205 lines
22 KiB
XML
205 lines
22 KiB
XML
<document id="8486E480A356FF54D3F14DFA27DDC85B" ID-DOI="10.11646/zootaxa.5028.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5453967" ID-ZooBank="C678F0FE-1B62-4F34-8A66-449CF9806B50" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630655426570" checkinUser="plazi" docAuthor="Santos, Bruna S., Marques, Mariana P., Bauer, Aaron M. & Ceríaco, Luis M. P." docDate="2021" docId="B007F528FFC7FFCA16D7FE17FAC75D71" docLanguage="en" docName="zootaxa.5028.1.1.pdf" docOrigin="Zootaxa 5028 (1)" docSource="http://zoobank.org/c678f0fe-1b62-4f34-8a66-449cf9806b50" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Psammophis leopardinus Bocage 1887" docType="treatment" docVersion="3" lastPageNumber="32" masterDocId="4C3E8D50FFD8FFD51640FFA1FFD2597B" masterDocTitle="Herpetological results of Francisco Newton’s Zoological Expedition to Angola (1903 – 1906): a taxonomic revision and new records of a forgotten collection" masterLastPageNumber="80" masterPageNumber="1" pageNumber="32" updateTime="1699281765633" updateUser="plazi">
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<mods:title id="2BF8676C666D1595499BB9A504BC9060">Herpetological results of Francisco Newton’s Zoological Expedition to Angola (1903 – 1906): a taxonomic revision and new records of a forgotten collection</mods:title>
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<mods:namePart id="AD62765FDB89D9031F32A3C1214AE8FC">Santos, Bruna S.</mods:namePart>
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<mods:roleTerm id="32E0BDEF368496D1AB4CE13621680218">Author</mods:roleTerm>
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<mods:namePart id="FED3A72D3AA8546FB1E32CF500CC2DFF">Marques, Mariana P.</mods:namePart>
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<mods:nameIdentifier id="1AD1D11BE44055EEF55D45FEF41105EC" type="email">mptlmarques@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="652AFD95D26B47F35E710DFF6D0823D6">Author</mods:roleTerm>
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<mods:namePart id="C0D5777FD088E09BB1F017508D50623E">Bauer, Aaron M.</mods:namePart>
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<mods:nameIdentifier id="C16FA9CA73FF908146E4517022B2EECE" type="email">aaron.bauer@villanova.edu</mods:nameIdentifier>
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<mods:namePart id="23112E980495248898258C25FBA3733D">Ceríaco, Luis M. P.</mods:namePart>
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<mods:nameIdentifier id="A0FB42AA97081595A4BFC0773B1C439B" type="email">brunascesantos@gmail.com</mods:nameIdentifier>
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<mods:title id="230BA0131F9B392633919BC5C6DBBD3B">Zootaxa</mods:title>
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<mods:part id="E080B69FAD54734913EE423D80F8BBAA">
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<mods:date id="94B19DFB062326A524CCB5BAD0585544">2021</mods:date>
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<mods:number id="99A54618AD562A718F4E5303C5337B93">2021-09-03</mods:number>
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<mods:number id="C29B3D295DA20EBE0085F2C296399F0D">5028</mods:number>
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<subSubSection id="70B417B5FFC7FFCA16D7FE17FDA158AB" box="[151,627,438,465]" pageId="31" pageNumber="32" type="nomenclature">
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<paragraph id="3811443EFFC7FFCA16D7FE17FDA158AB" blockId="31.[151,627,438,465]" box="[151,627,438,465]" pageId="31" pageNumber="32">
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<heading id="6359F352FFC7FFCA16D7FE17FDA158AB" bold="true" box="[151,627,438,465]" fontSize="11" level="2" pageId="31" pageNumber="32" reason="4">
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<taxonomicName id="FFAE3FBDFFC7FFCA16D7FE17FDA158AB" authority="Bocage, 1887" authorityName="Bocage" authorityYear="1887" box="[151,627,438,465]" class="Tubothalamea" family="Ammodiscidae" genus="Psammophis" kingdom="Chromista" order="Spirillinida" pageId="31" pageNumber="32" phylum="Foraminifera" rank="species" species="leopardinus">
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<emphasis id="0ADA982CFFC7FFCA16D7FE17FDA158AB" bold="true" box="[151,627,438,465]" pageId="31" pageNumber="32">
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<emphasis id="0ADA982CFFC7FFCA16D7FE17FE1A58AA" bold="true" box="[151,456,438,465]" italics="true" pageId="31" pageNumber="32">Psammophis leopardinus</emphasis>
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Bocage, 1887
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="70B417B5FFC7FFCA16D7FE59FB875BBD" pageId="31" pageNumber="32" type="materials_examined">
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<paragraph id="3811443EFFC7FFCA16D7FE59FB875BBD" blockId="31.[151,1437,504,1035]" pageId="31" pageNumber="32">
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<emphasis id="0ADA982CFFC7FFCA16D7FE59FE325B69" bold="true" box="[151,480,504,530]" pageId="31" pageNumber="32">
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Material (
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<specimenCount id="2EA88FB7FFC7FFCA1757FE59FE015B69" box="[279,467,504,530]" pageId="31" pageNumber="32" type="generic">five specimens</specimenCount>
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):
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</emphasis>
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One juvenile specimen (MHNCUP/
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<collectionCode id="5EBFDCFBFFC7FFCA15D9FE59FC1E5B69" box="[921,972,504,530]" country="Turkmenistan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12935" lsid="urn:lsid:biocol.org:col:12935" name="Desert Experiment Station of the W.I.R." pageId="31" pageNumber="32" type="Herbarium">REP</collectionCode>
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171, formerly UP-MHNFCP-094982), collected from África Ocidental [Occidental Africa] (impossible to georeference), date unknown;
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<materialsCitation id="88C64E63FFC7FFCA136EFDBCFD665B05" collectionCode="REP" elevation="45" location="Coroca" pageId="31" pageNumber="32" specimenCode="REP 173" specimenCount="1" specimenCount-adult="1" stateProvince="Rivers">
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<specimenCount id="2EA88FB7FFC7FFCA136EFDBCFA4E5B4D" box="[1326,1436,540,567]" pageId="31" pageNumber="32" type="adult">one adult</specimenCount>
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specimen (MHNCUP/
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<specimenCode id="6808EC45FFC7FFCA17CFFDE1FE265B21" box="[399,500,576,602]" collectionCode="REP" country="Turkmenistan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12935" lsid="urn:lsid:biocol.org:col:12935" name="Desert Experiment Station of the W.I.R." pageId="31" pageNumber="32" type="Herbarium">REP 173</specimenCode>
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, formerly UP-MHNFCP-094988;
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<figureCitation id="A09558BBFFC7FFCA1531FDE1FC005B20" box="[881,978,576,603]" captionStart="FIGURE S" captionStartId="74.[151,250,1069,1093]" captionTargetBox="[170,1416,164,1044]" captionTargetId="figure-33@74.[170,1416,164,1044]" captionTargetPageId="74" captionText="FIGURE S65. Specimen of Psammophis leopardinus (MHNCUP/REP 173), from Rio Coróca. Photo by Bruna Santos." figureDoi="http://doi.org/10.5281/zenodo.5454113" httpUri="https://zenodo.org/record/5454113/files/figure.png" pageId="31" pageNumber="32">Fig. S65</figureCitation>
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), collected from
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<collectingRegion id="FA6A8ADCFFC7FFCA12CCFDE1FB665B21" box="[1164,1204,576,602]" country="Nigeria" name="Rivers" pageId="31" pageNumber="32">Rio</collectingRegion>
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<location id="3D7112E5FFC7FFCA12FAFDE1FAD95B21" LSID="urn:lsid:plazi:treatment:B007F528FFC7FFCA16D7FE17FAC75D71:3D7112E5FFC7FFCA12FAFDE1FAD95B21" box="[1210,1291,576,602]" name="Coroca" pageId="31" pageNumber="32" stateProvince="Rivers">Coróca</location>
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[-15.78333º, 12.06667º,
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<elevation id="B383A30DFFC7FFCA1755FDC5FE575B05" box="[277,389,611,639]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="31" pageNumber="32" unit="m" value="45.0">
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<quantity id="FF56E9DBFFC7FFCA1755FDC5FE995B04" box="[277,331,612,639]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="31" pageNumber="32" unit="m" value="45.0">45 m</quantity>
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a.s.l.
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</elevation>
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]
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<collectingRegion id="FA6A8ADCFFC7FFCA17D2FDC5FD875B05" box="[402,597,612,638]" country="Angola" name="Namibe" pageId="31" pageNumber="32">Namibe Province</collectingRegion>
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, in 1904
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</materialsCitation>
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;
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<materialsCitation id="88C64E63FFC7FFCA14FEFDC4FCC25BD9" collectionCode="REP" pageId="31" pageNumber="32" specimenCode="REP 174" specimenCount="1" specimenCount-adult="1" stateProvince="Rivers">
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||
<specimenCount id="2EA88FB7FFC7FFCA14FEFDC4FCF15B05" box="[702,803,612,639]" pageId="31" pageNumber="32" type="adult">one adult</specimenCount>
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specimen (MHNCUP/
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<specimenCode id="6808EC45FFC7FFCA125EFDC5FB505B05" box="[1054,1154,612,638]" collectionCode="REP" country="Turkmenistan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12935" lsid="urn:lsid:biocol.org:col:12935" name="Desert Experiment Station of the W.I.R." pageId="31" pageNumber="32" type="Herbarium">REP 174</specimenCode>
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||
, formerly UP-MHNFCP- 094989), collected from unknow locality, date unknown
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</materialsCitation>
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||
;
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||
<materialsCitation id="88C64E63FFC7FFCA155BFD28FB835BBD" collectionCode="REP" pageId="31" pageNumber="32" specimenCode="REP 176-177" specimenCount="1" specimenCount-adult="1" stateProvince="Rivers">
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<specimenCount id="2EA88FB7FFC7FFCA155BFD28FC575BD9" box="[795,901,648,675]" pageId="31" pageNumber="32" type="adult">two adult</specimenCount>
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specimens (MHNCUP/
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<specimenCode id="6808EC45FFC7FFCA12D0FD29FAFC5BD9" box="[1168,1326,648,674]" collectionCode="REP" country="Turkmenistan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12935" lsid="urn:lsid:biocol.org:col:12935" name="Desert Experiment Station of the W.I.R." pageId="31" pageNumber="32" type="Herbarium">REP 176–177</specimenCode>
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, formerly UP-MHNFCP-017422 and 040697), collected from unknown locality, date unknown
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</materialsCitation>
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.
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="70B417B5FFC7FFCA1687FD71FAC75D71" pageId="31" pageNumber="32" type="discussion">
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<paragraph id="3811443EFFC7FFCA1687FD71FAC75D71" blockId="31.[151,1437,504,1035]" pageId="31" pageNumber="32">
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<emphasis id="0ADA982CFFC7FFCA1687FD71FE805B91" bold="true" box="[199,338,720,746]" pageId="31" pageNumber="32">Comments:</emphasis>
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<taxonomicName id="FFAE3FBDFFC7FFCA1717FD71FDA35B91" authorityName="Bocage" authorityYear="1887" box="[343,625,720,746]" class="Tubothalamea" family="Ammodiscidae" genus="Psammophis" kingdom="Chromista" order="Spirillinida" pageId="31" pageNumber="32" phylum="Foraminifera" rank="species" species="leopardinus">
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<emphasis id="0ADA982CFFC7FFCA1717FD71FDA35B91" box="[343,625,720,746]" italics="true" pageId="31" pageNumber="32">Psammophis leopardinus</emphasis>
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</taxonomicName>
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is found in dry savanna and semi-desert habitats of southwestern
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<collectingCountry id="40B904AEFFC7FFCA1303FD71FA4A5B91" box="[1347,1432,720,746]" name="Angola" pageId="31" pageNumber="32">Angola</collectingCountry>
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, as well as central and northern
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<collectingCountry id="40B904AEFFC7FFCA1440FD55FDB25A75" box="[512,608,756,782]" name="Namibia" pageId="31" pageNumber="32">Namibia</collectingCountry>
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||
(
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<bibRefCitation id="5C3F39CFFFC7FFCA1431FD55FC8C5A75" author="Marques, M. P. & Ceriaco, L. M. P. & Blackburn, D. C. & Bauer, A. M." box="[625,862,756,782]" pageId="31" pageNumber="32" pagination="1 - 501" refId="ref38562" refString="Marques, M. P., Ceriaco, L. M. P., Blackburn, D. C. & Bauer, A. M. (2018) Diversity and distribution of the amphibians and terrestrial reptiles of Angola atlas of historical and bibliographic records (1840 - 2017). Proceedings of the California Academy of Sciences, Series 4, 65 (Supplement 2), 1 - 501." type="journal article" year="2018">
|
||
Marques
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||
<emphasis id="0ADA982CFFC7FFCA149EFD54FCCB5A75" box="[734,793,756,782]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
|
||
2018
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||
</bibRefCitation>
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||
). It is currently treated as a full species (
|
||
<bibRefCitation id="5C3F39CFFFC7FFCA1374FD55FF015A49" author="Broadley, D. G." pageId="31" pageNumber="32" pagination="83 - 119" refId="ref34809" refString="Broadley, D. G. (2002) A review of the species of Psammophis Boie found south of latitude 12 ° S (Serpentes: Psammophiinae). African Journal of Herpetology, 51 (2), 83 - 119. https: // doi. org / 10.1080 / 21564574.2002.9635466" type="journal article" year="2002">Broadley 2002</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="5C3F39CFFFC7FFCA169DFCB9FE495A49" author="Kelly, C. M. R. & Barker, N. P. & Villet, M. H. & Broadley, D. G. & Branch, W. R." box="[221,411,792,818]" pageId="31" pageNumber="32" pagination="1045 - 1060" refId="ref38161" refString="Kelly, C. M. R., Barker, N. P., Villet, M. H., Broadley, D. G. & Branch, W. R. (2008) The snake family Psammophiidae (Reptilia: Serpentes): phylogenetics and species delimitation in the African sand snakes (Psammophis Boie, 1825) and allied genera. Molecular Phylogenetics and Evolution, 47, 1045 - 1060. https: // doi. org / 10.1016 / j. ympev. 2008.03.025" type="journal article" year="2008">
|
||
Kelly
|
||
<emphasis id="0ADA982CFFC7FFCA1762FCB8FE885A49" box="[290,346,792,818]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
|
||
2008
