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<document id="ED823AFAF87796E4C5B2CBF8F5BD0659" ID-DOI="10.11646/zootaxa.3964.1.9" ID-GBIF-Dataset="c7142014-7e5b-4908-b55d-c6652d8df338" ID-ISSN="1175-5326" ID-Zenodo-Dep="238349" ID-ZooBank="BF11430F-83C3-499A-9526-C04B83912557" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461073651569" checkinUser="plazi" docAuthor="Naglis, Stefan &amp; Barták, Miroslav" docDate="2015" docId="0B4F5B5D3E33AD0AFF5E957D55C95A98" docLanguage="en" docName="zt03964p137.pdf" docOrigin="Zootaxa 3964 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sciapus negrobovi Naglis &amp; Barták, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="130" masterDocId="F77623253E37AD0FFFC996225052581B" masterDocTitle="Dolichopodidae (Diptera) from the Iberian Peninsula, with description of three new species" masterLastPageNumber="137" masterPageNumber="125" pageNumber="129" updateTime="1698585576818" updateUser="plazi">
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<mods:title id="D2C6DB335617B1444D5BDBEDF465AE5C">Dolichopodidae (Diptera) from the Iberian Peninsula, with description of three new species</mods:title>
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<mods:namePart id="908F0CF6FB9AD4FB5881F3C1E4053524">Naglis, Stefan</mods:namePart>
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<mods:namePart id="94F4E39A0DC49D81E6B914E878FE3841">Barták, Miroslav</mods:namePart>
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<mods:date id="238A36236C8CD4D8B62BD26B81C84FBD">2015</mods:date>
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<treatment id="0B4F5B5D3E33AD0AFF5E957D55C95A98" ID-DOI="http://doi.org/10.5281/zenodo.6120085" ID-GBIF-Taxon="119617052" ID-Zenodo-Dep="6120085" LSID="urn:lsid:plazi:treatment:0B4F5B5D3E33AD0AFF5E957D55C95A98" httpUri="http://treatment.plazi.org/id/0B4F5B5D3E33AD0AFF5E957D55C95A98" lastPageId="5" lastPageNumber="130" pageId="4" pageNumber="129">
<subSubSection id="CBFCB9C03E33AD0BFF5E957D50B95B81" pageId="4" pageNumber="129" type="nomenclature">
<paragraph id="8359EA4B3E33AD0BFF5E957D51885B62" blockId="4.[151,474,863,922]" box="[151,474,863,889]" pageId="4" pageNumber="129">
<heading id="D8115D273E33AD0BFF5E957D51885B62" bold="true" box="[151,474,863,889]" fontSize="11" level="1" pageId="4" pageNumber="129" reason="1">
<emphasis id="B19236593E33AD0BFF5E957D51885B62" bold="true" box="[151,474,863,889]" pageId="4" pageNumber="129">
<taxonomicName id="44E691C83E33AD0BFF5E957D51245B62" ID-CoL="9HVP3" box="[151,374,863,889]" class="Insecta" family="Dolichopodidae" genus="Sciapus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="negrobovi" status="sp. nov.">
<emphasis id="B19236593E33AD0BFF5E957D51245B62" bold="true" box="[151,374,863,889]" italics="true" pageId="4" pageNumber="129">Sciapus negrobovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AAA18B223E33AD0BFEB5957D51885B62" box="[380,474,863,889]" pageId="4" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8359EA4B3E33AD0BFF5E95A350B95B81" blockId="4.[151,474,863,922]" box="[151,235,897,922]" pageId="4" pageNumber="129">
(
<figureCitation id="1BDDF6CE3E33AD0BFF5695A350B05B81" box="[159,226,897,922]" captionStart="FIGURE 3" captionStartId="5.[151,249,1538,1560]" captionTargetBox="[164,1420,710,1498]" captionTargetId="figure@5.[151,1436,695,1516]" captionTargetPageId="5" captionText="FIGURE 3. Sciapus negrobovi sp. nov.: hypopygium lateral. Aed = aedeagus; Cer = cercus; Epl = epandrial lobe; Hyp = hypandrium; OrgX = Organ X; Sur = surstylus (scale bar = 0.1 mm)." httpUri="https://zenodo.org/record/238352/files/figure.png" pageId="4" pageNumber="129">Fig. 3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CBFCB9C03E33AD0BFF5E95EB542F5C1D" pageId="4" pageNumber="129" type="materials_examined">
<paragraph id="8359EA4B3E33AD0BFF5E95EB542F5C1D" blockId="4.[151,1437,969,2014]" pageId="4" pageNumber="129">
<emphasis id="B19236593E33AD0BFF5E95EB51BB5BF9" bold="true" box="[151,489,969,994]" pageId="4" pageNumber="129">
<typeStatus id="5C5D54E93E33AD0BFF5E95EB50805BF9" box="[151,210,969,994]" pageId="4" pageNumber="129">Type</typeStatus>
material.
<typeStatus id="5C5D54E93E33AD0BFE8295EB51BB5BF9" box="[331,489,969,994]" pageId="4" pageNumber="129" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
♂:
<collectingCountry id="FBF1AADB3E33AD0BFDDA95EB52E25BF9" box="[531,688,969,994]" name="Portugal" pageId="4" pageNumber="129">
<emphasis id="B19236593E33AD0BFDDA95EB52E25BF9" bold="true" box="[531,688,969,994]" pageId="4" pageNumber="129">PORTUGAL</emphasis>
</collectingCountry>
, Valhelhas,
<quantity id="441E47AE3E33AD0BFCF095EB53D35BFA" box="[825,897,969,994]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="129" unit="m" value="500.0">500 m</quantity>
, near river, SW+PT,
<date id="F758CC8B3E33AD0BFBA295EB55725BF9" box="[1131,1312,969,994]" pageId="4" pageNumber="129" value="2009-07-16" valueMax="2009-07-17" valueMin="2009-07-16">1617.VII.2009</date>
, M. Barták (
<collectionCode id="E5F7728E3E33AD0BFF5695CC50AE5C1D" box="[159,252,1006,1030]" pageId="4" pageNumber="129">CULSP</collectionCode>
).
<typeStatus id="5C5D54E93E33AD0BFEC795CF51F25C1D" box="[270,416,1005,1030]" pageId="4" pageNumber="129" type="paratype">
<emphasis id="B19236593E33AD0BFEC795CF51F25C1D" bold="true" box="[270,416,1005,1030]" pageId="4" pageNumber="129">PARATYPE</emphasis>
</typeStatus>
:
<collectingCountry id="FBF1AADB3E33AD0BFE7B95CF521D5C1D" box="[434,591,1005,1030]" name="Portugal" pageId="4" pageNumber="129">
<emphasis id="B19236593E33AD0BFE7B95CF521D5C1D" bold="true" box="[434,591,1005,1030]" pageId="4" pageNumber="129">PORTUGAL</emphasis>
</collectingCountry>
, 1 ♀ same data as
<typeStatus id="5C5D54E93E33AD0BFCEA95CF53DA5C1D" box="[803,904,1005,1030]" pageId="4" pageNumber="129" type="holotype">holotype</typeStatus>
, M. Barták (
<collectionCode id="E5F7728E3E33AD0BFBDA95CC54225C1D" box="[1043,1136,1006,1030]" pageId="4" pageNumber="129">CULSP</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="CBFCB9C03E33AD0BFF0E923355215C8D" pageId="4" pageNumber="129" type="diagnosis">
<paragraph id="8359EA4B3E33AD0BFF0E923355215C8D" blockId="4.[151,1437,969,2014]" pageId="4" pageNumber="129">
<emphasis id="B19236593E33AD0BFF0E923351685C31" bold="true" box="[199,314,1041,1066]" pageId="4" pageNumber="129">Diagnosis</emphasis>
. Scape and pedicel yellow, first flagellomere dark brown; frons with dense yellowish-grey pruinosity, face with dense silvery-white pruinosity; mesonotum with dense yellowish-grey pruinosity; abdomen metallic green, tergites 25 with large yellow dorsolateral patch; legs yellow, tarsomeres simple; lower calypter with white setae; halter pale yellow; hypopygium mainly yellow; “Organ X” with long, S-shaped ventral setae.
</paragraph>
</subSubSection>
<subSubSection id="CBFCB9C03E33AD0AFF0E928351AB58E3" lastPageId="5" lastPageNumber="130" pageId="4" pageNumber="129" type="description">
<paragraph id="8359EA4B3E33AD0AFF0E928351AB58E3" blockId="4.[151,1437,969,2014]" lastBlockId="5.[151,1437,151,429]" lastPageId="5" lastPageNumber="130" pageId="4" pageNumber="129">
<emphasis id="B19236593E33AD0BFF0E928351035CA1" bold="true" box="[199,337,1185,1210]" pageId="4" pageNumber="129">Description</emphasis>
.
<emphasis id="B19236593E33AD0BFEA9928351CE5CA1" bold="true" box="[352,412,1185,1210]" pageId="4" pageNumber="129">Male</emphasis>
. Body length (
<typeStatus id="5C5D54E93E33AD0BFD8D928352FF5CA1" box="[580,685,1185,1210]" pageId="4" pageNumber="129" type="holotype">holotype</typeStatus>
):
<quantity id="441E47AE3E33AD0BFD099283534A5CA2" box="[704,792,1185,1210]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="4" pageNumber="129" unit="mm" value="3.5">3.5 mm</quantity>
, wing length
<quantity id="441E47AE3E33AD0BFC78928054595CA2" box="[945,1035,1186,1210]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="4" pageNumber="129" unit="mm" value="3.4">3.4 mm</quantity>
.
<emphasis id="B19236593E33AD0BFBDC928354075CA1" bold="true" box="[1045,1109,1185,1210]" pageId="4" pageNumber="129">Head</emphasis>
: frons with dense yellowishgrey pruinosity, face with dense silvery-white pruinosity, below antennae about as wide as diameter of eyes; clypeus bulging; pair of strong vertical and ocellar setae; lower postoculars white; palpus pale yellow, with white apical setae; proboscis yellow. Antenna: scape and pedicel yellow, first flagellomere dark brown, as long as high; arista apicodorsal, bare.
<emphasis id="B19236593E33AD0BFE7F9313525D5D51" bold="true" box="[438,527,1329,1354]" pageId="4" pageNumber="129">Thorax</emphasis>
: mesonotum metallic green, with dense yellowish-grey pruinosity; ac strong, comprising 78 pairs; 5 pairs of strong dc, with additional small pair in front; 2 strong scutellar setae and small, hair-like lateral seta; pleura with dense greyish-white pruinosity; 2 small pale ppls.
<emphasis id="B19236593E33AD0BFB8F935B54DA5D89" bold="true" box="[1094,1160,1401,1426]" pageId="4" pageNumber="129">Legs:</emphasis>
including coxae yellow, except hind tarsomeres slightly infuscated. Fore leg: coxa with white hairs and row of strong yellow lateral and some apical setae; femur with 45 white ventral setae, half as long as diameter of femur; tibia and tarsomeres bare; relative podomere ratios: 42:54:37:16:12:8:5. Mid leg: coxa with several strong yellow apical setae; femur bare; tibia with 1 strong black ad seta at 1/5; tarsomeres bare; relative podomere ratios: 48:75:50:21:16:9:6.
<collectingCountry id="FBF1AADB3E33AD0BFAF8902B553B5E39" box="[1329,1385,1545,1570]" name="India" pageId="4" pageNumber="129">Hind</collectingCountry>
leg: coxa with 2 strong white lateral setae; femur with 1 small black anterior preapical seta; tibia with row of small black pd setae, and some small black apical setae; basitarsus with small black pv seta basally; relative podomere ratios: 65:92:33:31:17:10:5. All claws and pulvilli present but small.
<emphasis id="B19236593E33AD0BFC75905754545E95" bold="true" box="[956,1030,1653,1678]" pageId="4" pageNumber="129">Wing:</emphasis>
hyaline, veins yellow; R1 and R2+3 straight; R4+5 strongly curved posteriad at distal 4/5; M1 with almost right-angle bend at 1/3, then running straight and joining wing margin anteriad of apex close to R4+5; M2 straight, not reaching wing margin, as long as dm-cu; section of M between dm-cu and M2 as long as dm-cu; CuAx ratio: 1.1; RMx ratio: 8.0; lower calypter white, with white setae; halter pale yellow.
<emphasis id="B19236593E33AD0BFE329133522B5F31" bold="true" box="[507,633,1809,1834]" pageId="4" pageNumber="129">Abdomen:</emphasis>
dark metallic green, tergites 25 with brown basal band and with large yellow dorsolateral patch, tergites 7 and 8 entirely yellow; all hairs and setae black. Hypopygium (
<figureCitation id="1BDDF6CE3E33AD0BFA8A911655DF5F56" box="[1347,1421,1844,1869]" captionStart="FIGURE 3" captionStartId="5.[151,249,1538,1560]" captionTargetBox="[164,1420,710,1498]" captionTargetId="figure@5.[151,1436,695,1516]" captionTargetPageId="5" captionText="FIGURE 3. Sciapus negrobovi sp. nov.: hypopygium lateral. Aed = aedeagus; Cer = cercus; Epl = epandrial lobe; Hyp = hypandrium; OrgX = Organ X; Sur = surstylus (scale bar = 0.1 mm)." httpUri="https://zenodo.org/record/238352/files/figure.png" pageId="4" pageNumber="129">Fig. 3</figureCitation>
): epandrium, cercus, and “Organ X” pale yellow, base of hypandrium and apical part of surstylus dark brown. Epandrial lobe with 1 long apical and 1 small basoventral setae; surstylus with 2 strong apical and 1 curved subapical setae; “Organ X” with midventral lobe bearing long, S-shaped setae.
<emphasis id="B19236593E33AD0BFBD79183542D5FA1" bold="true" box="[1054,1151,1953,1978]" pageId="4" pageNumber="129">Female:</emphasis>
similar to male but: fore coxa with yellow spine-like apical setae; fore femur with ventral row of 45 strong, yellow spine-like setae, longer than diameter of femur; fore tibia with some small ad and pd setae; mid femur with 1 anterior preapical seta; mid tibia with 1 strong ad, 3 pd, and 1 av setae; hind tibia with 1 strong ad seta; tergite 1 yellow, tergites 24 metallic green with yellow lateral patch.
</paragraph>
</subSubSection>
<subSubSection id="CBFCB9C03E32AD0AFF0E972654885906" box="[199,1242,260,285]" pageId="5" pageNumber="130" type="etymology">
<paragraph id="8359EA4B3E32AD0AFF0E972654885906" blockId="5.[151,1437,151,429]" box="[199,1242,260,285]" pageId="5" pageNumber="130">
<emphasis id="B19236593E32AD0AFF0E972651155906" bold="true" box="[199,327,260,285]" pageId="5" pageNumber="130">Etymology</emphasis>
. The new species is dedicated to the Russian dipterist, Oleg Pavlovich Negrobov.
</paragraph>
</subSubSection>
<subSubSection id="CBFCB9C03E32AD0AFF0E970555C95A98" pageId="5" pageNumber="130" type="discussion">
<paragraph id="8359EA4B3E32AD0AFF0E970554E059B6" blockId="5.[151,1437,151,429]" pageId="5" pageNumber="130">
<emphasis id="B19236593E32AD0AFF0E97055166595B" bold="true" box="[199,308,295,320]" pageId="5" pageNumber="130">Remarks</emphasis>
. The new species belongs to the
<taxonomicName id="44E691C83E32AD0AFD67970A5341595B" box="[686,787,296,320]" class="Insecta" family="Dolichopodidae" genus="Sciapus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="aberrans">
<emphasis id="B19236593E32AD0AFD67970A5341595B" box="[686,787,296,320]" italics="true" pageId="5" pageNumber="130">aberrans</emphasis>
</taxonomicName>
speciesgroup: tarsomeres simple; frons and thorax with dense pruinosity; cerci fused; “Organ X” fused, with long ventral setae; abdominal segments partly yellow. In the Palaearctic key (
<bibRefCitation id="E77797BA3E32AD0AFE9D974D52FA5993" author="Grichanov" box="[340,680,367,392]" pageId="5" pageNumber="130" refString="Grichanov, I. Ya. &amp; Negrobov, O. P. (2014) Palaearctic species of the genus Sciapus Zeller (Diptera: Dolichopodidae). Plant Protection News, 13 (Supplement), 4 - 85." type="journal article" year="2014">Grichanov &amp; Negrobov 2014</bibRefCitation>
)
<taxonomicName id="44E691C83E32AD0AFD71975253155993" box="[696,839,368,392]" class="Insecta" family="Dolichopodidae" genus="Sciapus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="negrobovi">
<emphasis id="B19236593E32AD0AFD71975253155993" box="[696,839,368,392]" italics="true" pageId="5" pageNumber="130">S. negrobovi</emphasis>
</taxonomicName>
runs to
<taxonomicName id="44E691C83E32AD0AFC609752549F5993" authority="Becker, 1918" authorityName="Becker" authorityYear="1918" box="[937,1229,367,392]" class="Insecta" family="Dolichopodidae" genus="Sciapus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="aberrans">
<emphasis id="B19236593E32AD0AFC609752547E5993" box="[937,1068,368,392]" italics="true" pageId="5" pageNumber="130">S. aberrans</emphasis>
Becker, 1918
</taxonomicName>
and
<taxonomicName id="44E691C83E32AD0AFACE9752510659B7" authority="Grichanov, 2007" authorityName="Grichanov" authorityYear="2007" class="Insecta" family="Dolichopodidae" genus="Sciapus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="subvicinus">
<emphasis id="B19236593E32AD0AFACE975255CE5993" box="[1287,1436,368,392]" italics="true" pageId="5" pageNumber="130">S. subvicinus</emphasis>
Grichanov, 2007
</taxonomicName>
. The three species can be separated by the characters given in the key below.
</paragraph>
<paragraph id="8359EA4B3E32AD0AFF5E97F955C959EA" blockId="5.[151,1436,475,644]" box="[151,1435,475,497]" pageId="5" pageNumber="130">
1 “Organ X” with midventral lobe bearing long, S-shaped setae (
<figureCitation id="1BDDF6CE3E32AD0AFC8D97F9532C59EA" box="[836,894,475,497]" captionStart="FIGURE 3" captionStartId="5.[151,249,1538,1560]" captionTargetBox="[164,1420,710,1498]" captionTargetId="figure@5.[151,1436,695,1516]" captionTargetPageId="5" captionText="FIGURE 3. Sciapus negrobovi sp. nov.: hypopygium lateral. Aed = aedeagus; Cer = cercus; Epl = epandrial lobe; Hyp = hypandrium; OrgX = Organ X; Sur = surstylus (scale bar = 0.1 mm)." httpUri="https://zenodo.org/record/238352/files/figure.png" pageId="5" pageNumber="130">Fig. 3</figureCitation>
A); epandrial lobe with 1 strong apical and 1 small bas-
</paragraph>
<paragraph id="8359EA4B3E32AD0AFF2897DA55C95A7D" blockId="5.[151,1436,475,644]" pageId="5" pageNumber="130">
oventral seta.........................................................................
<emphasis id="B19236593E32AD0AFB0497DA55CE5A15" bold="true" box="[1229,1436,504,526]" pageId="5" pageNumber="130">
<taxonomicName id="44E691C83E32AD0AFB0497DA551B5A15" box="[1229,1353,504,526]" class="Insecta" family="Dolichopodidae" genus="Sciapus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="negrobovi" status="sp. nov.">
<emphasis id="B19236593E32AD0AFB0497DA551B5A15" bold="true" box="[1229,1353,504,526]" italics="true" pageId="5" pageNumber="130">S. negrobovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AAA18B223E32AD0AFA8697DB55CE5A15" box="[1359,1436,505,526]" pageId="5" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
- “Organ X” without midventral lobe, setae not S-shaped....................................................... 2 2 Cercus small and rounded; epandrial lobe with 2 apical setae (
<bibRefCitation id="E77797BA3E32AD0AFCFA941154DF5A52" author="Grichanov" box="[819,1165,563,585]" pageId="5" pageNumber="130" refString="Grichanov, I. Ya. &amp; Negrobov, O. P. (2014) Palaearctic species of the genus Sciapus Zeller (Diptera: Dolichopodidae). Plant Protection News, 13 (Supplement), 4 - 85." type="journal article" year="2014">Grichanov &amp; Negrobov 2014, fig. 1</bibRefCitation>
).....
<taxonomicName id="44E691C83E32AD0AFB15941155CE5A53" authority="Becker" authorityName="Becker" box="[1244,1436,563,585]" class="Insecta" family="Dolichopodidae" genus="Sciapus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="aberrans">
<emphasis id="B19236593E32AD0AFB15941155CE5A53" bold="true" box="[1244,1436,563,585]" pageId="5" pageNumber="130">
<emphasis id="B19236593E32AD0AFB159411551C5A52" bold="true" box="[1244,1358,563,585]" italics="true" pageId="5" pageNumber="130">S. aberrans</emphasis>
Becker
</emphasis>
</taxonomicName>
- Cercus twice as long as wide; epandrial lobe with 1 apical and 1 midventral seta (
<bibRefCitation id="E77797BA3E32AD0AFC23947255C95A7D" author="Bulli" box="[1002,1435,592,614]" pageId="5" pageNumber="130" refString="Bulli, A. F. &amp; Negrobov, O. P. (1987) A new species of the genus Sciapus (Diptera, Dolichopodidae). Vestnik Zoologii, 3, 81 - 82. [in Russian]" type="journal article" year="1987">
Bulli &amp; Negrobov 1987, figs 13, as
<emphasis id="B19236593E32AD0AFA84947255C95A7D" box="[1357,1435,592,614]" italics="true" pageId="5" pageNumber="130">S. medi-</emphasis>
</bibRefCitation>
</paragraph>
<paragraph id="8359EA4B3E32AD0AFF28944C55C95A98" blockId="5.[151,1436,475,644]" box="[225,1435,621,644]" pageId="5" pageNumber="130">
<emphasis id="B19236593E32AD0AFF28944C516F5A98" box="[225,317,622,643]" italics="true" pageId="5" pageNumber="130">terraneus</emphasis>
Bulli &amp; Negrobov)........................................................
<taxonomicName id="44E691C83E32AD0AFB6C944C55C95A98" authority="Grichanov" authorityName="Grichanov" box="[1189,1435,622,644]" class="Insecta" family="Dolichopodidae" genus="Sciapus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="subvicinus">
<emphasis id="B19236593E32AD0AFB6C944C55C95A98" bold="true" box="[1189,1435,622,644]" pageId="5" pageNumber="130">
<emphasis id="B19236593E32AD0AFB6C944C55755A9F" bold="true" box="[1189,1319,622,644]" italics="true" pageId="5" pageNumber="130">S. subvicinus</emphasis>
Grichanov
</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>