C63777FE33DFE" authorityName="Kemal &amp; Kocak" authorityYear="2006" box="[587,683,874,899]" class="Insecta" family="Arctiidae" genus="Lerautia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="5A85663B9F2D1C36FD8C63777FE33DFE" box="[587,683,874,899]" italics="true" pageId="3" pageNumber="404">Lerautia</emphasis>
</taxonomicName>
are assigned to the subtribe
<taxonomicName id="AFF1C1AA9F2D1C36FC21637479C03DFE" authorityName="Walker" authorityYear="1865" box="[998,1160,873,899]" class="Insecta" family="Erebidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="subTribe" subTribe="Callimorphina">Callimorphina</taxonomicName>
along with
<taxonomicName id="AFF1C1AA9F2D1C36FACD637778263DF9" baseAuthorityName="Marmet &amp; Schmid" baseAuthorityYear="2000" box="[1290,1390,874,900]" class="Insecta" family="Arctiidae" genus="Coscinia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="5A85663B9F2D1C36FACD637778263DF9" box="[1290,1390,874,900]" italics="true" pageId="3" pageNumber="404">Coscinia</emphasis>
</taxonomicName>
and
<taxonomicName id="AFF1C1AA9F2D1C36FF5063937D903DD5" authorityName="Hubner" authorityYear="1819" box="[151,216,910,936]" class="Insecta" family="Erebidae" genus="Spiris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="5A85663B9F2D1C36FF5063937D903DD5" box="[151,216,910,936]" italics="true" pageId="3" pageNumber="404">Spiris</emphasis>
</taxonomicName>
. Although we have not been able to study the larval stages of
<emphasis id="5A85663B9F2D1C36FC4B63937EBB3DDA" box="[908,1011,910,935]" italics="true" pageId="3" pageNumber="404">
<taxonomicName id="AFF1C1AA9F2D1C36FC4B63937EA73DDA" authorityName="Kemal &amp; Kocak" authorityYear="2006" box="[908,1007,910,935]" class="Insecta" family="Arctiidae" genus="Lerautia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">Lerautia</taxonomicName>
,
</emphasis>
given its great similarity in the rest of studied characters, we assume that larval characteristics should not differ significantly.
</paragraph>
</subSubSection>
<subSubSection id="20EBE9A29F2D1C36FF0063C87FB538BD" pageId="3" pageNumber="404" type="diagnosis">
<paragraph id="684EBA299F2D1C36FF0063C87FA33A96" blockId="3.[151,1437,549,1872]" pageId="3" pageNumber="404">
<emphasis id="5A85663B9F2D1C36FF0063C87CD63D8D" bold="true" box="[199,414,981,1008]" pageId="3" pageNumber="404">Diagnosis. Imago.</emphasis>
Imagines of
<emphasis id="5A85663B9F2D1C36FDFE63CB7EC83D92" box="[569,896,981,1008]" italics="true" pageId="3" pageNumber="404">
<taxonomicName id="AFF1C1AA9F2D1C36FDFE63CB7FD73D8D" baseAuthorityName="Marmet &amp; Schmid" baseAuthorityYear="2000" box="[569,671,982,1008]" class="Insecta" family="Arctiidae" genus="Coscinia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">Coscinia</taxonomicName>
,
<taxonomicName id="AFF1C1AA9F2D1C36FD6963CB7FBA3D8D" authorityName="Hubner" authorityYear="1819" box="[686,754,982,1008]" class="Insecta" family="Erebidae" genus="Spiris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">Spiris</taxonomicName>
, Sagarriella
</emphasis>
and
<taxonomicName id="AFF1C1AA9F2D1C36FC7C63CB79533D92" authorityName="Kemal &amp; Kocak" authorityYear="2006" box="[955,1051,982,1007]" class="Insecta" family="Arctiidae" genus="Lerautia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="5A85663B9F2D1C36FC7C63CB79533D92" box="[955,1051,982,1007]" italics="true" pageId="3" pageNumber="404">Lerautia</emphasis>
</taxonomicName>
have elongated and narrow forewings, and antennae that are bipectinate in males and filiform in females. Typical members of
<taxonomicName id="AFF1C1AA9F2D1C36FB7963E478283A6E" authorityName="Walker" authorityYear="1865" box="[1214,1376,1017,1043]" class="Insecta" family="Erebidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="subTribe" subTribe="Callimorphina">Callimorphina</taxonomicName>
have a broader wing shape, and the antennae are filiform in both sexes. The male genitalia of
<taxonomicName id="AFF1C1AA9F2D1C36FBB1640078503A4A" authorityName="Walker" authorityYear="1865" box="[1142,1304,1053,1079]" class="Insecta" family="Erebidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="subTribe" subTribe="Callimorphina">Callimorphina</taxonomicName>
have a sacculus with a prominent process, always separated from the valve to form a sort of bilobed valve. The male genitalia of
<emphasis id="5A85663B9F2D1C36FF71647B7CBF3A02" box="[182,503,1125,1152]" italics="true" pageId="3" pageNumber="404">
<taxonomicName id="AFF1C1AA9F2D1C36FF71647B7C543AFD" baseAuthorityName="Marmet &amp; Schmid" baseAuthorityYear="2000" box="[182,284,1126,1152]" class="Insecta" family="Arctiidae" genus="Coscinia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">Coscinia</taxonomicName>
,
<taxonomicName id="AFF1C1AA9F2D1C36FEE0647B7C233AFD" authorityName="Hubner" authorityYear="1819" box="[295,363,1126,1152]" class="Insecta" family="Erebidae" genus="Spiris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">Spiris</taxonomicName>
, Sagarriella
</emphasis>
and
<taxonomicName id="AFF1C1AA9F2D1C36FDEA647B7FC53A02" authorityName="Kemal &amp; Kocak" authorityYear="2006" box="[557,653,1126,1151]" class="Insecta" family="Arctiidae" genus="Lerautia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="5A85663B9F2D1C36FDEA647B7FC53A02" box="[557,653,1126,1151]" italics="true" pageId="3" pageNumber="404">Lerautia</emphasis>
</taxonomicName>
lack a sacculus with similar processes, but always have a characteristic fold in the central part of the valve, which is absent in
<taxonomicName id="AFF1C1AA9F2D1C36FCEF64947E863ADE" authorityName="Walker" authorityYear="1865" box="[808,974,1161,1187]" class="Insecta" family="Erebidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="subTribe" subTribe="Callimorphina">Callimorphina</taxonomicName>
. The tip of the uncus in
<emphasis id="5A85663B9F2D1C36FB2664977C5F3ABA" italics="true" pageId="3" pageNumber="404">
<taxonomicName id="AFF1C1AA9F2D1C36FB266497780F3AD9" baseAuthorityName="Marmet &amp; Schmid" baseAuthorityYear="2000" box="[1249,1351,1162,1188]" class="Insecta" family="Arctiidae" genus="Coscinia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">Coscinia</taxonomicName>
,
<taxonomicName id="AFF1C1AA9F2D1C36FA93649778D03AD9" authorityName="Hubner" authorityYear="1819" box="[1364,1432,1162,1188]" class="Insecta" family="Erebidae" genus="Spiris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">Spiris</taxonomicName>
, Sagarriella
</emphasis>
and
<taxonomicName id="AFF1C1AA9F2D1C36FE9564B37CFA3ABA" authorityName="Kemal &amp; Kocak" authorityYear="2006" box="[338,434,1198,1223]" class="Insecta" family="Arctiidae" genus="Lerautia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="5A85663B9F2D1C36FE9564B37CFA3ABA" box="[338,434,1198,1223]" italics="true" pageId="3" pageNumber="404">Lerautia</emphasis>
</taxonomicName>
is usually tubular or slightly pointed, or spatulate; in
<taxonomicName id="AFF1C1AA9F2D1C36FBE364B079813ABA" authorityName="Walker" authorityYear="1865" box="[1060,1225,1197,1223]" class="Insecta" family="Erebidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="subTribe" subTribe="Callimorphina">Callimorphina</taxonomicName>
, the uncus widens subapically and resembles the shape of a bird´s beak.
</paragraph>
<paragraph id="684EBA299F2D1C36FF0064EB7FB538BD" blockId="3.[151,1437,549,1872]" pageId="3" pageNumber="404">
<emphasis id="5A85663B9F2D1C36FF0064EB7C5F3B6D" bold="true" box="[199,279,1270,1296]" pageId="3" pageNumber="404">Larva.</emphasis>
The larvae of
<emphasis id="5A85663B9F2D1C36FE7C64EB7F263B6D" box="[443,622,1270,1296]" italics="true" pageId="3" pageNumber="404">
<taxonomicName id="AFF1C1AA9F2D1C36FE7C64EB7F693B6D" baseAuthorityName="Marmet &amp; Schmid" baseAuthorityYear="2000" box="[443,545,1270,1296]" class="Insecta" family="Arctiidae" genus="Coscinia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">Coscinia</taxonomicName>
,
<taxonomicName id="AFF1C1AA9F2D1C36FDEA64EB7F263B6D" authorityName="Hubner" authorityYear="1819" box="[557,622,1270,1296]" class="Insecta" family="Erebidae" genus="Spiris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">Spiris</taxonomicName>
</emphasis>
and
<taxonomicName id="AFF1C1AA9F2D1C36FD6264E87E6D3B72" authority="Macià &amp; Mally &amp; Ylla &amp; Gastón &amp; Huertas, 2019" authorityName="Macià &amp; Mally &amp; Ylla &amp; Gastón &amp; Huertas" authorityYear="2019" box="[677,805,1269,1295]" class="Insecta" family="Erebidae" genus="Sagarriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5A85663B9F2D1C36FD6264E87E6D3B72" box="[677,805,1269,1295]" italics="true" pageId="3" pageNumber="404">Sagarriella</emphasis>
</taxonomicName>
(
<taxonomicName id="AFF1C1AA9F2D1C36FCF264EB7EDD3B72" authorityName="Kemal &amp; Kocak" authorityYear="2006" box="[821,917,1270,1295]" class="Insecta" family="Arctiidae" genus="Lerautia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="5A85663B9F2D1C36FCF264EB7EDD3B72" box="[821,917,1270,1295]" italics="true" pageId="3" pageNumber="404">Lerautia</emphasis>
</taxonomicName>
, larva not studied) have four to five P setae (P = posterior) in the superior area of each hemisphere epicranium of the head capsule. Setae P1 and P2 are common in most
<taxonomicName id="AFF1C1AA9F2D1C36FF1465207C123B2A" authorityName="Linnaeus" authorityYear="1758" box="[211,346,1341,1367]" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="order">Lepidoptera</taxonomicName>
larvae, but the other three seem to be exclusive to these genera. There is no reference to these setae in
<bibRefCitation id="0C60C7D89F2D1C36FF74657C7CC83B01" author="Gerasimov, A. M." box="[179,384,1377,1404]" pageId="3" pageNumber="404" pagination="177 - 194" refId="ref23853" refString="Gerasimov, A. M. (1935) Zur Frage der Homodynamie der Borsten von Schmetterlingsraupen. Zoologischer Anzeiger, 112 (7 / 8), 177 - 194." type="journal article" year="1935">Gerasimov (1935)</bibRefCitation>
,
<bibRefCitation id="0C60C7D89F2D1C36FE4C657C7F613B01" author="Hinton, H. E." box="[395,553,1377,1404]" pageId="3" pageNumber="404" pagination="1 - 37" refId="ref24471" refString="Hinton, H. E. (1946) On the homology and nomenclature of the setae of lepidopterous larvae, with some notes on the phylogeny of the Lepidoptera. Transactions of the Entomological Society of London, 97, 1 - 37. https: // doi. org / 10.1111 / j. 1365 - 2311.1946. tb 00372. x" type="journal article" year="1946">Hinton (1946)</bibRefCitation>
or
<bibRefCitation id="0C60C7D89F2D1C36FD8B657C7F903B01" author="Beck, H." box="[588,728,1377,1404]" pageId="3" pageNumber="404" pagination="1 - 406" refId="ref22947" refString="Beck, H. (1960) Die Larvalsystematik der Eulen. Abhandlungen zur Larvalsystematik der Insekten, 4, 1 - 406." type="journal article" year="1960">Beck (1960)</bibRefCitation>
, but they are mentioned in Garcia-Barros (1992) for
<emphasis id="5A85663B9F2D1C36FADA657C7D973BE2" italics="true" pageId="3" pageNumber="404">
<taxonomicName id="AFF1C1AA9F2D1C36FADA657C7D933BE2" authorityName="Sagarra" authorityYear="1924" class="Insecta" family="Erebidae" genus="Sagarriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="romei">Sagarriella romei</taxonomicName>
.
</emphasis>
These setae are not present in the species of the subtribe
<taxonomicName id="AFF1C1AA9F2D1C36FC9B659879493BE2" authorityName="Walker" authorityYear="1865" box="[860,1025,1413,1439]" class="Insecta" family="Erebidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="subTribe" subTribe="Callimorphina">Callimorphina</taxonomicName>
, in which only P1 and P2 are present in
<taxonomicName id="AFF1C1AA9F2D1C36FF7265B77F6D3BB9" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[181,549,1449,1476]" class="Insecta" family="Arctiidae" genus="Tyria" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="jacobeae">
<emphasis id="5A85663B9F2D1C36FF7265B77C153BBE" box="[181,349,1449,1475]" italics="true" pageId="3" pageNumber="404">Tyria jacobeae</emphasis>
(
<bibRefCitation id="0C60C7D89F2D1C36FEAA65B47F553BB9" author="Linnaeus, C." box="[365,541,1449,1476]" pageId="3" pageNumber="404" refId="ref25398" refString="Linnaeus, C. (1758) Systema Naturae per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, cum characteribus, difierentiis, synonymis, loeis. Edito Decima, Reformata. Impensis Direct. Laurentii Salvii, Holmiae, 824 pp." type="book" year="1758">Linnaeus, 1758</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="AFF1C1AA9F2D1C36FD9965B77FA13BBE" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[606,745,1449,1475]" class="Insecta" family="Arctiidae" genus="Utetheisa" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="pulchella">
<emphasis id="5A85663B9F2D1C36FD9965B77FA13BBE" box="[606,745,1449,1475]" italics="true" pageId="3" pageNumber="404">U. pulchella</emphasis>
</taxonomicName>
. In
<taxonomicName id="AFF1C1AA9F2D1C36FCDF65B479FF3BB9" authority="(Esper, 1785)" baseAuthorityName="Esper" baseAuthorityYear="1785" box="[792,1207,1449,1476]" class="Insecta" family="Arctiidae" genus="Cymbalophora" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="pudica">
<emphasis id="5A85663B9F2D1C36FCDF65B4795C3BBE" box="[792,1044,1449,1475]" italics="true" pageId="3" pageNumber="404">Cymbalophora pudica</emphasis>
(
<bibRefCitation id="0C60C7D89F2D1C36FBE365B479E63BB9" author="Esper, E. J." box="[1060,1198,1449,1476]" pageId="3" pageNumber="404" pagination="177" refId="ref23473" refString="Esper, E. J. (1785) Insecten mit bestaubten Ilugeln. Lepidoptera order Schmetterlinge. Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen, 3, 177." type="journal article" year="1785">Esper, 1785</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="5A85663B9F2D1C36FB0365B47D813B9A" italics="true" pageId="3" pageNumber="404">Callimorpha dominula</emphasis>
(
<bibRefCitation id="0C60C7D89F2D1C36FF1065D07CCE3B95" author="Linnaeus, C." box="[215,390,1485,1512]" pageId="3" pageNumber="404" refId="ref25398" refString="Linnaeus, C. (1758) Systema Naturae per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, cum characteribus, difierentiis, synonymis, loeis. Edito Decima, Reformata. Impensis Direct. Laurentii Salvii, Holmiae, 824 pp." type="book" year="1758">Linnaeus, 1758</bibRefCitation>
) and
<emphasis id="5A85663B9F2D1C36FE0465D07FA03B9A" box="[451,744,1485,1511]" italics="true" pageId="3" pageNumber="404">Euplagia quadripunctaria</emphasis>
(
<bibRefCitation id="0C60C7D89F2D1C36FD3165D07E303B95" author="Poda, N." box="[758,888,1485,1512]" pageId="3" pageNumber="404" pagination="89" refId="ref26667" refString="Poda, N. (1761) Insecta Musei Graecensis, quae in ordinis, genera et species juxta Systema Naturae Caroli Linnaei digessit Nocolaus Poda. Insecta Musei Graecensis, 8, 89." type="journal article" year="1761">Poda, 1761</bibRefCitation>
), these setae are not distinguished. The immature stages of
<emphasis id="5A85663B9F2D1C36FEC565EF7CFD3871" box="[258,437,1522,1548]" italics="true" pageId="3" pageNumber="404">
<taxonomicName id="AFF1C1AA9F2D1C36FEC565EF7C203871" baseAuthorityName="Marmet &amp; Schmid" baseAuthorityYear="2000" box="[258,360,1522,1548]" class="Insecta" family="Arctiidae" genus="Coscinia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">Coscinia</taxonomicName>
,
<taxonomicName id="AFF1C1AA9F2D1C36FEB365EF7CFD3871" authorityName="Hubner" authorityYear="1819" box="[372,437,1522,1548]" class="Insecta" family="Erebidae" genus="Spiris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">Spiris</taxonomicName>
</emphasis>
and
<taxonomicName id="AFF1C1AA9F2D1C36FE2A65EC7F253876" authority="Macià &amp; Mally &amp; Ylla &amp; Gastón &amp; Huertas, 2019" authorityName="Macià &amp; Mally &amp; Ylla &amp; Gastón &amp; Huertas" authorityYear="2019" box="[493,621,1521,1547]" class="Insecta" family="Erebidae" genus="Sagarriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5A85663B9F2D1C36FE2A65EC7F253876" box="[493,621,1521,1547]" italics="true" pageId="3" pageNumber="404">Sagarriella</emphasis>
</taxonomicName>
(
<taxonomicName id="AFF1C1AA9F2D1C36FDB965EF7F963876" authorityName="Kemal &amp; Kocak" authorityYear="2006" box="[638,734,1522,1547]" class="Insecta" family="Arctiidae" genus="Lerautia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="5A85663B9F2D1C36FDB965EF7F963876" box="[638,734,1522,1547]" italics="true" pageId="3" pageNumber="404">Lerautia</emphasis>
</taxonomicName>
, larva not studied) are very similar, and specific identification relies on breeding them to the adult stage. So, when breeding larvae from eggs, the best diagnostic characters for species distinction are the shape of the nails of the thoracic legs and abdominal prolegs and the chaetotaxy of the ocellar zone, where setae O1 and O2 are always in the same place, whereas seta O3 can approach more or less setae O1 or O2 depending on the species. Larva cylindrical and dark, with a white or orange dorsal line; verrucae D1, D2, SD1, L1, L2 and L3, with short, black and white setae.
</paragraph>
</subSubSection>
<subSubSection id="20EBE9A29F2D1C36FF0066D7799B392D" pageId="3" pageNumber="404" type="description">
<paragraph id="684EBA299F2D1C36FF0066D7799B392D" blockId="3.[151,1437,549,1872]" pageId="3" pageNumber="404">
<emphasis id="5A85663B9F2D1C36FF0066D77C453899" bold="true" box="[199,269,1738,1764]" pageId="3" pageNumber="404">Pupa.</emphasis>
Straight or slightly curved; fusion of the tips of the pterothecae and the tips of the antennae protruding or not; hair coverage of variable length; there are circular depressions in the abdomen and groups of setae of different length that coincide with the verruca of the larva; posterior end of pupa rounded, cremaster absent in
<emphasis id="5A85663B9F2D1C36FAF5670F7D90392D" italics="true" pageId="3" pageNumber="404">
<taxonomicName id="AFF1C1AA9F2D1C36FAF5670F78D03951" baseAuthorityName="Marmet &amp; Schmid" baseAuthorityYear="2000" box="[1330,1432,1810,1836]" class="Insecta" family="Arctiidae" genus="Coscinia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">Coscinia</taxonomicName>
,
<taxonomicName id="AFF1C1AA9F2D1C36FF50672B7D90392D" authorityName="Hubner" authorityYear="1819" box="[151,216,1846,1872]" class="Insecta" family="Erebidae" genus="Spiris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">Spiris</taxonomicName>
</emphasis>
and
<taxonomicName id="AFF1C1AA9F2D1C36FEC867287CC73932" authority="Macià &amp; Mally &amp; Ylla &amp; Gastón &amp; Huertas, 2019" authorityName="Macià &amp; Mally &amp; Ylla &amp; Gastón &amp; Huertas" authorityYear="2019" box="[271,399,1845,1871]" class="Insecta" family="Erebidae" genus="Sagarriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5A85663B9F2D1C36FEC867287CC73932" box="[271,399,1845,1871]" italics="true" pageId="3" pageNumber="404">Sagarriella</emphasis>
</taxonomicName>
, (
<taxonomicName id="AFF1C1AA9F2D1C36FE61672B7F4E3932" authorityName="Kemal &amp; Kocak" authorityYear="2006" box="[422,518,1846,1871]" class="Insecta" family="Arctiidae" genus="Lerautia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="5A85663B9F2D1C36FE61672B7F4E3932" box="[422,518,1846,1871]" italics="true" pageId="3" pageNumber="404">Lerautia</emphasis>
</taxonomicName>
, pupa not studied) present in
<taxonomicName id="AFF1C1AA9F2D1C36FC9567287EBC3932" authorityName="Walker" authorityYear="1865" box="[850,1012,1845,1871]" class="Insecta" family="Erebidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="subTribe" subTribe="Callimorphina">Callimorphina</taxonomicName>
except
<taxonomicName id="AFF1C1AA9F2D1C36FB8C672B79843932" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1099,1228,1845,1871]" class="Insecta" family="Arctiidae" genus="Tyria" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="jacobeae">
<emphasis id="5A85663B9F2D1C36FB8C672B79843932" box="[1099,1228,1845,1871]" italics="true" pageId="3" pageNumber="404">T. jacobeae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>