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="5C3F39CFFFC7FFCA17E6FCB9FD505A49" author="Wallach, V. & Williams, K. L. & Boundy, J." box="[422,642,792,818]" pageId="31" pageNumber="32" refId="ref40677" refString="Wallach, V., Williams, K. L. & Boundy, J. (2014) Snakes of the World: A Catalogue of Living and Extinct Species. CRC Press, Boca Raton, Florida, xxvi + [2] + 1209 pp." type="book" year="2014">
|
||
Wallach
|
||
<emphasis id="0ADA982CFFC7FFCA1448FCB8FDED5A49" box="[520,575,792,818]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
), after being initially described as a subspecies of
|
||
<taxonomicName id="FFAE3FBDFFC7FFCA12EFFCB9FE8A5A2C" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Tubothalamea" family="Ammodiscidae" genus="Psammophis" kingdom="Chromista" order="Spirillinida" pageId="31" pageNumber="32" phylum="Foraminifera" rank="species" species="sibilans">
|
||
<emphasis id="0ADA982CFFC7FFCA12EFFCB9FA4E5A49" box="[1199,1436,792,818]" italics="true" pageId="31" pageNumber="32">Psammophis sibilans</emphasis>
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||
(Linnaeus, 1758)
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||
</taxonomicName>
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||
by
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<bibRefCitation id="5C3F39CFFFC7FFCA17C6FC9CFDEC5A2C" author="Bocage, J. V. B." box="[390,574,828,855]" pageId="31" pageNumber="32" pagination="201 - 208" refId="ref34158" refString="Bocage, J. V. B. (1887 b) Melanges erpetologiques. IV. Reptiles du dernier voyage de MM. Capello et Ivens a travers l'Afrique. Jornal de Sciencias Mathematicas, Physicas e Naturaes, 11, 201 - 208." type="journal article" year="1887">Bocage (1887b)</bibRefCitation>
|
||
. The aforementioned specimens were previously identified as “
|
||
<taxonomicName id="FFAE3FBDFFC7FFCA1354FC9CFA5B5A2D" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1300,1417,828,854]" class="Tubothalamea" family="Ammodiscidae" genus="Psammophis" kingdom="Chromista" order="Spirillinida" pageId="31" pageNumber="32" phylum="Foraminifera" rank="species" species="sibilans">
|
||
<emphasis id="0ADA982CFFC7FFCA1354FC9CFA5B5A2D" box="[1300,1417,828,854]" italics="true" pageId="31" pageNumber="32">P. sibilans</emphasis>
|
||
</taxonomicName>
|
||
”, with the exception of MHNCUP/REP 171 which was labeled as “
|
||
<taxonomicName id="FFAE3FBDFFC7FFCA1539FCC0FBAA5A01" box="[889,1144,864,890]" class="Tubothalamea" family="Ammodiscidae" genus="Psammophis" kingdom="Chromista" order="Spirillinida" pageId="31" pageNumber="32" phylum="Foraminifera" rank="subSpecies" species="sibilans" subSpecies="leopardinus">
|
||
<emphasis id="0ADA982CFFC7FFCA1539FCC0FBAA5A01" box="[889,1144,864,890]" italics="true" pageId="31" pageNumber="32">P. sibilans leopardinus</emphasis>
|
||
</taxonomicName>
|
||
”.
|
||
<bibRefCitation id="5C3F39CFFFC7FFCA12D2FCC1FAEF5A01" author="Ferreira, J. B." box="[1170,1341,864,890]" pageId="31" pageNumber="32" pagination="111 - 117" refId="ref37152" refString="Ferreira, J. B. (1904) Reptis e amphibios de Angola da regiao ao norte do Quanza (Colleccao Newton 1903). Jornal de Sciencias Mathematicas Physicas e Naturaes, Segunda Serie, 7 (25), 111 - 117." type="journal article" year="1904">Ferreira (1904)</bibRefCitation>
|
||
cited an unspecified number of adult and juvenile specimens of “Bocage’s var D.” of
|
||
<taxonomicName id="FFAE3FBDFFC7FFCA15AAFC24FB8E5AE5" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1002,1116,900,926]" class="Tubothalamea" family="Ammodiscidae" genus="Psammophis" kingdom="Chromista" order="Spirillinida" pageId="31" pageNumber="32" phylum="Foraminifera" rank="species" species="sibilans">
|
||
<emphasis id="0ADA982CFFC7FFCA15AAFC24FB8E5AE5" box="[1002,1116,900,926]" italics="true" pageId="31" pageNumber="32">P. sibilans</emphasis>
|
||
</taxonomicName>
|
||
from “Cazengo”, “
|
||
<collectingCountry id="40B904AEFFC7FFCA1370FC25FA415AE5" box="[1328,1427,900,926]" name="Angola" pageId="31" pageNumber="32">Cabinda</collectingCountry>
|
||
” (in error, Newton did not travel to the
|
||
<collectingCountry id="40B904AEFFC7FFCA1401FC09FD4C5AB9" box="[577,670,936,962]" name="Angola" pageId="31" pageNumber="32">Cabinda</collectingCountry>
|
||
enclave) and “Chingo”.
|
||
<bibRefCitation id="5C3F39CFFFC7FFCA15F3FC08FB565AB9" author="Trape, J. - F. & Crochet, P. - A. & Broadley, D. G. & Sourouille, P. & Mane, Y. & Burger, M. & Bohme, W. & Saleh, M. & Karan, A. & Lanza, B. & Mediannikov, O." box="[947,1156,936,963]" pageId="31" pageNumber="32" pagination="61 - 91" refId="ref40435" refString="Trape, J. - F., Crochet, P. - A., Broadley, D. G., Sourouille, P., Mane, Y., Burger, M., Bohme, W., Saleh, M., Karan, A., Lanza, B. & Mediannikov, O. (2019) On the Psammophis sibilans group (Serpentes, Lamprophiidae, Psammophiinae) north of 12 ° S, with the description of a new species from West Africa. Bonn zoological Bulletin, 68 (1), 61 - 91. https: // doi. org / 10.20363 / BZB- 2019.68.1.061" type="journal article" year="2019">
|
||
Trape
|
||
<emphasis id="0ADA982CFFC7FFCA15BAFC08FBE05AB9" box="[1018,1074,936,962]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
|
||
(2019)
|
||
</bibRefCitation>
|
||
referred to the published specimens as
|
||
<taxonomicName id="FFAE3FBDFFC7FFCA1778FC6CFE0E5A9D" authorityName="Bocage" authorityYear="1887" box="[312,476,972,998]" class="Tubothalamea" family="Ammodiscidae" genus="Psammophis" kingdom="Chromista" order="Spirillinida" pageId="31" pageNumber="32" phylum="Foraminifera" rank="species" species="leopardinus">
|
||
<emphasis id="0ADA982CFFC7FFCA1778FC6CFE0E5A9D" box="[312,476,972,998]" italics="true" pageId="31" pageNumber="32">P. leopardinus</emphasis>
|
||
</taxonomicName>
|
||
and noted that they were presumably lost in the MBL fire, which we here refute, whereas
|
||
<bibRefCitation id="5C3F39CFFFC7FFCA16BAFC50FE3F5D71" author="Marques, M. P. & Ceriaco, L. M. P. & Blackburn, D. C. & Bauer, A. M." box="[250,493,1008,1035]" pageId="31" pageNumber="32" pagination="1 - 501" refId="ref38562" refString="Marques, M. P., Ceriaco, L. M. P., Blackburn, D. C. & Bauer, A. M. (2018) Diversity and distribution of the amphibians and terrestrial reptiles of Angola atlas of historical and bibliographic records (1840 - 2017). Proceedings of the California Academy of Sciences, Series 4, 65 (Supplement 2), 1 - 501." type="journal article" year="2018">
|
||
Marques
|
||
<emphasis id="0ADA982CFFC7FFCA1723FC50FE495D71" box="[355,411,1008,1034]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
|
||
(2018)
|
||
</bibRefCitation>
|
||
had previously treated them as
|
||
<taxonomicName id="FFAE3FBDFFC7FFCA1511FC51FAC35D71" authority="Peters, 1882" authorityName="Peters" authorityYear="1882" box="[849,1297,1008,1035]" class="Tubothalamea" family="Ammodiscidae" genus="Psammophis" kingdom="Chromista" order="Spirillinida" pageId="31" pageNumber="32" phylum="Foraminifera" rank="species" species="mossambicus">
|
||
<emphasis id="0ADA982CFFC7FFCA1511FC51FBAE5D71" box="[849,1148,1008,1034]" italics="true" pageId="31" pageNumber="32">Psammophis mossambicus</emphasis>
|
||
Peters, 1882
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |