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<document id="CA0004204411BC9E1D797168B3ACBCFB" ID-CLB-Dataset="26487" ID-DOI="10.11646/zootaxa.4615.3.1" ID-GBIF-Dataset="72c56097-337c-42fa-90ee-2da90529b2d2" ID-ISSN="1175-5326" ID-Zenodo-Dep="3246048" ID-ZooBank="4D816AA2-2AEA-470F-A79D-45452CFAE9F2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1560509144762" checkinUser="plazi" docAuthor="Macià, Ramon, Mally, Richard, Ylla, Josep, Gastón, Javier &amp; Huertas, Manuel" docDate="2019" docId="E0580B3F9F221C38FF5064987C4D393A" docLanguage="en" docName="zootaxa.4615.3.1.pdf" docOrigin="Zootaxa 4615 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coscinia cribraria subsp. benderi Macià &amp; Mally &amp; Ylla &amp; Gastón &amp; Huertas 2019, stat. rev." docType="treatment" docVersion="15" lastPageNumber="414" masterDocId="1C6173479F2E1C35FFC7601D7D483E7D" masterDocTitle="Integrative revision of the Iberian species of Coscinia Hübner, [1819] sensu lato and Spiris Hübner, [1819], (Lepidoptera: Erebidae, Arctiinae)" masterLastPageNumber="449" masterPageNumber="401" pageNumber="413" updateTime="1698734823461" updateUser="ExternalLinkService">
<mods:mods id="AB8681BD2CCB305E0524FA867A328722" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B1091B6D01F5E6F206EA06FDB133478D">
<mods:title id="12D91CD01174ED1FBCA3A94D0C3F06D2">Integrative revision of the Iberian species of Coscinia Hübner, [1819] sensu lato and Spiris Hübner, [1819], (Lepidoptera: Erebidae, Arctiinae)</mods:title>
</mods:titleInfo>
<mods:name id="B8411E8241566AF0B49127063644778A" type="personal">
<mods:role id="CFC1C9619717AAB7FCB55E56622838B8">
<mods:roleTerm id="DCFAD11C2098ACD2B76EE3E343F5EBAC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EF3A8FB4C4BDEFB370DFFAC01E1F89F9">Macià, Ramon</mods:namePart>
</mods:name>
<mods:name id="66D57DBC86FBCFD493F03E76A7D0A84B" type="personal">
<mods:role id="8AAD9D99363BE792ACEB1562923B5B2E">
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</mods:role>
<mods:namePart id="F6A102A344DBD0CB917E6780F611DD4F">Mally, Richard</mods:namePart>
</mods:name>
<mods:name id="A60BE3266816D3079F9985C46B42977E" type="personal">
<mods:role id="44EA2CC6F29E3BDE476A1C2D74224BF0">
<mods:roleTerm id="719A1AD4A151085428CEE89F4CF6BCED">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DF3906576510645DE6A9F34AAC1B5DAF">Ylla, Josep</mods:namePart>
</mods:name>
<mods:name id="852C0E90579BDD58E301E7B704D137C4" type="personal">
<mods:role id="3F36D672FCAC2D041E13692F86E903F9">
<mods:roleTerm id="DCB78183E6EDD3113795911674AB9E17">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="08FE44C155DE87CD70CA53F279A3D308">Gastón, Javier</mods:namePart>
</mods:name>
<mods:name id="09E6126246D3AE99151AA018A6162932" type="personal">
<mods:role id="0D1921ACC08C5311DE4FA7569448AB25">
<mods:roleTerm id="9FA95629D624E828C0A5EB0905EAECDB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F6EAD3EAFB41F4A10A585062F94B7C34">Huertas, Manuel</mods:namePart>
</mods:name>
<mods:typeOfResource id="E4B760BFC8D51491E95AB8EDCF1FAD63">text</mods:typeOfResource>
<mods:relatedItem id="AC898C733EDD3BEE3954598C9AFF3C7F" type="host">
<mods:titleInfo id="720CFC81E4D5F249F32B506821FA8D6D">
<mods:title id="A51CA1DBE7E75430B6A0C8A15BC96BC7">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A25411A67B0502EFADC950C384900C89">
<mods:date id="BA99B40F4A6BCF907976E735EE32BB99">2019</mods:date>
<mods:detail id="F74BFE96C944DA5EECCAA628C9A00F82" type="pubDate">
<mods:number id="3C0479777D6D14581D8CE6D111C2B5E5">2019-06-14</mods:number>
</mods:detail>
<mods:detail id="DA15E215328A8C3AB60629EA9ABAEB80" type="volume">
<mods:number id="4520567A2AF0B527846B11B209BC7ADA">4615</mods:number>
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<mods:number id="A08C2065B1957D2BB472F9346AA2EF0C">3</mods:number>
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</mods:extent>
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<mods:classification id="61420DECF88587A613DA752BA8797BEF">journal article</mods:classification>
<mods:identifier id="4C0E184890D750A219AE36CB9E74270A" type="CLB-Dataset">26487</mods:identifier>
<mods:identifier id="6FC69E5665AF3561A35DA4AC4216AA86" type="DOI">10.11646/zootaxa.4615.3.1</mods:identifier>
<mods:identifier id="75899181004882BD60C0CE6A2F00C161" type="GBIF-Dataset">72c56097-337c-42fa-90ee-2da90529b2d2</mods:identifier>
<mods:identifier id="80823AD7D3FCF560FA49A33908D33242" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
<treatment id="E0580B3F9F221C38FF5064987C4D393A" ID-GBIF-Taxon="158223111" LSID="urn:lsid:plazi:treatment:E0580B3F9F221C38FF5064987C4D393A" httpUri="http://treatment.plazi.org/id/E0580B3F9F221C38FF5064987C4D393A" lastPageId="13" lastPageNumber="414" pageId="12" pageNumber="413">
<subSubSection id="20EBE9A29F221C39FF50649879183A9C" pageId="12" pageNumber="413" type="nomenclature">
<paragraph id="684EBA299F221C39FF5064987FBF3AE2" blockId="12.[151,759,1157,1184]" box="[151,759,1157,1184]" pageId="12" pageNumber="413">
<heading id="33060D459F221C39FF5064987FBF3AE2" bold="true" box="[151,759,1157,1184]" fontSize="11" level="1" pageId="12" pageNumber="413" reason="1">
<emphasis id="5A85663B9F221C39FF5064987FBF3AE2" bold="true" box="[151,759,1157,1184]" pageId="12" pageNumber="413">
<taxonomicName id="AFF1C1AA9F221C39FF5064987FCE3AE2" authority="Marten, 1957" authorityName="Macià &amp; Mally &amp; Ylla &amp; Gastón &amp; Huertas" authorityYear="2019" baseAuthorityName="Marten" baseAuthorityYear="1957" box="[151,646,1157,1184]" class="Insecta" family="Arctiidae" genus="Coscinia" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="413" phylum="Arthropoda" rank="subSpecies" species="cribraria" status="stat. nov." subSpecies="benderi">
<emphasis id="5A85663B9F221C39FF5064987C9E3ADD" bold="true" box="[151,470,1157,1184]" italics="true" pageId="12" pageNumber="413">Coscinia cribraria benderi</emphasis>
<bibRefCitation id="0C60C7D89F221C39FE1964987FCE3AE2" author="Marten, W." box="[478,646,1157,1183]" pageId="12" pageNumber="413" pagination="89 - 91" refId="ref26012" refString="Marten, W. (1957) Coscinia benderi, eine neue Arctiide aus Andalusien (Lep. Arctiidae). Entomologische Zeitschrift, 67 (8), 89 - 91." type="journal article" year="1957">Marten, 1957</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="41B6DB409F221C39FD4A64987FBF3AE2" box="[653,759,1157,1183]" pageId="12" pageNumber="413">stat. rev.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="684EBA299F221C39FF5064D479183A9C" blockId="12.[151,1104,1225,1280]" box="[151,1104,1225,1250]" pageId="12" pageNumber="413">
Original combination:
<taxonomicName id="AFF1C1AA9F221C39FEB964D77FF63A9F" authority="Marten, 1957" authorityName="Marten" authorityYear="1957" box="[382,702,1225,1250]" class="Insecta" family="Arctiidae" genus="Coscinia" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="benderi">
<emphasis id="5A85663B9F221C39FEB964D77F633A9C" box="[382,555,1225,1249]" italics="true" pageId="12" pageNumber="413">Coscinia benderi</emphasis>
<bibRefCitation id="0C60C7D89F221C39FDF564D77FF63A9F" author="Marten, W." box="[562,702,1225,1250]" pageId="12" pageNumber="413" pagination="89 - 91" refId="ref26012" refString="Marten, W. (1957) Coscinia benderi, eine neue Arctiide aus Andalusien (Lep. Arctiidae). Entomologische Zeitschrift, 67 (8), 89 - 91." type="journal article" year="1957">Marten, 1957</bibRefCitation>
</taxonomicName>
.
<emphasis id="5A85663B9F221C39FD0064D47E913A9C" box="[711,985,1225,1249]" italics="true" pageId="12" pageNumber="413">Entomologische Zeitschrift</emphasis>
,
<emphasis id="5A85663B9F221C39FC2164D47EB73A9F" bold="true" box="[998,1023,1225,1250]" pageId="12" pageNumber="413">67</emphasis>
(8), 89.
</paragraph>
</subSubSection>
<subSubSection id="20EBE9A29F221C39FF5064F57C113B7D" box="[151,345,1255,1280]" pageId="12" pageNumber="413" type="description">
<paragraph id="684EBA299F221C39FF5064F57C113B7D" blockId="12.[151,1104,1225,1280]" box="[151,345,1255,1280]" pageId="12" pageNumber="413">
LT. Huelva (
<collectingCountry id="10E6FAB99F221C39FED264FA7C193B7D" box="[277,337,1255,1280]" name="Spain" pageId="12" pageNumber="413">Spain</collectingCountry>
)
</paragraph>
</subSubSection>
<subSubSection id="20EBE9A29F221C39FF5065307E053BC9" pageId="12" pageNumber="413" type="materials_examined">
<paragraph id="684EBA299F221C39FF5065307E053BC9" blockId="12.[151,1436,1325,2037]" pageId="12" pageNumber="413">
<emphasis id="5A85663B9F221C39FF5065307C2E3B35" bold="true" box="[151,358,1325,1352]" pageId="12" pageNumber="413">Material studied.</emphasis>
<materialsCitation id="D899B0749F221C39FEA96533784C3B35" ID-GBIF-Occurrence="2265605490" box="[366,1284,1326,1353]" collectingDate="1999-10-11" collectorName="El Abalario &amp; R. Macia &amp; J. Ylla" elevation="60" location="Huelva" pageId="12" pageNumber="413" specimenCount="3" specimenCount-female="3">
<specimenCount id="7EF771A09F221C39FEA965337CEE3B35" box="[366,422,1326,1352]" pageId="12" pageNumber="413" type="female">3♀♀</specimenCount>
<collectorName id="C504DFFF9F221C39FE6865337F783B35" box="[431,560,1326,1352]" pageId="12" pageNumber="413">El Abalario</collectorName>
(
<location id="6D2EECF29F221C39FD8765337FDE3B34" LSID="urn:lsid:plazi:treatment:E0580B3F9F221C38FF5064987C4D393A:6D2EECF29F221C39FD8765337FDE3B34" box="[576,662,1326,1353]" name="Huelva" pageId="12" pageNumber="413">Huelva</location>
),
<quantity id="AF0917CC9F221C39FD6C65337FA03B34" box="[683,744,1326,1353]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="413" unit="m" value="60.0">
<elevation id="E3DC5D1A9F221C39FD6C65337FA03B34" box="[683,744,1326,1353]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="413" unit="m" value="60.0">60 m</elevation>
</quantity>
, 29SQB00,
<date id="1C4F9CE99F221C39FCB465337EA23B35" box="[883,1002,1326,1352]" pageId="12" pageNumber="413" value="1999-10-11">
<collectingDate id="0C0B65019F221C39FCB465337EA23B35" box="[883,1002,1326,1352]" pageId="12" pageNumber="413" value="1999-10-11">11.X.1999</collectingDate>
</date>
,
<collectorName id="C504DFFF9F221C39FC30653279173B35" box="[1015,1119,1326,1353]" pageId="12" pageNumber="413">R. Macià</collectorName>
&amp;
<collectorName id="C504DFFF9F221C39FB41653279873B35" box="[1158,1231,1326,1353]" pageId="12" pageNumber="413">J. Ylla</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="D899B0749F221C39FAD665337EEC3B11" ID-GBIF-Occurrence="2265605462" collectingDate="2001-10-02" collectorName="R. Macia &amp; J. Ylla" elevation="60" location="El" municipality="Huelva" pageId="12" pageNumber="413" specimenCount="3" specimenCount-male="3">
<specimenCount id="7EF771A09F221C39FAD6653378013B35" box="[1297,1353,1326,1352]" pageId="12" pageNumber="413" type="male">3♂♂</specimenCount>
<location id="6D2EECF29F221C39FA96653378223B35" LSID="urn:lsid:plazi:treatment:E0580B3F9F221C38FF5064987C4D393A:6D2EECF29F221C39FA96653378223B35" box="[1361,1386,1326,1352]" municipality="Huelva" name="El" pageId="12" pageNumber="413">El</location>
Ab- alario (
<collectingMunicipality id="882A20539F221C39FF2F654F7C763B10" box="[232,318,1362,1389]" pageId="12" pageNumber="413">Huelva</collectingMunicipality>
),
<quantity id="AF0917CC9F221C39FE94654F7CD83B10" box="[339,400,1362,1389]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="413" unit="m" value="60.0">
<elevation id="E3DC5D1A9F221C39FE94654F7CD83B10" box="[339,400,1362,1389]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="413" unit="m" value="60.0">60 m</elevation>
</quantity>
, 29SQB00,
<date id="1C4F9CE99F221C39FDDB654F7FCF3B11" box="[540,647,1362,1388]" pageId="12" pageNumber="413" value="2001-10-02">
<collectingDate id="0C0B65019F221C39FDDB654F7FCF3B11" box="[540,647,1362,1388]" pageId="12" pageNumber="413" value="2001-10-02">2.X.2001</collectingDate>
</date>
,
<collectorName id="C504DFFF9F221C39FD53654E7FB53B11" box="[660,765,1362,1389]" pageId="12" pageNumber="413">R. Macià</collectorName>
&amp;
<collectorName id="C504DFFF9F221C39FCE2654E7E263B11" box="[805,878,1362,1389]" pageId="12" pageNumber="413">J. Ylla</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="D899B0749F221C39FC75654F7F473BED" ID-GBIF-Occurrence="2265605439" collectingDate="2002-10-03" collectorName="El Abalario &amp; R. Macia &amp; J. Ylla" elevation="60" location="Huelva" pageId="12" pageNumber="413" specimenCount="1" specimenCount-male="1">
<specimenCount id="7EF771A09F221C39FC75654F7E9D3B11" box="[946,981,1362,1388]" pageId="12" pageNumber="413" type="male">1♂</specimenCount>
<collectorName id="C504DFFF9F221C39FC19654F79283B11" box="[990,1120,1362,1388]" pageId="12" pageNumber="413">El Abalario</collectorName>
(
<location id="6D2EECF29F221C39FBB6654F798F3B10" LSID="urn:lsid:plazi:treatment:E0580B3F9F221C38FF5064987C4D393A:6D2EECF29F221C39FBB6654F798F3B10" box="[1137,1223,1362,1389]" name="Huelva" pageId="12" pageNumber="413">Huelva</location>
),
<quantity id="AF0917CC9F221C39FB1B654F78513B10" box="[1244,1305,1362,1389]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="413" unit="m" value="60.0">
<elevation id="E3DC5D1A9F221C39FB1B654F78513B10" box="[1244,1305,1362,1389]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="413" unit="m" value="60.0">60 m</elevation>
</quantity>
, 29SQB00,
<date id="1C4F9CE99F221C39FF50656B7C4A3BED" box="[151,258,1398,1424]" pageId="12" pageNumber="413" value="2002-10-03">
<collectingDate id="0C0B65019F221C39FF50656B7C4A3BED" box="[151,258,1398,1424]" pageId="12" pageNumber="413" value="2002-10-03">3.X.2002</collectingDate>
</date>
,
<collectorName id="C504DFFF9F221C39FECA656A7C3B3BED" box="[269,371,1398,1425]" pageId="12" pageNumber="413">R. Macià</collectorName>
&amp;
<collectorName id="C504DFFF9F221C39FE52656A7C943BED" box="[405,476,1398,1425]" pageId="12" pageNumber="413">J. Ylla</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="D899B0749F221C39FDDD656B78773BED" ID-GBIF-Occurrence="2265605440" box="[538,1343,1398,1425]" collectingDate="2017-05-01" collectorName="M. Huertas" country="Iceland" elevation="10" location="Saltes" municipality="Huelva" pageId="12" pageNumber="413" specimenCount="2" specimenCount-larva="2">
<specimenCount id="7EF771A09F221C39FDDD656B7F3A3BED" box="[538,626,1398,1424]" pageId="12" pageNumber="413" type="larva">2 larvae</specimenCount>
<collectingCountry id="10E6FAB99F221C39FDBE656B7FE93BED" box="[633,673,1398,1424]" name="Iceland" pageId="12" pageNumber="413">Isla</collectingCountry>
<location id="6D2EECF29F221C39FD60656B7FA23BED" LSID="urn:lsid:plazi:treatment:E0580B3F9F221C38FF5064987C4D393A:6D2EECF29F221C39FD60656B7FA23BED" box="[679,746,1398,1424]" country="Iceland" municipality="Huelva" name="Saltes" pageId="12" pageNumber="413">Saltés</location>
(
<collectingMunicipality id="882A20539F221C39FD3E656B7E073BEC" box="[761,847,1398,1425]" pageId="12" pageNumber="413">Huelva</collectingMunicipality>
),
<quantity id="AF0917CC9F221C39FCA6656B7ED43BEC" box="[865,924,1398,1425]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="413" unit="m" value="10.0">
<elevation id="E3DC5D1A9F221C39FCA6656B7ED43BEC" box="[865,924,1398,1425]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="413" unit="m" value="10.0">10 m</elevation>
</quantity>
, 29SPB81,
<date id="1C4F9CE99F221C39FBD9656B79CD3BED" box="[1054,1157,1398,1424]" pageId="12" pageNumber="413" value="2017-05-01">
<collectingDate id="0C0B65019F221C39FBD9656B79CD3BED" box="[1054,1157,1398,1424]" pageId="12" pageNumber="413" value="2017-05-01">1.V.2017</collectingDate>
</date>
,
<collectorName id="C504DFFF9F221C39FB57656A78453BEC" box="[1168,1293,1399,1425]" pageId="12" pageNumber="413">M. Huertas</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="D899B0749F221C39FA8D656B7E053BC9" ID-GBIF-Occurrence="2265605466" collectingDate="1986-09-22" collectorName="M. Huertas" country="Iceland" elevation="10" location="Saltes" municipality="Huelva" pageId="12" pageNumber="413" specimenCount="1" specimenCount-male="1">
<specimenCount id="7EF771A09F221C39FA8D656B78253BED" box="[1354,1389,1398,1424]" pageId="12" pageNumber="413" type="male">1♂</specimenCount>
<collectingCountry id="10E6FAB99F221C39FAB3656B78D43BED" box="[1396,1436,1398,1424]" name="Iceland" pageId="12" pageNumber="413">Isla</collectingCountry>
<location id="6D2EECF29F221C39FF5065877D923BC9" LSID="urn:lsid:plazi:treatment:E0580B3F9F221C38FF5064987C4D393A:6D2EECF29F221C39FF5065877D923BC9" box="[151,218,1434,1460]" country="Iceland" municipality="Huelva" name="Saltes" pageId="12" pageNumber="413">Saltés</location>
(
<collectingMunicipality id="882A20539F221C39FF2E65877C773BC8" box="[233,319,1434,1461]" pageId="12" pageNumber="413">Huelva</collectingMunicipality>
),
<quantity id="AF0917CC9F221C39FE9465877CC63BC8" box="[339,398,1434,1461]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="413" unit="m" value="10.0">
<elevation id="E3DC5D1A9F221C39FE9465877CC63BC8" box="[339,398,1434,1461]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="413" unit="m" value="10.0">10 m</elevation>
</quantity>
, 29SPB81,
<date id="1C4F9CE99F221C39FDD665877FDB3BC9" box="[529,659,1434,1460]" pageId="12" pageNumber="413" value="1986-09-22">
<collectingDate id="0C0B65019F221C39FDD665877FDB3BC9" box="[529,659,1434,1460]" pageId="12" pageNumber="413" value="1986-09-22">22.IX.1986</collectingDate>
</date>
,
<collectorName id="C504DFFF9F221C39FD5865867E553BC8" box="[671,797,1435,1461]" pageId="12" pageNumber="413">M. Huertas</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="20EBE9A29F221C39FF0065A07E493B81" pageId="12" pageNumber="413" type="diagnosis">
<paragraph id="684EBA299F221C39FF0065A07E493B81" blockId="12.[151,1436,1325,2037]" pageId="12" pageNumber="413">
<emphasis id="5A85663B9F221C39FF0065A07C0A3BA5" bold="true" box="[199,322,1469,1496]" pageId="12" pageNumber="413">Diagnosis.</emphasis>
This subspecies is characterized by the melanic, dark grey-brown forewings with a narrow costal band and a long dirty white stripe, parallel to the veins.
</paragraph>
</subSubSection>
<subSubSection id="20EBE9A29F221C38FF00661878CC3BEA" lastPageId="13" lastPageNumber="414" pageId="12" pageNumber="413" type="description">
<paragraph id="684EBA299F221C39FF0066187917385C" blockId="12.[151,1436,1325,2037]" box="[199,1119,1541,1569]" pageId="12" pageNumber="413">
<emphasis id="5A85663B9F221C39FF0066187F67385D" bold="true" box="[199,559,1541,1568]" pageId="12" pageNumber="413">
Description. Imago (
<figureCitation id="F0CAA6AC9F221C39FE7D66187F69385D" box="[442,545,1541,1568]" captionStart="FIGURES 1–8" captionStartId="9.[151,264,1350,1375]" captionTargetBox="[235,1351,205,1296]" captionTargetId="figure@9.[235,1355,200,1300]" captionTargetPageId="9" captionText="FIGURES 1–8. 1. C. cribraria cribraria, male, Villanueva de Sigena, Huesca, (Spain) (coll. RMV); 2. C. cribraria cribraria, female, Villanueva de Sigena, Huesca, (Spain) (coll. RMV); 3. C. cribraria benderi, male, El Abalario, Huelva, (Spain) (coll. RMV); 4. C. cribraria benderi, female, El Abalario, Huelva, (Spain) (coll. RMV); 5. C. cribraria rippertii, male, Llac de Colomers, Val d´Aran, Lleida, (Spain) (coll. RMV); 6. C.cribraria rippertii, female, Llac de Colomers, Val d´Aran, Lleida, (Spain) (coll. RMV); 7. C. cribraria ibicenca, male, Torrent des Fornàs, Eivissa, Balears, (Spain) (coll. RMV); 8. C. cribraria ibicenca, female, Torrent des Fornàs, Eivissa, Balears, (Spain) (coll. RMV)." httpUri="https://zenodo.org/record/3246050/files/figure.png" pageId="12" pageNumber="413">Figs 3–4</figureCitation>
).
</emphasis>
Average wingspan
<quantity id="AF0917CC9F221C39FCCC661B7E35385C" box="[779,893,1542,1569]" metricMagnitude="-2" metricUnit="m" metricValue="3.193" pageId="12" pageNumber="413" unit="mm" value="31.93">31.93 mm</quantity>
(n=15;
<quantity id="AF0917CC9F221C39FC12661B791A385C" box="[981,1106,1542,1569]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" metricValueMax="3.8" metricValueMin="2.6" pageId="12" pageNumber="413" unit="mm" value="32.0" valueMax="38.0" valueMin="26.0">26–38 mm</quantity>
).
</paragraph>
<paragraph id="684EBA299F221C39FF0066347C573814" blockId="12.[151,1436,1325,2037]" pageId="12" pageNumber="413">
<emphasis id="5A85663B9F221C39FF0066347CEF3839" bold="true" box="[199,423,1577,1604]" pageId="12" pageNumber="413">
Genitalia (
<figureCitation id="F0CAA6AC9F221C39FE8266347CD23839" box="[325,410,1577,1604]" captionStart="FIGURE 30" captionStartId="13.[151,250,1189,1214]" captionTargetBox="[170,1420,188,1145]" captionTargetId="figure@13.[164,1424,181,1147]" captionTargetPageId="13" captionText="FIGURE 30. C. cribraria benderi, male and female genitalia, gen. prep. 5729JG m. El Abalario, Huelva, (Spain), 5712JG f. El Abalario, Huelva, (Spain)" httpUri="https://zenodo.org/record/3246058/files/figure.png" pageId="12" pageNumber="413">Fig. 30</figureCitation>
).
</emphasis>
There are no structural differences in either sex from those of the nominate subspecies,
<taxonomicName id="AFF1C1AA9F221C39FA4466367C513815" authorityName="cribraria (Linnaeus" authorityYear="1758" class="Insecta" family="Arctiidae" genus="Coscinia" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="413" phylum="Arthropoda" rank="subSpecies" species="cribraria" subSpecies="cribraria">
<emphasis id="5A85663B9F221C39FA4466367C513815" italics="true" pageId="12" pageNumber="413">C. c. cribraria</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="684EBA299F221C39FF00666F7C173919" blockId="12.[151,1436,1325,2037]" pageId="12" pageNumber="413">
<emphasis id="5A85663B9F221C39FF00666F7CB538F1" bold="true" box="[199,509,1649,1676]" pageId="12" pageNumber="413">
Immature stages (
<figureCitation id="F0CAA6AC9F221C39FE5B666C7CB838F1" box="[412,496,1649,1676]" captionStart="FIGURE 46" captionStartId="30.[151,250,1916,1941]" captionTargetBox="[239,1347,1127,1855]" captionTargetId="figure@30.[239,1347,1127,1855]" captionTargetPageId="30" captionText="FIGURE 46. Immature state Coscinia cribraria benderi (picture MH)" httpUri="https://zenodo.org/record/3246076/files/figure.png" pageId="12" pageNumber="413">Fig. 46</figureCitation>
).
</emphasis>
The last instar larva measures
<quantity id="AF0917CC9F221C39FC94666F7E8738F0" box="[851,975,1650,1677]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="12" pageNumber="413" unit="mm" value="20.5" valueMax="21.0" valueMin="20.0">20–21 mm</quantity>
. Head capsule brownish, with the edge of the epicranial suture wide and whitish. Dorsal area dark brown, mixed with dark grey lines and spots, with a narrow white central line, bordered by a black zone; lateral sides brownish and ventral surface light grey with small dark spots; verruca D1, D2, SD1 dark grey; L1, L2, L3 light grey; thoracic legs dark brown; longer nails on abdominal prolegs. The pupae of both male and female straight, brown (not shiny); tips of the antennae touching each other in males, but not in females; the pterotheca tips are curved and touch each other along a shorter distance than in the nominate species.
</paragraph>
<paragraph id="684EBA299F221C39FF006773781E3989" blockId="12.[151,1436,1325,2037]" pageId="12" pageNumber="413">
<emphasis id="5A85663B9F221C39FF0067737CCB39F5" bold="true" box="[199,387,1902,1928]" pageId="12" pageNumber="413">Molecular data.</emphasis>
The three genetically investigated specimens differ by 0–0.22% in the COI Barcode from each other. The p-distances to other
<taxonomicName id="AFF1C1AA9F221C39FE32678E7F3639D1" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[501,638,1938,1965]" class="Insecta" family="Arctiidae" genus="Coscinia" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="cribraria">
<emphasis id="5A85663B9F221C39FE32678E7F3639D1" box="[501,638,1938,1965]" italics="true" pageId="12" pageNumber="413">C. cribraria</emphasis>
</taxonomicName>
specimens range from 1.30% to 7.56%; the lowest p-distance (1.30%) is with three
<taxonomicName id="AFF1C1AA9F221C39FEEF67AA7CF839AD" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[296,432,1974,2001]" class="Insecta" family="Arctiidae" genus="Coscinia" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="cribraria">
<emphasis id="5A85663B9F221C39FEEF67AA7CF839AD" box="[296,432,1974,2001]" italics="true" pageId="12" pageNumber="413">C. cribraria</emphasis>
</taxonomicName>
specimens from
<collectingCountry id="10E6FAB99F221C39FDA867AB7F8639AD" box="[623,718,1974,2000]" name="Grenada" pageId="12" pageNumber="413">Granada</collectingCountry>
in S.
<collectingCountry id="10E6FAB99F221C39FCC867AB7E0739AD" box="[783,847,1974,2000]" name="Spain" pageId="12" pageNumber="413">Spain</collectingCountry>
(ZMBN Lep478–480), the highest p-distance with a specimen of
<emphasis id="5A85663B9F221C39FEE167C67C883989" box="[294,448,2010,2037]" italics="true" pageId="12" pageNumber="413">C. c. ibicenca</emphasis>
from Formentera, the smaller of the two major Pityusic Islands (ZMBN Lep585).
</paragraph>
<caption id="3C8EEAA19F231C38FF5064B87CF03A9F" httpUri="https://zenodo.org/record/3246058/files/figure.png" pageId="13" pageNumber="414" startId="13.[151,250,1189,1214]" targetBox="[170,1420,188,1145]" targetPageId="13">
<paragraph id="684EBA299F231C38FF5064B87CF03A9F" blockId="13.[151,1437,1189,1250]" pageId="13" pageNumber="414">
<emphasis id="5A85663B9F231C38FF5064B87C603AC3" bold="true" box="[151,296,1189,1214]" pageId="13" pageNumber="414">FIGURE 30.</emphasis>
<taxonomicName id="AFF1C1AA9F231C38FEE864BB7CB73AC3" authorityName="Macià &amp; Mally &amp; Ylla &amp; Gastón &amp; Huertas" authorityYear="2019" baseAuthorityName="Marten" baseAuthorityYear="1957" box="[303,511,1190,1214]" class="Insecta" family="Arctiidae" genus="Coscinia" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="subSpecies" species="cribraria" subSpecies="benderi">
<emphasis id="5A85663B9F231C38FEE864BB7CB73AC3" box="[303,511,1190,1214]" italics="true" pageId="13" pageNumber="414">C. cribraria benderi</emphasis>
</taxonomicName>
, male and female genitalia, gen. prep. 5729JG m. El Abalario, Huelva, (Spain), 5712JG f. El Abalario, Huelva, (Spain)
</paragraph>
</caption>
<paragraph id="684EBA299F231C38FF00650C7C4D3B32" blockId="13.[151,1436,1296,1863]" pageId="13" pageNumber="414">
In the wingless gene, the three specimens are identical, and they differ by 0–0.30% from other specimens of
<taxonomicName id="AFF1C1AA9F231C38FA44650C7DB63B33" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Arctiidae" genus="Coscinia" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="cribraria">
<emphasis id="5A85663B9F231C38FA44650C7DB63B33" italics="true" pageId="13" pageNumber="414">C. cribraria</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="684EBA299F231C38FF00654478CC3BEA" blockId="13.[151,1436,1296,1863]" pageId="13" pageNumber="414">
<emphasis id="5A85663B9F231C38FF0065447CEB3B0F" box="[199,419,1368,1395]" italics="true" pageId="13" pageNumber="414">
<taxonomicName id="AFF1C1AA9F231C38FF0065447C633B0E" authorityName="Hubner" authorityYear="1819" box="[199,299,1369,1395]" class="Insecta" family="Arctiidae" genus="Coscinia" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="genus">Coscinia</taxonomicName>
<taxonomicName id="AFF1C1AA9F231C38FEF565447CEB3B0F" authorityName="Marten" authorityYear="1957" box="[306,419,1368,1394]" class="Insecta" family="Arctiidae" genus="Coscinia" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="benderi">c. benderi</taxonomicName>
</emphasis>
is well supported as monophyletic (1.00 PP, 99% BS). It is well-supported sister (1.00 PP, 94% BS) to a subclade of
<taxonomicName id="AFF1C1AA9F231C38FE7B65607F0B3BEB" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[444,579,1404,1431]" class="Insecta" family="Arctiidae" genus="Coscinia" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="cribraria">
<emphasis id="5A85663B9F231C38FE7B65607F0B3BEB" box="[444,579,1404,1431]" italics="true" pageId="13" pageNumber="414">C. cribraria</emphasis>
</taxonomicName>
comprising specimens from
<collectingCountry id="10E6FAB99F231C38FC4F65617E833BEB" box="[904,971,1404,1430]" name="Spain" pageId="13" pageNumber="414">Spain</collectingCountry>
,
<collectingCountry id="10E6FAB99F231C38FC11656179423BEB" box="[982,1034,1404,1430]" name="Italy" pageId="13" pageNumber="414">Italy</collectingCountry>
,
<collectingCountry id="10E6FAB99F231C38FBD2656079293BEA" box="[1045,1121,1405,1431]" name="France" pageId="13" pageNumber="414">France</collectingCountry>
and
<collectingCountry id="10E6FAB99F231C38FB51656179A03BEB" box="[1174,1256,1404,1430]" name="Austria" pageId="13" pageNumber="414">Austria</collectingCountry>
(see
<figureCitation id="F0CAA6AC9F231C38FAE36561783F3BEA" box="[1316,1399,1404,1431]" captionStart="FIGURE 62" captionStartId="43.[151,250,1865,1890]" captionTargetBox="[206,1235,194,1744]" captionTargetId="figure@43.[196,1250,194,1752]" captionTargetPageId="43" captionText="FIGURE 62. Phylogenetic tree (General Time Reversible model + Gamma distribution) obtained from Bayesian analysis of combined COI Barcode and wingless gene data of the Arctiinae specimens studied; numbers above branches are Posterior Probabilities ≥ 0.90, numbers below branches are Bootstrap values obtained from 1,000 Bootstrap replicates of a Maximum Likelihood analysis conducted with RaxML. Within the C. cribraria clade, the subspecies C. c. benderi, C. c. rippertii and C. c. ibicenca are marked in slightly darker green. The black vertical bars on the right mark BOLD BINs; the specimens of BIN ACE7017 are nested within BIN AAD9042. The scale bar (upper left) represents substitutions per nucleotide position." httpUri="https://zenodo.org/record/3246102/files/figure.png" pageId="13" pageNumber="414">Fig. 62</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="20EBE9A29F231C38FF0065BD7E69385B" pageId="13" pageNumber="414" type="biology_ecology">
<paragraph id="684EBA299F231C38FF0065BD79563BA3" blockId="13.[151,1436,1296,1863]" pageId="13" pageNumber="414">
<emphasis id="5A85663B9F231C38FF0065BD7C6F3BC6" bold="true" box="[199,295,1440,1467]" pageId="13" pageNumber="414">Biology.</emphasis>
Univoltine, flying from September to mid-November. Both sexes are attracted by artificial light. In its univoltinism, it differs from
<taxonomicName id="AFF1C1AA9F231C38FE1165D87FD23BA3" baseAuthorityName="Hübner" baseAuthorityYear="1804" box="[470,666,1476,1503]" class="Insecta" family="Arctiidae" genus="Coscinia" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="chrysocephala">
<emphasis id="5A85663B9F231C38FE1165D87FD23BA3" box="[470,666,1476,1503]" italics="true" pageId="13" pageNumber="414">C. chrysocephala</emphasis>
</taxonomicName>
, which is bivoltine and sympatric.
</paragraph>
<paragraph id="684EBA299F231C38FF0065F57E69385B" blockId="13.[151,1436,1296,1863]" pageId="13" pageNumber="414">
The larvae feed on a great variety of plants including
<emphasis id="5A85663B9F231C38FCD565F47CDD385B" italics="true" pageId="13" pageNumber="414">
<taxonomicName id="AFF1C1AA9F231C38FCD565F47E9F387F" authority=", Halimium" authorityName="Halimium" box="[786,983,1512,1539]" class="Magnoliopsida" family="Cistaceae" genus="Cistus" kingdom="Plantae" order="Malvales" pageId="13" pageNumber="414" phylum="Tracheophyta" rank="genus">Cistus, Halimium</taxonomicName>
,
<taxonomicName id="AFF1C1AA9F231C38FC2665F47922387F" box="[993,1130,1513,1538]" class="Magnoliopsida" family="Lamiaceae" genus="Rosmarinus" kingdom="Plantae" order="Lamiales" pageId="13" pageNumber="414" phylum="Tracheophyta" rank="genus">Rosmarinus</taxonomicName>
,
<taxonomicName id="AFF1C1AA9F231C38FBB465F47CDD385B" authority=", Rhamnus, Linaria, Salsola, Limoniastrum" authorityName="Rhamnus, Linaria, Salsola, Limoniastrum" class="Magnoliopsida" family="Polygonaceae" genus="Rumex" kingdom="Plantae" order="Polygonales" pageId="13" pageNumber="414" phylum="Tracheophyta" rank="genus">Rumex, Rhamnus, Linaria, Salsola, Limoniastrum</taxonomicName>
</emphasis>
, and
<taxonomicName id="AFF1C1AA9F231C38FE1566117E55385B" authority="(Ylla et al. 2010)" authorityName="Ylla" authorityYear="2010" baseAuthorityName="Ylla et al." baseAuthorityYear="2010" box="[466,797,1548,1574]" class="Magnoliopsida" family="Polygonaceae" genus="Polygonum" kingdom="Plantae" order="Polygonales" pageId="13" pageNumber="414" phylum="Tracheophyta" rank="genus">
<emphasis id="5A85663B9F231C38FE1566117F1A385B" box="[466,594,1548,1574]" italics="true" pageId="13" pageNumber="414">Polygonum</emphasis>
(
<bibRefCitation id="0C60C7D89F231C38FDA666117E5D385B" author="Ylla, J. &amp; Macia, R. &amp; Gaston, F. J." box="[609,789,1548,1574]" pageId="13" pageNumber="414" refId="ref27969" refString="Ylla, J., Macia, R. &amp; Gaston, F. J. (2010) Manual de identificacion y guia de campo de los Arctidos de la Peninsula Iberica y Baleares. Arganda Editio, Barcelona, 290 pp." type="book" year="2010">
Ylla
<emphasis id="5A85663B9F231C38FD5E66107F9A385B" box="[665,722,1548,1574]" italics="true" pageId="13" pageNumber="414">et al.</emphasis>
2010
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="20EBE9A29F231C38FF00662D7FC93813" pageId="13" pageNumber="414" type="distribution">
<paragraph id="684EBA299F231C38FF00662D7FC93813" blockId="13.[151,1436,1296,1863]" pageId="13" pageNumber="414">
<emphasis id="5A85663B9F231C38FF00662D7C843836" bold="true" box="[199,460,1584,1611]" pageId="13" pageNumber="414">
Distribution (
<figureCitation id="F0CAA6AC9F231C38FEAE662D7CF73836" box="[361,447,1584,1611]" captionStart="FIGURE 52" captionStartId="34.[151,250,1073,1098]" captionTargetBox="[234,1356,191,1020]" captionTargetId="figure@34.[227,1360,188,1024]" captionTargetPageId="34" captionText="FIGURE 52. Distribution map of C. cribraria benderi." httpUri="https://zenodo.org/record/3246088/files/figure.png" pageId="13" pageNumber="414">Fig. 52</figureCitation>
).
</emphasis>
Iberian endemic, known from a few coastal localities in the Province of Huelva, from Punta
<collectingRegion id="AA3574CB9F231C38FF1966497C7C3813" box="[222,308,1620,1646]" country="Italy" name="Umbria" pageId="13" pageNumber="414">Umbría</collectingRegion>
to the Doñana National Park.
</paragraph>
</subSubSection>
<subSubSection id="20EBE9A29F231C38FF0066647C4D393A" pageId="13" pageNumber="414" type="discussion">
<paragraph id="684EBA299F231C38FF0066647C4D393A" blockId="13.[151,1436,1296,1863]" pageId="13" pageNumber="414">
<emphasis id="5A85663B9F231C38FF0066647C7338EE" bold="true" box="[199,315,1657,1683]" pageId="13" pageNumber="414">Remarks.</emphasis>
The specific validity of
<taxonomicName id="AFF1C1AA9F231C38FD8E66647FF738EF" authorityName="Marten" authorityYear="1957" box="[585,703,1656,1683]" class="Insecta" family="Arctiidae" genus="Coscinia" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="benderi">
<emphasis id="5A85663B9F231C38FD8E66647FF738EF" box="[585,703,1656,1683]" italics="true" pageId="13" pageNumber="414">C. benderi</emphasis>
</taxonomicName>
has been discussed by several authors for years. The results of the present molecular study and the genitalia do not allow us to consider it as a
<emphasis id="5A85663B9F231C38FC146681792938CB" box="[979,1121,1692,1718]" italics="true" pageId="13" pageNumber="414">bona species</emphasis>
, different from
<taxonomicName id="AFF1C1AA9F231C38FAC8668078DE38CB" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1295,1430,1692,1719]" class="Insecta" family="Arctiidae" genus="Coscinia" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="cribraria">
<emphasis id="5A85663B9F231C38FAC8668078DE38CB" box="[1295,1430,1692,1719]" italics="true" pageId="13" pageNumber="414">C. cribraria</emphasis>
</taxonomicName>
. Nevertheless, its univoltinism, the allopatry with other forms of this species, the climatic conditions of the biotope and the particular geological terrain where it lives (sandy alluvial deposits associated with marshy areas), as well as the peculiarities of the biology and differences in the immature stages lead us to place it as a subspecies of
<taxonomicName id="AFF1C1AA9F231C38FA4467147DB6393B" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Arctiidae" genus="Coscinia" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="cribraria">
<emphasis id="5A85663B9F231C38FA4467147DB6393B" italics="true" pageId="13" pageNumber="414">C. cribraria</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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