264 lines
32 KiB
XML
264 lines
32 KiB
XML
<document id="C45FA038C778F60D2CF014E320EF72F1" ID-CLB-Dataset="29265" ID-DOI="10.11646/zootaxa.4496.1.19" ID-GBIF-Dataset="1cf4a9f3-3009-4abe-a993-5bacf7508661" ID-ISSN="1175-5326" ID-Zenodo-Dep="1446783" ID-ZooBank="2BA480FD-2657-4740-9C71-4469FB27E3C1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538747212016" checkinUser="plazi" docAuthor="Hong, Yong, Inkhavilay, Khamla & James, Samuel W." docDate="2018" docId="D15387F9FF91FF869CBB3744FEF00ADB" docLanguage="en" docName="zootaxa.4496.1.19.pdf" docOrigin="Zootaxa 4496 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amynthas fleischmani Hong & Inkhavilay & James 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="247" masterDocId="2D6AFF81FF99FF8F9C2C314DFF8E0C43" masterDocTitle="Nine new species of the " aeruginosus " - group in Amynthas (Clitellata: Megascolecidae) from Nam Et-Phouley National protected Area, Laos" masterLastPageNumber="250" masterPageNumber="238" pageNumber="246" updateTime="1698565735469" updateUser="ExternalLinkService">
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<mods:mods id="EF9EF78ABD69E5338ED32F1C7A947E81" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A9D298CF2797C4C757BFADBE49ECFA13">
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<mods:title id="9360CA52F12315BBBA023FEDA29A598F">Nine new species of the " aeruginosus " - group in Amynthas (Clitellata: Megascolecidae) from Nam Et-Phouley National protected Area, Laos</mods:title>
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<mods:name id="3EC2119BF834709E2BC042BB575BF6BE" type="personal">
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<mods:roleTerm id="77BDA1D5A8CF5CF3807E3E1DF0C14185">Author</mods:roleTerm>
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<mods:namePart id="4BA3D5682685AB73441A2AEA7FEDEEF8">Hong, Yong</mods:namePart>
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<mods:namePart id="40617F69E13A438D441DE19FD50EF9D6">Inkhavilay, Khamla</mods:namePart>
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<mods:roleTerm id="9F67CC91BEA0E5CD55CC5E54C5BFD175">Author</mods:roleTerm>
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<mods:namePart id="7B7CB2D642C9BBE63878E0D4FE95426D">James, Samuel W.</mods:namePart>
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<mods:typeOfResource id="0F11A2BBC873CBC13D7050F5DABA3D46">text</mods:typeOfResource>
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<mods:title id="E60079F03519FEC5F85C6DF7E8FEE25E">Zootaxa</mods:title>
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<mods:part id="C212A94739624E9607AF6172B0CF158B">
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<mods:date id="8B85C5EE452851076B50116241CB4ED5">2018</mods:date>
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<mods:number id="0091BFDD31E98DD048C8085F95C84CB8">2018-10-05</mods:number>
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<mods:number id="99CA18EE3439425123C47D148BD638DE">4496</mods:number>
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<mods:identifier id="355AB2A509A8A2359750106D5F176725" type="DOI">10.11646/zootaxa.4496.1.19</mods:identifier>
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<treatment id="D15387F9FF91FF869CBB3744FEF00ADB" ID-DOI="http://doi.org/10.5281/zenodo.5989259" ID-GBIF-Taxon="148639833" ID-Zenodo-Dep="5989259" LSID="urn:lsid:plazi:treatment:D15387F9FF91FF869CBB3744FEF00ADB" httpUri="http://treatment.plazi.org/id/D15387F9FF91FF869CBB3744FEF00ADB" lastPageId="9" lastPageNumber="247" pageId="8" pageNumber="246">
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<subSubSection id="11E06564FF91FF879CBB3744FEB80A07" pageId="8" pageNumber="246" type="nomenclature">
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<paragraph id="594536EFFF91FF879CBB3744FD850A60" blockId="8.[151,523,1545,1605]" box="[151,523,1545,1571]" pageId="8" pageNumber="246">
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<heading id="020D8183FF91FF879CBB3744FD850A60" bold="true" box="[151,523,1545,1571]" fontSize="11" level="1" pageId="8" pageNumber="246" reason="1">
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<emphasis id="6B8EEAFDFF91FF879CBB3744FD850A60" bold="true" box="[151,523,1545,1571]" pageId="8" pageNumber="246">
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<taxonomicName id="9EFA4D6CFF91FF879CBB3744FE2B0A60" authority="Hong & Inkhavilay & James, 2018" authorityName="Hong & Inkhavilay & James" authorityYear="2018" box="[151,421,1545,1571]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="246" phylum="Annelida" rank="species" species="fleischmani" status="sp. nov.">
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<emphasis id="6B8EEAFDFF91FF879CBB3744FE2B0A60" bold="true" box="[151,421,1545,1571]" italics="true" pageId="8" pageNumber="246">Amynthas fleischmani</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="70BD5786FF91FF879D813744FD850A60" box="[429,523,1545,1571]" pageId="8" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="594536EFFF91FF879CBB3761FEB80A07" blockId="8.[151,523,1545,1605]" box="[151,310,1580,1605]" pageId="8" pageNumber="246">
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(
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<figureCitation id="C1C12A6AFF91FF879CB23761FEA20A07" box="[158,300,1580,1605]" captionStart="FIGURE 3" captionStartId="10.[151,250,1969,1991]" captionTargetBox="[195,1404,209,1924]" captionTargetId="figure@10.[167,1419,205,1936]" captionTargetPageId="10" captionText="FIGURE 3. Amynthas fleischmani sp. nov. (A,B), Amynthas antethecus sp. nov. (C,D), and Amynthas elenabondae sp. nov. (E,F). A,C,E: ventral view; B,D,F: spermatheca. Scale bars 2 mm." httpUri="https://zenodo.org/record/1446789/files/figure.png" pageId="8" pageNumber="246">Figure 3A,B</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="11E06564FF91FF879CBB3723FA9A0B26" pageId="8" pageNumber="246" type="materials_examined">
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<paragraph id="594536EFFF91FF879CBB3723FA9A0B26" blockId="8.[151,1437,1646,2037]" pageId="8" pageNumber="246">
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<materialsCitation id="E9923CB2FF91FF879CBB3723FF410AB5" ID-GBIF-Occurrence="1914265328" collectingDate="2005-07-21" collectionCode="NPA" collectorName="Phimphet & Xay & Ammaline & Phoang & K. Inkhavilay" country="Laos" county="Thad Ang" elevation="792" location="Hoaphane Province" municipality="Viengthong district" pageId="8" pageNumber="246" specimenCode="BDNUL0047" specimenCount="1" typeStatus="holotype">
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<emphasis id="6B8EEAFDFF91FF879CBB3723FE880AC4" bold="true" box="[151,262,1646,1671]" pageId="8" pageNumber="246">Material.</emphasis>
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<typeStatus id="8641884DFF91FF879D3F3723FE0E0AC4" box="[275,384,1646,1671]" pageId="8" pageNumber="246" type="holotype">Holotype</typeStatus>
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: clitellate (
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<specimenCode id="095C9E94FF91FF879E2A3722FD280AC4" box="[518,678,1647,1671]" collectionCode="BDNUL" pageId="8" pageNumber="246">BDNUL0047</specimenCode>
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):
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<collectingCountry id="21ED767FFF91FF879E933722FD760AC5" box="[703,760,1647,1670]" name="Laos" pageId="8" pageNumber="246">Laos</collectingCountry>
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,
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<location id="5C256034FF91FF879F263723FC600AC4" LSID="urn:lsid:plazi:treatment:D15387F9FF91FF869CBB3744FEF00ADB:5C256034FF91FF879F263723FC600AC4" box="[778,1006,1646,1671]" country="Laos" county="Thad Ang" municipality="Viengthong district" name="Hoaphane Province" pageId="8" pageNumber="246">Hoaphane Province</location>
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,
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<location id="5C256034FF91FF87982C3722FB4C0AC4" LSID="urn:lsid:plazi:treatment:D15387F9FF91FF869CBB3744FEF00ADB:5C256034FF91FF87982C3722FB4C0AC4" box="[1024,1218,1646,1671]" country="Laos" county="Thad Ang" municipality="Viengthong district" name="Nam Et-Phouley" pageId="8" pageNumber="246">Nam Et-Phouley</location>
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<collectionCode id="3FEBAE2AFF91FF8798FC3722FA860AC4" box="[1232,1288,1647,1671]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13690" name="Nanjing Institute of Geology and Paleontology, Academia Sinica" pageId="8" pageNumber="246">NPA</collectionCode>
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,
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<collectingMunicipality id="B921AC95FF91FF8799353723FF660AED" pageId="8" pageNumber="246">Viengthong district</collectingMunicipality>
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,
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<collectingCounty id="B0244E63FF91FF879CD837D8FEE80AED" box="[244,358,1685,1710]" pageId="8" pageNumber="246">Thad Ang</collectingCounty>
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||
waterfall area,
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||
<elevation id="D2D7D1DCFF91FF879E3637DBFDEC0AEE" box="[538,610,1686,1710]" metricMagnitude="2" metricUnit="m" metricValue="7.92" pageId="8" pageNumber="246" unit="m" value="792.0">
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<quantity id="9E029B0AFF91FF879E3637DBFDEC0AEE" box="[538,610,1686,1710]" metricMagnitude="2" metricUnit="m" metricValue="7.92" pageId="8" pageNumber="246" unit="m" value="792.0">792 m</quantity>
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</elevation>
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(20o 15. 446' N, 103o 08– 921' E), good condition mossy forest with big trees, dark soil and thick litter layer,
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||
<collectingDate id="3D00E9C7FF91FF879E6837F7FD520A91" box="[580,732,1721,1746]" pageId="8" pageNumber="246" value="2005-07-21">
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||
<date id="2D44102FFF91FF879E6837F7FD520A91" box="[580,732,1721,1746]" pageId="8" pageNumber="246" value="2005-07-21">21 July 2005</date>
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||
</collectingDate>
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||
,
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||
<collectorName id="F40F5339FF91FF879EC737F4FCD70A91" box="[747,857,1721,1746]" pageId="8" pageNumber="246">Phimphet</collectorName>
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||
,
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<collectorName id="F40F5339FF91FF879F4B37F7FC190A91" box="[871,919,1722,1746]" pageId="8" pageNumber="246">Xay</collectorName>
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,
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<collectorName id="F40F5339FF91FF879F8937F4FB900A91" box="[933,1054,1721,1746]" pageId="8" pageNumber="246">Ammaline</collectorName>
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,
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<collectorName id="F40F5339FF91FF87980737F4FBF10A91" box="[1067,1151,1721,1746]" pageId="8" pageNumber="246">Phoang</collectorName>
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||
sone, and
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<collectorName id="F40F5339FF91FF87992C37F7FA130A91" box="[1280,1437,1721,1746]" pageId="8" pageNumber="246">K. Inkhavilay</collectorName>
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colls
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</materialsCitation>
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||
.
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<materialsCitation id="E9923CB2FF91FF879CCE3793FB230AB5" ID-GBIF-Occurrence="1914265329" box="[226,1197,1757,1782]" collectingDate="2005-07-21" collectionCode="BDNUL" collectorName="Phimphet & Xay & Ammaline & Phoang & K. Inkhavilay" country="Laos" county="Thad Ang" elevation="792" location="Hoaphane Province" municipality="Viengthong district" pageId="8" pageNumber="246" specimenCode="BDNUL0048, BDNUL0049" specimenCount="2" typeStatus="paratype">
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<typeStatus id="8641884DFF91FF879CCE3793FEDD0AB5" box="[226,339,1758,1782]" pageId="8" pageNumber="246" type="paratype">Paratypes</typeStatus>
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: 2 clitellates (
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||
<specimenCode id="095C9E94FF91FF879E283793FD2F0AB5" box="[516,673,1758,1782]" collectionCode="BDNUL" pageId="8" pageNumber="246">BDNUL0048</specimenCode>
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,
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||
<specimenCode id="095C9E94FF91FF879E983793FCDF0AB5" box="[692,849,1758,1782]" collectionCode="BDNUL" pageId="8" pageNumber="246">BDNUL0049</specimenCode>
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||
), same data as for
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<typeStatus id="8641884DFF91FF87986B3790FB230AB5" box="[1095,1197,1757,1782]" pageId="8" pageNumber="246" type="holotype">holotype</typeStatus>
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</materialsCitation>
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.
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<materialsCitation id="E9923CB2FF91FF8798933790FA9A0B26" ID-GBIF-Occurrence="1914265347" collectingDate="2005-07-22" collectionCode="NPA" collectorName="Phimphet & Xay & Ammaline & Phoang & K. Inkhavilay" country="Laos" county="Hoauy Kanang Mountain" elevation="1075" latitude="20.259533" location="Other" municipality="Viengthong district" pageId="8" pageNumber="246" specimenCount="1" typeStatus="holotype">
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<location id="5C256034FF91FF8798933790FB710AB5" LSID="urn:lsid:plazi:treatment:D15387F9FF91FF869CBB3744FEF00ADB:5C256034FF91FF8798933790FB710AB5" box="[1215,1279,1757,1782]" country="Laos" county="Hoauy Kanang Mountain" latitude="20.259533" municipality="Viengthong district" name="Other" pageId="8" pageNumber="246">Other</location>
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material: 21 clitellates, 1 semiclitellate, 1 aclitellate, same data as for
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<typeStatus id="8641884DFF91FF879F34364CFCF00B59" box="[792,894,1793,1818]" pageId="8" pageNumber="246" type="holotype">holotype</typeStatus>
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.
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<location id="5C256034FF91FF879FA5364CFB8C0B59" LSID="urn:lsid:plazi:treatment:D15387F9FF91FF869CBB3744FEF00ADB:5C256034FF91FF879FA5364CFB8C0B59" box="[905,1026,1793,1818]" country="Laos" county="Hoauy Kanang Mountain" latitude="20.259533" municipality="Viengthong district" name="Aclitellate" pageId="8" pageNumber="246">Aclitellate</location>
|
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:
|
||
<collectingCountry id="21ED767FFF91FF879820364FFBCB0B5A" box="[1036,1093,1794,1817]" name="Laos" pageId="8" pageNumber="246">Laos</collectingCountry>
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,
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<location id="5C256034FF91FF87987D364CFABF0B59" LSID="urn:lsid:plazi:treatment:D15387F9FF91FF869CBB3744FEF00ADB:5C256034FF91FF87987D364CFABF0B59" box="[1105,1329,1793,1818]" country="Laos" county="Hoauy Kanang Mountain" latitude="20.259533" municipality="Viengthong district" name="Hoaphane Province" pageId="8" pageNumber="246">Hoaphane Province</location>
|
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,
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<location id="5C256034FF91FF879910364FFAFC0B5A" LSID="urn:lsid:plazi:treatment:D15387F9FF91FF869CBB3744FEF00ADB:5C256034FF91FF879910364FFAFC0B5A" box="[1340,1394,1794,1817]" country="Laos" county="Hoauy Kanang Mountain" latitude="20.259533" municipality="Viengthong district" name="Nam" pageId="8" pageNumber="246">Nam</location>
|
||
Et-
|
||
<location id="5C256034FF91FF879CBB3665FF7D0B02" LSID="urn:lsid:plazi:treatment:D15387F9FF91FF869CBB3744FEF00ADB:5C256034FF91FF879CBB3665FF7D0B02" box="[151,243,1832,1857]" country="Laos" county="Hoauy Kanang Mountain" latitude="20.259533" municipality="Viengthong district" name="Phouley" pageId="8" pageNumber="246">Phouley</location>
|
||
<collectionCode id="3FEBAE2AFF91FF879CD13664FEB80B02" box="[253,310,1833,1857]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13690" name="Nanjing Institute of Geology and Paleontology, Academia Sinica" pageId="8" pageNumber="246">NPA</collectionCode>
|
||
,
|
||
<collectingMunicipality id="B921AC95FF91FF879D693665FDAD0B02" box="[325,547,1832,1857]" pageId="8" pageNumber="246">Viengthong district</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="B0244E63FF91FF879E1E3664FCD40B02" box="[562,858,1832,1857]" pageId="8" pageNumber="246">Hoauy Kanang Mountain</collectingCounty>
|
||
,
|
||
<elevation id="D2D7D1DCFF91FF879F453665FC390B02" box="[873,951,1832,1857]" metricMagnitude="3" metricUnit="m" metricValue="1.075" pageId="8" pageNumber="246" unit="m" value="1075.0">
|
||
<quantity id="9E029B0AFF91FF879F453665FC390B02" box="[873,951,1832,1857]" metricMagnitude="3" metricUnit="m" metricValue="1.075" pageId="8" pageNumber="246" unit="m" value="1075.0">1075m</quantity>
|
||
</elevation>
|
||
(
|
||
<geoCoordinate id="3CCE5028FF91FF879FE53665FBDB0B03" box="[969,1109,1829,1857]" direction="north" orientation="latitude" pageId="8" pageNumber="246" precision="1" value="20.259533">20o15.572'N</geoCoordinate>
|
||
, 103o09–629'E), good forest condition in dark soil,
|
||
<collectingDate id="3D00E9C7FF91FF879DB83600FDA90B26" box="[404,551,1868,1893]" pageId="8" pageNumber="246" value="2005-07-22">
|
||
<date id="2D44102FFF91FF879DB83600FDA90B26" box="[404,551,1868,1893]" pageId="8" pageNumber="246" value="2005-07-22">22 July 2005</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="F40F5339FF91FF879E1E3601FD2E0B26" box="[562,672,1868,1893]" pageId="8" pageNumber="246">Phimphet</collectorName>
|
||
,
|
||
<collectorName id="F40F5339FF91FF879E803600FD520B26" box="[684,732,1869,1893]" pageId="8" pageNumber="246">Xay</collectorName>
|
||
,
|
||
<collectorName id="F40F5339FF91FF879ECB3601FCEE0B26" box="[743,864,1868,1893]" pageId="8" pageNumber="246">Ammaline</collectorName>
|
||
,
|
||
<collectorName id="F40F5339FF91FF879F473601FC310B26" box="[875,959,1868,1893]" pageId="8" pageNumber="246">Phoang</collectorName>
|
||
sone, and
|
||
<collectorName id="F40F5339FF91FF87981B3600FB5F0B26" box="[1079,1233,1868,1893]" pageId="8" pageNumber="246">K. Inkhavilay</collectorName>
|
||
colls.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="11E06564FF91FF879CEB363DFBE40BCB" box="[199,1130,1904,1929]" pageId="8" pageNumber="246" type="etymology">
|
||
<paragraph id="594536EFFF91FF879CEB363DFBE40BCB" blockId="8.[151,1437,1646,2037]" box="[199,1130,1904,1929]" pageId="8" pageNumber="246">
|
||
<emphasis id="6B8EEAFDFF91FF879CEB363DFEC00BCB" bold="true" box="[199,334,1904,1928]" pageId="8" pageNumber="246">Etymology.</emphasis>
|
||
The species is named after Daren Fleischman of Fairfield,
|
||
<collectingRegion id="9B3EF80DFF91FF879FC9363CFBAE0BCB" box="[997,1056,1905,1928]" country="United States of America" name="Iowa" pageId="8" pageNumber="246">Iowa</collectingRegion>
|
||
,
|
||
<collectingCountry id="21ED767FFF91FF879807363DFBE90BCB" box="[1067,1127,1904,1928]" name="United States of America" pageId="8" pageNumber="246">USA</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="11E06564FF91FF869CEB36DEFBBE0CBB" lastPageId="9" lastPageNumber="247" pageId="8" pageNumber="246" type="diagnosis">
|
||
<paragraph id="594536EFFF91FF869CEB36DEFBBE0CBB" blockId="8.[151,1437,1646,2037]" lastBlockId="9.[151,1437,151,1688]" lastPageId="9" lastPageNumber="247" pageId="8" pageNumber="246">
|
||
<emphasis id="6B8EEAFDFF91FF879CEB36DEFECF0BEF" bold="true" box="[199,321,1939,1964]" pageId="8" pageNumber="246">Diagnosis.</emphasis>
|
||
Secondary male pores near lateral margins of ventrum in XVIII at 7th setal lines, 0.24 circumference apart ventrally, transverse sphincter-like slit; primary male pore on slender cylindrical penis within shallow parietal invagination; paired conical genital papillae within equatorial invaginations on XVII, XIX; paired circular papillae within invaginations on 17/18, 18/19, all just medial to level of secondary male pores; spermathecal pores paired in 7/8/9 at 9th setal lines, pores large with internal grooves, 0.28–0.32 circumference apart ventrally; first dorsal pore in 5/6; muscular prostatic ducts irregularly coiled.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="11E06564FF90FF869CEB304EFC970853" pageId="9" pageNumber="247" type="description">
|
||
<paragraph id="594536EFFF90FF869CEB304EFE4E0EEB" blockId="9.[151,1437,151,1688]" pageId="9" pageNumber="247">
|
||
<emphasis id="6B8EEAFDFF90FF869CEB304EFED70D5F" bold="true" box="[199,345,259,284]" pageId="9" pageNumber="247">Description.</emphasis>
|
||
Dimensions
|
||
<quantity id="9E029B0AFF90FF869DC03049FDEE0D5E" box="[492,608,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="7.65" metricValueMax="9.5" metricValueMin="5.8" pageId="9" pageNumber="247" unit="mm" value="76.5" valueMax="95.0" valueMin="58.0">58–95mm</quantity>
|
||
, by
|
||
<quantity id="9E029B0AFF90FF869EA23049FC9B0D5F" box="[654,789,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="3.7" metricValueMin="3.3" pageId="9" pageNumber="247" unit="mm" value="3.5" valueMax="3.7" valueMin="3.3">3.3–3.7 mm</quantity>
|
||
at segment X,
|
||
<quantity id="9E029B0AFF90FF869F923049FBCB0D5F" box="[958,1093,260,285]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" metricValueMax="3.5" metricValueMin="2.9" pageId="9" pageNumber="247" unit="mm" value="3.2" valueMax="3.5" valueMin="2.9">2.9–3.5 mm</quantity>
|
||
at segment XXX,
|
||
<quantity id="9E029B0AFF90FF86993A3049FA120D5F" box="[1302,1436,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="3.5999999999999996" metricValueMax="3.9" metricValueMin="3.3" pageId="9" pageNumber="247" unit="mm" value="3.5999999999999996" valueMax="3.9" valueMin="3.3">3.3–3.9 mm</quantity>
|
||
at clitellum; body cylindrical, segments 78–101. Setae evenly spaced, numbering 37–38 at VII, 30–35 at XX, 3–9 between secondary male pores (openings of copulatory pouches), setal formula AA:AB:ZZ:YZ= 3:2:4:2 at XIII. Female pore single in XIV, on a circular porophore
|
||
<quantity id="9E029B0AFF90FF869EDA303DFC0F0DC4" box="[758,897,368,392]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="9" pageNumber="247" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 mm</quantity>
|
||
in diameter. Prostomium epilobic with tongue open, clitellum light brown, anterior dorsum light brown, post-clitellum setal rings lighter, formalin preservation. First dorsal pore at 5/6. Clitellum annular XIV–XVI, setae invisible externally. Secondary male pores near lateral margins of ventrum in XVIII at 7th setal lines, 0.24 circumference apart ventrally, transverse sphincter-like slit; primary male pores on slender cylindrical penis within shallow parietal invagination. Paired conical genital papillae within equatorial invaginations on XVII, XIX; paired circular papillae within invaginations on 17/18, 18/ 19, all just medial to level of secondary male pores. Spermathecal pores paired in 7/8/9 at 8th setal lines, pores large with internal grooves, 0.28–0.32 circumference apart ventrally. A single genital marking in 10/
|
||
<quantity id="9E029B0AFF90FF8699183321FAE10EC6" box="[1332,1391,620,645]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="9" pageNumber="247" unit="in" value="11.0">11 in</quantity>
|
||
the Hoauy Kanang individual.
|
||
</paragraph>
|
||
<paragraph id="594536EFFF90FF869CEB33F9FC970853" blockId="9.[151,1437,151,1688]" pageId="9" pageNumber="247">
|
||
Septa 5/6/7 thick, 7/8 thin,
|
||
<date id="2D44102FFF90FF869E3933F9FDD20E8F" box="[533,604,692,716]" pageId="9" pageNumber="247" value="2010-09-08">8/9/10</date>
|
||
absent,
|
||
<date id="2D44102FFF90FF869EE933F9FCDF0E8F" box="[709,849,692,716]" pageId="9" pageNumber="247" value="2014-10-11" valueMax="2014-10-13" valueMin="2014-10-11">10/11–13/14</date>
|
||
thin. Gizzard large globular in VIII–X. Intestine enlarged from XV, lymph glands not found. Typhlosole low fold to vestigial from XXVII. Intestinal caeca originating in XXVII extending anteriorly about to XXIV, simple elongate triangle. Hearts esophageal X–XIII. Male sexual system holandric, testes and funnels in ventrally joined sacs in X, XI. Seminal vesicles paired, welldeveloped in XI, XII; with small round apical dorsal lobes on pair of XII. Prostates in XVIII, extending from XVII–XXI; stout muscular ducts irregularly coiled; glandular portions consisted of 2 main lobes, each divided into 4–6 lobes. Genital papilla glands dense sessile masses, corresponding to circular papillae of 17/18, 18/19; no glands for conical papillae in XVII and XIX. Ovaries in XIII. Spermathecae two pairs in VII, IX; ampulla cordate, muscular ducts shorter than ampulla, iridescent seminal chamber ovate, with slender stalk shorter than ampulla; stalk joins duct near ampulla base, no nephridia on ducts.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="11E06564FF90FF869CEB3551FEF00ADB" pageId="9" pageNumber="247" type="discussion">
|
||
<paragraph id="594536EFFF90FF869CEB3551FEF00ADB" blockId="9.[151,1437,151,1688]" pageId="9" pageNumber="247">
|
||
<emphasis id="6B8EEAFDFF90FF869CEB3551FEB50877" bold="true" box="[199,315,1052,1076]" pageId="9" pageNumber="247">Remarks.</emphasis>
|
||
Noting that the male pores of this and the next species are in shallow parietal invaginations we need to explain why we do not place the species in
|
||
<taxonomicName id="9EFA4D6CFF90FF869E93350DFBB7081B" authority="Sims & Easton, 1972" authorityName="Sims & Easton" authorityYear="1972" box="[703,1081,1087,1112]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="247" phylum="Annelida" rank="genus">
|
||
<emphasis id="6B8EEAFDFF90FF869E93350DFCBD081B" box="[703,819,1088,1112]" italics="true" pageId="9" pageNumber="247">Metaphire</emphasis>
|
||
<bibRefCitation id="3D6B4B1EFF90FF869F113572FBB7081B" box="[829,1081,1087,1112]" pageId="9" pageNumber="247" refId="ref9412" refString="Sims, R. W. & Easton, E. G. (1972) A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. The Biological Journal of the Linnean Society, London, 4, 169 - 268." type="journal article">Sims & Easton, 1972</bibRefCitation>
|
||
</taxonomicName>
|
||
. It is simply because all other characters agree closely with other species unequivocally assignable to
|
||
<taxonomicName id="9EFA4D6CFF90FF869FE13529FBB5083F" authorityName="Hong & Inkhavilay & James" authorityYear="2018" box="[973,1083,1124,1148]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="247" phylum="Annelida" rank="genus">
|
||
<emphasis id="6B8EEAFDFF90FF869FE13529FBB5083F" box="[973,1083,1124,1148]" italics="true" pageId="9" pageNumber="247">Amynthas</emphasis>
|
||
</taxonomicName>
|
||
, and to the
|
||
<taxonomicName id="9EFA4D6CFF90FF8698E43528FADF083F" authorityName="Kinberg" authorityYear="1867" box="[1224,1361,1125,1148]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="247" phylum="Annelida" rank="species" species="aeruginosus">
|
||
<emphasis id="6B8EEAFDFF90FF8698E43528FADF083F" box="[1224,1361,1125,1148]" italics="true" pageId="9" pageNumber="247">aeruginosus</emphasis>
|
||
</taxonomicName>
|
||
group within that genus. Laotian species we report here share a number of apomorphic character states as noted in remarks above and below. Placement in
|
||
<taxonomicName id="9EFA4D6CFF90FF869E4935E1FD570887" authorityName="Sims & Easton" authorityYear="1972" box="[613,729,1196,1220]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="247" phylum="Annelida" rank="genus">
|
||
<emphasis id="6B8EEAFDFF90FF869E4935E1FD570887" box="[613,729,1196,1220]" italics="true" pageId="9" pageNumber="247">Metaphire</emphasis>
|
||
</taxonomicName>
|
||
would certainly render that genus polyphyletic and
|
||
<taxonomicName id="9EFA4D6CFF90FF86990135E1FA120887" authorityName="Hong & Inkhavilay & James" authorityYear="2018" box="[1325,1436,1196,1220]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="247" phylum="Annelida" rank="genus">
|
||
<emphasis id="6B8EEAFDFF90FF86990135E1FA120887" box="[1325,1436,1196,1220]" italics="true" pageId="9" pageNumber="247">Amynthas</emphasis>
|
||
</taxonomicName>
|
||
paraphyletic. This species is in the
|
||
<taxonomicName id="9EFA4D6CFF90FF869E1E359DFD3508A4" authorityName="Kinberg" authorityYear="1867" box="[562,699,1232,1255]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="247" phylum="Annelida" rank="species" species="aeruginosus">
|
||
<emphasis id="6B8EEAFDFF90FF869E1E359DFD3508A4" box="[562,699,1232,1255]" italics="true" pageId="9" pageNumber="247">aeruginosus</emphasis>
|
||
</taxonomicName>
|
||
-group Laotian section generally characterized by long prostatic ducts often in mirror-image coils, first dorsal pores in 5/6 or nearby, and lack of pre-clitellar genital markings. However the immature individual from Hoauy Kanang has a single pre-clitellar genital marking with an associated stalked gland, calling into question the generality of that character within the section. That worm also had slightly more setae per segment and the spermathecae were in VIII and IX, but is otherwise the same except for immaturity. Within this section of the
|
||
<taxonomicName id="9EFA4D6CFF90FF869DEE34C8FDC509DF" authorityName="Kinberg" authorityYear="1867" box="[450,587,1413,1436]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="247" phylum="Annelida" rank="species" species="aeruginosus">
|
||
<emphasis id="6B8EEAFDFF90FF869DEE34C8FDC509DF" box="[450,587,1413,1436]" italics="true" pageId="9" pageNumber="247">aeruginosus</emphasis>
|
||
</taxonomicName>
|
||
-group,
|
||
<taxonomicName id="9EFA4D6CFF90FF869E8934C9FCC909DF" authority="Hong & Inkhavilay & James, 2018" authorityName="Hong & Inkhavilay & James" authorityYear="2018" box="[677,839,1412,1436]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="247" phylum="Annelida" rank="species" species="fleischmani" status="sp. nov.">
|
||
<emphasis id="6B8EEAFDFF90FF869E8934C9FCC909DF" box="[677,839,1412,1436]" italics="true" pageId="9" pageNumber="247">A. fleischmani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="70BD5786FF90FF869F7C34C9FC2209DF" box="[848,940,1412,1436]" pageId="9" pageNumber="247" rank="species">
|
||
<emphasis id="6B8EEAFDFF90FF869F7C34C9FC2209DF" bold="true" box="[848,940,1412,1436]" pageId="9" pageNumber="247">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is the only one known to have invaginated male pores and four pairs of invaginated genital papillae in the following locations: equatorial XVII, 17/18, 18/19, equatorial XIX. The papillae are of two
|
||
<typeStatus id="8641884DFF90FF869E513480FD3209A6" box="[637,700,1485,1509]" pageId="9" pageNumber="247">types</typeStatus>
|
||
: conical without associated glandular tissue, and circular with associated sessile glands. The closest other species known is
|
||
<taxonomicName id="9EFA4D6CFF90FF869F6034BDFC6A0A4B" authority="Hong & Inkhavilay & James, 2018" authorityName="Hong & Inkhavilay & James" authorityYear="2018" box="[844,996,1520,1544]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="247" phylum="Annelida" rank="species" species="antethecus" status="sp. nov.">
|
||
<emphasis id="6B8EEAFDFF90FF869F6034BDFC6A0A4B" box="[844,996,1520,1544]" italics="true" pageId="9" pageNumber="247">A. antethecus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="70BD5786FF90FF869FC034BDFBC80A4B" box="[1004,1094,1520,1544]" pageId="9" pageNumber="247" rank="species">
|
||
<emphasis id="6B8EEAFDFF90FF869FC034BDFBC80A4B" bold="true" box="[1004,1094,1520,1544]" pageId="9" pageNumber="247">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
from the same location as the majority of the specimens of
|
||
<taxonomicName id="9EFA4D6CFF90FF869DDA3759FD150A6F" authority="Hong & Inkhavilay & James, 2018" authorityName="Hong & Inkhavilay & James" authorityYear="2018" box="[502,667,1556,1580]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="247" phylum="Annelida" rank="species" species="fleischmani" status="sp. nov.">
|
||
<emphasis id="6B8EEAFDFF90FF869DDA3759FD150A6F" box="[502,667,1556,1580]" italics="true" pageId="9" pageNumber="247">A. fleischmani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="70BD5786FF90FF869E8B3759FC8B0A6F" box="[679,773,1556,1580]" pageId="9" pageNumber="247" rank="species">
|
||
<emphasis id="6B8EEAFDFF90FF869E8B3759FC8B0A6F" bold="true" box="[679,773,1556,1580]" pageId="9" pageNumber="247">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
It differs by lacking the conical equatorial papillae, the intersegmental papillae are not invaginated on the fully clitellate specimens, there are more setae per segment, the spermathecae are in VII and VIII, the prostatic ducts are in a single loop rather than coiled, and the male pores are more widely spaced.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |