481 lines
62 KiB
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481 lines
62 KiB
XML
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<mods:title id="5516ED85436FC75F71B6379760DFB72B">A taxonomic revision of two local endemic Radix spp. (Gastropoda: Lymnaeidae) from Khodutka geothermal area, Kamchatka, Russian Far East</mods:title>
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<treatment id="B31787903555FFA2FF6DF96A4BE601B9" ID-DOI="http://doi.org/10.5281/zenodo.5662764" ID-GBIF-Taxon="119589466" ID-Zenodo-Dep="5662764" LSID="urn:lsid:plazi:treatment:B31787903555FFA2FF6DF96A4BE601B9" httpUri="http://treatment.plazi.org/id/B31787903555FFA2FF6DF96A4BE601B9" lastPageId="4" lastPageNumber="588" pageId="1" pageNumber="586">
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<paragraph id="3B0136863555FFA7FF6DF96A4A1E06DF" blockId="1.[151,582,1697,1757]" box="[151,582,1697,1723]" pageId="1" pageNumber="586">
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<heading id="604981EA3555FFA7FF6DF96A4A1E06DF" bold="true" box="[151,582,1697,1723]" fontSize="11" level="1" pageId="1" pageNumber="586" reason="1">
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<taxonomicName id="FCBE4D053555FFA7FF6DF96A4A1E06DF" ID-CoL="78S5G" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[151,582,1697,1723]" class="Gastropoda" family="Lymnaeidae" genus="Radix" kingdom="Animalia" order="Hygrophila" pageId="1" pageNumber="586" phylum="Mollusca" rank="species" species="auricularia">
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<emphasis id="09CAEA943555FFA7FF6DF96A4A1E06DF" bold="true" box="[151,582,1697,1723]" pageId="1" pageNumber="586">
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<emphasis id="09CAEA943555FFA7FF6DF96A493706DF" bold="true" box="[151,367,1697,1723]" italics="true" pageId="1" pageNumber="586">Radix auricularia</emphasis>
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(Linnaeus, 1758)
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</emphasis>
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</paragraph>
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<paragraph id="3B0136863555FFA7FF6DF90F492A06B8" blockId="1.[151,582,1697,1757]" box="[151,370,1732,1757]" pageId="1" pageNumber="586">
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(
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<figureCitation id="A3852A033555FFA7FF65F90F495306B8" box="[159,267,1732,1757]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1692,1715]" captionTargetBox="[230,1360,184,1674]" captionTargetId="figure@2.[223,1367,178,1687]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Radix auricularia (Linnaeus, 1758) from Russian Far East. Figs. 1 - 4. Shell and proximal part of the female genitalia. Scale bar 2 mm. Fig. 1. Holotype of Lymnaea (R.) hadutkae Kruglov & Starobogatov, 1989 syn. n. (shell photo: M. V. Vinarski; genitalia redrawn from the protologue, Fig. 2.7). Fig. 2. Holotype of L. (R.) thermokamtschatica Kruglov & Starobogatov, 1989 syn. n. (shell photo: M. V. Vinarski; genitalia redrawn from the protologue, Fig. 2.9). Fig. 3. Newly collected specimen from the Khodutka geothermal area, Kamchatka (photos: O. V. Aksenova). Fig. 4. Specimen from Malkinskie hot springs, Kamchatka (photos: O. V. Aksenova). Fig. 5. Bayesian phylogram of haplotypes based on mitochondrial COI gene dataset (Appendixes 1 & 2). The scale bar indicates the branch length. Asterisks: Posterior probabilities ≥ 0.95; other significant node support values are mentioned in the figure. Haplotypes exclusively from NCBI’s Genbank are marked with circumflex accents (˄). The well-supported clade with two haplotypes from the Khodutka geothermal area is highlighted in blue. Fig. 6. General view of the Khodutka warm lake. Fig. 7. Map of observed localities on the Russian Far East. The red circles represent species records. Locality codes are given in Appendix 1. The digital elevation model and other layers of the map were added from Esri Data & Maps 10 dataset (Map: M. Yu. Gofarov)." httpUri="https://zenodo.org/record/225342/files/figure.png" pageId="1" pageNumber="586">Figs. 1–7</figureCitation>
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,
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<tableCitation id="763C033D3555FFA7FEECF90F493F06B8" box="[278,359,1732,1757]" captionStart="TABLE 1" captionStartId="3.[151,239,673,696]" captionTargetBox="[159,1424,755,1475]" captionTargetPageId="3" captionText="TABLE 1. The comparison of type specimens of endemic Lymnaea (Radix) species from Khodutka geothermal area with R. auricularia based on morphological and anatomical distinguishing features (Glöer & Pešić 2008)" httpUri="http://table.plazi.org/id/6FC1660E3557FFA5FF6DFD6A4C9902BC" pageId="1" pageNumber="586" tableUuid="6FC1660E3557FFA5FF6DFD6A4C9902BC">Table 1</tableCitation>
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)
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</paragraph>
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<taxonomicName id="FCBE4D053555FFA7FF6DF8C64ABD0746" ID-CoL="3WS3P" authority="Kruglov & Starobogatov, 1989" authorityName="Kruglov & Starobogatov" authorityYear="1989" box="[151,741,1804,1826]" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" kingdom="Animalia" order="Hygrophila" pageId="1" pageNumber="586" phylum="Mollusca" rank="species" species="hadutkae" subGenus="Radix">
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<emphasis id="09CAEA943555FFA7FF6DF8C648AB0746" box="[151,243,1805,1826]" italics="true" pageId="1" pageNumber="586">Lymnaea</emphasis>
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(
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<emphasis id="09CAEA943555FFA7FEFBF8C749640745" box="[257,316,1804,1825]" italics="true" pageId="1" pageNumber="586">Radix</emphasis>
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)
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<emphasis id="09CAEA943555FFA7FEB1F8C749F00745" box="[331,424,1804,1825]" italics="true" pageId="1" pageNumber="586">hadutkae</emphasis>
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<bibRefCitation id="5F2F4B773555FFA7FE57F8C74ABD0746" author="Kruglov" box="[429,741,1804,1826]" pageId="1" pageNumber="586" refString="Kruglov, N. D. & Starobogatov, Ya. I. (1989) Morphology and taxonomy of the molluscs from the subgenus Radix of the genus Lymnaea (Gastropoda, Pulmonata, Lymnaeidae) from Siberia and Far East of the USSR. Zoologicheskii Zhurnal, 68 (5), 17 - 30. [in Russian]" type="journal article" year="1989">Kruglov & Starobogatov, 1989</bibRefCitation>
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</taxonomicName>
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<emphasis id="09CAEA943555FFA7FD16F8C74B690745" bold="true" box="[748,817,1804,1825]" pageId="1" pageNumber="586">
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<taxonomicNameLabel id="12F957EF3555FFA7FD16F8C74B690745" box="[748,817,1804,1825]" pageId="1" pageNumber="586">syn. n.</taxonomicNameLabel>
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</emphasis>
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||
: 22, figs. 1.12 & 2.7;
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<treatmentCitation id="BA1F10973555FFA7FBF5F8C74D370746" author="Kruglov" box="[1039,1391,1804,1826]" page="85" pageId="1" pageNumber="586" year="1993">
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<bibRefCitation id="5F2F4B773555FFA7FBF5F8C74D120746" author="Kruglov" box="[1039,1354,1804,1826]" pageId="1" pageNumber="586" refString="Kruglov, N. D. & Starobogatov, Ya. I. (1993) Guide to recent molluscs of northern Eurasia. 3. Annotated and illustrated catalogue of species of the family Lymnaeidae (Gastropoda Pulmonata Lymnaeiformes) of Palaearctic and adjacent river drainage areas. Part 1. Ruthenica, 3 (1), 65 - 92." type="journal article" year="1993">Kruglov & Starobogatov, 1993</bibRefCitation>
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: 85
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</treatmentCitation>
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, fig. 11D
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<paragraph id="3B0136863555FFA7FF6DF8814925071A" blockId="1.[151,1437,1804,1918]" pageId="1" pageNumber="586">
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<treatmentCitationGroup id="1BAE11A83555FFA7FF6DF8814925071A" pageId="1" pageNumber="586">
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<taxonomicName id="FCBE4D053555FFA7FF6DF8814B3E073B" ID-CoL="3WS6X" authority="Kruglov & Starobogatov, 1989" authorityName="Kruglov & Starobogatov" authorityYear="1989" box="[151,870,1865,1887]" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" kingdom="Animalia" order="Hygrophila" pageId="1" pageNumber="586" phylum="Mollusca" rank="species" species="thermokamtschatica" subGenus="Radix">
