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<document ID-CLB-Dataset="9584" ID-DOI="http://dx.doi.org/10.3897/evolsyst.4.59893" ID-GBIF-Dataset="62c58258-c691-427e-9bf3-081a946616ee" ID-Pensoft-Pub="2535-0730-2-71" ID-Pensoft-UUID="C2F8869E8A18583F93087CC85413B251" ID-ZooBank="4EEF928329EF47AEA1A5ED43ECF87299" ModsDocID="2535-0730-2-71" checkinTime="1605571711922" checkinUser="pensoft" docAuthor="Pellinen, Markku J., Zahiri, Reza &amp; Sihvonen, Pasi" docDate="2020" docId="878A3589AF665E3C82FC27A3DABF80D4" docLanguage="en" docName="EvolutSyst 4(2): 71-77" docOrigin="Evolutionary Systematics 4 (2)" docSource="http://dx.doi.org/10.3897/evolsyst.4.59893" docTitle="Sacada chaehomensis Pellinen &amp; Zahiri 2020, sp. nov." docType="treatment" docUuid="D4C3AC48-60D0-4E02-9C37-FF09BA7099EA" docUuidSource="ZooBank" docVersion="5" id="C2F8869E8A18583F93087CC85413B251" lastPageNumber="71" masterDocId="C2F8869E8A18583F93087CC85413B251" masterDocTitle="A new species of Sacada Walker, 1862 from Thailand (Lepidoptera, Pyralidae, Pyralinae)" masterLastPageNumber="77" masterPageNumber="71" pageNumber="71" updateTime="1732876361434" updateUser="ExternalLinkService">
<mods:mods id="AAFD85ECDECBDCA8C15687AB213E0B46" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F2F4CAC9A76DD70E3D6ABE57412F84F6">
<mods:title id="0FA32025A1438E1C6B636BDBC09F289B">A new species of Sacada Walker, 1862 from Thailand (Lepidoptera, Pyralidae, Pyralinae)</mods:title>
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<mods:name id="6919BA074D00269C118542212ADFD063" type="personal">
<mods:role id="2D264404D7FE5F248341DBFB0786DFC0">
<mods:roleTerm id="B15FE77D3CBAC6462350817BE24E61C7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="124AFA836473B384D32A6CE9FF29396F">Pellinen, Markku J.</mods:namePart>
<mods:affiliation id="1709EDF02B44D25641E6CD0517729623">175, M 1 Muban Phichai, Mueang Lampang, Lampang, 52000, Thailand</mods:affiliation>
<mods:nameIdentifier id="400F9F5E3963E6DA9F92F3616280FB5F" type="email">markku.j.pellinen@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="6E9C4A1A9D4413FAF578581279108E4C" type="personal">
<mods:role id="C216F042F3D8532146FD5DFD5841087D">
<mods:roleTerm id="1F803B90B526FDCAA9F90D79E61FA353">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="56A883A3ECF3417DB8C14D18DEEE8BAF">Zahiri, Reza</mods:namePart>
<mods:affiliation id="FFC0DDBD403617111B3C7AC7FFBFF1E3">Universitaet Hamburg, Centrum fuer Naturkunde, Martin-Luther-King Platz 3, 20146, Hamburg, Germany &amp; Canadian Food Inspection Agency, Ottawa Plant Laboratory, Entomology Unit, Bldg. 18, 960 Carling Ave., Ottawa, Ontario, K 1 A 0 C 6 Canada</mods:affiliation>
<mods:nameIdentifier id="FB81D830A192DE4167BEA99697F6961C" type="email">reza.zahiri@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="F711CA14800F0EE0478732DB68C9C5AB" type="personal">
<mods:role id="74466DEB8CCA89499E10BB8BDEEC19F1">
<mods:roleTerm id="9E00D4536D5EEFDA6D9C48E23FE7EDC8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9FB199EAD2321B8556AA81CF59CE6AB3">Sihvonen, Pasi</mods:namePart>
<mods:affiliation id="4E56B1FC70F92DD1DB35C4B6A1514335">Finnish Museum of Natural History, Pohjoinen Rautatiekatu 13, FI- 00014, University of Helsinki, Helsinki, Finland</mods:affiliation>
<mods:nameIdentifier id="B07005B75505FAD901F34B6F6F68AC0E" type="orcid">https://orcid.org/0000-0003-2237-9325</mods:nameIdentifier>
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<mods:typeOfResource id="59BA9DE1D048861D90F131A83D0DEA41">text</mods:typeOfResource>
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<mods:title id="67A45EF80A971BF7AD7B63EE82A1468C">Evolutionary Systematics</mods:title>
</mods:titleInfo>
<mods:part id="DA82A8C1A055F82E6B92F6C2846B64D9">
<mods:date id="EEFEB73489BD9890404CB573142FDC65">2020</mods:date>
<mods:detail id="B0B2762F0D1F9A2CA3F4C1339FD0F4E2" type="volume">
<mods:number id="9925650F5727B36DB0C19FC3EF9B4F7F">4</mods:number>
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<mods:number id="F1540C87BE2BC6207C8BC8DCF1E124A2">2</mods:number>
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<mods:start id="F1C2EECF19E911D7104DDF5CB69E5279">71</mods:start>
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<mods:url id="2B384689BA84B599EE33F36C440C50C1">http://dx.doi.org/10.3897/evolsyst.4.59893</mods:url>
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<mods:classification id="03A4FB7F2D762B796F8114000AACA57C">journal article</mods:classification>
<mods:identifier id="8FC5495490E74F220C83024C43EB357D" type="DOI">http://dx.doi.org/10.3897/evolsyst.4.59893</mods:identifier>
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<treatment id="878A3589AF665E3C82FC27A3DABF80D4" ID-GBIF-Taxon="169091723" LSID="urn:lsid:zoobank.org:act:D4C3AC48-60D0-4E02-9C37-FF09BA7099EA" httpUri="http://treatment.plazi.org/id/878A3589AF665E3C82FC27A3DABF80D4" lastPageNumber="71" pageId="0" pageNumber="71" scope_family="Pyralidae" scope_order="Lepidoptera" scope_subFamily="Pyralinae">
<subSubSection id="C4E6626DA16904E37BE845EE78C78AC7" pageId="0" pageNumber="71" type="nomenclature">
<paragraph id="99895143CBE5C578AF8CC042BC1EDCD9" pageId="0" pageNumber="71">
<taxonomicName id="EC719841EE87EB20180B9168AC4704EA" LSID="878A3589-AF66-5E3C-82FC-27A3DABF80D4" authority="Pellinen &amp; Zahiri" authorityName="Pellinen &amp; Zahiri" authorityYear="2020" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada chaehomensis" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chaehomensis" status="sp. nov.">Sacada chaehomensis Pellinen &amp; Zahiri</taxonomicName>
<taxonomicNameLabel id="ED265F18A9751707EAE62DD01BD9E20A" pageId="0" pageNumber="71">sp. nov.</taxonomicNameLabel>
<figureCitation id="75EA1ADAEF65CAFA98C3A767F621DFD5" captionStart="Figures 2–4" captionStartId="F2" captionText="Figures 2 - 4. Adults of Sacada spp. with diagnostic characters indicated 2 S. dzonguensis Singh, Kirti &amp; Ranjan (male) 3 S. umtasorensis Singh, Kirti &amp; Ranjan (male) 4 a S. chaehomensis sp. nov. (male, dorsal view) 4 b S. chaehomensis sp. nov. (male, ventral view). Figs 2, 3 based on Singh et al. (2020), used with permission." httpUri="https://binary.pensoft.net/fig/475843" pageId="0" pageNumber="71">Figs 4</figureCitation>
<figureCitation id="7396A6E5C23A11D32CEF9FD0DD3160ED" captionStart="Figures 5–7" captionStartId="F3" captionText="Figures 5 - 7. Male genitalia of Sacada with diagnostic characters indicated 5 S. dzonguensis Singh, Kirti &amp; Ranjan (holotype) 5 a aedeagus 5 b genitalia 6 S. umtasorensis Singh, Kirti &amp; Ranjan (holotype) 6 a aedeagus 6 b genitalia 7 S. chaehomensis sp. nov. (holotype, slide Sihvonen 2828) 7 a aedeagus with vesica everted 7 b aedeagus 7 c genitalia. Figs 5, 6 based on Singh et al. (2020), used with permission." httpUri="https://binary.pensoft.net/fig/475844" pageId="0" pageNumber="71">, 7</figureCitation>
<figureCitation id="B7D8859A9E40C2FF10648393A6662AFB" captionStart="Figure 8" captionStartId="F4" captionText="Figure 8. Descaled male abdomen of Sacada chaehomensis sp. nov. (holotype) 8 a abdomen 8 b 8 th tergite 8 c tympanal organs (slide Sihvonen 2828)." httpUri="https://binary.pensoft.net/fig/475845" pageId="0" pageNumber="71">, 8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="6AE83D9869B7CFA791841351897B69F8" pageId="0" pageNumber="71" type="type material">
<paragraph id="ED41779981BD7C33BE4B0410FAFEA435" pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph id="F82262861E9C8F9E5CE38FDA8DFEDF60" pageId="0" pageNumber="71">
<emphasis id="7C8B0F550A1654C27C79E7FBAE932B53" bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
♂ (label data given verbatim,
<normalizedToken id="0F577E8311BA89A52C712E9E8D6290B3" originalValue="“/”">&quot;/&quot;</normalizedToken>
indicates new line,
<normalizedToken id="7C3BB6E629575D6F9E5525BF660D43DD" originalValue="“;”">&quot;;&quot;</normalizedToken>
indicates new label]: Thailand Lampang/ Chae Hom 340 M/
<geoCoordinate id="A4005B68F5489858C68F2145762E9D19" degrees="18" direction="north" minutes="43" orientation="latitude" precision="15" seconds="19" value="18.721945">18°43'19&quot;N</geoCoordinate>
,
<geoCoordinate id="45F05FE855C1F6DA1B58428F7F0F40AB" degrees="99" direction="east" minutes="33" orientation="longitude" precision="15" seconds="11" value="99.553055">99°33'11&quot;E</geoCoordinate>
/ 23.5.2020/ Leg. Pellinen, Markku [white label]; MJPT0020 [green label, indicates M. J. Pellinen DNA tissue sample]; ZMH-DNA0199 [indicates
<normalizedToken id="FEB3D8B34D4E8A391EE011139A624F4C" originalValue="Universität">Universitaet</normalizedToken>
Hamburg DNA process number]; 0146 [pink label, indicates M. J. Pellinen photograph number]; Pasi Sihvonen,/prep. number 2828 [blue label]; http://id.luomus.fi/GBT.1;
<emphasis id="C38D682C102E57B6D8A81FB82463DA01" italics="true" pageId="0" pageNumber="71">
<taxonomicName id="5F3119E3CFDF456715F382FC0F3610A3" authorityName="Walker" authorityYear="1862" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Sacada</taxonomicName>
/
<taxonomicName id="A5D64ACB3CD15F073D190452FF7D4DDE" authorityName="Pellinen &amp; Zahiri" authorityYear="2020" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="chaehomensis" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chaehomensis">chaehomensis</taxonomicName>
</emphasis>
/ Pellinen &amp; Zahiri/ HOLOTYPE [red label] (in coll. Finnish Museum of Natural History, Helsinki, Finland).
</paragraph>
</subSubSection>
<subSubSection id="A12A39A5DD2C24E9105C68E3DC76A3AE" pageId="0" pageNumber="71" type="materials_examined">
<paragraph id="64D3F69E73E20382FD824A1C98FB7519" pageId="0" pageNumber="71">Other material examined.</paragraph>
<paragraph id="5CCFD433F0E0E1EF4FF997A7566CA12F" pageId="0" pageNumber="71">Only holotype male is known.</paragraph>
</subSubSection>
<subSubSection id="CC4BD36CD540B0490DDEECE9E9DDFA24" pageId="0" pageNumber="71" type="diagnosis">
<paragraph id="31C509076FDEFC314F702FC1FE7BF7FE" pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph id="907BD97BA9451577AE142BC1137D1503" pageId="0" pageNumber="71">
<taxonomicName id="7445523A92505DD74BA1335E9CE5BBC1" authorityName="Pellinen &amp; Zahiri" authorityYear="2020" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada chaehomensis" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chaehomensis">
<emphasis id="AD45567767CCD44A7235A76840F54C44" italics="true" pageId="0" pageNumber="71">Sacada chaehomensis</emphasis>
</taxonomicName>
sp. nov. is morphologically similar to
<taxonomicName id="0EF13BAEDBD54B2C189E495FFFCCCE51" lsidName="S. dzonguensis" pageId="0" pageNumber="71" rank="species" species="dzonguensis">
<emphasis id="55A60BA14CAE0A03B18B4A9B13D33318" italics="true" pageId="0" pageNumber="71">S. dzonguensis</emphasis>
</taxonomicName>
Singh, Kirti &amp; Ranjan, 2020 and
<taxonomicName id="400DE15C660BD7657E2AB8A88BAA8FDA" lsidName="S. umtasorensis" pageId="0" pageNumber="71" rank="species" species="umtasorensis">
<emphasis id="DFBA75C49B5EEC5C3A9EDB1C71FF5E63" italics="true" pageId="0" pageNumber="71">S. umtasorensis</emphasis>
</taxonomicName>
Singh, Kirti &amp; Ranjan, 2020. Diagnostic male characters of these three species are indicated on Figs
<figureCitation id="B421795FB4C8CCDCE29F1B92141E5329" captionStart="Figures 2–4" captionStartId="F2" captionText="Figures 2 - 4. Adults of Sacada spp. with diagnostic characters indicated 2 S. dzonguensis Singh, Kirti &amp; Ranjan (male) 3 S. umtasorensis Singh, Kirti &amp; Ranjan (male) 4 a S. chaehomensis sp. nov. (male, dorsal view) 4 b S. chaehomensis sp. nov. (male, ventral view). Figs 2, 3 based on Singh et al. (2020), used with permission." httpUri="https://binary.pensoft.net/fig/475843" pageId="0" pageNumber="71">2</figureCitation>
-
<figureCitation id="DE7BFD20B0FC6323AE79690995493597" captionStart="Figures 5–7" captionStartId="F3" captionText="Figures 5 - 7. Male genitalia of Sacada with diagnostic characters indicated 5 S. dzonguensis Singh, Kirti &amp; Ranjan (holotype) 5 a aedeagus 5 b genitalia 6 S. umtasorensis Singh, Kirti &amp; Ranjan (holotype) 6 a aedeagus 6 b genitalia 7 S. chaehomensis sp. nov. (holotype, slide Sihvonen 2828) 7 a aedeagus with vesica everted 7 b aedeagus 7 c genitalia. Figs 5, 6 based on Singh et al. (2020), used with permission." httpUri="https://binary.pensoft.net/fig/475844" pageId="0" pageNumber="71">7</figureCitation>
(females of these three species are unknown). Forewing postmedial line is straight on costa and medial line is straight in
<taxonomicName id="54F90C79CA2D8E2065D6F27890755A4E" lsidName="S. chaehomensis" pageId="0" pageNumber="71" rank="species" species="chaehomensis">
<emphasis id="D5631AA94394356187C82A4EA6228C33" italics="true" pageId="0" pageNumber="71">S. chaehomensis</emphasis>
</taxonomicName>
sp. nov. (postmedial line curved and medial line angled in
<taxonomicName id="ED3288F66A10E6F9E9F46CD523B37121" lsidName="S. dzonguensis" pageId="0" pageNumber="71" rank="species" species="dzonguensis">
<emphasis id="E38E69D75449631910BC0DCAD85C09FE" italics="true" pageId="0" pageNumber="71">S. dzonguensis</emphasis>
</taxonomicName>
; postmedial line straight and medial line angled in
<taxonomicName id="62E490FFC691FBB9C127696F541DFF60" lsidName="S. umtasorensis" pageId="0" pageNumber="71" rank="species" species="umtasorensis">
<emphasis id="AF90CBE6C0701EFFAE88EC33CF2678DF" italics="true" pageId="0" pageNumber="71">S. umtasorensis</emphasis>
</taxonomicName>
). With regard to diagnostic male genitalia characters, see Figs
<figureCitation id="8E914174832ABC29C60F093F811B14F0" captionStart="Figures 5–7" captionStartId="F3" captionText="Figures 5 - 7. Male genitalia of Sacada with diagnostic characters indicated 5 S. dzonguensis Singh, Kirti &amp; Ranjan (holotype) 5 a aedeagus 5 b genitalia 6 S. umtasorensis Singh, Kirti &amp; Ranjan (holotype) 6 a aedeagus 6 b genitalia 7 S. chaehomensis sp. nov. (holotype, slide Sihvonen 2828) 7 a aedeagus with vesica everted 7 b aedeagus 7 c genitalia. Figs 5, 6 based on Singh et al. (2020), used with permission." httpUri="https://binary.pensoft.net/fig/475844" pageId="0" pageNumber="71">5-7</figureCitation>
.
</paragraph>
<caption id="49AFFC136F62835A3A3F78B61CE6D1E1" httpUri="https://binary.pensoft.net/fig/475843" pageId="0" pageNumber="71" start="Figures 2–4" startId="F2">
<paragraph id="BAAC08F2D27CF094F428EAF2F1FA1AA8" pageId="0" pageNumber="71">
<emphasis id="BB0E3D6315F82C882ACA46FF716CCBA2" bold="true" pageId="0" pageNumber="71">Figures 2-4.</emphasis>
Adults of
<taxonomicName id="3B9755866B5B9DE0CF66B6832642CF36" authorityName="Walker" authorityYear="1862" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="E7BDCBF9927705EE017C02327C640963" italics="true" pageId="0" pageNumber="71">Sacada</emphasis>
</taxonomicName>
spp. with diagnostic characters indicated
<emphasis id="79F34F18104A0A3945C1B67DF1BC425D" bold="true" pageId="0" pageNumber="71">2</emphasis>
<taxonomicName id="125A2AE8E38885BF3865BA764BD48638" lsidName="S. dzonguensis" pageId="0" pageNumber="71" rank="species" species="dzonguensis">
<emphasis id="F9431BA5C110125023603F61D3AABF8A" italics="true" pageId="0" pageNumber="71">S. dzonguensis</emphasis>
</taxonomicName>
Singh, Kirti &amp; Ranjan (male)
<emphasis id="465B3238F134916612D2AAA36C0D7AE5" bold="true" pageId="0" pageNumber="71">3</emphasis>
<taxonomicName id="F7A0E81A50A3D2AB317E89935CEE901B" lsidName="S. umtasorensis" pageId="0" pageNumber="71" rank="species" species="umtasorensis">
<emphasis id="BD9CF0D6FD43E6CC5C245A1149A6590E" italics="true" pageId="0" pageNumber="71">S. umtasorensis</emphasis>
</taxonomicName>
Singh, Kirti &amp; Ranjan (male)
<emphasis id="C3353BB4D6C06BC2367D1E3BEDF9D19C" bold="true" pageId="0" pageNumber="71">4a</emphasis>
<taxonomicName id="98CFA7AD374368D6BEFBC0F1A700BE1A" lsidName="S. chaehomensis" pageId="0" pageNumber="71" rank="species" species="chaehomensis">
<emphasis id="48AAE9998AF894B69009357570B6D865" italics="true" pageId="0" pageNumber="71">S. chaehomensis</emphasis>
</taxonomicName>
sp. nov. (male, dorsal view)
<emphasis id="D7AC502D344AEA5ED92C7A50F1677F71" bold="true" pageId="0" pageNumber="71">4b</emphasis>
<taxonomicName id="40AE2037516ED6A918473E2A04A4708D" lsidName="S. chaehomensis" pageId="0" pageNumber="71" rank="species" species="chaehomensis">
<emphasis id="2EA4CCA3CBE95E8011E9B5368A405E03" italics="true" pageId="0" pageNumber="71">S. chaehomensis</emphasis>
</taxonomicName>
sp. nov. (male, ventral view). Figs
<emphasis id="B89570A39383F8F3B6182BAC2681577E" bold="true" pageId="0" pageNumber="71">2, 3</emphasis>
based on
<bibRefCitation id="02ED9F003F6D3E8B4041AD4FE5B91C7F" DOI="https://doi.org/10.3897/zookeys.962.51194" author="Singh, N" journalOrPublisher="ZooKeys" pageId="0" pageNumber="71" pagination="139 - 163" refId="B12" refString="Singh, N, Kriti, JS, Ranjan, R, Chandra, K, Speidel, W, 2020. On the taxonomy of the genus Sacada Walker, 1862 from India, with descriptions of a new genus and two new species (Pyralinae, Pyralidae, Lepidoptera). ZooKeys 962: 139 - 163, DOI: https://doi.org/10.3897/zookeys.962.51194" title="On the taxonomy of the genus Sacada Walker, 1862 from India, with descriptions of a new genus and two new species (Pyralinae, Pyralidae, Lepidoptera)." url="https://doi.org/10.3897/zookeys.962.51194" volume="962" year="2020">Singh et al. (2020)</bibRefCitation>
, used with permission.
</paragraph>
</caption>
<caption id="9F364253AF70049AD18860EF7EE4ED68" httpUri="https://binary.pensoft.net/fig/475844" pageId="0" pageNumber="71" start="Figures 5–7" startId="F3">
<paragraph id="72AFDEF12CB06854D97F5F33F4E4C9FF" pageId="0" pageNumber="71">
<emphasis id="C1FDBC404B9EA669BD7511EA53219527" bold="true" pageId="0" pageNumber="71">Figures 5-7.</emphasis>
Male genitalia of
<taxonomicName id="6BDEAD4B60E9F7198841412303552868" authorityName="Walker" authorityYear="1862" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="DC72F6B89BCE9A471BEE434604574B12" italics="true" pageId="0" pageNumber="71">Sacada</emphasis>
</taxonomicName>
with diagnostic characters indicated
<emphasis id="DC6FB188026F769C4F5F0055BD868045" bold="true" pageId="0" pageNumber="71">5</emphasis>
<taxonomicName id="8678D1067BC347A1D019F05D0AAF8D98" lsidName="S. dzonguensis" pageId="0" pageNumber="71" rank="species" species="dzonguensis">
<emphasis id="ECC2066489321220E464030CD12801A1" italics="true" pageId="0" pageNumber="71">S. dzonguensis</emphasis>
</taxonomicName>
Singh, Kirti &amp; Ranjan (holotype)
<emphasis id="167A3E90798240568832FA7B4CE9CFF6" bold="true" pageId="0" pageNumber="71">5a</emphasis>
aedeagus
<emphasis id="49E671A1D8AF4E60190D6EF3F809C487" bold="true" pageId="0" pageNumber="71">5b</emphasis>
genitalia
<emphasis id="BCC86AA2B78911B6907DDE63F85D7AEF" bold="true" pageId="0" pageNumber="71">6</emphasis>
<taxonomicName id="B54BEC28E8BD688E7F7D20C15E5BD458" lsidName="S. umtasorensis" pageId="0" pageNumber="71" rank="species" species="umtasorensis">
<emphasis id="7B6EE171A2AF3D8BF9EE9B91549CD921" italics="true" pageId="0" pageNumber="71">S. umtasorensis</emphasis>
</taxonomicName>
Singh, Kirti &amp; Ranjan (holotype)
<emphasis id="DC168A1186E804AAF18AF62BA593F9EC" bold="true" pageId="0" pageNumber="71">6a</emphasis>
aedeagus
<emphasis id="7CFEF855E8CAAC3B5926F9DD09338C94" bold="true" pageId="0" pageNumber="71">6b</emphasis>
genitalia
<emphasis id="BE0FD26218CCE4E7D674DCDC24871468" bold="true" pageId="0" pageNumber="71">7</emphasis>
<taxonomicName id="20839A9267915E9DF06805F8633F1EDD" lsidName="S. chaehomensis" pageId="0" pageNumber="71" rank="species" species="chaehomensis">
<emphasis id="32A3B4234C53EBC0D9029B780490B3D3" italics="true" pageId="0" pageNumber="71">S. chaehomensis</emphasis>
</taxonomicName>
sp. nov. (holotype, slide Sihvonen 2828)
<emphasis id="FBA58BD3BA560E44D5533DD1DB7998EB" bold="true" pageId="0" pageNumber="71">7a</emphasis>
aedeagus with vesica everted
<emphasis id="E2C6258149AB7A0BEA69D4BAB7AA6373" bold="true" pageId="0" pageNumber="71">7b</emphasis>
aedeagus
<emphasis id="67B79049CE708642BE85CC7E3B5E6EA8" bold="true" pageId="0" pageNumber="71">7c</emphasis>
genitalia. Figs
<emphasis id="311999BF360846D6674422311312A277" bold="true" pageId="0" pageNumber="71">5, 6</emphasis>
based on
<bibRefCitation id="ED5FB6BF34C266AAC1AA42F816186766" DOI="https://doi.org/10.3897/zookeys.962.51194" author="Singh, N" journalOrPublisher="ZooKeys" pageId="0" pageNumber="71" pagination="139 - 163" refId="B12" refString="Singh, N, Kriti, JS, Ranjan, R, Chandra, K, Speidel, W, 2020. On the taxonomy of the genus Sacada Walker, 1862 from India, with descriptions of a new genus and two new species (Pyralinae, Pyralidae, Lepidoptera). ZooKeys 962: 139 - 163, DOI: https://doi.org/10.3897/zookeys.962.51194" title="On the taxonomy of the genus Sacada Walker, 1862 from India, with descriptions of a new genus and two new species (Pyralinae, Pyralidae, Lepidoptera)." url="https://doi.org/10.3897/zookeys.962.51194" volume="962" year="2020">Singh et al. (2020)</bibRefCitation>
, used with permission.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="12BABD849B4D951425310B5A5616339A" pageId="0" pageNumber="71" type="description">
<paragraph id="BFDD202F1DC76246EDD3135997690111" pageId="0" pageNumber="71">Description.</paragraph>
<paragraph id="B4839FA6F016B6FD024BA2BDC166F61A" pageId="0" pageNumber="71">
Description is based on male, female is unknown. Wingspan 40 mm. Antennae with basal two thirds bipectinate, apical third fasciculate. Head, thorax and abdomen rufous. Tegula with iridescent scales of purplish-grey hue. Labial palps short. Base of proboscis covered densely with scales. Fore- and midleg tibia with massive hair tuft, spur formula of legs 0-2-4. Forewing narrow, reddish brown, medial, postmedial and terminal lines dry straw colored (fuscous). Medial line angled, postmedial line straight, forming triangular area on midwing. Terminal line narrow, fringes concolorous with wings. Hind wing wide, grey, weakly rufous near margin, veins darker. Terminal line fuscous, fringes concolorous with wings. Forewing underside rufous grey-brown, medially darker and inner margin pale. Hindwing underside rufous grey-brown on costa, otherwise paler. Postmedial line weakly visible on both wings. Tympanal organs large, semi-circular, fornix tympani separate but connected via narrow sclerotised ridge (in Fig.
<figureCitation id="7710CFDF83B2772B3DD0DE56019EE047" captionStart="Figures 5–7" captionStartId="F3" captionText="Figures 5 - 7. Male genitalia of Sacada with diagnostic characters indicated 5 S. dzonguensis Singh, Kirti &amp; Ranjan (holotype) 5 a aedeagus 5 b genitalia 6 S. umtasorensis Singh, Kirti &amp; Ranjan (holotype) 6 a aedeagus 6 b genitalia 7 S. chaehomensis sp. nov. (holotype, slide Sihvonen 2828) 7 a aedeagus with vesica everted 7 b aedeagus 7 c genitalia. Figs 5, 6 based on Singh et al. (2020), used with permission." httpUri="https://binary.pensoft.net/fig/475844" pageId="0" pageNumber="71">7c</figureCitation>
metathorax sclerotisations and metafurca are visible between tympanal organs). Abdominal segments weakly sclerotised, 8th tergite with two posterior lobes, 8th sternite with round posterior margin.
</paragraph>
<paragraph id="62BF0FF563306A7B6F055C23D1AC2170" pageId="0" pageNumber="71">
<emphasis id="8827C997A01238E46FF551ACCB2B558B" bold="true" pageId="0" pageNumber="71">Male genitalia.</emphasis>
Uncus hood-shaped, curved, densely setose, with rounded lobes basolaterally. Gnathos sclerotised, arms forming narrow bands, arms fused medially forming upwards directed weakly dentate hook; basolateral parts curved inwards, lobe-shaped. Tegumen formed of two narrow plates. Valva simple, setose, pointing weakly outwards; costa straight. Transtilla large, sclerotised, plate-like, with distinct elongated, upwards directed projection medially, its apex blunt. Juxta broad, weakly constricted laterally, anterior margin even, posterior margin with two heavily sclerotised arms bearing few spikes. Vinculum narrow. Saccus elongated, apex round. Aedeagus stout and weakly undulating, caecum round, apical apodeme ventrally with small group of dentate spikes. Vesica large, simple, without distinct diverticula, basal part wider, apical part narrow and weakly curved; base covered with minute spikes, followed by minute sclerotisations, those become fewer towards vesica apex.
</paragraph>
</subSubSection>
<subSubSection id="0014E4BA87C324ACB53BFCEC39CE476B" pageId="0" pageNumber="71" type="etymology">
<paragraph id="8EF95D9D5CBF2E8B94C1BDA4A7D3C1A0" pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph id="3D81D47EA1437BD36EAF91FB35336810" pageId="0" pageNumber="71">The species is named after its type locality, Chae Hom, in Lampang province, Thailand.</paragraph>
</subSubSection>
<subSubSection id="D8F9BF95191129D1D56D4EA966520E18" pageId="0" pageNumber="71" type="biology_ecology">
<paragraph id="2495A134082134BECBE3F8363B11EC21" pageId="0" pageNumber="71">Biology.</paragraph>
<paragraph id="E2DC452A5C4CBC05335217EFCB87AF20" pageId="0" pageNumber="71">
The single known male was collected in May 2020 at 340 m altitude; it was attracted to light in forest with diverse tree species, interspersed with small vegetable plantations (Fig.
<figureCitation id="C4B691AE6040B8A68C5EC49C5BBC9B5F" captionStart="Figure 9" captionStartId="F5" captionText="Figure 9. Collecting site of Sacada chaehomensis sp. nov. Thailand, Lampang, Chae Hom, 8.9.2013, photo by Markku Pellinen." httpUri="https://binary.pensoft.net/fig/475846" pageId="0" pageNumber="71">9</figureCitation>
). Immature stages are unknown.
</paragraph>
<caption id="149B76659A34F879C94C98E9D0F9E37F" httpUri="https://binary.pensoft.net/fig/475845" pageId="0" pageNumber="71" start="Figure 8" startId="F4">
<paragraph id="B9D7E9914E5D2E0741465C32D42CE379" pageId="0" pageNumber="71">
<emphasis id="7184F2DC0FD8A03317CCC189060427A1" bold="true" pageId="0" pageNumber="71">Figure 8.</emphasis>
Descaled male abdomen of
<taxonomicName id="1F6FD6244A34373A05335D4BB99C113D" authorityName="Pellinen &amp; Zahiri" authorityYear="2020" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada chaehomensis" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chaehomensis">
<emphasis id="58E3B699F83B02A1F38B31D38A5704D1" italics="true" pageId="0" pageNumber="71">Sacada chaehomensis</emphasis>
</taxonomicName>
sp. nov. (holotype)
<emphasis id="2A1C896E02320EBD2DD73BB2CAC62D36" bold="true" pageId="0" pageNumber="71">8a</emphasis>
abdomen
<emphasis id="129422F46F4F4C3AF6B3919934098EC9" bold="true" pageId="0" pageNumber="71">8b</emphasis>
8th tergite
<emphasis id="BD1C4B2560ABEB20E7DC172CE92E57B7" bold="true" pageId="0" pageNumber="71">8c</emphasis>
tympanal organs (slide Sihvonen 2828).
</paragraph>
</caption>
<caption id="51C76B127CA42369D6F00BD0872E1383" httpUri="https://binary.pensoft.net/fig/475846" pageId="0" pageNumber="71" start="Figure 9" startId="F5">
<paragraph id="476BDAD53377166FEFCA0140EC9218FE" pageId="0" pageNumber="71">
<emphasis id="8047D987DDB02260FD2FA5DD79519562" bold="true" pageId="0" pageNumber="71">Figure 9.</emphasis>
Collecting site of
<taxonomicName id="44E9E2A4BF032ECA41EA4C3AD29D2F26" authorityName="Pellinen &amp; Zahiri" authorityYear="2020" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada chaehomensis" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chaehomensis">
<emphasis id="8F13D8A61571E3B511981FF944EF016E" italics="true" pageId="0" pageNumber="71">Sacada chaehomensis</emphasis>
</taxonomicName>
sp. nov. Thailand, Lampang, Chae Hom, 8.9.2013, photo by Markku Pellinen.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C50F6BF0A7D1F998ED01551CC1A3BF1D" pageId="0" pageNumber="71" type="distribution">
<paragraph id="FBAD8F5BC127774745739B31654850F9" pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph id="F2364C82EAD7D77824CBE1186B8E6824" pageId="0" pageNumber="71">The only specimen was collected in Lampang (Chae Hom) in northern Thailand.</paragraph>
</subSubSection>
<subSubSection id="7331A5AF5CB053A92432889D1CF0AB65" pageId="0" pageNumber="71" type="barcode analysis">
<paragraph id="B6826543D09668282CF16EE3B91FF519" pageId="0" pageNumber="71">Barcode analysis.</paragraph>
<paragraph id="59390D486AC95C146583AF8BAAEF6F27" pageId="0" pageNumber="71">
A 642 bp barcode of
<taxonomicName id="D54D9376592FBCE58A5F2C634EDB6313" authorityName="Pellinen &amp; Zahiri" authorityYear="2020" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada chaehomensis" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chaehomensis">
<emphasis id="92329A11CE2FA0820C18E20951008639" italics="true" pageId="0" pageNumber="71">Sacada chaehomensis</emphasis>
</taxonomicName>
sp. nov. was submitted to the identification engine on BOLD. Genetically nearest neighbors are unidentified pyralids from Malaysia, Sabah (North of Borneo) with a minimum divergence of 6.17% and another from Malaysia, North of Kuala Lumpur with 6.50%. The other pre-existing data for
<taxonomicName id="AAC308197755A14B3C195EA5D9AB1B66" authorityName="Walker" authorityYear="1862" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="C9956C28A1554CBFFE0C6A422BBDEA4B" italics="true" pageId="0" pageNumber="71">Sacada</emphasis>
</taxonomicName>
on BOLD were
<taxonomicName id="7DBD3E3F35CD5563E5D0A84E4A47BEE1" authorityName="Snellen" authorityYear="1892" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada ragonotalis" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ragonotalis">
<emphasis id="DA89E494DD9BFE91F3A8DBAF99D116CC" italics="true" pageId="0" pageNumber="71">Sacada ragonotalis</emphasis>
</taxonomicName>
(6.54%) and
<taxonomicName id="54E73522A3065B1304E4EE9C56AB1E64" authorityName="Hampson" authorityYear="1917" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada albioculalis" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="albioculalis">
<emphasis id="C7B154340310589038D4CDE3AF4A33BB" italics="true" pageId="0" pageNumber="71">Sacada albioculalis</emphasis>
</taxonomicName>
(8.93%), thus indicating that the unidentified pyralids may be
<taxonomicName id="8EEDB588C1EDEF125604D9B2B93A04E7" authorityName="Walker" authorityYear="1862" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="9A6A97413E2E365430FA37453C21BC07" italics="true" pageId="0" pageNumber="71">Sacada</emphasis>
</taxonomicName>
sp. The species morphologically most similar (i.e.,
<taxonomicName id="32A218CADB40D47900E027B569E32AB3" lsidName="S. dzonguensis" pageId="0" pageNumber="71" rank="species" species="dzonguensis">
<emphasis id="5D6900ED9B5A28EB718295461313CF7B" italics="true" pageId="0" pageNumber="71">S. dzonguensis</emphasis>
</taxonomicName>
and
<taxonomicName id="6BE030E1EB36350C5CB9721CFE80BCD7" lsidName="S. umtasorensis" pageId="0" pageNumber="71" rank="species" species="umtasorensis">
<emphasis id="99AFD55D725ECFC23FE7A438B25B7482" italics="true" pageId="0" pageNumber="71">S. umtasorensis</emphasis>
</taxonomicName>
) to the new taxon lacked representations on the BOLD DNA library. The tree topology generated by the BI analysis (Fig.
<figureCitation id="EBC288947D73318C208A6DACD3A56E55" captionStart="Figure 10" captionStartId="F6" captionText="Figure 10. The Bayesian tree from analysis of the COI 5 P sequences of Sacada chaehomensis sp. nov. and 17 public Sacada DNA sequences. The nodes are scored by posterior probabilities (PP)." httpUri="https://binary.pensoft.net/fig/475847" pageId="0" pageNumber="71">10</figureCitation>
) was very similar to the NJ-tree confirming the association (PP = 0.87) of the new species (ZMH199) with the sister pair of unidentified pyralids from Malaysia, Borneo (LPMLY526-14+ LPMLY208-14) and Malaysia, North of Kuala Lumpur (LPMAL244-14+LEPML271-13). The relationships between analyzed
<taxonomicName id="D10BC4729730731FF3EB1B255364B5D4" authorityName="Walker" authorityYear="1862" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="A2CBC8F219F50C6EA3C5FC6305BA032A" italics="true" pageId="0" pageNumber="71">Sacada</emphasis>
</taxonomicName>
must be treated with caution, because those are based on short DNA barcode sequences only, and are likely to change with addition of more molecular data and expanding taxon sampling.
</paragraph>
<caption id="23A8352015A96B4474FA946F1B95008B" httpUri="https://binary.pensoft.net/fig/475847" pageId="0" pageNumber="71" start="Figure 10" startId="F6">
<paragraph id="B514733286442F899D1783BA17CE5FA6" pageId="0" pageNumber="71">
<emphasis id="B01D8790B0B491E74DF9D286B51B1921" bold="true" pageId="0" pageNumber="71">Figure 10.</emphasis>
The Bayesian tree from analysis of the COI5P sequences of
<taxonomicName id="F782A755BA6F4BB202CA2078967241C0" authorityName="Pellinen &amp; Zahiri" authorityYear="2020" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada chaehomensis" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chaehomensis">
<emphasis id="4DBE96FCE0A0F66B9DAA889E1CE8C1A0" italics="true" pageId="0" pageNumber="71">Sacada chaehomensis</emphasis>
</taxonomicName>
sp. nov. and 17 public
<taxonomicName id="CF924AB9648DA162FED7C7B5D624FDD0" authorityName="Walker" authorityYear="1862" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="C6C5F9BF498018190A2D09D9FD40D56C" italics="true" pageId="0" pageNumber="71">Sacada</emphasis>
</taxonomicName>
DNA sequences. The nodes are scored by posterior probabilities (PP).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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