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<document id="A185EDF86D1B4E917A93E1C58C08C0CF" ID-DOI="10.11646/zootaxa.3893.4.7" ID-GBIF-Dataset="f1bee12c-0039-4daf-9a8c-13f72eca6253" ID-ISSN="1175-5326" ID-Zenodo-Dep="230192" ID-ZooBank="1BACFDA5-7457-496F-BDE3-0F509F29879A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460996611568" checkinUser="plazi" docAuthor="Sabella, Giorgio, Viglianisi, Fabio Massimo, D’Urso, Vera &amp; Nouira, Said" docDate="2014" docId="03F63E5293439A4EFF1CFE4FFEDEFC98" docLanguage="en" docName="zt03893p586.pdf" docOrigin="Zootaxa 3893 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tychobythinus atlanticus Besuchet 1963" docType="treatment" docVersion="10" lastPageNumber="585" masterDocId="FFCF462A93459A48FF8BFFDBFFAAFF96" masterDocTitle="A review of the Tychobythinus algiricus species group from North Africa (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="586" masterPageNumber="579" pageNumber="585" updateTime="1698562906218" updateUser="plazi">
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<mods:title id="4DC1C413C9EE5330144427BF6EDAD025">A review of the Tychobythinus algiricus species group from North Africa (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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<mods:namePart id="923B59DCF882B65E5D1748F49074D499">Sabella, Giorgio</mods:namePart>
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<mods:namePart id="182EFADFB0525C996ACA095D793C0E01">Viglianisi, Fabio Massimo</mods:namePart>
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<mods:namePart id="80D73BF6769C4137727461F7E29A1E3F">D’Urso, Vera</mods:namePart>
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<mods:namePart id="92D6EE01A05DB643D2BA8FFC924E7AAE">Nouira, Said</mods:namePart>
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<mods:title id="831C65895E4A19F4C15AF4A6EC0CC6B4">Zootaxa</mods:title>
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<mods:date id="E31483A213014FB87D9931F993186EDE">2014</mods:date>
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<mods:number id="FAD53B5E4DCB2D1E85AFFDEB5812DF54">2014-12-10</mods:number>
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<mods:number id="94C8DA98290A7FE075240C81609FB145">3893</mods:number>
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<mods:identifier id="572210F75AD005C67E031657AF8A3FBB" type="DOI">10.11646/zootaxa.3893.4.7</mods:identifier>
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<treatment id="03F63E5293439A4EFF1CFE4FFEDEFC98" ID-DOI="http://doi.org/10.5281/zenodo.6130137" ID-GBIF-Taxon="119597428" ID-Zenodo-Dep="6130137" LSID="urn:lsid:plazi:treatment:03F63E5293439A4EFF1CFE4FFEDEFC98" httpUri="http://treatment.plazi.org/id/03F63E5293439A4EFF1CFE4FFEDEFC98" lastPageNumber="585" pageId="6" pageNumber="585">
<subSubSection id="C345DCCF93439A4EFF1CFE4FFD22FE38" box="[151,648,404,430]" pageId="6" pageNumber="585" type="nomenclature">
<paragraph id="8BE08F4493439A4EFF1CFE4FFD22FE38" blockId="6.[151,648,404,463]" box="[151,648,404,430]" pageId="6" pageNumber="585">
<heading id="D0A8382893439A4EFF1CFE4FFD22FE38" bold="true" box="[151,648,404,430]" fontSize="11" level="1" pageId="6" pageNumber="585" reason="1">
<taxonomicName id="4C5FF4C793439A4EFF1CFE4FFD22FE38" ID-CoL="59SD5" authority="Besuchet, 1963" authorityName="Besuchet" authorityYear="1963" box="[151,648,404,430]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="atlanticus">
<emphasis id="B92B535693439A4EFF1CFE4FFD22FE38" bold="true" box="[151,648,404,430]" pageId="6" pageNumber="585">
<emphasis id="B92B535693439A4EFF1CFE4FFE6DFE38" bold="true" box="[151,455,404,430]" italics="true" pageId="6" pageNumber="585">Tychobythinus atlanticus</emphasis>
<bibRefCitation id="EFCEF2B593439A4EFE45FE4FFD22FE38" author="Besuchet, C." box="[462,648,404,430]" firstAuthor="Besuchet" journalOrPublisher="Mitteilungen der Schweizerischen Entomologischen Gesellschaft" pageId="6" pageNumber="585" pagination="227 - 232" part="35" refId="ref4601" refString="Besuchet, C. (1963 a) Pselaphides recoltes au Maroc par M. R. Mussard (Coleoptera). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 35, 227 - 232." title="Pselaphides recoltes au Maroc par M. R. Mussard (Coleoptera)" type="journal article" year="1963">Besuchet, 1963</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C345DCCF93439A4EFF1CFE6DFEE7FE59" box="[151,333,438,463]" pageId="6" pageNumber="585" type="description">
<paragraph id="8BE08F4493439A4EFF1CFE6DFEE7FE59" blockId="6.[151,648,404,463]" box="[151,333,438,463]" pageId="6" pageNumber="585">
(
<figureCitation id="136493C193439A4EFF14FE6DFF56FE59" box="[159,252,438,463]" captionStart="FIGURES 1 A – F" captionStartId="3.[151,264,1718,1741]" captionTargetBox="[158,1426,197,1688]" captionTargetId="figure@3.[151,1436,193,1698]" captionTargetPageId="3" captionText="FIGURES 1 A – F. Head of Tychobythinus. Scale: 0, 1 mm. A— Tychobythinus algiricus male, head dorsal view, schematic drawing. B— Tychobythinus algiricus male, head lateral view. C— Tychobythinus algiricus male, head ventral view, schematic drawing. D— Tychobythinus algiricus female, head dorsal view, schematic drawing. E— Tychobythinus escalerai male, head dorsal view, schematic drawing. F— Tychobythinus atlanticus male, head dorsal view, schematic drawing." httpUri="https://zenodo.org/record/230193/files/figure.png" pageId="6" pageNumber="585">Figs. 1F</figureCitation>
,
<figureCitation id="136493C193439A4EFE83FE6CFE81FE59" box="[264,299,439,463]" captionStart="FIGURES 2 A – E" captionStartId="4.[151,264,1636,1659]" captionTargetBox="[251,1320,413,1602]" captionTargetId="figure@4.[245,1341,409,1615]" captionTargetPageId="4" captionText="FIGURES 2 A – E. Scale: 0, 1 mm. A— Tychobythinus escalerai, paratype male, right antenna. B— Tychobythinus atlanticus, holotype male, right antenna. C— Tychobythinus algiricus male, right antenna. D— Tychobythinus algiricus male, right maxillary palpus. E— Tychobythinus algiricus male, aedeagus, dorsal view." httpUri="https://zenodo.org/record/230194/files/figure.png" pageId="6" pageNumber="585">2B</figureCitation>
,
<figureCitation id="136493C193439A4EFEBDFE6CFEEEFE59" box="[310,324,439,463]" captionStart="FIGURE 3" captionStartId="5.[151,250,1455,1477]" captionTargetBox="[151,1435,951,1433]" captionTargetId="figure@5.[151,1435,951,1433]" captionTargetPageId="5" captionText="FIGURE 3. Distribution map of Tychobythinus algiricus species group. • = Tychobythinus algiricus. ★ = Tychobythinus escalerai. ˔ = Tychobythinus atlanticus." httpUri="https://zenodo.org/record/230195/files/figure.png" pageId="6" pageNumber="585">3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C345DCCF93439A4EFF1CFE25FB57FD82" box="[151,1277,510,532]" pageId="6" pageNumber="585" type="reference_group">
<paragraph id="8BE08F4493439A4EFF1CFE25FB57FD82" blockId="6.[151,1277,510,532]" box="[151,1277,510,532]" pageId="6" pageNumber="585">
<treatmentCitationGroup id="AB4FA86A93439A4EFF1CFE25FB57FD82" box="[151,1277,510,532]" pageId="6" pageNumber="585">
<taxonomicName id="4C5FF4C793439A4EFF1CFE25FDDBFD82" ID-CoL="59SD5" authority="Besuchet 1963b: 218" authorityName="Besuchet" authorityPageNumber="218" authorityYear="1963" box="[151,625,510,532]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="atlanticus">
<emphasis id="B92B535693439A4EFF1CFE25FE39FD82" box="[151,403,510,532]" italics="true" pageId="6" pageNumber="585">Tychobythinus atlanticus</emphasis>
<treatmentCitation id="0AFEA95593439A4EFE12FE25FDDBFD82" author="Besuchet" box="[409,625,510,532]" page="218" pageId="6" pageNumber="585" year="1963">
<bibRefCitation id="EFCEF2B593439A4EFE12FE25FDDBFD82" author="Besuchet, C." box="[409,625,510,532]" firstAuthor="Besuchet" journalOrPublisher="Archives des Sciences" page="218" pageId="6" pageNumber="585" pagination="217 - 223" part="16" refId="ref4635" refString="Besuchet, C. (1963 b) Troisieme contribution a l'etude des pselaphides du Maroc (Coleoptera). Archives des Sciences, 16, 217 - 223." title="Troisieme contribution a l'etude des pselaphides du Maroc (Coleoptera)" type="journal article" year="1963" yearSuffix="b">Besuchet 1963b: 218</bibRefCitation>
</treatmentCitation>
</taxonomicName>
, fig. 1 (aedeagus);
<treatmentCitation id="0AFEA95593439A4EFCB9FE25FC78FD82" author="Kocher" box="[818,978,510,532]" page="5" pageId="6" pageNumber="585" year="1969">
<bibRefCitation id="EFCEF2B593439A4EFCB9FE25FC78FD82" author="Kocher, L." box="[818,978,510,532]" firstAuthor="Kocher" journalOrPublisher="Travaux de l'Institut scientifique cherifien, Serie zoologique" page="5" pageId="6" pageNumber="585" pagination="1 - 15" part="34" refId="ref4851" refString="Kocher, L. (1969) Catalogue commente des coleopteres du Maroc. Feuillet rectificatif n ° 7 (fascicule II, p. 206 a 214) (Fam. Pselaphidae). Travaux de l'Institut scientifique cherifien, Serie zoologique, 34 (1968), 1 - 15." title="Catalogue commente des coleopteres du Maroc. Feuillet rectificatif n ° 7 (fascicule II, p. 206 a 214) (Fam. Pselaphidae)" type="journal article" year="1969">Kocher 1969: 5</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AFEA95593439A4EFC56FE25FB53FD82" author="Lobl" box="[989,1273,510,532]" page="314" pageId="6" pageNumber="585" year="2004">
<bibRefCitation id="EFCEF2B593439A4EFC56FE25FB53FD82" author="Lobl, I. &amp; Besuchet, C." bookContentInfo="942 pp." box="[989,1273,510,532]" editor="Lobl, I. &amp; Smetana, A." firstAuthor="Lobl" journalOrPublisher="Apollo Books, Stenstrup" page="314" pageId="6" pageNumber="585" pagination="272 - 329" refId="ref4904" refString="Lobl, I. &amp; Besuchet, C. (2004) Subfamily Pselaphinae Latreille, 1802, In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea- Histeroidea - Staphylinoidea. Apollo Books, Stenstrup, pp. 272 - 329. [942 pp.]" title="Subfamily Pselaphinae Latreille, 1802" type="book chapter" volumeTitle="Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea- Histeroidea - Staphylinoidea" year="2004">Löbl &amp; Besuchet 2004: 314</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C345DCCF93439A4EFF1CFD9AFDC5FDE8" pageId="6" pageNumber="585" type="materials_examined">
<paragraph id="8BE08F4493439A4EFF1CFD9AFDC5FDE8" blockId="6.[151,1436,577,782]" pageId="6" pageNumber="585">
<emphasis id="B92B535693439A4EFF1CFD9AFED3FDCC" bold="true" box="[151,377,577,602]" pageId="6" pageNumber="585">Material examined</emphasis>
.
<materialsCitation id="3B37851993439A4EFE0CFD9AFDC5FDE8" ID-GBIF-Occurrence="3923149321" collectionCode="MHNG" collectorName="R. Mussard" country="Morocco" elevation="2" location="Djebel Tazzeka" pageId="6" pageNumber="585" specimenCount="1" specimenCount-male="1" stateProvince="Taza-Al Hoceima-Taounate" typeStatus="holotype">
<emphasis id="B92B535693439A4EFE0CFD9AFD6DFDCC" bold="true" box="[391,711,577,602]" pageId="6" pageNumber="585">
NORTHERN
<collectingCountry id="F348CFD493439A4EFDA4FD9AFD6DFDCC" box="[559,711,577,602]" name="Morocco" pageId="6" pageNumber="585">MOROCCO</collectingCountry>
</emphasis>
:
<emphasis id="B92B535693439A4EFD5DFD9AFBDAFDCC" bold="true" box="[726,1136,577,602]" pageId="6" pageNumber="585">
<collectingRegion id="499B41A693439A4EFD5DFD9AFBB7FDCC" box="[726,1053,577,602]" pageId="6" pageNumber="585">Taza-Al Hoceïma-Taounate</collectingRegion>
region
</emphasis>
:
<location id="8E80D99F93439A4EFBF5FD9AFA84FDCC" LSID="urn:lsid:plazi:treatment:03F63E5293439A4EFF1CFE4FFEDEFC98:8E80D99F93439A4EFBF5FD9AFA84FDCC" box="[1150,1326,577,602]" country="Morocco" name="Djebel Tazzeka" pageId="6" pageNumber="585" stateProvince="Taza-Al Hoceima-Taounate">Djebel Tazzeka</location>
,
<quantity id="4CA722A193439A4EFAB2FD9AFA3DFDCF" box="[1337,1431,577,601]" metricMagnitude="0" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="585" unit="m" value="1.7">
<elevation id="0072687793439A4EFAB2FD9AFA3DFDCF" box="[1337,1431,577,601]" metricMagnitude="0" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="585" unit="m" value="1.7">1.700 m</elevation>
</quantity>
,
<specimenCount id="9D5944CD93439A4EFF1CFDBDFF48FDE8" box="[151,226,613,638]" pageId="6" pageNumber="585" type="male">1 male</specimenCount>
(
<typeStatus id="54E431E693439A4EFF7AFDBEFEFCFDE8" box="[241,342,613,638]" pageId="6" pageNumber="585" type="holotype">holotype</typeStatus>
) (
<collectorName id="26AAEA9293439A4EFEE5FDBDFE5BFDE8" box="[366,497,613,638]" pageId="6" pageNumber="585">R. Mussard</collectorName>
) (
<collectionCode id="ED4E178193439A4EFD83FDBDFDCEFDE8" box="[520,612,614,638]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="6" pageNumber="585">MHNG</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C345DCCF93439A4EFF4CFD52FE43FD50" pageId="6" pageNumber="585" type="discussion">
<paragraph id="8BE08F4493439A4EFF4CFD52FE43FD50" blockId="6.[151,1436,577,782]" pageId="6" pageNumber="585">
<emphasis id="B92B535693439A4EFF4CFD52FE04FD34" bold="true" box="[199,430,649,674]" pageId="6" pageNumber="585">Comparative notes</emphasis>
. The characters that distinguish this species from
<taxonomicName id="4C5FF4C793439A4EFB87FD51FB39FD34" box="[1036,1171,649,674]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="escalerai">
<emphasis id="B92B535693439A4EFB87FD51FB39FD34" box="[1036,1171,649,674]" italics="true" pageId="6" pageNumber="585">T. escalerai</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5FF4C793439A4EFB5DFD51FAF1FD34" box="[1238,1371,649,674]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="algiricus">
<emphasis id="B92B535693439A4EFB5DFD51FAF1FD34" box="[1238,1371,649,674]" italics="true" pageId="6" pageNumber="585">T. algiricus</emphasis>
</taxonomicName>
have already been discussed above.
</paragraph>
</subSubSection>
<subSubSection id="C345DCCF93439A4EFF4CFD0AFEDEFC98" pageId="6" pageNumber="585" type="distribution">
<paragraph id="8BE08F4493439A4EFF4CFD0AFEDEFC98" blockId="6.[151,1436,577,782]" pageId="6" pageNumber="585">
<emphasis id="B92B535693439A4EFF4CFD0AFEF3FD7C" bold="true" box="[199,345,721,746]" pageId="6" pageNumber="585">Distribution</emphasis>
(
<figureCitation id="136493C193439A4EFEE1FD0AFE1AFD7C" box="[362,432,721,746]" captionStart="FIGURE 3" captionStartId="5.[151,250,1455,1477]" captionTargetBox="[151,1435,951,1433]" captionTargetId="figure@5.[151,1435,951,1433]" captionTargetPageId="5" captionText="FIGURE 3. Distribution map of Tychobythinus algiricus species group. • = Tychobythinus algiricus. ★ = Tychobythinus escalerai. ˔ = Tychobythinus atlanticus." httpUri="https://zenodo.org/record/230195/files/figure.png" pageId="6" pageNumber="585">Fig. 3</figureCitation>
). The species is known only from the
<typeStatus id="54E431E693439A4EFCE7FD09FC36FD7C" box="[876,924,722,746]" pageId="6" pageNumber="585">type</typeStatus>
locality (
<collectingCountry id="F348CFD493439A4EFB82FD09FBD9FD7C" box="[1033,1139,722,746]" name="Morocco" pageId="6" pageNumber="585">Morocco</collectingCountry>
, Djebel Tazzeka) where
<specimenCount id="9D5944CD93439A4EFA05FD09FF67FC98" pageId="6" pageNumber="585" type="male">1 male</specimenCount>
was collected.
</paragraph>
</subSubSection>
</treatment>
</document>
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