956 lines
106 KiB
XML
956 lines
106 KiB
XML
<document id="0F7181049AF1D3A2635BBD89E2F34C56" ID-CLB-Dataset="33888" ID-DOI="http://dx.doi.org/10.3897/zookeys.572.7377" ID-GBIF-Dataset="64701a2d-8e04-4941-a300-30fee945dd42" ID-PMC="PMC4843988" ID-Pensoft-Pub="1313-2970-572-155" ID-PubMed="28050161" ID-ZooBank="F09699B73D334D4C9AF673C4B4FDD63C" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-572-155" ModsDocOrigin="ZooKeys 572" ModsDocTitle="Capoetacoadi, a new species of cyprinid fish from the Karun River drainage, Iran based on morphological and molecular evidences (Teleostei, Cyprinidae)" checkinTime="1458185884506" checkinUser="pensoft" docAuthor="Alwan, Nisreen H., Zareian, Halimeh & Esmaeili, Hamid Reza" docDate="2016" docId="4B6AE239127EC345A5A3CB71FE985B0D" docLanguage="en" docName="ZooKeys 572: 155-180" docOrigin="ZooKeys 572" docSource="http://dx.doi.org/10.3897/zookeys.572.7377" docTitle="Capoeta coadi Alwan, Zareian & Esmaeili, 2016, sp. n." docType="treatment" docUuid="4B5B0984-0C65-4B6D-97CC-31245E179D13" docUuidSource="ZooBank" docVersion="7" lastPageNumber="165" masterDocId="FFE3FFD5FF09FF802128FF83A45C1374" masterDocTitle="Capoetacoadi, a new species of cyprinid fish from the Karun River drainage, Iran based on morphological and molecular evidences (Teleostei, Cyprinidae)" masterLastPageNumber="180" masterPageNumber="155" pageNumber="157" updateTime="1701384575521" updateUser="ExternalLinkService">
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<mods:title id="10D9C23003FAD7EB18767440E4FC3C41">Capoetacoadi, a new species of cyprinid fish from the Karun River drainage, Iran based on morphological and molecular evidences (Teleostei, Cyprinidae)</mods:title>
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<mods:namePart id="82249176E21DC3EC03E38B225FD5A110">Alwan, Nisreen H.</mods:namePart>
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<mods:namePart id="7122237AECFE909254A1BECC3068DD4B">Zareian, Halimeh</mods:namePart>
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<mods:namePart id="6B6FCB4386ACEEBAF4904DFFD1BCB9E4">Esmaeili, Hamid Reza</mods:namePart>
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<mods:typeOfResource id="30CE408487090E0D8C608902280703A0">text</mods:typeOfResource>
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<mods:date id="B15CB42B00A433BF32CC7501B9FC130C">2016</mods:date>
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<mods:detail id="589BF24BD05338ED0C09F6D5E98A6E50" type="volume">
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<mods:number id="CFF1B200F661FAED4B02C5EA0187B7D4">572</mods:number>
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<mods:url id="229F7E235BB8525A3D71D11369B63B49">http://dx.doi.org/10.3897/zookeys.572.7377</mods:url>
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<mods:classification id="D92551F44F99E13F5287233D5AFD6589">journal article</mods:classification>
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<mods:identifier id="DD3E9756484FDC13131950BB573D66D9" type="DOI">http://dx.doi.org/10.3897/zookeys.572.7377</mods:identifier>
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<mods:identifier id="C3D01BABC21586887D318BBC61A823E7" type="Pensoft-Pub">1313-2970-572-155</mods:identifier>
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<treatment id="4B6AE239127EC345A5A3CB71FE985B0D" ID-GBIF-Taxon="127894878" LSID="urn:lsid:zoobank.org:act:4B5B0984-0C65-4B6D-97CC-31245E179D13" httpUri="http://treatment.plazi.org/id/4B6AE239127EC345A5A3CB71FE985B0D" lastPageId="10" lastPageNumber="165" pageId="2" pageNumber="157">
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<subSubSection id="090CB7E417389912837FEE67337A4497" pageId="2" pageNumber="157" type="multiple">
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<paragraph id="20F0FF22C06E6A38A1C23A5A2F068906" pageId="2" pageNumber="157">Taxon classification Animalia Cypriniformes Cyprinidae</paragraph>
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</subSubSection>
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<subSubSection id="10E15E08B50DBFF65CD98448369F54CF" pageId="2" pageNumber="157" type="nomenclature">
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<paragraph id="7D75CCFCB7B521FC9D98752447BAF508" pageId="2" pageNumber="157">
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<taxonomicName id="37680ED2F335ABA40EFB54B7E9FAE97A" ID-CoL="68XM3" LSID="http://zoobank.org/4B5B0984-0C65-4B6D-97CC-31245E179D13" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="2" pageNumber="157" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
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<taxonomicNameLabel id="DA9BA9FC0FCB6485847B9B89CD3C08CB" pageId="2" pageNumber="157">sp. n.</taxonomicNameLabel>
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Figs 1, 2, 3
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</paragraph>
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</subSubSection>
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<subSubSection id="5A605EBD8EE5EA60C384736DFAC0D3E8" pageId="2" pageNumber="157" type="holotype">
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<paragraph id="FB814C2E4372DEACD781562619959FC9" pageId="2" pageNumber="157">Holotype.</paragraph>
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<paragraph id="0D4EF6C49441E6D543884D6F4A246B14" pageId="2" pageNumber="157">
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ZM-CBSU Z190, 157 mm SL; Iran, Kohgiluyeh and Boyer Ahmad prov., Beshar (Bashar) River at Tale Gah village, Karun River drainage,
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<geoCoordinate id="866A0704053069C66F8BC3F07CEDE0E9" direction="north" orientation="latitude" precision="15" value="30.790833">30°47'27"N</geoCoordinate>
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,
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<geoCoordinate id="5F159D568F4C2A3EA1FBE06B8656BFDE" direction="east" orientation="longitude" precision="15" value="51.42028">51°25'13"E</geoCoordinate>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="A9720E60BB5D0C46A191BD1853F56369" lastPageId="3" lastPageNumber="158" pageId="2" pageNumber="157" type="paratypes">
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<paragraph id="E035F7F5B31E88414D94F8A4037765C5" pageId="2" pageNumber="157">Paratypes.</paragraph>
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<paragraph id="55CD008B0C7FF530C55DE2DA404B0926" lastPageId="3" lastPageNumber="158" pageId="2" pageNumber="157">
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ZM-CBSU Z191, 6, 91-157 mm SL; same data as holotype. ZM-CBSU J520, 1, 107 mm SL; ZM-CBSU Z275, 12, 105-152 mm SL; Iran, Koh
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<pageBreakToken id="29558C7E8B86BD73E0FDE27BEF989B1D" pageId="3" pageNumber="158" start="start">giluyeh</pageBreakToken>
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and Boyer Ahmad prov., Beshar (Bashar) River at Tale Gah village, Karun River drainage,
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<geoCoordinate id="AF9E8320ADB6E0F0110258881CC221AF" direction="north" orientation="latitude" precision="15" value="30.790833">30°47'27"N</geoCoordinate>
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,
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<geoCoordinate id="A8FA8249B416935BF3061FCE3E98BBB2" direction="east" orientation="longitude" precision="15" value="51.42028">51°25'13"E</geoCoordinate>
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. 15 December 2014, G. Sayyadzadeh, R. Khaefi, A. Khajehpanah. ZM-CBSU J526, 1, 98 mm SL; ZM-CBSU J533, 1, 114 mm SL; ZM-CBSU J535, 1, 97 mm SL; ZM-CBSU J540, 1, 67 mm SL; All from Iran, Kohgiluyeh and Boyer Ahmad prov., Beshar River at Tange sorkh, Karun River drainage,
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<geoCoordinate id="04EDFBE9001017EADD4182175DACB368" direction="north" orientation="latitude" precision="15" value="30.437222">30°26'14"N</geoCoordinate>
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,
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<geoCoordinate id="CF74949AB71E6E94AC14F51BF92AEE89" direction="east" orientation="longitude" precision="15" value="51.763332">51°45'48"E</geoCoordinate>
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. 24 July 2011, R. Zamaneian Nejad, S. Mirgheiasi, S. Ghasemian. ZM-CBSU J444, 2, 73-90 mm SL; ZM-CBSU J447, 2, 76-111 mm SL; ZM-CBSU J450, 1, 86 mm SL; ZM-CBSU J452, 1, 107 mm SL; ZM-CBSU J459, 2, 104-120 mm SL; ZM-CBSU J464, 1, 110 mm SL; all from Iran, Kohgiluyeh and Boyer Ahmad prov., Beshar River at Mokhtar village, Karun River drainage,
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<geoCoordinate id="9A52D186ECCED28F7607AF49C864B890" direction="north" orientation="latitude" precision="15" value="30.675278">30°40'31"N</geoCoordinate>
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,
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<geoCoordinate id="EBBB5F41125FCDFC7A6FDCDE778626CC" direction="east" orientation="longitude" precision="15" value="51.523888">51°31'26"E</geoCoordinate>
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. 25 May 2011, R. Zamaneian Nejad.
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</paragraph>
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</subSubSection>
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<subSubSection id="C491FA77EAD948462386669CD4DB05D7" pageId="3" pageNumber="158" type="materials_examined">
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<paragraph id="4E365178647B59AB8A064B2CEDF17032" pageId="3" pageNumber="158">Additional material.</paragraph>
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<paragraph id="3FB27008DC95A3923A86B396C9076EB5" pageId="3" pageNumber="158">
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ZM-CBSU 7880-7881, 2, 96.69-158.12 mm SL; Iran, Fars prov., Sepidan city, Gorgu River, a tributary of Beshar River, north of Sepidan city, Karun River drainage,
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<geoCoordinate id="F92FFB63D0CDD89A3CAE8C41372FE3F7" direction="north" orientation="latitude" precision="1" value="30.354717">30°21.283'N</geoCoordinate>
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,
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<geoCoordinate id="FF6B39E8AE334A7E3D22AAA49FE94C4D" direction="east" orientation="longitude" precision="1" value="51.762566">51°45.754'E</geoCoordinate>
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. 2006. H.R. Esmaeili, A. Teimori, M. Ebrahimi and A. Gholamhoseini. SMF 33337, 1, 48.86 mm SL; Iran, Lorestan prov., Hadi River between Zagheh and Polehoru,
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<geoCoordinate id="C88B9F4E1F9DABD200B0A9E9AA88032D" direction="north" orientation="latitude" precision="1" value="33.518967">33°31.138'N</geoCoordinate>
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,
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<geoCoordinate id="3148C87FE836AA8E57F7A8C9BE716B3A" direction="east" orientation="longitude" precision="1" value="48.772335">48°46.340'E</geoCoordinate>
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. 04 March 2008. N. Alwan, K. Borkenhagen, M. Ghanbari Fardi and A. Kazemi. FSJF 2213, 11, 107.92-143.94 mm SL; Iran, Chaharmahal and Bakhtiari Prov., Sandgan River (Sandgan stream) at Sandgan,
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<geoCoordinate id="BE59997CE6E03F421728364416BBA9FE" direction="north" orientation="latitude" precision="1" value="31.261534">31°15.692'N</geoCoordinate>
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,
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<geoCoordinate id="9300ABCAB32AC9D9AF7C094EFF412014" direction="east" orientation="longitude" precision="1" value="51.28583">51°17.150'E</geoCoordinate>
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. 19 April 2007, A. Abdoli and J. Freyhof. FSJF 2233, 2, 156.22-162.23 mm SL; Iran, Kohgiluyeh and Boyer Ahmad prov., Beshar River, 20 km northeast of Yasuj,
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<geoCoordinate id="59B375A6AAAF2F695B5545C4AA8A7ED9" direction="north" orientation="latitude" precision="1" value="30.735867">30°44.152'N</geoCoordinate>
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,
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<geoCoordinate id="EFEDE2F94FD541FCCE0127A6A1A2E499" direction="east" orientation="longitude" precision="1" value="51.492035">51°29.522'E</geoCoordinate>
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. 19 April 2007. A. Abdoli and J. Freyhof. SMF 30865, 1, 26.94 mm SL; Iran, Kohgiluyeh and Boyer Ahmad prov., Beshar River at Tang-e Sorkh,
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<geoCoordinate id="E62259151AF7FB6B981D2D950ECFA84A" direction="north" orientation="latitude" precision="1" value="30.461334">30°27.680'N</geoCoordinate>
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,
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<geoCoordinate id="24D85E331EE7542FB19F1B8F6428BDBE" direction="east" orientation="longitude" precision="1" value="51.74845">51°44.907'E</geoCoordinate>
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. 28 November 2007, K. Borkenhagen, H. R. Esmaeili and F. Wicker (in 96% alcohol). SMF 30871, 1, 28.34 mm SL; Iran, Kohgiluyeh and Boyer Ahmad prov., Beshar River at Tang-e Sorkh,
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<geoCoordinate id="47DF5DB521E0BF43DA896EF71F1CDB68" direction="north" orientation="latitude" precision="1" value="30.461334">30°27.680'N</geoCoordinate>
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,
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<geoCoordinate id="80B7422D0BD793FEC3361CDDA9CFCA9B" direction="east" orientation="longitude" precision="1" value="51.74845">51°44.907'E</geoCoordinate>
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. 28 November 2007. K. Borkenhagen, H. R. Esmaeili and F. Wicker (in 96% alcohol). SMF 33316, 7, 35.22-166.87 mm SL; Iran, Kohgiluyeh and Boyer Ahmad prov., Beshar River at Tang-e Sorkh,
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<geoCoordinate id="1111DE28CB2EEEF92D50B38A9B638B4B" direction="north" orientation="latitude" precision="1" value="30.461334">30°27.680'N</geoCoordinate>
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,
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<geoCoordinate id="AE755CBC10CDD2F6766F3850D821345F" direction="east" orientation="longitude" precision="1" value="51.74845">51°44.907'E</geoCoordinate>
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. 28 November 2007, K. Borkenhagen, H. R. Esmaeili and F. Wicker. SMF 30872, 1, 29.70 mm SL; Iran, Fars prov., Sepidan, Tang-e Tizab,
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<geoCoordinate id="601332D884A192CC3D525AA1528EB978" direction="north" orientation="latitude" precision="1" value="30.391167">30°23.470'N</geoCoordinate>
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,
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<geoCoordinate id="A7AF4A62537BCD5812C2D5656E56FF45" direction="east" orientation="longitude" precision="1" value="51.7785">51°46.710'E</geoCoordinate>
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, 28 November 2007, K. Borkenhagen, H. R. Esmaeili and F. Wicker (in 96% alcohol).
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</paragraph>
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</subSubSection>
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<subSubSection id="6A95AAA553CF9E3288C8DA24121812E7" pageId="4" pageNumber="159" type="capoeta coadi specimens used for molecular genetic analysis">
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<paragraph id="B5E00BC3C947E41350083D292694CB7B" pageId="4" pageNumber="159">
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<taxonomicName id="57782AA9B2BA69DCC47D04B6817D6951" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="4" pageNumber="159" phylum="Chordata" rank="species" species="coadi">
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<pageBreakToken id="41455CE3233D4481ADE3DF1817D94B13" pageId="4" pageNumber="159" start="start">Capoeta</pageBreakToken>
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coadi
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</taxonomicName>
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specimens used for molecular genetic analysis.
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</paragraph>
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<paragraph id="CB6530D48534ABA629CDAC6FC2E2BEB7" pageId="4" pageNumber="159">
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ZM-CBSU M1275,1, Iran, Kohgiluyeh and Boyer Ahmad prov., Beshar River at Dehno village, Karun River drainage,
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<geoCoordinate id="06B760083757846D671C1798462BE757" direction="north" orientation="latitude" precision="15" value="30.648611">30°38'55"N</geoCoordinate>
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,
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<geoCoordinate id="3079CCE75B0517CF25758D2955783C7F" direction="east" orientation="longitude" precision="15" value="51.618053">51°37'05"E</geoCoordinate>
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. 16 January 2014, H.R. Esmaeili, G. Sayyadzadeh, H.R. Mehraban, M. Razbani. GenBank accession number: (COI: KU564296); ZM-CBSU M1447, 2, GenBank accession number: (COI: KU564297, KU564298; cytb: KU564303, KU564304) ZM-CBSU M1458, 2); Iran, Kohgiluyeh and Boyer Ahmad prov., Beshar River at Tale Gah village, Karun River drainage,
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<geoCoordinate id="7A8334E7BA322D2BAD6C21204707A8A6" direction="north" orientation="latitude" precision="15" value="30.790833">30°47'27"N</geoCoordinate>
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,
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<geoCoordinate id="5C1FD2E6DEE0CE1A1C9CD207E9811D31" direction="east" orientation="longitude" precision="15" value="51.42028">51°25'13"E</geoCoordinate>
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. 14 December 2013. G. Sayyadzadeh, A. Khajehpanah, R. Khaefi. GenBank accession number: (COI: KU564294, KU564295; cytb: KU564305, KU564306).
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</paragraph>
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</subSubSection>
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<subSubSection id="65ACAED4AC28348D5810758B4359C7B0" pageId="4" pageNumber="159" type="diagnosis">
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<paragraph id="34841A7F3E90C9C36354A3901F3399D4" pageId="4" pageNumber="159">Diagnosis.</paragraph>
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<paragraph id="77E7F5C905AE1B6E45565533046A5812" pageId="4" pageNumber="159">
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<taxonomicName id="7D5F57117A6E769CB0FA24953F2875BF" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="4" pageNumber="159" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
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sp. n. is distinguished from all other species of
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<taxonomicName id="3CE4B5BC6D5C735C39FBD7EC07D99A1E" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta" order="Cypriniformes" pageId="4" pageNumber="159" phylum="Chordata" rank="genus">Capoeta</taxonomicName>
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by the following combination of characters: last unbranched dorsal-fin ray weakly to moderately ossified and serrated in 1/3-2/3 of its length; scales small, 70-84 total lateral line scales (84 in holotype), 12-17 scales between dorsal-fin origin and lateral line (16 in holotype), 9-11 scales between anal-fin origin and lateral line (11 in holotype), 26-32 encircling least circumference of caudal peduncle (31 in holotype); total gill rakers 14-18 (17 in holotype), 10-13 gill rakers on lower limb of first gill arch (12 in holotype); 45-47 total vertebrae; one posterior pair of barbels; length of the longest dorsal-fin ray 14.92-21.58% SL (18.90 in holotype); head length 22.87-26.33% SL (23.76 in holotype); mouth width 7.48-9.77% SL (8.65 in holotype); bright golden-greenish or silvery body coloration in life.
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</paragraph>
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</subSubSection>
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<subSubSection id="C8F5093F5F2BD245D8C0FE15D425F4CB" lastPageId="6" lastPageNumber="161" pageId="5" pageNumber="160" type="description">
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<paragraph id="24F2AF6CD603ACE65A02922D71FD557A" pageId="5" pageNumber="160">
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<pageBreakToken id="C498E73206D7FF857EFCE612C533CDAE" pageId="5" pageNumber="160" start="start">Description</pageBreakToken>
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.
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</paragraph>
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<paragraph id="0B0759861BC776D553CD4F583CC960E3" pageId="5" pageNumber="160">General body shape and appearance are shown in Figs 1-3, morphometric data in Table 1 and meristic data are summarized in Tables 2-9. Body elongate and cylindrical; predorsal body profile smoothly convex with no marked discontinuity between head and body except when a nuchal hump is present in few specimens; greatest body depth at level of dorsal-fin origin; snout rounded (in 20 specimens) or pointed (in 14 specimens) and not size dependent; mouth inferior; lips slightly fleshy, especially at the mouth corners; lower lip covered with a sharp-edged horny sheath, its anterior margin straight in adult specimens and rounded to almost crescent-shaped in juveniles, with a considerable degree of individual variation.</paragraph>
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<caption id="7892EE4C7433F949B0BDBC8498C96CDD" pageId="5" pageNumber="160">
|
|
<paragraph id="0ED3A1317BFFA509E6BFB4ADDF1C7C80" pageId="5" pageNumber="160">
|
|
Figure 1.
|
|
<taxonomicName id="21DF82481E4271E6D99FB75A7441BB27" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
sp. n., ZM-CBSU Z190, holotype, 157 mm SL; Iran: Kohgiluyeh and Boyer Ahmad, Beshar River, Karun River drainage.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="23882801912CCC3AE75EB11CDCEFB2B8" pageId="5" pageNumber="160">
|
|
<paragraph id="F2F657463AC2C6EAD011A0678DD53EA1" pageId="5" pageNumber="160">
|
|
Figure 2.
|
|
<taxonomicName id="897AD274F26DE6D42FCA62B03EF710C9" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
sp. n., paratypes: aZM-CBSU Z191; 157 mm SLbZM-CBSU Z192, 148 mm SL; Iran: Kohgiluyeh and Boyer Ahmad, Beshar River, Karun River drainage.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="89B510DB8ABE2BA899A21D0D800BB628" pageId="5" pageNumber="160">
|
|
<paragraph id="190D27B66ACF4EFAF5692535B9F19DED" pageId="5" pageNumber="160">
|
|
Figure 3. Live specimen of
|
|
<taxonomicName id="78BD366AC2DC6F6B276018191329B3EF" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
sp. n, Iran: Kohgiluyeh and Boyer Ahmad, Beshar River, Karun River drainage.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="02A9A375D49DBCA33D5C2E882030C2B9" pageId="5" pageNumber="160">
|
|
<paragraph id="8ED95B92EAA748044DC5F9F0F5921DFB" pageId="5" pageNumber="160">
|
|
Table 1. Morphometric data of
|
|
<taxonomicName id="069D6F1CA1BB6588CF8C4055F32FB542" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
sp. n. (holotype ZM-CBSU Z190, and 33 paratypes),
|
|
<taxonomicName id="96565398C58457C10DEF765AFEA9AE40" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta buhsei" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="buhsei">Capoeta buhsei</taxonomicName>
|
|
and
|
|
<taxonomicName id="B028021979E3C3952DF3906E1A1B7D4A" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta saadii" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="saadii">Capoeta saadii</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="5EED94F5F9BBE72717C519D950985828" pageId="5" pageNumber="160">
|
|
<table id="1686CDD55CCC6138A58C9DE8E3AD53B0" pageId="5" pageNumber="160">
|
|
<tr id="092F881A0D2AAB7CA4B7BF91D7FC26A5" pageId="5" pageNumber="160">
|
|
<th id="ADDBBF79FACA5BE1A87D630A444B8092" colspan="1" pageId="5" pageNumber="160" rowspan="1">Holotype</th>
|
|
<th id="89518EC926B2F27ACFF0C985F7BA47C9" colspan="3" pageId="5" pageNumber="160" rowspan="1">Paratypes (n=33)</th>
|
|
<th id="92B097BD321281635BF6F40D920690FE" colspan="3" pageId="5" pageNumber="160" rowspan="1">
|
|
<taxonomicName id="B787B73A2C33CAFCEE631BC5B261185F" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta buhsei" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="buhsei">Capoeta buhsei</taxonomicName>
|
|
(n=27)
|
|
</th>
|
|
<th id="A5D3374EAF18408804F2159BF30D503A" colspan="3" pageId="5" pageNumber="160" rowspan="1">
|
|
<taxonomicName id="C51FCDE410C7A56CBABBB893F33A82D1" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta saadii" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="saadii">Capoeta saadii</taxonomicName>
|
|
(n=20)
|
|
</th>
|
|
</tr>
|
|
<tr id="796D84FC8A33BE34265809A0FD9574FA" pageId="5" pageNumber="160">
|
|
<th id="126EB8DDB30D301A309BD1BEEA292906" colspan="1" pageId="5" pageNumber="160" rowspan="1">Range</th>
|
|
<th id="3E490EEC358CD154A684340F44394C7C" colspan="1" pageId="5" pageNumber="160" rowspan="1">Mean</th>
|
|
<th id="6FCC21BDE0647D11BEE4B8F7D03B2DB5" colspan="1" pageId="5" pageNumber="160" rowspan="1">SD</th>
|
|
<th id="BF1C2787AF40F22E8ED54EDD491AB95A" colspan="1" pageId="5" pageNumber="160" rowspan="1">Range</th>
|
|
<th id="2543D73178FDACE404B728FC130AE7FC" colspan="1" pageId="5" pageNumber="160" rowspan="1">Mean</th>
|
|
<th id="F2E0F72AE74DC279DED83C7D8038EA1E" colspan="1" pageId="5" pageNumber="160" rowspan="1">SD</th>
|
|
<th id="92F88F09D8BB47F0CEB274D3E861BBCF" colspan="1" pageId="5" pageNumber="160" rowspan="1">Range</th>
|
|
<th id="EAF4F4E38E120CC439DCF85E145108BF" colspan="1" pageId="5" pageNumber="160" rowspan="1">Mean</th>
|
|
<th id="E8B72F754430B2A24994164BBE3E89E3" colspan="1" pageId="5" pageNumber="160" rowspan="1">SD</th>
|
|
</tr>
|
|
<tr id="0FE266061B2F6B2739F0B6C0280A4590" pageId="5" pageNumber="160">
|
|
<td id="173657116C03E4A819BB664A08705B25" colspan="10" pageId="5" pageNumber="160" rowspan="1">In percent of standard length</td>
|
|
</tr>
|
|
<tr id="E71C0EBAED6A2EF46BE5558950084334" pageId="5" pageNumber="160">
|
|
<td id="8A5355F94950A1245BABE626873DAD2B" colspan="11" pageId="5" pageNumber="160" rowspan="1">In percent of head length</td>
|
|
</tr>
|
|
</table>
|
|
</paragraph>
|
|
<paragraph id="E1BE9393038C096A74509E10C61A638E" pageId="5" pageNumber="160">Dorsal-fin origin anterior to pelvic-fin origin, its outer margin usually straight to concave with 3-5 unbranched and 8-9 branched rays (3 and 8 in holotype, respectively); last unbranched dorsal-fin ray weakly to moderately ossified, flexible and soft at the tip, serrated in 1/2-2/3 of its length (Fig. 4); pectoral fins not extending to pelvic-fin base; their outer margins usually slightly convex with 16-22 rays in total (19 in holotype) (Table 2); pelvic fins not extending to anal fin base, their outer margin straight or slightly convex and blunt with 7-11 rays in total (8 in holotype) (Table 2); pelvic axillary scale present; anal fin with 3 unbranched and 5 branched rays, outer margin straight or slightly convex; caudal fin forked with 16-19 branched rays (17 in holotype) (Table 3), its tip pointed and its upper lobe often longer than lower one.</paragraph>
|
|
<caption id="2C5B17D8D309AFE00D8B99F18704DFA5" pageId="5" pageNumber="160">
|
|
<paragraph id="B4570C1DCE4AF566DD8DD7F6D55675B7" pageId="5" pageNumber="160">
|
|
Figure 4. Dorsal fins of
|
|
<taxonomicName id="324B7F000C5B977A61ADF3CD98DD42CD" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
sp. n. aZM-CBSU J 444; 73 mm SLbZM-CBSU Z195; 104 mm SLcZM-CBSU Z192; 148 mm SL; Iran: Kohgiluyeh and Boyer Ahmad, Beshar River, Karun River drainage, to show size-dependent variability of the last simple dorsal-fin ray serration.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="FBC6D10B9CE0AA2F0CC0037C318A4C82" pageId="5" pageNumber="160">
|
|
<paragraph id="D2493BC618379EB1EBD55DAC59DC887E" pageId="5" pageNumber="160">
|
|
Table 2. Number of pectoral and pelvic fin rays in examined
|
|
<taxonomicName id="6BB23605714887179E8B4782A6F1B4CD" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="genus">Capoeta</taxonomicName>
|
|
species.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="E7B32658857971970869BB65A4082D31" pageId="5" pageNumber="160">
|
|
<table id="2D15F70F1241598998C3DCB2678FDB0A" pageId="5" pageNumber="160">
|
|
<tr id="3F3C18A151BAE00969C0E780C3C9F976" pageId="5" pageNumber="160">
|
|
<th id="AAB5509ECF58634569D886C6FF4D0D2A" colspan="10" pageId="5" pageNumber="160" rowspan="1">Pectoral fin rays</th>
|
|
<th id="AFF9F2BA47F247CC0C88246D2D4D4C69" colspan="6" pageId="5" pageNumber="160" rowspan="1">Pelvic fin rays</th>
|
|
</tr>
|
|
<tr id="7F836A5487786451FB943D53B2DCF5DB" pageId="5" pageNumber="160">
|
|
<th id="0A473019145E00CE9AB45902FBFB206C" colspan="1" pageId="5" pageNumber="160" rowspan="1">13</th>
|
|
<th id="090B3DC91A61FD97FBD720AAF06E37BC" colspan="1" pageId="5" pageNumber="160" rowspan="1">14</th>
|
|
<th id="C9E22EB4D9B92F77BA18776A9581CA25" colspan="1" pageId="5" pageNumber="160" rowspan="1">15</th>
|
|
<th id="94CEC6047BA839D2AC4D6CD6DDA31139" colspan="1" pageId="5" pageNumber="160" rowspan="1">16</th>
|
|
<th id="B884D3C56D6E6FC6A04022C2E6F7DDC8" colspan="1" pageId="5" pageNumber="160" rowspan="1">17</th>
|
|
<th id="9052B1B6A2A185528F82FF5929B5D06A" colspan="1" pageId="5" pageNumber="160" rowspan="1">18</th>
|
|
<th id="142DD0DC5FEF1986FEF3C46A5783078A" colspan="1" pageId="5" pageNumber="160" rowspan="1">19</th>
|
|
<th id="C3E010D56F3C62AC79DB6506EC9EABFD" colspan="1" pageId="5" pageNumber="160" rowspan="1">20</th>
|
|
<th id="FE60D9F6DE23858259B7B258B3FA0512" colspan="1" pageId="5" pageNumber="160" rowspan="1">21</th>
|
|
<th id="16453328A7E9606B4B9D38CBBF37FD2E" colspan="1" pageId="5" pageNumber="160" rowspan="1">22</th>
|
|
<th id="B4664C8AEBBFD3B24450E66B93843FEA" colspan="1" pageId="5" pageNumber="160" rowspan="1">7</th>
|
|
<th id="599937D9907BCFC2D3EC538C8C5E5AB3" colspan="1" pageId="5" pageNumber="160" rowspan="1">8</th>
|
|
<th id="1094C79A76ABB2710A43C6BAB53B88F3" colspan="1" pageId="5" pageNumber="160" rowspan="1">9</th>
|
|
<th id="A0FBCE0A9A0ABC405CD04CEB9B1314B5" colspan="1" pageId="5" pageNumber="160" rowspan="1">10</th>
|
|
<th id="D48734CCC365A6AE3F5A7EECD6D074D7" colspan="1" pageId="5" pageNumber="160" rowspan="1">11</th>
|
|
</tr>
|
|
<tr id="9F58A665ABD3A4816033CD21519F91BB" pageId="5" pageNumber="160">
|
|
<td id="3C6AC274B262551DB9236A81842BD97A" colspan="1" pageId="5" pageNumber="160" rowspan="1">
|
|
<taxonomicName id="8F2B15B0851A73DEE5E92FD031B30CA1" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta buhsei" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="buhsei">Capoeta buhsei</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="BFFAC3609419D06ED1EC2E2265B48259" pageId="5" pageNumber="160">
|
|
<td id="006A6716BDB0363A301E9405760927C7" colspan="1" pageId="5" pageNumber="160" rowspan="1">
|
|
<taxonomicName id="E40FE368A2022657E876576CA37BEF83" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="7D6B6B21CD24EC7B1B4B74C40DD5E471" pageId="5" pageNumber="160">
|
|
<td id="F7D152BA5D0FB3993EF0E72A0F35C7FF" colspan="1" pageId="5" pageNumber="160" rowspan="1">
|
|
<taxonomicName id="E722C62B5BDE2C6EEAF48CAF15817D23" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta mandica" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="mandica">Capoeta mandica</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="84B1E512049D4758DC6801690F6FFF05" pageId="5" pageNumber="160">
|
|
<td id="A55650AF23360FC96A5815C4A830907F" colspan="1" pageId="5" pageNumber="160" rowspan="1">
|
|
<taxonomicName id="9F4120D47A5347E3D1E77C3B70849DE6" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta saadii" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="saadii">Capoeta saadii</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="EE94106F13819CE87B93A23BA749C239" pageId="5" pageNumber="160">
|
|
<td id="4F7C4F4581DB64BC85ABD03B67F1E8B8" colspan="1" pageId="5" pageNumber="160" rowspan="1">
|
|
<taxonomicName id="880061E6DAD6ADD26A2D6C12067CD980" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta trutta" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="trutta">Capoeta trutta</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
</table>
|
|
</paragraph>
|
|
<caption id="C22686550F2DD53EB586A3C0821442F6" pageId="5" pageNumber="160">
|
|
<paragraph id="BABC1D4189E614E2071365500B7907C0" pageId="5" pageNumber="160">
|
|
Table 3. Number of branched caudal fin rays in examined
|
|
<taxonomicName id="1EBA24B344D9C7FCB19D52E601FB0709" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="genus">Capoeta</taxonomicName>
|
|
species.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="812C95741332A8A540F9B673DE2527CA" pageId="5" pageNumber="160">
|
|
<table id="5D146B39AD42619AAD9DE92CAC7153F2" pageId="5" pageNumber="160">
|
|
<tr id="F3D63A59427CF83B686D6FC5669B9FBA" pageId="5" pageNumber="160">
|
|
<th id="1D88F564EBB01073E44F5C3ECC38E507" colspan="1" pageId="5" pageNumber="160" rowspan="1">Branched caudal fin rays</th>
|
|
<th id="2571302ADC3DFC52EF7342F4A54E7AB8" colspan="1" pageId="5" pageNumber="160" rowspan="1">15</th>
|
|
<th id="5A735372B63C13F06A9A90B1A1FECC7F" colspan="1" pageId="5" pageNumber="160" rowspan="1">16</th>
|
|
<th id="C0BEA97633665646170110B1AD7F67F0" colspan="1" pageId="5" pageNumber="160" rowspan="1">17</th>
|
|
<th id="BD59DFB04AC2F313085D47298F6EE94B" colspan="1" pageId="5" pageNumber="160" rowspan="1">18</th>
|
|
<th id="EAC6D0656DC554C1C070C821ACF1E4C0" colspan="1" pageId="5" pageNumber="160" rowspan="1">19</th>
|
|
<th id="A7D40CFC7DCAB357A54A237FAACDF53F" colspan="1" pageId="5" pageNumber="160" rowspan="1">20</th>
|
|
</tr>
|
|
<tr id="3188E2105AF67962C3293F66F15459CC" pageId="5" pageNumber="160">
|
|
<td id="5936CE4E2D7C03EA0AFF39D0533FCA50" colspan="1" pageId="5" pageNumber="160" rowspan="1">
|
|
<taxonomicName id="27B7A511CA46866D7EA02A7058696CF5" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta buhsei" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="buhsei">Capoeta buhsei</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="0D06C4CBF2E5993BB0EC832C991E27F2" pageId="5" pageNumber="160">
|
|
<td id="EA48277C1D2DBF4850F8808D05D2C9F9" colspan="1" pageId="5" pageNumber="160" rowspan="1">
|
|
<taxonomicName id="61F0CD08D77AE1774594AEB45FE3701B" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta saadii" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="saadii">Capoeta saadii</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="FC1AF35DC12A25F117915055908EFC5F" pageId="5" pageNumber="160">
|
|
<td id="B3A39EE92B1E147AAAD220E05C93FA49" colspan="1" pageId="5" pageNumber="160" rowspan="1">
|
|
<taxonomicName id="A1E326E99F67B29CBB2FB80B0D096390" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta mandica" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="mandica">Capoeta mandica</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="AE22D6CC326B77A06229B51C26761520" pageId="5" pageNumber="160">
|
|
<td id="A75E2F000E15683C3DDC0B020008E579" colspan="1" pageId="5" pageNumber="160" rowspan="1">
|
|
<taxonomicName id="1DF5AA44D7F3B0B4D13FFE2648607E8E" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta trutta" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="trutta">Capoeta trutta</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
</table>
|
|
</paragraph>
|
|
<paragraph id="EEA7F4A21B36E4AC253629836D6086C8" pageId="5" pageNumber="160">
|
|
Scales small, total lateral-line scales 70-84; 12-17 scales between dorsal-fin origin and lateral line (Table 4); 9-11 scales between anal-fin origin and lateral line (Table 4); 26-32 circum-peduncle scales (Table 5); ventral midline and pectoral region covered with deeply embedded scales of reduced size; gill rakers slightly hooked, total gill rakers 14-18 (10-13 gill rakers on lower limb) of first gill arch (Table 8-9); 45-47 total vertebrae; usually one posterior pair of barbels present (very rarely two, 1 out of 51 individual); pharyngeal teeth arranged in 3 rows in the following manner: 2.3.5-5.3.2 and very similar in shape to those of
|
|
<taxonomicName id="1B66027AE20AFA6924D72660912EFADF" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta damascina" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="damascina">Capoeta damascina</taxonomicName>
|
|
; teeth in the main row spatulate or spoon-shaped and crowns flat, narrow and curved.
|
|
</paragraph>
|
|
<caption id="5E9D630543E50D84E28A6095A1AC4E01" pageId="5" pageNumber="160">
|
|
<paragraph id="EB00F9ABF759C61C19558B492E6F4B21" pageId="5" pageNumber="160">
|
|
Table 4. Number of scales above (between dorsal-fin origin and lateral line) and below (between dorsal-fin origin and lateral line) lateral line in examined
|
|
<taxonomicName id="91335E08B5C1EC881CC436EE13CA61D2" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="genus">Capoeta</taxonomicName>
|
|
species.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="D86D847FC170478B0C98DD631D02883C" pageId="5" pageNumber="160">
|
|
<table id="1494A844797764B0DD538D41B867CFE8" pageId="5" pageNumber="160">
|
|
<tr id="90BE18E49BE76194E974C9E348C0662C" pageId="5" pageNumber="160">
|
|
<th id="43ED024B531087028BE36C6D278DB424" colspan="12" pageId="5" pageNumber="160" rowspan="1">Above lateral line</th>
|
|
<th id="F75517F1A668393430C130FFA7A9B671" colspan="9" pageId="5" pageNumber="160" rowspan="1">Below lateral line</th>
|
|
</tr>
|
|
<tr id="79A177A467ED4E32EE02C2F590663991" pageId="5" pageNumber="160">
|
|
<th id="A81EFBD64746DB26A14BE35C77F59314" colspan="1" pageId="5" pageNumber="160" rowspan="1">6</th>
|
|
<th id="D3BEA9424CAE0BD74EDFCED7A56E737A" colspan="1" pageId="5" pageNumber="160" rowspan="1">7</th>
|
|
<th id="A639135B324F1934613A2ECCB9B7339A" colspan="1" pageId="5" pageNumber="160" rowspan="1">8</th>
|
|
<th id="D857F075FCF827D53A47EED7A049868E" colspan="1" pageId="5" pageNumber="160" rowspan="1">9</th>
|
|
<th id="4A5E56BA7538178BC7458F7B88E9B91E" colspan="1" pageId="5" pageNumber="160" rowspan="1">10</th>
|
|
<th id="4F9D5AE261E48CED083FCF1EEB30E7B2" colspan="1" pageId="5" pageNumber="160" rowspan="1">11</th>
|
|
<th id="3E2409ACA4F9BDEEBC3AC244D50DD3C7" colspan="1" pageId="5" pageNumber="160" rowspan="1">12</th>
|
|
<th id="939DF51C9BE6CA71A2FEB40F051641EA" colspan="1" pageId="5" pageNumber="160" rowspan="1">13</th>
|
|
<th id="4042CE7F6437A9E80580680391CE0937" colspan="1" pageId="5" pageNumber="160" rowspan="1">14</th>
|
|
<th id="E904C630E5ABF6F275F896121EC63D54" colspan="1" pageId="5" pageNumber="160" rowspan="1">15</th>
|
|
<th id="31CCAA25C2A221FFF895B9E4E0A31BFF" colspan="1" pageId="5" pageNumber="160" rowspan="1">16</th>
|
|
<th id="60E3FC3A8FFCD3F505D98E6CF10BA712" colspan="1" pageId="5" pageNumber="160" rowspan="1">17</th>
|
|
<th id="EC7E7949F55489E11794CEB9B3721411" colspan="1" pageId="5" pageNumber="160" rowspan="1">5</th>
|
|
<th id="C36DC9F896A1B2A2B18915104BCD7A3F" colspan="1" pageId="5" pageNumber="160" rowspan="1">6</th>
|
|
<th id="61F4565EE83AD79E16B3F148682D682F" colspan="1" pageId="5" pageNumber="160" rowspan="1">7</th>
|
|
<th id="03BC75594E4199791C308F6EDE3549E3" colspan="1" pageId="5" pageNumber="160" rowspan="1">8</th>
|
|
<th id="83F370B5286F89D3CACB87A2DD7CC862" colspan="1" pageId="5" pageNumber="160" rowspan="1">9</th>
|
|
<th id="BC8F1711AB452B03C035B047B7CE8BE2" colspan="1" pageId="5" pageNumber="160" rowspan="1">10</th>
|
|
<th id="75ABD48AA7A39EE080D40B6E80570967" colspan="1" pageId="5" pageNumber="160" rowspan="1">11</th>
|
|
<th id="36148B2AA1BCCD832B42288202A944F1" colspan="1" pageId="5" pageNumber="160" rowspan="1">12</th>
|
|
<th id="85F3A41AD87A0D183E699343D2AEB125" colspan="1" pageId="5" pageNumber="160" rowspan="1">13</th>
|
|
</tr>
|
|
<tr id="74A012981E3993A9634528F8767B9584" pageId="5" pageNumber="160">
|
|
<td id="729141B3A3F13BF56B4B6CF3EB08A41C" colspan="1" pageId="5" pageNumber="160" rowspan="1">
|
|
<taxonomicName id="D07825C717B96644F473354A04C6A7C3" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta buhsei" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="buhsei">Capoeta buhsei</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="951443F9ADAA8D6A74C676547BA0CE65" pageId="5" pageNumber="160">
|
|
<td id="32542CF3AC04C3DAE59045D4EF752516" colspan="1" pageId="5" pageNumber="160" rowspan="1">
|
|
<taxonomicName id="CC76563D8C0A0573BF2EB0C422948EBE" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="EA300A56B52C168F890928E1666B011A" pageId="5" pageNumber="160">
|
|
<td id="3E00C47FAB0FFE6A66EA1E6FC88F571E" colspan="1" pageId="5" pageNumber="160" rowspan="1">
|
|
<taxonomicName id="6B242E66368576726D51FB8EA53C75C4" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta mandica" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="mandica">Capoeta mandica</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="24A3B243B687AEF944B30AEA98C03783" pageId="5" pageNumber="160">
|
|
<td id="B3A2DB33AC7469A21813C2A0D39DA76A" colspan="1" pageId="5" pageNumber="160" rowspan="1">
|
|
<taxonomicName id="2DE06E833B8ACC2DDE4CCFEE920DAB96" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta saadii" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="saadii">Capoeta saadii</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="E23F2EFFD09DA9EF1AC8281C905FDB99" pageId="5" pageNumber="160">
|
|
<td id="1BF6D73D09DDBA95CE05E1EE0E5C74BF" colspan="1" pageId="5" pageNumber="160" rowspan="1">
|
|
<taxonomicName id="372A1458A20F4622ED0CD826B7168836" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta trutta" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="trutta">Capoeta trutta</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
</table>
|
|
</paragraph>
|
|
<caption id="8781602A44318E6B5822E540D8BB78B1" pageId="5" pageNumber="160">
|
|
<paragraph id="A4A17AEBFEADCC4499E9B7230C630759" pageId="5" pageNumber="160">
|
|
Table 5. Number of circum-pendicular scales in examined
|
|
<taxonomicName id="132AE3C89B4C7128E41E9D3A7E08EC85" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="genus">Capoeta</taxonomicName>
|
|
species.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="0A6355804C6D513083E85E8A01A09E96" pageId="5" pageNumber="160">
|
|
<table id="3A0C7DE52A1F6A9DB04AE1AF51C3B55C" pageId="5" pageNumber="160">
|
|
<tr id="647C375C2EEEAA679A945620B020C477" pageId="5" pageNumber="160">
|
|
<th id="B04E4BCA18C44F6B9A0C7AC432CCBB0C" colspan="1" pageId="5" pageNumber="160" rowspan="1">19</th>
|
|
<th id="C54991E32C344F4B611A6B96A3E27660" colspan="1" pageId="5" pageNumber="160" rowspan="1">21</th>
|
|
<th id="95963E8BDBCF1CE2837BCFE5FBDA8706" colspan="1" pageId="5" pageNumber="160" rowspan="1">22</th>
|
|
<th id="0B75874D9AE8EE850AEE3D6B162D78CA" colspan="1" pageId="5" pageNumber="160" rowspan="1">23</th>
|
|
<th id="F2415DAB98CA996556B07C501819C0F6" colspan="1" pageId="5" pageNumber="160" rowspan="1">24</th>
|
|
<th id="6896D22AF34F4CF5048F27855752091D" colspan="1" pageId="5" pageNumber="160" rowspan="1">25</th>
|
|
<th id="6CA3982201E3375794AF424E818BFF69" colspan="1" pageId="5" pageNumber="160" rowspan="1">26</th>
|
|
<th id="5394635B99DD5BCDB8E79C2505F0156B" colspan="1" pageId="5" pageNumber="160" rowspan="1">27</th>
|
|
<th id="3D2918616D49242880C91BCF0176D672" colspan="1" pageId="5" pageNumber="160" rowspan="1">28</th>
|
|
<th id="D46E08A5E5137647D34D4ABE2F19BE63" colspan="1" pageId="5" pageNumber="160" rowspan="1">29</th>
|
|
<th id="EE95B362178C5689E71FB9B99CFCBA17" colspan="1" pageId="5" pageNumber="160" rowspan="1">30</th>
|
|
<th id="D8FAC43FBC666056AA8C17C1276EBA3F" colspan="1" pageId="5" pageNumber="160" rowspan="1">31</th>
|
|
<th id="674348BF3EFF9D9C4C21143FFAC901C5" colspan="1" pageId="5" pageNumber="160" rowspan="1">32</th>
|
|
<th id="12A955695021865954DE65749D4718B9" colspan="1" pageId="5" pageNumber="160" rowspan="1">33</th>
|
|
</tr>
|
|
<tr id="BB6893C1981A312605178C2E41AA8CBF" pageId="5" pageNumber="160">
|
|
<td id="DD4871FA6A41FDC9E4F32DC163D2E7FD" colspan="1" pageId="5" pageNumber="160" rowspan="1">
|
|
<taxonomicName id="4858ED729ADB42C4B9991D42F0358795" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta buhsei" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="buhsei">Capoeta buhsei</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="8FDD26146E8DCF977C775DF7B9B60E73" pageId="5" pageNumber="160">
|
|
<td id="920169400E3EF5FF376725581256892E" colspan="1" pageId="5" pageNumber="160" rowspan="1">
|
|
<taxonomicName id="1E0AAE68233736BCFA9D0650E9CD94B4" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="E7E4F546833B954A8157CC217A82E2C0" pageId="5" pageNumber="160">
|
|
<td id="67F8FDD6C3D2EF9A6A9E0FCBADB9D42A" colspan="1" pageId="5" pageNumber="160" rowspan="1">
|
|
<taxonomicName id="69A7106190765F4A4AE3E9E11EC67592" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta fusca" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="fusca">Capoeta fusca</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="0E5F2E71675BCCD545976BEF7BB310D4" pageId="5" pageNumber="160">
|
|
<td id="9F27B51CE3CB64CC69FEB1967A1214FA" colspan="1" pageId="5" pageNumber="160" rowspan="1">
|
|
<taxonomicName id="725F9E34C54817D9779C240CF0BE618F" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta mandica" order="Cypriniformes" pageId="5" pageNumber="160" phylum="Chordata" rank="species" species="mandica">Capoeta mandica</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
</table>
|
|
</paragraph>
|
|
<paragraph id="99A449DEFA5C45A3CB5259C21053687C" pageId="6" pageNumber="161">
|
|
<pageBreakToken id="2B60388A02F6AC44B95EB05A4F96E936" pageId="6" pageNumber="161" start="start">Coloration</pageBreakToken>
|
|
. Live specimens. Dorsum and sides bright golden-green or silvery, darker dorsally and lighter below the lateral line; dorsal head bright golden-green or light pink-brown; dorsal, anal and caudal fins beige to light brown with light pink to red tinge; pectoral and pelvic-fins beige to light brown or golden with brown tinge on the first few rays (Fig. 3); few large black blotches present on the body of some specimens whereas small diffuse black spots are present only on the body of some juveniles (above the lateral line).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C3842437104778C76F081D2270D3C7CF" pageId="6" pageNumber="161" type="preserved specimens">
|
|
<paragraph id="0D361952B779CE1F83F9372A926CCC26" pageId="6" pageNumber="161">Preserved specimens.</paragraph>
|
|
<paragraph id="66EF35B2D52353A53C30ECAF52E58CA2" pageId="6" pageNumber="161">Dorsum, head and sides grey or brownish-grey dorsally and beige or yellow ventrally; dorsal and caudal fins dusky grey; pectoral, pelvic and anal fins white or beige with or without grey tinge; blotches and spots well discernible (Figs 1-2).</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="CE67CEA90E30DC015BE23F6F86DC8D84" pageId="6" pageNumber="161" type="sexual dimorphism">
|
|
<paragraph id="E8AF89E68B20E71CF802084891D917BB" pageId="6" pageNumber="161">Sexual dimorphism.</paragraph>
|
|
<paragraph id="7762A20C4EB1C81FB4681B1AE310B041" pageId="6" pageNumber="161">Breeding tubercles present in both sexes, being bigger and more pronounced in males. Tubercles present on the sides of the snout but may also cover the entire body surface, on and above the lateral line with one or two tubercles per scale but not on each scale, below the lateral line especially in the area above the anal fin and on the branched anal-fin rays; tip of anal fin reaching to or beyond the vertical of the caudal-fin base in females and to about 2/3 of the caudal peduncle in males.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="CC05D019C8EF786720A363E656FC58F5" pageId="7" pageNumber="162" type="habitat and distribution">
|
|
<paragraph id="0A8A928B87A260892469D63E7EE503FE" pageId="7" pageNumber="162">
|
|
<pageBreakToken id="4BEBCB4B2526A055ACAC47EB0F43E2B0" pageId="7" pageNumber="162" start="start">Habitat</pageBreakToken>
|
|
and distribution.
|
|
</paragraph>
|
|
<paragraph id="F1FEED5DD6A09EDA0A578FE1F60D78F8" pageId="7" pageNumber="162">
|
|
<taxonomicName id="BD56334F3A9FEB77B3A98273D9E69E46" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="7" pageNumber="162" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
sp. n. occurs in medium-fast flowing rivers with usually gravel substrates and clear waters (Fig. 5). At the Beshar River sampling site, the river is about 25 m wide, with substrate consisting of coarse gravel and boulders, and fast-flowing and semi-transparent waters. The physicochemical parameters at the spot were: dissolved oxygen, 9.89 mg/L; total dissolved solids, 190.2 mg/L; salinity, 0.19‰; conductivity, 395
|
|
<normalizedToken id="97A50D9182AC4D758032C93B5BCA1429" originalValue="µs">µs</normalizedToken>
|
|
/cm; pH: 8.5 and water temperature 23.4 °C. It is known only from the Karun River drainage, a system that constitutes the southeastern part of the Tigris-Euphrates River system.
|
|
</paragraph>
|
|
<caption id="BE748804D63F72760EA925631411C967" pageId="7" pageNumber="162">
|
|
<paragraph id="16FA4E869B6728A18777EF1F854DA4E7" pageId="7" pageNumber="162">
|
|
Figure 5. Beshar River at Taleh Gah village, Karun River drainage, type locality of
|
|
<taxonomicName id="D826F429F1CD2FEED1DE2B6E96A9E9F6" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="7" pageNumber="162" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="94FC382AA509262C45BA8EB57373EAF0" pageId="7" pageNumber="162" type="etymology">
|
|
<paragraph id="AE412545B2882280476DAF0987E25BA6" pageId="7" pageNumber="162">Etymology.</paragraph>
|
|
<paragraph id="11A726CD01D5FD545778B0B19C0AC519" pageId="7" pageNumber="162">The new species is named after Brian W. Coad, a well-known ichthyologist for his valuable contribution to the knowledge of freshwater fishes of Iran.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="D62BBFD8363279CC6D3C839BA7D13D23" lastPageId="10" lastPageNumber="165" pageId="7" pageNumber="162" type="comparative remarks">
|
|
<paragraph id="0B94C01DD61DCB1DAFD41E40F377227E" pageId="7" pageNumber="162">Comparative remarks.</paragraph>
|
|
<paragraph id="F735AF5F5A0485CAE4BFD95045598066" lastPageId="8" lastPageNumber="163" pageId="7" pageNumber="162">
|
|
The presence of one pair of barbels in
|
|
<taxonomicName id="40CE1F9189569FEA2F3C469D4695DDBB" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="7" pageNumber="162" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
sets the species apart from
|
|
<taxonomicName id="3A10015CA117AD8416B4A80AF1B63A61" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta antalyensis" order="Cypriniformes" pageId="7" pageNumber="162" phylum="Chordata" rank="species" species="antalyensis">Capoeta antalyensis</taxonomicName>
|
|
,
|
|
<taxonomicName id="BDCBB75C2A92E06DC52298234750E82E" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta baliki" order="Cypriniformes" pageId="7" pageNumber="162" phylum="Chordata" rank="species" species="baliki">Capoeta baliki</taxonomicName>
|
|
,
|
|
<taxonomicName id="46BC25682F114D9EA2E74AD1E8E44887" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta banarescui" order="Cypriniformes" pageId="7" pageNumber="162" phylum="Chordata" rank="species" species="banarescui">Capoeta banarescui</taxonomicName>
|
|
,
|
|
<taxonomicName id="C328F4C6967AD7E728A8460A27E59ED0" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta tinca" order="Cypriniformes" pageId="7" pageNumber="162" phylum="Chordata" rank="species" species="tinca">Capoeta tinca</taxonomicName>
|
|
, and
|
|
<taxonomicName id="D11B4506658AD20F5CBFEDD1A2825A16" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta heratensis" order="Cypriniformes" pageId="7" pageNumber="162" phylum="Chordata" rank="species" species="heratensis">Capoeta heratensis</taxonomicName>
|
|
, all of which have two pairs of barbels based on data from
|
|
<bibRefCitation id="BD144D0AE2C8EF4F5FA160E65BB6958C" author="Turan, D" journalOrPublisher="Revue Suisse de Zoologie" pageId="17" pageNumber="172" pagination="421 - 436" title="A review of Capoetatinca, with descriptions of two new species from Turkey (Teleostei: Cyprinidae)." url="10.5962/bhl.part.80358" volume="113" year="2006 a">Turan et al. (2006a)</bibRefCitation>
|
|
and this study. The new species is further distinguished from
|
|
<taxonomicName id="0C9D47D91AD3CE81C33ABC10BD1ADE05" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta antalyensis" order="Cypriniformes" pageId="7" pageNumber="162" phylum="Chordata" rank="species" species="antalyensis">Capoeta antalyensis</taxonomicName>
|
|
by the presence of serrae on the last unbranched dorsal-fin ray (vs. absence) (Fig. 4), and by number of scales between dorsal-fin origin and lateral line (12-17 vs. 10-12 in
|
|
<taxonomicName id="695D2654986350A6FD2A4BA4EB9396F2" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta antalyensis" order="Cypriniformes" pageId="7" pageNumber="162" phylum="Chordata" rank="species" species="antalyensis">Capoeta antalyensis</taxonomicName>
|
|
)
|
|
<pageBreakToken id="D3B77053290BE84A213E642111C5550A" pageId="8" pageNumber="163" start="start">(</pageBreakToken>
|
|
Table 4), between anal-fin origin and lateral line (9-11 vs. 7), and by total number of the lateral-line scales (70-84 vs. 51-57) (Table 7).
|
|
<taxonomicName id="75D1134DB78E4C16BD2615AAE3FA0619" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
is distinguished from
|
|
<taxonomicName id="E7219BAE2863ADC5DD95B93DB5A3D870" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta banarescui" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="banarescui">Capoeta banarescui</taxonomicName>
|
|
by number of scales between anal-fin origin and lateral line (9-11 vs. 8-9) (Table 4). Data for
|
|
<taxonomicName id="4B539064A657869E296BE16DEA41721A" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta antalyensis" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="antalyensis">Capoeta antalyensis</taxonomicName>
|
|
and
|
|
<taxonomicName id="3B589F11A62C00EDF835FB6610686080" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta banarescui" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="banarescui">Capoeta banarescui</taxonomicName>
|
|
are from
|
|
<bibRefCitation id="2335B5A1A7A219275EA540B6E18F1A11" author="Turan, D" journalOrPublisher="Revue Suisse de Zoologie" pageId="17" pageNumber="172" pagination="421 - 436" title="A review of Capoetatinca, with descriptions of two new species from Turkey (Teleostei: Cyprinidae)." url="10.5962/bhl.part.80358" volume="113" year="2006 a">Turan et al. (2006a)</bibRefCitation>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="B16CB0621F48CE5C8101690EC17148BF" lastPageId="9" lastPageNumber="164" pageId="8" pageNumber="163">
|
|
<taxonomicName id="93ED074EAE1E464DE6E63AF7170BE72E" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
is distinguished from
|
|
<taxonomicName id="91F9630004C4149CC5864C5C9808A585" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta mandica" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="mandica">Capoeta mandica</taxonomicName>
|
|
,
|
|
<taxonomicName id="CA1A45799F2540F1AE7AB39E36B61231" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta erhani" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="erhani">Capoeta erhani</taxonomicName>
|
|
, and
|
|
<taxonomicName id="52A8BC28DB4736B0FE1DB63591CCC070" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta trutta" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="trutta">Capoeta trutta</taxonomicName>
|
|
by having 10-13 gill rakers on the lower limb of the first gill arch (vs. 17-24 in
|
|
<taxonomicName id="CD409E6BBEB06CFF859891720AA6C9BD" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta mandica" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="mandica">Capoeta mandica</taxonomicName>
|
|
, 20-22 in
|
|
<taxonomicName id="45DDCC4C257B94BE273470658EEB3F4D" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta erhani" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="erhani">Capoeta erhani</taxonomicName>
|
|
and 18-25 in
|
|
<taxonomicName id="3C6A0560C0343CE6E99E5A65C585FA72" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta trutta" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="trutta">Capoeta trutta</taxonomicName>
|
|
[data from
|
|
<bibRefCitation id="93E20AD34B76D0499D412D3D69CF0EE6" author="Krupp, F" journalOrPublisher="Universitaet in Mainz, Mainz" pageId="16" pageNumber="171" title="Systematik und Zoogeographie der Suesswasserfische des levantinischen Grabenbruchsystems und der Ostkueste des Mittelmeeres Dissertation zur Erlangung des Grades " Doktor der Naturwissenschaften " am Fachbereich Biologie der Johannes Gutenberg" year="1985">Krupp 1985</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="B66063723FA5CB682B1A14E03BB537C5" author="Turan, D" journalOrPublisher="Ichthyological Exploration of Freshwaters" pageId="17" pageNumber="172" pagination="263 - 270" title="Capoetaerhani, a new species of cyprinid fish from Ceyhan River, Turkey (Teleostei: Cyprinidae)." volume="19" year="2008">Turan et al. 2008</bibRefCitation>
|
|
, Table 8]). The total number of gill rakers in
|
|
<taxonomicName id="3F3E291AA31F82463087DFEE6A587F37" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
specimens is 14-18 that is lower than in
|
|
<taxonomicName id="DE82DB0D7E7311574E1BA7B13168F144" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta mandica" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="mandica">Capoeta mandica</taxonomicName>
|
|
(23-27),
|
|
<taxonomicName id="84AF8BAFAC8B0BA9E5478F15C2259824" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta barroisi" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="barroisi">Capoeta barroisi</taxonomicName>
|
|
(28-30),
|
|
<taxonomicName id="35E415923359E8BC125DC7D35E8A941C" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta turani" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="turani">Capoeta turani</taxonomicName>
|
|
(25-30) and
|
|
<taxonomicName id="B047CE8284A51E0D22DF6E98D90B4E97" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta trutta" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="trutta">Capoeta trutta</taxonomicName>
|
|
(21-31) [data from
|
|
<bibRefCitation id="8310415AF059E566F43FCA8253F1BA77" author="Turan, D" journalOrPublisher="Ichthyological Exploration of Freshwaters" pageId="17" pageNumber="172" pagination="147 - 156" title="Capoetaekmekciae, a new species of cyprinid fish from northeastern Anatolia (Teleostei: Cyprinidae)." volume="17" year="2006 b">Turan et al. (2006b)</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="D664997A429E4926043B176B5DB09BA5" author="Oezulug, M" journalOrPublisher="Ichthyological Exploration of Freshwaters" pageId="17" pageNumber="172" pagination="289 - 296" title="Capoetaturani, a new species of barbel from River Seyhan, Turkey (Teleostei: Cyprinidae)." volume="19" year="2008">
|
|
<normalizedToken id="1F2F8BDC5A1C3608A39457EB9943103B" originalValue="Özuluğ">Oezulug</normalizedToken>
|
|
and Freyhof (2008)
|
|
</bibRefCitation>
|
|
, and this study] Table 9.
|
|
<taxonomicName id="2B5A6CB4E01BB446924D15CE487CB744" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
is further distinguished from
|
|
<taxonomicName id="DBDECE64921EAB39A5D6446FE20B566F" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta mandica" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="mandica">Capoeta mandica</taxonomicName>
|
|
by having fewer pectoral fin rays (16-22 vs. 13-16) (Table 2).
|
|
<taxonomicName id="BB50374B8B64CAB9007293F1D24CF5FE" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
is distinguished from
|
|
<taxonomicName id="D63079E075FB3AC3927F7845C9F7DAB8" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta bergamae" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="bergamae">Capoeta bergamae</taxonomicName>
|
|
,
|
|
<taxonomicName id="4CFE9004ED5355FEB6B97C05F01C7CD8" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta capoeta" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="capoeta">Capoeta capoeta</taxonomicName>
|
|
and
|
|
<taxonomicName id="963127A35E34238FAD61603798AAE168" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta sieboldii" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="sieboldii">Capoeta sieboldii</taxonomicName>
|
|
by number of scales between dorsal-fin origin and lateral line (12-17 in
|
|
<taxonomicName id="52AC6F769D796F5BF66BDCB946437A87" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
vs. 8-10 in
|
|
<taxonomicName id="6FBCD6E8A08E63A8A5B11448C24DD18F" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta capoeta" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="capoeta">Capoeta capoeta</taxonomicName>
|
|
and 9-11 in
|
|
<taxonomicName id="9193B42202B3C7220C240F37564C7E25" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta sieboldii" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="sieboldii">Capoeta sieboldii</taxonomicName>
|
|
); number of scales between anal-fin origin and lateral line (9-11 in
|
|
<taxonomicName id="01500339420142455109CFBF71802D25" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
vs. 7-9 in
|
|
<taxonomicName id="9982B183DF354F2CD635D2FB45C2BA74" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta bergamae" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="bergamae">Capoeta bergamae</taxonomicName>
|
|
, 6-10 in
|
|
<taxonomicName id="6C36280ACB3CB6BF824C118126A7A0BE" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta capoeta" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="capoeta">Capoeta capoeta</taxonomicName>
|
|
and 8-10 in
|
|
<taxonomicName id="1D850829CD8ABE4C4C0CD2A7B5EF420A" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta sieboldii" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="sieboldii">Capoeta sieboldii</taxonomicName>
|
|
); total lateral line scales (70-84 in
|
|
<taxonomicName id="2DA64D4BCE2C6881A18DE5642829CC41" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
vs. 48-66 in
|
|
<taxonomicName id="AD8846CB2EA26CC8869911DC2CAABC8B" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta capoeta" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="capoeta">Capoeta capoeta</taxonomicName>
|
|
and 52-60 in
|
|
<taxonomicName id="18EC1D009C80C80684F5D68843BA449D" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta sieboldii" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="sieboldii">Capoeta sieboldii</taxonomicName>
|
|
) [data from
|
|
<bibRefCitation id="5101D46108CF09C9C9B7C40C8E1514DF" author="Banarescu, PM" journalOrPublisher="AULA-Verlag, Wiebelsheim" pageId="15" pageNumber="170" title="The Freshwater Fishes of Europe. Cyprinidae 2. Part I: Rhodeus to Capoeta" year="1999">Banarescu 1999</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="4F069203590DB9148578CBEECD51328A" author="Turan, D" journalOrPublisher="Ichthyological Exploration of Freshwaters" pageId="17" pageNumber="172" pagination="147 - 156" title="Capoetaekmekciae, a new species of cyprinid fish from northeastern Anatolia (Teleostei: Cyprinidae)." volume="17" year="2006 b">Turan et al. 2006b</bibRefCitation>
|
|
, Tables 4, 7]. In addition to the presence of serrae on the unbranched dorsal-fin ray,
|
|
<taxonomicName id="0975BFEFAE0CFB4604E8AC8A9C139221" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
is set apart from
|
|
<taxonomicName id="1087D9683777A18CAEA636B5EA5822A1" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta caelestis" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="caelestis">Capoeta caelestis</taxonomicName>
|
|
by the number of scales between the dorsal-fin origin and lateral line (12-17 in
|
|
<taxonomicName id="B262A5DAA59EA5A5B434A30E1551197E" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="8" pageNumber="163" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
vs. 10-13.5
|
|
<pageBreakToken id="A9DCEC6115A993CA155CD61C53CA6FC3" pageId="9" pageNumber="164" start="start">in</pageBreakToken>
|
|
<taxonomicName id="0CB92CFF4DDCAE528EF92652CABE3F33" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta caelestis" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="caelestis">Capoeta caelestis</taxonomicName>
|
|
); scales between anal-fin origin and lateral line (9-11 in
|
|
<taxonomicName id="431A6F808E26EDBF7FC59AF3DFFC078E" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
vs. 7-8 in
|
|
<taxonomicName id="476137F3059A05DAD1F177F47BC3F53B" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta caelestis" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="caelestis">Capoeta caelestis</taxonomicName>
|
|
); circum-peduncular scales (26-32 in
|
|
<taxonomicName id="3B8F22436280FF9A4A7B7BB5234D5D33" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
vs. 23-24 in
|
|
<taxonomicName id="369FE4E1253445D4B8EE0C572D957C90" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta caelestis" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="caelestis">Capoeta caelestis</taxonomicName>
|
|
) (Tables 4-5) and probably vertebral counts (45-47 in
|
|
<taxonomicName id="C2C9BA3B3172F8EBE8D8FE4F053415BF" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
vs. 44 in
|
|
<taxonomicName id="0A4A8643F2591CD80E4DF79A117E5095" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta caelestis" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="caelestis">Capoeta caelestis</taxonomicName>
|
|
) [data from
|
|
<bibRefCitation id="6DB06B317C3F84A1C86382A011EA2043" author="Schoeter, C" journalOrPublisher="Ichthyological Exploration of Freshwaters" pageId="17" pageNumber="172" pagination="229 - 236" title="Capoetacaelestis, a new species from Goeksu River, Turkey (Teleostei: Cyprinidae)." volume="20" year="2009">
|
|
<normalizedToken id="849FA56B38D76ED67AA1C65EC2F33A04" originalValue="Schöter">Schoeter</normalizedToken>
|
|
et al. 2009
|
|
</bibRefCitation>
|
|
].
|
|
</paragraph>
|
|
<caption id="A808481E0F833255C941E8796BFFA9A4" pageId="9" pageNumber="164">
|
|
<paragraph id="97855A9616DDB87ECACA9C518F189387" pageId="9" pageNumber="164">
|
|
Table 6. Number of caudal-peduncle scales in examined
|
|
<taxonomicName id="7EA7C05657409E2A14BB23B1F6B8B235" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="genus">Capoeta</taxonomicName>
|
|
species.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="089C06B3282E613A400EC66BA63FB297" pageId="9" pageNumber="164">
|
|
<table id="9A4E9E1DDD968172CF05042981DEA8D1" pageId="9" pageNumber="164">
|
|
<tr id="BC7DCC48564125618E9444E4E82B0414" pageId="9" pageNumber="164">
|
|
<th id="9E0D723058B1E62F9795063D3B372519" colspan="1" pageId="9" pageNumber="164" rowspan="1">10</th>
|
|
<th id="B2B2F937FC6CE9209930EBFC16488907" colspan="1" pageId="9" pageNumber="164" rowspan="1">11</th>
|
|
<th id="2493950AA9E56DF395D62AD3E5A04DD6" colspan="1" pageId="9" pageNumber="164" rowspan="1">12</th>
|
|
<th id="71F14EBA481AE2CF3BEC546482FAE235" colspan="1" pageId="9" pageNumber="164" rowspan="1">13</th>
|
|
<th id="DB82E11B7AA2DA3C9D96C75FA223C964" colspan="1" pageId="9" pageNumber="164" rowspan="1">14</th>
|
|
<th id="61F3B2F1BD1FB727B82DB1F0E37E5197" colspan="1" pageId="9" pageNumber="164" rowspan="1">15</th>
|
|
<th id="638AE7D57D0D97E1FC52BF66CD6FE5DD" colspan="1" pageId="9" pageNumber="164" rowspan="1">16</th>
|
|
<th id="2AEEC0E69303492B47AACA282DA2E485" colspan="1" pageId="9" pageNumber="164" rowspan="1">17</th>
|
|
<th id="0014C63CBC4C4A3D013B2AE6A33B2313" colspan="1" pageId="9" pageNumber="164" rowspan="1">18</th>
|
|
<th id="8C50721AF5E72ADB496F7E1752EC88A9" colspan="1" pageId="9" pageNumber="164" rowspan="1">19</th>
|
|
<th id="A28856F2D9EFCC636EB8F8AD05E66803" colspan="1" pageId="9" pageNumber="164" rowspan="1">20</th>
|
|
<th id="BC89F91960624EE717DDA52034BB81ED" colspan="1" pageId="9" pageNumber="164" rowspan="1">21</th>
|
|
<th id="BE8AD550A44DB34AD4E721A9BF17D945" colspan="1" pageId="9" pageNumber="164" rowspan="1">22</th>
|
|
</tr>
|
|
<tr id="D1C64D0A8E637B5D07AFB8D05168C176" pageId="9" pageNumber="164">
|
|
<td id="6743717385EAD106551EED38B961D081" colspan="1" pageId="9" pageNumber="164" rowspan="1">
|
|
<taxonomicName id="B3B7E644891AFD620FFAD1ADAA689D8A" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta buhsei" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="buhsei">Capoeta buhsei</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="77809AC4D11144F5732D22B4E8F27477" pageId="9" pageNumber="164">
|
|
<td id="FD8A124381D5D726011AB3E19A9674EC" colspan="1" pageId="9" pageNumber="164" rowspan="1">
|
|
<taxonomicName id="2828B2AAE4EA1B90C765F87C3CAD5EC8" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="CF72641A873D7E2DDA3FF1BB7F3CCE5A" pageId="9" pageNumber="164">
|
|
<td id="75845187629D6E66FD4A353A1F941E2D" colspan="1" pageId="9" pageNumber="164" rowspan="1">
|
|
<taxonomicName id="7202BC1E282CC9372800502503FCA5FD" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta mandica" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="mandica">Capoeta mandica</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="D1B1A00689560E0091DAB3106819D6BF" pageId="9" pageNumber="164">
|
|
<td id="EF584AF0D402EAEC67B225F3C8121E58" colspan="1" pageId="9" pageNumber="164" rowspan="1">
|
|
<taxonomicName id="E50422343FE6479183120BCA792888EE" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta saadii" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="saadii">Capoeta saadii</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="089C1CF8171CDCCB69DE7FABF35A2F84" pageId="9" pageNumber="164">
|
|
<td id="2ED39E9B9B157B47F4F4E327DF2FC280" colspan="1" pageId="9" pageNumber="164" rowspan="1">
|
|
<taxonomicName id="D6D1A76C9B8E1106618DC195A7817367" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta trutta" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="trutta">Capoeta trutta</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
</table>
|
|
</paragraph>
|
|
<caption id="478E6CAADF3F6FEF9F87B79C9F3C948A" pageId="9" pageNumber="164">
|
|
<paragraph id="3290AAD662F84501F7635183505DF93F" pageId="9" pageNumber="164">
|
|
Table 7. Number of lateral-line scales in examined
|
|
<taxonomicName id="DA8B0B5C08C55484BD3FCD1205CBC365" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="genus">Capoeta</taxonomicName>
|
|
species.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="339603A2B90F70D4D8D17E8F4F64D2E2" pageId="9" pageNumber="164">
|
|
<table id="C9D75EEEA3D3D805C9C74B25C49EEC62" pageId="9" pageNumber="164">
|
|
<tr id="54AC40C9A79BA700D6C9A5EE7438BBDC" pageId="9" pageNumber="164">
|
|
<th id="5F0C59D4CF4A002F650458DBB37061C5" colspan="1" pageId="9" pageNumber="164" rowspan="1">58</th>
|
|
<th id="5294836D77424B08F7F24510AF1A40AE" colspan="1" pageId="9" pageNumber="164" rowspan="1">59</th>
|
|
<th id="6FE4B6C37A748C24277B56AEA24B5C02" colspan="1" pageId="9" pageNumber="164" rowspan="1">61</th>
|
|
<th id="22CFDF8B7806F7E2868A0240F6F1215F" colspan="1" pageId="9" pageNumber="164" rowspan="1">62</th>
|
|
<th id="E42AC49A444F801FC55910194C670328" colspan="1" pageId="9" pageNumber="164" rowspan="1">63</th>
|
|
<th id="78C78C44E54575E7315637F291EF23A1" colspan="1" pageId="9" pageNumber="164" rowspan="1">64</th>
|
|
<th id="1E90557CBB4C483843B425CFF6123989" colspan="1" pageId="9" pageNumber="164" rowspan="1">65</th>
|
|
<th id="78CA9EBEB760A8BB2F9BC129ABE41742" colspan="1" pageId="9" pageNumber="164" rowspan="1">66</th>
|
|
<th id="4C7473B8BFB19F8AA102CF5AC72A7D6F" colspan="1" pageId="9" pageNumber="164" rowspan="1">67</th>
|
|
<th id="5DD39AB8A39A56FA1D244A4DB0672222" colspan="1" pageId="9" pageNumber="164" rowspan="1">68</th>
|
|
<th id="001C3ADE3FFBA863B5A00472F1C71589" colspan="1" pageId="9" pageNumber="164" rowspan="1">69</th>
|
|
<th id="2847B416B4FEB2F1FF3C72FA5DF5ED1F" colspan="1" pageId="9" pageNumber="164" rowspan="1">70</th>
|
|
<th id="13FDA1CDCE3B05777D4BB74ED1D45E62" colspan="1" pageId="9" pageNumber="164" rowspan="1">71</th>
|
|
<th id="D5B94178FE3F2447F2B640D41215320C" colspan="1" pageId="9" pageNumber="164" rowspan="1">72</th>
|
|
<th id="56FAB5CB8E7C741049AB9FF385BAB8D2" colspan="1" pageId="9" pageNumber="164" rowspan="1">73</th>
|
|
<th id="A1ADDFCAE499D2B802EE6358CE772AA3" colspan="1" pageId="9" pageNumber="164" rowspan="1">74</th>
|
|
<th id="81A7C68FFE2567445DD118887FFA996E" colspan="1" pageId="9" pageNumber="164" rowspan="1">75</th>
|
|
<th id="CF57C6D983B471C0A9CBC8AC77176071" colspan="1" pageId="9" pageNumber="164" rowspan="1">76</th>
|
|
<th id="F48B035D145A242572B88AAA91D538FD" colspan="1" pageId="9" pageNumber="164" rowspan="1">77</th>
|
|
<th id="7C62CEDFAA89CD5078BE0376C9625DCF" colspan="1" pageId="9" pageNumber="164" rowspan="1">78</th>
|
|
<th id="BF62090D3B5DDEF1A4CBEDB9EEEB65E5" colspan="1" pageId="9" pageNumber="164" rowspan="1">79</th>
|
|
<th id="FCACA71AF13E97826AC801A98C55BFD3" colspan="1" pageId="9" pageNumber="164" rowspan="1">80</th>
|
|
<th id="715DE126A2F259464A61D631A9DDF6A4" colspan="1" pageId="9" pageNumber="164" rowspan="1">81</th>
|
|
<th id="3F0CBD5E680CF7465F7A9459799D293C" colspan="1" pageId="9" pageNumber="164" rowspan="1">82</th>
|
|
<th id="326AF1EB6BC18155334595F429F15D3B" colspan="1" pageId="9" pageNumber="164" rowspan="1">83</th>
|
|
<th id="BB1256A25F2158183EF9488E5FC0966B" colspan="1" pageId="9" pageNumber="164" rowspan="1">84</th>
|
|
<th id="A56E5F712BB59211D134EDA219B1DBA6" colspan="1" pageId="9" pageNumber="164" rowspan="1">85</th>
|
|
<th id="67CB5899C780FAB146FB1114FD379251" colspan="1" pageId="9" pageNumber="164" rowspan="1">87</th>
|
|
<th id="138C5DAB4B623466C961CEEA4E8FD94A" colspan="1" pageId="9" pageNumber="164" rowspan="1">89</th>
|
|
</tr>
|
|
<tr id="3B82443FACE5C6B800D0414F1496CC65" pageId="9" pageNumber="164">
|
|
<td id="6068F381BC6521636EF7AFD92989D968" colspan="1" pageId="9" pageNumber="164" rowspan="1">
|
|
<taxonomicName id="B482E16D48A5DD76605E590C39AA8F8F" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta buhsei" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="buhsei">Capoeta buhsei</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="61E5C9D1BB566493B2E6C9AFA239A711" pageId="9" pageNumber="164">
|
|
<td id="AB0E8E7C0CDF4786CE449A36F018D6C9" colspan="1" pageId="9" pageNumber="164" rowspan="1">
|
|
<taxonomicName id="7697407F211D3F7269FA6074BF4A7E6E" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="618F2E72E374091E3519FB571191EB64" pageId="9" pageNumber="164">
|
|
<td id="661F180BFC948125326A59CE67D0ED6B" colspan="1" pageId="9" pageNumber="164" rowspan="1">
|
|
<taxonomicName id="E65CB1081EE5844F572AC161E48C44F0" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta mandica" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="mandica">Capoeta mandica</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="8F80D7E5CA831375FEE4D9EFBB46E972" pageId="9" pageNumber="164">
|
|
<td id="61D5C9F50567336F87C8DFA7CDF4C406" colspan="1" pageId="9" pageNumber="164" rowspan="1">
|
|
<taxonomicName id="F5A2129E6ECF1B4E113A5EBEBBE248C5" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta saadii" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="saadii">Capoeta saadii</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="FB40C5DE2311A5818445F623AD0F3B6E" pageId="9" pageNumber="164">
|
|
<td id="3C863DE44E3F0FDAE4290663D0FFF3AE" colspan="1" pageId="9" pageNumber="164" rowspan="1">
|
|
<taxonomicName id="C2CD0A291F83DD341B29EDAA1AC0527D" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta trutta" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="trutta">Capoeta trutta</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
</table>
|
|
</paragraph>
|
|
<caption id="FBDC97D29AA2A005AE12095E95FCC0FF" pageId="9" pageNumber="164">
|
|
<paragraph id="07A304DD88E81303421C4EB158DD1B85" pageId="9" pageNumber="164">
|
|
Table 8. Gill rakers on the lower limb of the first gill arch in studied
|
|
<taxonomicName id="3780F0F279D891FF78F0AFA5EA292706" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="genus">Capoeta</taxonomicName>
|
|
species.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="13669E9789CC344B6C551FC4F92EB54B" pageId="9" pageNumber="164">
|
|
<table id="B7D6B8F6917602C8435761D605020C57" pageId="9" pageNumber="164">
|
|
<tr id="CD931D891FA70AC5C1925F2C0A4495CB" pageId="9" pageNumber="164">
|
|
<th id="D5ED3916BA40096E0D4AA77975FDAF87" colspan="1" pageId="9" pageNumber="164" rowspan="1">GR</th>
|
|
<th id="DB8F1BB3E7D3CAC45F47E9F8619DB4A6" colspan="1" pageId="9" pageNumber="164" rowspan="1">8</th>
|
|
<th id="2DB105775DC9D6FC3BD5541DEA5F0916" colspan="1" pageId="9" pageNumber="164" rowspan="1">9</th>
|
|
<th id="B506333614A0ED1C5DACDE80DE12463D" colspan="1" pageId="9" pageNumber="164" rowspan="1">10</th>
|
|
<th id="77444B98BE2BE4A7F8E9B3AC35EEB081" colspan="1" pageId="9" pageNumber="164" rowspan="1">11</th>
|
|
<th id="55D775EF852F914AF9B6353B23456B60" colspan="1" pageId="9" pageNumber="164" rowspan="1">12</th>
|
|
<th id="18B013D0376C31926F8AB830EF1A14FE" colspan="1" pageId="9" pageNumber="164" rowspan="1">13</th>
|
|
<th id="164556723B0231A19617519B2D9AB248" colspan="1" pageId="9" pageNumber="164" rowspan="1">17</th>
|
|
<th id="09A528D2F0434AEC2CD84C7B06821489" colspan="1" pageId="9" pageNumber="164" rowspan="1">18</th>
|
|
<th id="E7BD279660DF49FB0E4121E69359218E" colspan="1" pageId="9" pageNumber="164" rowspan="1">19</th>
|
|
<th id="C4BB9FBD5D7C032937970C79949CA751" colspan="1" pageId="9" pageNumber="164" rowspan="1">20</th>
|
|
<th id="1A95CD086AAF3F70B2699004FECAA419" colspan="1" pageId="9" pageNumber="164" rowspan="1">22</th>
|
|
<th id="01B6D1311E56AA2F10AE8E8220522C2F" colspan="1" pageId="9" pageNumber="164" rowspan="1">24</th>
|
|
</tr>
|
|
<tr id="2CCF1CDECE5F09016C89C3516162560F" pageId="9" pageNumber="164">
|
|
<td id="B8C8435FB1E7468B1CAABDADDEA5D619" colspan="1" pageId="9" pageNumber="164" rowspan="1">
|
|
<taxonomicName id="21DB722AD5C719528E60B3715FDA32CD" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta buhsei" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="buhsei">Capoeta buhsei</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="CA9DF9F2DC0A68DB29C80C617A6E0108" pageId="9" pageNumber="164">
|
|
<td id="2ADAF8B0B4F183FFCA1E620C5B8A70BA" colspan="1" pageId="9" pageNumber="164" rowspan="1">
|
|
<taxonomicName id="733597310E6F9DE7D9CEC4B00310B37A" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="7B746D5F33946C573A6E58AB2D6659A8" pageId="9" pageNumber="164">
|
|
<td id="7C58FE86C9A8836D625B84D2FAF539D7" colspan="1" pageId="9" pageNumber="164" rowspan="1">
|
|
<taxonomicName id="BD3B94166BB76140DFC0F8AA006498D4" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta mandica" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="mandica">Capoeta mandica</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
</table>
|
|
</paragraph>
|
|
<caption id="1521B662029EA9048A8C5A7F61BA8299" pageId="9" pageNumber="164">
|
|
<paragraph id="BB1B28EAABBC309421D8FAA56284DDC8" pageId="9" pageNumber="164">
|
|
Table 9. Number of total gill rakers on the first gill arch in examined
|
|
<taxonomicName id="67D72F37C967D37977CC60C023EE7385" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="genus">Capoeta</taxonomicName>
|
|
species.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="5C0D8D2489920DFE156F330589991AF4" pageId="9" pageNumber="164">
|
|
<table id="981C60591F76DEEEF25C747D4FF4E732" pageId="9" pageNumber="164">
|
|
<tr id="CDADD3CEB8F71612619FF8A7209A2551" pageId="9" pageNumber="164">
|
|
<th id="FFAC7D3354A4F2C565BFACD384AC16C6" colspan="1" pageId="9" pageNumber="164" rowspan="1">12</th>
|
|
<th id="ADF057207FC41DB4F437CF9B08D903B8" colspan="1" pageId="9" pageNumber="164" rowspan="1">13</th>
|
|
<th id="985DC7E97B2CC9244880BFBA2D58398D" colspan="1" pageId="9" pageNumber="164" rowspan="1">14</th>
|
|
<th id="86985599AB70542269CE1C4D932E5FE5" colspan="1" pageId="9" pageNumber="164" rowspan="1">15</th>
|
|
<th id="F2A2B88F973A9AC1671EF32DF10E9EB0" colspan="1" pageId="9" pageNumber="164" rowspan="1">16</th>
|
|
<th id="D29E1BE852F30C55F2EB91A23E1BC6D4" colspan="1" pageId="9" pageNumber="164" rowspan="1">17</th>
|
|
<th id="92B084AB7ADC10BC8F821047538E2653" colspan="1" pageId="9" pageNumber="164" rowspan="1">18</th>
|
|
<th id="900798B31C8885A0BCFBF3428644F0F1" colspan="1" pageId="9" pageNumber="164" rowspan="1">21</th>
|
|
<th id="A60A3E6630B45CC7D9D977B2F92B6664" colspan="1" pageId="9" pageNumber="164" rowspan="1">22</th>
|
|
<th id="9EA4CE29868EA461A74C676BA9961F07" colspan="1" pageId="9" pageNumber="164" rowspan="1">23</th>
|
|
<th id="96BFCE9A4159825BA2158BF37673280B" colspan="1" pageId="9" pageNumber="164" rowspan="1">24</th>
|
|
<th id="A7D374914971EBB7A11419C1A366D836" colspan="1" pageId="9" pageNumber="164" rowspan="1">25</th>
|
|
<th id="67AC5FE251BF20E76A72ACC298CA861C" colspan="1" pageId="9" pageNumber="164" rowspan="1">26</th>
|
|
<th id="0F424DE1528338303073AF21E19288E4" colspan="1" pageId="9" pageNumber="164" rowspan="1">27</th>
|
|
<th id="59C1BAB19A923F0726F14356A8122C89" colspan="1" pageId="9" pageNumber="164" rowspan="1">28</th>
|
|
<th id="6CB0AA4ED7E5BB7EA2700A7332A782AB" colspan="1" pageId="9" pageNumber="164" rowspan="1">31</th>
|
|
</tr>
|
|
<tr id="5A3F1262B89D96AF71E15FBC214A38D5" pageId="9" pageNumber="164">
|
|
<td id="0EB29DDB0562B9286401E27D6CFFDD53" colspan="1" pageId="9" pageNumber="164" rowspan="1">
|
|
<taxonomicName id="B9FCC3465629E263296067276A303B84" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta buhsei" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="buhsei">Capoeta buhsei</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="3C7085E5109D3DFB5E5D410817B85FD0" pageId="9" pageNumber="164">
|
|
<td id="17047A46E3B418EE3984027E928D94D7" colspan="1" pageId="9" pageNumber="164" rowspan="1">
|
|
<taxonomicName id="0E4805A0844FD59B6FC20AC85A09CC81" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="E9FC7B732EFA7644CC96F5146F230EC1" pageId="9" pageNumber="164">
|
|
<td id="B70511F7867F25FEE6A1842FB1450812" colspan="1" pageId="9" pageNumber="164" rowspan="1">
|
|
<taxonomicName id="89F084483B8B20F7AB1E2E2A3B1DB5A4" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta mandica" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="mandica">Capoeta mandica</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="35548A30CB2BF8ABF116EFD2A1555BB4" pageId="9" pageNumber="164">
|
|
<td id="2EC323C074713950C99E515059B2BC55" colspan="1" pageId="9" pageNumber="164" rowspan="1">
|
|
<taxonomicName id="2224774CBF70F9470D2069A2BD330866" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta saadii" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="saadii">Capoeta saadii</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr id="9CCC7D29B2C40C18FF7D210390D30937" pageId="9" pageNumber="164">
|
|
<td id="CD7A21240100B5327EF1AB57C2B82C4A" colspan="1" pageId="9" pageNumber="164" rowspan="1">
|
|
<taxonomicName id="DE67C4B2112459017D2392264EA629A6" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta trutta" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="trutta">Capoeta trutta</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
</table>
|
|
</paragraph>
|
|
<paragraph id="43E6845C0A29F0ABA3EDB3E1F14A8119" pageId="9" pageNumber="164">
|
|
It is distinguished from
|
|
<taxonomicName id="144CE7F059C96E8ABE9E7230612F26FE" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta damascina" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="damascina">Capoeta damascina</taxonomicName>
|
|
by having 11-13, modally 13, gill rakers on the lower limb of the first gill arch (vs. 12-18, modally 14-15) (
|
|
<bibRefCitation id="D19856361EEBA56E16615CEC9B81F509" author="Alwan, N" journalOrPublisher="Goethe Universitaet, Frankfurt a. M., Germany" pageId="15" pageNumber="170" title="Systematics, taxonomy, phylogeny and zoogeography of the Capoetadamascina species complex (Pisces: Teleostei: Cyprinidae) inferred from comparative morphology and molecular markers" year="2011">Alwan 2011</bibRefCitation>
|
|
, Table 8).
|
|
<taxonomicName id="EDE092A1DEBDE24FE22E34362DFCA6CA" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
is clearly distinguished from
|
|
<taxonomicName id="F1BE80BBFBE89E41A920EDB5A6A144A3" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta ekmekciae" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="ekmekciae">Capoeta ekmekciae</taxonomicName>
|
|
by number of scales between dorsal-fin origin and lateral line (12-17 in
|
|
<taxonomicName id="5DB7C621A0474FD4AB4898A8736ECAA3" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
vs. 9-10 in
|
|
<taxonomicName id="40B0F846F66ED404C67D78276977AF1E" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta ekmekciae" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="ekmekciae">Capoeta ekmekciae</taxonomicName>
|
|
); number of scales between anal-fin origin and lateral line (9-11 in
|
|
<taxonomicName id="D6AC1E7B3B83C6C1D5BBF9945A075458" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
vs. 6-7 in
|
|
<taxonomicName id="7EDA5FB075580FEB446EF986434AB902" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta ekmekciae" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="ekmekciae">Capoeta ekmekciae</taxonomicName>
|
|
) (Table 4); number of lateral line scales (70-84 in
|
|
<taxonomicName id="1C33885A374426AE58BF414F2441C7A2" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
vs. 55-61 in
|
|
<taxonomicName id="9B8F1B75E0BEEE464FE350273EF287D6" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta ekmekciae" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="ekmekciae">Capoeta ekmekciae</taxonomicName>
|
|
[data from
|
|
<bibRefCitation id="9A47E1AE00C44F1EA773C302D3E6B480" author="Turan, D" journalOrPublisher="Ichthyological Exploration of Freshwaters" pageId="17" pageNumber="172" pagination="147 - 156" title="Capoetaekmekciae, a new species of cyprinid fish from northeastern Anatolia (Teleostei: Cyprinidae)." volume="17" year="2006 b">Turan et al. 2006b</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="08B97AFEBCA97C420C767FC42D696D00" author="Alwan, N" journalOrPublisher="Goethe Universitaet, Frankfurt a. M., Germany" pageId="15" pageNumber="170" title="Systematics, taxonomy, phylogeny and zoogeography of the Capoetadamascina species complex (Pisces: Teleostei: Cyprinidae) inferred from comparative morphology and molecular markers" year="2011">Alwan 2011</bibRefCitation>
|
|
].
|
|
</paragraph>
|
|
<paragraph id="D5089B76D5422E50F18EA4423349F9D6" pageId="9" pageNumber="164">
|
|
<taxonomicName id="685B29B7C8FEEF642EC5EDFDA1DD79DB" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
is distinguished from
|
|
<taxonomicName id="E1132CB5BBAAADEFBB467A7FB4D55E69" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta kosswigi" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="kosswigi">Capoeta kosswigi</taxonomicName>
|
|
by total number of gill rakers (Table 9): 14-18 in
|
|
<taxonomicName id="D52F49DBFF47C2919ADF1CF04F4C99CC" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
vs. 19-28 in
|
|
<taxonomicName id="01940C1E030DA6D21F59B51AF8C75C84" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta kosswigi" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="kosswigi">Capoeta kosswigi</taxonomicName>
|
|
(see
|
|
<bibRefCitation id="CD802D1A5F1D2A03DDC56C31FCA1CB8B" author="Karaman, MS" journalOrPublisher="Mitteilungen aus dem hamburgischen Zoologischen Museum und Institut" pageId="16" pageNumber="171" pagination="17 - 54" title="Suesswasserfische der Tuerkei. 7. Teil. Revision der kleinasiatischen und vorderasiatischen Arten des Genus Capoeta (Varicorhinus, Partim)." volume="66" year="1969">Karaman 1969</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="446035B4841152F150E3B170998C3C58" author="Turan, D" journalOrPublisher="Ichthyological Exploration of Freshwaters" pageId="17" pageNumber="172" pagination="147 - 156" title="Capoetaekmekciae, a new species of cyprinid fish from northeastern Anatolia (Teleostei: Cyprinidae)." volume="17" year="2006 b">Turan et al. 2006b</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="6FA6C50EE2057028254744410572843E" author="Turan, C" journalOrPublisher="Acta Zoologica Cracoviensia" pageId="17" pageNumber="172" pagination="1 - 14" title="Molecular systematics of Capoeta (Cyprinidae) species complex inferred from mitochondrial 16 s rDNA sequence data." url="10.3409/azc.51a_1-2.1-14" volume="51 A" year="2008">Turan 2008</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="4B4208141B7F53C17CEBA378E418B2DC" pageId="9" pageNumber="164">
|
|
<taxonomicName id="8A505B5D3769B7BC34B9C0EA2B400E95" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
is distinguished from
|
|
<taxonomicName id="ADA2424340310DB80200F5EE06E134A0" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta mauricii" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="mauricii">Capoeta mauricii</taxonomicName>
|
|
and
|
|
<taxonomicName id="67C1003FAF66FFDA0AF4896A99E6B3B7" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta pestai" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="pestai">Capoeta pestai</taxonomicName>
|
|
by having a weaker, thinner and less ossified last unbranched dorsal-fin ray in juveniles and adults and fewer scales between dorsal-fin origin and lateral line (12-17 in
|
|
<taxonomicName id="AFF24286FA7866F11C82C56B09AB723D" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
vs. 18-22 in
|
|
<taxonomicName id="1444494E62452284F06F73DE5105D01A" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta mauricii" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="mauricii">Capoeta mauricii</taxonomicName>
|
|
and 16-19 in
|
|
<taxonomicName id="E503638A50D3D00BBE8A2DFAE90E05FD" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta pestai" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="pestai">Capoeta pestai</taxonomicName>
|
|
[data from
|
|
<bibRefCitation id="EDBC9DD618CC439307D79F4CC8388E88" author="Oezulug, M" journalOrPublisher="Ichthyological Exploration of Freshwaters" pageId="17" pageNumber="172" pagination="289 - 296" title="Capoetaturani, a new species of barbel from River Seyhan, Turkey (Teleostei: Cyprinidae)." volume="19" year="2008">
|
|
<normalizedToken id="FC0593CA685C66640AE920A2BEA6CA7F" originalValue="Özuluğ">Oezulug</normalizedToken>
|
|
and Freyhof 2008
|
|
</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="9FEFD7FC0BB75D786F5827F3E0E9AF0F" author="Kuecuek, F" journalOrPublisher="Zoology in the Middle East" pageId="16" pageNumber="171" pagination="71 - 82" title="Capoetamauricii n. sp., a new species of cyprinid fish from Lake Beysehir, Turkey (Osteichthyes: Cyprinidae)." url="10.1080/09397140.2009.10638349" volume="47" year="2009">
|
|
<normalizedToken id="A777E268D439B0D3B92523BDC60F5DE2" originalValue="Küçük">Kuecuek</normalizedToken>
|
|
et al. 2009
|
|
</bibRefCitation>
|
|
]). It is further distinguished from
|
|
<taxonomicName id="1EACD6BC374267252373F41A99FB3EE6" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta pestai" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="pestai">Capoeta pestai</taxonomicName>
|
|
by the absence of spots on the body except in juveniles (vs. presence of many on the body [see
|
|
<bibRefCitation id="1E192A6233F4515A12E19F1B5D6DEAD1" author="Oezulug, M" journalOrPublisher="Ichthyological Exploration of Freshwaters" pageId="17" pageNumber="172" pagination="289 - 296" title="Capoetaturani, a new species of barbel from River Seyhan, Turkey (Teleostei: Cyprinidae)." volume="19" year="2008">
|
|
<normalizedToken id="21D1A8BC3933E9A6A1FFB178C4F75797" originalValue="Özuluğ">Oezulug</normalizedToken>
|
|
and Freyhof 2008
|
|
</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="89F1B0B7CC32534AEF7CC54196CDDFF6" author="Kuecuek, F" journalOrPublisher="Zoology in the Middle East" pageId="16" pageNumber="171" pagination="71 - 82" title="Capoetamauricii n. sp., a new species of cyprinid fish from Lake Beysehir, Turkey (Osteichthyes: Cyprinidae)." url="10.1080/09397140.2009.10638349" volume="47" year="2009">
|
|
<normalizedToken id="A939AD2613BBD7EC53ECF61F8D99B949" originalValue="Küçük">Kuecuek</normalizedToken>
|
|
et al. 2009
|
|
</bibRefCitation>
|
|
]).
|
|
<taxonomicName id="BA0E4E6EBE9FA601F56D8C0CFB1B6F1B" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
is distinguished from
|
|
<taxonomicName id="C5EC1DF927581359A6422093BBF92274" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta umbla" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="umbla">Capoeta umbla</taxonomicName>
|
|
by total number of lateral line scales (70-84 vs. 86-104), number of scales between dorsal-fin origin and lateral line (12-17 vs. 18-24), number of scales between anal-fin origin and lateral line (9-11 vs. 11.5-15.5), and circum-pendicular scales (26-32 vs. 32-39) (see
|
|
<bibRefCitation id="8C82D03B73B49E392C30590EB352ED47" author="Alwan, N" journalOrPublisher="Goethe Universitaet, Frankfurt a. M., Germany" pageId="15" pageNumber="170" title="Systematics, taxonomy, phylogeny and zoogeography of the Capoetadamascina species complex (Pisces: Teleostei: Cyprinidae) inferred from comparative morphology and molecular markers" year="2011">Alwan 2011</bibRefCitation>
|
|
, Tables 4-7).
|
|
</paragraph>
|
|
<paragraph id="9E6D5F62EF7E11B5AEC9FA32C5780994" lastPageId="10" lastPageNumber="165" pageId="9" pageNumber="164">
|
|
Compared to other Iranian species of
|
|
<taxonomicName id="D74EBE5AC9396B627EF324CA04DDD965" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="genus">Capoeta</taxonomicName>
|
|
,
|
|
<taxonomicName id="45C1C65369419F49FEA38DE602A2A0D5" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
has more scales and fewer gill rakers than
|
|
<taxonomicName id="A7986DE6968F18F83B8C3FEF49FC7201" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta aculeata" order="Cypriniformes" pageId="9" pageNumber="164" phylum="Chordata" rank="species" species="aculeata">Capoeta aculeata</taxonomicName>
|
|
(number of scales between dorsal-fin origin and lateral line:
|
|
<pageBreakToken id="8105275F4B625EB78BC9706B80A699D3" pageId="10" pageNumber="165" start="start">12</pageBreakToken>
|
|
-17 vs. 6-10; number of scales between anal-fin origin and lateral line: 9-11 vs. 5-8; circum-peduncular scales: 26-32 vs. 13-23; total number of lateral line scales: 70-84 vs. 36-52; caudal peduncle scales: 14-18 vs. 10-12; gill rakers on the lower limb of the first gill arch: 10-13 vs. 15-18 [data from
|
|
<bibRefCitation id="F12BB3F68C67F85DBD7B09A07BB784EF" author="Coad, BW" journalOrPublisher="Zoology in the Middle East" pageId="15" pageNumber="170" pagination="63 - 72" title="Capoetaaculeata (Valenciennes in Cuv. & Val. 1844), a valid species of cyprinid fish from Iran (Teleostei: Cyprinidae)." url="10.1080/09397140.1994.10637662" volume="10" year="1994">Coad and Krupp 1994</bibRefCitation>
|
|
] and this study (Tables 4-9)).
|
|
<taxonomicName id="9FEE5C4062B7821BB865B014C8F9349B" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="10" pageNumber="165" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
is distinguished from
|
|
<taxonomicName id="CCB4F7CD7BCEA25B86619F4AE2FF93C2" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta fusca" order="Cypriniformes" pageId="10" pageNumber="165" phylum="Chordata" rank="species" species="fusca">Capoeta fusca</taxonomicName>
|
|
by more total vertebrae (45-47 vs. 44), and more total lateral-line scales (70-84 vs. 40-62) (see
|
|
<bibRefCitation id="D3919439BE705DC2559EB6E56001FAE3" author="Coad, BW" journalOrPublisher="Shabpareh Publications, Tehran" pageId="15" pageNumber="170" title="Fishes of Tehran Province and adjacent areas." year="2008">Coad 2008</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="0FD1A1E76E41F1A624F5472E36A4E0C5" author="Johari, SA" journalOrPublisher="Zoology in the Middle East" pageId="16" pageNumber="171" pagination="63 - 70" title="Biological and morphometric characteristics of Capoetafusca, a cyprinid fish living in the qanats of south Khorasan, Iran (Osteichthyes: Cyprinidae)." url="10.1080/09397140.2009.10638348" volume="47" year="2009">Johari et al. 2009</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="5B1C5DFC76159189617AE55B0607E150" pageId="10" pageNumber="165">
|
|
<taxonomicName id="65FC5068D7D3BBA8A8483515C647B4C7" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="10" pageNumber="165" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
differs from its sister species (see Figs 6-7),
|
|
<taxonomicName id="27289C3C3AF818FE32DF02E2B8CD5AF7" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta buhsei" order="Cypriniformes" pageId="10" pageNumber="165" phylum="Chordata" rank="species" species="buhsei">Capoeta buhsei</taxonomicName>
|
|
in having more gill rakers on lower limb of first gill arch (10-13 vs. 8-10), more gill rakers on the whole first gill arch (14-18 vs. 12-14, see Tables 8-9) and by depth of caudal peduncle in percent of standard length (10.03-11.61 vs. 8.58-10.84).
|
|
<taxonomicName id="57DC455D2107FCD11644311B733F36C3" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta coadi" order="Cypriniformes" pageId="10" pageNumber="165" phylum="Chordata" rank="species" species="coadi">Capoeta coadi</taxonomicName>
|
|
is distinguished from another closely related species,
|
|
<taxonomicName id="72D852227E71F92C16D063E08F2B572C" family="Cyprinidae" genus="Capoeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Capoeta saadii" order="Cypriniformes" pageId="10" pageNumber="165" phylum="Chordata" rank="species" species="saadii">Capoeta saadii</taxonomicName>
|
|
by having more scales below the lateral line (9-11 vs. 6-10, modally 9) (Table 4) and more circum-pendicular scales (26-32 vs. 23-28, modally 25-26) [data from
|
|
<bibRefCitation id="F4B3DDA4D7D8FB686B6DBFDF5D801EB7" author="Alwan, N" journalOrPublisher="Goethe Universitaet, Frankfurt a. M., Germany" pageId="15" pageNumber="170" title="Systematics, taxonomy, phylogeny and zoogeography of the Capoetadamascina species complex (Pisces: Teleostei: Cyprinidae) inferred from comparative morphology and molecular markers" year="2011">Alwan (2011)</bibRefCitation>
|
|
].
|
|
</paragraph>
|
|
<caption id="1912AFEBE447A6CED5F8BE5959D14046" pageId="10" pageNumber="165">
|
|
<paragraph id="3C1EABDC174C8E6CCB9C72B7576341C9" pageId="10" pageNumber="165">Figure 6. Bayesian tree inferred from cyt b. Numbers left of the slash, indicate the posterior probabilities of the Bayesian analysis, using MrBayes, while numbers right of the slash are the bootstrap support for 10,000 replicates in the Maximum Likelihood tree, using RaxML. Asterisks (*) indicate less than 50% Maximum Likelihood support for the node.</paragraph>
|
|
</caption>
|
|
<caption id="CA4567EB21B9AED5FFB32C285F4E4C1E" pageId="10" pageNumber="165">
|
|
<paragraph id="954DD4E21570734D0FFCAE0D171AFCB5" pageId="10" pageNumber="165">Figure 7. Bayesian tree inferred from COI. Numbers left of the slash, indicate the posterior probabilities of the Bayesian analysis, using MrBayes, while numbers right of the slash are the bootstrap support for 10,000 replicates in the Maximum Likelihood tree, using RaxML. Asterisks (*) indicate less than 50% Maximum Likelihood support and (-) indicates less than 0.50 Bayesian posterior probabilities for the node.</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |