238 lines
31 KiB
XML
238 lines
31 KiB
XML
<document id="673965C27F4B531A880E103175ED504E" ID-CLB-Dataset="27764" ID-DOI="10.11646/zootaxa.4532.1.1" ID-GBIF-Dataset="b3fb460f-20b4-4c62-98ea-a07d901bbfc0" ID-ISSN="1175-5326" ID-Zenodo-Dep="2614967" ID-ZooBank="4E9D0908-0933-48AF-A6ED-F3B8D39E8994" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553787614041" checkinUser="plazi" docAuthor="Cairns, Stephen D." docDate="2018" docId="03B0147FFFF8FFDC76CC6D4D45C3FB55" docLanguage="en" docName="zootaxa.4532.1.1.pdf" docOrigin="Zootaxa 4532 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Narella merga Cairns 2018, n. sp." docType="treatment" docVersion="13" lastPageNumber="25" masterDocId="FF896C07FFEEFFC4765B6A1F4732FF92" masterDocTitle="Primnoidae (Cnidaria: Octocorallia: Calcaxonia) of the Okeanos Explorer expeditions (CAPSTONE) to the central Pacific" masterLastPageNumber="43" masterPageNumber="1" pageNumber="23" updateTime="1698714965598" updateUser="ExternalLinkService">
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<mods:titleInfo id="47DACC329FA014B5B2B6306B7BC1FE97">
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<mods:title id="61DE23BC0638B6EA3EBF95BC859D1A45">Primnoidae (Cnidaria: Octocorallia: Calcaxonia) of the Okeanos Explorer expeditions (CAPSTONE) to the central Pacific</mods:title>
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<mods:name id="A3663AC1954769E02D4135EC2D31408E" type="personal">
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<mods:roleTerm id="40CD3E521EA7C713625D23F401C12571">Author</mods:roleTerm>
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<mods:namePart id="831616EF4692FD0937B6EED1B62470F7">Cairns, Stephen D.</mods:namePart>
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<mods:typeOfResource id="FEA6C03365EE8A32A286901DA44E99CE">text</mods:typeOfResource>
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<mods:title id="D1DA69E09CC24FE46F5295BE29C90FDA">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="90D0F349DC3E0472FC3FB15EAABB1D06">
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<mods:date id="7DC8698222D610DCAB440349CD208257">2018</mods:date>
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<mods:number id="18684584C5A69DC77C7517E1D04C8CD6">2018-12-17</mods:number>
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<mods:detail id="F76FE26024EB631EE1DD827F8637A6A0" type="volume">
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<mods:number id="4A3003366B6C6BD471A8564CFBBA6FCF">4532</mods:number>
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<mods:detail id="49602AC765A36F59085DC3B6922FAFAA" type="issue">
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<mods:classification id="B1A8925EFE34478826EE6860D51D30FA">journal article</mods:classification>
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<mods:identifier id="C8E66008B2A15196CB2C72A404473ABC" type="DOI">10.11646/zootaxa.4532.1.1</mods:identifier>
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<treatment id="03B0147FFFF8FFDC76CC6D4D45C3FB55" ID-DOI="http://doi.org/10.5281/zenodo.5951529" ID-GBIF-Taxon="154521080" ID-Zenodo-Dep="5951529" LSID="urn:lsid:plazi:treatment:03B0147FFFF8FFDC76CC6D4D45C3FB55" httpUri="http://treatment.plazi.org/id/03B0147FFFF8FFDC76CC6D4D45C3FB55" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23">
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<subSubSection id="C303F6E2FFF8FFD276CC6D4D46A5F8FF" box="[151,407,1874,1901]" pageId="22" pageNumber="23" type="nomenclature">
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<paragraph id="8BA6A569FFF8FFD276CC6D4D46A5F8FF" blockId="22.[151,407,1874,1937]" box="[151,407,1874,1901]" pageId="22" pageNumber="23">
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<heading id="D0EE1205FFF8FFD276CC6D4D46A5F8FF" bold="true" box="[151,407,1874,1901]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
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<emphasis id="B96D797BFFF8FFD276CC6D4D46A5F8FF" bold="true" box="[151,407,1874,1901]" pageId="22" pageNumber="23">
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<taxonomicName id="4C19DEEAFFF8FFD276CC6D4D4675F8FF" ID-CoL="45PN7" authority="Cairns, 2018" authorityName="Cairns" authorityYear="2018" box="[151,327,1874,1901]" class="Anthozoa" family="Primnoidae" genus="Narella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="22" pageNumber="23" phylum="Cnidaria" rank="species" species="merga" status="sp. nov.">
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<emphasis id="B96D797BFFF8FFD276CC6D4D4675F8FF" bold="true" box="[151,327,1874,1901]" italics="true" pageId="22" pageNumber="23">Narella merga</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25EC400FFF8FFD2770E6D4C46A5F8FF" box="[341,407,1875,1901]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C303F6E2FFF8FFD276CC6D684660F803" box="[151,338,1911,1937]" pageId="22" pageNumber="23" type="description">
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<paragraph id="8BA6A569FFF8FFD276CC6D684660F803" blockId="22.[151,407,1874,1937]" box="[151,338,1911,1937]" pageId="22" pageNumber="23">
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<figureCitation id="1322B9ECFFF8FFD276CC6D6847CDF803" box="[151,255,1911,1937]" captionStart="FIGURE 1" captionStartId="3.[151,250,1774,1799]" captionTargetBox="[174,1414,166,1753]" captionTargetId="figure@3.[174,1414,166,1753]" captionTargetPageId="3" captionText="FIGURE 1. In situ images of various species: A, Callogorgia cracentis, holotype. B, Calyptrophora lyra, holotype (not in situ). C, C. diaphana, EX1606-9-1. D, C. carinata, holotype. E, C. clarki, EX1605-L3-6. F, C. pourtalesi, holotype. G, C. distolos, holotype. H, Candidella gigantea, EX1606-6. I, Macroprimnoa ornata, holotype. J, Narella virgosa, holotype. K-L, N. aurantiaca, holotype. M, N. merga, holotype. N, N. horrida, EX1703-2-3. O, N. fordi, holotype. P, N. calamus, holotype. Q, N. virgosa, paratype, EX1504-L4-9-4. R, N. macrocalyx, EX1606-7. S, N. hawaiiensis, EX1504-9-2. T, Paracalyotrophora hawaiiensis, EX1703-2-1. U, P. echinata, EX1703-10-1. V, P. spiralis, holotype. W, Plumarella circumoperculum, EX1603-2-3. X, Thouarella hilgendorfi, EX1605-L1-17." httpUri="https://zenodo.org/record/2614969/files/figure.png" pageId="22" pageNumber="23">Figs. 1M</figureCitation>
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,
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<figureCitation id="1322B9ECFFF8FFD277516D684610F803" box="[266,290,1911,1937]" captionStart="FIGURE 11" captionStartId="23.[151,250,1810,1835]" captionTargetBox="[170,1410,191,1778]" captionTargetId="figure@23.[170,1410,191,1778]" captionTargetPageId="23" captionText="FIGURE 11. Narella merga, holotype: A–C, Stereo views of lateral and adaxial sides of a polyp. D, basal scales. E, outer surface of medial scales. F, buccal scales. G, adaxial buccal scales. H, opercular scales. I, coenenchymal scales. J, pinnular scale. Scale bars in mm." httpUri="https://zenodo.org/record/2614987/files/figure.png" pageId="22" pageNumber="23">11</figureCitation>
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A–J
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</paragraph>
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</subSubSection>
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<subSubSection id="C303F6E2FFF8FFD276CC6DA0454DF86F" pageId="22" pageNumber="23" type="etymology">
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<paragraph id="8BA6A569FFF8FFD276CC6DA0454DF86F" blockId="22.[151,1437,1983,2045]" pageId="22" pageNumber="23">
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<emphasis id="B96D797BFFF8FFD276CC6DA0462AF84B" bold="true" box="[151,280,1983,2009]" pageId="22" pageNumber="23">Etymology</emphasis>
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. Named
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<taxonomicName id="4C19DEEAFFF8FFD277DA6DDF46FAF84B" authorityName="Cairns" authorityYear="2018" box="[385,456,1984,2009]" class="Anthozoa" family="Primnoidae" genus="Narella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="22" pageNumber="23" phylum="Cnidaria" rank="species" species="merga">
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<emphasis id="B96D797BFFF8FFD277DA6DDF46FAF84B" box="[385,456,1984,2009]" italics="true" pageId="22" pageNumber="23">merga</emphasis>
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</taxonomicName>
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(Latin for a two-pronged pitchfork), in allusion to the growth form of the colony. The name is considered as a noun in apposition.
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</paragraph>
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</subSubSection>
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<caption id="DF66F5E1FFF9FFD376CC6D0D467CF8E0" httpUri="https://zenodo.org/record/2614987/files/figure.png" pageId="23" pageNumber="24" startId="23.[151,250,1810,1835]" targetBox="[170,1410,191,1778]" targetPageId="23">
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<paragraph id="8BA6A569FFF9FFD376CC6D0D467CF8E0" blockId="23.[151,1437,1810,1907]" pageId="23" pageNumber="24">
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<emphasis id="B96D797BFFF9FFD376CC6D0D462DF8B9" bold="true" box="[151,287,1810,1835]" pageId="23" pageNumber="24">FIGURE 11.</emphasis>
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<taxonomicName id="4C19DEEAFFF9FFD3777E6D0D468AF8B8" authorityName="Cairns" authorityYear="2018" box="[293,440,1810,1834]" class="Anthozoa" family="Primnoidae" genus="Narella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="23" pageNumber="24" phylum="Cnidaria" rank="species" species="merga">
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<emphasis id="B96D797BFFF9FFD3777E6D0D468AF8B8" box="[293,440,1810,1834]" italics="true" pageId="23" pageNumber="24">Narella merga</emphasis>
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</taxonomicName>
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, holotype:
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<emphasis id="B96D797BFFF9FFD374706D0D456EF8B9" bold="true" box="[555,604,1810,1835]" pageId="23" pageNumber="24">A–C</emphasis>
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, Stereo views of lateral and adaxial sides of a polyp.
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<emphasis id="B96D797BFFF9FFD372216D0C43BEF8B9" bold="true" box="[1146,1164,1811,1835]" pageId="23" pageNumber="24">D</emphasis>
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, basal scales.
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<emphasis id="B96D797BFFF9FFD373466D0C421CF8B9" bold="true" box="[1309,1326,1811,1835]" pageId="23" pageNumber="24">E</emphasis>
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, outer surface of medial scales.
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<emphasis id="B96D797BFFF9FFD377236D2846B5F8DD" bold="true" box="[376,391,1847,1871]" pageId="23" pageNumber="24">F</emphasis>
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, buccal scales.
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<emphasis id="B96D797BFFF9FFD3747D6D294508F8DD" bold="true" box="[550,570,1846,1871]" pageId="23" pageNumber="24">G</emphasis>
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, adaxial buccal scales.
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<emphasis id="B96D797BFFF9FFD3757C6D284409F8DD" bold="true" box="[807,827,1847,1871]" pageId="23" pageNumber="24">H</emphasis>
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, opercular scales.
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<emphasis id="B96D797BFFF9FFD375AC6D284333F8DD" bold="true" box="[1015,1025,1847,1871]" pageId="23" pageNumber="24">I</emphasis>
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, coenenchymal scales.
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<emphasis id="B96D797BFFF9FFD372B46D2843CEF8DD" bold="true" box="[1263,1276,1847,1871]" pageId="23" pageNumber="24">J</emphasis>
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, pinnular scale. Scale bars in mm.
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</paragraph>
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</caption>
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<subSubSection id="C303F6E2FFF9FFDC769C6DBE456BFF21" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="materials_examined">
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||
<paragraph id="8BA6A569FFF9FFD3769C6DBE42AEF829" blockId="23.[151,1436,1952,2016]" box="[199,1436,1952,1979]" pageId="23" pageNumber="24">
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<emphasis id="B96D797BFFF9FFD3769C6DBE46D2F829" bold="true" box="[199,480,1952,1979]" pageId="23" pageNumber="24">Type and Type Locality</emphasis>
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.
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<materialsCitation id="3B71AF34FFF9FFD377B56DBE42AEF829" ID-GBIF-Occurrence="2117768082" box="[494,1436,1952,1979]" collectingDate="2436" collectingDateMax="2440" collectingDateMin="2436" collectionCode="SEM" pageId="23" pageNumber="24" specimenCode="USNM 1424202" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54A21BCBFFF9FFD377B56DBE4564F829" box="[494,598,1953,1979]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
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and
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<collectionCode id="ED083DACFFF9FFD374D66DBE45F5F829" box="[653,711,1953,1979]" pageId="23" pageNumber="24">SEM</collectionCode>
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stubs
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<collectingDate id="EFE37A41FFF9FFD375556DBE44B8F829" box="[782,906,1953,1979]" pageId="23" pageNumber="24" value="2436" valueMax="2440">
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<date id="FFA783A9FFF9FFD375556DBE44B8F829" box="[782,906,1953,1979]" pageId="23" pageNumber="24" value="2436" valueMax="2440">2436-2440</date>
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</collectingDate>
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,
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<specimenCode id="DBBF0D12FFF9FFD375CE6DBE4361F829" box="[917,1107,1953,1979]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="23" pageNumber="24" type="Museum">USNM 1424202</specimenCode>
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. Type Locality:
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<emphasis id="B96D797BFFF9FFD373536DBD4218F829" box="[1288,1322,1954,1979]" italics="true" pageId="23" pageNumber="24">EX</emphasis>
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1606-502,
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</materialsCitation>
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</paragraph>
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<paragraph id="8BA6A569FFF9FFD376CC6DDB427BF84D" blockId="23.[151,1436,1952,2016]" box="[151,1353,1988,2016]" pageId="23" pageNumber="24">
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19.15048˚N, 164.5587˚ E (
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<collectingCountry id="F30EE5F9FFF9FFD3779E6DDA450DF84D" box="[453,575,1989,2015]" name="Heard Island and McDonald Islands" pageId="23" pageNumber="24">McDonald</collectingCountry>
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Guyot, Mid-Pacific Mountains Range west of
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<collectingCountry id="F30EE5F9FFF9FFD372106DDA43E8F872" box="[1099,1242,1989,2016]" name="Wake Island" pageId="23" pageNumber="24">Wake Island</collectingCountry>
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),
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<quantity id="4CE1088CFFF9FFD372B66DDB4277F84D" box="[1261,1349,1988,2015]" metricMagnitude="3" metricUnit="m" metricValue="2.575" pageId="23" pageNumber="24" unit="m" value="2575.0">2575 m</quantity>
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.
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</paragraph>
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<paragraph id="8BA6A569FFF6FFDC769C6A87456BFF21" blockId="24.[151,1437,152,1223]" box="[199,601,152,179]" pageId="24" pageNumber="25">
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||
<emphasis id="B96D797BFFF6FFDC769C6A87469DFF21" bold="true" box="[199,431,152,179]" pageId="24" pageNumber="25">Material Examined</emphasis>
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||
. The
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<typeStatus id="54A21BCBFFF6FFDC77AB6A864564FF21" box="[496,598,153,179]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C303F6E2FFF6FFDC769C6AA34599FB31" pageId="24" pageNumber="25" type="description">
|
||
<paragraph id="8BA6A569FFF6FFDC769C6AA34261FEF5" blockId="24.[151,1437,152,1223]" pageId="24" pageNumber="25">
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<emphasis id="B96D797BFFF6FFDC769C6AA34663FF45" bold="true" box="[199,337,188,215]" pageId="24" pageNumber="25">Description</emphasis>
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. Only the distal
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||
<quantity id="4CE1088CFFF6FFDC744F6AA34550FF45" box="[532,610,188,215]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" pageId="24" pageNumber="25" unit="cm" value="9.5">9.5 cm</quantity>
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||
(19 whorls) of the
|
||
<typeStatus id="54A21BCBFFF6FFDC751F6AA24494FF45" box="[836,934,189,215]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
|
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was collected, representing just part of one of the two branches, but
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<emphasis id="B96D797BFFF6FFDC77E96AFD46CAFF69" box="[434,504,226,251]" italics="true" pageId="24" pageNumber="25">in situ</emphasis>
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||
photographs (
|
||
<figureCitation id="1322B9ECFFF6FFDC74C76AFE45CBFF69" box="[668,761,225,251]" captionStart="FIGURE 1" captionStartId="3.[151,250,1774,1799]" captionTargetBox="[174,1414,166,1753]" captionTargetId="figure@3.[174,1414,166,1753]" captionTargetPageId="3" captionText="FIGURE 1. In situ images of various species: A, Callogorgia cracentis, holotype. B, Calyptrophora lyra, holotype (not in situ). C, C. diaphana, EX1606-9-1. D, C. carinata, holotype. E, C. clarki, EX1605-L3-6. F, C. pourtalesi, holotype. G, C. distolos, holotype. H, Candidella gigantea, EX1606-6. I, Macroprimnoa ornata, holotype. J, Narella virgosa, holotype. K-L, N. aurantiaca, holotype. M, N. merga, holotype. N, N. horrida, EX1703-2-3. O, N. fordi, holotype. P, N. calamus, holotype. Q, N. virgosa, paratype, EX1504-L4-9-4. R, N. macrocalyx, EX1606-7. S, N. hawaiiensis, EX1504-9-2. T, Paracalyotrophora hawaiiensis, EX1703-2-1. U, P. echinata, EX1703-10-1. V, P. spiralis, holotype. W, Plumarella circumoperculum, EX1603-2-3. X, Thouarella hilgendorfi, EX1605-L1-17." httpUri="https://zenodo.org/record/2614969/files/figure.png" pageId="24" pageNumber="25">Fig. 1M</figureCitation>
|
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) show the colony to have a short, stout stem about
|
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<quantity id="4CE1088CFFF6FFDC731D6AFE424CFF69" box="[1350,1406,225,251]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="25" unit="cm" value="2.0">2 cm</quantity>
|
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in height that bifurcates into two long branches, each about
|
||
<quantity id="4CE1088CFFF6FFDC75406B1A4452FE8D" box="[795,864,261,287]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="25" unit="cm" value="20.0">20 cm</quantity>
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long. The color of the colony is white. The whorls are relatively closely spaced (about 2 whorls/cm), each whorl having three polyps; the whorl diameter is about
|
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<quantity id="4CE1088CFFF6FFDC73226B3647F4FEF5" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="24" pageNumber="25" unit="mm" value="4.4">4.4 mm</quantity>
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. The horizontal length of a contracted polyp is about 4.0 mm. Polychaete commensalism was not noted.
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</paragraph>
|
||
<paragraph id="8BA6A569FFF6FFDC769C6B6E44A5FD39" blockId="24.[151,1437,152,1223]" pageId="24" pageNumber="25">
|
||
The basal body wall scales (
|
||
<figureCitation id="1322B9ECFFF6FFDC745E6B6E4558FE19" box="[517,618,369,395]" captionStart="FIGURE 11" captionStartId="23.[151,250,1810,1835]" captionTargetBox="[170,1410,191,1778]" captionTargetId="figure@23.[170,1410,191,1778]" captionTargetPageId="23" captionText="FIGURE 11. Narella merga, holotype: A–C, Stereo views of lateral and adaxial sides of a polyp. D, basal scales. E, outer surface of medial scales. F, buccal scales. G, adaxial buccal scales. H, opercular scales. I, coenenchymal scales. J, pinnular scale. Scale bars in mm." httpUri="https://zenodo.org/record/2614987/files/figure.png" pageId="24" pageNumber="25">Fig. 11D</figureCitation>
|
||
) are
|
||
<quantity id="4CE1088CFFF6FFDC74F86B6E4419FE19" box="[675,811,369,395]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="24" pageNumber="25" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.8–2.2 mm</quantity>
|
||
in height, their distal margin rounded (not spinose, serrate or undulate), forming a short cowl around the medial scales. The dorsolateral edge of the basal scale is rounded but on its lower half bears one or two short but tall (up to
|
||
<quantity id="4CE1088CFFF6FFDC756C6BA744A2FE41" box="[823,912,440,467]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="25" unit="mm" value="0.5">0.5 mm</quantity>
|
||
) thin ridges. The basal scales are open on the adaxial side. The medials (
|
||
<figureCitation id="1322B9ECFFF6FFDC779E6BC24515FE65" box="[453,551,477,503]" captionStart="FIGURE 11" captionStartId="23.[151,250,1810,1835]" captionTargetBox="[170,1410,191,1778]" captionTargetId="figure@23.[170,1410,191,1778]" captionTargetPageId="23" captionText="FIGURE 11. Narella merga, holotype: A–C, Stereo views of lateral and adaxial sides of a polyp. D, basal scales. E, outer surface of medial scales. F, buccal scales. G, adaxial buccal scales. H, opercular scales. I, coenenchymal scales. J, pinnular scale. Scale bars in mm." httpUri="https://zenodo.org/record/2614987/files/figure.png" pageId="24" pageNumber="25">Fig. 11E</figureCitation>
|
||
) are short (
|
||
<quantity id="4CE1088CFFF6FFDC74F26BC34404FE65" box="[681,822,476,503]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="24" pageNumber="25" unit="mm" value="1.4" valueMax="1.5" valueMin="1.3">1.3–1.5 mm</quantity>
|
||
), narrow, and upturned at their distal edge. The buccal scales (
|
||
<figureCitation id="1322B9ECFFF6FFDC76B1681E4678FD89" box="[234,330,513,539]" captionStart="FIGURE 11" captionStartId="23.[151,250,1810,1835]" captionTargetBox="[170,1410,191,1778]" captionTargetId="figure@23.[170,1410,191,1778]" captionTargetPageId="23" captionText="FIGURE 11. Narella merga, holotype: A–C, Stereo views of lateral and adaxial sides of a polyp. D, basal scales. E, outer surface of medial scales. F, buccal scales. G, adaxial buccal scales. H, opercular scales. I, coenenchymal scales. J, pinnular scale. Scale bars in mm." httpUri="https://zenodo.org/record/2614987/files/figure.png" pageId="24" pageNumber="25">Fig. 11F</figureCitation>
|
||
) range from
|
||
<quantity id="4CE1088CFFF6FFDC7786681E4557FD89" box="[477,613,513,539]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.2" metricValueMin="1.6" pageId="24" pageNumber="25" unit="mm" value="1.9" valueMax="2.2" valueMin="1.6">1.6–2.2 mm</quantity>
|
||
in length and are quite wide, also having a rounded distal edge. All body wall scales bear a low granulation and are not ridged except for the dorsolateral edge of the basals. The ratio of the lengths of the major body wall scales is: 1: 0.7: 0.95. There is one pair of flat adaxial buccal scales, shaped as rectangles or ellipses, measuring
|
||
<quantity id="4CE1088CFFF6FFDC778468724557FD15" box="[479,613,621,647]" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="24" pageNumber="25" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.6–0.7 mm</quantity>
|
||
in greater diameter. The adaxial polyp side is also covered with numerous smaller elliptical scales (
|
||
<figureCitation id="1322B9ECFFF6FFDC77F7688E4522FD39" box="[428,528,657,683]" captionStart="FIGURE 11" captionStartId="23.[151,250,1810,1835]" captionTargetBox="[170,1410,191,1778]" captionTargetId="figure@23.[170,1410,191,1778]" captionTargetPageId="23" captionText="FIGURE 11. Narella merga, holotype: A–C, Stereo views of lateral and adaxial sides of a polyp. D, basal scales. E, outer surface of medial scales. F, buccal scales. G, adaxial buccal scales. H, opercular scales. I, coenenchymal scales. J, pinnular scale. Scale bars in mm." httpUri="https://zenodo.org/record/2614987/files/figure.png" pageId="24" pageNumber="25">Fig. 11G</figureCitation>
|
||
) measuring
|
||
<quantity id="4CE1088CFFF6FFDC74C0688F441AFD39" box="[667,808,656,683]" metricMagnitude="-3" metricUnit="m" metricValue="7.62" metricValueMax="8.889999999999999" metricValueMin="6.35" pageId="24" pageNumber="25" unit="in" value="0.3" valueMax="0.35" valueMin="0.25">0.25–0.35 in</quantity>
|
||
diameter.
|
||
</paragraph>
|
||
<paragraph id="8BA6A569FFF6FFDC769C68AA4529FC11" blockId="24.[151,1437,152,1223]" pageId="24" pageNumber="25">
|
||
The abaxial opercular scales (
|
||
<figureCitation id="1322B9ECFFF6FFDC744568AA4540FD5D" box="[542,626,693,719]" captionStart="FIGURE 11" captionStartId="23.[151,250,1810,1835]" captionTargetBox="[170,1410,191,1778]" captionTargetId="figure@23.[170,1410,191,1778]" captionTargetPageId="23" captionText="FIGURE 11. Narella merga, holotype: A–C, Stereo views of lateral and adaxial sides of a polyp. D, basal scales. E, outer surface of medial scales. F, buccal scales. G, adaxial buccal scales. H, opercular scales. I, coenenchymal scales. J, pinnular scale. Scale bars in mm." httpUri="https://zenodo.org/record/2614987/files/figure.png" pageId="24" pageNumber="25">Fig. 11</figureCitation>
|
||
, top) are symmetrical, each bearing broad lateral lobes and measuring
|
||
<quantity id="4CE1088CFFF6FFDC76CC68C6462CFD61" box="[151,286,729,755]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="24" pageNumber="25" unit="mm" value="1.85" valueMax="1.9" valueMin="1.8">1.8–1.9 mm</quantity>
|
||
in length (L:W = 1.2). The lateral opercular scales are asymmetrical, having only one lateral lobe, and measure
|
||
<quantity id="4CE1088CFFF6FFDC76A568E246B5FC85" box="[254,391,765,791]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="24" pageNumber="25" unit="mm" value="1.85" valueMax="1.9" valueMin="1.8">1.8–1.9 mm</quantity>
|
||
in length (L:W = 1.5–2.0). The symmetrical (no lateral lobes) adaxial opercular scales are the smallest (
|
||
<figureCitation id="1322B9ECFFF6FFDC7775693E46A7FCA9" box="[302,405,801,827]" captionStart="FIGURE 11" captionStartId="23.[151,250,1810,1835]" captionTargetBox="[170,1410,191,1778]" captionTargetId="figure@23.[170,1410,191,1778]" captionTargetPageId="23" captionText="FIGURE 11. Narella merga, holotype: A–C, Stereo views of lateral and adaxial sides of a polyp. D, basal scales. E, outer surface of medial scales. F, buccal scales. G, adaxial buccal scales. H, opercular scales. I, coenenchymal scales. J, pinnular scale. Scale bars in mm." httpUri="https://zenodo.org/record/2614987/files/figure.png" pageId="24" pageNumber="25">Fig. 11H</figureCitation>
|
||
, lower), measuring
|
||
<quantity id="4CE1088CFFF6FFDC742C693E45CDFCA9" box="[631,767,801,827]" metricMagnitude="-3" metricUnit="m" metricValue="1.2999999999999998" metricValueMax="1.4" metricValueMin="1.2" pageId="24" pageNumber="25" unit="mm" value="1.2999999999999998" valueMax="1.4" valueMin="1.2">1.2–1.4 mm</quantity>
|
||
in length (L:W = 1.9–2.0). The adaxial operculars are flat, whereas the other operculars have highly concave outer surfaces. The pinnular scales are about
|
||
<quantity id="4CE1088CFFF6FFDC7293695A4219FCCD" box="[1224,1323,837,863]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="24" pageNumber="25" unit="mm" value="0.11">0.11 mm</quantity>
|
||
in length, flat, and slightly curved (
|
||
<figureCitation id="1322B9ECFFF6FFDC77EB6976453CFC11" box="[432,526,873,899]" captionStart="FIGURE 11" captionStartId="23.[151,250,1810,1835]" captionTargetBox="[170,1410,191,1778]" captionTargetId="figure@23.[170,1410,191,1778]" captionTargetPageId="23" captionText="FIGURE 11. Narella merga, holotype: A–C, Stereo views of lateral and adaxial sides of a polyp. D, basal scales. E, outer surface of medial scales. F, buccal scales. G, adaxial buccal scales. H, opercular scales. I, coenenchymal scales. J, pinnular scale. Scale bars in mm." httpUri="https://zenodo.org/record/2614987/files/figure.png" pageId="24" pageNumber="25">Fig. 11J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA6A569FFF6FFDC769C699247C2FC7D" blockId="24.[151,1437,152,1223]" pageId="24" pageNumber="25">
|
||
The coenenchymal scales (
|
||
<figureCitation id="1322B9ECFFF6FFDC77A869924579FC35" box="[499,587,909,935]" captionStart="FIGURE 11" captionStartId="23.[151,250,1810,1835]" captionTargetBox="[170,1410,191,1778]" captionTargetId="figure@23.[170,1410,191,1778]" captionTargetPageId="23" captionText="FIGURE 11. Narella merga, holotype: A–C, Stereo views of lateral and adaxial sides of a polyp. D, basal scales. E, outer surface of medial scales. F, buccal scales. G, adaxial buccal scales. H, opercular scales. I, coenenchymal scales. J, pinnular scale. Scale bars in mm." httpUri="https://zenodo.org/record/2614987/files/figure.png" pageId="24" pageNumber="25">
|
||
Fig.
|
||
<date id="FFA783A9FFF6FFDC747D69924579FC35" box="[550,587,909,935]" pageId="24" pageNumber="25" value="2011-01">11I</date>
|
||
</figureCitation>
|
||
) are a mixture of elongate (L:W = 3.1) to elliptical (L:W = 1.4) in shape and up to
|
||
<quantity id="4CE1088CFFF6FFDC768169AE4600FC59" box="[218,306,945,971]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="24" pageNumber="25" unit="mm" value="2.2">2.2 mm</quantity>
|
||
in length. They are thin and imbricate, most bearing a short and relatively low ridge on their outer surface.
|
||
</paragraph>
|
||
<paragraph id="8BA6A569FFF6FFDC769C69E74599FB31" blockId="24.[151,1437,152,1223]" pageId="24" pageNumber="25">
|
||
<emphasis id="B96D797BFFF6FFDC769C69E74654FB81" bold="true" box="[199,358,1016,1043]" pageId="24" pageNumber="25">Comparisons</emphasis>
|
||
. The single bifurcation growth mode of
|
||
<taxonomicName id="4C19DEEAFFF6FFDC757269E544A3FB81" authorityName="Cairns" authorityYear="2018" box="[809,913,1018,1043]" class="Anthozoa" family="Primnoidae" genus="Narella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="24" pageNumber="25" phylum="Cnidaria" rank="species" species="merga">
|
||
<emphasis id="B96D797BFFF6FFDC757269E544A3FB81" box="[809,913,1018,1043]" italics="true" pageId="24" pageNumber="25">N. merga</emphasis>
|
||
</taxonomicName>
|
||
appears to be unique within the genus. Among those species that have sparse dichotomous branching and a basal scale dorsolateral ridge,
|
||
<taxonomicName id="4C19DEEAFFF6FFDC72D16E0143C0FBA5" authorityName="Cairns" authorityYear="2018" box="[1162,1266,1054,1079]" class="Anthozoa" family="Primnoidae" genus="Narella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="24" pageNumber="25" phylum="Cnidaria" rank="species" species="merga">
|
||
<emphasis id="B96D797BFFF6FFDC72D16E0143C0FBA5" box="[1162,1266,1054,1079]" italics="true" pageId="24" pageNumber="25">N. merga</emphasis>
|
||
</taxonomicName>
|
||
is most similar to
|
||
<taxonomicName id="4C19DEEAFFF6FFDC76EF6E5D450EFBC9" authority="Cairns & Baco, 2007" authorityName="Cairns & Baco" authorityYear="2007" box="[180,572,1089,1115]" class="Anthozoa" family="Primnoidae" genus="Narella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="24" pageNumber="25" phylum="Cnidaria" rank="species" species="alaskensis">
|
||
<emphasis id="B96D797BFFF6FFDC76EF6E5D467AFBC9" box="[180,328,1089,1115]" italics="true" pageId="24" pageNumber="25">N. alaskensis</emphasis>
|
||
<bibRefCitation id="EF88D898FFF6FFDC77146E5E450EFBC9" author="Cairns, S. D. & Baco, A." box="[335,572,1089,1115]" pageId="24" pageNumber="25" pagination="391 - 407" refId="ref27693" refString="Cairns, S. D. & Baco, A. (2007) Review of five new Alaskan species of the deep-water octocoral Narella (Octocorallia: Primnoidae). Systematics and Biodiversity, 5 (4), 391 - 407. https: // doi. org / 10.1017 / S 1477200007002472" type="journal article" year="2007">Cairns & Baco, 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
(Gulf of
|
||
<collectingRegion id="49DD6B8BFFF6FFDC74FF6E5E45C7FBC9" box="[676,757,1089,1115]" country="United States of America" name="Alaska" pageId="24" pageNumber="25">Alaska</collectingRegion>
|
||
,
|
||
<quantity id="4CE1088CFFF6FFDC74A46E5F44AEFBC9" box="[767,924,1088,1115]" metricMagnitude="3" metricUnit="m" metricValue="2.726" metricValueMax="3.075" metricValueMin="2.377" pageId="24" pageNumber="25" unit="m" value="2726.0" valueMax="3075.0" valueMin="2377.0">2377–3075 m</quantity>
|
||
) and even shares an overlapping depth range. But,
|
||
<taxonomicName id="4C19DEEAFFF6FFDC76966E79466DFBED" authorityName="Cairns & Baco" authorityYear="2007" box="[205,351,1125,1151]" class="Anthozoa" family="Primnoidae" genus="Narella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="24" pageNumber="25" phylum="Cnidaria" rank="species" species="alaskensis">
|
||
<emphasis id="B96D797BFFF6FFDC76966E79466DFBED" box="[205,351,1125,1151]" italics="true" pageId="24" pageNumber="25">N. alaskensis</emphasis>
|
||
</taxonomicName>
|
||
differs in having more polyps per whorl (4–9), smaller polyps, and much longer medial body wall scales that sometimes bear a longitudinal ridge.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C303F6E2FFF6FFDC769C6EB345C3FB55" box="[199,753,1196,1223]" pageId="24" pageNumber="25" type="distribution">
|
||
<paragraph id="8BA6A569FFF6FFDC769C6EB345C3FB55" blockId="24.[151,1437,152,1223]" box="[199,753,1196,1223]" pageId="24" pageNumber="25">
|
||
<emphasis id="B96D797BFFF6FFDC769C6EB3466BFB55" bold="true" box="[199,345,1196,1223]" pageId="24" pageNumber="25">Distribution</emphasis>
|
||
. Known only from the
|
||
<typeStatus id="54A21BCBFFF6FFDC743B6EB245A2FB55" box="[608,656,1197,1223]" pageId="24" pageNumber="25">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |