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<document id="5E0E2674651EE484F1E3B15E3600028A" ID-CLB-Dataset="5896" ID-DOI="10.11646/zootaxa.4981.3.7" ID-GBIF-Dataset="ceca425f-e47a-4fe0-8c4c-f0abaae680e4" ID-ISSN="1175-5326" ID-Zenodo-Dep="5046838" ID-ZooBank="23C81666-C440-489C-AF53-6C37FB68AFEB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623283102752" checkinUser="plazi" docAuthor="Sáfián, Szabolcs, Collins, Steve, Warren-Gash, Haydon &amp; Belcastro, Claudio" docDate="2021" docId="03BD9F74FA412A1031DD37AF0BACF85F" docLanguage="en" docName="zootaxa.4981.3.7.pdf" docOrigin="Zootaxa 4981 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cephetola wologizi Safian 2021, sp. nov." docType="treatment" docVersion="8" lastPageNumber="573" masterDocId="FF84E70CFA502A03314A35080812FFB7" masterDocTitle="Description of five new species of Epitola sensu lato (Lepidoptera: Lycaenidae Poritiinae) from West and Central Africa" masterLastPageNumber="576" masterPageNumber="554" pageNumber="571" updateTime="1699009154658" updateUser="ExternalLinkService">
<mods:mods id="9DB55818872CE524B1A529ED53E3872F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="523170852D5E5A2C9C044FAE41D5E3BD">
<mods:title id="688C97200BB129557C6E6DA83D6B2012">Description of five new species of Epitola sensu lato (Lepidoptera: Lycaenidae Poritiinae) from West and Central Africa</mods:title>
</mods:titleInfo>
<mods:name id="3DAD99750CCB5482A15A0CC6B0BD11B9" type="personal">
<mods:role id="EF01D70A092890EC16501B072517866D">
<mods:roleTerm id="68548914EA94979F8D3892DB11EC8A76">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F7E6FB311519FB6306A409FD98B10616">Sáfián, Szabolcs</mods:namePart>
<mods:nameIdentifier id="88FEF0B537B20062D3F55989DBA96170" type="ORCID">0000-0002-0614-4203</mods:nameIdentifier>
<mods:affiliation id="A490A053D85C6A21418A6310A2DB9A9F">African Butterfly Research Institute, P. O. Box 14308, Nairobi, Kenya.</mods:affiliation>
<mods:nameIdentifier id="FEB36612086674941269D0E86FA13DF1" type="email">szsafian@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="A6FF89E4AF193569F9FC9202F495FC76" type="personal">
<mods:role id="E90D3F53C20572D86BE1BC0224D43155">
<mods:roleTerm id="89B6510893A595886CE0705DB2CD8294">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="54C8C13A1A1ABB5E8B885BB7E416EE55">Collins, Steve</mods:namePart>
<mods:affiliation id="DE470631B12C194BCA0968DD150C01A5">African Butterfly Research Institute, P. O. Box 14308, Nairobi, Kenya.</mods:affiliation>
</mods:name>
<mods:name id="F186707663DFCA5D0E424AF42423AA5B" type="personal">
<mods:role id="6792B6613B35E2B9A02A94C9423611BE">
<mods:roleTerm id="05CF971269071D4559F82D61524331D0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="94BC051166F3C30D70C014281855638F">Warren-Gash, Haydon</mods:namePart>
<mods:affiliation id="96394F144ADC7BC727609167F48BF877">98 Overstrand Mansions, Prince of Wales Drive, London SW 11 4 EU, UK.</mods:affiliation>
</mods:name>
<mods:name id="E1D34BBBA3718EF750F221DBCC860E1F" type="personal">
<mods:role id="F4D1A33228B4BD91432FA69A12A92EC6">
<mods:roleTerm id="F32FF0086D8C4421BC5FA49C8FEFFE0B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2DAA96B688B6C5D74BCCA990B24A254A">Belcastro, Claudio</mods:namePart>
<mods:nameIdentifier id="8081E19F5CC6D8FAFEE43DE2A6CDEF43" type="ORCID">0000-0003-0102-5106</mods:nameIdentifier>
<mods:affiliation id="05AD2580BFF4ADA07066261074118706">Museo della Biodiversita, Parco Nationale della Sila, Loc Cupone, I- 87058 Spezzano della Sila, Italy. belcastroclaudio @ yahoo. com; https: // orcid. org / 0000 - 0003 - 0102 - 5106</mods:affiliation>
<mods:nameIdentifier id="649570A045EACFBDDA5717A99E979A52" type="email">belcastroclaudio@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="80948D32D6DC55CF769070AF9E2133DC">text</mods:typeOfResource>
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<mods:titleInfo id="23207077A6F9D9FA8B42F8CC2142C3F9">
<mods:title id="934CC4F21C3578299F3732209B41EF6A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="667BA5762F0C9CFEA45653F1658FFD46">
<mods:date id="36D603F152C376811903F1D00777563B">2021</mods:date>
<mods:detail id="11D5C0655A74E85D777A2399EBC1C3D9" type="pubDate">
<mods:number id="D3667D214420FD150B9753CEB912C39C">2021-06-08</mods:number>
</mods:detail>
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<mods:number id="A275F84AE927EF370CD99A7EE561473F">4981</mods:number>
</mods:detail>
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<mods:number id="79F521E2EA402AE78DFFC4263459AEE6">3</mods:number>
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<mods:start id="29042A0386CD2EBC30FDACF2BC469846">554</mods:start>
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<mods:classification id="A15143FB7A6F4B1081D4F7DB848BFE6B">journal article</mods:classification>
<mods:identifier id="160E303DCAA408F51D21E3DE002DF77A" type="CLB-Dataset">5896</mods:identifier>
<mods:identifier id="822F778AFD1FB774F8EA3F707C7A2BA2" type="DOI">10.11646/zootaxa.4981.3.7</mods:identifier>
<mods:identifier id="F49180F89F08471BD7BEE9891C580C56" type="GBIF-Dataset">ceca425f-e47a-4fe0-8c4c-f0abaae680e4</mods:identifier>
<mods:identifier id="DA6674C33B1C255E6B404F88133B38E8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="193E0AEE87C4566CE4CB34FF73A1454B" type="Zenodo-Dep">5046838</mods:identifier>
<mods:identifier id="35B55B84E1ABA9294DB368D2DDCC83D8" type="ZooBank">23C81666-C440-489C-AF53-6C37FB68AFEB</mods:identifier>
</mods:mods>
<treatment id="03BD9F74FA412A1031DD37AF0BACF85F" ID-DOI="http://doi.org/10.5281/zenodo.4923640" ID-GBIF-Taxon="183228186" ID-Zenodo-Dep="4923640" LSID="urn:lsid:plazi:treatment:03BD9F74FA412A1031DD37AF0BACF85F" httpUri="http://treatment.plazi.org/id/03BD9F74FA412A1031DD37AF0BACF85F" lastPageId="19" lastPageNumber="573" pageId="17" pageNumber="571">
<subSubSection id="C30E7DE9FA412A1231DD37AF09A2FD51" pageId="17" pageNumber="571" type="nomenclature">
<paragraph id="8BAB2E62FA412A1231DD37AF0A2AFD75" blockId="17.[151,568,679,779]" box="[151,568,679,706]" pageId="17" pageNumber="571">
<heading id="D0E3990EFA412A1231DD37AF0A2AFD75" bold="true" box="[151,568,679,706]" fontSize="11" level="1" pageId="17" pageNumber="571" reason="1">
<emphasis id="B960F270FA412A1231DD37AF0A2AFD75" bold="true" box="[151,568,679,706]" pageId="17" pageNumber="571">
<taxonomicName id="4C1455E1FA412A1231DD37AF09C1FD75" authority="Safian" authorityName="Safian" authorityYear="2021" box="[151,467,679,706]" class="Insecta" family="Lycaenidae" genus="Cephetola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="571" phylum="Arthropoda" rank="species" species="wologizi" status="sp. nov.">
<emphasis id="B960F270FA412A1231DD37AF096EFD75" bold="true" box="[151,380,679,706]" italics="true" pageId="17" pageNumber="571">Cephetola wologizi</emphasis>
Sáfián
</taxonomicName>
<taxonomicNameLabel id="A2534F0BFA412A12309137A00A2AFD75" box="[475,568,680,706]" pageId="17" pageNumber="571" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAB2E62FA412A1231DD37C409A2FD51" blockId="17.[151,568,679,779]" box="[151,432,716,742]" pageId="17" pageNumber="571">urn:lsid:zoobank.org:act:</paragraph>
</subSubSection>
<subSubSection id="C30E7DE9FA412A1231DD37F809ACFCBC" box="[151,446,752,779]" pageId="17" pageNumber="571" type="description">
<paragraph id="8BAB2E62FA412A1231DD37F809ACFCBC" blockId="17.[151,568,679,779]" box="[151,446,752,779]" pageId="17" pageNumber="571">
(
<figureCitation id="132F32E7FA412A1231D537F8093FFCBD" box="[159,301,752,778]" captionStart="FIGURE 13" captionStartId="18.[151,250,1865,1889]" captionTargetBox="[164,1416,249,1826]" captionTargetId="figure-55@18.[151,1436,234,1839]" captionTargetPageId="18" captionText="FIGURE 13. Cephetola males. C. wologizi, holotype. A, upperside; B, underside. C. wologizi, paratype. C, upperside; D, underside. C. aurealiae, holotype, DRC. E, upperside, F, underside. The latter was accessed through the online type collection of RMCA (see link at the list of examined material)." figureDoi="http://doi.org/10.5281/zenodo.5046876" httpUri="https://zenodo.org/record/5046876/files/figure.png" pageId="17" pageNumber="571">Figs 13A–D</figureCitation>
;
<figureCitation id="132F32E7FA412A12307337F8099FFCBD" box="[313,397,752,778]" captionStart="FIGURE 14" captionStartId="19.[151,250,1460,1484]" captionTargetBox="[164,1426,210,1424]" captionTargetId="figure-28@19.[151,1436,179,1434]" captionTargetPageId="19" captionText="FIGURE 14. Cephetola male genitalia. C. wologizi, holotype, gen. prep.: SAFI00385.A, lateral view; C, aedeagus. C. aureliae, holotype. B, lateral view, D, aedeagus (reproduced from Libert’s original drawing (Libert 1999)." figureDoi="http://doi.org/10.5281/zenodo.5046880" httpUri="https://zenodo.org/record/5046880/files/figure.png" pageId="17" pageNumber="571">14A, C</figureCitation>
;
<figureCitation id="132F32E7FA412A1230D337F809A4FCBC" box="[409,438,752,779]" captionStart="FIGURE 15" captionStartId="20.[151,250,1142,1166]" captionTargetBox="[155,1432,183,1114]" captionTargetId="figure-20@20.[151,1436,179,1117]" captionTargetPageId="20" captionText="FIGURE 15. Type localities of C. wologizi and C. praecox and known localities of C. obscuralis, C. obscuralis ssp. (sensu Libert 2020) and C. aureliae." figureDoi="http://doi.org/10.5281/zenodo.5046884" httpUri="https://zenodo.org/record/5046884/files/figure.png" pageId="17" pageNumber="571">15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30E7DE9FA412A1231DD36300B0EFBB1" pageId="17" pageNumber="571" type="materials_examined">
<paragraph id="8BAB2E62FA412A1231DD36300C17FC2D" blockId="17.[151,1437,824,2039]" pageId="17" pageNumber="571">
<materialsCitation id="3B7C243FFA412A1231DD36300C17FC2D" ID-GBIF-Occurrence="3312256301" collectingDate="2018-11-04" collectingDateMax="2018-11-09" collectingDateMin="2018-11-04" collectionCode="ANHRT" country="Liberia" elevation="1043" latitude="8.124233" location="Belegizi Ridge and summit Mountains" longLatPrecision="1" longitude="-9.936364" municipality="Wologizi Mountains" pageId="17" pageNumber="571" specimenCode="SAFI00385" specimenCount="1" specimenCount-male="1" stateProvince="Lofa County" typeStatus="holotype">
<emphasis id="B960F270FA412A1231DD3630091FFCE5" bold="true" box="[151,269,824,850]" pageId="17" pageNumber="571">
<typeStatus id="54AF90C0FA412A1231DD3630091AFCE5" box="[151,264,824,850]" pageId="17" pageNumber="571" type="holotype">Holotype</typeStatus>
:
</emphasis>
•
<specimenCount id="9D12E5EBFA412A1230623631092FFCE4" box="[296,317,825,851]" pageId="17" pageNumber="571" type="male">♂</specimenCount>
<collectingCountry id="F3036EF2FA412A12300C363009A5FCE5" box="[326,439,824,850]" name="Liberia" pageId="17" pageNumber="571">LIBERIA</collectingCountry>
,
<collectingRegion id="49D0E080FA412A12308E36300A47FCE5" box="[452,597,824,850]" country="Liberia" name="Lofa" pageId="17" pageNumber="571">Lofa County</collectingRegion>
,
<collectingMunicipality id="6BCFB418FA412A12332B36300B58FCE5" box="[609,842,824,850]" pageId="17" pageNumber="571">Wologizi Mountains</collectingMunicipality>
,
<location id="8ECB78B9FA412A12321D36300D1FFCE5" LSID="urn:lsid:plazi:treatment:03BD9F74FA412A1031DD37AF0BACF85F:8ECB78B9FA412A12321D36300D1FFCE5" box="[855,1293,824,850]" country="Liberia" latitude="8.124233" longLatPrecision="1" longitude="-9.936364" municipality="Wologizi Mountains" name="Belegizi Ridge and summit Mountains" pageId="17" pageNumber="571" stateProvince="Lofa County">Belegizi Ridge and summit Mountains</location>
general collecting,
<geoCoordinate id="EE2048A5FA412A1231BA36540997FCC1" box="[240,389,860,886]" degrees="8" direction="north" minutes="7" orientation="latitude" pageId="17" pageNumber="571" precision="1" seconds="27.24" value="8.124233">8°7’27.24»N</geoCoordinate>
,
<geoCoordinate id="EE2048A5FA412A1230C536540A27FCC0" box="[399,565,860,887]" degrees="9" direction="west" minutes="56" orientation="longitude" pageId="17" pageNumber="571" precision="1" seconds="10.91" value="-9.936364">9°56’10.91»W</geoCoordinate>
,
<quantity id="4CEC8387FA412A12330A36540AC5FCC0" box="[576,727,860,887]" metricMagnitude="3" metricUnit="m" metricValue="1.043" metricValueMax="1.0859999999999999" metricValueMin="1.0" pageId="17" pageNumber="571" unit="m" value="1043.0" valueMax="1086.0" valueMin="1000.0">
<elevation id="0039C951FA412A12330A36540AC5FCC0" box="[576,727,860,887]" metricMagnitude="3" metricUnit="m" metricValue="1.043" metricValueMax="1.0859999999999999" metricValueMin="1.0" pageId="17" pageNumber="571" unit="m" value="1043.0" valueMax="1086.0" valueMin="1000.0">1000-1086 m</elevation>
</quantity>
,
<date id="FFAA08A2FA412A1233A836540B62FCC1" box="[738,880,860,886]" pageId="17" pageNumber="571" value="2018-11-04" valueMax="2018-11-09" valueMin="2018-11-04">
<collectingDate id="EFEEF14AFA412A1233A836540B62FCC1" box="[738,880,860,886]" pageId="17" pageNumber="571" value="2018-11-04" valueMax="2018-11-09" valueMin="2018-11-04">4–9.XI.2018</collectingDate>
</date>
<location id="8ECB78B9FA412A12323C36540BFDFCC1" LSID="urn:lsid:plazi:treatment:03BD9F74FA412A1031DD37AF0BACF85F:8ECB78B9FA412A12323C36540BFDFCC1" box="[886,1007,860,886]" country="Liberia" latitude="8.124233" longLatPrecision="1" longitude="-9.936364" municipality="Wologizi Mountains" name="Safian, Sz." pageId="17" pageNumber="571" stateProvince="Lofa County">Sáfián, Sz.</location>
,
<location id="8ECB78B9FA412A1232B036540CD6FCC0" LSID="urn:lsid:plazi:treatment:03BD9F74FA412A1031DD37AF0BACF85F:8ECB78B9FA412A1232B036540CD6FCC0" box="[1018,1220,860,887]" country="Liberia" latitude="8.124233" longLatPrecision="1" longitude="-9.936364" municipality="Wologizi Mountains" name="Simonics, G. Leg." pageId="17" pageNumber="571" stateProvince="Lofa County">Simonics, G. Leg.</location>
<collectionCode id="ED05B6A7FA412A12358236540D34FCC1" box="[1224,1318,860,886]" pageId="17" pageNumber="571">ANHRT</collectionCode>
unique ID number: ANHRTUK00058086. Gen. prep.
<specimenCode id="DBB28619FA412A12333636880B10FC2C" box="[636,770,896,923]" pageId="17" pageNumber="571">SAFI00385</specimenCode>
.
<location id="8ECB78B9FA412A12324636880B6CFC2D" LSID="urn:lsid:plazi:treatment:03BD9F74FA412A1031DD37AF0BACF85F:8ECB78B9FA412A12324636880B6CFC2D" box="[780,894,896,922]" country="Liberia" latitude="8.124233" longLatPrecision="1" longitude="-9.936364" municipality="Wologizi Mountains" name="Deposited" pageId="17" pageNumber="571" stateProvince="Lofa County">Deposited</location>
in
<collectionCode id="ED05B6A7FA412A1232EA36880C13FC2D" box="[928,1025,896,922]" pageId="17" pageNumber="571">ANHRT</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BAB2E62FA412A12318D36AC0B0EFBB1" blockId="17.[151,1437,824,2039]" pageId="17" pageNumber="571">
<emphasis id="B960F270FA412A12318D36AC092FFC09" bold="true" box="[199,317,932,958]" pageId="17" pageNumber="571">
<typeStatus id="54AF90C0FA412A12318D36AC092AFC09" box="[199,312,932,958]" pageId="17" pageNumber="571" type="paratype">Paratype</typeStatus>
:
</emphasis>
•
<specimenCount id="9D12E5EBFA412A12301336AD097CFC08" box="[345,366,933,959]" pageId="17" pageNumber="571" type="male">♂</specimenCount>
<collectingCountry id="F3036EF2FA412A12303D36AC09FAFC09" box="[375,488,932,958]" name="Liberia" pageId="17" pageNumber="571">LIBERIA</collectingCountry>
,
<collectingRegion id="49D0E080FA412A1230BF36AC0A95FC09" box="[501,647,932,958]" country="Liberia" name="Lofa" pageId="17" pageNumber="571">Lofa County</collectingRegion>
, Wologizi Mountains, Belegizi Ridge and summit Mountains general collecting,
<geoCoordinate id="EE2048A5FA412A12305936C009B5FC55" box="[275,423,968,994]" degrees="8" direction="north" minutes="7" orientation="latitude" pageId="17" pageNumber="571" precision="1" seconds="27.24" value="8.124233">8°7’27.24»N</geoCoordinate>
,
<geoCoordinate id="EE2048A5FA412A1230F836C00A4AFC54" box="[434,600,968,995]" degrees="9" direction="west" minutes="56" orientation="longitude" pageId="17" pageNumber="571" precision="1" seconds="10.91" value="-9.936364">9°56’10.91»W</geoCoordinate>
,
<quantity id="4CEC8387FA412A12332836C00AE8FC54" box="[610,762,968,995]" metricMagnitude="3" metricUnit="m" metricValue="1.043" metricValueMax="1.0859999999999999" metricValueMin="1.0" pageId="17" pageNumber="571" unit="m" value="1043.0" valueMax="1086.0" valueMin="1000.0">1000-1086 m</quantity>
,
<date id="FFAA08A2FA412A12324F36C00B81FC55" box="[773,915,968,994]" pageId="17" pageNumber="571" value="2018-11-04" valueMax="2018-11-09" valueMin="2018-11-04">4–9.XI.2018</date>
Sáfián, Sz., Simonics, G. Leg.
<collectionCode id="ED05B6A7FA412A1235A636C00D58FC55" box="[1260,1354,968,994]" pageId="17" pageNumber="571">ANHRT</collectionCode>
unique ID number: ANHRTUK00058083. Deposited in
<collectionCode id="ED05B6A7FA412A1233FD36E40B0AFBB1" box="[695,792,1004,1030]" pageId="17" pageNumber="571">ANHRT</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E7DE9FA412A12318D31180C66FA6D" pageId="17" pageNumber="571" type="description">
<paragraph id="8BAB2E62FA412A12318D31180C6AFAB5" blockId="17.[151,1437,824,2039]" pageId="17" pageNumber="571">
<emphasis id="B960F270FA412A12318D3118094BFB9D" bold="true" box="[199,345,1040,1066]" pageId="17" pageNumber="571">Description.</emphasis>
Forewing length: ca.
<quantity id="4CEC8387FA412A12330131180A8FFB9C" box="[587,669,1040,1067]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="571" unit="mm" value="15.0">15 mm</quantity>
. Wingspan: ca.
<quantity id="4CEC8387FA412A12320131180BA0FB9C" box="[843,946,1040,1067]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" pageId="17" pageNumber="571" unit="mm" value="28.5">28.5 mm</quantity>
. Wing shape typical of
<taxonomicName id="4C1455E1FA412A1235FC31180D3BFB9D" authorityName="Libert" authorityYear="1999" box="[1206,1321,1040,1066]" class="Insecta" family="Lycaenidae" genus="Cephetola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="571" phylum="Arthropoda" rank="genus">
<emphasis id="B960F270FA412A1235FC31180D3BFB9D" box="[1206,1321,1040,1066]" italics="true" pageId="17" pageNumber="571">Cephetola</emphasis>
</taxonomicName>
triangular with acute apex and slightly convex, curving outer margin. Upperside completely black with extensive deep metallic blue sheen on hindwing. Only scattered blue scales present on forewing (
<figureCitation id="132F32E7FA412A1232A731500C44FBC5" box="[1005,1110,1112,1138]" captionStart="FIGURE 13" captionStartId="18.[151,250,1865,1889]" captionTargetBox="[164,1416,249,1826]" captionTargetId="figure-55@18.[151,1436,234,1839]" captionTargetPageId="18" captionText="FIGURE 13. Cephetola males. C. wologizi, holotype. A, upperside; B, underside. C. wologizi, paratype. C, upperside; D, underside. C. aurealiae, holotype, DRC. E, upperside, F, underside. The latter was accessed through the online type collection of RMCA (see link at the list of examined material)." figureDoi="http://doi.org/10.5281/zenodo.5046876" httpUri="https://zenodo.org/record/5046876/files/figure.png" pageId="17" pageNumber="571">Fig. 13A</figureCitation>
). Underside dark brown with extensive graphite-grey in post-median on hindwing, also two zig-zagging grey submarginal lines and other grey spotting at hindwing base. Forewing with grey gently curving submarginal line, connected to an inner zig-zagging line. Further grey dusting present in median and base (
<figureCitation id="132F32E7FA412A1233BF31CC0B4EFB69" box="[757,860,1220,1246]" captionStart="FIGURE 13" captionStartId="18.[151,250,1865,1889]" captionTargetBox="[164,1416,249,1826]" captionTargetId="figure-55@18.[151,1436,234,1839]" captionTargetPageId="18" captionText="FIGURE 13. Cephetola males. C. wologizi, holotype. A, upperside; B, underside. C. wologizi, paratype. C, upperside; D, underside. C. aurealiae, holotype, DRC. E, upperside, F, underside. The latter was accessed through the online type collection of RMCA (see link at the list of examined material)." figureDoi="http://doi.org/10.5281/zenodo.5046876" httpUri="https://zenodo.org/record/5046876/files/figure.png" pageId="17" pageNumber="571">Fig. 13B</figureCitation>
). Legs and palpi dark brown, ringed with grey, eyes black, hairless, smooth. Antennae black ringed with grey, their club black with ochre tip.
</paragraph>
<paragraph id="8BAB2E62FA412A12318D30040C66FA6D" blockId="17.[151,1437,824,2039]" pageId="17" pageNumber="571">
<emphasis id="B960F270FA412A12318D3004096AFA91" bold="true" box="[199,376,1292,1318]" pageId="17" pageNumber="571">Male genitalia.</emphasis>
Medium,
<quantity id="4CEC8387FA412A1230A030040A2DFA90" box="[490,575,1292,1319]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="571" unit="mm" value="1.5">1.5 mm</quantity>
along dorsoventral axis. Slender and rather simple, similar to other species in the group. Coremata missing. Tegumen narrow, uncus triangular in lateral view, sub-unci angled, very slender, tip of longer arm very slightly upcurving. Valva triangular, broadening from base, broadest in the middle, narrows down to the dorsally pointy tip. The valva’s ventral edge slightly sinuous. Saccus slender, shorter than half the length of the valva (
<figureCitation id="132F32E7FA412A1230443094096AFA01" box="[270,376,1436,1462]" captionStart="FIGURE 14" captionStartId="19.[151,250,1460,1484]" captionTargetBox="[164,1426,210,1424]" captionTargetId="figure-28@19.[151,1436,179,1434]" captionTargetPageId="19" captionText="FIGURE 14. Cephetola male genitalia. C. wologizi, holotype, gen. prep.: SAFI00385.A, lateral view; C, aedeagus. C. aureliae, holotype. B, lateral view, D, aedeagus (reproduced from Libert’s original drawing (Libert 1999)." figureDoi="http://doi.org/10.5281/zenodo.5046880" httpUri="https://zenodo.org/record/5046880/files/figure.png" pageId="17" pageNumber="571">Fig. 14A</figureCitation>
). Aedeagus typical to many species in the genus, spear-shaped with triangular tooth on dorsal edge, broader in the middle. Anterior end rounded, tip narrows down acutely (
<figureCitation id="132F32E7FA412A12354B30C80C7AFA6D" box="[1025,1128,1472,1498]" captionStart="FIGURE 14" captionStartId="19.[151,250,1460,1484]" captionTargetBox="[164,1426,210,1424]" captionTargetId="figure-28@19.[151,1436,179,1434]" captionTargetPageId="19" captionText="FIGURE 14. Cephetola male genitalia. C. wologizi, holotype, gen. prep.: SAFI00385.A, lateral view; C, aedeagus. C. aureliae, holotype. B, lateral view, D, aedeagus (reproduced from Libert’s original drawing (Libert 1999)." figureDoi="http://doi.org/10.5281/zenodo.5046880" httpUri="https://zenodo.org/record/5046880/files/figure.png" pageId="17" pageNumber="571">Fig. 14C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30E7DE9FA412A12318D30EC09B0F9DD" pageId="17" pageNumber="571" type="etymology">
<paragraph id="8BAB2E62FA412A12318D30EC091CF9F1" blockId="17.[151,1437,824,2039]" pageId="17" pageNumber="571">
<emphasis id="B960F270FA412A12318D30EC095CFA49" bold="true" box="[199,334,1508,1534]" pageId="17" pageNumber="571">Etymology.</emphasis>
Masculine noun in apposition. The name refers to the species’
<typeStatus id="54AF90C0FA412A12355430ED0C5CFA48" box="[1054,1102,1509,1535]" pageId="17" pageNumber="571">type</typeStatus>
locality, the Wologizi Mountains. The mountains and the surrounding lowland forests are proposed to be protected to conserve the rich wildlife of
<collectingCountry id="F3036EF2FA412A1231FC33240919F9F1" box="[182,267,1580,1606]" name="Liberia" pageId="17" pageNumber="571">Liberia</collectingCountry>
.
</paragraph>
<paragraph id="8BAB2E62FA412A12318D335809B0F9DD" blockId="17.[151,1437,824,2039]" box="[199,418,1616,1643]" pageId="17" pageNumber="571">
<emphasis id="B960F270FA412A12318D3358090CF9DD" bold="true" box="[199,286,1616,1642]" pageId="17" pageNumber="571">Female</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30E7DE9FA412A11318D337C0C9EFF61" lastPageId="18" lastPageNumber="572" pageId="17" pageNumber="571" type="diagnosis">
<paragraph id="8BAB2E62FA412A12318D337C0BAEF8F5" blockId="17.[151,1437,824,2039]" pageId="17" pageNumber="571">
<emphasis id="B960F270FA412A12318D337C09DDF939" bold="true" box="[199,463,1652,1678]" pageId="17" pageNumber="571">Differential diagnosis.</emphasis>
The only
<taxonomicName id="4C1455E1FA412A123309337C0AA4F939" authorityName="Libert" authorityYear="1999" box="[579,694,1652,1678]" class="Insecta" family="Lycaenidae" genus="Cephetola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="571" phylum="Arthropoda" rank="genus">
<emphasis id="B960F270FA412A123309337C0AA4F939" box="[579,694,1652,1678]" italics="true" pageId="17" pageNumber="571">Cephetola</emphasis>
</taxonomicName>
with similar silvery-grey dusting on the hindwing underside with limited blue spotting on the forewing upperside is
<taxonomicName id="4C1455E1FA412A12339D33900B47F905" authorityName="Libert" authorityYear="1999" box="[727,853,1688,1714]" class="Insecta" family="Lycaenidae" genus="Cephetola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="571" phylum="Arthropoda" rank="species" species="aureliae">
<emphasis id="B960F270FA412A12339D33900B47F905" box="[727,853,1688,1714]" italics="true" pageId="17" pageNumber="571">C. aureliae</emphasis>
</taxonomicName>
, which is known only from Eastern
<collectingCountry id="F3036EF2FA412A1235BC33900D21F905" box="[1270,1331,1688,1714]" name="Democratic Republic of the Congo" pageId="17" pageNumber="571">DRC</collectingCountry>
. The two rather prominent blue spots on the forewing of
<taxonomicName id="4C1455E1FA412A1233E333B40B34F961" authorityName="Libert" authorityYear="1999" box="[681,806,1724,1750]" class="Insecta" family="Lycaenidae" genus="Cephetola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="571" phylum="Arthropoda" rank="species" species="aureliae">
<emphasis id="B960F270FA412A1233E333B40B34F961" box="[681,806,1724,1750]" italics="true" pageId="17" pageNumber="571">C. aureliae</emphasis>
</taxonomicName>
immediately sets it aside from the Liberian
<taxonomicName id="4C1455E1FA412A12345C33B40D87F961" authorityName="Safian" authorityYear="2021" box="[1302,1429,1724,1750]" class="Insecta" family="Lycaenidae" genus="Cephetola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="571" phylum="Arthropoda" rank="species" species="wologizi">
<emphasis id="B960F270FA412A12345C33B40D87F961" box="[1302,1429,1724,1750]" italics="true" pageId="17" pageNumber="571">C. wologizi</emphasis>
</taxonomicName>
, which lacks any blue spots on its forewing with only a few blue scattered scales present. The recently described, similar species
<taxonomicName id="4C1455E1FA412A12300F320C0A5EF8A8" authority="Safian, 2015" authorityName="Safian" authorityYear="2015" box="[325,588,1796,1823]" class="Insecta" family="Lycaenidae" genus="Cephetola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="571" phylum="Arthropoda" rank="species" species="wingae">
<emphasis id="B960F270FA412A12300F320C09A5F8A9" box="[325,439,1796,1822]" italics="true" pageId="17" pageNumber="571">C. wingae</emphasis>
<bibRefCitation id="EF855393FA412A1230F4320C0A5EF8A8" author="Safian, Sz." box="[446,588,1796,1823]" pageId="17" pageNumber="571" pagination="12 - 19" refId="ref10538" refString="Safian, Sz. (2015) Two new Epitolini from Liberia in the genera Stempfferia Jackson, 1962 and Cephetola Libert, 1999 (Lepidoptera: Lycaenidae). Metamorphosis, 26, 12 - 19." type="journal article" year="2015">Sáfián, 2015</bibRefCitation>
</taxonomicName>
lacks silvery-grey scaling on its underside. The unique shape of valva with the strongly hooked tip sets it aside from any known
<taxonomicName id="4C1455E1FA412A1233A332200BAEF8F5" box="[745,956,1832,1858]" class="Insecta" family="Lycaenidae" genus="Cephetola" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="571" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B960F270FA412A1233A332200B4EF8F5" box="[745,860,1832,1858]" italics="true" pageId="17" pageNumber="571">Cephetola</emphasis>
species.
</taxonomicName>
</paragraph>
<paragraph id="8BAB2E62FA412A11318D32440C9EFF61" blockId="17.[151,1437,824,2039]" lastBlockId="18.[151,1436,150,214]" lastPageId="18" lastPageNumber="572" pageId="17" pageNumber="571">
<emphasis id="B960F270FA412A12318D32440959F8D1" bold="true" box="[199,331,1868,1894]" pageId="17" pageNumber="571">Bionomics.</emphasis>
The
<typeStatus id="54AF90C0FA412A1230CE324409F4F8D1" box="[388,486,1868,1894]" pageId="17" pageNumber="571" type="holotype">holotype</typeStatus>
and the only
<typeStatus id="54AF90C0FA412A12333532450ACDF8D0" box="[639,735,1869,1895]" pageId="17" pageNumber="571" type="paratype">paratype</typeStatus>
were captured during display flight on a hilltop at
<quantity id="4CEC8387FA412A12345032440D7DF8D0" box="[1306,1391,1868,1895]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="571" unit="m" value="1000.0">1000 m</quantity>
asl. in the Wologizi Mountains (
<figureCitation id="132F32E7FA412A12309F32780A3AF83C" box="[469,552,1904,1931]" captionStart="FIGURE 15" captionStartId="20.[151,250,1142,1166]" captionTargetBox="[155,1432,183,1114]" captionTargetId="figure-20@20.[151,1436,179,1117]" captionTargetPageId="20" captionText="FIGURE 15. Type localities of C. wologizi and C. praecox and known localities of C. obscuralis, C. obscuralis ssp. (sensu Libert 2020) and C. aureliae." figureDoi="http://doi.org/10.5281/zenodo.5046884" httpUri="https://zenodo.org/record/5046884/files/figure.png" pageId="17" pageNumber="571">Fig. 15</figureCitation>
) displaying rather high at lower canopy level in a fire-disturbed upland forest between 9.30 and 10.00 a.m. The species was replaced by the slightly larger
<taxonomicName id="4C1455E1FA412A123544329C0D08F819" authority="(Kirby, 1890)" baseAuthorityName="Kirby" baseAuthorityYear="1890" box="[1038,1306,1940,1966]" class="Insecta" family="Lycaenidae" genus="Cephetola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="571" phylum="Arthropoda" rank="species" species="doleta">
<emphasis id="B960F270FA412A123544329C0C64F819" box="[1038,1142,1940,1966]" italics="true" pageId="17" pageNumber="571">C. doleta</emphasis>
(
<bibRefCitation id="EF855393FA412A1235CD329C0D00F819" author="Kirby, W. F." box="[1159,1298,1940,1966]" pageId="17" pageNumber="571" pagination="261 - 274" refId="ref9975" refString="Kirby, W. F. (1890) Descriptions of new species of African Lycaenidae chiefly from the collections of Dr. Staudinger and Mr. Henley Grose-Smith. Annals and Magazine of Natural History, 6, 261 - 274. https: // doi. org / 10.1080 / 00222939008694032" type="journal article" year="1890">Kirby, 1890</bibRefCitation>
)
</taxonomicName>
just before 10.00 with slight overlap in their display time. Males of other
<taxonomicName id="4C1455E1FA412A12321932B00BB7F865" authorityName="Westwood" authorityYear="1851" box="[851,933,1976,2002]" class="Insecta" family="Lycaenidae" genus="Epitola" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="571" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B960F270FA412A12321932B00BB7F865" box="[851,933,1976,2002]" italics="true" pageId="17" pageNumber="571">Epitola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2534F0BFA412A1232E732B00BDCF865" box="[941,974,1976,2002]" pageId="17" pageNumber="571" sensu="lato">s.l.</taxonomicNameLabel>
species also frequented the same hilltop;
<taxonomicName id="4C1455E1FA412A1231DD32D409A9F841" authority="(Druce, 1890)" baseAuthorityName="Druce" baseAuthorityYear="1890" box="[151,443,2012,2038]" class="Insecta" family="Lycaenidae" genus="Cephetola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="571" phylum="Arthropoda" rank="species" species="pinodes">
<emphasis id="B960F270FA412A1231DD32D40903F841" box="[151,273,2012,2038]" italics="true" pageId="17" pageNumber="571">C. pinodes</emphasis>
(
<bibRefCitation id="EF855393FA412A12306B32D409A1F841" author="Druce, H. H." box="[289,435,2012,2038]" pageId="17" pageNumber="571" pagination="24 - 31" refId="ref9499" refString="Druce, H. H. (1890) Descriptions of twelve new species of Lycaenidae from West Africa and one from the Solomon Islands in the collection of Herbert Druce. Annals and Magazine of Natural History, Series 6, 5 (25), 24 - 31. https: // doi. org / 10.1080 / 00222939009460774" type="journal article" year="1890">Druce, 1890</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C1455E1FA412A12308332D40B33F840" authority="(Roche, 1954)" baseAuthorityName="Roche" baseAuthorityYear="1954" box="[457,801,2012,2039]" class="Insecta" family="Lycaenidae" genus="Cephetola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="571" phylum="Arthropoda" rank="species" species="subcoerulea">
<emphasis id="B960F270FA412A12308332D40A66F841" box="[457,628,2012,2038]" italics="true" pageId="17" pageNumber="571">C. subcoerulea</emphasis>
(
<bibRefCitation id="EF855393FA412A1233CF32D40B08F840" author="Roche, P." box="[645,794,2012,2039]" pageId="17" pageNumber="571" pagination="489 - 501" refId="ref10442" refString="Roche, P. (1954) Revisional notes on the genus Epitola Westwood / Lepidoptera: Lycaenidae. Bulletin of the British Museum (Natural History, Entomology), 3, 489 - 501. https: // doi. org / 10.5962 / bhl. part. 1060" type="journal article" year="1954">Roche, 1954</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C1455E1FA412A12326532D40D68F841" authority="(Grose-Smith &amp; Kirby, [1892])" baseAuthorityName="Grose-Smith &amp; Kirby" baseAuthorityYear="1892" box="[815,1402,2011,2038]" class="Insecta" family="Lycaenidae" genus="Stempfferia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="571" phylum="Arthropoda" rank="species" species="ciconia">
<emphasis id="B960F270FA412A12326532D40C19F841" box="[815,1035,2012,2038]" italics="true" pageId="17" pageNumber="571">Stempfferia ciconia</emphasis>
(Grose-Smith &amp; Kirby, [1892])
</taxonomicName>
,
<taxonomicName id="4C1455E1FA412A1134CD32D40908FF06" baseAuthorityName="Bethune-Baker" baseAuthorityYear="1903" class="Insecta" family="Lycaenidae" genus="Stempfferia" kingdom="Animalia" lastPageId="18" lastPageNumber="572" order="Lepidoptera" pageId="17" pageNumber="571" phylum="Arthropoda" rank="species" species="moyambina">
<emphasis id="B960F270FA412A1134CD32D40908FF06" italics="true" lastPageId="18" lastPageNumber="572" pageId="17" pageNumber="571">S. moyambina</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1455E1FA422A113060359F0A23FF06" authority="Safian, 2015" authorityName="Safian" authorityYear="2015" box="[298,561,150,177]" class="Insecta" family="Lycaenidae" genus="Stempfferia" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="572" phylum="Arthropoda" rank="species" species="katikae">
<emphasis id="B960F270FA422A113060359F098BFF06" box="[298,409,151,177]" italics="true" pageId="18" pageNumber="572">S. katikae</emphasis>
<bibRefCitation id="EF855393FA422A1130EB359F0A23FF06" author="Safian, Sz." box="[417,561,150,177]" pageId="18" pageNumber="572" pagination="12 - 19" refId="ref10538" refString="Safian, Sz. (2015) Two new Epitolini from Liberia in the genera Stempfferia Jackson, 1962 and Cephetola Libert, 1999 (Lepidoptera: Lycaenidae). Metamorphosis, 26, 12 - 19." type="journal article" year="2015">Sáfián, 2015</bibRefCitation>
</taxonomicName>
were all collected on the same day (
<bibRefCitation id="EF855393FA422A113292359F0CA4FF06" author="Safian, Sz. &amp; Coache, A. &amp; Rainon, B." box="[984,1206,151,177]" pageId="18" pageNumber="572" pagination="1 - 3" refId="ref10664" refString="Safian, Sz., Coache, A. &amp; Rainon, B. (2020 a) New data on the distribution of Iridana agneshorvathae Collins, Larsen &amp; Safian, 2008 with description of the previously unknown female (Lepidoptera, Lycaenidae, Poritiinae). Faunitaxys, 8 (3), 1 - 3." type="journal article" year="2020">
Sáfián
<emphasis id="B960F270FA422A113560359F0C76FF06" box="[1066,1124,151,177]" italics="true" pageId="18" pageNumber="572">et al.</emphasis>
2020a
</bibRefCitation>
), as well as a single specimen of
<taxonomicName id="4C1455E1FA422A11306C35B30B8AFF62" authority="Collins, Larsen &amp; Safian, 2008" authorityName="Collins, Larsen &amp; Safian" authorityYear="2008" box="[294,920,187,213]" class="Insecta" family="Lycaenidae" genus="Iridana" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="572" phylum="Arthropoda" rank="species" species="agneshorvathae">
<emphasis id="B960F270FA422A11306C35B30A26FF62" box="[294,564,187,213]" italics="true" pageId="18" pageNumber="572">Iridana agneshorvathae</emphasis>
Collins, Larsen &amp; Sáfián, 2008
</taxonomicName>
(
<bibRefCitation id="EF855393FA422A1132EC35B30C92FF61" author="Safian, Sz. &amp; Koyvogui, S. &amp; Simonics, G. &amp; Florczyk, K." box="[934,1152,187,214]" pageId="18" pageNumber="572" pagination="104 - 128" refId="ref10769" refString="Safian, Sz., Koyvogui, S., Simonics, G. &amp; Florczyk, K. (2020 b) Butterfly diversity (Lepidoptera, Papilionoidea) in the Ziama Massif in Guinea and the adjacent Wologizi and Wonegizi Mountains in Liberia (West Africa). A landscape conservation approach. Metamorphosis, 31 (1), 104 - 128." type="journal article" year="2020">
Sáfián
<emphasis id="B960F270FA422A1132BC35B30C3DFF62" box="[1014,1071,187,213]" italics="true" pageId="18" pageNumber="572">et al.</emphasis>
2020b
</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="DF6B7EEAFA422A1131DD32410A9CF81E" ID-DOI="http://doi.org/10.5281/zenodo.5046876" ID-Zenodo-Dep="5046876" httpUri="https://zenodo.org/record/5046876/files/figure.png" pageId="18" pageNumber="572" startId="18.[151,250,1865,1889]" targetBox="[164,1416,249,1826]" targetPageId="18">
<paragraph id="8BAB2E62FA422A1131DD32410A9CF81E" blockId="18.[151,1436,1865,1961]" pageId="18" pageNumber="572">
<emphasis id="B960F270FA422A1131DD32410933F8D6" bold="true" box="[151,289,1865,1889]" pageId="18" pageNumber="572">FIGURE 13.</emphasis>
<taxonomicName id="4C1455E1FA422A11306232410982F8D6" authorityName="Libert" authorityYear="1999" box="[296,400,1865,1889]" class="Insecta" family="Lycaenidae" genus="Cephetola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="572" phylum="Arthropoda" rank="genus">
<emphasis id="B960F270FA422A11306232410982F8D6" box="[296,400,1865,1889]" italics="true" pageId="18" pageNumber="572">Cephetola</emphasis>
</taxonomicName>
males.
<taxonomicName id="4C1455E1FA422A1130AA32420A46F8D6" authorityName="Safian" authorityYear="2021" box="[480,596,1865,1889]" class="Insecta" family="Lycaenidae" genus="Cephetola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="572" phylum="Arthropoda" rank="species" species="wologizi">
<emphasis id="B960F270FA422A1130AA32420A46F8D6" box="[480,596,1865,1889]" italics="true" pageId="18" pageNumber="572">C. wologizi</emphasis>
</taxonomicName>
, holotype. A, upperside; B, underside.
<taxonomicName id="4C1455E1FA422A1132AF32420C4BF8D6" authorityName="Safian" authorityYear="2021" box="[997,1113,1865,1889]" class="Insecta" family="Lycaenidae" genus="Cephetola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="572" phylum="Arthropoda" rank="species" species="wologizi">
<emphasis id="B960F270FA422A1132AF32420C4BF8D6" box="[997,1113,1865,1889]" italics="true" pageId="18" pageNumber="572">C. wologizi</emphasis>
</taxonomicName>
, paratype. C, upperside; D, underside.
<emphasis id="B960F270FA422A1131A732660960F832" box="[237,370,1901,1925]" italics="true" pageId="18" pageNumber="572">
<taxonomicName id="4C1455E1FA422A1131A73266097CF832" box="[237,366,1901,1925]" class="Insecta" family="Lycaenidae" genus="Cephetola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="572" phylum="Arthropoda" rank="species" species="aurealiae">C. aurealiae</taxonomicName>
,
</emphasis>
holotype, DRC. E, upperside, F, underside. The latter was accessed through the online type collection of RMCA (see link at the list of examined material).
</paragraph>
</caption>
<caption id="DF6B7EEAFA432A1031DD30BC0C7BFA47" ID-DOI="http://doi.org/10.5281/zenodo.5046880" ID-Zenodo-Dep="5046880" httpUri="https://zenodo.org/record/5046880/files/figure.png" pageId="19" pageNumber="573" startId="19.[151,250,1460,1484]" targetBox="[164,1426,210,1424]" targetPageId="19">
<paragraph id="8BAB2E62FA432A1031DD30BC0C7BFA47" blockId="19.[151,1436,1460,1520]" pageId="19" pageNumber="573">
<emphasis id="B960F270FA432A1031DD30BC090DFA7B" bold="true" box="[151,287,1460,1484]" pageId="19" pageNumber="573">FIGURE 14.</emphasis>
<taxonomicName id="4C1455E1FA432A10306E30BC099EFA7B" authorityName="Libert" authorityYear="1999" box="[292,396,1460,1484]" class="Insecta" family="Lycaenidae" genus="Cephetola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="573" phylum="Arthropoda" rank="genus">
<emphasis id="B960F270FA432A10306E30BC099EFA7B" box="[292,396,1460,1484]" italics="true" pageId="19" pageNumber="573">Cephetola</emphasis>
</taxonomicName>
male genitalia.
<taxonomicName id="4C1455E1FA432A10336630BC0A8CFA7B" authorityName="Safian" authorityYear="2021" box="[556,670,1460,1484]" class="Insecta" family="Lycaenidae" genus="Cephetola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="573" phylum="Arthropoda" rank="species" species="wologizi">
<emphasis id="B960F270FA432A10336630BC0A8CFA7B" box="[556,670,1460,1484]" italics="true" pageId="19" pageNumber="573">C. wologizi</emphasis>
</taxonomicName>
, holotype, gen. prep.: SAFI00385. A, lateral view; C, aedeagus.
<taxonomicName id="4C1455E1FA432A10346C30BC0D87FA7B" authorityName="Libert" authorityYear="1999" box="[1318,1429,1460,1484]" class="Insecta" family="Lycaenidae" genus="Cephetola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="573" phylum="Arthropoda" rank="species" species="aureliae">
<emphasis id="B960F270FA432A10346C30BC0D87FA7B" box="[1318,1429,1460,1484]" italics="true" pageId="19" pageNumber="573">C. aureliae</emphasis>
</taxonomicName>
, holotype. B, lateral view, D, aedeagus (reproduced from Libert’s original drawing (
<bibRefCitation id="EF855393FA432A1032AB30D00C4CFA47" author="Libert, M." box="[993,1118,1496,1520]" pageId="19" pageNumber="573" refId="ref10087" refString="Libert, M. (1999) Revision des genres Epitola Westwood, Hypophytala Clench et Stempfferia Jackson, et description de trois nouveaux genres (Lepidoptera Lycaenidae). African Butterfly Research Institute, Nairobi &amp; Lambillionea, Tervuren, 219 pp." type="book" year="1999">Libert 1999</bibRefCitation>
).
</paragraph>
</caption>
<subSubSection id="C30E7DE9FA432A10318D33160BACF85F" pageId="19" pageNumber="573" type="discussion">
<paragraph id="8BAB2E62FA432A10318D331608EFF97E" blockId="19.[151,1437,1566,2025]" pageId="19" pageNumber="573">
<emphasis id="B960F270FA432A10318D33160926F98F" bold="true" box="[199,308,1566,1592]" pageId="19" pageNumber="573">Remarks</emphasis>
.
<taxonomicName id="4C1455E1FA432A103009331709D6F98F" authorityName="Safian" authorityYear="2021" box="[323,452,1566,1593]" class="Insecta" family="Lycaenidae" genus="Cephetola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="573" phylum="Arthropoda" rank="species" species="wologizi">
<emphasis id="B960F270FA432A103009331709D6F98F" box="[323,452,1566,1593]" italics="true" pageId="19" pageNumber="573">C. wologizi</emphasis>
</taxonomicName>
has no close relative in West Africa. Actually, the only similar-looking species,
<taxonomicName id="4C1455E1FA432A10341F331708C5F9EB" authorityName="Libert" authorityYear="1999" class="Insecta" family="Lycaenidae" genus="Cephetola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="573" phylum="Arthropoda" rank="species" species="aureliae">
<emphasis id="B960F270FA432A10341F331708C5F9EB" italics="true" pageId="19" pageNumber="573">C. aureliae</emphasis>
</taxonomicName>
with very different genitalia, is known from a few specimens: The
<typeStatus id="54AF90C0FA432A103299334A0C27F9EB" box="[979,1077,1602,1628]" pageId="19" pageNumber="573" type="holotype">holotype</typeStatus>
of that species was collected in savannah-forest mosaic in the Semuliki Valley at the foothills of Rwenzori in the
<collectingCountry id="F3036EF2FA432A10357B336E0C79F937" box="[1073,1131,1638,1664]" name="Democratic Republic of the Congo" pageId="19" pageNumber="573">DRC</collectingCountry>
(
<bibRefCitation id="EF855393FA432A103530336E0D16F937" author="Libert, M." box="[1146,1284,1638,1664]" pageId="19" pageNumber="573" refId="ref10087" refString="Libert, M. (1999) Revision des genres Epitola Westwood, Hypophytala Clench et Stempfferia Jackson, et description de trois nouveaux genres (Lepidoptera Lycaenidae). African Butterfly Research Institute, Nairobi &amp; Lambillionea, Tervuren, 219 pp." type="book" year="1999">Libert 1999</bibRefCitation>
), other specimens were captured in mid-altitude forest near Djugu (
<bibRefCitation id="EF855393FA432A10324833820BBEF913" author="Ducarme, R." box="[770,940,1674,1700]" pageId="19" pageNumber="573" pagination="23 - 37" refId="ref9560" refString="Ducarme, R. (2018) The butterflies (Lepidoptera: Papilionoidea) of the north-eastern Democratic Republic of Congo. Metamorphosis, 29, 23 - 37." type="journal article" year="2018">Ducarme 2018</bibRefCitation>
) some
<quantity id="4CEC8387FA432A1032B433820C73F913" box="[1022,1121,1674,1700]" metricMagnitude="6" metricUnit="m" metricValue="4.6" pageId="19" pageNumber="573" unit="km" value="4600.0">4600 km</quantity>
from the type locality of
<taxonomicName id="4C1455E1FA432A1034C9338308E4F97F" authorityName="Safian" authorityYear="2021" class="Insecta" family="Lycaenidae" genus="Cephetola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="573" phylum="Arthropoda" rank="species" species="wologizi">
<emphasis id="B960F270FA432A1034C9338308E4F97F" italics="true" pageId="19" pageNumber="573">C. wologizi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BAB2E62FA432A10318D33DA0BACF85F" blockId="19.[151,1437,1566,2025]" pageId="19" pageNumber="573">
Since most species of smaller
<taxonomicName id="4C1455E1FA432A10335C33DA0A7AF95B" authorityName="Westwood" authorityYear="1851" box="[534,616,1746,1772]" class="Insecta" family="Lycaenidae" genus="Epitola" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="573" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B960F270FA432A10335C33DA0A7AF95B" box="[534,616,1746,1772]" italics="true" pageId="19" pageNumber="573">Epitola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2534F0BFA432A10332533DA0A82F95B" box="[623,656,1746,1772]" pageId="19" pageNumber="573" sensu="lato">s.l.</taxonomicNameLabel>
are often difficult to collect, it would not be appropriate to even guess about the distribution of
<taxonomicName id="4C1455E1FA432A1030FC33FF0A2AF8A7" authorityName="Safian" authorityYear="2021" box="[438,568,1782,1809]" class="Insecta" family="Lycaenidae" genus="Cephetola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="573" phylum="Arthropoda" rank="species" species="wologizi">
<emphasis id="B960F270FA432A1030FC33FF0A2AF8A7" box="[438,568,1782,1809]" italics="true" pageId="19" pageNumber="573">C. wologizi</emphasis>
</taxonomicName>
. However, its occurrence in the upland zone of Wologizi Mountains would predict its presence in other mountainous localities in the
<collectingCountry id="F3036EF2FA432A10326B32120B60F883" box="[801,882,1818,1844]" name="Guinea-Bissau" pageId="19" pageNumber="573">Guinea</collectingCountry>
Highlands, particularly in the Nimba Mountains, where another undescribed species of
<taxonomicName id="4C1455E1FA432A10330F32360AAAF8EF" authorityName="Libert" authorityYear="1999" box="[581,696,1854,1880]" class="Insecta" family="Lycaenidae" genus="Cephetola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="573" phylum="Arthropoda" rank="genus">
<emphasis id="B960F270FA432A10330F32360AAAF8EF" box="[581,696,1854,1880]" italics="true" pageId="19" pageNumber="573">Cephetola</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1455E1FA432A10338D32370BDDF8EE" authority="Safian, 2015" authorityName="Safian" authorityYear="2015" box="[711,975,1854,1881]" class="Insecta" family="Lycaenidae" genus="Cephetola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="573" phylum="Arthropoda" rank="species" species="wingae">
<emphasis id="B960F270FA432A10338D32370B2BF8EF" box="[711,825,1855,1881]" italics="true" pageId="19" pageNumber="573">C. wingae</emphasis>
<bibRefCitation id="EF855393FA432A10320B32360BDDF8EE" author="Safian, Sz." box="[833,975,1854,1881]" pageId="19" pageNumber="573" pagination="12 - 19" refId="ref10538" refString="Safian, Sz. (2015) Two new Epitolini from Liberia in the genera Stempfferia Jackson, 1962 and Cephetola Libert, 1999 (Lepidoptera: Lycaenidae). Metamorphosis, 26, 12 - 19." type="journal article" year="2015">Sáfián, 2015</bibRefCitation>
</taxonomicName>
was recently found.
<taxonomicName id="4C1455E1FA432A1035F632370D3CF8EF" authorityName="Safian" authorityYear="2015" box="[1212,1326,1855,1881]" class="Insecta" family="Lycaenidae" genus="Cephetola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="573" phylum="Arthropoda" rank="species" species="wingae">
<emphasis id="B960F270FA432A1035F632370D3CF8EF" box="[1212,1326,1855,1881]" italics="true" pageId="19" pageNumber="573">C. wingae</emphasis>
</taxonomicName>
has since been recorded also in the Wologizi Mountains, not far from the
<typeStatus id="54AF90C0FA432A103238326B0BB0F8CA" box="[882,930,1891,1917]" pageId="19" pageNumber="573">type</typeStatus>
locality of
<taxonomicName id="4C1455E1FA432A103562326B0D42F8CB" authority="(Safian 2020)" baseAuthorityName="Safian" baseAuthorityYear="2020" box="[1064,1360,1890,1917]" class="Insecta" family="Lycaenidae" genus="Cephetola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="573" phylum="Arthropoda" rank="species" species="wologizi">
<emphasis id="B960F270FA432A103562326B0CBBF8CB" box="[1064,1193,1890,1917]" italics="true" pageId="19" pageNumber="573">C. wologizi</emphasis>
(
<bibRefCitation id="EF855393FA432A1035F3326A0D5AF8CB" author="Safian, Sz." box="[1209,1352,1890,1916]" pageId="19" pageNumber="573" pagination="219 - 230" refId="ref10612" refString="Safian, Sz. (2020) Description of the previously unknown sexes in four West African Lycaenidae (Lepidoptera) with new data on their distribution. Zootaxa. 4834 (2), 219 - 230. https: // doi. org / 10.11646 / zootaxa. 4834.2.4." type="journal article" year="2020">Sáfián 2020</bibRefCitation>
)
</taxonomicName>
. Other ‘Nimba specials’ such as
<taxonomicName id="4C1455E1FA432A1030F2328E0B50F817" authority="Hecq, 1987" authorityName="Hecq" authorityYear="1987" box="[440,834,1926,1953]" class="Insecta" family="Nymphalidae" genus="Hypolimnas" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="573" phylum="Arthropoda" rank="species" species="aubergeri">
<emphasis id="B960F270FA432A1030F2328E0AA6F817" box="[440,692,1926,1952]" italics="true" pageId="19" pageNumber="573">Hypolimnas aubergeri</emphasis>
<bibRefCitation id="EF855393FA432A1033F7328F0B50F817" author="Hecq, J." box="[701,834,1926,1953]" pageId="19" pageNumber="573" pagination="26 - 28" refId="ref9729" refString="Hecq, J. (1987) Une nouvelle espece d' Hypolymnas [Hypolimnas] Hubner (Nymphalidae africain). Lambillionea, 87 (3 - 4), 26 - 28." type="journal article" year="1987">Hecq, 1987</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1455E1FA432A103204328E0CDFF817" authority="Hecq, 1977" authorityName="Hecq" authorityYear="1977" box="[846,1229,1926,1953]" class="Insecta" family="Nymphalidae" genus="Euphaedra" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="573" phylum="Arthropoda" rank="species" species="aubergeri">
<emphasis id="B960F270FA432A103204328E0C52F817" box="[846,1088,1926,1952]" italics="true" pageId="19" pageNumber="573">Euphaedra aubergeri</emphasis>
<bibRefCitation id="EF855393FA432A103502328F0CDFF817" author="Hecq, J." box="[1096,1229,1926,1953]" pageId="19" pageNumber="573" pagination="51 - 56" refId="ref9690" refString="Hecq, J. (1977) Trois nouvelles espece d' Euphaedra (Lepid. Nymph. afr.). Note no. 10. Lambillionea, 77 (7 - 8), 51 - 56." type="journal article" year="1977">Hecq, 1977</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1455E1FA432A103593328E0A12F873" authority="(Collins &amp; Larsen, 2000)" baseAuthorityName="Collins &amp; Larsen" baseAuthorityYear="2000" class="Insecta" family="Hesperiidae" genus="Bettonula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="573" phylum="Arthropoda" rank="subSpecies" species="bettoni" subSpecies="nimba">
<emphasis id="B960F270FA432A103593328E08CFF873" italics="true" pageId="19" pageNumber="573">Bettonula bettoni nimba</emphasis>
(
<bibRefCitation id="EF855393FA432A1031A632A209EBF873" author="Collins, S. C. &amp; Larsen, T. B." box="[236,505,1962,1989]" pageId="19" pageNumber="573" pagination="57 - 75" refId="ref9226" refString="Collins, S. C. &amp; Larsen, T. B. (2000) Eight new species and five new subspecies of African butterflies (Rhopalocera) - an ABRI research paper. Metamorphosis, 11 (2), 57 - 75." type="journal article" year="2000">Collins &amp; Larsen, 2000</bibRefCitation>
)
</taxonomicName>
have also been found in Wologizi (
<bibRefCitation id="EF855393FA432A1032DA32A20C79F872" author="Safian, Sz. &amp; Koyvogui, S. &amp; Simonics, G. &amp; Florczyk, K." box="[912,1131,1962,1989]" pageId="19" pageNumber="573" pagination="104 - 128" refId="ref10769" refString="Safian, Sz., Koyvogui, S., Simonics, G. &amp; Florczyk, K. (2020 b) Butterfly diversity (Lepidoptera, Papilionoidea) in the Ziama Massif in Guinea and the adjacent Wologizi and Wonegizi Mountains in Liberia (West Africa). A landscape conservation approach. Metamorphosis, 31 (1), 104 - 128." type="journal article" year="2020">
Sáfián
<emphasis id="B960F270FA432A1032AB32A30C0BF873" box="[993,1049,1962,1988]" italics="true" pageId="19" pageNumber="573">et al.</emphasis>
2020b
</bibRefCitation>
), indicating wider distribution of such pre-montane and upland elements of the
<collectingCountry id="F3036EF2FA432A1033A632C60B2FF85F" box="[748,829,1998,2024]" name="Guinea-Bissau" pageId="19" pageNumber="573">Guinea</collectingCountry>
Highlands.
</paragraph>
</subSubSection>
</treatment>
</document>
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