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<emphasis id="09CAEA943555FFA7FF6DF88148AB073B" box="[151,243,1866,1887]" italics="true" pageId="1" pageNumber="586">Lymnaea</emphasis>
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(
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<emphasis id="09CAEA943555FFA7FEFEF8814967073B" box="[260,319,1866,1887]" italics="true" pageId="1" pageNumber="586">Radix</emphasis>
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)
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<emphasis id="09CAEA943555FFA7FEAAF8814A45073B" box="[336,541,1866,1887]" italics="true" pageId="1" pageNumber="586">thermokamtschatica</emphasis>
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<bibRefCitation id="5F2F4B773555FFA7FDDCF8824B3E073B" author="Kruglov" box="[550,870,1865,1887]" pageId="1" pageNumber="586" refString="Kruglov, N. D. & Starobogatov, Ya. I. (1989) Morphology and taxonomy of the molluscs from the subgenus Radix of the genus Lymnaea (Gastropoda, Pulmonata, Lymnaeidae) from Siberia and Far East of the USSR. Zoologicheskii Zhurnal, 68 (5), 17 - 30. [in Russian]" type="journal article" year="1989">Kruglov & Starobogatov, 1989</bibRefCitation>
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</taxonomicName>
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<emphasis id="09CAEA943555FFA7FC94F8814BEE073B" bold="true" box="[878,950,1866,1887]" pageId="1" pageNumber="586">
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<taxonomicNameLabel id="12F957EF3555FFA7FC94F8814BEE073B" box="[878,950,1866,1887]" pageId="1" pageNumber="586">syn. n.</taxonomicNameLabel>
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</emphasis>
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||
: 22, figs. 1.9 & 2.9;
|
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<treatmentCitation id="BA1F10973555FFA7FB6DF882497A071A" author="Kruglov" page="85" pageId="1" pageNumber="586" year="1993">
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<bibRefCitation id="5F2F4B773555FFA7FB6DF88248A5071A" author="Kruglov" pageId="1" pageNumber="586" refString="Kruglov, N. D. & Starobogatov, Ya. I. (1993) Guide to recent molluscs of northern Eurasia. 3. Annotated and illustrated catalogue of species of the family Lymnaeidae (Gastropoda Pulmonata Lymnaeiformes) of Palaearctic and adjacent river drainage areas. Part 1. Ruthenica, 3 (1), 65 - 92." type="journal article" year="1993">Kruglov & Starobogatov, 1993</bibRefCitation>
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: 85
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</treatmentCitation>
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, fig. 11E
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<paragraph id="3B0136863556FFA4FF6DF9574C980791" blockId="2.[151,1437,1692,2037]" pageId="2" pageNumber="587">
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<emphasis id="09CAEA943556FFA4FF6DF957496406D5" bold="true" box="[151,316,1692,1715]" pageId="2" pageNumber="587">FIGURES 1–7.</emphasis>
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<taxonomicName id="FCBE4D053556FFA4FEBEF9574AE806D6" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[324,688,1692,1714]" class="Gastropoda" family="Lymnaeidae" genus="Radix" kingdom="Animalia" order="Hygrophila" pageId="2" pageNumber="587" phylum="Mollusca" rank="species" species="auricularia">
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<emphasis id="09CAEA943556FFA4FEBEF957492706D5" box="[324,383,1692,1713]" italics="true" pageId="2" pageNumber="587">Radix</emphasis>
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<emphasis id="09CAEA943556FFA4FE7CF95749A106D5" box="[390,505,1692,1713]" italics="true" pageId="2" pageNumber="587">auricularia</emphasis>
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(Linnaeus, 1758)
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</taxonomicName>
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from Russian Far East.
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<emphasis id="09CAEA943556FFA4FC57F9574C4806D5" bold="true" box="[941,1040,1692,1714]" pageId="2" pageNumber="587">Figs. 1-4.</emphasis>
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Shell and proximal part of the female genitalia. Scale bar 2 mm.
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<emphasis id="09CAEA943556FFA4FE5EF97149BE06AB" bold="true" box="[420,486,1722,1744]" pageId="2" pageNumber="587">Fig. 1.</emphasis>
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Holotype of
|
||
<taxonomicName id="FCBE4D053556FFA4FD91F9704CCC06B4" authority="Kruglov & Starobogatov, 1989" authorityName="Kruglov & Starobogatov" authorityYear="1989" box="[619,1172,1722,1744]" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" kingdom="Animalia" order="Hygrophila" pageId="2" pageNumber="587" phylum="Mollusca" rank="species" species="hadutkae" subGenus="Radix">
|
||
<emphasis id="09CAEA943556FFA4FD91F9704A9F06B4" box="[619,711,1723,1744]" italics="true" pageId="2" pageNumber="587">Lymnaea</emphasis>
|
||
(
|
||
<emphasis id="09CAEA943556FFA4FD2FF9704AB406B4" box="[725,748,1723,1744]" italics="true" pageId="2" pageNumber="587">R.</emphasis>
|
||
)
|
||
<emphasis id="09CAEA943556FFA4FD03F9714B0D06AB" box="[761,853,1722,1743]" italics="true" pageId="2" pageNumber="587">hadutkae</emphasis>
|
||
Kruglov & Starobogatov, 1989
|
||
</taxonomicName>
|
||
<emphasis id="09CAEA943556FFA4FB60F9714C8706AB" bold="true" box="[1178,1247,1722,1743]" pageId="2" pageNumber="587">
|
||
<taxonomicNameLabel id="12F957EF3556FFA4FB60F9714C8706AB" box="[1178,1247,1722,1743]" pageId="2" pageNumber="587">syn. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(shell photo: M.V. Vinarski; genitalia redrawn from the protologue, Fig. 2.7).
|
||
<emphasis id="09CAEA943556FFA4FCE0F91C4B3A0688" bold="true" box="[794,866,1751,1773]" pageId="2" pageNumber="587">Fig. 2.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="FCBE4D053556FFA4FC01F9134938076E" authority="Kruglov & Starobogatov, 1989" authorityName="Kruglov & Starobogatov" authorityYear="1989" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" kingdom="Animalia" order="Hygrophila" pageId="2" pageNumber="587" phylum="Mollusca" rank="species" species="thermokamtschatica" subGenus="Radix">
|
||
<emphasis id="09CAEA943556FFA4FC01F9134C480689" box="[1019,1040,1752,1773]" italics="true" pageId="2" pageNumber="587">L.</emphasis>
|
||
(
|
||
<emphasis id="09CAEA943556FFA4FBDEF9134C620689" box="[1060,1082,1752,1773]" italics="true" pageId="2" pageNumber="587">R.</emphasis>
|
||
)
|
||
<emphasis id="09CAEA943556FFA4FBB4F91C4D430688" box="[1102,1307,1751,1772]" italics="true" pageId="2" pageNumber="587">thermokamtschatica</emphasis>
|
||
Kruglov & Starobogatov, 1989
|
||
</taxonomicName>
|
||
<emphasis id="09CAEA943556FFA4FE91F93E49ED076E" bold="true" box="[363,437,1781,1802]" pageId="2" pageNumber="587">
|
||
<taxonomicNameLabel id="12F957EF3556FFA4FE91F93E49ED076E" box="[363,437,1781,1802]" pageId="2" pageNumber="587">syn. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(shell photo: M.V. Vinarski; genitalia redrawn from the protologue, Fig. 2.9).
|
||
<emphasis id="09CAEA943556FFA4FAFCF93F4D15076D" bold="true" box="[1286,1357,1780,1802]" pageId="2" pageNumber="587">Fig. 3.</emphasis>
|
||
Newly collected specimen from the Khodutka geothermal area, Kamchatka (photos: O.V. Aksenova).
|
||
<emphasis id="09CAEA943556FFA4FB59F8D94CB40743" bold="true" box="[1187,1260,1810,1832]" pageId="2" pageNumber="587">Fig. 4.</emphasis>
|
||
Specimen from Malkinskie hot springs, Kamchatka (photos: O.V. Aksenova).
|
||
<emphasis id="09CAEA943556FFA4FCAAF8E44BC40720" bold="true" box="[848,924,1839,1861]" pageId="2" pageNumber="587">Fig. 5.</emphasis>
|
||
Bayesian phylogram of haplotypes based on mitochondrial COI gene dataset (Appendixes 1 & 2). The scale bar indicates the branch length. Asterisks: Posterior probabilities ≥0.95; other significant node support values are mentioned in the figure. Haplotypes exclusively from NCBI’s Genbank are marked with circumflex accents (
|
||
<emphasis id="09CAEA943556FFA4FD66F84D4AF207F9" bold="true" box="[668,682,1926,1949]" pageId="2" pageNumber="587">˄</emphasis>
|
||
). The well-supported clade with two haplotypes from the Khodutka geothermal area is highlighted in blue.
|
||
<emphasis id="09CAEA943556FFA4FDDFF86F4A3007DD" bold="true" box="[549,616,1956,1978]" pageId="2" pageNumber="587">Fig. 6.</emphasis>
|
||
General view of the Khodutka warm lake.
|
||
<emphasis id="09CAEA943556FFA4FBD9F86F4C3E07DE" bold="true" box="[1059,1126,1956,1978]" pageId="2" pageNumber="587">Fig. 7.</emphasis>
|
||
Map of observed localities on the Russian Far East. The red circles represent species records. Locality codes are given in Appendix 1. The digital elevation model and other layers of the map were added from Esri Data & Maps 10 dataset (Map: M.Yu. Gofarov).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="73A4650D3557FFA5FF6DFF5C49790213" pageId="3" pageNumber="588" type="materials_examined">
|
||
<paragraph id="3B0136863557FFA5FF6DFF5C49790213" blockId="3.[151,1436,151,631]" pageId="3" pageNumber="588">
|
||
<emphasis id="09CAEA943557FFA5FF6DFF5C49DD00D4" bold="true" box="[151,389,151,176]" pageId="3" pageNumber="588">Material examined.</emphasis>
|
||
<typeStatus id="E40588243557FFA5FE6BFF5C49A100D4" box="[401,505,151,176]" pageId="3" pageNumber="588" type="holotype">Holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="FCBE4D053557FFA5FDD2FF524B3800D4" box="[552,864,151,176]" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" kingdom="Animalia" order="Hygrophila" pageId="3" pageNumber="588" phylum="Mollusca" rank="species" species="hadutkae" subGenus="Radix">
|
||
<emphasis id="09CAEA943557FFA5FDD2FF524AD600D4" box="[552,654,153,176]" italics="true" pageId="3" pageNumber="588">Lymnaea</emphasis>
|
||
(
|
||
<emphasis id="09CAEA943557FFA5FD59FF5C4ABC00D4" box="[675,740,151,176]" italics="true" pageId="3" pageNumber="588">Radix</emphasis>
|
||
)
|
||
<emphasis id="09CAEA943557FFA5FD03FF5C4B3800D4" box="[761,864,151,176]" italics="true" pageId="3" pageNumber="588">hadutkae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<collectionCode id="5DAFAE433557FFA5FC8EFF534BF000D4" box="[884,936,152,176]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="3" pageNumber="588">ZIN</collectionCode>
|
||
),
|
||
<emphasis id="09CAEA943557FFA5FC45FF5C4C0D00D4" bold="true" box="[959,1109,151,176]" pageId="3" pageNumber="588">Kamchatka:</emphasis>
|
||
“Khodutkinskiy hot spring, gradient t° = 15–41°[C], opt. ~ 25–27°[C],
|
||
<date id="4F0010463557FFA5FD84FF774AB200B1" box="[638,746,188,213]" pageId="3" pageNumber="588" value="1979-09-08">8.ix.1979</date>
|
||
, N.N. Khmeleva leg.” (no. 510–1982).
|
||
<typeStatus id="E40588243557FFA5FB52FF774D4800B1" box="[1192,1296,188,213]" pageId="3" pageNumber="588" type="holotype">Holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="FCBE4D053557FFA5FACCFF764982009C" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" kingdom="Animalia" order="Hygrophila" pageId="3" pageNumber="588" phylum="Mollusca" rank="species" species="thermokamtschatica" subGenus="Radix">
|
||
<emphasis id="09CAEA943557FFA5FACCFF764DC400B0" box="[1334,1436,189,212]" italics="true" pageId="3" pageNumber="588">Lymnaea</emphasis>
|
||
(
|
||
<emphasis id="09CAEA943557FFA5FF5AFF1448B9009C" box="[160,225,223,248]" italics="true" pageId="3" pageNumber="588">Radix</emphasis>
|
||
)
|
||
<emphasis id="09CAEA943557FFA5FF0DFF144982009C" box="[247,474,223,248]" italics="true" pageId="3" pageNumber="588">thermokamtschatica</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<collectionCode id="5DAFAE433557FFA5FE15FF2B4A7B009C" box="[495,547,224,248]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="3" pageNumber="588">ZIN</collectionCode>
|
||
),
|
||
<emphasis id="09CAEA943557FFA5FDC6FF144A89009C" bold="true" box="[572,721,223,248]" pageId="3" pageNumber="588">Kamchatka:</emphasis>
|
||
“Khodutkinskiy hot spring, gradient t° = 15–41°[C], opt. ~ 25–27°[C],
|
||
<date id="4F0010463557FFA5FED8FECF49D50179" box="[290,397,260,285]" pageId="3" pageNumber="588" value="1979-09-08">8.ix.1979</date>
|
||
, N.N. Khmeleva leg.” (no. 510–1982). Topotypes (
|
||
<collectionCode id="5DAFAE433557FFA5FC0DFECF4C630178" box="[1015,1083,260,284]" pageId="3" pageNumber="588">INEP</collectionCode>
|
||
),
|
||
<specimenCount id="2DB8FD0F3557FFA5FBAFFECF4CAD0179" box="[1109,1269,260,285]" pageId="3" pageNumber="588" type="generic">31 specimens</specimenCount>
|
||
,
|
||
<emphasis id="09CAEA943557FFA5FAFDFECF4DC40179" bold="true" box="[1287,1436,260,285]" pageId="3" pageNumber="588">Kamchatka:</emphasis>
|
||
Khodutka geothermal area, warm lake and warm river source,
|
||
<date id="4F0010463557FFA5FCA2FEE14BB70127" box="[856,1007,295,323]" pageId="3" pageNumber="588" value="2013-06-06">6th June 2013</date>
|
||
, Aksenova, Bespalaya & Bolotov leg. (
|
||
<collectionCode id="5DAFAE433557FFA5FF65FE9948BB010E" box="[159,227,338,362]" pageId="3" pageNumber="588">INEP</collectionCode>
|
||
).
|
||
<emphasis id="09CAEA943557FFA5FF0CFE9A49D3010E" bold="true" box="[246,395,337,362]" pageId="3" pageNumber="588">Kamchatka:</emphasis>
|
||
<specimenCount id="2DB8FD0F3557FFA5FE68FE9A4A46010E" box="[402,542,337,362]" pageId="3" pageNumber="588" type="generic">6 specimens</specimenCount>
|
||
, Paratunskie hot springs,
|
||
<date id="4F0010463557FFA5FCC7FE994B8C010E" box="[829,980,334,362]" pageId="3" pageNumber="588" value="2013-06-08">8th June 2013</date>
|
||
, Aksenova, Bespalaya & Bolotov leg.;
|
||
<specimenCount id="2DB8FD0F3557FFA5FA74FE99495701F5" pageId="3" pageNumber="588" type="generic">7 specimens</specimenCount>
|
||
, Malkinskie hot springs,
|
||
<date id="4F0010463557FFA5FDCFFEB24A9601F5" box="[565,718,373,401]" pageId="3" pageNumber="588" value="2012-07-03">3rd July 2012</date>
|
||
, Bespalaya, Bolotov & Vikhrev leg.;
|
||
<specimenCount id="2DB8FD0F3557FFA5FB7FFEB24D4C01F5" box="[1157,1300,376,401]" pageId="3" pageNumber="588" type="generic">4 specimens</specimenCount>
|
||
, Plotnikova River (Bolshaya River Basin),
|
||
<date id="4F0010463557FFA5FE02FE544AD601D3" box="[504,654,412,440]" pageId="3" pageNumber="588" value="2012-07-03">3rd July 2012</date>
|
||
, Bespalaya, Bolotov & Vikhrev leg. (
|
||
<collectionCode id="5DAFAE433557FFA5FBC1FE544C2701D3" box="[1083,1151,415,439]" pageId="3" pageNumber="588">INEP</collectionCode>
|
||
).
|
||
<emphasis id="09CAEA943557FFA5FB6EFE544DDD01DC" bold="true" box="[1172,1413,415,440]" pageId="3" pageNumber="588">Kurile Archipelago:</emphasis>
|
||
<specimenCount id="2DB8FD0F3557FFA5FA74FE54495701BA" pageId="3" pageNumber="588" type="generic">9 specimens</specimenCount>
|
||
, Kunashir Island, Aliger Lake,
|
||
<date id="4F0010463557FFA5FD89FE0D4B4B01BA" box="[627,787,451,478]" pageId="3" pageNumber="588" value="2011-07-27">27th July 2011</date>
|
||
, Bespalaya & Aksenova leg. (
|
||
<collectionCode id="5DAFAE433557FFA5FB92FE0D4CF401BA" box="[1128,1196,454,478]" pageId="3" pageNumber="588">INEP</collectionCode>
|
||
).
|
||
<emphasis id="09CAEA943557FFA5FB3AFE0E4DDE01BA" bold="true" box="[1216,1414,453,478]" pageId="3" pageNumber="588">Sakhalin Island:</emphasis>
|
||
<specimenCount id="2DB8FD0F3557FFA5FA77FE0D49570261" pageId="3" pageNumber="588" type="generic">2 specimens</specimenCount>
|
||
, Tym River,
|
||
<date id="4F0010463557FFA5FE67FE264A670261" box="[413,575,489,517]" pageId="3" pageNumber="588" value="2012-06-14">14th June 2012</date>
|
||
, Bespalaya, Bolotov & Vikhrev leg. (
|
||
<collectionCode id="5DAFAE433557FFA5FC25FE264C7B0261" box="[991,1059,493,517]" pageId="3" pageNumber="588">INEP</collectionCode>
|
||
).
|
||
<emphasis id="09CAEA943557FFA5FBCFFE274CF70261" bold="true" box="[1077,1199,492,517]" pageId="3" pageNumber="588">Primorye:</emphasis>
|
||
<specimenCount id="2DB8FD0F3557FFA5FB4CFE264D1A0261" box="[1206,1346,492,517]" pageId="3" pageNumber="588" type="generic">3 specimens</specimenCount>
|
||
, a small pond near Razdolnaya River,
|
||
<date id="4F0010463557FFA5FE1EFDD84AD00248" box="[484,648,528,556]" pageId="3" pageNumber="588" value="2012-06-25">25th June 2012</date>
|
||
, Bespalaya, Bolotov & Vikhrev leg;
|
||
<specimenCount id="2DB8FD0F3557FFA5FBDEFDDF4CE80248" box="[1060,1200,531,556]" pageId="3" pageNumber="588" type="generic">4 specimens</specimenCount>
|
||
, Ussuri River (Amur River Basin),
|
||
<date id="4F0010463557FFA5FEC5FDF049B70237" box="[319,495,567,595]" pageId="3" pageNumber="588" value="2012-06-20">20th June 2012</date>
|
||
, Bespalaya, Bolotov & Vikhrev leg. (
|
||
<collectionCode id="5DAFAE433557FFA5FC4DFDF04BA30237" box="[951,1019,571,595]" pageId="3" pageNumber="588">INEP</collectionCode>
|
||
). Locality coordinates are given in Appendix 1.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="6FC1660E3557FFA5FF6DFD6A4C9902BC" ID-Table-UUID="6FC1660E3557FFA5FF6DFD6A4C9902BC" httpUri="http://table.plazi.org/id/6FC1660E3557FFA5FF6DFD6A4C9902BC" pageId="3" pageNumber="588" targetBox="[159,1424,755,1475]" targetIsTable="true" targetPageId="3">
|
||
<paragraph id="3B0136863557FFA5FF6DFD6A4C9902BC" blockId="3.[151,1435,672,728]" pageId="3" pageNumber="588">
|
||
<emphasis id="09CAEA943557FFA5FF6DFD6A495102DC" bold="true" box="[151,265,673,696]" pageId="3" pageNumber="588">TABLE 1.</emphasis>
|
||
The comparison of type specimens of endemic
|
||
<taxonomicName id="FCBE4D053557FFA5FCF9FD6A4BE202DC" authority="(Radix)" baseAuthorityName="Radix" box="[771,954,672,696]" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" kingdom="Animalia" order="Hygrophila" pageId="3" pageNumber="588" phylum="Mollusca" rank="genus">
|
||
<emphasis id="09CAEA943557FFA5FCF9FD6A4B3C02DC" box="[771,868,673,696]" italics="true" pageId="3" pageNumber="588">Lymnaea</emphasis>
|
||
(
|
||
<emphasis id="09CAEA943557FFA5FC89FD6B4BE902DC" box="[883,945,672,696]" italics="true" pageId="3" pageNumber="588">Radix</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
species from Khodutka geothermal area with
|
||
<taxonomicName id="FCBE4D053557FFA5FF6DFD0A497602BC" box="[151,302,704,728]" class="Gastropoda" family="Lymnaeidae" genus="Radix" kingdom="Animalia" order="Hygrophila" pageId="3" pageNumber="588" phylum="Mollusca" rank="species" species="auricularia">
|
||
<emphasis id="09CAEA943557FFA5FF6DFD0A497602BC" box="[151,302,704,728]" italics="true" pageId="3" pageNumber="588">R. auricularia</emphasis>
|
||
</taxonomicName>
|
||
based on morphological and anatomical distinguishing features (Glöer & Pešić 2008)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3B0136863557FFA5FF65FD384D300503" pageId="3" pageNumber="588">
|
||
<table id="49BEC42635570059FF65FD384DC805A7" box="[159,1424,755,1475]" gridcols="8" gridrows="6" pageId="3" pageNumber="588">
|
||
<tr id="858E34C435570059FF65FD384DC803E0" box="[159,1424,755,900]" gridrow="0" pageId="3" pageNumber="588">
|
||
<th id="C65F5DB835570059FF65FD38493403E0" box="[159,364,755,900]" gridcol="0" gridrow="0" pageId="3" pageNumber="588">Species</th>
|
||
<th id="C65F5DB835570059FE7BFD384A2E03E0" box="[385,630,755,900]" gridcol="1" gridrow="0" pageId="3" pageNumber="588">Region and reference</th>
|
||
<th id="C65F5DB835570059FD70FD384AAF03E0" box="[650,759,755,900]" gridcol="2" gridrow="0" pageId="3" pageNumber="588">Aperture ear-shaped</th>
|
||
<th id="C65F5DB835570059FCF4FD384BD803E0" box="[782,896,755,900]" gridcol="3" gridrow="0" pageId="3" pageNumber="588">Upper margin of aperture (frontal view)</th>
|
||
<th id="C65F5DB835570059FC6EFD384BAF03E0" box="[916,1015,755,900]" gridcol="4" gridrow="0" pageId="3" pageNumber="588">Upper margin of aperture (apical view)</th>
|
||
<th id="C65F5DB835570059FBE0FD384C2403E0" box="[1050,1148,755,900]" gridcol="5" gridrow="0" pageId="3" pageNumber="588">Juveniles look like the adults</th>
|
||
<th id="C65F5DB835570059FB69FD384D5203E0" box="[1171,1290,755,900]" gridcol="6" gridrow="0" pageId="3" pageNumber="588">Bursa duct</th>
|
||
<th id="C65F5DB835570059FAE4FD384DC803E0" box="[1310,1424,755,900]" gridcol="7" gridrow="0" pageId="3" pageNumber="588">“Freckles” on foot and preputium</th>
|
||
</tr>
|
||
<tr id="858E34C435570059FF65FC5C4DC803A8" box="[159,1424,919,972]" gridrow="1" pageId="3" pageNumber="588">
|
||
<th id="C65F5DB835570059FF65FC5C493403A8" box="[159,364,919,972]" gridcol="0" gridrow="1" pageId="3" pageNumber="588">
|
||
<taxonomicName id="FCBE4D053557FFA5FF65FC53497703C9" box="[159,303,920,941]" class="Gastropoda" family="Lymnaeidae" genus="Radix" kingdom="Animalia" order="Hygrophila" pageId="3" pageNumber="588" phylum="Mollusca" rank="species" species="auricularia">
|
||
<emphasis id="09CAEA943557FFA5FF65FC53497703C9" box="[159,303,920,941]" italics="true" pageId="3" pageNumber="588">R. auricularia</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="C65F5DB835570059FE7BFC5C4A2E03A8" box="[385,630,919,972]" gridcol="1" gridrow="1" pageId="3" pageNumber="588">Southern Europe (Glöer & Pešić 2008)</td>
|
||
<td id="C65F5DB835570059FD70FC5C4AAF03A8" box="[650,759,919,972]" gridcol="2" gridrow="1" pageId="3" pageNumber="588">yes</td>
|
||
<td id="C65F5DB835570059FCF4FC5C4BD803A8" box="[782,896,919,972]" gridcol="3" gridrow="1" pageId="3" pageNumber="588">90º or>90º</td>
|
||
<td id="C65F5DB835570059FC6EFC5C4BAF03A8" box="[916,1015,919,972]" gridcol="4" gridrow="1" pageId="3" pageNumber="588">straight</td>
|
||
<td id="C65F5DB835570059FBE0FC5C4C2403A8" box="[1050,1148,919,972]" gridcol="5" gridrow="1" pageId="3" pageNumber="588">no</td>
|
||
<td id="C65F5DB835570059FB69FC5C4D5203A8" box="[1171,1290,919,972]" gridcol="6" gridrow="1" pageId="3" pageNumber="588">very long</td>
|
||
<td id="C65F5DB835570059FAE4FC5C4DC803A8" box="[1310,1424,919,972]" gridcol="7" gridrow="1" pageId="3" pageNumber="588">yes</td>
|
||
</tr>
|
||
<tr id="858E34C435570059FF65FC144DC80414" box="[159,1424,991,1136]" gridrow="2" pageId="3" pageNumber="588">
|
||
<th id="C65F5DB835570059FF65FC1449340414" box="[159,364,991,1136]" gridcol="0" gridrow="2" pageId="3" pageNumber="588">
|
||
<taxonomicName id="FCBE4D053557FFA5FF65FC2B49770391" box="[159,303,992,1013]" class="Gastropoda" family="Lymnaeidae" genus="Radix" kingdom="Animalia" order="Hygrophila" pageId="3" pageNumber="588" phylum="Mollusca" rank="species" species="auricularia">
|
||
<emphasis id="09CAEA943557FFA5FF65FC2B49770391" box="[159,303,992,1013]" italics="true" pageId="3" pageNumber="588">R. auricularia</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="C65F5DB835570059FE7BFC144A2E0414" box="[385,630,991,1136]" gridcol="1" gridrow="2" pageId="3" pageNumber="588">Russian Far East, excluding Khodutka area (our data, list of localities see. in “Material examined”)</td>
|
||
<td id="C65F5DB835570059FD70FC144AAF0414" box="[650,759,991,1136]" gridcol="2" gridrow="2" pageId="3" pageNumber="588">yes</td>
|
||
<td id="C65F5DB835570059FCF4FC144BD80414" box="[782,896,991,1136]" gridcol="3" gridrow="2" pageId="3" pageNumber="588">90º or>90º</td>
|
||
<td id="C65F5DB835570059FC6EFC144BAF0414" box="[916,1015,991,1136]" gridcol="4" gridrow="2" pageId="3" pageNumber="588">straight</td>
|
||
<td id="C65F5DB835570059FBE0FC144C240414" box="[1050,1148,991,1136]" gridcol="5" gridrow="2" pageId="3" pageNumber="588">no</td>
|
||
<td id="C65F5DB835570059FB69FC144D520414" box="[1171,1290,991,1136]" gridcol="6" gridrow="2" pageId="3" pageNumber="588">very long</td>
|
||
<td id="C65F5DB835570059FAE4FC144DC80414" box="[1310,1424,991,1136]" gridcol="7" gridrow="2" pageId="3" pageNumber="588">yes</td>
|
||
</tr>
|
||
<tr id="858E34C435570059FF65FB4F4DC804DC" box="[159,1424,1156,1208]" gridrow="3" pageId="3" pageNumber="588">
|
||
<th id="C65F5DB835570059FF65FB4F493404DC" box="[159,364,1156,1208]" gridcol="0" gridrow="3" pageId="3" pageNumber="588">
|
||
<taxonomicName id="FCBE4D053557FFA5FF65FB4F497704FD" box="[159,303,1156,1177]" class="Gastropoda" family="Lymnaeidae" genus="Radix" kingdom="Animalia" order="Hygrophila" pageId="3" pageNumber="588" phylum="Mollusca" rank="species" species="auricularia">
|
||
<emphasis id="09CAEA943557FFA5FF65FB4F497704FD" box="[159,303,1156,1177]" italics="true" pageId="3" pageNumber="588">R. auricularia</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="C65F5DB835570059FE7BFB4F4A2E04DC" box="[385,630,1156,1208]" gridcol="1" gridrow="3" pageId="3" pageNumber="588">Khodutka area, Kamchatka (our data)</td>
|
||
<td id="C65F5DB835570059FD70FB4F4AAF04DC" box="[650,759,1156,1208]" gridcol="2" gridrow="3" pageId="3" pageNumber="588">yes</td>
|
||
<td id="C65F5DB835570059FCF4FB4F4BD804DC" box="[782,896,1156,1208]" gridcol="3" gridrow="3" pageId="3" pageNumber="588">90º or>90º</td>
|
||
<td id="C65F5DB835570059FC6EFB4F4BAF04DC" box="[916,1015,1156,1208]" gridcol="4" gridrow="3" pageId="3" pageNumber="588">straight</td>
|
||
<td id="C65F5DB835570059FBE0FB4F4C2404DC" box="[1050,1148,1156,1208]" gridcol="5" gridrow="3" pageId="3" pageNumber="588">no</td>
|
||
<td id="C65F5DB835570059FB69FB4F4D5204DC" box="[1171,1290,1156,1208]" gridcol="6" gridrow="3" pageId="3" pageNumber="588">very long or long</td>
|
||
<td id="C65F5DB835570059FAE4FB4F4DC804DC" box="[1310,1424,1156,1208]" gridcol="7" gridrow="3" pageId="3" pageNumber="588">yes</td>
|
||
</tr>
|
||
<tr id="858E34C435570059FF65FB074DC80559" box="[159,1424,1228,1341]" gridrow="4" pageId="3" pageNumber="588">
|
||
<th id="C65F5DB835570059FF65FB0749340559" box="[159,364,1228,1341]" gridcol="0" gridrow="4" pageId="3" pageNumber="588">
|
||
Holotype of
|
||
<taxonomicName id="FCBE4D053557FFA5FEE5FB0748A4049B" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" kingdom="Animalia" order="Hygrophila" pageId="3" pageNumber="588" phylum="Mollusca" rank="species" species="hadutkae" subGenus="Radix">
|
||
<emphasis id="09CAEA943557FFA5FEE5FB0748A4049B" italics="true" pageId="3" pageNumber="588">L. (R.) hadutkae</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="C65F5DB835570059FE7BFB074A2E0559" box="[385,630,1228,1341]" gridcol="1" gridrow="4" pageId="3" pageNumber="588">Khodutka area, Kamchatka (Kruglov & Starobogatov 1989; Kruglov 2005; our data)</td>
|
||
<td id="C65F5DB835570059FD70FB074AAF0559" box="[650,759,1228,1341]" gridcol="2" gridrow="4" pageId="3" pageNumber="588">yes</td>
|
||
<td id="C65F5DB835570059FCF4FB074BD80559" box="[782,896,1228,1341]" gridcol="3" gridrow="4" pageId="3" pageNumber="588">90º</td>
|
||
<td id="C65F5DB835570059FC6EFB074BAF0559" box="[916,1015,1228,1341]" gridcol="4" gridrow="4" pageId="3" pageNumber="588">straight</td>
|
||
<td id="C65F5DB835570059FBE0FB074C240559" box="[1050,1148,1228,1341]" gridcol="5" gridrow="4" pageId="3" pageNumber="588">--</td>
|
||
<td id="C65F5DB835570059FB69FB074D520559" box="[1171,1290,1228,1341]" gridcol="6" gridrow="4" pageId="3" pageNumber="588">very long</td>
|
||
<td id="C65F5DB835570059FAE4FB074DC80559" box="[1310,1424,1228,1341]" gridcol="7" gridrow="4" pageId="3" pageNumber="588">no data</td>
|
||
</tr>
|
||
<tr id="858E34C435570059FF65FA9A4DC805A7" box="[159,1424,1361,1475]" gridrow="5" pageId="3" pageNumber="588">
|
||
<th id="C65F5DB835570059FF65FA9A493405A7" box="[159,364,1361,1475]" gridcol="0" gridrow="5" pageId="3" pageNumber="588">
|
||
Holotype of
|
||
<taxonomicName id="FCBE4D053557FFA5FEE5FA99493405E1" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" kingdom="Animalia" order="Hygrophila" pageId="3" pageNumber="588" phylum="Mollusca" rank="species" species="thermokamtschatica" subGenus="Radix">
|
||
<emphasis id="09CAEA943557FFA5FEE5FA99493405E1" italics="true" pageId="3" pageNumber="588">L. (R.) thermokamtschatica</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="C65F5DB835570059FE7BFA9A4A2E05A7" box="[385,630,1361,1475]" gridcol="1" gridrow="5" pageId="3" pageNumber="588">Khodutka area, Kamchatka (Kruglov & Starobogatov 1989; Kruglov 2005; our data)</td>
|
||
<td id="C65F5DB835570059FD70FA9A4AAF05A7" box="[650,759,1361,1475]" gridcol="2" gridrow="5" pageId="3" pageNumber="588">yes</td>
|
||
<td id="C65F5DB835570059FCF4FA9A4BD805A7" box="[782,896,1361,1475]" gridcol="3" gridrow="5" pageId="3" pageNumber="588">>90º</td>
|
||
<td id="C65F5DB835570059FC6EFA9A4BAF05A7" box="[916,1015,1361,1475]" gridcol="4" gridrow="5" pageId="3" pageNumber="588">straight</td>
|
||
<td id="C65F5DB835570059FBE0FA9A4C2405A7" box="[1050,1148,1361,1475]" gridcol="5" gridrow="5" pageId="3" pageNumber="588">--</td>
|
||
<td id="C65F5DB835570059FB69FA9A4D5205A7" box="[1171,1290,1361,1475]" gridcol="6" gridrow="5" pageId="3" pageNumber="588">long</td>
|
||
<td id="C65F5DB835570059FAE4FA9A4DC805A7" box="[1310,1424,1361,1475]" gridcol="7" gridrow="5" pageId="3" pageNumber="588">no data</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<subSubSection id="73A4650D3557FFA5FF3DF9C94CD1073A" pageId="3" pageNumber="588" type="description">
|
||
<paragraph id="3B0136863557FFA5FF3DF9C94CD1073A" blockId="3.[151,1437,1538,2023]" pageId="3" pageNumber="588">
|
||
<emphasis id="09CAEA943557FFA5FF3DF9C94901067F" bold="true" box="[199,345,1538,1563]" pageId="3" pageNumber="588">Description.</emphasis>
|
||
Specimens from the Khodutka geothermal area are very small with shell height 7.0–
|
||
<quantity id="FC469B633557FFA5FACAF9C94DC0067E" box="[1328,1432,1538,1563]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="3" pageNumber="588" unit="mm" value="14.6">14.6 mm</quantity>
|
||
, width 4.0–
|
||
<quantity id="FC469B633557FFA5FEEAF9ED4930065A" box="[272,360,1574,1599]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="3" pageNumber="588" unit="mm" value="9.5">9.5 mm</quantity>
|
||
. Shell is glossy, dark-yellow or light brownish, elongated, with 4 whorls enlarging slowly (
|
||
<figureCitation id="A3852A033557FFA5FA9EF9ED489B0607" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1692,1715]" captionTargetBox="[230,1360,184,1674]" captionTargetId="figure@2.[223,1367,178,1687]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Radix auricularia (Linnaeus, 1758) from Russian Far East. Figs. 1 - 4. Shell and proximal part of the female genitalia. Scale bar 2 mm. Fig. 1. Holotype of Lymnaea (R.) hadutkae Kruglov & Starobogatov, 1989 syn. n. (shell photo: M. V. Vinarski; genitalia redrawn from the protologue, Fig. 2.7). Fig. 2. Holotype of L. (R.) thermokamtschatica Kruglov & Starobogatov, 1989 syn. n. (shell photo: M. V. Vinarski; genitalia redrawn from the protologue, Fig. 2.9). Fig. 3. Newly collected specimen from the Khodutka geothermal area, Kamchatka (photos: O. V. Aksenova). Fig. 4. Specimen from Malkinskie hot springs, Kamchatka (photos: O. V. Aksenova). Fig. 5. Bayesian phylogram of haplotypes based on mitochondrial COI gene dataset (Appendixes 1 & 2). The scale bar indicates the branch length. Asterisks: Posterior probabilities ≥ 0.95; other significant node support values are mentioned in the figure. Haplotypes exclusively from NCBI’s Genbank are marked with circumflex accents (˄). The well-supported clade with two haplotypes from the Khodutka geothermal area is highlighted in blue. Fig. 6. General view of the Khodutka warm lake. Fig. 7. Map of observed localities on the Russian Far East. The red circles represent species records. Locality codes are given in Appendix 1. The digital elevation model and other layers of the map were added from Esri Data & Maps 10 dataset (Map: M. Yu. Gofarov)." httpUri="https://zenodo.org/record/225342/files/figure.png" pageId="3" pageNumber="588">Figs. 1–3</figureCitation>
|
||
). The spire is long, widely conical, in adult shells often destroyed on the top (likely due to the geothermal water influence). The aperture is ear-shaped, relatively narrow with an angle of 90° or>90° on the top. The inner lip is folded at the columella, with weakly-sinuated fold. The foot and dorsal part of the preputium bear “freckles”. The bursa copulatrix is oval or rounded; the bursa duct is thin and very long or long (
|
||
<figureCitation id="A3852A033557FFA5FB71F97D4CA006AB" box="[1163,1272,1718,1743]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1692,1715]" captionTargetBox="[230,1360,184,1674]" captionTargetId="figure@2.[223,1367,178,1687]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Radix auricularia (Linnaeus, 1758) from Russian Far East. Figs. 1 - 4. Shell and proximal part of the female genitalia. Scale bar 2 mm. Fig. 1. Holotype of Lymnaea (R.) hadutkae Kruglov & Starobogatov, 1989 syn. n. (shell photo: M. V. Vinarski; genitalia redrawn from the protologue, Fig. 2.7). Fig. 2. Holotype of L. (R.) thermokamtschatica Kruglov & Starobogatov, 1989 syn. n. (shell photo: M. V. Vinarski; genitalia redrawn from the protologue, Fig. 2.9). Fig. 3. Newly collected specimen from the Khodutka geothermal area, Kamchatka (photos: O. V. Aksenova). Fig. 4. Specimen from Malkinskie hot springs, Kamchatka (photos: O. V. Aksenova). Fig. 5. Bayesian phylogram of haplotypes based on mitochondrial COI gene dataset (Appendixes 1 & 2). The scale bar indicates the branch length. Asterisks: Posterior probabilities ≥ 0.95; other significant node support values are mentioned in the figure. Haplotypes exclusively from NCBI’s Genbank are marked with circumflex accents (˄). The well-supported clade with two haplotypes from the Khodutka geothermal area is highlighted in blue. Fig. 6. General view of the Khodutka warm lake. Fig. 7. Map of observed localities on the Russian Far East. The red circles represent species records. Locality codes are given in Appendix 1. The digital elevation model and other layers of the map were added from Esri Data & Maps 10 dataset (Map: M. Yu. Gofarov)." httpUri="https://zenodo.org/record/225342/files/figure.png" pageId="3" pageNumber="588">Figs. 1–3</figureCitation>
|
||
and
|
||
<tableCitation id="763C033D3557FFA5FACCF97D4DD306AB" box="[1334,1419,1718,1743]" captionStart="TABLE 1" captionStartId="3.[151,239,673,696]" captionTargetBox="[159,1424,755,1475]" captionTargetPageId="3" captionText="TABLE 1. The comparison of type specimens of endemic Lymnaea (Radix) species from Khodutka geothermal area with R. auricularia based on morphological and anatomical distinguishing features (Glöer & Pešić 2008)" httpUri="http://table.plazi.org/id/6FC1660E3557FFA5FF6DFD6A4C9902BC" pageId="3" pageNumber="588" tableUuid="6FC1660E3557FFA5FF6DFD6A4C9902BC">Table 1</tableCitation>
|
||
). According to morphological and anatomical data, specimens from other localities of the Russian Far East are similar to those found from the Khodutka area (
|
||
<figureCitation id="A3852A033557FFA5FD22F9354B7B0773" box="[728,803,1790,1815]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1692,1715]" captionTargetBox="[230,1360,184,1674]" captionTargetId="figure@2.[223,1367,178,1687]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Radix auricularia (Linnaeus, 1758) from Russian Far East. Figs. 1 - 4. Shell and proximal part of the female genitalia. Scale bar 2 mm. Fig. 1. Holotype of Lymnaea (R.) hadutkae Kruglov & Starobogatov, 1989 syn. n. (shell photo: M. V. Vinarski; genitalia redrawn from the protologue, Fig. 2.7). Fig. 2. Holotype of L. (R.) thermokamtschatica Kruglov & Starobogatov, 1989 syn. n. (shell photo: M. V. Vinarski; genitalia redrawn from the protologue, Fig. 2.9). Fig. 3. Newly collected specimen from the Khodutka geothermal area, Kamchatka (photos: O. V. Aksenova). Fig. 4. Specimen from Malkinskie hot springs, Kamchatka (photos: O. V. Aksenova). Fig. 5. Bayesian phylogram of haplotypes based on mitochondrial COI gene dataset (Appendixes 1 & 2). The scale bar indicates the branch length. Asterisks: Posterior probabilities ≥ 0.95; other significant node support values are mentioned in the figure. Haplotypes exclusively from NCBI’s Genbank are marked with circumflex accents (˄). The well-supported clade with two haplotypes from the Khodutka geothermal area is highlighted in blue. Fig. 6. General view of the Khodutka warm lake. Fig. 7. Map of observed localities on the Russian Far East. The red circles represent species records. Locality codes are given in Appendix 1. The digital elevation model and other layers of the map were added from Esri Data & Maps 10 dataset (Map: M. Yu. Gofarov)." httpUri="https://zenodo.org/record/225342/files/figure.png" pageId="3" pageNumber="588">Fig. 4</figureCitation>
|
||
). However, specimens from non-geothermal localities (Plotnikova River, Aliger Lake, a pond near Razdolnaya River and Ussuri River) are larger with shell height up to
|
||
<quantity id="FC469B633557FFA5FF6DF88D48A5073A" box="[151,253,1862,1887]" metricMagnitude="-2" metricUnit="m" metricValue="2.57" pageId="3" pageNumber="588" unit="mm" value="25.7">25.7 mm</quantity>
|
||
, width up to
|
||
<quantity id="FC469B633557FFA5FE75F88C49AD073A" box="[399,501,1863,1887]" metricMagnitude="-2" metricUnit="m" metricValue="2.09" pageId="3" pageNumber="588" unit="mm" value="20.9">20.9 mm</quantity>
|
||
, and had a more compact shell shape with a wide aperture.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="73A4650D3557FFA2FF3DF8A14BE601B9" lastPageId="4" lastPageNumber="589" pageId="3" pageNumber="588" type="discussion">
|
||
<paragraph id="3B0136863557FFA2FF3DF8A14BE601B9" blockId="3.[151,1437,1538,2023]" lastBlockId="4.[151,1436,151,477]" lastPageId="4" lastPageNumber="589" pageId="3" pageNumber="588">
|
||
<emphasis id="09CAEA943557FFA5FF3DF8A1496307E7" bold="true" box="[199,315,1898,1923]" pageId="3" pageNumber="588">Remarks.</emphasis>
|
||
Both of the endemics from the Khodutka geothermal area (
|
||
<figureCitation id="A3852A033557FFA5FC2DF8A14C4207E7" box="[983,1050,1898,1923]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1692,1715]" captionTargetBox="[230,1360,184,1674]" captionTargetId="figure@2.[223,1367,178,1687]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Radix auricularia (Linnaeus, 1758) from Russian Far East. Figs. 1 - 4. Shell and proximal part of the female genitalia. Scale bar 2 mm. Fig. 1. Holotype of Lymnaea (R.) hadutkae Kruglov & Starobogatov, 1989 syn. n. (shell photo: M. V. Vinarski; genitalia redrawn from the protologue, Fig. 2.7). Fig. 2. Holotype of L. (R.) thermokamtschatica Kruglov & Starobogatov, 1989 syn. n. (shell photo: M. V. Vinarski; genitalia redrawn from the protologue, Fig. 2.9). Fig. 3. Newly collected specimen from the Khodutka geothermal area, Kamchatka (photos: O. V. Aksenova). Fig. 4. Specimen from Malkinskie hot springs, Kamchatka (photos: O. V. Aksenova). Fig. 5. Bayesian phylogram of haplotypes based on mitochondrial COI gene dataset (Appendixes 1 & 2). The scale bar indicates the branch length. Asterisks: Posterior probabilities ≥ 0.95; other significant node support values are mentioned in the figure. Haplotypes exclusively from NCBI’s Genbank are marked with circumflex accents (˄). The well-supported clade with two haplotypes from the Khodutka geothermal area is highlighted in blue. Fig. 6. General view of the Khodutka warm lake. Fig. 7. Map of observed localities on the Russian Far East. The red circles represent species records. Locality codes are given in Appendix 1. The digital elevation model and other layers of the map were added from Esri Data & Maps 10 dataset (Map: M. Yu. Gofarov)." httpUri="https://zenodo.org/record/225342/files/figure.png" pageId="3" pageNumber="588">Fig. 6</figureCitation>
|
||
) are conspecific to
|
||
<taxonomicName id="FCBE4D053557FFA5FB0CF8A04DCC07E7" box="[1270,1428,1898,1923]" class="Gastropoda" family="Lymnaeidae" genus="Radix" kingdom="Animalia" order="Hygrophila" pageId="3" pageNumber="588" phylum="Mollusca" rank="species" species="auricularia">
|
||
<emphasis id="09CAEA943557FFA5FB0CF8A04DCC07E7" box="[1270,1428,1898,1923]" italics="true" pageId="3" pageNumber="588">R. auricularia</emphasis>
|
||
</taxonomicName>
|
||
, based on morphological, anatomical and molecular studies (
|
||
<tableCitation id="763C033D3557FFA5FCCEF8404BD107C0" box="[820,905,1931,1956]" captionStart="TABLE 1" captionStartId="3.[151,239,673,696]" captionTargetBox="[159,1424,755,1475]" captionTargetPageId="3" captionText="TABLE 1. The comparison of type specimens of endemic Lymnaea (Radix) species from Khodutka geothermal area with R. auricularia based on morphological and anatomical distinguishing features (Glöer & Pešić 2008)" httpUri="http://table.plazi.org/id/6FC1660E3557FFA5FF6DFD6A4C9902BC" pageId="3" pageNumber="588" tableUuid="6FC1660E3557FFA5FF6DFD6A4C9902BC">Table 1</tableCitation>
|
||
and
|
||
<figureCitation id="A3852A033557FFA5FC45F8404C7107C0" box="[959,1065,1931,1956]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1692,1715]" captionTargetBox="[230,1360,184,1674]" captionTargetId="figure@2.[223,1367,178,1687]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Radix auricularia (Linnaeus, 1758) from Russian Far East. Figs. 1 - 4. Shell and proximal part of the female genitalia. Scale bar 2 mm. Fig. 1. Holotype of Lymnaea (R.) hadutkae Kruglov & Starobogatov, 1989 syn. n. (shell photo: M. V. Vinarski; genitalia redrawn from the protologue, Fig. 2.7). Fig. 2. Holotype of L. (R.) thermokamtschatica Kruglov & Starobogatov, 1989 syn. n. (shell photo: M. V. Vinarski; genitalia redrawn from the protologue, Fig. 2.9). Fig. 3. Newly collected specimen from the Khodutka geothermal area, Kamchatka (photos: O. V. Aksenova). Fig. 4. Specimen from Malkinskie hot springs, Kamchatka (photos: O. V. Aksenova). Fig. 5. Bayesian phylogram of haplotypes based on mitochondrial COI gene dataset (Appendixes 1 & 2). The scale bar indicates the branch length. Asterisks: Posterior probabilities ≥ 0.95; other significant node support values are mentioned in the figure. Haplotypes exclusively from NCBI’s Genbank are marked with circumflex accents (˄). The well-supported clade with two haplotypes from the Khodutka geothermal area is highlighted in blue. Fig. 6. General view of the Khodutka warm lake. Fig. 7. Map of observed localities on the Russian Far East. The red circles represent species records. Locality codes are given in Appendix 1. The digital elevation model and other layers of the map were added from Esri Data & Maps 10 dataset (Map: M. Yu. Gofarov)." httpUri="https://zenodo.org/record/225342/files/figure.png" pageId="3" pageNumber="588">Figs. 1–5</figureCitation>
|
||
). The same elongated shells with narrow apertures were reported for several
|
||
<taxonomicName id="FCBE4D053557FFA5FD80F8654B4107A2" box="[634,793,1965,1990]" class="Gastropoda" family="Lymnaeidae" genus="Radix" kingdom="Animalia" order="Hygrophila" pageId="3" pageNumber="588" phylum="Mollusca" rank="species" species="auricularia">
|
||
<emphasis id="09CAEA943557FFA5FD80F8654B4107A2" box="[634,793,1965,1990]" italics="true" pageId="3" pageNumber="588">R. auricularia</emphasis>
|
||
</taxonomicName>
|
||
populations from the Baikal Lake area (
|
||
<bibRefCitation id="5F2F4B773557FFA5FB26F8664DC907A1" author="Stift" box="[1244,1425,1965,1990]" pageId="3" pageNumber="588" refString="Stift, M., Michel, E., Sitnikova, T. Y., Mamonova, E. Y. & Sherbakov, D. Y. (2004) Palaearctic gastropod gains a foothold in the dominion of endemics: range expansion and morphological change of Lymnaea (Radix) auricularia in Lake Baikal. Hydrobiologia, 513 (1 - 3), 101 - 108. http: // dx. doi. org / 10.1023 / b: hydr. 0000018175.37771. d 6" type="journal article" year="2004">
|
||
Stift
|
||
<emphasis id="09CAEA943557FFA5FAEEF8654D1507A2" box="[1300,1357,1965,1990]" italics="true" pageId="3" pageNumber="588">et al.</emphasis>
|
||
2004
|
||
</bibRefCitation>
|
||
). According to our data,
|
||
<taxonomicName id="FCBE4D053557FFA5FE4EF8044A020783" box="[436,602,1998,2023]" class="Gastropoda" family="Lymnaeidae" genus="Radix" kingdom="Animalia" order="Hygrophila" pageId="3" pageNumber="588" phylum="Mollusca" rank="species" species="auricularia">
|
||
<emphasis id="09CAEA943557FFA5FE4EF8044A020783" box="[436,602,1998,2023]" italics="true" pageId="3" pageNumber="588">R. auricularia</emphasis>
|
||
</taxonomicName>
|
||
is widespread on the Russian Far East, where it was recorded from Kamchatka, Primorye, Sakhalin, and Kuriles (
|
||
<figureCitation id="A3852A033550FFA2FD61FF5C4A8700D4" box="[667,735,151,176]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1692,1715]" captionTargetBox="[230,1360,184,1674]" captionTargetId="figure@2.[223,1367,178,1687]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Radix auricularia (Linnaeus, 1758) from Russian Far East. Figs. 1 - 4. Shell and proximal part of the female genitalia. Scale bar 2 mm. Fig. 1. Holotype of Lymnaea (R.) hadutkae Kruglov & Starobogatov, 1989 syn. n. (shell photo: M. V. Vinarski; genitalia redrawn from the protologue, Fig. 2.7). Fig. 2. Holotype of L. (R.) thermokamtschatica Kruglov & Starobogatov, 1989 syn. n. (shell photo: M. V. Vinarski; genitalia redrawn from the protologue, Fig. 2.9). Fig. 3. Newly collected specimen from the Khodutka geothermal area, Kamchatka (photos: O. V. Aksenova). Fig. 4. Specimen from Malkinskie hot springs, Kamchatka (photos: O. V. Aksenova). Fig. 5. Bayesian phylogram of haplotypes based on mitochondrial COI gene dataset (Appendixes 1 & 2). The scale bar indicates the branch length. Asterisks: Posterior probabilities ≥ 0.95; other significant node support values are mentioned in the figure. Haplotypes exclusively from NCBI’s Genbank are marked with circumflex accents (˄). The well-supported clade with two haplotypes from the Khodutka geothermal area is highlighted in blue. Fig. 6. General view of the Khodutka warm lake. Fig. 7. Map of observed localities on the Russian Far East. The red circles represent species records. Locality codes are given in Appendix 1. The digital elevation model and other layers of the map were added from Esri Data & Maps 10 dataset (Map: M. Yu. Gofarov)." httpUri="https://zenodo.org/record/225342/files/figure.png" pageId="4" pageNumber="589">Fig. 7</figureCitation>
|
||
).
|
||
<bibRefCitation id="5F2F4B773550FFA2FD0EFF5C4BCD00D4" author="Zhadin" box="[756,917,151,176]" pageId="4" pageNumber="589" refString="Zhadin, V. I. (1952) Molluscs of fresh and brackish water of the USSR. Identification Guides on the Fauna of the USSR, 46, 1 - 374. [in Russian]" type="book chapter" year="1952">Zhadin (1952)</bibRefCitation>
|
||
reported that this species is widespread on the Pacific Basin, including Kamchatka, Amur Basin and Primorye, which is in agreement with our records. However,
|
||
<bibRefCitation id="5F2F4B773550FFA2FF6DFF114A5C0097" author="Kruglov" box="[151,516,218,243]" pageId="4" pageNumber="589" refString="Kruglov, N. D. & Starobogatov, Ya. I. (1993) Guide to recent molluscs of northern Eurasia. 3. Annotated and illustrated catalogue of species of the family Lymnaeidae (Gastropoda Pulmonata Lymnaeiformes) of Palaearctic and adjacent river drainage areas. Part 1. Ruthenica, 3 (1), 65 - 92." type="journal article" year="1993">Kruglov & Starobogatov (1993)</bibRefCitation>
|
||
noted that
|
||
<taxonomicName id="FCBE4D053550FFA2FD72FF174B700097" box="[648,808,218,243]" class="Gastropoda" family="Lymnaeidae" genus="Radix" kingdom="Animalia" order="Hygrophila" pageId="4" pageNumber="589" phylum="Mollusca" rank="species" species="auricularia">
|
||
<emphasis id="09CAEA943550FFA2FD72FF174B700097" box="[648,808,218,243]" italics="true" pageId="4" pageNumber="589">R. auricularia</emphasis>
|
||
</taxonomicName>
|
||
ranged in Europe, the western part of the Middle East and Siberia, but not mentioned in the species records from the Far East. Thereafter, Starobogatov
|
||
<emphasis id="09CAEA943550FFA2FAF6FF364D1F0171" box="[1292,1351,252,277]" italics="true" pageId="4" pageNumber="589">et al.</emphasis>
|
||
(2004) again reported that species ranged on the Far East, where it was recorded in rivers of the Pacific coast from Kamchatka to Primorye. Meanwhile, Kruglov & Starobogatov (1989, 1993), Starobogatov
|
||
<emphasis id="09CAEA943550FFA2FB2CFEF44D480133" box="[1238,1296,318,343]" italics="true" pageId="4" pageNumber="589">et al.</emphasis>
|
||
(2004) and
|
||
<bibRefCitation id="5F2F4B773550FFA2FF6DFE94491F011C" author="Kruglov" box="[151,327,351,376]" pageId="4" pageNumber="589" refString="Kruglov, N. D. (2005) Molluscs of the family Lymnaeidae (Gastropoda: Pulmonata) in Europe and Northern Asia. SGPU Publishing, Smolensk, 507 pp. [in Russian]" type="book" year="2005">Kruglov (2005)</bibRefCitation>
|
||
listed many local endemic
|
||
<taxonomicName id="FCBE4D053550FFA2FD86FE944AE6011C" box="[636,702,351,376]" class="Gastropoda" family="Lymnaeidae" genus="Radix" kingdom="Animalia" order="Hygrophila" pageId="4" pageNumber="589" phylum="Mollusca" rank="genus">
|
||
<emphasis id="09CAEA943550FFA2FD86FE944AE6011C" box="[636,702,351,376]" italics="true" pageId="4" pageNumber="589">Radix</emphasis>
|
||
</taxonomicName>
|
||
species for different areas of the Russian Far East, which belong to the sections
|
||
<taxonomicName id="FCBE4D053550FFA2FEB7FE4A4B1401FD" authority="Kruglov & Starobogatov, 1989" authorityName="Kruglov & Starobogatov" authorityYear="1989" box="[333,844,385,410]" class="Gastropoda" family="Lymnaeidae" genus="Ussuriradix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Basommatophora" pageId="4" pageNumber="589" phylum="Mollusca" rank="genus">
|
||
<emphasis id="09CAEA943550FFA2FEB7FE4A498B01FE" box="[333,467,385,410]" italics="true" pageId="4" pageNumber="589">Ussuriradix</emphasis>
|
||
<bibRefCitation id="5F2F4B773550FFA2FE25FE4A4B1401FD" author="Kruglov" box="[479,844,385,410]" pageId="4" pageNumber="589" refString="Kruglov, N. D. & Starobogatov, Ya. I. (1989) Morphology and taxonomy of the molluscs from the subgenus Radix of the genus Lymnaea (Gastropoda, Pulmonata, Lymnaeidae) from Siberia and Far East of the USSR. Zoologicheskii Zhurnal, 68 (5), 17 - 30. [in Russian]" type="journal article" year="1989">Kruglov & Starobogatov, 1989</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="FCBE4D053550FFA2FCA6FE4A4D3F01FD" authority="Kruglov & Starobogatov, 1989" authorityName="Kruglov & Starobogatov" authorityYear="1989" box="[860,1383,385,410]" class="Gastropoda" family="Lymnaeidae" genus="Nipponiradix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Basommatophora" pageId="4" pageNumber="589" phylum="Mollusca" rank="genus">
|
||
<emphasis id="09CAEA943550FFA2FCA6FE4A4BA901FE" box="[860,1009,385,410]" italics="true" pageId="4" pageNumber="589">Nipponiradix</emphasis>
|
||
<bibRefCitation id="5F2F4B773550FFA2FC04FE4A4D3F01FD" author="Kruglov" box="[1022,1383,385,410]" pageId="4" pageNumber="589" refString="Kruglov, N. D. & Starobogatov, Ya. I. (1989) Morphology and taxonomy of the molluscs from the subgenus Radix of the genus Lymnaea (Gastropoda, Pulmonata, Lymnaeidae) from Siberia and Far East of the USSR. Zoologicheskii Zhurnal, 68 (5), 17 - 30. [in Russian]" type="journal article" year="1989">Kruglov & Starobogatov, 1989</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="FCBE4D053550FFA2FF6DFE694ACA01DF" authority="Kruglov & Starobogatov, 1989" authorityName="Kruglov & Starobogatov" authorityYear="1989" box="[151,658,418,443]" class="Gastropoda" family="Lymnaeidae" genus="Okhotiradix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Basommatophora" pageId="4" pageNumber="589" phylum="Mollusca" rank="genus">
|
||
<emphasis id="09CAEA943550FFA2FF6DFE69494601DF" box="[151,286,418,443]" italics="true" pageId="4" pageNumber="589">Okhotiradix</emphasis>
|
||
<bibRefCitation id="5F2F4B773550FFA2FED3FE694ACA01DF" author="Kruglov" box="[297,658,418,443]" pageId="4" pageNumber="589" refString="Kruglov, N. D. & Starobogatov, Ya. I. (1989) Morphology and taxonomy of the molluscs from the subgenus Radix of the genus Lymnaea (Gastropoda, Pulmonata, Lymnaeidae) from Siberia and Far East of the USSR. Zoologicheskii Zhurnal, 68 (5), 17 - 30. [in Russian]" type="journal article" year="1989">Kruglov & Starobogatov, 1989</bibRefCitation>
|
||
</taxonomicName>
|
||
. Taxonomic position of these taxa is uncertain and they need in a current revision based on mitochondrial and nuclear molecular markers.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |