496 lines
70 KiB
XML
496 lines
70 KiB
XML
<document ID-DOI="10.11646/zootaxa.5256.5.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7758872" ID-ZooBank="E5B9708E-D626-4923-825A-A9FD59AE4024" checkinTime="1679468565563" checkinUser="plazi" docAuthor="Matalin, Andrey V. & Wiesner, Jürgen" docDate="2023" docId="050B87D78D4E247AE1B81A68FBB4771C" docLanguage="en" docName="zootaxa.5256.5.1.pdf" docOrigin="Zootaxa 5256 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5256.5.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Therates hunanensis Matalin et Wiesner 2023, sp. n." docType="treatment" docVersion="3" lastPageNumber="421" masterDocId="F932FFAF8D40246EE12F1976FFC07601" masterDocTitle="Revision of the Therates fruhstorferi complex (Coleoptera, Cicindelidae)" masterLastPageNumber="433" masterPageNumber="401" pageNumber="415" updateTime="1679505860750" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the Therates fruhstorferi complex (Coleoptera, Cicindelidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Matalin, Andrey V.</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0002-7790-8709</mods:nameIdentifier>
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<mods:affiliation>Moscow State Pedagogical University, Department of Zoology & Ecology, Kibalchicha str. 6, build. 3, Moscow 129164, Russia. andrei-matalin @ yandex. ru; https: // orcid. org / 0000 - 0002 - 7790 - 8709 & Pirogov National Research Medical University, Department of Biology, Ostrovitianova str. 1, Moscow 117997, Russia.</mods:affiliation>
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<mods:nameIdentifier type="email">andrei-matalin@yandex.ru</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wiesner, Jürgen</mods:namePart>
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<mods:affiliation>Dresdener Ring 11, D- 38444 Wolfsburg, Germany.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2023</mods:date>
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||
<mods:detail type="pubDate">
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||
<mods:number>2023-03-22</mods:number>
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</mods:detail>
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||
<mods:detail type="volume">
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<mods:number>5256</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>5</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>401</mods:start>
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<mods:end>433</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5256.5.1</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5256.5.1</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7758872</mods:identifier>
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<mods:identifier type="ZooBank">E5B9708E-D626-4923-825A-A9FD59AE4024</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:050B87D78D4E247AE1B81A68FBB4771C" httpUri="http://treatment.plazi.org/id/050B87D78D4E247AE1B81A68FBB4771C" lastPageId="20" lastPageNumber="421" pageId="14" pageNumber="415">
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<subSubSection box="[151,730,798,825]" pageId="14" pageNumber="415" type="nomenclature">
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<paragraph blockId="14.[151,730,798,861]" box="[151,730,798,825]" pageId="14" pageNumber="415">
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<heading bold="true" box="[151,730,798,825]" fontSize="11" level="1" pageId="14" pageNumber="415" reason="1">
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<emphasis bold="true" box="[151,730,798,825]" pageId="14" pageNumber="415">
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<taxonomicName authority="Matalin et Wiesner" authorityName="Matalin et Wiesner" authorityYear="2023" box="[151,651,798,825]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="415" phylum="Arthropoda" rank="species" species="hunanensis" status="sp. n.">
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<emphasis bold="true" box="[151,404,798,825]" italics="true" pageId="14" pageNumber="415">Therates hunanensis</emphasis>
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Matalin et Wiesner
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</taxonomicName>
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,
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<taxonomicNameLabel box="[664,730,799,825]" pageId="14" pageNumber="415" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,634,834,861]" pageId="14" pageNumber="415" type="description">
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<paragraph blockId="14.[151,730,798,861]" box="[151,634,834,861]" pageId="14" pageNumber="415">
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<figureCitation box="[151,236,834,861]" captionStart="FIGURES 15–28" captionStartId="9.[152,270,1641,1665]" captionTargetBox="[156,1418,712,1612]" captionTargetId="figure-217@9.[151,1435,709,1617]" captionTargetPageId="9" captionText="FIGURES 15–28. Therates spp., habitus: 15–18—T. vitalisi vitalisi (15, 16—Vietnam, Lao Cai; 17—Vietnam, Fan Si Pan Mnt; 18—Vietnam, Sa Pa); 19–21—T. vitalisi ida (China, Kuatun); 22–24—T. biserratus (22, 23—China, Yinjiang County; 24—China, Fanjing Shan-Kuaichang); 25—T. hunnanensis sp. n. (China, Shunhuangshan); 26–28—T. sauteri (26, 27—China, Tiawan, Taihorin; 28—China, Taiwan, Ilan Hsien); 17–20, 24–28—males; 15, 16, 21–23 —females; 15, 16, 19, 20, 25— holotypes; 22, 23—paratype; 26, 27—lectotype; 15, 17–19, 21, 22, 24–26, 28—habitus; 16, 20, 23, 27—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758876" httpUri="https://zenodo.org/record/7758876/files/figure.png" pageId="14" pageNumber="415">Figs 25</figureCitation>
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,
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<figureCitation box="[248,320,835,861]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="14" pageNumber="415">42, 43</figureCitation>
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,
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<figureCitation box="[332,405,834,861]" captionStart="FIGURES 73–80" captionStartId="18.[152,270,1909,1933]" captionTargetBox="[302,1276,200,1868]" captionTargetId="figure-28@18.[296,1292,181,1885]" captionTargetPageId="18" captionText="FIGURES 73–80. Therates spp. labrum: 73, 74—T. biserratus (73—China, Yinjiang County; 74—China, Fanjing Shan- Kuaichang); 75, 76—T. hunnanensis sp. n. (China, Shunhuangshan); 77–80—T. sauteri (77, 78 —China, Taiwan, Taihorin; 79—China, Taiwan, Ilan Hsien; 80—China, Taiwan, Taipei Hsien); 74, 75, 77, 79—males; 73, 76, 78, 80—females; 75— holotype; 73, 76—paratype; 77, 78—paralectotypes." figureDoi="http://doi.org/10.5281/zenodo.7758890" httpUri="https://zenodo.org/record/7758890/files/figure.png" pageId="14" pageNumber="415">75, 76</figureCitation>
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,
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<figureCitation box="[416,516,835,861]" captionStart="FIGURES 99–114" captionStartId="21.[152,273,1418,1442]" captionTargetBox="[155,1421,210,1377]" captionTargetId="figure-17@21.[151,1435,181,1394]" captionTargetPageId="21" captionText="FIGURES 99–114. Therates spp., left elytron: 99–102—T. vitalisi vitalisi (99 —Vietnam, Lao Cai; 100, 101—Vietnam, Sa Pa; 102—Laos, 15 km NW Louang Namtha); 103, 104—T. vitalisi ida (China, Kuatun); 105, 106—T. biserratus (105— China, Yinjiang County; 106—China, Fanjing Shan-Kuaichang); 107, 108—T. hunnanensis sp. n. (China, Shunhuangshan); 109–114—T. sauteri (109, 110, 112, 113—China, Taiwan, Taihorin; 111—China, Taiwan, Ilan Hsien; 114—China, Taiwan, Taipei Hsien); 100, 103, 106, 107, 109–111—males; 99, 101, 102, 104, 105, 108, 112–114—females; 99, 103, 107— holotypes; 105, 108—paratypes; 109, 110, 112, 113—paralectotypes" figureDoi="http://doi.org/10.5281/zenodo.7758896" httpUri="https://zenodo.org/record/7758896/files/figure.png" pageId="14" pageNumber="415">107, 108</figureCitation>
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,
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<figureCitation box="[528,630,835,861]" captionStart="FIGURES 152–174" captionStartId="24.[152,270,1210,1234]" captionTargetBox="[163,1428,181,1165]" captionTargetId="figure-28@24.[151,1435,181,1186]" captionTargetPageId="24" captionText="FIGURES 152–174. Therates spp., aedeagus: 152–166—T. vitalisi vitalisi (152, 153—Vietnam, Xuan Son N.P.; 154— Vietnam, Sa Pa; 155, 156—Laos, Ban Nape—Kaev Nua pass; 157–163—Laos, 15–20 km NW Louang Namtha; 164, 165— Laos, 8 km NE Ban Nape; 166—Laos, Thong Khan); 167—T. vitalisi ida (China, Kuatun); 168—T. biserratus (China, Fanjing Shan-Kuaichang); 169–171—T. hunnanensis sp. n. (China, Shunhuangshan); 172—T. motoensis (China, Hanmi); 173, 174—T. sauteri (173—China, Taiwan, Taihorin; 174—China, Taiwan, Ilan Hsien); 167, 169—holotypes; 170, 171—paratypes; 173— lectotype; 152—dorsal view; 153–174—left lateral view; ac—apical carinae; sp—spring; fl—flagellum. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758906" httpUri="https://zenodo.org/record/7758906/files/figure.png" pageId="14" pageNumber="415">169–171</figureCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection box="[151,820,903,927]" pageId="14" pageNumber="415" type="reference_group">
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||
<paragraph blockId="14.[151,820,903,927]" box="[151,820,903,927]" pageId="14" pageNumber="415">
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||
<treatmentCitationGroup box="[151,820,903,927]" pageId="14" pageNumber="415">
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||
<taxonomicName ID-CoL="5F7VM" authority="W. Horn, 1902" authorityName="W. Horn" authorityYear="1902" box="[151,639,903,927]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="415" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subSpecies="fruhstorferi">
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<emphasis box="[151,486,903,927]" italics="true" pageId="14" pageNumber="415">Therates fruhstorferi fruhstorferi</emphasis>
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W.
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<bibRefCitation author="Horn, W." box="[526,639,903,927]" pageId="14" pageNumber="415" pagination="65 - 75" refId="ref25658" refString="Horn, W. (1902) Neue Cicindeliden gesammelt von Fruhstorfer in Tonkin 1900. Deutsche Entomologische Zeitschrift, 1902, 65 - 75." type="journal article" year="1902">Horn, 1902</bibRefCitation>
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||
</taxonomicName>
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||
—
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||
<treatmentCitation author="Tan, J. - J." box="[666,816,903,927]" page="257" pageId="14" pageNumber="415" year="1993">
|
||
<bibRefCitation author="Tan, J. - J." box="[666,816,903,927]" pageId="14" pageNumber="415" pagination="257 - 258" refId="ref27248" refString="Tan, J. - J. (1993) Coleoptera: Cicindelidae. In: Huang, F. - S. (Ed.), Insects of Wuling mountains area, southeastern China. Science Press, Beijing, pp. 257 - 258 [in Chinese, with English summary]" type="book chapter" year="1993">Tan 1993: 257</bibRefCitation>
|
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</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="14" pageNumber="415" type="materials_examined">
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||
<paragraph blockId="14.[151,1437,973,2007]" pageId="14" pageNumber="415">
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||
<emphasis bold="true" box="[151,331,973,999]" pageId="14" pageNumber="415">Type material.</emphasis>
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||
<materialsCitation collectingDate="2013-06-20" collectionCode="JW" collectorName="Jatua" country="China" elevation="1100" latitude="26.22" location="SHUNHUANGSHAN" longitude="-111.01" pageId="14" pageNumber="415" specimenCount="1" stateProvince="Hunan" typeStatus="holotype">
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||
<typeStatus box="[343,461,973,999]" pageId="14" pageNumber="415">HOLOTYPE</typeStatus>
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||
,
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||
<specimenCount box="[481,500,975,1002]" pageId="14" pageNumber="415" type="male">♂</specimenCount>
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||
—
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||
<collectingCountry box="[530,599,975,1002]" name="China" pageId="14" pageNumber="415">China</collectingCountry>
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, S
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||
<collectingRegion box="[645,722,974,1000]" country="China" name="Hunan" pageId="14" pageNumber="415">Hunan</collectingRegion>
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,
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<quantity box="[739,888,973,1000]" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.3" metricValueMin="0.9" pageId="14" pageNumber="415" unit="m" value="1100.0" valueMax="1300.0" valueMin="900.0">
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<elevation box="[739,888,973,1000]" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.3" metricValueMin="0.9" pageId="14" pageNumber="415" unit="m" value="1100.0" valueMax="1300.0" valueMin="900.0">900–1300 m</elevation>
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</quantity>
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,
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<location LSID="urn:lsid:plazi:treatment:050B87D78D4E247AE1B81A68FBB4771C:887D601A8D4E2460E2A51ABBFB5375E6" box="[906,1171,973,999]" country="China" name="SHUNHUANGSHAN" pageId="14" pageNumber="415" stateProvince="Hunan">SHUNHUANGSHAN</location>
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for. park,
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<geoCoordinate box="[1305,1436,973,999]" degrees="26" direction="north" minutes="22" orientation="latitude" pageId="14" pageNumber="415" value="26.22">
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26
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22-24’N
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<geoCoordinate box="[151,294,1009,1035]" degrees="111" direction="east" minutes="01" orientation="longitude" pageId="14" pageNumber="415" value="-111.01">
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111
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<superScript attach="none" box="[191,199,1009,1023]" fontSize="6" pageId="14" pageNumber="415">o</superScript>
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01-02’E
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</geoCoordinate>
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,
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<date box="[306,436,1009,1035]" pageId="14" pageNumber="415" value="2013-06-20">
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<collectingDate box="[306,436,1009,1035]" pageId="14" pageNumber="415" value="2013-06-20">20.VI.2013</collectingDate>
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</date>
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,
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<collectorName box="[448,506,1010,1036]" pageId="14" pageNumber="415">Jatua</collectorName>
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leg. (
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<collectionCode box="[570,613,1009,1035]" collectionName="Jan Wieringa" pageId="14" pageNumber="415">JW</collectionCode>
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||
)
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||
</materialsCitation>
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||
;
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||
<materialsCitation collectionCode="JW, MSPU" country="China" pageId="14" pageNumber="415" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Hunan" typeStatus="paratype">
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||
<typeStatus box="[633,750,1009,1035]" pageId="14" pageNumber="415">PARATYPES</typeStatus>
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||
,
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||
<specimenCount box="[764,822,1011,1038]" pageId="14" pageNumber="415" type="male">2♂♂</specimenCount>
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<specimenCount box="[822,845,1011,1038]" pageId="14" pageNumber="415" type="female">1♀</specimenCount>
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—same labeled as the holotype (
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<specimenCount box="[1223,1256,1011,1038]" pageId="14" pageNumber="415" type="male">1♂</specimenCount>
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<specimenCount box="[1256,1281,1011,1038]" pageId="14" pageNumber="415" type="female">1♀</specimenCount>
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||
—
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||
<collectionCode box="[1315,1361,1011,1038]" collectionName="Jan Wieringa" pageId="14" pageNumber="415">JW</collectionCode>
|
||
;
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||
<specimenCount box="[1373,1403,1011,1038]" pageId="14" pageNumber="415" type="male">1♂</specimenCount>
|
||
—
|
||
<collectionCode box="[151,232,1045,1071]" collectionName="MSPU" pageId="14" pageNumber="415">MSPU</collectionCode>
|
||
)
|
||
</materialsCitation>
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||
.
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||
</paragraph>
|
||
<paragraph blockId="14.[151,1437,973,2007]" pageId="14" pageNumber="415">
|
||
<emphasis bold="true" box="[199,445,1081,1107]" pageId="14" pageNumber="415">Additional material.</emphasis>
|
||
<materialsCitation collectingDate="1998-08-12" collectionCode="XX" collectorName="Wang Shu Yong" country="China" elevation="1400" location="Tiang Ping Shan" pageId="14" pageNumber="415" specimenCount="1" specimenCount-male="1" stateProvince="Hunan Prov.">
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||
<collectingCountry box="[455,531,1081,1107]" name="China" pageId="14" pageNumber="415">CHINA</collectingCountry>
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||
,
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||
<emphasis box="[546,691,1082,1107]" italics="true" pageId="14" pageNumber="415">
|
||
<collectingRegion box="[546,691,1082,1107]" country="China" name="Hunan" pageId="14" pageNumber="415">Hunan Prov.</collectingRegion>
|
||
</emphasis>
|
||
:
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||
<specimenCount box="[708,737,1083,1110]" pageId="14" pageNumber="415" type="male">1♂</specimenCount>
|
||
—
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||
<location LSID="urn:lsid:plazi:treatment:050B87D78D4E247AE1B81A68FBB4771C:887D601A8D4E2460E3D01D4DFC077252" box="[767,967,1081,1110]" country="China" name="Tiang Ping Shan" pageId="14" pageNumber="415" stateProvince="Hunan Prov.">Tiang Ping Shan</location>
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||
,
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||
<date box="[981,1130,1081,1107]" pageId="14" pageNumber="415" value="1998-08-12">
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||
<collectingDate box="[981,1130,1081,1107]" pageId="14" pageNumber="415" value="1998-08-12">12.VIII.1998</collectingDate>
|
||
</date>
|
||
,
|
||
<quantity box="[1144,1235,1081,1108]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="14" pageNumber="415" unit="m" value="1400.0">
|
||
<elevation box="[1144,1235,1081,1108]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="14" pageNumber="415" unit="m" value="1400.0">1400 m</elevation>
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||
</quantity>
|
||
,
|
||
<collectorName box="[1249,1436,1081,1108]" pageId="14" pageNumber="415">Wang Shu Yong</collectorName>
|
||
(
|
||
<collectionCode box="[160,206,1117,1143]" pageId="14" pageNumber="415">XX</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="14" pageNumber="415" type="reference_group">
|
||
<paragraph blockId="14.[151,1437,973,2007]" pageId="14" pageNumber="415">
|
||
<emphasis bold="true" box="[199,336,1153,1179]" pageId="14" pageNumber="415">References.</emphasis>
|
||
<materialsCitation country="China" county="Sangzhi County" elevation="1470" location="Tianping Mt." pageId="14" pageNumber="415" specimenCount="1" stateProvince="Hunan">
|
||
<collectingCountry box="[355,428,1153,1179]" name="China" pageId="14" pageNumber="415">CHINA</collectingCountry>
|
||
,
|
||
<emphasis box="[454,608,1154,1179]" italics="true" pageId="14" pageNumber="415">
|
||
<collectingRegion box="[454,608,1154,1179]" country="China" name="Hunan" pageId="14" pageNumber="415">Hunan Prov.</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<collectingCounty box="[634,827,1153,1179]" pageId="14" pageNumber="415">Sangzhi County</collectingCounty>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:050B87D78D4E247AE1B81A68FBB4771C:887D601A8D4E2460E27E1DF7FC33729D" box="[849,1011,1153,1180]" country="China" county="Sangzhi County" name="Tianping Mt." pageId="14" pageNumber="415" stateProvince="Hunan">Tianping Mt.</location>
|
||
,
|
||
<quantity box="[1034,1201,1153,1180]" metricMagnitude="3" metricUnit="m" metricValue="1.47" metricValueMax="1.5699999999999998" metricValueMin="1.3699999999999999" pageId="14" pageNumber="415" unit="m" value="1470.0" valueMax="1570.0" valueMin="1370.0">
|
||
<elevation box="[1034,1201,1153,1180]" metricMagnitude="3" metricUnit="m" metricValue="1.47" metricValueMax="1.5699999999999998" metricValueMin="1.3699999999999999" pageId="14" pageNumber="415" unit="m" value="1470.0" valueMax="1570.0" valueMin="1370.0">1370–1570 m</elevation>
|
||
</quantity>
|
||
(
|
||
<bibRefCitation author="Tan, J. - J." box="[1227,1345,1153,1179]" pageId="14" pageNumber="415" pagination="257 - 258" refId="ref27248" refString="Tan, J. - J. (1993) Coleoptera: Cicindelidae. In: Huang, F. - S. (Ed.), Insects of Wuling mountains area, southeastern China. Science Press, Beijing, pp. 257 - 258 [in Chinese, with English summary]" type="book chapter" year="1993">Tan 1993</bibRefCitation>
|
||
—as
|
||
<taxonomicName authority="W. Horn, 1902" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="415" phylum="Arthropoda" rank="species" species="fruhstorferi">
|
||
<emphasis italics="true" pageId="14" pageNumber="415">T. fruhstorferi</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="14" pageNumber="415" type="diagnosis">
|
||
<paragraph blockId="14.[151,1437,973,2007]" pageId="14" pageNumber="415">
|
||
<emphasis bold="true" box="[199,322,1225,1251]" pageId="14" pageNumber="415">Diagnosis.</emphasis>
|
||
From related
|
||
<taxonomicName authorityName="W. Horn" authorityYear="1913" box="[477,580,1225,1251]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="415" phylum="Arthropoda" rank="species" species="vitalisi">
|
||
<emphasis box="[477,580,1225,1251]" italics="true" pageId="14" pageNumber="415">T. vitalisi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="W. Horn" authorityYear="1912" box="[593,698,1226,1251]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="415" phylum="Arthropoda" rank="species" species="sauteri">
|
||
<emphasis box="[593,698,1226,1251]" italics="true" pageId="14" pageNumber="415">T. sauteri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Matalin & Wiesner" authorityYear="2023" box="[750,891,1225,1251]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="415" phylum="Arthropoda" rank="species" species="biserratus">
|
||
<emphasis box="[750,891,1225,1251]" italics="true" pageId="14" pageNumber="415">T. biserratus</emphasis>
|
||
</taxonomicName>
|
||
a new species is easily recognised by the smaller size (mean)—
|
||
<quantity box="[309,420,1261,1288]" metricMagnitude="-2" metricUnit="m" metricValue="1.093" pageId="14" pageNumber="415" unit="mm" value="10.93">10.93 mm</quantity>
|
||
<emphasis box="[426,456,1262,1287]" italics="true" pageId="14" pageNumber="415">vs.</emphasis>
|
||
<quantity box="[462,574,1261,1288]" metricMagnitude="-2" metricUnit="m" metricValue="1.244" pageId="14" pageNumber="415" unit="mm" value="12.44">12.44 mm</quantity>
|
||
in
|
||
<taxonomicName authorityName="W. Horn" authorityYear="1913" box="[607,709,1261,1287]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="415" phylum="Arthropoda" rank="species" species="vitalisi">
|
||
<emphasis box="[607,709,1261,1287]" italics="true" pageId="14" pageNumber="415">T. vitalisi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity box="[722,820,1261,1288]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="14" pageNumber="415" unit="mm" value="11.8">11.8 mm</quantity>
|
||
in
|
||
<taxonomicName authorityName="W. Horn" authorityYear="1912" box="[853,957,1262,1287]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="415" phylum="Arthropoda" rank="species" species="sauteri">
|
||
<emphasis box="[853,957,1262,1287]" italics="true" pageId="14" pageNumber="415">T. sauteri</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<quantity box="[1015,1113,1261,1288]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="14" pageNumber="415" unit="mm" value="11.1">11.1 mm</quantity>
|
||
in
|
||
<taxonomicName authorityName="Matalin & Wiesner" authorityYear="2023" box="[1146,1286,1261,1287]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="415" phylum="Arthropoda" rank="species" species="biserratus">
|
||
<emphasis box="[1146,1286,1261,1287]" italics="true" pageId="14" pageNumber="415">T. biserratus</emphasis>
|
||
</taxonomicName>
|
||
, by distinctly striated orbital plates in posterior third, by the shape of antennomeres 9–11 (
|
||
<figureCitation box="[1019,1148,1297,1323]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="14" pageNumber="415">Figs 42, 43</figureCitation>
|
||
<emphasis box="[1156,1187,1298,1323]" italics="true" pageId="14" pageNumber="415">vs.</emphasis>
|
||
<figureCitation box="[1195,1325,1297,1323]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="14" pageNumber="415">Figs 44–49</figureCitation>
|
||
), by wide apical dot (
|
||
<figureCitation box="[282,372,1333,1360]" captionStart="FIGURES 15–28" captionStartId="9.[152,270,1641,1665]" captionTargetBox="[156,1418,712,1612]" captionTargetId="figure-217@9.[151,1435,709,1617]" captionTargetPageId="9" captionText="FIGURES 15–28. Therates spp., habitus: 15–18—T. vitalisi vitalisi (15, 16—Vietnam, Lao Cai; 17—Vietnam, Fan Si Pan Mnt; 18—Vietnam, Sa Pa); 19–21—T. vitalisi ida (China, Kuatun); 22–24—T. biserratus (22, 23—China, Yinjiang County; 24—China, Fanjing Shan-Kuaichang); 25—T. hunnanensis sp. n. (China, Shunhuangshan); 26–28—T. sauteri (26, 27—China, Tiawan, Taihorin; 28—China, Taiwan, Ilan Hsien); 17–20, 24–28—males; 15, 16, 21–23 —females; 15, 16, 19, 20, 25— holotypes; 22, 23—paratype; 26, 27—lectotype; 15, 17–19, 21, 22, 24–26, 28—habitus; 16, 20, 23, 27—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758876" httpUri="https://zenodo.org/record/7758876/files/figure.png" pageId="14" pageNumber="415">Figs 25</figureCitation>
|
||
,
|
||
<figureCitation box="[387,488,1333,1359]" captionStart="FIGURES 99–114" captionStartId="21.[152,273,1418,1442]" captionTargetBox="[155,1421,210,1377]" captionTargetId="figure-17@21.[151,1435,181,1394]" captionTargetPageId="21" captionText="FIGURES 99–114. Therates spp., left elytron: 99–102—T. vitalisi vitalisi (99 —Vietnam, Lao Cai; 100, 101—Vietnam, Sa Pa; 102—Laos, 15 km NW Louang Namtha); 103, 104—T. vitalisi ida (China, Kuatun); 105, 106—T. biserratus (105— China, Yinjiang County; 106—China, Fanjing Shan-Kuaichang); 107, 108—T. hunnanensis sp. n. (China, Shunhuangshan); 109–114—T. sauteri (109, 110, 112, 113—China, Taiwan, Taihorin; 111—China, Taiwan, Ilan Hsien; 114—China, Taiwan, Taipei Hsien); 100, 103, 106, 107, 109–111—males; 99, 101, 102, 104, 105, 108, 112–114—females; 99, 103, 107— holotypes; 105, 108—paratypes; 109, 110, 112, 113—paralectotypes" figureDoi="http://doi.org/10.5281/zenodo.7758896" httpUri="https://zenodo.org/record/7758896/files/figure.png" pageId="14" pageNumber="415">107, 108</figureCitation>
|
||
<emphasis box="[499,529,1334,1359]" italics="true" pageId="14" pageNumber="415">vs.</emphasis>
|
||
<figureCitation box="[539,941,1333,1360]" captionStart="FIGURES 15–28" captionStartId="9.[152,270,1641,1665]" captionTargetBox="[156,1418,712,1612]" captionTargetId="figure-217@9.[151,1435,709,1617]" captionTargetPageId="9" captionText="FIGURES 15–28. Therates spp., habitus: 15–18—T. vitalisi vitalisi (15, 16—Vietnam, Lao Cai; 17—Vietnam, Fan Si Pan Mnt; 18—Vietnam, Sa Pa); 19–21—T. vitalisi ida (China, Kuatun); 22–24—T. biserratus (22, 23—China, Yinjiang County; 24—China, Fanjing Shan-Kuaichang); 25—T. hunnanensis sp. n. (China, Shunhuangshan); 26–28—T. sauteri (26, 27—China, Tiawan, Taihorin; 28—China, Taiwan, Ilan Hsien); 17–20, 24–28—males; 15, 16, 21–23 —females; 15, 16, 19, 20, 25— holotypes; 22, 23—paratype; 26, 27—lectotype; 15, 17–19, 21, 22, 24–26, 28—habitus; 16, 20, 23, 27—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758876" httpUri="https://zenodo.org/record/7758876/files/figure.png" pageId="14" pageNumber="415">Figs 15, 17–19, 21, 22, 24, 26, 28</figureCitation>
|
||
,
|
||
<figureCitation box="[956,1158,1333,1359]" captionStart="FIGURES 99–114" captionStartId="21.[152,273,1418,1442]" captionTargetBox="[155,1421,210,1377]" captionTargetId="figure-17@21.[151,1435,181,1394]" captionTargetPageId="21" captionText="FIGURES 99–114. Therates spp., left elytron: 99–102—T. vitalisi vitalisi (99 —Vietnam, Lao Cai; 100, 101—Vietnam, Sa Pa; 102—Laos, 15 km NW Louang Namtha); 103, 104—T. vitalisi ida (China, Kuatun); 105, 106—T. biserratus (105— China, Yinjiang County; 106—China, Fanjing Shan-Kuaichang); 107, 108—T. hunnanensis sp. n. (China, Shunhuangshan); 109–114—T. sauteri (109, 110, 112, 113—China, Taiwan, Taihorin; 111—China, Taiwan, Ilan Hsien; 114—China, Taiwan, Taipei Hsien); 100, 103, 106, 107, 109–111—males; 99, 101, 102, 104, 105, 108, 112–114—females; 99, 103, 107— holotypes; 105, 108—paratypes; 109, 110, 112, 113—paralectotypes" figureDoi="http://doi.org/10.5281/zenodo.7758896" httpUri="https://zenodo.org/record/7758896/files/figure.png" pageId="14" pageNumber="415">99–106, 109–114</figureCitation>
|
||
), by pale hind tibia and coloration of hind tarsi (
|
||
<figureCitation box="[427,509,1369,1396]" captionStart="FIGURES 15–28" captionStartId="9.[152,270,1641,1665]" captionTargetBox="[156,1418,712,1612]" captionTargetId="figure-217@9.[151,1435,709,1617]" captionTargetPageId="9" captionText="FIGURES 15–28. Therates spp., habitus: 15–18—T. vitalisi vitalisi (15, 16—Vietnam, Lao Cai; 17—Vietnam, Fan Si Pan Mnt; 18—Vietnam, Sa Pa); 19–21—T. vitalisi ida (China, Kuatun); 22–24—T. biserratus (22, 23—China, Yinjiang County; 24—China, Fanjing Shan-Kuaichang); 25—T. hunnanensis sp. n. (China, Shunhuangshan); 26–28—T. sauteri (26, 27—China, Tiawan, Taihorin; 28—China, Taiwan, Ilan Hsien); 17–20, 24–28—males; 15, 16, 21–23 —females; 15, 16, 19, 20, 25— holotypes; 22, 23—paratype; 26, 27—lectotype; 15, 17–19, 21, 22, 24–26, 28—habitus; 16, 20, 23, 27—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758876" httpUri="https://zenodo.org/record/7758876/files/figure.png" pageId="14" pageNumber="415">Fig. 25</figureCitation>
|
||
<emphasis box="[517,548,1370,1395]" italics="true" pageId="14" pageNumber="415">vs.</emphasis>
|
||
<figureCitation box="[556,944,1369,1396]" captionStart="FIGURES 15–28" captionStartId="9.[152,270,1641,1665]" captionTargetBox="[156,1418,712,1612]" captionTargetId="figure-217@9.[151,1435,709,1617]" captionTargetPageId="9" captionText="FIGURES 15–28. Therates spp., habitus: 15–18—T. vitalisi vitalisi (15, 16—Vietnam, Lao Cai; 17—Vietnam, Fan Si Pan Mnt; 18—Vietnam, Sa Pa); 19–21—T. vitalisi ida (China, Kuatun); 22–24—T. biserratus (22, 23—China, Yinjiang County; 24—China, Fanjing Shan-Kuaichang); 25—T. hunnanensis sp. n. (China, Shunhuangshan); 26–28—T. sauteri (26, 27—China, Tiawan, Taihorin; 28—China, Taiwan, Ilan Hsien); 17–20, 24–28—males; 15, 16, 21–23 —females; 15, 16, 19, 20, 25— holotypes; 22, 23—paratype; 26, 27—lectotype; 15, 17–19, 21, 22, 24–26, 28—habitus; 16, 20, 23, 27—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758876" httpUri="https://zenodo.org/record/7758876/files/figure.png" pageId="14" pageNumber="415">Figs 15, 17–19, 21, 22, 24, 26, 28</figureCitation>
|
||
), as well as by the shape of aedeagus (
|
||
<figureCitation captionStart="FIGURES 152–174" captionStartId="24.[152,270,1210,1234]" captionTargetBox="[163,1428,181,1165]" captionTargetId="figure-28@24.[151,1435,181,1186]" captionTargetPageId="24" captionText="FIGURES 152–174. Therates spp., aedeagus: 152–166—T. vitalisi vitalisi (152, 153—Vietnam, Xuan Son N.P.; 154— Vietnam, Sa Pa; 155, 156—Laos, Ban Nape—Kaev Nua pass; 157–163—Laos, 15–20 km NW Louang Namtha; 164, 165— Laos, 8 km NE Ban Nape; 166—Laos, Thong Khan); 167—T. vitalisi ida (China, Kuatun); 168—T. biserratus (China, Fanjing Shan-Kuaichang); 169–171—T. hunnanensis sp. n. (China, Shunhuangshan); 172—T. motoensis (China, Hanmi); 173, 174—T. sauteri (173—China, Taiwan, Taihorin; 174—China, Taiwan, Ilan Hsien); 167, 169—holotypes; 170, 171—paratypes; 173— lectotype; 152—dorsal view; 153–174—left lateral view; ac—apical carinae; sp—spring; fl—flagellum. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758906" httpUri="https://zenodo.org/record/7758906/files/figure.png" pageId="14" pageNumber="415">Figs 169–171</figureCitation>
|
||
<emphasis box="[255,286,1406,1431]" italics="true" pageId="14" pageNumber="415">vs.</emphasis>
|
||
<figureCitation box="[292,559,1405,1432]" captionStart="FIGURES 152–174" captionStartId="24.[152,270,1210,1234]" captionTargetBox="[163,1428,181,1165]" captionTargetId="figure-28@24.[151,1435,181,1186]" captionTargetPageId="24" captionText="FIGURES 152–174. Therates spp., aedeagus: 152–166—T. vitalisi vitalisi (152, 153—Vietnam, Xuan Son N.P.; 154— Vietnam, Sa Pa; 155, 156—Laos, Ban Nape—Kaev Nua pass; 157–163—Laos, 15–20 km NW Louang Namtha; 164, 165— Laos, 8 km NE Ban Nape; 166—Laos, Thong Khan); 167—T. vitalisi ida (China, Kuatun); 168—T. biserratus (China, Fanjing Shan-Kuaichang); 169–171—T. hunnanensis sp. n. (China, Shunhuangshan); 172—T. motoensis (China, Hanmi); 173, 174—T. sauteri (173—China, Taiwan, Taihorin; 174—China, Taiwan, Ilan Hsien); 167, 169—holotypes; 170, 171—paratypes; 173— lectotype; 152—dorsal view; 153–174—left lateral view; ac—apical carinae; sp—spring; fl—flagellum. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758906" httpUri="https://zenodo.org/record/7758906/files/figure.png" pageId="14" pageNumber="415">Figs 152–168, 173, 174</figureCitation>
|
||
); additionally from
|
||
<taxonomicName authorityName="Matalin & Wiesner" authorityYear="2023" box="[779,920,1405,1431]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="415" phylum="Arthropoda" rank="species" species="biserratus">
|
||
<emphasis box="[779,920,1405,1431]" italics="true" pageId="14" pageNumber="415">T. biserratus</emphasis>
|
||
</taxonomicName>
|
||
a new species is clearly distinguished by wide apical dot extend the suture (
|
||
<figureCitation box="[474,561,1441,1468]" captionStart="FIGURES 15–28" captionStartId="9.[152,270,1641,1665]" captionTargetBox="[156,1418,712,1612]" captionTargetId="figure-217@9.[151,1435,709,1617]" captionTargetPageId="9" captionText="FIGURES 15–28. Therates spp., habitus: 15–18—T. vitalisi vitalisi (15, 16—Vietnam, Lao Cai; 17—Vietnam, Fan Si Pan Mnt; 18—Vietnam, Sa Pa); 19–21—T. vitalisi ida (China, Kuatun); 22–24—T. biserratus (22, 23—China, Yinjiang County; 24—China, Fanjing Shan-Kuaichang); 25—T. hunnanensis sp. n. (China, Shunhuangshan); 26–28—T. sauteri (26, 27—China, Tiawan, Taihorin; 28—China, Taiwan, Ilan Hsien); 17–20, 24–28—males; 15, 16, 21–23 —females; 15, 16, 19, 20, 25— holotypes; 22, 23—paratype; 26, 27—lectotype; 15, 17–19, 21, 22, 24–26, 28—habitus; 16, 20, 23, 27—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758876" httpUri="https://zenodo.org/record/7758876/files/figure.png" pageId="14" pageNumber="415">Figs 25</figureCitation>
|
||
,
|
||
<figureCitation box="[573,671,1441,1467]" captionStart="FIGURES 99–114" captionStartId="21.[152,273,1418,1442]" captionTargetBox="[155,1421,210,1377]" captionTargetId="figure-17@21.[151,1435,181,1394]" captionTargetPageId="21" captionText="FIGURES 99–114. Therates spp., left elytron: 99–102—T. vitalisi vitalisi (99 —Vietnam, Lao Cai; 100, 101—Vietnam, Sa Pa; 102—Laos, 15 km NW Louang Namtha); 103, 104—T. vitalisi ida (China, Kuatun); 105, 106—T. biserratus (105— China, Yinjiang County; 106—China, Fanjing Shan-Kuaichang); 107, 108—T. hunnanensis sp. n. (China, Shunhuangshan); 109–114—T. sauteri (109, 110, 112, 113—China, Taiwan, Taihorin; 111—China, Taiwan, Ilan Hsien; 114—China, Taiwan, Taipei Hsien); 100, 103, 106, 107, 109–111—males; 99, 101, 102, 104, 105, 108, 112–114—females; 99, 103, 107— holotypes; 105, 108—paratypes; 109, 110, 112, 113—paralectotypes" figureDoi="http://doi.org/10.5281/zenodo.7758896" httpUri="https://zenodo.org/record/7758896/files/figure.png" pageId="14" pageNumber="415">107, 108</figureCitation>
|
||
<emphasis box="[678,708,1442,1467]" italics="true" pageId="14" pageNumber="415">vs.</emphasis>
|
||
<figureCitation box="[715,842,1441,1467]" captionStart="FIGURES 15–28" captionStartId="9.[152,270,1641,1665]" captionTargetBox="[156,1418,712,1612]" captionTargetId="figure-217@9.[151,1435,709,1617]" captionTargetPageId="9" captionText="FIGURES 15–28. Therates spp., habitus: 15–18—T. vitalisi vitalisi (15, 16—Vietnam, Lao Cai; 17—Vietnam, Fan Si Pan Mnt; 18—Vietnam, Sa Pa); 19–21—T. vitalisi ida (China, Kuatun); 22–24—T. biserratus (22, 23—China, Yinjiang County; 24—China, Fanjing Shan-Kuaichang); 25—T. hunnanensis sp. n. (China, Shunhuangshan); 26–28—T. sauteri (26, 27—China, Tiawan, Taihorin; 28—China, Taiwan, Ilan Hsien); 17–20, 24–28—males; 15, 16, 21–23 —females; 15, 16, 19, 20, 25— holotypes; 22, 23—paratype; 26, 27—lectotype; 15, 17–19, 21, 22, 24–26, 28—habitus; 16, 20, 23, 27—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758876" httpUri="https://zenodo.org/record/7758876/files/figure.png" pageId="14" pageNumber="415">Figs 22, 24</figureCitation>
|
||
,
|
||
<figureCitation box="[854,956,1441,1468]" captionStart="FIGURES 99–114" captionStartId="21.[152,273,1418,1442]" captionTargetBox="[155,1421,210,1377]" captionTargetId="figure-17@21.[151,1435,181,1394]" captionTargetPageId="21" captionText="FIGURES 99–114. Therates spp., left elytron: 99–102—T. vitalisi vitalisi (99 —Vietnam, Lao Cai; 100, 101—Vietnam, Sa Pa; 102—Laos, 15 km NW Louang Namtha); 103, 104—T. vitalisi ida (China, Kuatun); 105, 106—T. biserratus (105— China, Yinjiang County; 106—China, Fanjing Shan-Kuaichang); 107, 108—T. hunnanensis sp. n. (China, Shunhuangshan); 109–114—T. sauteri (109, 110, 112, 113—China, Taiwan, Taihorin; 111—China, Taiwan, Ilan Hsien; 114—China, Taiwan, Taipei Hsien); 100, 103, 106, 107, 109–111—males; 99, 101, 102, 104, 105, 108, 112–114—females; 99, 103, 107— holotypes; 105, 108—paratypes; 109, 110, 112, 113—paralectotypes" figureDoi="http://doi.org/10.5281/zenodo.7758896" httpUri="https://zenodo.org/record/7758896/files/figure.png" pageId="14" pageNumber="415">105, 106</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="20" lastPageNumber="421" pageId="14" pageNumber="415" type="description">
|
||
<paragraph blockId="14.[151,1437,973,2007]" box="[199,1196,1477,1504]" pageId="14" pageNumber="415">
|
||
<emphasis bold="true" box="[199,345,1477,1503]" pageId="14" pageNumber="415">Description.</emphasis>
|
||
TL =
|
||
<quantity box="[414,576,1477,1504]" metricMagnitude="-2" metricUnit="m" metricValue="1.085" metricValueMax="1.13" metricValueMin="1.04" pageId="14" pageNumber="415" unit="mm" value="10.85" valueMax="11.3" valueMin="10.4">10.4–11.3 mm</quantity>
|
||
in males (mean =
|
||
<quantity box="[783,885,1477,1504]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="14" pageNumber="415" unit="mm" value="10.7">10.7 mm</quantity>
|
||
, n = 3),
|
||
<quantity box="[978,1077,1477,1504]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="14" pageNumber="415" unit="mm" value="11.6">11.6 mm</quantity>
|
||
in female.
|
||
</paragraph>
|
||
<paragraph blockId="14.[151,1437,973,2007]" pageId="14" pageNumber="415">
|
||
Head black with blue-violet reflection; orbital plates distinctly striated in posterior third; frons slightly elevated in anterior third (see above), slightly downward anteriorly (in lateral view), frontal sulci deep, slightly convergent in anterior half and clearly divergent in posterior half; occiput flat. Mandibles in males light-brown underside, pale topside with light-brown teeth and apical molar; in female dark brown except pale basal half of topside. Labial palpi pale except brown apical palpomere; palpomeres 1 and 2 of maxillary palpi pale, palpomeres 3 and 4 light-brown and brown, respectively. Antennae extend posteriorly to the shoulders; scape pale on anterior side and brown-black on posterior side; antennomere 2 dark-brown or brown-black; antennomeres 3–6 black-brown topside and yellowbrown underside; antennomeres 9 and
|
||
<quantity box="[588,646,1765,1791]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="14" pageNumber="415" unit="in" value="10.0">10 in</quantity>
|
||
males with protruding downward anterior lower margins (
|
||
<figureCitation box="[1308,1394,1765,1791]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="14" pageNumber="415">Fig. 43</figureCitation>
|
||
); in female slightly dilated anteriorly; antennomere
|
||
<quantity box="[696,755,1801,1827]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="14" pageNumber="415" unit="in" value="11.0">11 in</quantity>
|
||
males oval with slightly elongated apex, in female sharply abrupt anteriorly (
|
||
<figureCitation box="[356,442,1837,1863]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="14" pageNumber="415">Fig. 42</figureCitation>
|
||
). Labrum indistinctly elongate, LL/LW = 1.0–1.08 (mean = 1.04, n = 4), trapezoid with straight lateral margins, black-brown with very large yellow apical spot unseparated from central apical teeth; in female apices of apical and lateral teeth red-brown (
|
||
<figureCitation box="[733,817,1909,1935]" captionStart="FIGURES 73–80" captionStartId="18.[152,270,1909,1933]" captionTargetBox="[302,1276,200,1868]" captionTargetId="figure-28@18.[296,1292,181,1885]" captionTargetPageId="18" captionText="FIGURES 73–80. Therates spp. labrum: 73, 74—T. biserratus (73—China, Yinjiang County; 74—China, Fanjing Shan- Kuaichang); 75, 76—T. hunnanensis sp. n. (China, Shunhuangshan); 77–80—T. sauteri (77, 78 —China, Taiwan, Taihorin; 79—China, Taiwan, Ilan Hsien; 80—China, Taiwan, Taipei Hsien); 74, 75, 77, 79—males; 73, 76, 78, 80—females; 75— holotype; 73, 76—paratype; 77, 78—paralectotypes." figureDoi="http://doi.org/10.5281/zenodo.7758890" httpUri="https://zenodo.org/record/7758890/files/figure.png" pageId="14" pageNumber="415">Fig. 76</figureCitation>
|
||
), in males brown-black (
|
||
<figureCitation box="[1095,1179,1909,1936]" captionStart="FIGURES 73–80" captionStartId="18.[152,270,1909,1933]" captionTargetBox="[302,1276,200,1868]" captionTargetId="figure-28@18.[296,1292,181,1885]" captionTargetPageId="18" captionText="FIGURES 73–80. Therates spp. labrum: 73, 74—T. biserratus (73—China, Yinjiang County; 74—China, Fanjing Shan- Kuaichang); 75, 76—T. hunnanensis sp. n. (China, Shunhuangshan); 77–80—T. sauteri (77, 78 —China, Taiwan, Taihorin; 79—China, Taiwan, Ilan Hsien; 80—China, Taiwan, Taipei Hsien); 74, 75, 77, 79—males; 73, 76, 78, 80—females; 75— holotype; 73, 76—paratype; 77, 78—paralectotypes." figureDoi="http://doi.org/10.5281/zenodo.7758890" httpUri="https://zenodo.org/record/7758890/files/figure.png" pageId="14" pageNumber="415">Fig. 75</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="14.[151,1437,973,2007]" pageId="14" pageNumber="415">Pronotum black with blue-violet reflection; as wide as long, PW/PL = 0.98–1.03 (mean = 1.0, n = 4), apical lobe practically equal wide throughout; thorax black with violet reflection.</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7758882" ID-Zenodo-Dep="7758882" httpUri="https://zenodo.org/record/7758882/files/figure.png" pageId="15" pageNumber="416" startId="15.[152,270,1909,1933]" targetBox="[320,1268,195,1860]" targetPageId="15">
|
||
<paragraph blockId="15.[151,1437,1909,2041]" pageId="15" pageNumber="416">
|
||
<emphasis bold="true" box="[152,355,1909,1933]" pageId="15" pageNumber="416">FIGURES 50–57.</emphasis>
|
||
<taxonomicName box="[370,591,1909,1933]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="fruhstorferi">
|
||
<emphasis box="[370,591,1909,1933]" italics="true" pageId="15" pageNumber="416">Therates fruhstorferi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[606,646,1909,1933]" pageId="15" pageNumber="416">ssp.</emphasis>
|
||
, labrum: 50–55—
|
||
<taxonomicName authorityName="W. Horn" authorityYear="1902" box="[850,1132,1909,1933]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subSpecies="fruhstorferi">
|
||
<emphasis box="[850,1132,1909,1933]" italics="true" pageId="15" pageNumber="416">T. fruhstorferi fruhstorferi</emphasis>
|
||
</taxonomicName>
|
||
(50, 51—Vietnam, Montes Mauson; 52—Vietnam, Na Hang; 53—Vietnam, Tam Dao; 54, 55—Laos, Ban Nape); 56, 57—
|
||
<taxonomicName authorityName="Matalin et Wiesner" authorityYear="2023" box="[1146,1388,1945,1969]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subSpecies="australis">
|
||
<emphasis box="[1146,1388,1945,1969]" italics="true" pageId="15" pageNumber="416">T. fruhstorferi australis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="15" pageNumber="416">ssp. n.</emphasis>
|
||
(56—Vietnam, Kon Ka Kinh N.P.; 57—Vietnam, Pak Khe River); 50, 52, 54, 56—males; 51, 53, 55, 57—females; 50, 51—paralectotypes; 56—holotype; 57—paratype.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7758884" ID-Zenodo-Dep="7758884" httpUri="https://zenodo.org/record/7758884/files/figure.png" pageId="16" pageNumber="417" startId="16.[152,270,1909,1933]" targetBox="[356,1241,203,1829]" targetPageId="16">
|
||
<paragraph blockId="16.[151,1436,1909,2041]" pageId="16" pageNumber="417">
|
||
<emphasis bold="true" box="[152,349,1909,1933]" pageId="16" pageNumber="417">FIGURES 58–64.</emphasis>
|
||
<taxonomicName box="[357,499,1909,1933]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis box="[357,446,1909,1933]" italics="true" pageId="16" pageNumber="417">Therates</emphasis>
|
||
<emphasis bold="true" box="[454,499,1909,1933]" pageId="16" pageNumber="417">spp.</emphasis>
|
||
</taxonomicName>
|
||
, labrum: 58, 59—
|
||
<taxonomicName authority="Matalin & Wiesner, 2023" authorityName="Matalin & Wiesner" authorityYear="2023" box="[692,1009,1909,1933]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="subSpecies" species="pseudovitalisi" status="ssp. n." subSpecies="pseudovitalisi">
|
||
<emphasis box="[692,1009,1909,1933]" italics="true" pageId="16" pageNumber="417">T. pseudovitalisi pseudovitalisi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1018,1087,1909,1933]" pageId="16" pageNumber="417">
|
||
<taxonomicNameLabel box="[1018,1087,1909,1933]" pageId="16" pageNumber="417" rank="subSpecies">ssp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(58—Laos, Ban Song Cha; 59— Laos, Ban Bun Neua); 60, 61—
|
||
<taxonomicName authorityName="Matalin et Wiesner" authorityYear="2023" box="[478,735,1945,1969]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="subSpecies" species="pseudovitalisi" subSpecies="tibetana">
|
||
<emphasis box="[478,735,1945,1969]" italics="true" pageId="16" pageNumber="417">T. pseudovitalisi tibetana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[742,810,1945,1969]" pageId="16" pageNumber="417">ssp. n.</emphasis>
|
||
(China, Xia-Cha-Yu); 62, 63—
|
||
<taxonomicName authorityName="Tan" authorityYear="1981" box="[1135,1262,1946,1969]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="motoensis">
|
||
<emphasis box="[1135,1262,1946,1969]" italics="true" pageId="16" pageNumber="417">T. motoensis</emphasis>
|
||
</taxonomicName>
|
||
(China, Hanmi); 64—
|
||
<taxonomicName authority="Matalin & Wiesner, 2023" authorityName="Matalin & Wiesner" authorityYear="2023" box="[202,371,1981,2005]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="ziyardamensis" status="sp. n.">
|
||
<emphasis box="[202,371,1981,2005]" italics="true" pageId="16" pageNumber="417">T. ziyardamensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[378,436,1981,2005]" pageId="16" pageNumber="417">
|
||
<taxonomicNameLabel box="[378,436,1981,2005]" pageId="16" pageNumber="417" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(Myanmar, Zi Yar Dam); 58, 60, 62, 64—males; 59, 61, 63—females; 58, 60, 64—holotypes; 59, 61—paratypes.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7758888" ID-Zenodo-Dep="7758888" httpUri="https://zenodo.org/record/7758888/files/figure.png" pageId="17" pageNumber="418" startId="17.[152,270,1944,1968]" targetBox="[283,1300,190,1909]" targetPageId="17">
|
||
<paragraph blockId="17.[151,1436,1944,2040]" pageId="17" pageNumber="418">
|
||
<emphasis bold="true" box="[152,347,1944,1968]" pageId="17" pageNumber="418">FIGURES 65–72.</emphasis>
|
||
<taxonomicName authorityName="W. Horn" authorityYear="1913" box="[354,519,1944,1968]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="vitalisi">
|
||
<emphasis box="[354,519,1944,1968]" italics="true" pageId="17" pageNumber="418">Therates vitalisi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName box="[527,650,1944,1968]" pageId="17" pageNumber="418" rank="subSpecies" subSpecies="labrum">
|
||
<emphasis bold="true" box="[527,567,1944,1968]" pageId="17" pageNumber="418">ssp.</emphasis>
|
||
labrum
|
||
</taxonomicName>
|
||
: 65–70—
|
||
<taxonomicName authorityName="W. Horn" authorityYear="1913" box="[749,920,1944,1968]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="subSpecies" species="vitalisi" subSpecies="vitalisi">
|
||
<emphasis box="[749,920,1944,1968]" italics="true" pageId="17" pageNumber="418">T. vitalisi vitalisi</emphasis>
|
||
</taxonomicName>
|
||
(65, 67—Vietnam, Sa Pa; 66—Vietnam, Lao Cai; 68—Laos, 15 km NW of Louang Namtha; 69—Vietnam, Pu Hoat N.P.; 70—Vietnam, Xuan Lien N.P.); 71, 72—
|
||
<taxonomicName authorityName="Mandl" authorityYear="1954" box="[1303,1436,1980,2004]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="subSpecies" species="vitalisi" subSpecies="ida">
|
||
<emphasis box="[1303,1436,1980,2004]" italics="true" pageId="17" pageNumber="418">T. vitalisi ida</emphasis>
|
||
</taxonomicName>
|
||
(China, Fukien); 65, 69–71—males; 66–68, 72—females; 66, 71—holotypes; 72 – paratype.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7758890" ID-Zenodo-Dep="7758890" httpUri="https://zenodo.org/record/7758890/files/figure.png" pageId="18" pageNumber="419" startId="18.[152,270,1909,1933]" targetBox="[302,1276,200,1868]" targetPageId="18">
|
||
<paragraph blockId="18.[151,1437,1909,2041]" pageId="18" pageNumber="419">
|
||
<emphasis bold="true" box="[152,350,1909,1933]" pageId="18" pageNumber="419">FIGURES 73–80.</emphasis>
|
||
<taxonomicName box="[360,504,1909,1933]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis box="[360,449,1909,1933]" italics="true" pageId="18" pageNumber="419">Therates</emphasis>
|
||
<emphasis bold="true" box="[459,504,1909,1933]" pageId="18" pageNumber="419">spp.</emphasis>
|
||
</taxonomicName>
|
||
labrum: 73, 74—
|
||
<taxonomicName authorityName="Matalin & Wiesner" authorityYear="2023" box="[695,827,1909,1933]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="biserratus">
|
||
<emphasis box="[695,827,1909,1933]" italics="true" pageId="18" pageNumber="419">T. biserratus</emphasis>
|
||
</taxonomicName>
|
||
(73—China, Yinjiang County; 74—China, Fanjing ShanKuaichang); 75, 76—
|
||
<taxonomicName authority="Matalin & Wiesner, 2023" authorityName="Matalin & Wiesner" authorityYear="2023" box="[376,531,1945,1969]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="hunnanensis" status="sp. n.">
|
||
<emphasis box="[376,531,1945,1969]" italics="true" pageId="18" pageNumber="419">T. hunnanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[539,599,1945,1969]" pageId="18" pageNumber="419">
|
||
<taxonomicNameLabel box="[539,599,1945,1969]" pageId="18" pageNumber="419" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(China, Shunhuangshan); 77–80—
|
||
<taxonomicName authorityName="W. Horn" authorityYear="1912" box="[965,1062,1946,1969]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="sauteri">
|
||
<emphasis box="[965,1062,1946,1969]" italics="true" pageId="18" pageNumber="419">T. sauteri</emphasis>
|
||
</taxonomicName>
|
||
(77, 78 —China, Taiwan, Taihorin; 79—China, Taiwan, Ilan Hsien; 80—China, Taiwan, Taipei Hsien); 74, 75, 77, 79—males; 73, 76, 78, 80—females; 75— holotype; 73, 76—paratype; 77, 78—paralectotypes.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7758894" ID-Zenodo-Dep="7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="19" pageNumber="420" startId="19.[152,270,1318,1342]" targetBox="[156,1427,198,1278]" targetPageId="19">
|
||
<paragraph blockId="19.[151,1437,1318,1559]" pageId="19" pageNumber="420">
|
||
<emphasis bold="true" box="[152,349,1318,1343]" pageId="19" pageNumber="420">FIGURES 81–98.</emphasis>
|
||
<taxonomicName box="[357,499,1319,1343]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis box="[357,446,1319,1343]" italics="true" pageId="19" pageNumber="420">Therates</emphasis>
|
||
<emphasis bold="true" box="[455,493,1319,1343]" pageId="19" pageNumber="420">spp</emphasis>
|
||
.
|
||
</taxonomicName>
|
||
, left elytron: 81–87—
|
||
<taxonomicName box="[734,994,1319,1343]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subSpecies="fruhtorferi">
|
||
<emphasis box="[734,994,1319,1343]" italics="true" pageId="19" pageNumber="420">T. fruhstorferi fruhtorferi</emphasis>
|
||
</taxonomicName>
|
||
(81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—
|
||
<taxonomicName authorityName="Matalin et Wiesner" authorityYear="2023" box="[370,612,1391,1415]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subSpecies="australis">
|
||
<emphasis box="[370,612,1391,1415]" italics="true" pageId="19" pageNumber="420">T. fruhstorferi australis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[621,690,1391,1415]" pageId="19" pageNumber="420">ssp. n.</emphasis>
|
||
(88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—
|
||
<taxonomicName authority="Matalin & Wiesner, 2023" authorityName="Matalin & Wiesner" authorityYear="2023" box="[202,515,1427,1451]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="subSpecies" species="pseudovitalisi" status="ssp. n." subSpecies="pseudovitalisi">
|
||
<emphasis box="[202,515,1427,1451]" italics="true" pageId="19" pageNumber="420">T. pseudovitalisi pseudovitalisi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[521,588,1427,1451]" pageId="19" pageNumber="420">
|
||
<taxonomicNameLabel box="[521,588,1427,1451]" pageId="19" pageNumber="420" rank="subSpecies">ssp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—
|
||
<taxonomicName authorityName="Matalin et Wiesner" authorityYear="2023" box="[243,505,1463,1487]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="subSpecies" species="pseudovitalisi" subSpecies="tibetana">
|
||
<emphasis box="[243,505,1463,1487]" italics="true" pageId="19" pageNumber="420">T. pseudovitalisi tibetana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[515,585,1463,1487]" pageId="19" pageNumber="420">ssp. n.</emphasis>
|
||
(China, Xia-Cha-Yu); 96—
|
||
<taxonomicName authority="Matalin & Wiesner, 2023" authorityName="Matalin & Wiesner" authorityYear="2023" box="[879,1051,1463,1487]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="species" species="ziyardamensis" status="sp. n.">
|
||
<emphasis box="[879,1051,1463,1487]" italics="true" pageId="19" pageNumber="420">T. ziyardamensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1061,1121,1463,1487]" pageId="19" pageNumber="420">
|
||
<taxonomicNameLabel box="[1061,1121,1463,1487]" pageId="19" pageNumber="420" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(Myanmar, Zi Yar Dam); 97, 98—
|
||
<taxonomicName authorityName="Tan" authorityYear="1981" box="[202,329,1499,1522]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="species" species="motoensis">
|
||
<emphasis box="[202,329,1499,1522]" italics="true" pageId="19" pageNumber="420">T. motoensis</emphasis>
|
||
</taxonomicName>
|
||
(China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="19.[151,1437,1605,2028]" pageId="19" pageNumber="420">
|
||
Fore femora pale with brown posterior side; middle femora pale with brown with violet tinge anterior side; hind femora pale or in basal third and black-brown with violet tinge in apical 2/3; fore tibiae pale with black apices and light-brown lateral inner side; middle tibiae pale with black apices; hind tibiae pale with brown-black apices and light-brown lateral inner side (in
|
||
<specimenCount box="[518,636,1713,1739]" pageId="19" pageNumber="420" type="male">two males</specimenCount>
|
||
), or brown with violet tinge (in
|
||
<specimenCount box="[988,1089,1713,1739]" pageId="19" pageNumber="420" type="male">one male</specimenCount>
|
||
and in female); fore tarsi black with light violet tinge except dark brown tarsomeres 1 and 2 with black apices; middle and hind tarsi black except yellow-brown or brown tarsomere 1 with black apex; HTbL/HTaL = 1.16–1.24 (mean = 1.19, n = 4).
|
||
</paragraph>
|
||
<paragraph blockId="19.[151,1437,1605,2028]" pageId="19" pageNumber="420">
|
||
Elytra black with blue-violet reflection, sub-parallel or indistinctly divergent toward apex, EL/EW = 2.0–2.09 (mean = 2.03, n = 4); punctuation deep and regular between basal humps and oblique central dot, apical quarter glabrous and smooth; scutellum black with light violet tinge; apical margin in males evenly rounded in female shortly cut. Elytral pattern includes four isolated yellow spots: large triangular basal portion of humeral lunule, basal spot large and prolonged to the suture in males, and very small not extend the suture in female, elongate oblique central dot as well as wide apical dot extend the suture (
|
||
<figureCitation box="[777,864,2001,2028]" captionStart="FIGURES 15–28" captionStartId="9.[152,270,1641,1665]" captionTargetBox="[156,1418,712,1612]" captionTargetId="figure-217@9.[151,1435,709,1617]" captionTargetPageId="9" captionText="FIGURES 15–28. Therates spp., habitus: 15–18—T. vitalisi vitalisi (15, 16—Vietnam, Lao Cai; 17—Vietnam, Fan Si Pan Mnt; 18—Vietnam, Sa Pa); 19–21—T. vitalisi ida (China, Kuatun); 22–24—T. biserratus (22, 23—China, Yinjiang County; 24—China, Fanjing Shan-Kuaichang); 25—T. hunnanensis sp. n. (China, Shunhuangshan); 26–28—T. sauteri (26, 27—China, Tiawan, Taihorin; 28—China, Taiwan, Ilan Hsien); 17–20, 24–28—males; 15, 16, 21–23 —females; 15, 16, 19, 20, 25— holotypes; 22, 23—paratype; 26, 27—lectotype; 15, 17–19, 21, 22, 24–26, 28—habitus; 16, 20, 23, 27—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758876" httpUri="https://zenodo.org/record/7758876/files/figure.png" pageId="19" pageNumber="420">Figs 25</figureCitation>
|
||
,
|
||
<figureCitation box="[876,978,2001,2027]" captionStart="FIGURES 99–114" captionStartId="21.[152,273,1418,1442]" captionTargetBox="[155,1421,210,1377]" captionTargetId="figure-17@21.[151,1435,181,1394]" captionTargetPageId="21" captionText="FIGURES 99–114. Therates spp., left elytron: 99–102—T. vitalisi vitalisi (99 —Vietnam, Lao Cai; 100, 101—Vietnam, Sa Pa; 102—Laos, 15 km NW Louang Namtha); 103, 104—T. vitalisi ida (China, Kuatun); 105, 106—T. biserratus (105— China, Yinjiang County; 106—China, Fanjing Shan-Kuaichang); 107, 108—T. hunnanensis sp. n. (China, Shunhuangshan); 109–114—T. sauteri (109, 110, 112, 113—China, Taiwan, Taihorin; 111—China, Taiwan, Ilan Hsien; 114—China, Taiwan, Taipei Hsien); 100, 103, 106, 107, 109–111—males; 99, 101, 102, 104, 105, 108, 112–114—females; 99, 103, 107— holotypes; 105, 108—paratypes; 109, 110, 112, 113—paralectotypes" figureDoi="http://doi.org/10.5281/zenodo.7758896" httpUri="https://zenodo.org/record/7758896/files/figure.png" pageId="19" pageNumber="420">107, 108</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="20.[151,1437,151,285]" pageId="20" pageNumber="421">
|
||
Aedeagus with slightly elongate and indistinctly shortly curved apex and long lateral carinae (
|
||
<figureCitation box="[1263,1423,151,177]" captionStart="FIGURES 152–174" captionStartId="24.[152,270,1210,1234]" captionTargetBox="[163,1428,181,1165]" captionTargetId="figure-28@24.[151,1435,181,1186]" captionTargetPageId="24" captionText="FIGURES 152–174. Therates spp., aedeagus: 152–166—T. vitalisi vitalisi (152, 153—Vietnam, Xuan Son N.P.; 154— Vietnam, Sa Pa; 155, 156—Laos, Ban Nape—Kaev Nua pass; 157–163—Laos, 15–20 km NW Louang Namtha; 164, 165— Laos, 8 km NE Ban Nape; 166—Laos, Thong Khan); 167—T. vitalisi ida (China, Kuatun); 168—T. biserratus (China, Fanjing Shan-Kuaichang); 169–171—T. hunnanensis sp. n. (China, Shunhuangshan); 172—T. motoensis (China, Hanmi); 173, 174—T. sauteri (173—China, Taiwan, Taihorin; 174—China, Taiwan, Ilan Hsien); 167, 169—holotypes; 170, 171—paratypes; 173— lectotype; 152—dorsal view; 153–174—left lateral view; ac—apical carinae; sp—spring; fl—flagellum. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758906" httpUri="https://zenodo.org/record/7758906/files/figure.png" pageId="20" pageNumber="421">Figs 169–171</figureCitation>
|
||
); AL =
|
||
<quantity box="[217,352,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.6" metricValueMin="2.4" pageId="20" pageNumber="421" unit="mm" value="2.5" valueMax="2.6" valueMin="2.4">2.4–2.6 mm</quantity>
|
||
(mean =
|
||
<quantity box="[459,547,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="421" unit="mm" value="2.5">2.5 mm</quantity>
|
||
, n = 3); EL/AL = 2.54—2.92 (mean = 2.74, n = 3).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[199,1109,223,249]" pageId="20" pageNumber="421" type="etymology">
|
||
<paragraph blockId="20.[151,1437,151,285]" box="[199,1109,223,249]" pageId="20" pageNumber="421">
|
||
<emphasis bold="true" box="[199,334,223,249]" pageId="20" pageNumber="421">Etymology.</emphasis>
|
||
A new species is named for the
|
||
<typeStatus box="[695,743,223,249]" pageId="20" pageNumber="421">type</typeStatus>
|
||
area—Chinese province
|
||
<collectingRegion box="[1027,1105,223,249]" country="China" name="Hunan" pageId="20" pageNumber="421">Hunan</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[199,1140,258,285]" pageId="20" pageNumber="421" type="distribution">
|
||
<paragraph blockId="20.[151,1437,151,285]" box="[199,1140,258,285]" pageId="20" pageNumber="421">
|
||
<emphasis bold="true" box="[199,352,258,285]" pageId="20" pageNumber="421">Distribution.</emphasis>
|
||
<collectingCountry box="[359,437,259,285]" name="China" pageId="20" pageNumber="421">CHINA</collectingCountry>
|
||
:
|
||
<collectingRegion box="[447,629,259,285]" country="China" name="Hunan" pageId="20" pageNumber="421">Hunan Province</collectingRegion>
|
||
(Dong’an and Chengbu Counties) (
|
||
<figureCitation box="[1029,1128,259,285]" captionStart="FIGURES 184" captionStartId="28.[152,270,1579,1603]" captionTargetBox="[151,1435,597,1556]" captionTargetId="figure-226@28.[151,1435,597,1556]" captionTargetPageId="28" captionText="FIGURES 184. Distribution of Therates spp.: dark blue solid circle—T. vitalisi vitalisi; dark blue half-solid circle—T. vitalisi ida; lilac solid circle—T. hunnanensis sp. n.; orange solid circle—T. biserratus; pink solid circle—T. motoensis; red open circle—T. sauteri; red solid quadrat—T. fruhstorferi fruhstorferi; red half-solid quadrat—T. fruhstorferi australis ssp. n.; blue solid quadrat—T. ziyardamensis sp. n.; green solid quadrat—T. pseudovitalisi sp. n. (nominotypical subspecies); green halfsolid quadrat—T. pseudovitalisi tibetana ssp. n.; L in the symbols—according only to the literature sources. Map source— https://commons.wikimedia.org/wiki/File:East_Asia_topographic_map.png." figureDoi="http://doi.org/10.5281/zenodo.7758916" httpUri="https://zenodo.org/record/7758916/files/figure.png" pageId="20" pageNumber="421">Fig. 184</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |