287 lines
30 KiB
XML
287 lines
30 KiB
XML
<document ID-CLB-Dataset="55738" ID-DOI="http://dx.doi.org/10.3897/zookeys.1098.73223" ID-GBIF-Dataset="8b26ea49-e662-41f7-a213-47f4b598c91c" ID-PMC="PMC9848896" ID-Pensoft-Pub="1313-2970-1098-1" ID-Pensoft-UUID="B22184E75B1A59C9B985C0DD9F092196" ID-PubMed="36761078" ID-ZooBank="B4F4033F296E43CCBE54B9413BC19268" ModsDocID="1313-2970-1098-1" checkinTime="1651618226429" checkinUser="pensoft" docAuthor="Rakotonirina, Jean Claude & Fisher, Brian L." docDate="2022" docId="42D9CC9F80945062806D04DCE1940D8F" docLanguage="en" docName="ZooKeys 1098: 1-180" docOrigin="ZooKeys 1098" docPubDate="2022-05-03" docSource="http://dx.doi.org/10.3897/zookeys.1098.73223" docTitle="Camponotus hagensii Forel 1886" docType="treatment" docVersion="5" id="B22184E75B1A59C9B985C0DD9F092196" lastPageNumber="1" masterDocId="B22184E75B1A59C9B985C0DD9F092196" masterDocTitle="Revision of the Malagasy Camponotus subgenus Myrmosaga (Hymenoptera, Formicidae) using qualitative and quantitative morphology" masterLastPageNumber="180" masterPageNumber="1" pageNumber="1" updateTime="1732660529870" updateUser="ExternalLinkService">
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<mods:title id="C1F937EBF938D6ABD0D3A5100022B5FE">Revision of the Malagasy Camponotus subgenus Myrmosaga (Hymenoptera, Formicidae) using qualitative and quantitative morphology</mods:title>
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<mods:name id="959E03F6D9C89DFB748C6511AF3FE2CD" type="personal">
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<mods:namePart id="8FA0A31F77EBD6BD4C361A9BE3432A20">Rakotonirina, Jean Claude</mods:namePart>
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<mods:affiliation id="BB6F3A4F732A23EE585E114F22B09888">Madagascar Biodiversity Center, BP 6257, Parc Botanique et Zoologique de Tsimbazaza, Antananarivo, Madagascar & Departement d'Entomologie-Culture, Elevage, Sante; Faculte des Sciences, BP 906, Universite d'Antananarivo, Antananarivo, Madagascar</mods:affiliation>
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<mods:nameIdentifier id="CF7F3272AC0E93630562E67A1EAC62D3" type="email">jcrakoto25@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="11811F0A369C49914CFC6BD5697A1DB7">Fisher, Brian L.</mods:namePart>
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<mods:affiliation id="435FB3595A9FE8C7A47619D5642C6E29">Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:title id="A4DAC1C6ED16BB616016385B99CE142A">ZooKeys</mods:title>
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<mods:date id="600D4431AA36FDF81C7D5499B451DCC8">2022</mods:date>
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<mods:number id="3B1DF636FA1DA030066998C2FE80BD7F">2022-05-03</mods:number>
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<mods:number id="98D050AC8237E330E012549DBA8B4924">1098</mods:number>
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<mods:classification id="D8132B50BDBDE39467F2225943661CFF">journal article</mods:classification>
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<mods:identifier id="03606F1F05DC64C7AE979D80524FF0E7" type="DOI">http://dx.doi.org/10.3897/zookeys.1098.73223</mods:identifier>
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<treatment id="42D9CC9F80945062806D04DCE1940D8F" ID-GBIF-Taxon="195190301" LSID="urn:lsid:plazi:treatment:42D9CC9F80945062806D04DCE1940D8F" httpUri="http://treatment.plazi.org/id/42D9CC9F80945062806D04DCE1940D8F" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
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<subSubSection id="D5B4EFD8525228FAF8955235171CA837" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="31AF8F21FBC105A4E4AB25B58FE95890" pageId="0" pageNumber="1">
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<taxonomicName id="9C4378B1A97020C8F552A7306CC87F02" ID-CoL="QDTZ" LSID="42D9CC9F-8094-5062-806D-04DCE1940D8F" authority="Forel" authorityName="Forel" authorityYear="1886" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus hagensii" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hagensii">Camponotus hagensii Forel</taxonomicName>
