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<document id="249251DE6D4C1A56383DEAD925D35114" ID-DOI="10.11646/zootaxa.1954.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5240917" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629589774350" checkinUser="felipe" docAuthor="Goldschmidt, Tom" docDate="2008" docId="B03B87976970FFADFF21FE515E7FFA57" docLanguage="en" docName="zt01954p120.pdf" docOrigin="Zootaxa 1954 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1954.1.1" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Nilotonia (Cookonia) avomonina Goldschmidt 2008, sp. nov." docType="treatment" docVersion="3" lastPageNumber="58" masterDocId="4C02FFEF6947FF94FFB6FFD65B27FFBA" masterDocTitle="Taxonomical, ecological and zoogeographical studies on anisitsiellid water mites (Acari: Hydrachnidia: Anisitsiellidae Koenike, 1910) from Madagascar" masterLastPageNumber="120" masterPageNumber="1" pageNumber="56" updateTime="1699081947219" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="C853BFAA8244E87EE5F070C555B1189D">Taxonomical, ecological and zoogeographical studies on anisitsiellid water mites (Acari: Hydrachnidia: Anisitsiellidae Koenike, 1910) from Madagascar</mods:title>
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<mods:namePart id="FA9DC17895DB0DEB9024DEB269DB1F7A">Goldschmidt, Tom</mods:namePart>
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<mods:number id="131B9CE2024E562BC07AC1F382FB35F2">1954</mods:number>
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<treatment id="B03B87976970FFADFF21FE515E7FFA57" ID-DOI="http://doi.org/10.5281/zenodo.5241967" ID-Zenodo-Dep="5241967" LSID="urn:lsid:plazi:treatment:B03B87976970FFADFF21FE515E7FFA57" httpUri="http://treatment.plazi.org/id/B03B87976970FFADFF21FE515E7FFA57" lastPageId="57" lastPageNumber="58" pageId="55" pageNumber="56">
<subSubSection id="7088650A6970FFA3FF21FE5159ABFE1B" box="[151,652,391,417]" pageId="55" pageNumber="56" type="nomenclature">
<paragraph id="382D36816970FFA3FF21FE5159ABFE1B" blockId="55.[151,652,391,452]" box="[151,652,391,417]" pageId="55" pageNumber="56">
<heading id="636581ED6970FFA3FF21FE5159ABFE1B" bold="true" box="[151,652,391,417]" fontSize="11" level="1" pageId="55" pageNumber="56" reason="1">
<emphasis id="0AE6EA936970FFA3FF21FE5159ABFE1B" bold="true" box="[151,652,391,417]" pageId="55" pageNumber="56">
<taxonomicName id="FF924D026970FFA3FF21FE515900FE1B" authority="Goldschmidt, 2008" authorityName="Goldschmidt" authorityYear="2008" box="[151,551,391,417]" class="Arachnida" family="Anisitsiellidae" genus="Nilotonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="avomonina" status="sp. nov." subGenus="Cookonia">
<emphasis id="0AE6EA936970FFA3FF21FE515A2EFE1B" bold="true" box="[151,265,391,417]" italics="true" pageId="55" pageNumber="56">Nilotonia</emphasis>
(
<emphasis id="0AE6EA936970FFA3FEACFE515AA8FE1B" bold="true" box="[282,399,391,417]" italics="true" pageId="55" pageNumber="56">Cookonia</emphasis>
)
<emphasis id="0AE6EA936970FFA3FE16FE515900FE1B" bold="true" box="[416,551,391,417]" italics="true" pageId="55" pageNumber="56">avomonina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="11D557E86970FFA3FD98FE5159ABFE1B" box="[558,652,391,417]" pageId="55" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7088650A6970FFA3FF21FE7C5A88FE7E" box="[151,431,426,452]" pageId="55" pageNumber="56" type="description">
<paragraph id="382D36816970FFA3FF21FE7C5A88FE7E" blockId="55.[151,652,391,452]" box="[151,431,426,452]" pageId="55" pageNumber="56">
(
<figureCitation id="A0A92A046970FFA3FF29FE7C5A65FE7E" box="[159,322,426,452]" captionStart="FIGURES 112120" captionStartId="56.[151,269,1907,1931]" captionTargetBox="[209,1347,200,1874]" captionTargetId="figure-78@56.[199,1386,194,1878]" captionTargetPageId="56" captionText="FIGURES 112120. Nilotonia (Cookonia) avomonina subgen. et sp. nov., female holotype (MD 106a): 112, idiosoma, ventral view; 113, idiosoma, dorsal view; 114, leg-II; 115, leg-III-6; 116, leg-IV (first segment broken); 117, capitulum with left palp, lateral view; 118, left chelicera, lateral view; 119, right chelicera, medial view; 120, right palp, medial view. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5240959" httpUri="https://zenodo.org/record/5240959/files/figure.png" pageId="55" pageNumber="56">Figs 112120</figureCitation>
,
<tableCitation id="7510033A6970FFA3FEF8FE7C5A82FE7E" box="[334,421,426,452]" captionStart="TABLE 6" captionStartId="52.[151,241,151,175]" captionText="TABLE 6. Measurement data for Nilotonia (Cookonia) anjozorobe subgen. et sp. nov. and Nilotonia (Cookonia)" pageId="55" pageNumber="56">Table 6</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="7088650A6970FFA3FF21FE2C5915FD36" pageId="55" pageNumber="56" type="materials_examined">
<paragraph id="382D36816970FFA3FF21FE2C5AD4FDDE" blockId="55.[151,1436,506,1972]" pageId="55" pageNumber="56">
<emphasis id="0AE6EA936970FFA3FF21FE2C5BFAFDAE" bold="true" box="[151,221,506,532]" pageId="55" pageNumber="56">Type:</emphasis>
<materialsCitation id="88FA3CDC6970FFA3FF50FE2C5AD4FDDE" collectingDate="2001-07-10" collectionCode="MD" elevation="2350" pageId="55" pageNumber="56" specimenCode="MD 106" specimenCount="1" typeStatus="holotype">
<emphasis id="0AE6EA936970FFA3FF50FE2C5A95FDAE" bold="true" box="[230,434,506,532]" pageId="55" pageNumber="56">
<typeStatus id="E72988236970FFA3FF50FE2C5A7FFDAE" box="[230,344,506,532]" pageId="55" pageNumber="56" type="holotype">Holotype</typeStatus>
female
</emphasis>
,
<specimenCode id="68349EFA6970FFA3FE77FE2C5900FDAE" box="[449,551,506,532]" collectionCode="MD" country="Germany" name="Museum Donaueschingen" pageId="55" pageNumber="56">MD 106</specimenCode>
a,
<taxonomicName id="FF924D026970FFA3FDF7FE2C5BFCFD86" authority=", Reserve Manjakatompo, Mont Ambohimirandrana" authorityName="Reserve Manjakatompo, Mont Ambohimirandrana" class="Arachnida" genus="Ankaratra" kingdom="Animalia" order="Opiliones" pageId="55" pageNumber="56" phylum="Arthropoda" rank="subGenus" subGenus="Antananarivo">Ankaratra (Antananarivo), Reserve Manjakatompo, Mont Ambohimirandrana</taxonomicName>
, spring area exposition south-east (drainage Rivière Mahiavona), upper spring,
<elevation id="B3BFD1B26970FFA3FB38FDF45E33FD86" box="[1166,1300,546,572]" metricMagnitude="3" metricUnit="m" metricValue="2.35" pageId="55" pageNumber="56" unit="m" value="2350.0">
<quantity id="FF6A9B646970FFA3FB38FDF45FCEFD86" box="[1166,1257,546,572]" metricMagnitude="3" metricUnit="m" metricValue="2.35" pageId="55" pageNumber="56" unit="m" value="2350.0">2350 m</quantity>
asl
</elevation>
, 13.4 °C, 2 µS/cm,
<date id="4C2C10416970FFA3FF45FD9C5A5EFDDE" box="[243,377,586,612]" pageId="55" pageNumber="56" value="2001-07-10">
<collectingDate id="5C68E9A96970FFA3FF45FD9C5A5EFDDE" box="[243,377,586,612]" pageId="55" pageNumber="56" value="2001-07-10">07.10.2001</collectingDate>
</date>
, mounted.
</materialsCitation>
</paragraph>
<paragraph id="382D36816970FFA3FF71FDA45915FD36" blockId="55.[151,1436,506,1972]" box="[199,562,626,652]" pageId="55" pageNumber="56">
<emphasis id="0AE6EA936970FFA3FF71FDA45A16FD36" bold="true" box="[199,305,626,652]" pageId="55" pageNumber="56">Habitat:</emphasis>
Spring at
<quantity id="FF6A9B646970FFA3FE1CFDA45924FD36" box="[426,515,626,652]" metricMagnitude="3" metricUnit="m" metricValue="2.35" pageId="55" pageNumber="56" unit="m" value="2350.0">2350 m</quantity>
asl.
</paragraph>
</subSubSection>
<subSubSection id="7088650A6970FFA3FF71FD4C58D8FD0E" box="[199,1023,666,692]" pageId="55" pageNumber="56" type="distribution">
<paragraph id="382D36816970FFA3FF71FD4C58D8FD0E" blockId="55.[151,1436,506,1972]" box="[199,1023,666,692]" pageId="55" pageNumber="56">
<emphasis id="0AE6EA936970FFA3FF71FD4C5A4EFD0E" bold="true" box="[199,361,666,692]" pageId="55" pageNumber="56">Distribution:</emphasis>
<collectingCountry id="408576116970FFA3FEC7FD4C5AD9FD0E" box="[369,510,666,692]" name="Madagascar" pageId="55" pageNumber="56">Madagascar</collectingCountry>
(Central Highland Ankaratra mountains).
</paragraph>
</subSubSection>
<subSubSection id="7088650A6970FFA3FF71FD145A5AFC96" pageId="55" pageNumber="56" type="etymology">
<paragraph id="382D36816970FFA3FF71FD145A5AFC96" blockId="55.[151,1436,506,1972]" pageId="55" pageNumber="56">
<emphasis id="0AE6EA936970FFA3FF71FD145A95FD66" bold="true" box="[199,434,706,732]" pageId="55" pageNumber="56">Derivatio nominis:</emphasis>
<emphasis id="0AE6EA936970FFA3FE0DFD145AC0FD61" box="[443,487,706,731]" italics="true" pageId="55" pageNumber="56">Avo</emphasis>
(Malagasy) — high;
<emphasis id="0AE6EA936970FFA3FD5FFD145866FD61" box="[745,833,706,731]" italics="true" pageId="55" pageNumber="56">monina</emphasis>
(Malagasy) — to live; referring to the high elevation (highest sample site within the water mite samples from
<collectingCountry id="408576116970FFA3FCC4FD3C5F26FCBE" box="[882,1025,746,772]" name="Madagascar" pageId="55" pageNumber="56">Madagascar</collectingCountry>
) where the
<typeStatus id="E72988236970FFA3FB39FD3C5FE6FCBE" box="[1167,1217,746,772]" pageId="55" pageNumber="56">type</typeStatus>
locality (and only locality) is situated.
</paragraph>
</subSubSection>
<subSubSection id="7088650A6970FFA3FF71FCEC5F52FBFE" pageId="55" pageNumber="56" type="diagnosis">
<paragraph id="382D36816970FFA3FF71FCEC5F52FBFE" blockId="55.[151,1436,506,1972]" pageId="55" pageNumber="56">
<emphasis id="0AE6EA936970FFA3FF71FCEC5A6EFCEE" bold="true" box="[199,329,826,852]" pageId="55" pageNumber="56">Diagnosis:</emphasis>
Characters of the genus; integument lined; caudal and lateral margins of Cx-IV widely extended; postgenital sclerite widely extended, separated from secondary extension of Cx-IV by very narrow stripe of soft integument; Vgl-2 and -4 individually on large platelets; Cx-I medially approximate but not fused, Cx-III medially with slight secondary sclerotization, approximate to each other; genital field elongated trapezoid, acetabula mid-sized, elongated-oval, touching each other; sub-terminal seta on leg-IV-6 relatively short, inserted far proximal, not reaching distal tip of segment; capitulum stout, nearly rectangular; palp relatively slender, P1 without dorsal seta, P4 ventrally with small protrusion, P5 slender.
</paragraph>
</subSubSection>
<subSubSection id="7088650A6970FFADFF71FB845AADFA9F" lastPageId="57" lastPageNumber="58" pageId="55" pageNumber="56" type="description">
<paragraph id="382D36816970FFADFF71FB845E7AFB47" blockId="55.[151,1436,506,1972]" lastBlockId="57.[151,1436,1051,1517]" lastPageId="57" lastPageNumber="58" pageId="55" pageNumber="56">
<emphasis id="0AE6EA936970FFA3FF71FB845A9EFBD6" bold="true" box="[199,441,1106,1132]" pageId="55" pageNumber="56">Description, female</emphasis>
(n = 1): Idiosoma rounded-oval (
<figureCitation id="A0A92A046970FFA3FC88FB845883FBD6" box="[830,932,1106,1132]" captionStart="FIGURES 112120" captionStartId="56.[151,269,1907,1931]" captionTargetBox="[209,1347,200,1874]" captionTargetId="figure-78@56.[199,1386,194,1878]" captionTargetPageId="56" captionText="FIGURES 112120. Nilotonia (Cookonia) avomonina subgen. et sp. nov., female holotype (MD 106a): 112, idiosoma, ventral view; 113, idiosoma, dorsal view; 114, leg-II; 115, leg-III-6; 116, leg-IV (first segment broken); 117, capitulum with left palp, lateral view; 118, left chelicera, lateral view; 119, right chelicera, medial view; 120, right palp, medial view. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5240959" httpUri="https://zenodo.org/record/5240959/files/figure.png" pageId="55" pageNumber="56">Fig. 112</figureCitation>
); dorsum mainly covered by large plate (L/ W 775/544), including Dgl-3 to -6, Lgl-4 and post-ocular setae, colour slightly dark-purple; lateral eyes small, oval, separated on both sides, lying free under integument; Dgl-1 far anterior; soft integument around dorsal plate lined, bearing slightly extended Dgl-2, Lgl-2 and l1-4; setae of Dgl-1, -3, -4 and -5 mid-sized, setae of other glandularia small (
<figureCitation id="A0A92A046970FFA3FE00FB24593AFAB6" box="[438,541,1266,1292]" captionStart="FIGURES 112120" captionStartId="56.[151,269,1907,1931]" captionTargetBox="[209,1347,200,1874]" captionTargetId="figure-78@56.[199,1386,194,1878]" captionTargetPageId="56" captionText="FIGURES 112120. Nilotonia (Cookonia) avomonina subgen. et sp. nov., female holotype (MD 106a): 112, idiosoma, ventral view; 113, idiosoma, dorsal view; 114, leg-II; 115, leg-III-6; 116, leg-IV (first segment broken); 117, capitulum with left palp, lateral view; 118, left chelicera, lateral view; 119, right chelicera, medial view; 120, right palp, medial view. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5240959" httpUri="https://zenodo.org/record/5240959/files/figure.png" pageId="55" pageNumber="56">Fig. 113</figureCitation>
); Lgl-1 fused with lateral secondary sclerotization of Cx-IV; Lgl-3 and l
<quantity id="FF6A9B646970FFA3FAD9FB245EBBFAB6" box="[1391,1436,1266,1292]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="55" pageNumber="56" unit="in" value="5.0">5 in</quantity>
soft integument on ventral side, secondary sclerotization of Cx-IV forming a nearly closed, rounded ventral shield (L/W 706/755), extended also latero-dorsally of the Cx-I/II/III; Cx-I medially approximate, posterior half overlapping, but separated, postero-lateral margin of Cx-II lying over Cx-III, postero-medial margin of Cx-II lying under Cx-III; Cx-III medially approximate, medial margin of Cx-III extended by fine secondary sclerotization; suture line between Cx-III and Cx-IV incomplete, medially curved towards anterior, medial margin of Cx-IV straight to slightly concave, forming very narrow genital bay, caudal margin short, rounded, latero-caudal margin oblique towards anterior, insertion of leg-IV far anterior (
<figureCitation id="A0A92A046970FFA3FBF2F9DC5F8BF99E" box="[1092,1196,1546,1572]" captionStart="FIGURES 112120" captionStartId="56.[151,269,1907,1931]" captionTargetBox="[209,1347,200,1874]" captionTargetId="figure-78@56.[199,1386,194,1878]" captionTargetPageId="56" captionText="FIGURES 112120. Nilotonia (Cookonia) avomonina subgen. et sp. nov., female holotype (MD 106a): 112, idiosoma, ventral view; 113, idiosoma, dorsal view; 114, leg-II; 115, leg-III-6; 116, leg-IV (first segment broken); 117, capitulum with left palp, lateral view; 118, left chelicera, lateral view; 119, right chelicera, medial view; 120, right palp, medial view. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5240959" httpUri="https://zenodo.org/record/5240959/files/figure.png" pageId="55" pageNumber="56">Fig. 112</figureCitation>
); medial and caudal margin of Cx-IV clearly visible, lateral margin unclear; setae at postero-lateral margin of Cx-I and medial margin of Cx-III/IV very small (
<figureCitation id="A0A92A046970FFA3FDA2F98C595EF9CE" box="[532,633,1626,1652]" captionStart="FIGURES 112120" captionStartId="56.[151,269,1907,1931]" captionTargetBox="[209,1347,200,1874]" captionTargetId="figure-78@56.[199,1386,194,1878]" captionTargetPageId="56" captionText="FIGURES 112120. Nilotonia (Cookonia) avomonina subgen. et sp. nov., female holotype (MD 106a): 112, idiosoma, ventral view; 113, idiosoma, dorsal view; 114, leg-II; 115, leg-III-6; 116, leg-IV (first segment broken); 117, capitulum with left palp, lateral view; 118, left chelicera, lateral view; 119, right chelicera, medial view; 120, right palp, medial view. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5240959" httpUri="https://zenodo.org/record/5240959/files/figure.png" pageId="55" pageNumber="56">Fig. 112</figureCitation>
); genital bay narrow, approximate to genital field, formed by medial margins and caudal secondary sclerotization of Cx-IV as well as post-genital sclerite; genital flaps elongated, anteriorly and postero-laterally smoothly rounded, laterally nearly straight, broadened towards cauda; acetabula mid-sized, elongated-oval, slender, touching each other, Ac1 distant from anterior margin of genital flaps, Ac3 slightly distant from posterior margin; pre-genital sclerite triangular, laterally protruding genital flaps, post-genital sclerite large, greatly extended by secondary sclerotization, including setae of Vgl-1, forming caudal segment of ventral shield (
<figureCitation id="A0A92A046970FFA3FDE1F89C5998F8DE" box="[599,703,1866,1892]" captionStart="FIGURES 112120" captionStartId="56.[151,269,1907,1931]" captionTargetBox="[209,1347,200,1874]" captionTargetId="figure-78@56.[199,1386,194,1878]" captionTargetPageId="56" captionText="FIGURES 112120. Nilotonia (Cookonia) avomonina subgen. et sp. nov., female holotype (MD 106a): 112, idiosoma, ventral view; 113, idiosoma, dorsal view; 114, leg-II; 115, leg-III-6; 116, leg-IV (first segment broken); 117, capitulum with left palp, lateral view; 118, left chelicera, lateral view; 119, right chelicera, medial view; 120, right palp, medial view. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5240959" httpUri="https://zenodo.org/record/5240959/files/figure.png" pageId="55" pageNumber="56">Fig. 112</figureCitation>
); Vgl-3 very close to caudal margin of Cx-IV, included in secondary sclerotization of Cx-IV; Vgl-2 and Vgl-4 on relatively large, irregular platelets; excretory pore without sclerotization (
<figureCitation id="A0A92A046970FFA3FEC7F84C5AF0F80E" box="[369,471,1946,1972]" captionStart="FIGURES 112120" captionStartId="56.[151,269,1907,1931]" captionTargetBox="[209,1347,200,1874]" captionTargetId="figure-78@56.[199,1386,194,1878]" captionTargetPageId="56" captionText="FIGURES 112120. Nilotonia (Cookonia) avomonina subgen. et sp. nov., female holotype (MD 106a): 112, idiosoma, ventral view; 113, idiosoma, dorsal view; 114, leg-II; 115, leg-III-6; 116, leg-IV (first segment broken); 117, capitulum with left palp, lateral view; 118, left chelicera, lateral view; 119, right chelicera, medial view; 120, right palp, medial view. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5240959" httpUri="https://zenodo.org/record/5240959/files/figure.png" pageId="55" pageNumber="56">Fig. 112</figureCitation>
); legs with several large setae; claws on leg-I to -III with one small dorsal and one larger ventral clawlet (
<figureCitation id="A0A92A04697EFFADFE16FBCD5963FB8F" box="[416,580,1051,1077]" captionStart="FIGURES 112120" captionStartId="56.[151,269,1907,1931]" captionTargetBox="[209,1347,200,1874]" captionTargetId="figure-78@56.[199,1386,194,1878]" captionTargetPageId="56" captionText="FIGURES 112120. Nilotonia (Cookonia) avomonina subgen. et sp. nov., female holotype (MD 106a): 112, idiosoma, ventral view; 113, idiosoma, dorsal view; 114, leg-II; 115, leg-III-6; 116, leg-IV (first segment broken); 117, capitulum with left palp, lateral view; 118, left chelicera, lateral view; 119, right chelicera, medial view; 120, right palp, medial view. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5240959" httpUri="https://zenodo.org/record/5240959/files/figure.png" pageId="57" pageNumber="58">Figs 114, 115</figureCitation>
), leg-IV-6 distally tapering, with two small, peg-like terminal setae (
<figureCitation id="A0A92A04697EFFADFADBFBCD5BE1FBE7" captionStart="FIGURES 112120" captionStartId="56.[151,269,1907,1931]" captionTargetBox="[209,1347,200,1874]" captionTargetId="figure-78@56.[199,1386,194,1878]" captionTargetPageId="56" captionText="FIGURES 112120. Nilotonia (Cookonia) avomonina subgen. et sp. nov., female holotype (MD 106a): 112, idiosoma, ventral view; 113, idiosoma, dorsal view; 114, leg-II; 115, leg-III-6; 116, leg-IV (first segment broken); 117, capitulum with left palp, lateral view; 118, left chelicera, lateral view; 119, right chelicera, medial view; 120, right palp, medial view. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5240959" httpUri="https://zenodo.org/record/5240959/files/figure.png" pageId="57" pageNumber="58">Fig. 116</figureCitation>
); capitulum very short, relatively high, dorsal and ventral margins nearly parallel (
<figureCitation id="A0A92A04697EFFADFB07FB955E3DFBE7" box="[1201,1306,1091,1117]" captionStart="FIGURES 112120" captionStartId="56.[151,269,1907,1931]" captionTargetBox="[209,1347,200,1874]" captionTargetId="figure-78@56.[199,1386,194,1878]" captionTargetPageId="56" captionText="FIGURES 112120. Nilotonia (Cookonia) avomonina subgen. et sp. nov., female holotype (MD 106a): 112, idiosoma, ventral view; 113, idiosoma, dorsal view; 114, leg-II; 115, leg-III-6; 116, leg-IV (first segment broken); 117, capitulum with left palp, lateral view; 118, left chelicera, lateral view; 119, right chelicera, medial view; 120, right palp, medial view. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5240959" httpUri="https://zenodo.org/record/5240959/files/figure.png" pageId="57" pageNumber="58">Fig. 117</figureCitation>
); chelicera strong, proximally high, claw very heavy, dorso-distally serrate (
<figureCitation id="A0A92A04697EFFADFC1EFBBD5F75FB3F" box="[936,1106,1131,1157]" captionStart="FIGURES 112120" captionStartId="56.[151,269,1907,1931]" captionTargetBox="[209,1347,200,1874]" captionTargetId="figure-78@56.[199,1386,194,1878]" captionTargetPageId="56" captionText="FIGURES 112120. Nilotonia (Cookonia) avomonina subgen. et sp. nov., female holotype (MD 106a): 112, idiosoma, ventral view; 113, idiosoma, dorsal view; 114, leg-II; 115, leg-III-6; 116, leg-IV (first segment broken); 117, capitulum with left palp, lateral view; 118, left chelicera, lateral view; 119, right chelicera, medial view; 120, right palp, medial view. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5240959" httpUri="https://zenodo.org/record/5240959/files/figure.png" pageId="57" pageNumber="58">Figs 118, 119</figureCitation>
); palps slender, P1 without setae, P2 with one latero-ventral seta and six dorsal setae, ventral margin straight, P3 with one lateral and three dorsal setae, P4 distally curved, dorso-distal corner rounded, ventral setae slightly in distal half, ventral margin with slight protrusion, P5 slender, cone-shaped with long slender terminal claws (
<figureCitation id="A0A92A04697EFFADFB1DFB355E77FB47" box="[1195,1360,1251,1277]" captionStart="FIGURES 112120" captionStartId="56.[151,269,1907,1931]" captionTargetBox="[209,1347,200,1874]" captionTargetId="figure-78@56.[199,1386,194,1878]" captionTargetPageId="56" captionText="FIGURES 112120. Nilotonia (Cookonia) avomonina subgen. et sp. nov., female holotype (MD 106a): 112, idiosoma, ventral view; 113, idiosoma, dorsal view; 114, leg-II; 115, leg-III-6; 116, leg-IV (first segment broken); 117, capitulum with left palp, lateral view; 118, left chelicera, lateral view; 119, right chelicera, medial view; 120, right palp, medial view. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5240959" httpUri="https://zenodo.org/record/5240959/files/figure.png" pageId="57" pageNumber="58">Figs 117, 120</figureCitation>
).
</paragraph>
<caption id="6CED6609697FFFACFF21F8A55A9EF848" ID-DOI="http://doi.org/10.5281/zenodo.5240959" ID-Zenodo-Dep="5240959" httpUri="https://zenodo.org/record/5240959/files/figure.png" pageId="56" pageNumber="57" startId="56.[151,269,1907,1931]" targetBox="[209,1347,200,1874]" targetPageId="56">
<paragraph id="382D3681697FFFACFF21F8A55A9EF848" blockId="56.[151,1436,1907,2034]" pageId="56" pageNumber="57">
<emphasis id="0AE6EA93697FFFACFF21F8A55A50F831" bold="true" box="[151,375,1907,1931]" pageId="56" pageNumber="57">FIGURES 112120.</emphasis>
<taxonomicName id="FF924D02697FFFACFEC8F8A559C7F830" authority="Goldschmidt, 2008" authorityName="Goldschmidt" authorityYear="2008" box="[382,736,1907,1930]" class="Arachnida" family="Anisitsiellidae" genus="Nilotonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="avomonina" status="subgen. et sp. nov." subGenus="Cookonia">
<emphasis id="0AE6EA93697FFFACFEC8F8A55AC4F830" box="[382,483,1907,1930]" italics="true" pageId="56" pageNumber="57">Nilotonia</emphasis>
(
<emphasis id="0AE6EA93697FFFACFE45F8A5597CF830" box="[499,603,1907,1930]" italics="true" pageId="56" pageNumber="57">Cookonia</emphasis>
)
<emphasis id="0AE6EA93697FFFACFDDCF8A559C7F830" box="[618,736,1907,1930]" italics="true" pageId="56" pageNumber="57">avomonina</emphasis>
</taxonomicName>
<emphasis id="0AE6EA93697FFFACFD5EF8A55890F831" bold="true" box="[744,951,1907,1931]" pageId="56" pageNumber="57">
<taxonomicNameLabel id="11D557E8697FFFACFD5EF8A55890F831" box="[744,951,1907,1931]" pageId="56" pageNumber="57" rank="species">subgen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, female holotype (MD 106a): 112, idiosoma, ventral view; 113, idiosoma, dorsal view; 114, leg-II; 115, leg-III-6; 116, leg-IV (first segment broken); 117, capitulum with left palp, lateral view; 118, left chelicera, lateral view; 119, right chelicera, medial view; 120, right palp, medial view. Scale bars = 100 µm.
</paragraph>
</caption>
<caption id="6CED6609697EFFADFF21FC7B5AD3FC5D" ID-DOI="http://doi.org/10.5281/zenodo.5240961" ID-Zenodo-Dep="5240961" httpUri="https://zenodo.org/record/5240961/files/figure.png" pageId="57" pageNumber="58" startId="57.[151,269,941,965]" targetBox="[166,1414,188,906]" targetPageId="57">
<paragraph id="382D3681697EFFADFF21FC7B5AD3FC5D" blockId="57.[151,1436,941,999]" pageId="57" pageNumber="58">
<emphasis id="0AE6EA93697EFFADFF21FC7B5A5FFC7F" bold="true" box="[151,376,941,965]" pageId="57" pageNumber="58">FIGURES 121, 122.</emphasis>
<taxonomicName id="FF924D02697EFFADFEC9FC785898FC7E" authority="Cook, 1966" authorityName="Cook" authorityYear="1966" box="[383,959,941,965]" class="Arachnida" family="Anisitsiellidae" genus="Mamersella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="tototaensis" subGenus="Neomamersella">
<emphasis id="0AE6EA93697EFFADFEC9FC78581EFC7F" box="[383,825,942,965]" italics="true" pageId="57" pageNumber="58">Mamersella (Neomamersella) tototaensis</emphasis>
<bibRefCitation id="5C034B70697EFFADFCF6FC7B5898FC7E" author="Cook, D. R." box="[832,959,941,964]" pageId="57" pageNumber="58" pagination="1 - 418" refId="ref55236" refString="Cook, D. R. (1966) The water mites of Liberia. Memoirs of the American Entomological Institute, 6, 1 - 418." type="journal article" year="1966">Cook, 1966</bibRefCitation>
</taxonomicName>
, female holotype (FMC Lib 82): 121, leg-II; 122, leg-IV. Scale bar = 100 µm.
</paragraph>
</caption>
<paragraph id="382D3681697EFFADFF71FADD5AADFA9F" blockId="57.[151,1436,1051,1517]" box="[199,394,1291,1317]" pageId="57" pageNumber="58">
<emphasis id="0AE6EA93697EFFADFF71FADD5A37FA9F" bold="true" box="[199,272,1291,1317]" pageId="57" pageNumber="58">Male:</emphasis>
Unknown
</paragraph>
</subSubSection>
<subSubSection id="7088650A697EFFADFF71FAE55E7FFA57" pageId="57" pageNumber="58" type="discussion">
<paragraph id="382D3681697EFFADFF71FAE55E7FFA57" blockId="57.[151,1436,1051,1517]" pageId="57" pageNumber="58">
<emphasis id="0AE6EA93697EFFADFF71FAE55A65FAF7" bold="true" box="[199,322,1331,1357]" pageId="57" pageNumber="58">Remarks:</emphasis>
The two species of
<taxonomicName id="FF924D02697EFFADFD9DFAE559BAFAF6" authority="Goldschmidt, 2008" authorityName="Goldschmidt" authorityYear="2008" box="[555,669,1331,1356]" class="Arachnida" family="Anisitsiellidae" genus="Nilotonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="57" pageNumber="118" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Cookonia">
<emphasis id="0AE6EA93697EFFADFD9DFAE559BAFAF6" box="[555,669,1331,1356]" italics="true" pageId="57" pageNumber="58">Cookonia</emphasis>
</taxonomicName>
are separated by the degree of fusion of their ventral shield (postgenital plate and Cx-IV fused in
<taxonomicName id="FF924D02697EFFADFDA5FA8D5863FACE" authorityName="Goldschmidt" authorityYear="2008" box="[531,836,1371,1397]" class="Arachnida" family="Anisitsiellidae" genus="Nilotonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="anjozorobe" subGenus="Cookonia">
<emphasis id="0AE6EA93697EFFADFDA5FA8D5909FACE" box="[531,558,1371,1396]" italics="true" pageId="57" pageNumber="58">N.</emphasis>
(
<emphasis id="0AE6EA93697EFFADFD89FA8D5996FACE" box="[575,689,1371,1396]" italics="true" pageId="57" pageNumber="58">Cookonia</emphasis>
)
<emphasis id="0AE6EA93697EFFADFD77FA8D5863FACE" box="[705,836,1371,1396]" italics="true" pageId="57" pageNumber="58">anjozorobe</emphasis>
</taxonomicName>
, not fused in
<taxonomicName id="FF924D02697EFFADFC54FA8D5E35FACE" authorityName="Goldschmidt" authorityYear="2008" box="[994,1298,1371,1397]" class="Arachnida" family="Anisitsiellidae" genus="Nilotonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="avomonina" subGenus="Cookonia">
<emphasis id="0AE6EA93697EFFADFC54FA8D58DAFACE" box="[994,1021,1371,1396]" italics="true" pageId="57" pageNumber="58">N.</emphasis>
(
<emphasis id="0AE6EA93697EFFADFBBBFA8D5F58FACE" box="[1037,1151,1371,1396]" italics="true" pageId="57" pageNumber="58">Cookonia</emphasis>
)
<emphasis id="0AE6EA93697EFFADFB26FA8D5E35FACE" box="[1168,1298,1371,1396]" italics="true" pageId="57" pageNumber="58">avomonina</emphasis>
</taxonomicName>
), the extent of the female dorsal plate (in
<taxonomicName id="FF924D02697EFFADFE5AFA55583AFA26" authorityName="Goldschmidt" authorityYear="2008" box="[492,797,1411,1437]" class="Arachnida" family="Anisitsiellidae" genus="Nilotonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="anjozorobe" subGenus="Cookonia">
<emphasis id="0AE6EA93697EFFADFE5AFA55592FFA26" box="[492,520,1411,1436]" italics="true" pageId="57" pageNumber="58">N.</emphasis>
(
<emphasis id="0AE6EA93697EFFADFDAEFA5559ADFA26" box="[536,650,1411,1436]" italics="true" pageId="57" pageNumber="58">Cookonia</emphasis>
)
<emphasis id="0AE6EA93697EFFADFD2CFA55583AFA26" box="[666,797,1411,1436]" italics="true" pageId="57" pageNumber="58">anjozorobe</emphasis>
</taxonomicName>
not including Dgl-6 and Lgl-4) and proportions of the palps and legs (slenderer in
<taxonomicName id="FF924D02697EFFADFE55FA7D5831FA7E" authorityName="Goldschmidt" authorityYear="2008" box="[483,790,1451,1477]" class="Arachnida" family="Anisitsiellidae" genus="Nilotonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="avomonina" subGenus="Cookonia">
<emphasis id="0AE6EA93697EFFADFE55FA7D5AD9FA7E" box="[483,510,1451,1476]" italics="true" pageId="57" pageNumber="58">N.</emphasis>
(
<emphasis id="0AE6EA93697EFFADFDA6FA7D59A5FA7E" box="[528,642,1451,1476]" italics="true" pageId="57" pageNumber="58">Cookonia</emphasis>
)
<emphasis id="0AE6EA93697EFFADFD22FA7D5831FA7E" box="[660,790,1451,1476]" italics="true" pageId="57" pageNumber="58">avomonina</emphasis>
</taxonomicName>
). A sexual dimorphism similar to
<taxonomicName id="FF924D02697EFFADFB1BFA7D5BC1FA56" authorityName="Goldschmidt" authorityYear="2008" class="Arachnida" family="Anisitsiellidae" genus="Nilotonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="anjozorobe" subGenus="Cookonia">
<emphasis id="0AE6EA93697EFFADFB1BFA7D5FEEFA7E" box="[1197,1225,1451,1476]" italics="true" pageId="57" pageNumber="58">N.</emphasis>
(
<emphasis id="0AE6EA93697EFFADFB6CFA7D5E6BFA7E" box="[1242,1356,1451,1476]" italics="true" pageId="57" pageNumber="58">Cookonia</emphasis>
)
<emphasis id="0AE6EA93697EFFADFAE8FA7D5BC1FA56" italics="true" pageId="57" pageNumber="58">anjozorobe</emphasis>
</taxonomicName>
in the extent of the ventral shield (larger in male) is found in
<taxonomicName id="FF924D02697EFFADFC01FA055FEBFA57" authority="Cook, 1967" authorityName="Cook" authorityYear="1967" box="[951,1228,1491,1517]" class="Arachnida" family="Nilotoniidae" genus="Bharatonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="0AE6EA93697EFFADFC01FA055F1AFA56" box="[951,1085,1491,1516]" italics="true" pageId="57" pageNumber="58">Bharatonia</emphasis>
<bibRefCitation id="5C034B70697EFFADFBF2FA055FEBFA57" author="Cook" box="[1092,1228,1491,1517]" pageId="57" pageNumber="58" pagination="1 - 411" refId="ref55264" refString="Cook D. R. (1967) Water mites from India. Memoirs of the American Entomological Institute, 9, 1 - 411." type="journal article" year="1967">Cook, 1967</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="40857611697EFFADFAA2FA055E73FA57" box="[1300,1364,1491,1517]" name="India" pageId="57" pageNumber="58">India</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>