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<document id="697E43882471AE2C6308D7BBA9236C96" ID-CLB-Dataset="29106" ID-DOI="10.11646/zootaxa.4455.2.2" ID-GBIF-Dataset="431a0907-2b69-4c52-86c6-6a0eb0c04d2e" ID-ISSN="1175-5326" ID-Zenodo-Dep="1457167" ID-ZooBank="595A9780-C604-465C-BDE2-589B87A4DB52" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539261406172" checkinUser="plazi" docAuthor="Antić, Dragan Ž., Dražina, Tvrtko, Rađa, Tonći, Lučić, Luka R. &amp; Makarov, Slobodan E." docDate="2018" docId="0383AF242005FFFD14DFFD59FC77FD71" docLanguage="en" docName="zootaxa.4455.2.2.pdf" docOrigin="Zootaxa 4455 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Typhloiulus gracilis Antić &amp; Dražina &amp; Rađa &amp; Lučić &amp; Makarov 2018, new species" docType="treatment" docVersion="8" lastPageNumber="277" masterDocId="FFBAD75C2008FFEE1448FFEDFFC3FFBC" masterDocTitle="Review of the genus Typhloiulus Latzel, 1884 in the Dinaric region, with a description of four new species and the first description of the male of Typhloiulus insularis Strasser, 1938 (Diplopoda: Julida: Julidae)" masterLastPageNumber="294" masterPageNumber="258" pageNumber="271" updateTime="1698575520044" updateUser="ExternalLinkService">
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<mods:title id="AFCD94ADAB43008918C1867C94F579AB">Review of the genus Typhloiulus Latzel, 1884 in the Dinaric region, with a description of four new species and the first description of the male of Typhloiulus insularis Strasser, 1938 (Diplopoda: Julida: Julidae)</mods:title>
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<mods:namePart id="2CCC0E84B338511F7D1FFFC0FD7A1AA3">Antić, Dragan Ž.</mods:namePart>
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<mods:namePart id="61E3C05C3B5FA5CBBDD675D69BC0E944">Dražina, Tvrtko</mods:namePart>
</mods:name>
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<mods:namePart id="DB3E8237B65945439889E515C7DB94D3">Rađa, Tonći</mods:namePart>
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<mods:namePart id="DACA3A8E537E9C20CBB534B14B9B56E8">Lučić, Luka R.</mods:namePart>
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<mods:namePart id="3CA65B76D256E3D297BF2400B3ADD783">Makarov, Slobodan E.</mods:namePart>
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<mods:title id="D88898BF9B112A908CF900BB35AE8219">Zootaxa</mods:title>
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<mods:date id="E82BA5715C3929D83BE28F9C37A86F26">2018</mods:date>
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<mods:number id="E028571A9C063C4726B3A640AB48CBA9">2018-08-02</mods:number>
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<mods:number id="1EE559262DFF91C11BBDCEAFFC27BE67">4455</mods:number>
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<subSubSection id="C3304DB92005FFE314DFFD59FE93FD53" pageId="13" pageNumber="271" type="nomenclature">
<paragraph id="8B951E322005FFE314DFFD59FDB1FD72" blockId="13.[151,626,692,751]" box="[151,626,692,718]" pageId="13" pageNumber="271">
<heading id="D0DDA95E2005FFE314DFFD59FDB1FD72" bold="true" box="[151,626,692,718]" fontSize="11" level="1" pageId="13" pageNumber="271" reason="1">
<emphasis id="B95EC2202005FFE314DFFD59FDB1FD72" bold="true" box="[151,626,692,718]" pageId="13" pageNumber="271">
<taxonomicName id="4C2A65B12005FFE314DFFD59FE16FD72" ID-CoL="84DYJ" authority="Antic" authorityName="Antić &amp; Dražina &amp; Rađa &amp; Lučić &amp; Makarov" authorityYear="2018" box="[151,469,692,718]" class="Diplopoda" family="Julidae" genus="Typhloiulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="13" pageNumber="271" phylum="Arthropoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="B95EC2202005FFE314DFFD59FE44FD72" bold="true" box="[151,391,692,718]" italics="true" pageId="13" pageNumber="271">Typhloiulus gracilis</emphasis>
Antić
</taxonomicName>
,
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</emphasis>
</heading>
</paragraph>
<paragraph id="8B951E322005FFE314DFFD3BFE93FD53" blockId="13.[151,626,692,751]" box="[151,336,726,751]" pageId="13" pageNumber="271">
<figureCitation id="131102B72005FFE314DFFD3BFF28FD53" box="[151,235,726,751]" captionStart="FIGURE 11" captionStartId="17.[151,250,1914,1936]" captionTargetBox="[203,1372,193,1879]" captionTargetId="figure@17.[203,1372,193,1879]" captionTargetPageId="17" captionText="FIGURE 11. Typhloiulus gracilis sp. nov. A. Paratype juvenile (IZB), habitus, lateral view. B. Holotype male (NHMSC), anterior part of the body, lateral view. C. Paratype male (IZB), body rings 7 and 8, lateral view. D. Holotype male (NHMSC), midbody rings, lateral view. E. Paratype female (NHMSC), telson, lateral view. Arrow indicates ventral margin of body ring 7. Scale lines: 1 mm." httpUri="https://zenodo.org/record/1457191/files/figure.png" pageId="13" pageNumber="271">Figs 11</figureCitation>
,
<figureCitation id="131102B72005FFE314BFFD3AFED6FD53" box="[247,277,727,751]" captionStart="FIGURE 12" captionStartId="18.[151,250,1941,1963]" captionTargetBox="[229,1316,245,1851]" captionTargetId="figure@18.[205,1383,193,1924]" captionTargetPageId="18" captionText="FIGURE 12. Typhloiulus gracilis sp. nov., paratype male (IZB). A. First pair of legs, anterior view. B. Promere, porsterior view. C. Penis, posterior view. D. Gonopods, mesal view. E. Gonopods, lateral view. Abbreviations: l: lamella; m: mesomere; o: opisthomere; p: promere; s: solenomere; t: telopodite; v: velum; ml: mesal lobe. Scale line: 0.3 mm." httpUri="https://zenodo.org/record/1457193/files/figure.png" pageId="13" pageNumber="271">12</figureCitation>
,
<figureCitation id="131102B72005FFE31569FD3BFE93FD53" box="[289,336,726,751]" captionStart="FIGURE 15" captionStartId="23.[151,250,1539,1561]" captionTargetBox="[199,1385,193,1520]" captionTargetId="figure@23.[199,1385,193,1520]" captionTargetPageId="23" captionText="FIGURE 15. Gonopods of representatives of the subgenus Inversotyphlus Strasser, 1962, mesal view. A. Typhloiulus clavatus sp. nov., holotype male (NHMSC). B. Typhloiulus gracilis sp. nov., paratype male (IZB). C. Typhloiulus opisthonodus sp. nov., holotype male (NHMSC). D. Typhloiulus lobifer Attems, 1951, male from Bitumen mine Minjera, island of Brač. E. Typhloiulus gellianae Makarov &amp; Rađa, 2006, holotype male (ZDR). Scale lines: 0.1 mm." httpUri="https://zenodo.org/record/1457199/files/figure.png" pageId="13" pageNumber="271">15B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3304DB92005FFE314DFFCF3FBBFFCC3" pageId="13" pageNumber="271" type="materials_examined">
<paragraph id="8B951E322005FFE314DFFCF3FBBFFCC3" blockId="13.[151,1437,798,2011]" pageId="13" pageNumber="271">
<materialsCitation id="3B42146F2005FFE314DFFCF3FC26FCE7" ID-GBIF-Occurrence="1917075749" collectingDate="2013-09-09" collectionCode="NHMSC" collectorName="T. Rada" country="Croatia" county="Jama na Boroviku Pit" latitude="43.135616" location="Material" longLatPrecision="1" longitude="16.6874" municipality="Pitve" pageId="13" pageNumber="271" specimenCount="2" typeStatus="holotype">
<emphasis id="B95EC2202005FFE314DFFCF3FEBFFC8B" bold="true" box="[151,380,798,823]" pageId="13" pageNumber="271">
<location id="8EF548E92005FFE314DFFCF3FF3CFC8B" LSID="urn:lsid:plazi:treatment:0383AF242005FFFD14DFFD59FC77FD71:8EF548E92005FFE314DFFCF3FF3CFC8B" box="[151,255,798,823]" country="Croatia" county="Jama na Boroviku Pit" latitude="43.135616" longLatPrecision="1" longitude="16.6874" municipality="Pitve" name="Material" pageId="13" pageNumber="271">Material</location>
examined
</emphasis>
:
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<typeStatus id="5491A0902005FFE315C7FCF2FE04FC8B" box="[399,455,799,823]" pageId="13" pageNumber="271">Type</typeStatus>
</location>
specimens.
<typeStatus id="5491A0902005FFE3161FFCF3FD7CFC8B" box="[599,703,798,823]" pageId="13" pageNumber="271" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="F33D5EA22005FFE31759FCF2FC4EFC8B" box="[785,909,799,823]" name="Croatia" pageId="13" pageNumber="271">CROATIA</collectingCountry>
,
<location id="8EF548E92005FFE317D4FCF3FB84FC8A" LSID="urn:lsid:plazi:treatment:0383AF242005FFFD14DFFD59FC77FD71:8EF548E92005FFE317D4FCF3FB84FC8A" box="[924,1095,798,823]" country="Croatia" county="Jama na Boroviku Pit" latitude="43.135616" longLatPrecision="1" longitude="16.6874" municipality="Pitve" name="Island of Hvar" pageId="13" pageNumber="271">Island of Hvar</location>
,
<collectingMunicipality id="6BF184482005FFE3101DFCF3FB52FC8B" box="[1109,1169,798,823]" pageId="13" pageNumber="271">Pitve</collectingMunicipality>
,
<collectingCounty id="62F466BE2005FFE310E8FCF2FA5FFC8B" box="[1184,1436,798,823]" pageId="13" pageNumber="271">Jama na Boroviku Pit</collectingCounty>
(
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,
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),
<collectingDate id="EFD0C11A2005FFE315BCFCAEFD04FCE7" box="[500,711,834,859]" pageId="13" pageNumber="271" value="2013-09-09">
<date id="FF9438F22005FFE315BCFCAEFD04FCE7" box="[500,711,834,859]" pageId="13" pageNumber="271" value="2013-09-09">9 September 2013</date>
</collectingDate>
,
<collectorName id="26DF7BE42005FFE3169EFCAEFCECFCE7" box="[726,815,834,859]" pageId="13" pageNumber="271">T. Rađa</collectorName>
leg. (
<collectionCode id="ED3B86F72005FFE3173BFCAEFC1EFCE7" box="[883,989,835,859]" pageId="13" pageNumber="271">NHMSC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B42146F2005FFE317BAFCAEFBBFFCC3" ID-GBIF-Occurrence="1917075785" collectingDate="2013-09-09" collectionCode="NHMSC" collectorName="T. Rada" country="Croatia" county="Jama na Boroviku Pit" latitude="43.135616" location="Material" longLatPrecision="1" longitude="16.6874" municipality="Pitve" pageId="13" pageNumber="271" specimenCount="8" specimenCount-female="1" specimenCount-juvenile="4" specimenCount-male="3" typeStatus="paratype">
<typeStatus id="5491A0902005FFE317BAFCAEFBA0FCE7" box="[1010,1123,835,859]" pageId="13" pageNumber="271" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D2CD5BB2005FFE31038FCAEFB0BFCE7" box="[1136,1224,834,859]" pageId="13" pageNumber="271" type="male">3 males</specimenCount>
(2 males—IZB, 1 male—NHMSC),
<specimenCount id="9D2CD5BB2005FFE3152CFC8AFE06FCC3" box="[356,453,870,895]" pageId="13" pageNumber="271" type="female">1 female</specimenCount>
(
<collectionCode id="ED3B86F72005FFE3159CFC8AFDFDFCC3" box="[468,574,871,895]" pageId="13" pageNumber="271">NHMSC</collectionCode>
),
<specimenCount id="9D2CD5BB2005FFE31618FC8AFD08FCC3" box="[592,715,870,895]" pageId="13" pageNumber="271" type="juvenile">4 juveniles</specimenCount>
(
<collectionCode id="ED3B86F72005FFE31691FC8AFC81FCC3" box="[729,834,871,895]" pageId="13" pageNumber="271">NHMSC</collectionCode>
), same data as for
<typeStatus id="5491A0902005FFE3105AFC8BFBBBFCC3" box="[1042,1144,870,895]" pageId="13" pageNumber="271" type="holotype">holotype</typeStatus>
.
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</paragraph>
</subSubSection>
<subSubSection id="C3304DB92005FFE3148FFC67FBA3FBC6" pageId="13" pageNumber="271" type="diagnosis">
<paragraph id="8B951E322005FFE3148FFC67FBA3FBC6" blockId="13.[151,1437,798,2011]" pageId="13" pageNumber="271">
<emphasis id="B95EC2202005FFE3148FFC67FE82FC1F" bold="true" box="[199,321,906,931]" pageId="13" pageNumber="271">Diagnosis.</emphasis>
<taxonomicName id="4C2A65B12005FFE31500FC67FDE4FC1F" authorityName="Antić &amp; Dražina &amp; Rađa &amp; Lučić &amp; Makarov" authorityYear="2018" box="[328,551,906,931]" class="Diplopoda" family="Julidae" genus="Typhloiulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="13" pageNumber="271" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B95EC2202005FFE31500FC67FDE4FC1F" box="[328,551,906,931]" italics="true" pageId="13" pageNumber="271">Typhloiulus gracilis</emphasis>
</taxonomicName>
is most similar to the members of the subgenus
<emphasis id="B95EC2202005FFE31003FC67FB34FC1F" box="[1099,1271,906,931]" italics="true" pageId="13" pageNumber="271">
<taxonomicName id="4C2A65B12005FFE31003FC67FB30FC1F" authorityName="Strasser" authorityYear="1962" box="[1099,1267,906,931]" class="Diplopoda" family="Julidae" genus="Inversotyphlus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="13" pageNumber="271" phylum="Arthropoda" rank="genus">Inversotyphlus</taxonomicName>
,
</emphasis>
viz.
<taxonomicName id="4C2A65B12005FFE31167FC61FA56FC1F" authorityName="Attems" authorityYear="1951" box="[1327,1429,906,931]" class="Diplopoda" family="Julidae" genus="Typhloiulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="13" pageNumber="271" phylum="Arthropoda" rank="species" species="lobifer">
<emphasis id="B95EC2202005FFE31167FC61FA56FC1F" box="[1327,1429,906,931]" italics="true" pageId="13" pageNumber="271">T. lobifer</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A65B12005FFE314DFFC42FED9FC7B" authorityName="Makarov &amp; Rada" authorityYear="2006" box="[151,282,942,967]" class="Diplopoda" family="Julidae" genus="Typhloiulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="13" pageNumber="271" phylum="Arthropoda" rank="species" species="gellianae">
<emphasis id="B95EC2202005FFE314DFFC42FED9FC7B" box="[151,282,942,967]" italics="true" pageId="13" pageNumber="271">T. gellianae</emphasis>
</taxonomicName>
, and herein described
<taxonomicName id="4C2A65B12005FFE3165DFC42FD4DFC7B" authorityName="Antić &amp; Dražina &amp; Rađa &amp; Lučić &amp; Makarov" authorityYear="2018" box="[533,654,942,967]" class="Diplopoda" family="Julidae" genus="Typhloiulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="13" pageNumber="271" phylum="Arthropoda" rank="species" species="clavatus">
<emphasis id="B95EC2202005FFE3165DFC42FD4DFC7B" box="[533,654,942,967]" italics="true" pageId="13" pageNumber="271">T. clavatus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C2A65B12005FFE31683FC42FCB8FC7B" authorityName="Antić &amp; Dražina &amp; Rađa &amp; Lučić &amp; Makarov" authorityYear="2018" box="[715,891,942,967]" class="Diplopoda" family="Julidae" genus="Typhloiulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="13" pageNumber="271" phylum="Arthropoda" rank="species" species="opisthonodus">
<emphasis id="B95EC2202005FFE31683FC42FCB8FC7B" box="[715,891,942,967]" italics="true" pageId="13" pageNumber="271">T. opisthonodus</emphasis>
</taxonomicName>
. These species are characterized by the presence of a very long opisthomere with a similarly shaped velum; the presence of a poorly developed, but visible opisthomeral lamella; and similar male sexual and habitus structures (see Discussion). The new species differs in the presence of very slender gonopods with a characteristic promere having a very slender apical half, curved mesad and slightly anteriad, with an almost lateral position in regard to mesomere.
<taxonomicName id="4C2A65B12005FFE31072FBD3FADAFBEB" authorityName="Antić &amp; Dražina &amp; Rađa &amp; Lučić &amp; Makarov" authorityYear="2018" box="[1082,1305,1086,1111]" class="Diplopoda" family="Julidae" genus="Typhloiulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="13" pageNumber="271" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B95EC2202005FFE31072FBD3FADAFBEB" box="[1082,1305,1086,1111]" italics="true" pageId="13" pageNumber="271">Typhloiulus gracilis</emphasis>
</taxonomicName>
also differs in the presence of a somewhat shorter and slightly curved downwards preanal process.
</paragraph>
</subSubSection>
<subSubSection id="C3304DB92005FFFD148FFB6BFF15FE73" lastPageId="19" lastPageNumber="277" pageId="13" pageNumber="271" type="description">
<paragraph id="8B951E322005FFE3148FFB6BFDA4F867" blockId="13.[151,1437,798,2011]" pageId="13" pageNumber="271">
<emphasis id="B95EC2202005FFE3148FFB6BFE9AFB23" bold="true" box="[199,345,1158,1183]" pageId="13" pageNumber="271">Description.</emphasis>
Based on
<typeStatus id="5491A0902005FFE31586FB6BFDF3FB23" box="[462,560,1158,1183]" pageId="13" pageNumber="271" type="holotype">holotype</typeStatus>
(largest male)
<quantity id="4CD2B3D72005FFE31691FB6AFCEBFB22" box="[729,808,1159,1183]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" pageId="13" pageNumber="271" unit="mm" value="47.0">47 mm</quantity>
long, vertical diameter of the largest body ring
<quantity id="4CD2B3D72005FFE31177FB6BFA54FB22" box="[1343,1431,1158,1183]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="13" pageNumber="271" unit="mm" value="2.6">2.6 mm</quantity>
, body with 57 podous rings + 0 apodous rings + telson.
<emphasis id="B95EC2202005FFE3175BFB47FC60FB7F" bold="true" box="[787,931,1194,1219]" pageId="13" pageNumber="271">Colouration</emphasis>
: (
<figureCitation id="131102B72005FFE317F4FB47FBE6FB7F" box="[956,1061,1194,1219]" captionStart="FIGURE 11" captionStartId="17.[151,250,1914,1936]" captionTargetBox="[203,1372,193,1879]" captionTargetId="figure@17.[203,1372,193,1879]" captionTargetPageId="17" captionText="FIGURE 11. Typhloiulus gracilis sp. nov. A. Paratype juvenile (IZB), habitus, lateral view. B. Holotype male (NHMSC), anterior part of the body, lateral view. C. Paratype male (IZB), body rings 7 and 8, lateral view. D. Holotype male (NHMSC), midbody rings, lateral view. E. Paratype female (NHMSC), telson, lateral view. Arrow indicates ventral margin of body ring 7. Scale lines: 1 mm." httpUri="https://zenodo.org/record/1457191/files/figure.png" pageId="13" pageNumber="271">Fig. 11A</figureCitation>
). Depigmented, yellowish white.
<emphasis id="B95EC2202005FFE314DFFB23FF14FB5B" bold="true" box="[151,215,1230,1255]" pageId="13" pageNumber="271">Head</emphasis>
: (
<figureCitation id="131102B72005FFE314A7FB23FE95FB5B" box="[239,342,1230,1255]" captionStart="FIGURE 11" captionStartId="17.[151,250,1914,1936]" captionTargetBox="[203,1372,193,1879]" captionTargetId="figure@17.[203,1372,193,1879]" captionTargetPageId="17" captionText="FIGURE 11. Typhloiulus gracilis sp. nov. A. Paratype juvenile (IZB), habitus, lateral view. B. Holotype male (NHMSC), anterior part of the body, lateral view. C. Paratype male (IZB), body rings 7 and 8, lateral view. D. Holotype male (NHMSC), midbody rings, lateral view. E. Paratype female (NHMSC), telson, lateral view. Arrow indicates ventral margin of body ring 7. Scale lines: 1 mm." httpUri="https://zenodo.org/record/1457191/files/figure.png" pageId="13" pageNumber="271">Fig. 11B</figureCitation>
). Labrum with three labral teeth, with 4 supralabral and 8+7 labral setae. Gnathochilarium with promentum in the form of an elongated kite; lamellae linguales with 4+4 setae in one row, stipites each with 3 setae. Antennae
<quantity id="4CD2B3D72005FFE31514FAFBFE04FA92" box="[348,455,1302,1327]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" pageId="13" pageNumber="271" unit="mm" value="3.55">3.55 mm</quantity>
long, their length 136% of vertical diameter of the largest body ring. Length of antennomeres IVIII (in mm): 0.2 (I), 0.67 (II), 0.62 (III), 0.62 (IV), 0.69 (V), 0.52 (VI), 0.15 (VII), and 0.08 (VIII). Length/width ratio of antennomeres IVII: 0.8 (I), 2.6 (II), 2.4 (III), 2.6 (IV), 2.4 (V), 1.8 (VI), and 0.75 (VII). Antennomeres V and VI each with a terminal corolla of large bacilliform sensilla; antennomere VII with a terminal corolla of microsetae.
<emphasis id="B95EC2202005FFE3164BFA4BFD4AFA03" bold="true" box="[515,649,1446,1471]" pageId="13" pageNumber="271">Body rings</emphasis>
: (
<figureCitation id="131102B72005FFE316EBFA4BFCCEFA03" box="[675,781,1446,1471]" captionStart="FIGURE 11" captionStartId="17.[151,250,1914,1936]" captionTargetBox="[203,1372,193,1879]" captionTargetId="figure@17.[203,1372,193,1879]" captionTargetPageId="17" captionText="FIGURE 11. Typhloiulus gracilis sp. nov. A. Paratype juvenile (IZB), habitus, lateral view. B. Holotype male (NHMSC), anterior part of the body, lateral view. C. Paratype male (IZB), body rings 7 and 8, lateral view. D. Holotype male (NHMSC), midbody rings, lateral view. E. Paratype female (NHMSC), telson, lateral view. Arrow indicates ventral margin of body ring 7. Scale lines: 1 mm." httpUri="https://zenodo.org/record/1457191/files/figure.png" pageId="13" pageNumber="271">Fig. 11D</figureCitation>
). The whole metazonal area with longitudinal striations. Midbody rings with
<emphasis id="B95EC2202005FFE315C8FA21FE62FA5F" box="[384,417,1484,1507]" italics="true" pageId="13" pageNumber="271">ca.</emphasis>
20 short metazonal setae. Length of midbody setae
<emphasis id="B95EC2202005FFE317BBFA21FBD7FA5F" box="[1011,1044,1484,1507]" italics="true" pageId="13" pageNumber="271">ca.</emphasis>
10% of vertical diameter of rings.
<emphasis id="B95EC2202005FFE314DFFA03FF22F9BB" bold="true" box="[151,225,1518,1543]" pageId="13" pageNumber="271">Telson</emphasis>
: (
<figureCitation id="131102B72005FFE314B7FA03FEA4F9BB" box="[255,359,1518,1543]" captionStart="FIGURE 11" captionStartId="17.[151,250,1914,1936]" captionTargetBox="[203,1372,193,1879]" captionTargetId="figure@17.[203,1372,193,1879]" captionTargetPageId="17" captionText="FIGURE 11. Typhloiulus gracilis sp. nov. A. Paratype juvenile (IZB), habitus, lateral view. B. Holotype male (NHMSC), anterior part of the body, lateral view. C. Paratype male (IZB), body rings 7 and 8, lateral view. D. Holotype male (NHMSC), midbody rings, lateral view. E. Paratype female (NHMSC), telson, lateral view. Arrow indicates ventral margin of body ring 7. Scale lines: 1 mm." httpUri="https://zenodo.org/record/1457191/files/figure.png" pageId="13" pageNumber="271">Fig. 11E</figureCitation>
). Epiproct with elongated preanal process, slightly curved downwards and covered by long dorsal and lateral setae. Paraprocts rounded, with numerous long setae. Hypoproct lenticular, with 10 setae.
<emphasis id="B95EC2202005FFE31114F9FFFA57F997" bold="true" box="[1372,1428,1554,1579]" pageId="13" pageNumber="271">Legs</emphasis>
: First pair of legs modified, hook-shaped (
<figureCitation id="131102B72005FFE31623F9DBFD17F9F3" box="[619,724,1590,1615]" captionStart="FIGURE 12" captionStartId="18.[151,250,1941,1963]" captionTargetBox="[229,1316,245,1851]" captionTargetId="figure@18.[205,1383,193,1924]" captionTargetPageId="18" captionText="FIGURE 12. Typhloiulus gracilis sp. nov., paratype male (IZB). A. First pair of legs, anterior view. B. Promere, porsterior view. C. Penis, posterior view. D. Gonopods, mesal view. E. Gonopods, lateral view. Abbreviations: l: lamella; m: mesomere; o: opisthomere; p: promere; s: solenomere; t: telopodite; v: velum; ml: mesal lobe. Scale line: 0.3 mm." httpUri="https://zenodo.org/record/1457193/files/figure.png" pageId="13" pageNumber="271">Fig. 12A</figureCitation>
); coxa, postfemur, and tibiotarsus each with one seta, prefemur with four or five setae, femur each with two setae. Tibiotarsus with a small dorsal lobe (tarsal remnant) and tuberculate on the tip. Second pair of legs with poorly developed ventral pads on the tibia.
<emphasis id="B95EC2202005FFE31082F993FAC9F92B" bold="true" box="[1226,1290,1662,1687]" pageId="13" pageNumber="271">Penis</emphasis>
: (
<figureCitation id="131102B72005FFE3116DF993FA4EF92B" box="[1317,1421,1662,1687]" captionStart="FIGURE 12" captionStartId="18.[151,250,1941,1963]" captionTargetBox="[229,1316,245,1851]" captionTargetId="figure@18.[205,1383,193,1924]" captionTargetPageId="18" captionText="FIGURE 12. Typhloiulus gracilis sp. nov., paratype male (IZB). A. First pair of legs, anterior view. B. Promere, porsterior view. C. Penis, posterior view. D. Gonopods, mesal view. E. Gonopods, lateral view. Abbreviations: l: lamella; m: mesomere; o: opisthomere; p: promere; s: solenomere; t: telopodite; v: velum; ml: mesal lobe. Scale line: 0.3 mm." httpUri="https://zenodo.org/record/1457193/files/figure.png" pageId="13" pageNumber="271">Fig. 12C</figureCitation>
). Bilobed. Lobes prominent, elongated.
<emphasis id="B95EC2202005FFE3161BF94FFC08F907" bold="true" box="[595,971,1698,1723]" pageId="13" pageNumber="271">Ventral margin of body ring 7</emphasis>
: (
<figureCitation id="131102B72005FFE317ADF94FFB88F907" box="[997,1099,1698,1723]" captionStart="FIGURE 11" captionStartId="17.[151,250,1914,1936]" captionTargetBox="[203,1372,193,1879]" captionTargetId="figure@17.[203,1372,193,1879]" captionTargetPageId="17" captionText="FIGURE 11. Typhloiulus gracilis sp. nov. A. Paratype juvenile (IZB), habitus, lateral view. B. Holotype male (NHMSC), anterior part of the body, lateral view. C. Paratype male (IZB), body rings 7 and 8, lateral view. D. Holotype male (NHMSC), midbody rings, lateral view. E. Paratype female (NHMSC), telson, lateral view. Arrow indicates ventral margin of body ring 7. Scale lines: 1 mm." httpUri="https://zenodo.org/record/1457191/files/figure.png" pageId="13" pageNumber="271">Fig. 11C</figureCitation>
). Well-developed, elongated, with rounded tip; almost right-angled to metazona.
<emphasis id="B95EC2202005FFE3169DF92BFC8DF963" bold="true" box="[725,846,1734,1759]" pageId="13" pageNumber="271">Gonopods</emphasis>
: (
<figureCitation id="131102B72005FFE3172DF92BFBCCF962" box="[869,1039,1734,1759]" captionStart="FIGURE 12" captionStartId="18.[151,250,1941,1963]" captionTargetBox="[229,1316,245,1851]" captionTargetId="figure@18.[205,1383,193,1924]" captionTargetPageId="18" captionText="FIGURE 12. Typhloiulus gracilis sp. nov., paratype male (IZB). A. First pair of legs, anterior view. B. Promere, porsterior view. C. Penis, posterior view. D. Gonopods, mesal view. E. Gonopods, lateral view. Abbreviations: l: lamella; m: mesomere; o: opisthomere; p: promere; s: solenomere; t: telopodite; v: velum; ml: mesal lobe. Scale line: 0.3 mm." httpUri="https://zenodo.org/record/1457193/files/figure.png" pageId="13" pageNumber="271">Figs 12B, D, E</figureCitation>
,
<figureCitation id="131102B72005FFE31053F92BFB8AF963" box="[1051,1097,1734,1759]" captionStart="FIGURE 15" captionStartId="23.[151,250,1539,1561]" captionTargetBox="[199,1385,193,1520]" captionTargetId="figure@23.[199,1385,193,1520]" captionTargetPageId="23" captionText="FIGURE 15. Gonopods of representatives of the subgenus Inversotyphlus Strasser, 1962, mesal view. A. Typhloiulus clavatus sp. nov., holotype male (NHMSC). B. Typhloiulus gracilis sp. nov., paratype male (IZB). C. Typhloiulus opisthonodus sp. nov., holotype male (NHMSC). D. Typhloiulus lobifer Attems, 1951, male from Bitumen mine Minjera, island of Brač. E. Typhloiulus gellianae Makarov &amp; Rađa, 2006, holotype male (ZDR). Scale lines: 0.1 mm." httpUri="https://zenodo.org/record/1457199/files/figure.png" pageId="13" pageNumber="271">15B</figureCitation>
). Very long and thin. Promere (
<emphasis id="B95EC2202005FFE314E8F907FF6CF8BF" bold="true" box="[160,175,1770,1795]" pageId="13" pageNumber="271">p</emphasis>
) and mesomere (
<emphasis id="B95EC2202005FFE3153CF907FE48F8BF" bold="true" box="[372,395,1770,1795]" pageId="13" pageNumber="271">m</emphasis>
) shorter than opisthomere (
<emphasis id="B95EC2202005FFE3168DF907FD10F8BF" bold="true" box="[709,723,1770,1795]" pageId="13" pageNumber="271">o</emphasis>
). Promere (
<emphasis id="B95EC2202005FFE31712F907FCAAF8BF" bold="true" box="[858,873,1770,1795]" pageId="13" pageNumber="271">p</emphasis>
) longer than
<emphasis id="B95EC2202005FFE3104AF907FBDAF8BF" bold="true" box="[1026,1049,1770,1795]" pageId="13" pageNumber="271">m</emphasis>
; wide at the base; apical half very slender, curved mesad and slightly anteriad. Mesal lobe (
<emphasis id="B95EC2202005FFE3175EF8E3FCF6F89B" bold="true" box="[790,821,1806,1831]" pageId="13" pageNumber="271">ml</emphasis>
) high and narrow, with three setae, one apical and two subapical. The telopodite (
<emphasis id="B95EC2202005FFE31599F8DFFE19F8F7" bold="true" box="[465,474,1842,1867]" pageId="13" pageNumber="271">t</emphasis>
) well-developed, tapering distad; apex almost at the same height as apex of
<emphasis id="B95EC2202005FFE3113EF8DFFA56F8F7" bold="true" box="[1398,1429,1842,1867]" pageId="13" pageNumber="271">ml</emphasis>
. Mesomere (
<emphasis id="B95EC2202005FFE31555F8BBFEF7F8D3" bold="true" box="[285,308,1878,1903]" pageId="13" pageNumber="271">m</emphasis>
) thin over all of its height, in the form of a rod, with antero-apical tuberculation and a small posterior node. Opisthomere (
<emphasis id="B95EC2202005FFE31536F897FE4FF82F" bold="true" box="[382,396,1914,1939]" pageId="13" pageNumber="271">o</emphasis>
) very slender, considerably longer than
<emphasis id="B95EC2202005FFE3171DF897FCA7F82F" bold="true" box="[853,868,1914,1939]" pageId="13" pageNumber="271">p</emphasis>
and
<emphasis id="B95EC2202005FFE317D6F897FC76F82F" bold="true" box="[926,949,1914,1939]" pageId="13" pageNumber="271">m</emphasis>
. With poorly developed lamella (
<emphasis id="B95EC2202005FFE3117BF897FAF8F82F" bold="true" box="[1331,1339,1914,1939]" pageId="13" pageNumber="271">l</emphasis>
) anteromesally; with one spine mesally; velum (
<emphasis id="B95EC2202005FFE316CCF873FD51F80B" bold="true" box="[644,658,1950,1975]" pageId="13" pageNumber="271">v</emphasis>
) unipartite, tapering anteriad, with fimbriated tip. Solenomere (
<emphasis id="B95EC2202005FFE311C0F873FA50F80B" bold="true" box="[1416,1427,1950,1975]" pageId="13" pageNumber="271">s</emphasis>
) narrow, distally with several short spines.
</paragraph>
<caption id="DF554EBA2006FFE014DFF865FBB9F867" httpUri="https://zenodo.org/record/1457185/files/figure.png" pageId="14" pageNumber="272" startId="14.[151,250,1928,1950]" targetBox="[208,1376,193,1911]" targetPageId="14">
<paragraph id="8B951E322006FFE014DFF865FBB9F867" blockId="14.[151,1437,1928,2011]" pageId="14" pageNumber="272">
<emphasis id="B95EC2202006FFE014DFF865FED1F821" bold="true" box="[151,274,1928,1950]" pageId="14" pageNumber="272">FIGURE 8.</emphasis>
<taxonomicName id="4C2A65B12006FFE01551F865FD50F822" authority="(Latzel, 1884)" authorityName="Latzel" authorityYear="1884" baseAuthorityName="Latzel" baseAuthorityYear="1884" box="[281,659,1928,1950]" class="Diplopoda" family="Julidae" genus="Typhloiulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="14" pageNumber="272" phylum="Arthropoda" rank="species" species="psilonotus">
<emphasis id="B95EC2202006FFE01551F865FE3EF822" box="[281,509,1928,1950]" italics="true" pageId="14" pageNumber="272">Typhloiulus psilonotus</emphasis>
(Latzel, 1884)
</taxonomicName>
specimens from Danilovgrad, Montenegro.
<emphasis id="B95EC2202006FFE0101CF865FBAEF821" bold="true" box="[1108,1133,1928,1949]" pageId="14" pageNumber="272">A.</emphasis>
Female, habitus, lateral view.
<emphasis id="B95EC2202006FFE014DFF84AFF6DF800" bold="true" box="[151,174,1959,1980]" pageId="14" pageNumber="272">B.</emphasis>
Female, anterior part of the body, lateral view.
<emphasis id="B95EC2202006FFE016D4F84BFD77F800" bold="true" box="[668,692,1958,1980]" pageId="14" pageNumber="272">C.</emphasis>
Female, telson, lateral view.
<emphasis id="B95EC2202006FFE017ACF84AFC3EF800" bold="true" box="[996,1021,1959,1980]" pageId="14" pageNumber="272">D.</emphasis>
Female, midbody rings, lateral view.
<emphasis id="B95EC2202006FFE011CDF84AFA5FF800" bold="true" box="[1413,1436,1959,1980]" pageId="14" pageNumber="272">E.</emphasis>
Male, body ring 7, lateral view. Arrow indicates ventral margin of body ring 7.
<emphasis id="B95EC2202006FFE017F2F829FBF0F867" bold="true" box="[954,1075,1988,2011]" pageId="14" pageNumber="272">Scale lines:</emphasis>
1 mm.
</paragraph>
</caption>
<caption id="DF554EBA2007FFE114DFF86CFAE7F868" httpUri="https://zenodo.org/record/1457187/files/figure.png" pageId="15" pageNumber="273" startId="15.[151,250,1921,1943]" targetBox="[151,1436,193,1899]" targetPageId="15">
<paragraph id="8B951E322007FFE114DFF86CFAE7F868" blockId="15.[151,1436,1921,2004]" pageId="15" pageNumber="273">
<emphasis id="B95EC2202007FFE114DFF86CFED7F82A" bold="true" box="[151,276,1921,1943]" pageId="15" pageNumber="273">FIGURE 9.</emphasis>
<taxonomicName id="4C2A65B12007FFE11554F86CFE2DF82B" authority="Antić &amp; Dražina &amp; Rađa &amp; Lučić &amp; Makarov, 2018" authorityName="Antić &amp; Dražina &amp; Rađa &amp; Lučić &amp; Makarov" authorityYear="2018" box="[284,494,1921,1943]" class="Diplopoda" family="Julidae" genus="Typhloiulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="15" pageNumber="273" phylum="Arthropoda" rank="species" species="clavatus" status="sp. nov.">
<emphasis id="B95EC2202007FFE11554F86CFE2DF82B" box="[284,494,1921,1943]" italics="true" pageId="15" pageNumber="273">Typhloiulus clavatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D7F5B2007FFE115BEF86CFD8BF82A" box="[502,584,1921,1942]" pageId="15" pageNumber="273" rank="species">
<emphasis id="B95EC2202007FFE115BEF86CFD8BF82A" bold="true" box="[502,584,1921,1942]" pageId="15" pageNumber="273">sp. nov.</emphasis>
</taxonomicNameLabel>
(NHMSC).
<emphasis id="B95EC2202007FFE11682F86CFD21F82A" bold="true" box="[714,738,1921,1942]" pageId="15" pageNumber="273">A.</emphasis>
Paratype female, habitus, lateral view.
<emphasis id="B95EC2202007FFE11032F86CFB52F82A" bold="true" box="[1146,1169,1921,1942]" pageId="15" pageNumber="273">B.</emphasis>
Paratype female, anterior part of the body, lateral view.
<emphasis id="B95EC2202007FFE1159DF872FE2DF809" bold="true" box="[469,494,1951,1973]" pageId="15" pageNumber="273">C.</emphasis>
Paratype female, telson, lateral view.
<emphasis id="B95EC2202007FFE117C8F84DFC5AF809" bold="true" box="[896,921,1952,1973]" pageId="15" pageNumber="273">D.</emphasis>
Paratype female, midbody rings, lateral view.
<emphasis id="B95EC2202007FFE111CCF84DFA58F809" bold="true" box="[1412,1435,1952,1973]" pageId="15" pageNumber="273">E.</emphasis>
Holotype male, body rings 6 and 7, lateral view. Arrow indicates ventral margin of body ring 7.
<emphasis id="B95EC2202007FFE1102CF850FB1FF868" bold="true" box="[1124,1244,1981,2004]" pageId="15" pageNumber="273">Scale lines:</emphasis>
1 mm.
</paragraph>
</caption>
<caption id="DF554EBA2018FFFE14DFF88BFE5BF864" httpUri="https://zenodo.org/record/1457189/files/figure.png" pageId="16" pageNumber="274" startId="16.[151,250,1894,1916]" targetBox="[164,1383,238,1796]" targetPageId="16">
<paragraph id="8B951E322018FFFE14DFF88BFE5BF864" blockId="16.[151,1436,1894,2008]" pageId="16" pageNumber="274">
<emphasis id="B95EC2202018FFFE14DFF88BFEE3F8C7" bold="true" box="[151,288,1894,1916]" pageId="16" pageNumber="274">FIGURE 10.</emphasis>
<taxonomicName id="4C2A65B12018FFFE1560F88BFE3AF8C0" authority="Antić &amp; Dražina &amp; Rađa &amp; Lučić &amp; Makarov, 2018" authorityName="Antić &amp; Dražina &amp; Rađa &amp; Lučić &amp; Makarov" authorityYear="2018" box="[296,505,1894,1916]" class="Diplopoda" family="Julidae" genus="Typhloiulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="16" pageNumber="274" phylum="Arthropoda" rank="species" species="clavatus" status="sp. nov.">
<emphasis id="B95EC2202018FFFE1560F88BFE3AF8C0" box="[296,505,1894,1916]" italics="true" pageId="16" pageNumber="274">Typhloiulus clavatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D7F5B2018FFFE1649F88BFD90F8C7" box="[513,595,1894,1915]" pageId="16" pageNumber="274" rank="species">
<emphasis id="B95EC2202018FFFE1649F88BFD90F8C7" bold="true" box="[513,595,1894,1915]" pageId="16" pageNumber="274">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype male (NHMSC).
<emphasis id="B95EC2202018FFFE173BF88BFC48F8C7" bold="true" box="[883,907,1894,1915]" pageId="16" pageNumber="274">A.</emphasis>
First pair of legs, anterior view.
<emphasis id="B95EC2202018FFFE1096F88BFB35F8C7" bold="true" box="[1246,1270,1894,1915]" pageId="16" pageNumber="274">B.</emphasis>
Penis, posterior view.
<emphasis id="B95EC2202018FFFE149CF868FF2EF827" bold="true" box="[212,237,1925,1947]" pageId="16" pageNumber="274">C.</emphasis>
Promere, posterior view.
<emphasis id="B95EC2202018FFFE15BCF868FDCEF826" bold="true" box="[500,525,1925,1946]" pageId="16" pageNumber="274">D.</emphasis>
Gonopods, mesal view.
<emphasis id="B95EC2202018FFFE1740F868FCDCF826" bold="true" box="[776,799,1925,1946]" pageId="16" pageNumber="274">E.</emphasis>
Mesomere and opisthomere, mesal view.
<emphasis id="B95EC2202018FFFE1085F868FB1DF826" bold="true" box="[1229,1246,1925,1946]" pageId="16" pageNumber="274">F.</emphasis>
Gonopods, lateral view.
<emphasis id="B95EC2202018FFFE149BF84EFE49F804" bold="true" box="[211,394,1955,1977]" pageId="16" pageNumber="274">Abbreviations: l:</emphasis>
lamella;
<emphasis id="B95EC2202018FFFE15A1F84EFDC5F804" bold="true" box="[489,518,1955,1976]" pageId="16" pageNumber="274">m:</emphasis>
mesomere;
<emphasis id="B95EC2202018FFFE16CBF84EFD54F804" bold="true" box="[643,663,1955,1976]" pageId="16" pageNumber="274">o:</emphasis>
opisthomere;
<emphasis id="B95EC2202018FFFE1761F84EFC83F804" bold="true" box="[809,832,1955,1976]" pageId="16" pageNumber="274">p:</emphasis>
promere;
<emphasis id="B95EC2202018FFFE17E0F84EFC79F804" bold="true" box="[936,954,1955,1976]" pageId="16" pageNumber="274">s:</emphasis>
solenomere;
<emphasis id="B95EC2202018FFFE100BF84EFB97F804" bold="true" box="[1091,1108,1955,1976]" pageId="16" pageNumber="274">t:</emphasis>
telopodite;
<emphasis id="B95EC2202018FFFE1085F84EFB20F804" bold="true" box="[1229,1251,1955,1976]" pageId="16" pageNumber="274">v:</emphasis>
velum;
<emphasis id="B95EC2202018FFFE117EF84EFA98F804" bold="true" box="[1334,1371,1955,1976]" pageId="16" pageNumber="274">ml:</emphasis>
mesal lobe.
<emphasis id="B95EC2202018FFFE1487F82CFEFDF864" bold="true" box="[207,318,1985,2008]" pageId="16" pageNumber="274">Scale line:</emphasis>
0.3 mm.
</paragraph>
</caption>
<caption id="DF554EBA2019FFFF14DFF897FE94F850" httpUri="https://zenodo.org/record/1457191/files/figure.png" pageId="17" pageNumber="275" startId="17.[151,250,1914,1936]" targetBox="[203,1372,193,1879]" targetPageId="17">
<paragraph id="8B951E322019FFFF14DFF897FE94F850" blockId="17.[151,1436,1914,2028]" pageId="17" pageNumber="275">
<emphasis id="B95EC2202019FFFF14DFF897FEE2F833" bold="true" box="[151,289,1914,1936]" pageId="17" pageNumber="275">FIGURE 11.</emphasis>
<taxonomicName id="4C2A65B12019FFFF1562F897FE35F82C" authority="Antić &amp; Dražina &amp; Rađa &amp; Lučić &amp; Makarov, 2018" authorityName="Antić &amp; Dražina &amp; Rađa &amp; Lučić &amp; Makarov" authorityYear="2018" box="[298,502,1914,1936]" class="Diplopoda" family="Julidae" genus="Typhloiulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="17" pageNumber="275" phylum="Arthropoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="B95EC2202019FFFF1562F897FE35F82C" box="[298,502,1914,1936]" italics="true" pageId="17" pageNumber="275">Typhloiulus gracilis</emphasis>
</taxonomicName>
<emphasis id="B95EC2202019FFFF1648F896FDB5F833" bold="true" box="[512,630,1914,1936]" pageId="17" pageNumber="275">
<taxonomicNameLabel id="A26D7F5B2019FFFF1648F896FD97F82C" box="[512,596,1915,1936]" pageId="17" pageNumber="275" rank="species">sp. nov.</taxonomicNameLabel>
A.
</emphasis>
Paratype juvenile (IZB), habitus, lateral view.
<emphasis id="B95EC2202019FFFF1020F896FBBCF82C" bold="true" box="[1128,1151,1915,1936]" pageId="17" pageNumber="275">B.</emphasis>
Holotype male (NHMSC), anterior part of the body, lateral view.
<emphasis id="B95EC2202019FFFF1657F874FDF4F813" bold="true" box="[543,567,1945,1967]" pageId="17" pageNumber="275">C.</emphasis>
Paratype male (IZB), body rings 7 and 8, lateral view.
<emphasis id="B95EC2202019FFFF1025F874FB45F812" bold="true" box="[1133,1158,1945,1966]" pageId="17" pageNumber="275">D.</emphasis>
Holotype male (NHMSC), midbody rings, lateral view.
<emphasis id="B95EC2202019FFFF15F0F855FE13F871" bold="true" box="[440,464,1976,1997]" pageId="17" pageNumber="275">E.</emphasis>
Paratype female (NHMSC), telson, lateral view. Arrow indicates ventral margin of body ring 7.
<emphasis id="B95EC2202019FFFF14DFF838FECCF850" bold="true" box="[151,271,2005,2028]" pageId="17" pageNumber="275">Scale lines:</emphasis>
1 mm.
</paragraph>
</caption>
<caption id="DF554EBA201AFFFC14DFF878FB75F854" httpUri="https://zenodo.org/record/1457193/files/figure.png" pageId="18" pageNumber="276" startId="18.[151,250,1941,1963]" targetBox="[229,1316,245,1851]" targetPageId="18">
<paragraph id="8B951E32201AFFFC14DFF878FB75F854" blockId="18.[151,1436,1941,2024]" pageId="18" pageNumber="276">
<emphasis id="B95EC220201AFFFC14DFF878FEE1F816" bold="true" box="[151,290,1941,1963]" pageId="18" pageNumber="276">FIGURE 12.</emphasis>
<taxonomicName id="4C2A65B1201AFFFC1562F878FE36F817" authority="Antić &amp; Dražina &amp; Rađa &amp; Lučić &amp; Makarov, 2018" authorityName="Antić &amp; Dražina &amp; Rađa &amp; Lučić &amp; Makarov" authorityYear="2018" box="[298,501,1941,1963]" class="Diplopoda" family="Julidae" genus="Typhloiulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="18" pageNumber="276" phylum="Arthropoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="B95EC220201AFFFC1562F878FE36F817" box="[298,501,1941,1963]" italics="true" pageId="18" pageNumber="276">Typhloiulus gracilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D7F5B201AFFFC15B6F878FD92F816" box="[510,593,1941,1962]" pageId="18" pageNumber="276" rank="species">
<emphasis id="B95EC220201AFFFC15B6F878FD92F816" bold="true" box="[510,593,1941,1962]" pageId="18" pageNumber="276">sp. nov.</emphasis>
</taxonomicNameLabel>
, paratype male (IZB).
<emphasis id="B95EC220201AFFFC170AF878FC98F816" bold="true" box="[834,859,1941,1962]" pageId="18" pageNumber="276">A.</emphasis>
First pair of legs, anterior view.
<emphasis id="B95EC220201AFFFC10FDF878FB0FF816" bold="true" box="[1205,1228,1941,1962]" pageId="18" pageNumber="276">B.</emphasis>
Promere, porsterior view.
<emphasis id="B95EC220201AFFFC149BF85EFF28F875" bold="true" box="[211,235,1971,1993]" pageId="18" pageNumber="276">C.</emphasis>
Penis, posterior view.
<emphasis id="B95EC220201AFFFC1599F859FE29F875" bold="true" box="[465,490,1972,1993]" pageId="18" pageNumber="276">D.</emphasis>
Gonopods, mesal view.
<emphasis id="B95EC220201AFFFC16A9F859FD3AF875" bold="true" box="[737,761,1972,1993]" pageId="18" pageNumber="276">E.</emphasis>
Gonopods, lateral view.
<emphasis id="B95EC220201AFFFC17BCF85EFB68F875" bold="true" box="[1012,1195,1971,1993]" pageId="18" pageNumber="276">Abbreviations: l:</emphasis>
lamella;
<emphasis id="B95EC220201AFFFC1141F859FAE4F875" bold="true" box="[1289,1319,1972,1993]" pageId="18" pageNumber="276">m:</emphasis>
mesomere;
<emphasis id="B95EC220201AFFFC14DFF83FFF6FF85B" bold="true" box="[151,172,2002,2023]" pageId="18" pageNumber="276">o:</emphasis>
opisthomere;
<emphasis id="B95EC220201AFFFC1575F83FFE97F85B" bold="true" box="[317,340,2002,2023]" pageId="18" pageNumber="276">p:</emphasis>
promere;
<emphasis id="B95EC220201AFFFC15F3F83FFE0EF85B" bold="true" box="[443,461,2002,2023]" pageId="18" pageNumber="276">s:</emphasis>
solenomere;
<emphasis id="B95EC220201AFFFC161EF83FFDA4F85B" bold="true" box="[598,615,2002,2023]" pageId="18" pageNumber="276">t:</emphasis>
telopodite;
<emphasis id="B95EC220201AFFFC16A8F83FFD36F85B" bold="true" box="[736,757,2002,2023]" pageId="18" pageNumber="276">v:</emphasis>
velum;
<emphasis id="B95EC220201AFFFC1700F83FFCAFF85B" bold="true" box="[840,876,2002,2023]" pageId="18" pageNumber="276">ml:</emphasis>
mesal lobe.
<emphasis id="B95EC220201AFFFC17A4F83CFB98F854" bold="true" box="[1004,1115,2001,2024]" pageId="18" pageNumber="276">Scale line:</emphasis>
0.3 mm.
</paragraph>
</caption>
<paragraph id="8B951E32201BFFFD148FFF75FD12FF68" blockId="19.[151,1436,151,717]" pageId="19" pageNumber="277">
<typeStatus id="5491A090201BFFFD148FFF75FEEAFF0C" box="[199,297,152,176]" pageId="19" pageNumber="277" type="paratype">Paratype</typeStatus>
(female)
<quantity id="4CD2B3D7201BFFFD15DDFF75FE27FF13" box="[405,484,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="277" unit="mm" value="40.0">40 mm</quantity>
long, vertical diameter of the largest body ring
<quantity id="4CD2B3D7201BFFFD17B1FF75FB93FF13" box="[1017,1104,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="19" pageNumber="277" unit="mm" value="2.4">2.4 mm</quantity>
, body with 54 podous rings + 0 apodous rings + telson. Hypoproct with 10 setae.
</paragraph>
<paragraph id="8B951E32201BFFFD148FFF32FF15FE73" blockId="19.[151,1436,151,717]" pageId="19" pageNumber="277">
<emphasis id="B95EC220201BFFFD148FFF32FEF6FF44" bold="true" box="[199,309,223,248]" pageId="19" pageNumber="277">Variation</emphasis>
. The smallest
<typeStatus id="5491A090201BFFFD1594FF0DFDFFFF44" box="[476,572,224,248]" pageId="19" pageNumber="277" type="paratype">paratype</typeStatus>
male
<quantity id="4CD2B3D7201BFFFD16CAFF32FD25FF4B" box="[642,742,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="19" pageNumber="277" unit="mm" value="17.5">17.5 mm</quantity>
long, vertical diameter of the largest body ring
<quantity id="4CD2B3D7201BFFFD1148FF0DFA9BFF4B" box="[1280,1368,224,248]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="19" pageNumber="277" unit="mm" value="1.7">1.7 mm</quantity>
, body with 40 podous rings + 3 apodous rings + telson. Antennae
<quantity id="4CD2B3D7201BFFFD177AFEE9FC4BFEA0" box="[818,904,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="19" pageNumber="277" unit="mm" value="2.2">2.2 mm</quantity>
long in the smallest
<typeStatus id="5491A090201BFFFD1038FEE8FB13FEA1" box="[1136,1232,261,285]" pageId="19" pageNumber="277" type="paratype">paratype</typeStatus>
male, their length 130% of vertical diameter of the largest body ring. Length of antennomeres IVIII (in mm): 0.15 (I), 0.4 (II), 0.41 (III), 0.33 (IV), 0.43 (V), 0.33 (VI), 0.1 (VII), and 0.05 (VIII). Length/width ratio of antennomeres IVII: 0.8 (I), 2.7 (II), 2.7 (III), 2 (IV), 2 (V), 1.7 (VI), and 0.8 (VII). Hypoproct with 1+1 long apical setae and 1+1 long, medial setae. The other two
<typeStatus id="5491A090201BFFFD15CAFE78FE21FE11" box="[386,482,405,429]" pageId="19" pageNumber="277" type="paratype">paratype</typeStatus>
males with 52 or 55 podous rings + 1 or 0 apodous rings + telson. Hypoproct with 8 setae.
</paragraph>
</subSubSection>
<subSubSection id="C3304DB9201BFFFD148FFE31FEB9FD14" pageId="19" pageNumber="277" type="distribution">
<paragraph id="8B951E32201BFFFD148FFE31FE63FDA4" blockId="19.[151,1436,151,717]" pageId="19" pageNumber="277">
<emphasis id="B95EC220201BFFFD148FFE31FE9AFE49" bold="true" box="[199,345,476,501]" pageId="19" pageNumber="277">Distribution</emphasis>
: This species is known only from its
<typeStatus id="5491A090201BFFFD1744FE30FCFFFE49" box="[780,828,477,501]" pageId="19" pageNumber="277">type</typeStatus>
locality, Jama na Boroviku Pit on the Island of Hvar (
<figureCitation id="131102B7201BFFFD14D7FE12FF31FDA4" box="[159,242,511,536]" captionStart="FIGURE 20" captionStartId="28.[151,250,1947,1969]" captionTargetBox="[151,1436,1121,1925]" captionTargetId="figure@28.[151,1436,1121,1925]" captionTargetPageId="28" captionText="FIGURE 20. Distribution of representatives of Inversotyphlus Strasser, 1962 and Typhloiulus edentulus Attems, 1951 (Attemsotyphlus Strasser, 1962)." httpUri="https://zenodo.org/record/1457209/files/figure.png" pageId="19" pageNumber="277">Fig. 20</figureCitation>
, yellow circle).
</paragraph>
<paragraph id="8B951E32201BFFFD148FFDC9FEB9FD14" blockId="19.[151,1436,151,717]" pageId="19" pageNumber="277">
<emphasis id="B95EC220201BFFFD148FFDC9FEC9FD81" bold="true" box="[199,266,548,573]" pageId="19" pageNumber="277">Notes</emphasis>
: Jama na Boroviku Pit is
<quantity id="4CD2B3D7201BFFFD160FFDC9FD47FD80" box="[583,644,548,572]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="19" pageNumber="277" unit="m" value="30.0">30 m</quantity>
deep and is inhabited by other invertebrates, such as
<taxonomicName id="4C2A65B1201BFFFD1140FDC9FD0BFDDC" authority="Muller, 1931" authorityName="Muller" authorityYear="1931" class="Insecta" family="Leiodidae" genus="Spelaeobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="277" phylum="Arthropoda" rank="subSpecies" species="pharensis" subGenus="Pretneriella" subSpecies="langhofferi">
<emphasis id="B95EC220201BFFFD1140FDC9FDEFFDDC" italics="true" pageId="19" pageNumber="277">Spelaeobates (Pretneriella) pharensis langhofferi</emphasis>
<bibRefCitation id="EFBB63C3201BFFFD167CFDAAFD0BFDDC" box="[564,712,583,608]" pageId="19" pageNumber="277" refId="ref15740" refString="Muller, J. G. (1931) Nuovi coleotteri cavernicoli e ipogei delle Alpi meridionali e del Carso Adriatico. Atti del museo civico di storia naturale di Trieste, 11, 179 - 205." type="journal article">Müller, 1931</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C2A65B1201BFFFD174CFDAAFA94FDDC" authority="(Reitter, 1881)" baseAuthorityName="Reitter" baseAuthorityYear="1881" box="[772,1367,583,608]" class="Insecta" family="Leiodidae" genus="Phaneropella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="277" phylum="Arthropoda" rank="species" species="lesinae" subGenus="Phaneropella">
<emphasis id="B95EC220201BFFFD174CFDAAFB64FDDC" box="[772,1191,583,608]" italics="true" pageId="19" pageNumber="277">Phaneropella (Phaneropella) lesinae</emphasis>
(
<bibRefCitation id="EFBB63C3201BFFFD10F0FDAAFA8CFDDC" box="[1208,1359,583,608]" pageId="19" pageNumber="277" refId="ref15974" refString="Reitter, E. (1881) Neue und seltene Coleopteren im Jahre 1880 in Suddalmatien und Montenegro gesammelt und beschrieben. Deutsche entomologische Zeitschrift, 25 (1), 177 - 230. https: // doi. org / 10.1002 / mmnd. 48018810122" type="journal article">Reitter, 1881</bibRefCitation>
)
</taxonomicName>
(both
<taxonomicName id="4C2A65B1201BFFFD14DFFD81FED4FD39" authorityName="" authorityYear="1758" box="[151,279,620,645]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="277" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
);
<taxonomicName id="4C2A65B1201BFFFD1565FD81FE63FD39" authorityName="Bolívar" authorityYear="1887" box="[301,416,620,645]" class="Insecta" family="Tetrigidae" genus="Chthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="277" phylum="Arthropoda" rank="genus">
<emphasis id="B95EC220201BFFFD1565FD81FE63FD39" box="[301,416,620,645]" italics="true" pageId="19" pageNumber="277">Chthonius</emphasis>
</taxonomicName>
sp.,
<taxonomicName id="4C2A65B1201BFFFD1594FD81FD4FFD39" box="[476,652,620,645]" class="Arachnida" family="Chthoniidae" genus="Microchthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="19" pageNumber="277" phylum="Arthropoda" rank="genus">
<emphasis id="B95EC220201BFFFD1594FD81FD4FFD39" box="[476,652,620,645]" italics="true" pageId="19" pageNumber="277">Microchthonius</emphasis>
</taxonomicName>
sp., and
<taxonomicName id="4C2A65B1201BFFFD16B2FD81FCB1FD39" authorityName="Chamberlin" authorityYear="1930" box="[762,882,620,645]" class="Arachnida" family="Neobisiidae" genus="Neobisium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="19" pageNumber="277" phylum="Arthropoda" rank="genus">
<emphasis id="B95EC220201BFFFD16B2FD81FCB1FD39" box="[762,882,620,645]" italics="true" pageId="19" pageNumber="277">Neobisium</emphasis>
</taxonomicName>
sp. (all
<taxonomicName id="4C2A65B1201BFFFD179EFD81FB67FD39" box="[982,1188,620,645]" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="19" pageNumber="277" phylum="Arthropoda" rank="order">Pseudoscorpiones</taxonomicName>
); and
<taxonomicName id="4C2A65B1201BFFFD10A5FD81FAB1FD39" box="[1261,1394,620,645]" class="Gastropoda" family="Agardhiellidae" genus="Agardhiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="277" phylum="Mollusca" rank="genus">
<emphasis id="B95EC220201BFFFD10A5FD81FAB1FD39" box="[1261,1394,620,645]" italics="true" pageId="19" pageNumber="277">Agardhiella</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="4C2A65B1201BFFFD14D6FD62FEADFD14" box="[158,366,655,680]" class="Gastropoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="277" phylum="Mollusca" rank="order">Stylommatophora</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3304DB9201BFFFD148FFD59FC77FD71" box="[199,948,692,717]" pageId="19" pageNumber="277" type="etymology">
<paragraph id="8B951E32201BFFFD148FFD59FC77FD71" blockId="19.[151,1436,151,717]" box="[199,948,692,717]" pageId="19" pageNumber="277">
<emphasis id="B95EC220201BFFFD148FFD59FE8DFD71" bold="true" box="[199,334,692,717]" pageId="19" pageNumber="277">Etymology.</emphasis>
To denote the slender, elongated gonopods. Adjective.
</paragraph>
</subSubSection>
</treatment>
</document>