197 lines
21 KiB
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197 lines
21 KiB
XML
<document id="66093257C7370E5B1AC0BE33CC928984" ID-CLB-Dataset="188710" ID-DOI="10.11646/zootaxa.5209.2.2" ID-GBIF-Dataset="87405f1b-232d-4e0c-a7d8-4d6824a801c5" ID-ISSN="1175-5326" ID-Zenodo-Dep="7325861" ID-ZooBank="40F0B633-691B-404E-95B3-0C23F5133791" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1668583694219" checkinUser="plazi" docAuthor="Zumbado Echavarria, Marco A., Barrantes Barrantes, Edwin A., Helmick, Ericka E., Bartlett, Charles R. & Bahder, Brian W." docDate="2022" docId="154B032E3561427AF2F47A37FA300DCB" docLanguage="en" docName="zootaxa.5209.2.2.pdf" docOrigin="Zootaxa 5209 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tico Bahder & Bartlett 2021" docType="key" docVersion="7" lastPageNumber="197" masterDocId="E9727B5635684270F2637D0DFFAC0C24" masterDocTitle="A new species of planthopper in the genus Tico (Hemiptera: Auchenorrhyncha: Derbidae) on palms in Costa Rica" masterLastPageNumber="198" masterPageNumber="187" pageNumber="196" updateTime="1699411561995" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="229ADB3BBE0C462925F9210B1B990DFD">
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<mods:title id="C8407048343EA7960CB5178CF6C3B336">A new species of planthopper in the genus Tico (Hemiptera: Auchenorrhyncha: Derbidae) on palms in Costa Rica</mods:title>
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<mods:name id="7C2D81E17B54CE0EE7FEC0BB8F24365D" type="personal">
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<mods:namePart id="E35ED549D9543E6AEBE0A70235AE4FC9">Zumbado Echavarria, Marco A.</mods:namePart>
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<mods:affiliation id="D3C6970D05BF1699F312C8EA9AC3DAF4">Universidad de Costa Rica-Sede San Ramón, Departamento de Ciencias Naturales, de la Iglesia el Tremedal 400 mts al Oeste carretera hacia San Pedro, San Ramón, Alajuela, Costa Rica.</mods:affiliation>
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<mods:namePart id="CA64E55CE095E8BAC61C83AC96C0A139">Barrantes Barrantes, Edwin A.</mods:namePart>
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<mods:affiliation id="15B6328B872B2C5FAF9B1C20442172C0">Universidad de Costa Rica-Sede San Ramón, Departamento de Ciencias Naturales, de la Iglesia el Tremedal 400 mts al Oeste carretera hacia San Pedro, San Ramón, Alajuela, Costa Rica.</mods:affiliation>
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<mods:namePart id="C5DFC91E31347C63F366E72A4FBA2F13">Helmick, Ericka E.</mods:namePart>
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<mods:affiliation id="032A1A074A891EFC7C9F6EC96DF18CAA">University of Florida, Department of Entomology and Nematology-Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314 - 7719, USA. E-mail: ehelmick @ ufl. edu</mods:affiliation>
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<mods:namePart id="BD40E784513E6585D30388992EFC8C5A">Bartlett, Charles R.</mods:namePart>
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<mods:affiliation id="75B916B39402AED2A3A22BCBD56B3778">University of Delaware, Department of Entomology and Wildlife Ecology, 250 Townsend Hall, Newark, DE 19716 - 2160, USA.</mods:affiliation>
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<mods:namePart id="F52C3FF35652882DAE63AB3B4661736B">Bahder, Brian W.</mods:namePart>
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<mods:affiliation id="1EC9C0010D1DD50DF13707A58B61E39E">University of Florida, Department of Entomology and Nematology-Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314 - 7719, USA. bbahder @ ufl. edu; https: // orcid. org / 0000 - 0002 - 1118 - 4832</mods:affiliation>
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<mods:nameIdentifier id="8DC0AE0319A65D26EDD982EC4723B915" type="email">bbahder@ufl.edu</mods:nameIdentifier>
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<mods:title id="16AD6C7E130DF778340026EB6D50F400">Zootaxa</mods:title>
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<mods:date id="B44707409138A36C0AFB97D33E0F7D36">2022</mods:date>
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<mods:number id="47426D709D0558F3FF8DDC64183A41CA">2022-11-16</mods:number>
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<treatment id="154B032E3561427AF2F47A37FA300DCB" ID-DOI="http://doi.org/10.5281/zenodo.7330845" ID-GBIF-Taxon="205079188" ID-Zenodo-Dep="7330845" LSID="urn:lsid:plazi:treatment:154B032E3561427AF2F47A37FA300DCB" httpUri="http://treatment.plazi.org/id/154B032E3561427AF2F47A37FA300DCB" lastPageId="10" lastPageNumber="197" pageId="9" pageNumber="196">
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<subSubSection id="D5F8E1B335614279F2F47A37FE0F0B70" box="[151,419,1850,1876]" pageId="9" pageNumber="196" type="nomenclature">
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<paragraph id="9D5DB23835614279F2F47A37FE0F0B70" blockId="9.[151,419,1850,1876]" box="[151,419,1850,1876]" pageId="9" pageNumber="196">
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<heading id="C615055435614279F2F47A37FE0F0B70" bold="true" box="[151,419,1850,1876]" fontSize="11" level="1" pageId="9" pageNumber="196" reason="1">
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<emphasis id="AF966E2A35614279F2F47A37FE0F0B70" bold="true" box="[151,419,1850,1876]" pageId="9" pageNumber="196">
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Key to species of
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<taxonomicName id="5AE2C9BB35614279F30D7A37FE0F0B70" ID-CoL="9DNFB" authorityName="Bahder & Bartlett" authorityYear="2021" box="[366,419,1850,1876]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="196" phylum="Arthropoda" rank="genus">
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<emphasis id="AF966E2A35614279F30D7A37FE0F0B70" bold="true" box="[366,419,1850,1876]" italics="true" pageId="9" pageNumber="196">Tico</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="D5F8E1B33561427AF2F47A72FA300DCB" lastPageId="10" lastPageNumber="197" pageId="9" pageNumber="196" type="key">
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<keyStep id="2616AA9D35614279F2F47A72FA300BEB" pageId="9" pageNumber="196">
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<paragraph id="9D5DB23835614279F2F47A72FA300BB0" blockId="9.[151,1437,1919,2029]" box="[151,1436,1919,1941]" pageId="9" pageNumber="196">
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<keyLead id="2613110D35614279F2F47A72FA300BB0" box="[151,1436,1919,1941]" pageId="9" pageNumber="196">
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1. Body brownish orange to fuscous, forewings dark, reddish, legs pale (
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<figureCitation id="05D9AEBD35614279F1E17A72FC170BB0" box="[898,955,1919,1941]" captionStart="FIGURE 2" captionStartId="4.[152,255,1373,1397]" captionTargetBox="[151,1436,273,1350]" captionTargetId="figure-65@4.[151,1436,273,1350]" captionTargetPageId="4" captionText="FIGURE 2. Adult male habitus of Tico villablanca sp. n.; A) lateral view, and B) dorsal view; scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7325867" httpUri="https://zenodo.org/record/7325867/files/figure.png" pageId="9" pageNumber="196">Fig. 2</figureCitation>
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)...................................... 2
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</keyLead>
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</paragraph>
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<paragraph id="9D5DB23835614279F2F47A90FA300BEB" blockId="9.[151,1437,1919,2029]" pageId="9" pageNumber="196">
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<keyLead id="2613110D35614279F2F47A90FA300BEB" pageId="9" pageNumber="196">
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- Body pale (body yellowish-white), wings clear except for varied fuscous to black markings (
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<bibRefCitation id="F973CFC935614279F6387A91FABC0B95" author="Bahder, B. W. & Echavarria, M. A. Z. & Barrantes, E. A. B. & Helmick, E. E. & Bartlett, C. R." box="[1115,1296,1948,1970]" pageId="9" pageNumber="196" pagination="369 - 392" refId="ref5842" refString="Bahder, B. W., Echavarria, M. A. Z., Barrantes, E. A. B., Helmick, E. E. & Bartlett, C. R. (2021) A new genus and two new species of planthopper in the tribe Cenchreini (Hemiptera: Auchenorrhyncha: Derbidae) from lowland tropical rainforest in Costa Rica. Zootaxa, 4908 (3), 369 - 392. https: // doi. org / 10.11646 / zootaxa. 4908.3.3" type="journal article" year="2021">
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Bahder
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<emphasis id="AF966E2A35614279F6CB7A90FB7E0B95" box="[1192,1234,1948,1970]" italics="true" pageId="9" pageNumber="196">et al</emphasis>
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. 2021
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</bibRefCitation>
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, figs 4, 12)..................................................................................................... 3
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="2616AA9D3561427AF2F47ADAFA300CCC" lastPageId="10" lastPageNumber="197" pageId="9" pageNumber="196">
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<paragraph id="9D5DB2383561427AF2F47ADAFA300C89" blockId="9.[151,1437,1919,2029]" lastBlockId="10.[151,1437,151,496]" lastPageId="10" lastPageNumber="197" pageId="9" pageNumber="196">
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<keyLead id="2613110D3561427AF2F47ADAFA300C89" lastPageId="10" lastPageNumber="197" pageId="9" pageNumber="196">
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2. Frons and vertex lacking sensorial pits, face (except genae below eyes) pale (
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<figureCitation id="05D9AEBD35614279F1897ADAFB8B0BC8" box="[1002,1063,2007,2029]" captionStart="FIGURE 3" captionStartId="5.[152,255,926,950]" captionTargetBox="[156,1430,390,895]" captionTargetId="figure-122@5.[152,1436,381,903]" captionTargetPageId="5" captionText="FIGURE 3. Adult Tico villablanca sp. n.; A) head, pronotum and mesonotum dorsal view, B) head, pronotum, and mesonotum right lateral view, and C) head and pronotum frontal view; scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7325869" httpUri="https://zenodo.org/record/7325869/files/figure.png" pageId="9" pageNumber="196">Fig. 3</figureCitation>
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); aedeagus bearing large, heart-shape flanges at apex, angled cephalad (
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<figureCitation id="05D9AEBD3562427AF0457D9AFDCC0C89" box="[550,608,151,173]" captionStart="FIGURE 7" captionStartId="8.[152,255,1424,1448]" captionTargetBox="[151,1437,181,1400]" captionTargetId="figure-24@8.[151,1437,181,1400]" captionTargetPageId="8" captionText="FIGURE 7. Aedeagus of Tico villablanca sp. n.; A) right lateral view, B) left lateral view, and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7325879" httpUri="https://zenodo.org/record/7325879/files/figure.png" pageId="10" pageNumber="197">Fig. 7</figureCitation>
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)...................................................
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<taxonomicName id="5AE2C9BB3562427AF6BE7D95FACC0C88" authority="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder, 2022" authorityName="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder" authorityYear="2022" box="[1245,1376,151,173]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="197" phylum="Arthropoda" rank="species" species="villablanca" status="sp. nov.">
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<emphasis id="AF966E2A3562427AF6BE7D95FACC0C88" box="[1245,1376,151,173]" italics="true" pageId="10" pageNumber="197">T. villablanca</emphasis>
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</taxonomicName>
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<emphasis id="AF966E2A3562427AF7057D95FA300C89" bold="true" box="[1382,1436,152,173]" pageId="10" pageNumber="197">
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<taxonomicNameLabel id="B4A5D3513562427AF7057D95FA300C89" box="[1382,1436,152,173]" pageId="10" pageNumber="197" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</keyLead>
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</paragraph>
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<paragraph id="9D5DB2383562427AF2F47DB8FA300CCC" blockId="10.[151,1437,151,496]" pageId="10" pageNumber="197">
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<keyLead id="2613110D3562427AF2F47DB8FA300CCC" pageId="10" pageNumber="197">
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- Frons and vertex with many sensorial pits of variable size; face fuscous red (
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<bibRefCitation id="F973CFC93562427AF1D07DB8FBC80CEE" author="Bahder, B. W. & Echavarria, M. A. Z. & Barrantes, E. A. B. & Helmick, E. E. & Bartlett, C. R." box="[947,1124,181,203]" pageId="10" pageNumber="197" pagination="283 - 294" refId="ref5934" refString="Bahder, B. W., Echavarria, M. A. Z., Barrantes, E. A. B., Helmick, E. E. & Bartlett, C. R. (2022) A new species of planthopper in the genus Tico (Hemiptera: Auchenorrhyncha: Derbidae) on palms from lowland tropical rainforest in Costa Rica. Zootaxa, 5125 (3), 283 - 294. https: // doi. org / 10.11646 / zootaxa. 5125.3.2" type="journal article" year="2022">
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Bahder
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<emphasis id="AF966E2A3562427AF19C7DB8FB840CEE" box="[1023,1064,181,202]" italics="true" pageId="10" pageNumber="197">et al</emphasis>
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. 2022
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</bibRefCitation>
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, fig. 3); aedeagus with serrations on dorsal and ventral margins
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<bibRefCitation id="F973CFC93562427AF0667DDFFD140CC3" author="Bahder, B. W. & Echavarria, M. A. Z. & Barrantes, E. A. B. & Helmick, E. E. & Bartlett, C. R." box="[517,696,210,232]" pageId="10" pageNumber="197" pagination="283 - 294" refId="ref5934" refString="Bahder, B. W., Echavarria, M. A. Z., Barrantes, E. A. B., Helmick, E. E. & Bartlett, C. R. (2022) A new species of planthopper in the genus Tico (Hemiptera: Auchenorrhyncha: Derbidae) on palms from lowland tropical rainforest in Costa Rica. Zootaxa, 5125 (3), 283 - 294. https: // doi. org / 10.11646 / zootaxa. 5125.3.2" type="journal article" year="2022">
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Bahder
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<emphasis id="AF966E2A3562427AF0337DDEFDD60CC3" box="[592,634,210,232]" italics="true" pageId="10" pageNumber="197">et al</emphasis>
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. 2022
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</bibRefCitation>
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, fig. 7)................................................
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<taxonomicName id="5AE2C9BB3562427AF7287DDEFA300CCC" box="[1355,1436,211,232]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="197" phylum="Arthropoda" rank="species" species="sierra">
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<emphasis id="AF966E2A3562427AF7287DDEFA300CCC" box="[1355,1436,211,232]" italics="true" pageId="10" pageNumber="197">T. sierra</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="2616AA9D3562427AF2F47DFDFA300D5E" pageId="10" pageNumber="197">
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<paragraph id="9D5DB2383562427AF2F47DFDFA300D06" blockId="10.[151,1437,151,496]" pageId="10" pageNumber="197">
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<keyLead id="2613110D3562427AF2F47DFDFA300D06" pageId="10" pageNumber="197">
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3. Forewing black spot at base and 4 large blacks spots distally on trailing margin (the dorsal part of the wing when in repose); head with dark marking anterior to eyes (
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<bibRefCitation id="F973CFC93562427AF00E7C00FC8E0D06" author="Bahder, B. W. & Echavarria, M. A. Z. & Barrantes, E. A. B. & Helmick, E. E. & Bartlett, C. R." box="[621,802,269,291]" pageId="10" pageNumber="197" pagination="369 - 392" refId="ref5842" refString="Bahder, B. W., Echavarria, M. A. Z., Barrantes, E. A. B., Helmick, E. E. & Bartlett, C. R. (2021) A new genus and two new species of planthopper in the tribe Cenchreini (Hemiptera: Auchenorrhyncha: Derbidae) from lowland tropical rainforest in Costa Rica. Zootaxa, 4908 (3), 369 - 392. https: // doi. org / 10.11646 / zootaxa. 4908.3.3" type="journal article" year="2021">
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Bahder
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<emphasis id="AF966E2A3562427AF0D97C00FD480D06" box="[698,740,269,290]" italics="true" pageId="10" pageNumber="197">et al</emphasis>
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. 2021
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</bibRefCitation>
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, figs 12, 13)......................................... 4
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</keyLead>
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</paragraph>
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<paragraph id="9D5DB2383562427AF2F47C26FA300D5E" blockId="10.[151,1437,151,496]" pageId="10" pageNumber="197">
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<keyLead id="2613110D3562427AF2F47C26FA300D5E" pageId="10" pageNumber="197">
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- Forewing without series of 4 dark spots, markings limited to well-separated single spots (including one in coastal cell, one near fork of CuA, and one in cell C3, proximad of an intramedial crossvein; head in front of eyes pale (
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<bibRefCitation id="F973CFC93562427AF6FD7C4AFAF90D79" author="Bahder, B. W. & Echavarria, M. A. Z. & Barrantes, E. A. B. & Helmick, E. E. & Bartlett, C. R." box="[1182,1365,327,349]" pageId="10" pageNumber="197" pagination="369 - 392" refId="ref5842" refString="Bahder, B. W., Echavarria, M. A. Z., Barrantes, E. A. B., Helmick, E. E. & Bartlett, C. R. (2021) A new genus and two new species of planthopper in the tribe Cenchreini (Hemiptera: Auchenorrhyncha: Derbidae) from lowland tropical rainforest in Costa Rica. Zootaxa, 4908 (3), 369 - 392. https: // doi. org / 10.11646 / zootaxa. 4908.3.3" type="journal article" year="2021">
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Bahder
|
|
<emphasis id="AF966E2A3562427AF6887C45FABA0D78" box="[1259,1302,327,349]" italics="true" pageId="10" pageNumber="197">et al</emphasis>
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|
. 2021
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</bibRefCitation>
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, figs 4, 5).......................................................................................
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<taxonomicName id="5AE2C9BB3562427AF76C7C68FA300D5E" ID-CoL="9FNPB" authorityName="Bahder & Bartlett" authorityYear="2021" box="[1295,1436,357,378]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="197" phylum="Arthropoda" rank="species" species="emmettcarri">
|
|
<emphasis id="AF966E2A3562427AF76C7C68FA300D5E" box="[1295,1436,357,378]" italics="true" pageId="10" pageNumber="197">T. emmettcarri</emphasis>
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</taxonomicName>
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|
</keyLead>
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|
</paragraph>
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|
</keyStep>
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|
<keyStep id="2616AA9D3562427AF2F47C8FFA300DCB" pageId="10" pageNumber="197">
|
|
<paragraph id="9D5DB2383562427AF2F47C8FFA300D91" blockId="10.[151,1437,151,496]" pageId="10" pageNumber="197">
|
|
<keyLead id="2613110D3562427AF2F47C8FFA300D91" pageId="10" pageNumber="197">
|
|
4. Ventral processes of first gonapophysis of ovipositor not twisted, apex of dorsal process angled dorsad, bifid; pregenital plate pyramid-shaped (Fennah 1951, fig. 11H)...........................................................
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|
<taxonomicName id="5AE2C9BB3562427AF7557CADFA300D91" ID-CoL="9FNPD" baseAuthorityName="Fennah" baseAuthorityYear="1952" box="[1334,1436,416,437]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="197" phylum="Arthropoda" rank="species" species="sororius">
|
|
<emphasis id="AF966E2A3562427AF7557CADFA300D91" box="[1334,1436,416,437]" italics="true" pageId="10" pageNumber="197">T. sororius</emphasis>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="9D5DB2383562427AF2F47CB0FA300DCB" blockId="10.[151,1437,151,496]" pageId="10" pageNumber="197">
|
|
<keyLead id="2613110D3562427AF2F47CB0FA300DCB" pageId="10" pageNumber="197">
|
|
- Ventral processes of first gonapophysis twisted into near-helix, apex of dorsal process not bifid, hooked anteriorly; pregenital plate spade-shaped (
|
|
<bibRefCitation id="F973CFC93562427AF3C77CD7FDF50DCB" author="Bahder, B. W. & Echavarria, M. A. Z. & Barrantes, E. A. B. & Helmick, E. E. & Bartlett, C. R." box="[420,601,474,496]" pageId="10" pageNumber="197" pagination="369 - 392" refId="ref5842" refString="Bahder, B. W., Echavarria, M. A. Z., Barrantes, E. A. B., Helmick, E. E. & Bartlett, C. R. (2021) A new genus and two new species of planthopper in the tribe Cenchreini (Hemiptera: Auchenorrhyncha: Derbidae) from lowland tropical rainforest in Costa Rica. Zootaxa, 4908 (3), 369 - 392. https: // doi. org / 10.11646 / zootaxa. 4908.3.3" type="journal article" year="2021">
|
|
Bahder
|
|
<emphasis id="AF966E2A3562427AF3927CD6FDB70DCB" box="[497,539,474,496]" italics="true" pageId="10" pageNumber="197">et al</emphasis>
|
|
. 2021
|
|
</bibRefCitation>
|
|
, fig. 11)................................................
|
|
<taxonomicName id="5AE2C9BB3562427AF6917CD6FA300DCB" ID-CoL="9FNPC" box="[1266,1436,474,496]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="197" phylum="Arthropoda" rank="species" species="pseudosororius">
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|
<emphasis id="AF966E2A3562427AF6917CD6FA300DCB" box="[1266,1436,474,496]" italics="true" pageId="10" pageNumber="197">T. pseudosororius</emphasis>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
</subSubSection>
|
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</treatment>
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</document> |