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</paragraph>
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<subSubSection id="4EAE1366FF9A528A079D14C7663961A1" pageId="0" pageNumber="1" type="description">
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<paragraph id="E3CDC70AA7FCB231F8B6402AE375F871" pageId="0" pageNumber="1">
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<figureCitation id="D3AF7E944C62FD518FD83298BCD47652" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Head in full-face view A C. aurosus (CASENT 0064815) B C. hagensii (CASENT 0191568)." figureDoi="10.3897/zookeys.1098.73223.figure33" httpUri="https://binary.pensoft.net/fig/678092" pageId="0" pageNumber="1">Figs 33A</figureCitation>
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<figureCitation id="5D3C52417DA5257041F68772192FEF07" captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Camponotus hagensii A lateral view B head in full-face view C dorsal view of minor worker CASENT 0191568 D distribution map." figureDoi="10.3897/zookeys.1098.73223.figure56" httpUri="https://binary.pensoft.net/fig/678115" pageId="0" pageNumber="1">, 56</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="0E9AD7471609A4B571835CF39885BFD2" pageId="0" pageNumber="1" type="reference_group">
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||
<paragraph id="BDBEDB8D0306005396429CC41399D114" pageId="0" pageNumber="1">
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||
<taxonomicName id="B40AF470A3B58A2240A265EE762C0605" authority="hagensii" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus rubripes subsp. r." order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="rubripes" subSpecies="r.">Camponotus rubripes r. hagensii</taxonomicName>
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Forel, 1886a: 158. Lectotype minor worker, by present designation, Centre de Madagascar (Hildebrandt) AntWeb CASENT0910112 (MHNG) [examined]. Paralectotypes: 2 minor workers and 3 major workers of same data as lectotype but respectively specimen coded as: FOCOL2423, FOCOL2424 (ZMHB), and CASENT0910111 (MHNG), CASENT0104632 and FOCOL2422 (ZMHB), CASENT0101494 (MNHN) [examined]. As subspecies of
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<taxonomicName id="2F3419344F8AB22059D7B4CA03A06AE9" authorityName="Rakotonirina & Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus maculatus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="maculatus">Camponotus maculatus</taxonomicName>
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, Forel, 1891: 27; of
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<taxonomicName id="9B488DA0998EF9DD42DA0CBAB855FC36" authorityName="Roger" authorityYear="1863" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus fumidus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fumidus">Camponotus fumidus</taxonomicName>
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, Santschi, 1922: 101. Raised to species by
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<bibRefCitation id="AD2286435A84255F08B767954502C850" DOI="https://doi.org/10.5962/bhl.title.10348" author="Dalla Torre, KW" journalOrPublisher="Engelmann, Leipzig" pageId="0" pageNumber="1" refId="B3" refString="Dalla Torre, KW, 1893. Catalogus Hymenopterorum hucusque descriptorum systematicus et synonymicus. Vol. 7. Formicidae (Heterogyna). W. Engelmann, Leipzig, DOI: https://doi.org/10.5962/bhl.title.10348" title="Catalogus Hymenopterorum hucusque descriptorum systematicus et synonymicus. Vol. 7. Formicidae (Heterogyna). W." url="https://doi.org/10.5962/bhl.title.10348" year="1893">Dalla Torre 1893</bibRefCitation>
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||
: 233;
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<bibRefCitation id="AEDAFFF655964CD474F3E5F0F411FA35" DOI="https://doi.org/10.5962/bhl.part.36672" author="Forel, A" journalOrPublisher="Revue Suisse de Zoologie" pageId="0" pageNumber="1" pagination="257 - 276" refId="B24" refString="Forel, A, 1914. Le genre Camponotus Mayr et les genres voisins. Revue Suisse de Zoologie 22: 257 - 276, DOI: https://doi.org/10.5962/bhl.part.36672" title="Le genre Camponotus Mayr et les genres voisins." url="https://doi.org/10.5962/bhl.part.36672" volume="22" year="1914">Forel 1914</bibRefCitation>
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||
: 267;
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<bibRefCitation id="00EA5273709BD79270F03BB0B16BE60C" DOI="https://doi.org/10.5281/zenodo.25547" author="Emery, C" journalOrPublisher="Revue Zoologique Africaine (Brussels)" pageId="0" pageNumber="1" refId="B10" refString="Emery, C, 1920b. Studi sui Camponotus. Bollettino della Societa Entomologica Italiana 52: 3-48. https://doi.org/10.5281/zenodo.25547" title="Studi sui Camponotus. Bollettino della Societa Entomologica Italiana 52: 3 - 48." url="https://doi.org/10.5281/zenodo.25547" year="1920 b">Emery 1920b</bibRefCitation>
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: 6;
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<bibRefCitation id="842EC78A3627AB4F22FA6108630EA9A0" author="Wheeler, WM" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="0" pageNumber="1" pagination="1005 - 1055" refId="B39" refString="Wheeler, WM, 1922. Ants of the American Museum Congo expedition. A contribution to the myrmecology of Africa. IX. A synonymic list of the ants of the Malagasy region. Bulletin of the American Museum of Natural History 45: 1005 - 1055" title="Ants of the American Museum Congo expedition. A contribution to the myrmecology of Africa. IX. A synonymic list of the ants of the Malagasy region." volume="45" year="1922">Wheeler 1922</bibRefCitation>
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||
: 1039;
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<bibRefCitation id="B030D957337914198EC958B1D6B08B8B" author="Emery, C" journalOrPublisher="Genera Insectorum" pageId="0" pageNumber="1" pagination="1 - 302" refId="B11" refString="Emery, C, 1925. Hymenoptera. Fam. Formicidae. Subfam. Formicinae. Genera Insectorum 183: 1 - 302" title="Hymenoptera. Fam. Formicidae. Subfam. Formicinae." volume="183" year="1925">Emery 1925</bibRefCitation>
|
||
: 92.
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||
</paragraph>
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||
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<subSubSection id="8B4061DC032918BBC19EB4745DE9AD6C" pageId="0" pageNumber="1" type="materials_examined">
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<paragraph id="5C089AFB8569DD7A6BB4D9CDAC4C0605" pageId="0" pageNumber="1">Additional material examined.</paragraph>
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<paragraph id="C4A7783F3763E8FE5CA6752037A2B149" pageId="0" pageNumber="1">
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<emphasis id="3C75E01D8283155FB19F0A6F759DBE5C" bold="true" pageId="0" pageNumber="1">
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<collectingCountry id="E9D133C90D49149C107F1BB5BFC1AA0C" name="Madagascar">Madagascar</collectingCountry>
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:
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<collectingRegion id="E8FC29927E2C94E4C06FD9C696E72F79" country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
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||
</emphasis>
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||
: Central Madagascar; Ambatomanjaka; Miarinarivo,
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<geoCoordinate id="98A30723C98116829D4C077404226D3D" degrees="18.876091" direction="south" orientation="latitude" precision="1" value="-18.876091">-18.876091</geoCoordinate>
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,
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<geoCoordinate id="4E20F3B0AA2E5E6F38120DA60360C509" degrees="46.865775" direction="east" orientation="longitude" precision="1" value="46.865776">46.865775</geoCoordinate>
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,
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<quantity id="FC17765B23C2A725C26D8C4BFBADB4A8" metricMagnitude="3" metricUnit="m" metricValue="1.343" unit="m" value="1343.0">1343 m</quantity>
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(Hildebrandt) (CAS).
|
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<emphasis id="FBFCF82441A37DEC1BDA63FEC16EBDE6" bold="true" pageId="0" pageNumber="1">
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||
<collectingRegion id="7D896A706609FEE5230EF24E30532870" country="Madagascar" name="Antsiranana">Antsiranana</collectingRegion>
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</emphasis>
|
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:
|
||
<quantity id="C11813020D26EF6E8ED4D23EFB287CA4" metricMagnitude="4" metricUnit="m" metricValue="1.22" unit="km" value="12.2">12.2 km</quantity>
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WSW Befingotra,
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<normalizedToken id="3BC5CCE5F8A623D3B4DD4F5517D712EF" originalValue="Réserve">Reserve</normalizedToken>
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Anjanaharibe-Sud, -14.75, 49.43333,
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<quantity id="393DC058AD75B9D8C1F78F14284CCCB5" metricMagnitude="3" metricUnit="m" metricValue="1.96" unit="m" value="1960.0">1960 m</quantity>
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, montane rainforest (B.L. Fisher) (CAS);
|
||
<quantity id="A22931C103C590E3E69B31534B0E1E8E" metricMagnitude="4" metricUnit="m" metricValue="1.22" unit="km" value="12.2">12.2 km</quantity>
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WSW Befingotra,
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<normalizedToken id="03723F1F5DC7C9B3DD26AD5CC9736D14" originalValue="Réserve">Reserve</normalizedToken>
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Anjanaharibe-Sud, -14.75, 49.43333,
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<quantity id="0FE54F3BD0387031E079A5EA838AC1F9" metricMagnitude="3" metricUnit="m" metricValue="1.985" unit="m" value="1985.0">1985 m</quantity>
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, montane rainforest (B.L. Fisher) (CAS);
|
||
<normalizedToken id="1DD792A183EB799805517F61FD462A72" originalValue="Nosi-Bé">Nosi-Be</normalizedToken>
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du Majunga; Nosy be; Nosibe,
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<geoCoordinate id="50130A5F3DB6A283D1AF6890F70B4390" degrees="13.315028" direction="south" orientation="latitude" precision="1" value="-13.315028">-13.315028</geoCoordinate>
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,
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<geoCoordinate id="694CE44B041C81C832EFAB0C036F52DC" degrees="48.25927" direction="east" orientation="longitude" precision="1" value="48.25927">48.25927</geoCoordinate>
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,
|
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<quantity id="62AF395A3B81EDC70544DBB668EDB769" metricMagnitude="2" metricUnit="m" metricValue="1.28" unit="m" value="128.0">128 m</quantity>
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(P.
|
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<normalizedToken id="858E2EA8FA65DB26ADB8E7B4B9B93284" originalValue="Carié">Carie</normalizedToken>
|
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) (CAS); PN Marojejy,
|
||
<quantity id="D996C62F37A70B19FE072C6393A516B2" metricMagnitude="4" metricUnit="m" metricValue="2.54" unit="km" value="25.4">25.4 km</quantity>
|
||
30° NNE Andapa,
|
||
<quantity id="9D400606E2F0C328C9F4F6B22283E948" metricMagnitude="4" metricUnit="m" metricValue="1.09" unit="km" value="10.9">10.9 km</quantity>
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311° NW Manantenina,
|
||
<geoCoordinate id="B810716E5EE44A551E5F422DE523492A" degrees="14.445" direction="south" orientation="latitude" precision="55" value="-14.445">-14.445</geoCoordinate>
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,
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<geoCoordinate id="6B51C8AC034C444C7694782EFA9EEBA4" degrees="49.735" direction="east" orientation="longitude" precision="55" value="49.735">49.735</geoCoordinate>
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,
|
||
<quantity id="D3B18B2B8D5A112F84F0668455CB29B1" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="m" value="2000.0">2000 m</quantity>
|
||
, montane shrubland (B.L. Fisher) (CAS).
|
||
<emphasis id="4D0CADB4E6D0E3F481816F3A9DB82FF5" bold="true" pageId="0" pageNumber="1">
|
||
<collectingRegion id="5C696C5111D7DA36159B287632E08AA0" country="Madagascar" name="Fianarantsoa">Fianarantsoa</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<quantity id="6AF1A439001E59A8C58EDF2E0E3E6BFE" metricMagnitude="4" metricUnit="m" metricValue="3.6" unit="km" value="36.0">36 km</quantity>
|
||
S Ambalavao, PN Andringitra, -22.2, 46.96667,
|
||
<quantity id="CDE2748C4E08D5A67F37BCF0D9A08D75" metricMagnitude="3" metricUnit="m" metricValue="1.9" unit="m" value="1900.0">1900 m</quantity>
|
||
, montane rainforest (B.L. Fisher) (CAS);
|
||
<quantity id="23B0DAE0C2B1DECB56B62A9E395CABCD" metricMagnitude="4" metricUnit="m" metricValue="3.6" unit="km" value="36.0">36 km</quantity>
|
||
S Ambalavao, PN Andringitra, -22.2, 46.96667,
|
||
<quantity id="8F9F623D7DE6C052406498D3FB2B8EA0" metricMagnitude="3" metricUnit="m" metricValue="1.975" unit="m" value="1975.0">1975 m</quantity>
|
||
, few trees, ericoid thicket (B.L. Fisher) (CAS);
|
||
<quantity id="242C0DACE7C1FFFA10ACFC30CEF49FEC" metricMagnitude="3" metricUnit="m" metricValue="8.5" unit="km" value="8.5">8.5 km</quantity>
|
||
SE Antanitotsy, Anjavidilava Forest,
|
||
<geoCoordinate id="8CEF88DCC4D08B591F2F9C82F74FE49B" degrees="22.16667" direction="south" orientation="latitude" precision="1" value="-22.16667">-22.16667</geoCoordinate>
|
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,
|
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<geoCoordinate id="3AAF83051129F7211BE81A3EA419F3CA" degrees="46.96667" direction="east" orientation="longitude" precision="1" value="46.96667">46.96667</geoCoordinate>
|
||
,
|
||
<quantity id="E1BEAB63713FE7D060A75895C0408AD6" metricMagnitude="3" metricUnit="m" metricValue="1.9899999999999998" unit="m" value="1990.0">1990 m</quantity>
|
||
, Philipia Forest (B.L. Fisher, Sylvain) (CAS); Forest
|
||
<normalizedToken id="56D65D75926B31D33436C85736D57BFF" originalValue="d’Ambalamanakana">d'Ambalamanakana</normalizedToken>
|
||
, -20.73333, 47.2 (A. Pauly) (CAS); PN Andringitra;
|
||
<quantity id="265A805C04657818C9FA4743DCFA99F9" metricMagnitude="3" metricUnit="m" metricValue="8.5" unit="km" value="8.5">8.5 km</quantity>
|
||
SE Antanitotsy,
|
||
<geoCoordinate id="6CE77A3F9671617BC2475E7EE4631192" degrees="22.16667" direction="south" orientation="latitude" precision="1" value="-22.16667">-22.16667</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="8F1921F3531BEF580BE1509632203850" degrees="46.96667" direction="east" orientation="longitude" precision="1" value="46.96667">46.96667</geoCoordinate>
|
||
,
|
||
<quantity id="692089881E33CED9064D16561E5CF8FE" metricMagnitude="3" metricUnit="m" metricValue="1.9899999999999998" unit="m" value="1990.0">1990 m</quantity>
|
||
, montane rainforest (Sylvain) (CAS); Soanierenana IV Non Protected Area,
|
||
<quantity id="F983440DCBF480CBEFF1D81D9C340F48" metricMagnitude="4" metricUnit="m" metricValue="2.522" unit="km" value="25.22">25.22 km</quantity>
|
||
SW Ambositra,
|
||
<geoCoordinate id="BBF4275DBEDADFB9382EC619E6840929" degrees="20.72389" direction="south" orientation="latitude" precision="1" value="-20.72389">-20.72389</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="637DFF8DD2B5CA1DF65DCD10F0400BF2" degrees="47.10705" direction="east" orientation="longitude" precision="1" value="47.10705">47.10705</geoCoordinate>
|
||
,
|
||
<quantity id="BF71EA4CACB755B4BA8E391F2DF343E5" metricMagnitude="3" metricUnit="m" metricValue="1.736" unit="m" value="1736.0">1736 m</quantity>
|
||
, savannah grassland (A. Ravelomanana) (CAS); Vohiparara,
|
||
<geoCoordinate id="7F4B675C75DC243E40B1DEF00B80DC1B" degrees="21.23333" direction="south" orientation="latitude" precision="1" value="-21.23333">-21.23333</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="3E25FEF53C055D0AEC4E0F202B052EAD" degrees="47.36667" direction="east" orientation="longitude" precision="1" value="47.36667">47.36667</geoCoordinate>
|
||
(A. Pauly) (CAS).
|
||
<emphasis id="74863F81880B8C8ABB1584E5C304742A" bold="true" pageId="0" pageNumber="1">
|
||
<collectingRegion id="AAEFC604E3551F12B0055CD6C8A9FBE2" country="Madagascar" name="Toliara">Toliara</collectingRegion>
|
||
</emphasis>
|
||
: Anosy Region, Anosyenne Mts,
|
||
<quantity id="A6F2578EA6F7585FFBFF3BE9E0219D1E" metricMagnitude="4" metricUnit="m" metricValue="3.25" unit="km" value="32.5">32.5 km</quantity>
|
||
NW Manantenina,
|
||
<geoCoordinate id="3D622EFA7155D92DB191102285143FDA" degrees="24.14098" direction="south" orientation="latitude" precision="1" value="-24.14098">-24.14098</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="FBE4916C272AA3A01A377AB2A218E0A3" degrees="47.03689" direction="east" orientation="longitude" precision="1" value="47.03689">47.03689</geoCoordinate>
|
||
,
|
||
<quantity id="76F2509E39C030D046A609E91033D84A" metricMagnitude="3" metricUnit="m" metricValue="1.9" unit="m" value="1900.0">1900 m</quantity>
|
||
, montane rainforest (B.L. Fisher, F.A. Esteves et al.) (CAS); near ANGAP office, PN Zombitse,
|
||
<geoCoordinate id="2F896CC05441B30854111A6DDE290F8B" degrees="22.8865" direction="south" orientation="latitude" precision="5" value="-22.8865">-22.8865</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="0B626BED22153AE15FFD23CD09DCC16D" degrees="44.69217" direction="east" orientation="longitude" precision="1" value="44.69217">44.69217</geoCoordinate>
|
||
,
|
||
<quantity id="5029C3B52FD54D5027DCE0F993D8C8CB" metricMagnitude="2" metricUnit="m" metricValue="8.4" unit="m" value="840.0">840 m</quantity>
|
||
, deciduous spiny forest (R.
|
||
<normalizedToken id="809C7B8F68FCF42CFB85E3630082FC04" originalValue="Harin’Hala">Harin'Hala</normalizedToken>
|
||
) (CAS); near road, PN Zombitse,
|
||
<geoCoordinate id="521EA23F306E9DBE76521F953A0941E3" degrees="22.8405" direction="south" orientation="latitude" precision="5" value="-22.8405">-22.8405</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="456E164BEF7B3C14C31307F861755CCB" degrees="44.73117" direction="east" orientation="longitude" precision="1" value="44.73117">44.73117</geoCoordinate>
|
||
,
|
||
<quantity id="F668AE7102A7757E5651C55C0C3D074A" metricMagnitude="2" metricUnit="m" metricValue="8.25" unit="m" value="825.0">825 m</quantity>
|
||
, spiny deciduous forest (R.
|
||
<normalizedToken id="63F221CE9CE74E4A4A86DB10F7E68429" originalValue="Harin’Hala">Harin'Hala</normalizedToken>
|
||
) (CAS).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ED836483DA0EF29C931870324AAE7DE9" pageId="0" pageNumber="1" type="diagnosis">
|
||
<paragraph id="A31C976188E71711127E8BEF587DB9E5" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||
<paragraph id="BEC89C0E393BA425662FC17ADB1DD0E1" pageId="0" pageNumber="1">In full-face view, lateral margin of head anterior to eye level diverging posteriorly and covered with erect hairs; anterior clypeal margin truncate; mesosoma in profile low and long; gastral tergites without abundant pubescence; head and gaster black, mesosoma reddish orange to brown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EEC98B7C5977F61A2B6CFF5E4D64F4F5" pageId="0" pageNumber="1" type="description">
|
||
<paragraph id="A0FB09AD12BC55D6309359723FA5FC62" pageId="0" pageNumber="1">Description.</paragraph>
|
||
<paragraph id="BFAAF996B48431EAD7DBB43A6C001161" pageId="0" pageNumber="1">
|
||
<emphasis id="F212041D602445297561D97935CA8687" bold="true" pageId="0" pageNumber="1">Minor worker.</emphasis>
|
||
In full-face view, head sides diverging towards broadly convex posterior margin or parallel anterior to eye level and rounding evenly to posterior border; eye protruding and large (EL/CS: 0.30
|
||
<normalizedToken id="F742EFEB29BB7961525B20272F47D78A" originalValue="±">+/-</normalizedToken>
|
||
0.01; 0.28-0.33), breaking lateral cephalic margin; level of its posterior margin located ca. at posterior 1/4 of head (PoOc/CL: 0.25
|
||
<normalizedToken id="AFB3E185EFA550C20B6525FBDF196138" originalValue="±">+/-</normalizedToken>
|
||
0.01; 0.23-0.25); frontal carinae wide (FR/CS: 0.34
|
||
<normalizedToken id="8877A3BF278792C0C27C433A1E2220A6" originalValue="±">+/-</normalizedToken>
|
||
0.01; 0.31-0.35), posteriorly diverging, distance between them larger than their smallest distance to eye; clypeus without well-defined anterolateral corner, anteromedian margin approximately truncate; mandible with two apical teeth distantly spaced; antennal scape relatively long (SL/CS: 1.24
|
||
<normalizedToken id="FCF9360061DBCA46410A7F1C99E79307" originalValue="±">+/-</normalizedToken>
|
||
0.06; 1.09-1.36). Promesonotum weakly convex, mesonotum with posterior portion flat immediately anterior to weakly visible metanotal groove; propodeal dorsum almost straight, its junction to declivity bluntly angulate; height of propodeal declivity 3/4 length of dorsum. Petiolar node short and high; its dorsal margin inclined posteriorly, rounding to anterior margin; height of anterior face 2/3 that of posterior; femur of hind leg rounded axially, not twisted basally.
|
||
</paragraph>
|
||
<paragraph id="F62FCE73A176EF619FE023468C029A5F" pageId="0" pageNumber="1">First and second gastral tergites without a pair of white spots; erect hairs on lateral margin of head lacking; more than six erect hairs present near posterior margin of head; antennal scape covered only with suberect hairs inclined at ca. 30° and appressed hairs; pronotum with few erect hairs, mesonotum bearing a pair of erect hairs; posterodorsal corner of propodeum with two pairs of erect hairs. Head, antenna, and gaster dark brown or reddish brown; mesosoma, petiolar node, and legs yellow to orange-yellow.</paragraph>
|
||
<paragraph id="E6EA5031413E52EB3CB8833D79F0E713" pageId="0" pageNumber="1">
|
||
<emphasis id="E6CD95E6E509A01EC9170B942C6B3558" bold="true" pageId="0" pageNumber="1">Major worker.</emphasis>
|
||
Differing from minor worker in having larger, heart-shaped head (CS: 2.37
|
||
<normalizedToken id="B0313C72B61AF41C73CE2AD647BF9AA9" originalValue="±">+/-</normalizedToken>
|
||
0.26; 2.00-2.70; CWb/CL: 0.97
|
||
<normalizedToken id="C888CF44F7C9911955788F4FEA38A7C2" originalValue="±">+/-</normalizedToken>
|
||
0.05; 0.92-1.04) with broadly concave posterior margin, anteromedian margin of clypeus slightly excised medially; apical 1/4 of antennal scape extending beyond posterior cephalic margin; robust mesosoma with distinct metanotum; propodeal dorsum convex immediately behind metanotum, <2
|
||
<normalizedToken id="BD183DCF9D5BFCC6AB63277851CAB03E" originalValue="×">x</normalizedToken>
|
||
the height of declivity surface; petiolar node much higher than long and tapering dorsally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B4C6611FFF9855DD03604830D55A6BE3" pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph id="25A782F92C1924CFA123A601A26B5F2D" pageId="0" pageNumber="1">Distribution and biology.</paragraph>
|
||
<paragraph id="5A6582CF4DC490F42F50DF41EF215741" pageId="0" pageNumber="1">
|
||
<taxonomicName id="15A8772E17487E3EE601F9D8851D791A" authorityName="Forel" authorityYear="1886" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus hagensii" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hagensii">
|
||
<emphasis id="D04D1EA1B49663A59370E368F5AA3BDB" italics="true" pageId="0" pageNumber="1">Camponotus hagensii</emphasis>
|
||
</taxonomicName>
|
||
is known mostly from montane rainforest, montane ericoid thicket, philipia forest, montane shrubland, and savannah grassland of the high plateau ranging from the PN Marojejy in the north to the Anosyenne Mountains in the south (Fig.
|
||
<figureCitation id="635860E6A8553FC643946B631EFF87D6" captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Camponotus hagensii A lateral view B head in full-face view C dorsal view of minor worker CASENT 0191568 D distribution map." figureDoi="10.3897/zookeys.1098.73223.figure56" httpUri="https://binary.pensoft.net/fig/678115" pageId="0" pageNumber="1">56D</figureCitation>
|
||
). The species has been found nesting in the ground, in root mat layers on the ground, and in rotting tree stumps. Workers have been found foraging frequently on the ground and leaf litter, rarely on low vegetation.
|
||
</paragraph>
|
||
<caption id="F6BEFA7D208F7D0FD3FF2A83DFF224EB" doi="10.3897/zookeys.1098.73223.figure56" httpUri="https://binary.pensoft.net/fig/678115" pageId="0" pageNumber="1" start="Figure 56" startId="F56">
|
||
<paragraph id="22498ADCA3F9450E21D9845723E99FAB" pageId="0" pageNumber="1">
|
||
<emphasis id="8215C248F87A7D45960E82056DAEF7D6" bold="true" pageId="0" pageNumber="1">Figure 56.</emphasis>
|
||
<taxonomicName id="07BE6C634FD19B5A1C947398CF0C4C0F" authorityName="Forel" authorityYear="1886" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus hagensii" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hagensii">
|
||
<emphasis id="A1FB239B89879E5A2EB94D889AAC806F" italics="true" pageId="0" pageNumber="1">Camponotus hagensii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D99F675E0B8E3E0A70BCBB43B2A0A16A" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
lateral view
|
||
<emphasis id="186103DA0FA150F9044E26D7CA2F605F" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
head in full-face view
|
||
<emphasis id="3BEAC10C6AFFF05EA0095C354610C888" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
dorsal view of minor worker CASENT0191568
|
||
<emphasis id="C9EFC68633ED98B2803690FFD56949D9" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
distribution map.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="987E87743EF329967F316B77621F6489" pageId="0" pageNumber="1" type="discussion">
|
||
<paragraph id="3681DE652D2DF210EBEFBEF8A44427F2" pageId="0" pageNumber="1">Discussion.</paragraph>
|
||
<paragraph id="9DFB181DA5ADEE191AA1699A458062D9" pageId="0" pageNumber="1">
|
||
<taxonomicName id="E823BBC05178781551890995CC6B83F0" authorityName="Forel" authorityYear="1886" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus hagensii" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hagensii">
|
||
<emphasis id="A115A3D9D4917E34F60FBEA48D9357F8" italics="true" pageId="0" pageNumber="1">Camponotus hagensii</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="4CDE7BB01B1756359E2395AA50711585" lsidName="C. aurosus" pageId="0" pageNumber="1" rank="species" species="aurosus">
|
||
<emphasis id="6BADFB442C232FEA311CE941BB9D05EE" italics="true" pageId="0" pageNumber="1">C. aurosus</emphasis>
|
||
</taxonomicName>
|
||
with respect to the bicolored body and the presence of erect hairs on the lateral margin of the head anterior to eye level, but in
|
||
<taxonomicName id="0A434233F816D881AC693572B970FF55" lsidName="C. aurosus" pageId="0" pageNumber="1" rank="species" species="aurosus">
|
||
<emphasis id="F4B9C1D99020DDE2F71F53106B7C6526" italics="true" pageId="0" pageNumber="1">C. aurosus</emphasis>
|
||
</taxonomicName>
|
||
the anterior clypeal margin is broadly triangular, the mesosoma is short and high in profile, and the gastral tergites are covered with abundant pubescence.
|
||
</paragraph>
|
||
<paragraph id="201FFFC7015A2EA13F3DDFEE6DFFC5E2" pageId="0" pageNumber="1">
|
||
The cluster of
|
||
<taxonomicName id="1036B60DB1BC430BE339D1BC4EE68CDE" lsidName="C. hagensii" pageId="0" pageNumber="1" rank="species" species="hagensii">
|
||
<emphasis id="2C9F32A8700B16A13BEAF66C0CA444F2" italics="true" pageId="0" pageNumber="1">C. hagensii</emphasis>
|
||
</taxonomicName>
|
||
samples presented by the dendrogram of multivariate morphometric analysis is supported by the cumulative LDA at 100% identification success. This finding is consistent with the results of the qualitative morphology-based study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |