treatments-xml/data/D1/53/87/D15387F9FF9CFF8A9CBB33C2FC3F0BD5.xml
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<document id="20DA7430E5F6B8A85523BADD35631193" ID-CLB-Dataset="29265" ID-DOI="10.11646/zootaxa.4496.1.19" ID-GBIF-Dataset="1cf4a9f3-3009-4abe-a993-5bacf7508661" ID-ISSN="1175-5326" ID-Zenodo-Dep="1446783" ID-ZooBank="2BA480FD-2657-4740-9C71-4469FB27E3C1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538747212016" checkinUser="plazi" docAuthor="Hong, Yong, Inkhavilay, Khamla &amp; James, Samuel W." docDate="2018" docId="D15387F9FF9CFF8A9CBB33C2FC3F0BD5" docLanguage="en" docName="zootaxa.4496.1.19.pdf" docOrigin="Zootaxa 4496 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amynthas nametensis Hong &amp; Inkhavilay &amp; James 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="243" masterDocId="2D6AFF81FF99FF8F9C2C314DFF8E0C43" masterDocTitle="Nine new species of the &quot; aeruginosus &quot; - group in Amynthas (Clitellata: Megascolecidae) from Nam Et-Phouley National protected Area, Laos" masterLastPageNumber="250" masterPageNumber="238" pageNumber="243" updateTime="1698565735469" updateUser="ExternalLinkService">
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<mods:title id="A4D2412C9FFEB37DC1E07CF7AF008994">Nine new species of the &quot; aeruginosus &quot; - group in Amynthas (Clitellata: Megascolecidae) from Nam Et-Phouley National protected Area, Laos</mods:title>
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<mods:name id="06E97EFFE3936DAAE3F0330CA4F19202" type="personal">
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<mods:namePart id="A21ED72FC7BA649370B69D2DEE37486B">Hong, Yong</mods:namePart>
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<mods:namePart id="FF8AA0CEF032AD333C5470AC3142DD15">Inkhavilay, Khamla</mods:namePart>
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<mods:namePart id="8F98D4B2D1903A3ACC9E72AB6A4EA4BD">James, Samuel W.</mods:namePart>
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<mods:title id="2D9642653EFEB31445DAD221F80FFE0A">Zootaxa</mods:title>
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<mods:date id="DA112DB19A8721778E054FC0312DF035">2018</mods:date>
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<mods:number id="C6092374BE1FD4A82DBBF313FADDE12F">2018-10-05</mods:number>
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<treatment id="D15387F9FF9CFF8A9CBB33C2FC3F0BD5" ID-DOI="http://doi.org/10.5281/zenodo.5989253" ID-GBIF-Taxon="148639829" ID-Zenodo-Dep="5989253" LSID="urn:lsid:plazi:treatment:D15387F9FF9CFF8A9CBB33C2FC3F0BD5" httpUri="http://treatment.plazi.org/id/D15387F9FF9CFF8A9CBB33C2FC3F0BD5" lastPageNumber="243" pageId="5" pageNumber="243">
<subSubSection id="11E06564FF9CFF8A9CBB33C2FEBB0E88" pageId="5" pageNumber="243" type="nomenclature">
<paragraph id="594536EFFF9CFF8A9CBB33C2FD8A0EE9" blockId="5.[151,516,655,715]" box="[151,516,655,682]" pageId="5" pageNumber="243">
<heading id="020D8183FF9CFF8A9CBB33C2FD8A0EE9" bold="true" box="[151,516,655,682]" fontSize="11" level="1" pageId="5" pageNumber="243" reason="1">
<emphasis id="6B8EEAFDFF9CFF8A9CBB33C2FD8A0EE9" bold="true" box="[151,516,655,682]" pageId="5" pageNumber="243">
<taxonomicName id="9EFA4D6CFF9CFF8A9CBB33C2FE110EEA" authority="Hong &amp; Inkhavilay &amp; James, 2018" authorityName="Hong &amp; Inkhavilay &amp; James" authorityYear="2018" box="[151,415,655,681]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="243" phylum="Annelida" rank="species" species="nametensis" status="sp. nov.">
<emphasis id="6B8EEAFDFF9CFF8A9CBB33C2FE110EEA" bold="true" box="[151,415,655,681]" italics="true" pageId="5" pageNumber="243">Amynthas nametensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70BD5786FF9CFF8A9D8B33DDFD8A0EE9" box="[423,516,656,682]" pageId="5" pageNumber="243" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="594536EFFF9CFF8A9CBB33FFFEBB0E88" blockId="5.[151,516,655,715]" box="[151,309,690,715]" pageId="5" pageNumber="243">
(
<figureCitation id="C1C12A6AFF9CFF8A9CB233FFFEA30E88" box="[158,301,690,715]" captionStart="FIGURE 2" captionStartId="7.[151,250,1956,1978]" captionTargetBox="[164,1389,203,1926]" captionTargetId="figure@7.[155,1431,193,1934]" captionTargetPageId="7" captionText="FIGURE 2. Amynthas nametensis sp. nov. (A,B), Amynthas somechanae sp. nov. (C,D), and Amynthas wiggeri sp. nov. (E,F). A,C,E: ventral view; B,D,F: spermatheca. Scale bars 2 mm." httpUri="https://zenodo.org/record/1446787/files/figure.png" pageId="5" pageNumber="243">Figure 2A,B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="11E06564FF9CFF8A9CBB33B7FDCD0FE5" pageId="5" pageNumber="243" type="materials_examined">
<paragraph id="594536EFFF9CFF8A9CBB33B7FDCD0FE5" blockId="5.[151,1437,762,1942]" pageId="5" pageNumber="243">
<materialsCitation id="E9923CB2FF9CFF8A9CBB33B7FA160F1D" ID-GBIF-Occurrence="1914265335" collectingDate="2005-07-21" collectionCode="NBCA" collectorName="Phimphet &amp; Xay &amp; Ammaline &amp; Phoang &amp; K. Inkhavilay" country="Laos" county="Thad Ang" elevation="792" latitude="20.257433" location="Hoaphane Province" longLatPrecision="1" longitude="103.00248" municipality="Viengthong district" pageId="5" pageNumber="243" specimenCode="BDNUL0038" specimenCount="1" typeStatus="holotype">
<emphasis id="6B8EEAFDFF9CFF8A9CBB33B7FE880F50" bold="true" box="[151,262,762,787]" pageId="5" pageNumber="243">Material.</emphasis>
<typeStatus id="8641884DFF9CFF8A9D3D33B7FEF00F50" box="[273,382,762,787]" pageId="5" pageNumber="243" type="holotype">Holotype</typeStatus>
: clitellate (
<specimenCode id="095C9E94FF9CFF8A9DD333B6FD110F50" box="[511,671,763,787]" collectionCode="BDNUL" pageId="5" pageNumber="243">BDNUL0038</specimenCode>
):
<collectingCountry id="21ED767FFF9CFF8A9E9933B6FD600F51" box="[693,750,763,786]" name="Laos" pageId="5" pageNumber="243">Laos</collectingCountry>
,
<location id="5C256034FF9CFF8A9ED233B7FC6F0F50" LSID="urn:lsid:plazi:treatment:D15387F9FF9CFF8A9CBB33C2FC3F0BD5:5C256034FF9CFF8A9ED233B7FC6F0F50" box="[766,993,762,787]" country="Laos" county="Thad Ang" latitude="20.257433" longLatPrecision="1" longitude="103.00248" municipality="Viengthong district" name="Hoaphane Province" pageId="5" pageNumber="243">Hoaphane Province</location>
,
<location id="5C256034FF9CFF8A9FDC33B6FB210F50" LSID="urn:lsid:plazi:treatment:D15387F9FF9CFF8A9CBB33C2FC3F0BD5:5C256034FF9CFF8A9FDC33B6FB210F50" box="[1008,1199,762,787]" country="Laos" county="Thad Ang" latitude="20.257433" longLatPrecision="1" longitude="103.00248" municipality="Viengthong district" name="Nam Et-Phouley" pageId="5" pageNumber="243">Nam Et-Phouley</location>
<collectionCode id="3FEBAE2AFF9CFF8A989633B6FA850F50" box="[1210,1291,763,787]" pageId="5" pageNumber="243">NBCA</collectionCode>
,
<collectingMunicipality id="B921AC95FF9CFF8A993633B7FF660F79" pageId="5" pageNumber="243">Viengthong district</collectingMunicipality>
,
<collectingCounty id="B0244E63FF9CFF8A9CDE326CFEEC0F79" box="[242,354,801,826]" pageId="5" pageNumber="243">Thad Ang</collectingCounty>
waterfall area,
<elevation id="D2D7D1DCFF9CFF8A9E3C326FFDD80F7A" box="[528,598,802,826]" metricMagnitude="2" metricUnit="m" metricValue="7.92" pageId="5" pageNumber="243" unit="m" value="792.0">
<quantity id="9E029B0AFF9CFF8A9E3C326FFDD80F7A" box="[528,598,802,826]" metricMagnitude="2" metricUnit="m" metricValue="7.92" pageId="5" pageNumber="243" unit="m" value="792.0">792 m</quantity>
</elevation>
(
<geoCoordinate id="3CCE5028FF9CFF8A9E49326FFD790F7A" box="[613,759,798,826]" direction="north" orientation="latitude" pageId="5" pageNumber="243" precision="1" value="20.257433">20o 15.446'N</geoCoordinate>
,
<geoCoordinate id="3CCE5028FF9CFF8A9F2E326FFC110F7A" box="[770,927,798,826]" direction="east" orientation="longitude" pageId="5" pageNumber="243" precision="1" value="103.00248">103o 0 8.921' E</geoCoordinate>
), good condition mossy forest with big trees, dark soil and thick litter layer,
<collectingDate id="3D00E9C7FF9CFF8A9DD3320BFD160F1D" box="[511,664,837,862]" pageId="5" pageNumber="243" value="2005-07-21">
<date id="2D44102FFF9CFF8A9DD3320BFD160F1D" box="[511,664,837,862]" pageId="5" pageNumber="243" value="2005-07-21">21 July 2005</date>
</collectingDate>
,
<collectorName id="F40F5339FF9CFF8A9E893208FC9A0F1D" box="[677,788,837,862]" pageId="5" pageNumber="243">Phimphet</collectorName>
,
<collectorName id="F40F5339FF9CFF8A9F0D320BFCDF0F1D" box="[801,849,838,862]" pageId="5" pageNumber="243">Xay</collectorName>
,
<collectorName id="F40F5339FF9CFF8A9F733208FC560F1D" box="[863,984,837,862]" pageId="5" pageNumber="243">Ammaline</collectorName>
,
<collectorName id="F40F5339FF9CFF8A9FCA3208FBB40F1D" box="[998,1082,837,862]" pageId="5" pageNumber="243">Phoang</collectorName>
sone, and
<collectorName id="F40F5339FF9CFF8A9896320BFAD90F1D" box="[1210,1367,837,862]" pageId="5" pageNumber="243">K. Inkhavilay</collectorName>
colls
</materialsCitation>
.
<materialsCitation id="E9923CB2FF9CFF8A9CBB3227FBB70FC1" ID-GBIF-Occurrence="1914265346" box="[151,1081,873,898]" collectingDate="2005-07-21" collectionCode="BDNUL" collectorName="Phimphet &amp; Xay &amp; Ammaline &amp; Phoang &amp; K. Inkhavilay" country="Laos" county="Thad Ang" elevation="792" latitude="20.257433" location="Hoaphane Province" longLatPrecision="1" longitude="103.00248" municipality="Viengthong district" pageId="5" pageNumber="243" specimenCode="BDNUL0039, BDNUL0040" specimenCount="2" typeStatus="paratype">
<typeStatus id="8641884DFF9CFF8A9CBB3227FE870FC1" box="[151,265,874,898]" pageId="5" pageNumber="243" type="paratype">Paratypes</typeStatus>
: 2 clitellates (
<specimenCode id="095C9E94FF9CFF8A9D843227FDCB0FC1" box="[424,581,874,898]" collectionCode="BDNUL" pageId="5" pageNumber="243">BDNUL0039</specimenCode>
, (
<specimenCode id="095C9E94FF9CFF8A9E763227FD770FC1" box="[602,761,874,898]" collectionCode="BDNUL" pageId="5" pageNumber="243">BDNUL0040</specimenCode>
), same data as for
<typeStatus id="8641884DFF9CFF8A9FFF3224FBB70FC1" box="[979,1081,873,898]" pageId="5" pageNumber="243" type="holotype">holotype</typeStatus>
</materialsCitation>
.
<materialsCitation id="E9923CB2FF9CFF8A98693224FDB10FE5" ID-GBIF-Occurrence="1914265341" collectingDate="2005-07-21" collectionCode="BDNUL" collectorName="Phimphet &amp; Xay &amp; Ammaline &amp; Phoang &amp; K. Inkhavilay" country="Laos" county="Thad Ang" elevation="792" latitude="20.257433" location="Hoaphane Province" longLatPrecision="1" longitude="103.00248" municipality="Viengthong district" pageId="5" pageNumber="243" specimenCount="1" typeStatus="holotype">
Other material: 3 clitellates, 6 aclitellates, same data as for
<typeStatus id="8641884DFF9CFF8A9DF532C0FDB10FE5" box="[473,575,909,934]" pageId="5" pageNumber="243" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="11E06564FF9CFF8A9CEB32FCFC340F8A" box="[199,954,945,970]" pageId="5" pageNumber="243" type="etymology">
<paragraph id="594536EFFF9CFF8A9CEB32FCFC340F8A" blockId="5.[151,1437,762,1942]" box="[199,954,945,970]" pageId="5" pageNumber="243">
<emphasis id="6B8EEAFDFF9CFF8A9CEB32FCFEC00F8A" bold="true" box="[199,334,945,969]" pageId="5" pageNumber="243">Etymology.</emphasis>
The species is named for its
<typeStatus id="8641884DFF9CFF8A9EBF32FFFD4D0F89" box="[659,707,946,970]" pageId="5" pageNumber="243">type</typeStatus>
locality near Nam Et.
</paragraph>
</subSubSection>
<subSubSection id="11E06564FF9CFF8A9CEB3299FD1D0AA1" pageId="5" pageNumber="243" type="diagnosis">
<paragraph id="594536EFFF9CFF8A9CEB3299FB760819" blockId="5.[151,1437,762,1942]" pageId="5" pageNumber="243">
<emphasis id="6B8EEAFDFF9CFF8A9CEB3299FECF0FAE" bold="true" box="[199,321,980,1005]" pageId="5" pageNumber="243">Diagnosis.</emphasis>
Spermathecal pores paired in 7/8, 8/9 at 12th13th setal lines; male pores superficial on thickened circular pads, surrounded by one white convex outer ring in XVIII at 10th setal line, 0.240.27 circumference ventrally apart, elevated genital papillae in line with male pores presetal in XVIII, XIX; first dorsal pore at 12/13, large typhlosole, intestinal origin in XVI, spermathecal diverticula coiled into dense irregular block.
</paragraph>
<paragraph id="594536EFFF9CFF8A9CEB3529FBD409DD" blockId="5.[151,1437,762,1942]" pageId="5" pageNumber="243">
<emphasis id="6B8EEAFDFF9CFF8A9CEB3529FEC0083E" bold="true" box="[199,334,1124,1149]" pageId="5" pageNumber="243">Decription.</emphasis>
Dimensions
<quantity id="9E029B0AFF9CFF8A9DCC352BFDE6083E" box="[480,616,1126,1150]" metricMagnitude="-2" metricUnit="m" metricValue="8.9" metricValueMax="10.0" metricValueMin="7.8" pageId="5" pageNumber="243" unit="mm" value="89.0" valueMax="100.0" valueMin="78.0">78100 mm</quantity>
, by 5.06.0 mm at segment X,
<quantity id="9E029B0AFF9CFF8A9FEE3528FBC6083E" box="[962,1096,1125,1150]" metricMagnitude="-3" metricUnit="m" metricValue="5.15" metricValueMax="5.5" metricValueMin="4.8" pageId="5" pageNumber="243" unit="mm" value="5.15" valueMax="5.5" valueMin="4.8">4.85.5 mm</quantity>
at segment XXX,
<quantity id="9E029B0AFF9CFF8A993A3528FA13083E" box="[1302,1437,1125,1150]" metricMagnitude="-3" metricUnit="m" metricValue="4.55" metricValueMax="4.6" metricValueMin="4.5" pageId="5" pageNumber="243" unit="mm" value="4.55" valueMax="4.6" valueMin="4.5">4.54.6 mm</quantity>
at clitellum; body cylindrical, segments 121136. Setae evenly distributed around segmental equators, numbering 5864 at VII, 6370 at XX, 26 between male pores, setal formula AA:AB:ZZ:YZ= 4:2:4:1 at XIII. Female pore single at XIV on an oval porophore
<quantity id="9E029B0AFF9CFF8A9E01359CFD3A08AA" box="[557,692,1233,1258]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="5" pageNumber="243" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.50.6 mm</quantity>
in diameter. Prostomium epilobic with tongue open, unpigmented except IVIII, clitellum light brown, formalin preservation. First dorsal pore at 12/13. Clitellum annular XIVXVI; setae invisible externally. Male pores superficial on circular thickened areas, surrounded by one white outer ring at lateral margins of ventrum in XVIII at 10th setal lines, 0.240.27 circumference apart ventrally. Elevated genital papillae in line with male pores presetal in XVIII, XIX. Spermathecal pores paired in 7/8, 8/9 at 12th13th setal lines, 0.290.37 circumference apart ventrally; conspicuous bumps below mid-lateral.
</paragraph>
<paragraph id="594536EFFF9CFF8A9CEB34E7FD1D0AA1" blockId="5.[151,1437,762,1942]" pageId="5" pageNumber="243">Septa 5/67/8 very thick, 8/9 thin, 9/10 muscular, 10/11 thin, 11/12 thick, 12/13 thin. Gizzard globular in VIII. Intestine enlarged from XVI, lymph glands not found. Typhlosole very deep fold from XXIV. Intestinal caeca simple originating in XXVII and extending anteriorly about to XXII, simple finger-shaped. Hearts esophageal X XIII; X small. Male sexual system holandric, testes and funnels in annular sacs in X, XI; sacs enclose all other segmental organs. Seminal vesicles paired in XI, XII, small. Prostates in XVIII, and extending from XVIIXXIII; long stout ducts coiled, glandular portions consisted of 2 main lobes each divided into 23 small lobes. Ovaries in XIII. Spermathecae paired in VII, VIII; ampulla medium size globular-shaped, duct much shorter than ampulla, diverticulum tubular, iridescent, coiled into irregular ovate block like a large oval chamber, stalk much shorter than ampulla, no nephridia on spermathecal ducts.</paragraph>
</subSubSection>
<subSubSection id="11E06564FF9CFF8A9CEB37A3FC3F0BD5" pageId="5" pageNumber="243" type="discussion">
<paragraph id="594536EFFF9CFF8A9CEB37A3FC3F0BD5" blockId="5.[151,1437,762,1942]" pageId="5" pageNumber="243">
<emphasis id="6B8EEAFDFF9CFF8A9CEB37A3FEB50B45" bold="true" box="[199,315,1774,1798]" pageId="5" pageNumber="243">Remarks.</emphasis>
<taxonomicName id="9EFA4D6CFF9CFF8A9D6D37A0FDBD0B46" authority="Hong &amp; Inkhavilay &amp; James, 2018" authorityName="Hong &amp; Inkhavilay &amp; James" authorityYear="2018" box="[321,563,1773,1797]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="243" phylum="Annelida" rank="species" species="nametensis" status="sp. nov.">
<emphasis id="6B8EEAFDFF9CFF8A9D6D37A0FDBD0B46" box="[321,563,1773,1797]" italics="true" pageId="5" pageNumber="243">Amynthas nametensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70BD5786FF9CFF8A9E1637A3FD1A0B45" box="[570,660,1774,1798]" pageId="5" pageNumber="243" rank="species">
<emphasis id="6B8EEAFDFF9CFF8A9E1637A3FD1A0B45" bold="true" box="[570,660,1774,1798]" pageId="5" pageNumber="243">sp. nov.</emphasis>
</taxonomicNameLabel>
is unusual in the present collection because it has annular testes sacs, first dorsal pore in the more typical
<taxonomicName id="9EFA4D6CFF9CFF8A9E00365CFD140B6A" authorityName="Hong &amp; Inkhavilay &amp; James" authorityYear="2018" box="[556,666,1809,1833]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="243" phylum="Annelida" rank="genus">
<emphasis id="6B8EEAFDFF9CFF8A9E00365CFD140B6A" box="[556,666,1809,1833]" italics="true" pageId="5" pageNumber="243">Amynthas</emphasis>
</taxonomicName>
location of 12/13, an intestinal origin in XVI rather than XV, a very well-developed typhlosole which starts before the caeca, and septa
<date id="2D44102FFF9CFF8A9FBB367BFC510B0D" box="[919,991,1846,1870]" pageId="5" pageNumber="243" value="2010-09-08">8/9/10</date>
present and thickened to some degree.
<taxonomicName id="9EFA4D6CFF9CFF8A9CBB3614FEE20B32" baseAuthorityName="Thai &amp; Samphon" baseAuthorityYear="1988" box="[151,364,1881,1905]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="243" phylum="Annelida" rank="species" species="choanus">
<emphasis id="6B8EEAFDFF9CFF8A9CBB3614FEE20B32" box="[151,364,1881,1905]" italics="true" pageId="5" pageNumber="243">Amynthas choanus</emphasis>
</taxonomicName>
and
<taxonomicName id="9EFA4D6CFF9CFF8A9D853617FD810B32" baseAuthorityName="Thai &amp; Samphon" baseAuthorityYear="1990" box="[425,527,1882,1905]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="243" phylum="Annelida" rank="species" species="uanus">
<emphasis id="6B8EEAFDFF9CFF8A9D853617FD810B32" box="[425,527,1882,1905]" italics="true" pageId="5" pageNumber="243">A. uanus</emphasis>
</taxonomicName>
have dorsal pores in
<date id="2D44102FFF9CFF8A9F213617FC440B31" box="[781,970,1881,1906]" pageId="5" pageNumber="243" value="2011-12-13" valueMax="2011-12-10" valueMin="2011-12-13">12/13 and 10/11</date>
, respectively, but the former has genital markings in VII and VIII and the latter has C-shaped male porophores.
</paragraph>
</subSubSection>
</treatment>
</document>