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<document ID-CLB-Dataset="209209" ID-DOI="http://dx.doi.org/10.3897/asp.80.e90570" ID-GBIF-Dataset="bac17de6-baf9-4fee-9ded-268adbb26a61" ID-Pensoft-Pub="1864-8312-80-649" ID-Pensoft-UUID="FAE6DFEF69735531A4CBA275E078320D" ID-ZooBank="AD0B1CC262414460B6B888E6B95F20D7" ModsDocID="1864-8312-80-649" checkinTime="1671114540108" checkinUser="pensoft" docAuthor="Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S. &amp; Harms, Danilo" docDate="2022" docId="91C8C7D43B7A56BDA76F3C11C0E278F2" docLanguage="en" docName="ArthropodSystPhyl 80: 649-691" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 80" docPubDate="2022-12-14" docSource="http://dx.doi.org/10.3897/asp.80.e90570" docTitle="Mahajanganella schwarzeneggeri Lorenz &amp; Loria &amp; Harvey &amp; Harms 2022, sp. nov." docType="treatment" docVersion="5" id="FAE6DFEF69735531A4CBA275E078320D" lastPageNumber="649" masterDocId="FAE6DFEF69735531A4CBA275E078320D" masterDocTitle="The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the &quot; hottest &quot; biodiversity hotspots" masterLastPageNumber="691" masterPageNumber="649" pageNumber="649" updateTime="1732733992417" updateUser="ExternalLinkService">
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<mods:title id="4E6D0E5A82A91424F63631D5AEDBF123">The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the &quot; hottest &quot; biodiversity hotspots</mods:title>
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<mods:namePart id="EFE4E2CACD8644AAAD116EE081431CFF">Lorenz, Michelle</mods:namePart>
<mods:affiliation id="7E667B14489594D8B4F69631C93474E9">Museum of Nature Hamburg - Zoology, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany &amp; University of Hamburg, Biology Department, Ohnhorststrasse 18, 22609 Hamburg, Germany</mods:affiliation>
<mods:nameIdentifier id="33EDB34669AF5C2448F10F97C1CFBBE4" type="email">michellelorenz@gmx.net</mods:nameIdentifier>
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<mods:namePart id="6A880F37D8AAC9A18B7EDE508C04A407">Loria, Stephanie F.</mods:namePart>
<mods:affiliation id="847B96413DD53CF4F58B1A00B6151E3B">Museum of Nature Hamburg - Zoology, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
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<mods:namePart id="F30E72D97F94A152FB06F9DE6C20193E">Harvey, Mark S.</mods:namePart>
<mods:nameIdentifier id="C6430F22D0BAB04B723E1F9355A5DFB4" type="ORCID">https://orcid.org/0000-0003-1482-0109</mods:nameIdentifier>
<mods:affiliation id="E321540C856A16A797751E08D5D47496">Western Australian Museum, Collections &amp; Research, 49 Kew Street, Welshpool, WA 6106, Australia &amp; University of Western Australia, School of Biological Sciences, Crawley WA 6009, Australia</mods:affiliation>
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<mods:namePart id="469F04AFD16335AD9376C57FE2FA5C95">Harms, Danilo</mods:namePart>
<mods:nameIdentifier id="A490535EB89C8E381B675578D4BDDF6F" type="ORCID">https://orcid.org/0000-0002-7189-5345</mods:nameIdentifier>
<mods:affiliation id="3062BAF2E07F56A75D1B3294CA29E26D">Museum of Nature Hamburg - Zoology, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
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<mods:date id="3C55F0A68DC250AD7A099FEFAE48FA5E">2022</mods:date>
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<mods:number id="6E8690A194768CC2E0B520D8566B2770">2022-12-14</mods:number>
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<treatment id="91C8C7D43B7A56BDA76F3C11C0E278F2" ID-GBIF-Taxon="206273796" LSID="urn:lsid:zoobank.org:act:NomEnClAturAlACts/8DD8CBE7-C8F5-46BE-8FBC-2ABDB7030F69" httpUri="http://treatment.plazi.org/id/91C8C7D43B7A56BDA76F3C11C0E278F2" lastPageNumber="649" pageId="0" pageNumber="649" scope_family="Feaellidae" scope_order="Pseudoscorpiones" scope_subFamily="Feaellinae" scope_superFamily="Feaelloidea">
<subSubSection id="D16FD7D394539F29C2FFDC3766E0A024" pageId="0" pageNumber="649" type="nomenclature">
<paragraph id="8CA461B0904DC415258F27E3AE1955D6" pageId="0" pageNumber="649">
<taxonomicName id="695586364D797C12E647E7B532215B4B" LSID="https://zoobank.org/NomenclaturalActs/8DD8CBE7-C8F5-46BE-8FBC-2ABDB7030F69" authority="Lorenz &amp; Loria &amp; Harvey &amp; Harms, 2022" authorityName="Lorenz &amp; Loria &amp; Harvey &amp; Harms" authorityYear="2022" class="Arachnida" family="Feaellidae" genus="Mahajanganella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mahajanganella schwarzeneggeri" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="species" species="schwarzeneggeri" status="sp. nov.">Mahajanganella schwarzeneggeri</taxonomicName>
<taxonomicNameLabel id="C770DEA080265B1EABBD669A80194ABC" pageId="0" pageNumber="649">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="9E5CD313041F73FC0BDD82265905DF70" pageId="0" pageNumber="649" type="type material">
<paragraph id="216343619C8BBC188C0DD210E9723D6C" pageId="0" pageNumber="649">Type material examined.</paragraph>
<paragraph id="811A27CD4210F852DCABF8BAC5B4C8AA" pageId="0" pageNumber="649">
MADAGASCAR: Mahajanga Province [Boeny Region]: Holotype ♂ (CAS 9071739 [BLF6812]), allotype ♀ (ZMH-A0016716), paratypes: 1 ♂ (ZMH-A0016717), 1 ♀ (CAS 9071739 [BLF6812]), Parc National de Baie de Baly;
<geoCoordinate id="9B32F1E75B47BF9738FFCF9F7ED3DC35" degrees="16" direction="south" minutes="00" orientation="latitude" precision="15" seconds="36" value="-16.01">16°0036″S</geoCoordinate>
<geoCoordinate id="AD38C8AFCB3CED3D82158691796149EF" degrees="45" direction="east" minutes="15" orientation="longitude" precision="15" seconds="54" value="45.265">45°1554″E</geoCoordinate>
, 26-30 November 2002, B.L. Fisher, Griswold et al.
</paragraph>
</subSubSection>
<subSubSection id="F36976C0F0997757BB74E81A062E9D3D" pageId="0" pageNumber="649" type="diagnosis">
<paragraph id="4A55FF5368DE740EF4AD514D37C27198" pageId="0" pageNumber="649">Diagnosis.</paragraph>
<paragraph id="51935698187090002E2319FB8726DB45" pageId="0" pageNumber="649">
Very similar to
<taxonomicName id="88CDB07927C8EBBAD797BC23A8F8C7AE" authorityName="Lorenz &amp; Loria &amp; Harvey &amp; Harms" authorityYear="2022" class="Arachnida" family="Feaellidae" genus="Mahajanganella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mahajanganella heraclis" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="species" species="heraclis">
<emphasis id="177FA771EBD4E24D709637653F40F3E7" italics="true" pageId="0" pageNumber="649">Mahajanganella heraclis</emphasis>
</taxonomicName>
<emphasis id="38D3249F0DF31617845BC9568BAB9792" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
but smaller female body size (body length 2.03 versus 2.15 in
<taxonomicName id="C8165E59AB4DE5BBB7DD43BE1F3F5EE3" lsidName="M. heraclis" pageId="0" pageNumber="649" rank="species" species="heraclis">
<emphasis id="9DDEBAFF27D21F748CCED7FB76BC31AE" italics="true" pageId="0" pageNumber="649">M. heraclis</emphasis>
</taxonomicName>
<emphasis id="B7F2011098A74053AE3C161469FDAF03" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
), and most slender carapace (1.47 (♂)/1.41 (♀)
<normalizedToken id="7F6C24E2A35C22F6D30EC68BEE86848D" originalValue="×">x</normalizedToken>
longer than broad versus 1.35 (♂)/1.29 (♀) in
<taxonomicName id="50D4D1ED0389342E74B6BB87FFB45B3F" lsidName="M. heraclis" pageId="0" pageNumber="649" rank="species" species="heraclis">
<emphasis id="AF444F22645202CDE3AF7047A0F33A80" italics="true" pageId="0" pageNumber="649">M. heraclis</emphasis>
</taxonomicName>
); most slender femora of
<taxonomicName id="375CD79E6E4B4881F21BF0BA66BDEF6D" authorityName="Lorenz &amp; Loria &amp; Harvey &amp; Harms" authorityYear="2022" class="Arachnida" family="Feaellidae" genus="Mahajanganella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mahajanganella" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="genus">
<emphasis id="93521EFF78E3C9930ABC7F02BE85B47D" italics="true" pageId="0" pageNumber="649">Mahajanganella</emphasis>
</taxonomicName>
<emphasis id="FB4B7C3C9BBB65AAC1D2F6D20AC1AB2F" bold="true" pageId="0" pageNumber="649">gen. nov.</emphasis>
(1.57/1.74 ♂/♀
<normalizedToken id="0CF117F26011C5C919CD83741050B644" originalValue="×">x</normalizedToken>
longer than broad versus 1.64/1.62 ♂/♀ in
<taxonomicName id="1DDCB3FB43EEA046CB93ECE6FD495BE2" lsidName="M. heraclis" pageId="0" pageNumber="649" rank="species" species="heraclis">
<emphasis id="8CE92FF8353E3CB438437C7D8FA99B6A" italics="true" pageId="0" pageNumber="649">M. heraclis</emphasis>
</taxonomicName>
<emphasis id="66E851687DF4F170C9181E7EEFCCF52F" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="10C61E16F5A05BA2F6C0AECE2F01DE32" pageId="0" pageNumber="649" type="etymology">
<paragraph id="AE49005125C1998B82287173C0BF3E0E" pageId="0" pageNumber="649">Etymology.</paragraph>
<paragraph id="79829F25FC10179CD91FBB0289B17397" pageId="0" pageNumber="649">
This species is a patronym honoring Arnold Schwarzenegger, a famous former bodybuilder from Austria, known as an actor in the movie
<normalizedToken id="FE35FA8721B7533E0C01786290271CA9" originalValue="“Terminator”">&quot;Terminator&quot;</normalizedToken>
, former governor of California and now supporting conservation programs.
</paragraph>
</subSubSection>
<subSubSection id="7E9954AF0BB900A5D580B1DB6C7DBE36" pageId="0" pageNumber="649" type="description">
<paragraph id="07CF595A99A82801A4715761EFD3D621" pageId="0" pageNumber="649">Description.</paragraph>
<paragraph id="355D7EB0C4678B4EA4220DD7364C020F" pageId="0" pageNumber="649">
The following description is based on holotype and allotype. -
<emphasis id="2CCF73F33ACFFE2D6572C62A65F8A316" italics="true" pageId="0" pageNumber="649">Carapace</emphasis>
(Fig.
<figureCitation id="48EFE2E45428A0CF90075946B0D8E189" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Drawings of Mahajanganella schwarzeneggeri sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure16" httpUri="https://binary.pensoft.net/fig/782008" pageId="0" pageNumber="649">16A</figureCitation>
): 1.47 (♂), 1.38-1.43 (♀) times longer than broad. -
<emphasis id="186B4F7D63735854E0BCB0CB6A9EE005" italics="true" pageId="0" pageNumber="649">Pedipalps</emphasis>
(Fig.
<figureCitation id="C6AE897BCEE99E36DB7D4B20FB649BFE" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Drawings of Mahajanganella schwarzeneggeri sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure16" httpUri="https://binary.pensoft.net/fig/782008" pageId="0" pageNumber="649">16C, D</figureCitation>
): see trichobothrial pattern in genus description. Chelal fixed finger with 8 teeth in the OR, 13-17 in the MR and 9-12 in the IR; movable finger with 8-9 teeth in the OR, 11-13 in the MR and 9-12 in the IR; terminal teeth build a group of 6 equally sized and 1 larger tooth on the fixed finger and 8 equally sized teeth on the movable finger. -
<taxonomicName id="FCBE13859F91E90A195A1D25EDEFE9F3" genus="Dimensions" lsidName="Dimensions" pageId="0" pageNumber="649" rank="genus">
<emphasis id="BFB2AAF8CC04A24812EB574E0A426D7E" italics="true" pageId="0" pageNumber="649">Dimensions</emphasis>
</taxonomicName>
(mm):
<emphasis id="5C65BF36CC2C52A08AFDA6C8D94E45C3" italics="true" pageId="0" pageNumber="649">Holotype</emphasis>
♂: Body length 1.93; abdomen length 1.34; width (with pleura) 1.17; (without pleura) 1.07; carapace length 0.56; width 0.38. Pedipalp: trochanter 0.23; femur length 0.56; width 0.32; patella 0.44; chela (without pedicel) 0.47; hand length 0.09; width 0.13; movable finger 0.36. Leg I: trochanter 0.09; femur 0.21; patella 0.14; tibia 0.15; tarsus 0.26. Leg IV: trochanter 0.15; femur 0.13; patella 0.29; tibia 0.24; tarsus 0.31.
<emphasis id="88D18F31CF543F5547AC6DE541C37927" italics="true" pageId="0" pageNumber="649">Allotype</emphasis>
♀: Body length 2.04; abdomen length 1.44; width (with pleura) 1.30; (without pleura) 1.18; carapace length 0.60; width 0.42. Pedipalp: trochanter 0.27; femur length 0.56; width 0.34; patella 0.48; chela (without pedicel) 0.51; hand length 0.10; width 0.15; movable finger 0.39. Leg I: trochanter 0.09; femur 0.21; patella 0.20; tibia 0.13; tarsus 0.28. Leg IV: trochanter 0.16; femur 0.13; patella 0.34; tibia 0.31; tarsus 0.33.
</paragraph>
<caption id="82B6955BB2EE7CCD51792E5D2F3BD71F" doi="10.3897/asp.80.e90570.figure16" httpUri="https://binary.pensoft.net/fig/782008" pageId="0" pageNumber="649" start="Figure 16" startId="F16">
<paragraph id="E1F4442DCBC03D1549264C60C037EC26" pageId="0" pageNumber="649">
<emphasis id="8F4690BDC2D932ED94AA5B992E4555A0" bold="true" pageId="0" pageNumber="649">Figure 16.</emphasis>
Drawings of
<taxonomicName id="C5D6B1423BCBE254EBEDAE3211EC29D9" authorityName="Lorenz &amp; Loria &amp; Harvey &amp; Harms" authorityYear="2022" class="Arachnida" family="Feaellidae" genus="Mahajanganella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mahajanganella schwarzeneggeri" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="species" species="schwarzeneggeri">
<emphasis id="10A0E3246682EA49E93E3AABAA27F84D" italics="true" pageId="0" pageNumber="649">Mahajanganella schwarzeneggeri</emphasis>
</taxonomicName>
<emphasis id="655B3E4FEAF45964B62DA23977BCEAF5" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
:
<emphasis id="14374FAB74D53DCEF5DF05007A83EDEC" bold="true" pageId="0" pageNumber="649">A</emphasis>
carapace from dorsal;
<emphasis id="C733F42F8ABB5FF03A148D292EB2330A" bold="true" pageId="0" pageNumber="649">B</emphasis>
leg IV;
<emphasis id="F5E35E65E893D4CE624CBD3941902518" bold="true" pageId="0" pageNumber="649">C</emphasis>
left chela from dorsal with trichobothrial pattern;
<emphasis id="F4000B9EB045B4F874A27094969AF9CE" bold="true" pageId="0" pageNumber="649">D</emphasis>
right pedipalp from dorsal. Scale bars: 0.2 mm (
<emphasis id="15B5677CE7E13B372ABB62269B0C2189" bold="true" pageId="0" pageNumber="649">A</emphasis>
-
<emphasis id="6DBAD4CCC94991B10C84E34759840C53" bold="true" pageId="0" pageNumber="649">C</emphasis>
); 0.4 mm (
<emphasis id="19A8CF9FED558FC79AEF83F6AE632891" bold="true" pageId="0" pageNumber="649">D</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="664CA8DBD3C8F12975FABDC0A8D46A12" pageId="0" pageNumber="649" type="variation">
<paragraph id="D2F06FB168955CEABAA3CACDF7A11A49" pageId="0" pageNumber="649">Variation.</paragraph>
<paragraph id="1FC9F9A9EE42E9910E99B3093C4F2CE6" pageId="0" pageNumber="649">Body length 1.85-1.93 (♂), 2.02-2.04 (♀); abdomen length 1.25-1.34 (♂), 1.43-1.44 (♀); width (with pleura) 1.14-1.17 (♂), 1.29-1.30 (♀); width (without pleura) 1.03-1.07 (♂), 1.17-1.18 (♀); 1.21-1.25 (♂), 1.22 (♀) times longer than broad; carapace length 0.56 (♂), 0.58-0.60 (♀); width 0.38 (♂), 0.42 (♀); 1.47 (♂), 1.38-1.43 (♂) times longer than broad. Pedipalp: trochanter 0.23 (♂), 0.25-0.27 (♀); femur length 0.52-0.56 (♂), 0.56-0.58 (♀); width 0.29-0.32 (♂), 0.33-0.34 (♀); 1.75-1.79 (♂), 1.65-1.76 (♀) times longer than broad; patella 0.42-0.44 (♂), 0.48 (♀); chela (without pedicel) 0.44-0.47 (♂), 0.51 (♀); hand length 0.09-0.10 (♂), 0.09-0.10 (♀); width 0.13-0.14 (♂), 0.15 (♀); 0.67-0.69 (♂), 0.67-0.71 (♀) times longer than broad; movable finger 0.36 (♂), 0.39-0.40 (♀); femur 1.24-1.27 (♂), 1.17-1.21 (♀) times longer than patella; femur 1.18-1.19 (♂), 1.10-1.14 (♀) times longer than chela.</paragraph>
</subSubSection>
<subSubSection id="211B9E3B247DA96FF0D7415C4211D622" pageId="0" pageNumber="649" type="genbank code">
<paragraph id="05B3271F991830AEEF64034E6197AA6C" pageId="0" pageNumber="649">GenBank Code.</paragraph>
<paragraph id="1CC5AAD6D107F4F053F5ADB8CDD12152" pageId="0" pageNumber="649">Genetic data is unavailable for this species.</paragraph>
</subSubSection>
<subSubSection id="EC5C83025E28DC6BB23241342115EFC9" pageId="0" pageNumber="649" type="habitat">
<paragraph id="96DE54A305D650D583C3117B455A5033" pageId="0" pageNumber="649">Habitat.</paragraph>
<paragraph id="23572C48D39CB094A77662F8BD60F8D3" pageId="0" pageNumber="649">Specimens were found in sifted litter in tropical dry forest at an elevation of 10 m very close to the coastline.</paragraph>
</subSubSection>
<subSubSection id="398904C0528607002A2BAA1C6803337A" pageId="0" pageNumber="649" type="distribution">
<paragraph id="47FDAAF02EEC8AB039479854121543A1" pageId="0" pageNumber="649">Distribution.</paragraph>
<paragraph id="D788D5C331A67F197FDA0A9E8E6D0A52" pageId="0" pageNumber="649">Known only from the type locality (BLF6812) in the Boeny Region (formerly Mahajanga Province) of western Madagascar.</paragraph>
</subSubSection>
</treatment>
</document>