599 lines
74 KiB
XML
599 lines
74 KiB
XML
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<mods:titleInfo id="010AB487E5505E2DACE409C7270A0397">
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<mods:title id="AF94034014CE5F0DF7171CAEC354545B">Tychobythinus inopinatus, a new troglobitic species from Sicily (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
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<mods:name id="955814811E340D7BD625CA1262D39F74" type="personal">
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<mods:namePart id="84A8D1EE3989DB8D4EF0C0960DB26833">Sabella, Giorgio</mods:namePart>
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<mods:affiliation id="EB885A327BCD5337C3D06228B3924933">Department of Biological, Geological and Environmental Sciences, section Animal Biology, University of Catania, Via Androne 81, 95124 Catania, Italy. sabellag @ unict. it; https: // orcid. org / 0000 - 0002 - 2234 - 1901</mods:affiliation>
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<mods:namePart id="5E93C9AE529D296D9730A62078E71C1D">Costanzo, Salvatore</mods:namePart>
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<mods:affiliation id="1F95139937FCC40B6125175A263212FB">Centro Universitario per la Tutela e la Gestione degli Ambienti Naturali e degli Agroecosistemi, University of Catania, Via S. Sofia 98, 95123 Catania, Italy .. scostanz @ unict. it; https: // orcid. org / 0000 - 0001 - 6103 - 6929</mods:affiliation>
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<mods:namePart id="B87534F06B4E15871FCAF527F822944C">Nicolosi, Giuseppe</mods:namePart>
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<mods:affiliation id="48CFEADAB540A59F6D21043CE2E89494">Department of Life Sciences and Systems Biology, University of Turin, Via Accademia Albertina 13, 10123, Turin, Italy & Centro Speleologico Etneo, via Valdisavoia 3, 95123 Catania, Italy</mods:affiliation>
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<mods:title id="ECE98494FA53748862F03C77CC11FCF6">Zootaxa</mods:title>
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<mods:part id="2AF412B83E1034CA7BA9C2A766A75DCF">
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<mods:date id="78AA38BABC2E2C97FC6FF658BF24754C">2020</mods:date>
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<treatment id="28348793FFD12069FF1CFCAAFE10A5D6" ID-DOI="http://doi.org/10.5281/zenodo.3851211" ID-GBIF-Taxon="164184084" ID-Zenodo-Dep="3851211" LSID="urn:lsid:zoobank.org:act:40039BFF-19D6-4966-B466-9D31DA9FEC5F" httpUri="http://treatment.plazi.org/id/28348793FFD12069FF1CFCAAFE10A5D6" lastPageId="7" lastPageNumber="280" pageId="3" pageNumber="276">
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<emphasis id="92E9EA97FFD1206DFF1CFCAAFC27A158" bold="true" box="[151,964,1258,1285]" pageId="3" pageNumber="276">
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<taxonomicName id="679D4D06FFD1206DFF1CFCAAFCBAA159" ID-CoL="8YC4P" authority="Sabella, Costanzo and Nicolosi" authorityName="Sabella, Costanzo and Nicolosi" authorityYear="2020" box="[151,857,1258,1285]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="276" phylum="Arthropoda" rank="species" species="inopinatus" status="sp. nov.">
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<emphasis id="92E9EA97FFD1206DFF1CFCAAFE31A159" bold="true" box="[151,466,1258,1285]" italics="true" pageId="3" pageNumber="276">Tychobythinus inopinatus</emphasis>
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Sabella, Costanzo and Nicolosi
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</taxonomicName>
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,
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<taxonomicNameLabel id="89DA57ECFFD1206DFCECFCABFC27A158" box="[871,964,1259,1285]" pageId="3" pageNumber="276" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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<subSubSection id="E887650EFFD1206DFF1CFD4EFC7CA111" pageId="3" pageNumber="276" type="description">
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<paragraph id="A0223685FFD1206DFF1CFD4EFEC3A174" blockId="3.[151,964,1258,1356]" box="[151,288,1294,1321]" pageId="3" pageNumber="276">
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(
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<figureCitation id="38A62A00FFD1206DFF14FD4EFEFBA174" box="[159,280,1294,1321]" captionStart-0="FIGURES 6–7" captionStart-1="FIGURES 8–11" captionStart-2="FIGURES 12–16" captionStartId-0="4.[151,264,1747,1772]" captionStartId-1="5.[151,264,1929,1954]" captionStartId-2="6.[151,264,949,974]" captionTargetBox-0="[170,1419,673,1714]" captionTargetBox-1="[222,1362,852,1901]" captionTargetBox-2="[178,1407,187,920]" captionTargetId-0="figure@4.[161,1427,669,1722]" captionTargetId-1="figure@5.[218,1369,849,1904]" captionTargetId-2="figure@6.[174,1413,181,923]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURES 6–7. Tychobythinus inopinatus sp. nov. habitus, dorsal view. 6. Holotype, male. 7. Paratype, female (Ph. G. Nicolosi). Scale: 0.1 mm." captionText-1="FIGURES 8–11. Tychobythinus inopinatus sp. nov. 8. Male head in dorsal view, schematic drawning. 9. Female head in dorsal view, schematic drawning. 10. Paratype male, head, lateral view (Ph. G. Nicolosi). 11. Paratype male, head, ventral view (Ph. G. Nicolosi). Scale: 0.1 mm." captionText-2="FIGURES 12–16. Tychobythinus inopinatus sp. nov. 12. Paratype female, left antenna. 13. Paratype female, right maxillary palpus. 14. Holotype male, aedeagus, dorsal view. 15. Paratype male, internal sac of aedeagus, dorsal view. 16. Paratype female, telisternite, dorsal view. Scale: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3833589" figureDoi-1="http://doi.org/10.5281/zenodo.3833591" figureDoi-2="http://doi.org/10.5281/zenodo.3833593" httpUri-0="https://zenodo.org/record/3833589/files/figure.png" httpUri-1="https://zenodo.org/record/3833591/files/figure.png" httpUri-2="https://zenodo.org/record/3833593/files/figure.png" pageId="3" pageNumber="276">Figs. 6–16</figureCitation>
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)
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</paragraph>
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urn:lsid:zoobank.org:act:
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<emphasis id="92E9EA97FFD1206DFF1CFD3AFEDCA1C9" bold="true" box="[151,319,1401,1428]" pageId="3" pageNumber="276">Type material</emphasis>
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.
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<materialsCitation id="10F53CD8FFD1206DFEC5FD3AFECEA1E5" ID-GBIF-Occurrence="2620591316" collectingDate="2019-10-22" collectionCode="DBUC" collectorName="G. Nicolosi" country="Italy" location="Monello Cave" municipality="Syracuse" pageId="3" pageNumber="276" specimenCount="1" specimenCount-male="1" stateProvince="Sicilia" typeStatus="holotype">
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<typeStatus id="7F268827FFD1206DFEC5FD3AFE58A1C9" box="[334,443,1402,1428]" pageId="3" pageNumber="276" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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:
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<collectingRegion id="6259F867FFD1206DFDA5FD3AFD2EA1C9" box="[558,717,1402,1428]" country="Italy" name="Sicilia" pageId="3" pageNumber="276">Sicily Region</collectingRegion>
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:
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<collectingMunicipality id="4046ACFFFFD1206DFD52FD3AFCA2A1C9" box="[729,833,1402,1428]" pageId="3" pageNumber="276">Syracuse</collectingMunicipality>
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,
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<location id="A542605EFFD1206DFCC6FD3AFC13A1C9" LSID="urn:lsid:plazi:treatment:28348793FFD12069FF1CFCAAFE10A5D6:A542605EFFD1206DFCC6FD3AFC13A1C9" box="[845,1008,1402,1428]" country="Italy" municipality="Syracuse" name="Monello Cave" pageId="3" pageNumber="276" stateProvince="Sicilia">Monello Cave</location>
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,
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<date id="D4231045FFD1206DFC77FD3AFB97A1C9" box="[1020,1140,1402,1428]" pageId="3" pageNumber="276" value="2019-10-22">
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<collectingDate id="C467E9ADFFD1206DFC77FD3AFB97A1C9" box="[1020,1140,1402,1428]" pageId="3" pageNumber="276" value="2019-10-22">22.X.2019</collectingDate>
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</date>
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, trap 17,
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,
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<collectorName id="0D685353FFD1206DFA91FD3AFA7FA1C9" box="[1306,1436,1402,1428]" pageId="3" pageNumber="276">G. Nicolosi</collectorName>
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leg. (
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<collectionCode id="C68CAE40FFD1206DFF5AFDDEFEC6A1E5" box="[209,293,1438,1464]" pageId="3" pageNumber="276">DBUC</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="10F53CD8FFD1206DFEB2FDDEFBF6A1E5" ID-GBIF-Occurrence="2620591315" box="[313,1045,1437,1465]" collectionCode="DBUC" country="Italy" location="Sicily Region" pageId="3" pageNumber="276" specimenCount="17" stateProvince="Sicilia" typeStatus="paratype">
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<typeStatus id="7F268827FFD1206DFEB2FDDEFE52A1E5" box="[313,433,1438,1464]" pageId="3" pageNumber="276" type="paratype">Paratypes</typeStatus>
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</emphasis>
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(all
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in
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):
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</emphasis>
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:
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<collectingRegion id="6259F867FFD1206DFCFEFDDEFBF6A1E5" box="[885,1045,1438,1464]" country="Italy" name="Sicilia" pageId="3" pageNumber="276">Sicily Region</collectingRegion>
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</materialsCitation>
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:
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<materialsCitation id="10F53CD8FFD1206DFBAAFDDEFABEA1E5" ID-GBIF-Occurrence="2620591303" box="[1057,1373,1438,1465]" collectionCode="DBUC" country="Italy" location="Sicily Region" pageId="3" pageNumber="276" specimenCount="1" specimenCount-female="1" stateProvince="Sicilia" typeStatus="holotype">
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<specimenCount id="B69BFD0CFFD1206DFBAAFDDEFBAEA1E4" box="[1057,1101,1438,1465]" pageId="3" pageNumber="276" type="female">1 ♀</specimenCount>
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, same data of
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<typeStatus id="7F268827FFD1206DFB7CFDDEFABEA1E5" box="[1271,1373,1438,1464]" pageId="3" pageNumber="276" type="holotype">holotype</typeStatus>
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</materialsCitation>
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;
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<materialsCitation id="10F53CD8FFD1206DFAE2FDDEFDE9A181" ID-GBIF-Occurrence="2620591301" collectionCode="DBUC" country="Italy" location="Sicily Region" pageId="3" pageNumber="276" specimenCount="1" specimenCount-female="1" stateProvince="Sicilia" typeStatus="holotype">
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<specimenCount id="B69BFD0CFFD1206DFAE2FDDEFA76A1E4" box="[1385,1429,1438,1465]" pageId="3" pageNumber="276" type="female">1 ♀</specimenCount>
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, same data of
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<typeStatus id="7F268827FFD1206DFEA7FD82FE72A181" box="[300,401,1474,1500]" pageId="3" pageNumber="276" type="holotype">holotype</typeStatus>
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,
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<collectingMethod id="79DC4E92FFD1206DFE17FD82FE10A180" box="[412,499,1474,1501]" pageId="3" pageNumber="276">but trap</collectingMethod>
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8
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</materialsCitation>
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;
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<materialsCitation id="10F53CD8FFD1206DFD9CFD82FC32A181" ID-GBIF-Occurrence="2620591307" box="[535,977,1474,1501]" collectionCode="DBUC" country="Italy" location="Sicily Region" pageId="3" pageNumber="276" specimenCount="1" specimenCount-female="1" stateProvince="Sicilia" typeStatus="holotype">
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<specimenCount id="B69BFD0CFFD1206DFD9CFD82FDA2A180" box="[535,577,1474,1501]" pageId="3" pageNumber="276" type="female">1 ♀</specimenCount>
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, same data of
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<typeStatus id="7F268827FFD1206DFD6FFD82FCAAA181" box="[740,841,1474,1500]" pageId="3" pageNumber="276" type="holotype">holotype</typeStatus>
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,
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<collectingMethod id="79DC4E92FFD1206DFCDFFD82FC48A180" box="[852,939,1474,1501]" pageId="3" pageNumber="276">but trap</collectingMethod>
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10
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||
</materialsCitation>
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;
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<materialsCitation id="10F53CD8FFD1206DFC56FD82FA74A181" ID-GBIF-Occurrence="2620591310" box="[989,1431,1474,1501]" collectionCode="DBUC" country="Italy" location="Sicily Region" pageId="3" pageNumber="276" specimenCount="1" specimenCount-male="1" stateProvince="Sicilia" typeStatus="holotype">
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<specimenCount id="B69BFD0CFFD1206DFC56FD82FBEAA180" box="[989,1033,1474,1501]" pageId="3" pageNumber="276" type="male">1 ♂</specimenCount>
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, same data of
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<typeStatus id="7F268827FFD1206DFB21FD82FAECA181" box="[1194,1295,1474,1500]" pageId="3" pageNumber="276" type="holotype">holotype</typeStatus>
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,
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<collectingMethod id="79DC4E92FFD1206DFA90FD82FA91A180" box="[1307,1394,1474,1501]" pageId="3" pageNumber="276">but trap</collectingMethod>
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12
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</materialsCitation>
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;
|
||
<materialsCitation id="10F53CD8FFD1206DFF1CFDA6FDA8A25D" ID-GBIF-Occurrence="2620591312" box="[151,587,1510,1537]" collectionCode="DBUC" country="Italy" location="Sicily Region" pageId="3" pageNumber="276" specimenCount="1" specimenCount-female="1" stateProvince="Sicilia" typeStatus="holotype">
|
||
<specimenCount id="B69BFD0CFFD1206DFF1CFDA6FF22A25C" box="[151,193,1510,1537]" pageId="3" pageNumber="276" type="female">1 ♀</specimenCount>
|
||
, same data of
|
||
<typeStatus id="7F268827FFD1206DFEEBFDA6FE26A25D" box="[352,453,1510,1536]" pageId="3" pageNumber="276" type="holotype">holotype</typeStatus>
|
||
,
|
||
<collectingMethod id="79DC4E92FFD1206DFE5BFDA6FDC5A25C" box="[464,550,1510,1537]" pageId="3" pageNumber="276">but trap</collectingMethod>
|
||
16
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="10F53CD8FFD1206DFDDDFDA6FB33A25D" ID-GBIF-Occurrence="2620591306" box="[598,1232,1510,1537]" collectingDate="2019-07-31" collectionCode="VII" collectorName="G. Nicolosi" country="Italy" location="Sicily Region" pageId="3" pageNumber="276" specimenCount="1" specimenCount-female="1" stateProvince="Sicilia" typeStatus="paratype">
|
||
<specimenCount id="B69BFD0CFFD1206DFDDDFDA6FD9CA25C" box="[598,639,1510,1537]" pageId="3" pageNumber="276" type="female">1 ♀</specimenCount>
|
||
, same locality,
|
||
<date id="D4231045FFD1206DFCA7FDA6FC54A25D" box="[812,951,1510,1536]" pageId="3" pageNumber="276" value="2019-07-31">
|
||
<collectingDate id="C467E9ADFFD1206DFCA7FDA6FC54A25D" box="[812,951,1510,1536]" pageId="3" pageNumber="276" value="2019-07-31">
|
||
31.
|
||
<collectionCode id="C68CAE40FFD1206DFCC0FDA6FC95A25D" box="[843,886,1510,1536]" pageId="3" pageNumber="276">VII</collectionCode>
|
||
.2019
|
||
</collectingDate>
|
||
</date>
|
||
, trap 10,
|
||
<collectorName id="0D685353FFD1206DFB96FDA6FB7DA25D" box="[1053,1182,1510,1536]" pageId="3" pageNumber="276">G. Nicolosi</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="10F53CD8FFD1206DFB50FDA6FEF7A279" ID-GBIF-Occurrence="2620591308" collectingDate="2019-07-31" collectionCode="VII" collectorName="G. Nicolosi" country="Italy" location="Sicily Region" pageId="3" pageNumber="276" specimenCount="2" specimenCount-female="2" stateProvince="Sicilia" typeStatus="paratype">
|
||
<specimenCount id="B69BFD0CFFD1206DFB50FDA6FAF9A25C" box="[1243,1306,1510,1537]" pageId="3" pageNumber="276" type="female">2 ♀♀</specimenCount>
|
||
, same data,
|
||
<collectingMethod id="79DC4E92FFD1206DFF1CFE4AFF0DA278" box="[151,238,1546,1573]" pageId="3" pageNumber="276">but trap</collectingMethod>
|
||
12
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="10F53CD8FFD1206DFEAAFE4AFC4CA279" ID-GBIF-Occurrence="2620591317" box="[289,943,1546,1573]" collectingDate="2019-08-22" collectionCode="VII" collectorName="G. Nicolosi" country="Italy" location="Sicily Region" pageId="3" pageNumber="276" specimenCount="1" specimenCount-female="1" stateProvince="Sicilia" typeStatus="paratype">
|
||
<specimenCount id="B69BFD0CFFD1206DFEAAFE4AFEA8A278" box="[289,331,1546,1573]" pageId="3" pageNumber="276" type="female">1 ♀</specimenCount>
|
||
, same locality,
|
||
<date id="D4231045FFD1206DFE70FE4AFD73A279" box="[507,656,1546,1572]" pageId="3" pageNumber="276" value="2019-08-22">
|
||
<collectingDate id="C467E9ADFFD1206DFE70FE4AFD73A279" box="[507,656,1546,1572]" pageId="3" pageNumber="276" value="2019-08-22">22.VIII.2019</collectingDate>
|
||
</date>
|
||
, trap 12,
|
||
<collectorName id="0D685353FFD1206DFD72FE4AFC98A279" box="[761,891,1546,1572]" pageId="3" pageNumber="276">G. Nicolosi</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="10F53CD8FFD1206DFC30FE4AFEC9A215" ID-GBIF-Occurrence="2620591305" collectingDate="2019-09-05" collectionCode="VII" collectorName="G. Nicolosi" country="Italy" location="Sicily Region" pageId="3" pageNumber="276" specimenCount="1" specimenCount-female="1" stateProvince="Sicilia" typeStatus="paratype">
|
||
<specimenCount id="B69BFD0CFFD1206DFC30FE4AFC06A278" box="[955,997,1546,1573]" pageId="3" pageNumber="276" type="female">1 ♀</specimenCount>
|
||
, same locality,
|
||
<date id="D4231045FFD1206DFB1DFE49FAF4A279" box="[1174,1303,1545,1572]" pageId="3" pageNumber="276" value="2019-09-05">
|
||
<collectingDate id="C467E9ADFFD1206DFB1DFE49FAF4A279" box="[1174,1303,1545,1572]" pageId="3" pageNumber="276" value="2019-09-05">05.IX.2019</collectingDate>
|
||
</date>
|
||
, trap 12,
|
||
<collectorName id="0D685353FFD1206DFA0AFE4AFF15A215" pageId="3" pageNumber="276">G. Nicolosi</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="10F53CD8FFD1206DFEBCFE6EFC56A215" ID-GBIF-Occurrence="2620591311" box="[311,949,1582,1609]" collectingDate="2019-09-19" collectionCode="VII" collectorName="G. Nicolosi" country="Italy" location="Sicily Region" pageId="3" pageNumber="276" specimenCount="1" specimenCount-female="1" stateProvince="Sicilia" typeStatus="paratype">
|
||
<specimenCount id="B69BFD0CFFD1206DFEBCFE6EFE81A214" box="[311,354,1582,1609]" pageId="3" pageNumber="276" type="female">1 ♀</specimenCount>
|
||
, same locality,
|
||
<date id="D4231045FFD1206DFD98FE6EFD77A215" box="[531,660,1582,1608]" pageId="3" pageNumber="276" value="2019-09-19">
|
||
<collectingDate id="C467E9ADFFD1206DFD98FE6EFD77A215" box="[531,660,1582,1608]" pageId="3" pageNumber="276" value="2019-09-19">19.IX.2019</collectingDate>
|
||
</date>
|
||
, trap 17,
|
||
<collectorName id="0D685353FFD1206DFD74FE6EFC62A215" box="[767,897,1582,1608]" pageId="3" pageNumber="276">G. Nicolosi</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="10F53CD8FFD1206DFC49FE6EFEC8A231" ID-GBIF-Occurrence="2620591309" collectingDate="2019-10-07" collectionCode="VII" collectorName="G. Nicolosi" country="Italy" location="Sicily Region" pageId="3" pageNumber="276" specimenCount="1" specimenCount-female="1" stateProvince="Sicilia" typeStatus="paratype">
|
||
<specimenCount id="B69BFD0CFFD1206DFC49FE6EFC0EA214" box="[962,1005,1582,1609]" pageId="3" pageNumber="276" type="female">1 ♀</specimenCount>
|
||
, same locality,
|
||
<date id="D4231045FFD1206DFB15FE6EFAF4A215" box="[1182,1303,1582,1608]" pageId="3" pageNumber="276" value="2019-10-07">
|
||
<collectingDate id="C467E9ADFFD1206DFB15FE6EFAF4A215" box="[1182,1303,1582,1608]" pageId="3" pageNumber="276" value="2019-10-07">07.X.2019</collectingDate>
|
||
</date>
|
||
, trap 10,
|
||
<collectorName id="0D685353FFD1206DFA0AFE6EFF15A231" pageId="3" pageNumber="276">G. Nicolosi</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="10F53CD8FFD1206DFEB3FE12FD90A231" ID-GBIF-Occurrence="2620591314" box="[312,627,1618,1645]" collectingDate="2019-10-07" collectionCode="VII" collectorName="G. Nicolosi" country="Italy" location="Sicily Region" pageId="3" pageNumber="276" specimenCount="1" specimenCount-female="1" stateProvince="Sicilia" typeStatus="paratype">
|
||
<specimenCount id="B69BFD0CFFD1206DFEB3FE12FE87A230" box="[312,356,1618,1645]" pageId="3" pageNumber="276" type="female">1 ♀</specimenCount>
|
||
, same data,
|
||
<collectingMethod id="79DC4E92FFD1206DFE7FFE12FDAFA230" box="[500,588,1618,1645]" pageId="3" pageNumber="276">but trap</collectingMethod>
|
||
17
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="10F53CD8FFD1206DFD0BFE12FAE6A231" ID-GBIF-Occurrence="2620591302" box="[640,1285,1618,1645]" collectingDate="2019-11-07" collectionCode="VII" collectorName="G. Nicolosi" country="Italy" location="Sicily Region" pageId="3" pageNumber="276" specimenCount="1" specimenCount-female="1" stateProvince="Sicilia" typeStatus="paratype">
|
||
<specimenCount id="B69BFD0CFFD1206DFD0BFE12FD4FA230" box="[640,684,1618,1645]" pageId="3" pageNumber="276" type="female">1 ♀</specimenCount>
|
||
, same locality,
|
||
<date id="D4231045FFD1206DFCD4FE12FC02A231" box="[863,993,1618,1644]" pageId="3" pageNumber="276" value="2019-11-07">
|
||
<collectingDate id="C467E9ADFFD1206DFCD4FE12FC02A231" box="[863,993,1618,1644]" pageId="3" pageNumber="276" value="2019-11-07">07.XI.2019</collectingDate>
|
||
</date>
|
||
, trap 10,
|
||
<collectorName id="0D685353FFD1206DFBC5FE12FB32A231" box="[1102,1233,1618,1644]" pageId="3" pageNumber="276">G. Nicolosi</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="10F53CD8FFD1206DFA98FE12FD7FA2CD" ID-GBIF-Occurrence="2620591313" collectingDate="2019-12-05" collectionCode="VII" collectorName="G. Nicolosi" country="Italy" location="Sicily Region" pageId="3" pageNumber="276" specimenCount="3" specimenCount-female="3" stateProvince="Sicilia" typeStatus="paratype">
|
||
<specimenCount id="B69BFD0CFFD1206DFA98FE12FAB7A230" box="[1299,1364,1618,1645]" pageId="3" pageNumber="276" type="female">3 ♀♀</specimenCount>
|
||
, same locality,
|
||
<date id="D4231045FFD1206DFF7CFE35FE60A2CD" box="[247,387,1653,1680]" pageId="3" pageNumber="276" value="2019-12-05">
|
||
<collectingDate id="C467E9ADFFD1206DFF7CFE35FE60A2CD" box="[247,387,1653,1680]" pageId="3" pageNumber="276" value="2019-12-05">05.XII.2019</collectingDate>
|
||
</date>
|
||
, trap 10,
|
||
<collectorName id="0D685353FFD1206DFE60FE36FD8FA2CD" box="[491,620,1654,1680]" pageId="3" pageNumber="276">G. Nicolosi</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E887650EFFD1206BFF4CFED9FC1BA766" lastPageId="5" lastPageNumber="278" pageId="3" pageNumber="276" type="description">
|
||
<paragraph id="A0223685FFD1206DFF4CFED9FD92A37D" blockId="3.[151,1437,1401,2041]" pageId="3" pageNumber="276">
|
||
<emphasis id="92E9EA97FFD1206DFF4CFED9FEB2A2E9" bold="true" box="[199,337,1689,1716]" pageId="3" pageNumber="276">Description</emphasis>
|
||
.
|
||
<emphasis id="92E9EA97FFD1206DFEE9FEDAFE7FA2E9" box="[354,412,1690,1716]" italics="true" pageId="3" pageNumber="276">Male</emphasis>
|
||
(
|
||
<figureCitation id="38A62A00FFD1206DFE26FEDAFE16A2E8" box="[429,501,1690,1717]" captionStart="FIGURES 6–7" captionStartId="4.[151,264,1747,1772]" captionTargetBox="[170,1419,673,1714]" captionTargetId="figure@4.[161,1427,669,1722]" captionTargetPageId="4" captionText="FIGURES 6–7. Tychobythinus inopinatus sp. nov. habitus, dorsal view. 6. Holotype, male. 7. Paratype, female (Ph. G. Nicolosi). Scale: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3833589" httpUri="https://zenodo.org/record/3833589/files/figure.png" pageId="3" pageNumber="276">Fig. 6</figureCitation>
|
||
): Length
|
||
<quantity id="67659B60FFD1206DFDEEFED9FCEFA2E8" box="[613,780,1689,1717]" metricMagnitude="-3" metricUnit="m" metricValue="1.325" metricValueMax="1.35" metricValueMin="1.3" pageId="3" pageNumber="276" unit="mm" value="1.325" valueMax="1.35" valueMin="1.3">1.30–1.35 mm</quantity>
|
||
, apterous and anophthalmous. Pale brown or brown with lighter antennae, palpi, and legs. Pubescence consists of long and flattened setae (length:
|
||
<quantity id="67659B60FFD1206DFB03FEFEFACEA284" box="[1160,1325,1726,1753]" metricMagnitude="-5" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7" pageId="3" pageNumber="276" unit="mm" value="0.075" valueMax="0.08" valueMin="0.07">0.07–0.08 mm</quantity>
|
||
) on head, pronotum, elytra, and abdomen, other suberect shorter setae (length:
|
||
<quantity id="67659B60FFD1206DFC1FFEA2FBDBA2A0" box="[916,1080,1762,1789]" metricMagnitude="-5" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="3" pageNumber="276" unit="mm" value="0.035" valueMax="0.04" valueMin="0.03">0.03–0.04 mm</quantity>
|
||
) on antennae, and legs, and thin fluff on apical segment of maxillary palpi.
|
||
</paragraph>
|
||
<paragraph id="A0223685FFD1206AFF4CFF6AFE02A6DA" blockId="3.[151,1437,1401,2041]" lastBlockId="4.[151,1437,153,647]" lastPageId="4" lastPageNumber="277" pageId="3" pageNumber="276">
|
||
Head (
|
||
<figureCitation id="38A62A00FFD1206DFE99FF6AFEACA319" box="[274,335,1834,1860]" captionStart="FIGURES 8–11" captionStartId="5.[151,264,1929,1954]" captionTargetBox="[222,1362,852,1901]" captionTargetId="figure@5.[218,1369,849,1904]" captionTargetPageId="5" captionText="FIGURES 8–11. Tychobythinus inopinatus sp. nov. 8. Male head in dorsal view, schematic drawning. 9. Female head in dorsal view, schematic drawning. 10. Paratype male, head, lateral view (Ph. G. Nicolosi). 11. Paratype male, head, ventral view (Ph. G. Nicolosi). Scale: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3833591" httpUri="https://zenodo.org/record/3833591/files/figure.png" pageId="3" pageNumber="276">Fig 8</figureCitation>
|
||
) wider (
|
||
<quantity id="67659B60FFD1206DFE3AFF6AFDFAA318" box="[433,537,1834,1861]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="276" unit="mm" value="0.3">0.30 mm</quantity>
|
||
) than long (
|
||
<quantity id="67659B60FFD1206DFD28FF6AFCE8A318" box="[675,779,1834,1861]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="276" unit="mm" value="0.24">0.24 mm</quantity>
|
||
) narrower than pronotum. Frontal lobe wider (
|
||
<quantity id="67659B60FFD1206DFA95FF69FA77A318" box="[1310,1428,1833,1861]" metricMagnitude="-4" metricUnit="m" metricValue="1.75" pageId="3" pageNumber="276" unit="mm" value="0.175">0.175 mm</quantity>
|
||
) than long (
|
||
<quantity id="67659B60FFD1206DFE9DFF0EFE63A334" box="[278,384,1870,1897]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="276" unit="mm" value="0.08">0.08 mm</quantity>
|
||
) with subparallel, protruding and sharp sides; antennal tubercle protruding. Frons between antennal tubercles with large median sulcus reaching anterior edge of vertexal foveae. Median clypeal carina welldefined, equally visible in dorsal as well as lateral view, extending to ocular region. Tempora rounded, convex occipital region traversed by median longitudinal sulcus, extending to posterior edge of vertexal foveae, these last wide and well-impressed. Gular region (
|
||
<figureCitation id="38A62A00FFD1206DFDD1FF9EFD3BA3A5" box="[602,728,2014,2040]" captionStart="FIGURES 8–11" captionStartId="5.[151,264,1929,1954]" captionTargetBox="[222,1362,852,1901]" captionTargetId="figure@5.[218,1369,849,1904]" captionTargetPageId="5" captionText="FIGURES 8–11. Tychobythinus inopinatus sp. nov. 8. Male head in dorsal view, schematic drawning. 9. Female head in dorsal view, schematic drawning. 10. Paratype male, head, lateral view (Ph. G. Nicolosi). 11. Paratype male, head, ventral view (Ph. G. Nicolosi). Scale: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3833591" httpUri="https://zenodo.org/record/3833591/files/figure.png" pageId="3" pageNumber="276">Figs 10–11</figureCitation>
|
||
) behind labium with deep and broad impression margined pos- teriorly by transverse ridge projecting and laterally recurved backwards on each side of head. This ridge with long and acute median process, process projecting ventrally and ending with a bifid tuft of setae on the tips. Base of median process enlarged posteriorly in an oval impression, at base with two long and sturdy median bristles. Antennae
|
||
<quantity id="67659B60FFD6206AFF1CF944FEDAA542" box="[151,313,260,287]" metricMagnitude="-4" metricUnit="m" metricValue="6.55" metricValueMax="6.6" metricValueMin="6.5" pageId="4" pageNumber="277" unit="mm" value="0.655" valueMax="0.66" valueMin="0.65">0.65–0.66 mm</quantity>
|
||
long, scape more than 4 times longer (
|
||
<quantity id="67659B60FFD6206AFD6EF945FC69A542" box="[741,906,261,287]" metricMagnitude="-4" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="4" pageNumber="277" unit="mm" value="0.195" valueMax="0.2" valueMin="0.19">0.19–0.20 mm</quantity>
|
||
) than wide (
|
||
<quantity id="67659B60FFD6206AFB98F944FB69A542" box="[1043,1162,260,287]" metricMagnitude="-5" metricUnit="m" metricValue="4.5" pageId="4" pageNumber="277" unit="mm" value="0.045">0.045 mm</quantity>
|
||
), narrowed and flattened with sharp medial margin on basal third, wider at middle. Pedicel ovoid, slightly asymmetric, about 1.5 times longer than wide, about as wide as scape, and wider than funicular segments. Antennomere III distinctly longer than wide and narrowed at base; antennomeres IV and V about as long as wide, antennomeres VI–VII wider than long, antennomeres VI shorter and closer than VII; antennomeres VIII wider than long, and wider than VII. Antennal club consisting of last three antennomeres which are widening progressively from IX to XI. Antennomeres IX and X distinctly wider than long, antennomere XI distinctly longer than wide and twice as long as combined length of antennomeres IX and X. Maxillary palpi (
|
||
<figureCitation id="38A62A00FFD6206AFDECFA41FD55A646" box="[615,694,513,539]" captionStart="FIGURES 12–16" captionStartId="6.[151,264,949,974]" captionTargetBox="[178,1407,187,920]" captionTargetId="figure@6.[174,1413,181,923]" captionTargetPageId="6" captionText="FIGURES 12–16. Tychobythinus inopinatus sp. nov. 12. Paratype female, left antenna. 13. Paratype female, right maxillary palpus. 14. Holotype male, aedeagus, dorsal view. 15. Paratype male, internal sac of aedeagus, dorsal view. 16. Paratype female, telisternite, dorsal view. Scale: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3833593" httpUri="https://zenodo.org/record/3833593/files/figure.png" pageId="4" pageNumber="277">Fig. 13</figureCitation>
|
||
) with palpomere IIs elongate and gradually expanded from base to apex, their surface covered by 8-10 tubercles. Palpomeres III slightly longer than wide, surface with 4–6 tubercles; last palpomere about 5 times as long (
|
||
<quantity id="67659B60FFD6206AFDB4FA09FD45A63E" box="[575,678,585,611]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="4" pageNumber="277" unit="mm" value="0.29">0.29 mm</quantity>
|
||
) as wide (
|
||
<quantity id="67659B60FFD6206AFC93FA08FC73A63E" box="[792,912,584,611]" metricMagnitude="-5" metricUnit="m" metricValue="5.5" pageId="4" pageNumber="277" unit="mm" value="0.055">0.055 mm</quantity>
|
||
), widest at basal third, lateral margin distinctly curved and sinuate at middle.
|
||
</paragraph>
|
||
<caption id="F4E2660DFFD6206AFF1CFE93FE89A352" ID-DOI="http://doi.org/10.5281/zenodo.3833589" ID-Zenodo-Dep="3833589" httpUri="https://zenodo.org/record/3833589/files/figure.png" pageId="4" pageNumber="277" startId="4.[151,264,1747,1772]" targetBox="[170,1419,673,1714]" targetPageId="4">
|
||
<paragraph id="A0223685FFD6206AFF1CFE93FE89A352" blockId="4.[151,1437,1747,1808]" pageId="4" pageNumber="277">
|
||
<emphasis id="92E9EA97FFD6206AFF1CFE93FED6A2B1" bold="true" box="[151,309,1747,1772]" pageId="4" pageNumber="277">FIGURES 6–7</emphasis>
|
||
.
|
||
<taxonomicName id="679D4D06FFD6206AFEC8FE94FDA4A2B1" authority="Sabella & Costanzo & Nicolosi, 2020" authorityName="Sabella & Costanzo & Nicolosi" authorityYear="2020" box="[323,583,1748,1772]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="inopinatus" status="sp. nov.">
|
||
<emphasis id="92E9EA97FFD6206AFEC8FE94FDA4A2B1" box="[323,583,1748,1772]" italics="true" pageId="4" pageNumber="277">Tychobythinus inopinatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="92E9EA97FFD6206AFDC5FE94FD7CA2B1" bold="true" box="[590,671,1748,1772]" pageId="4" pageNumber="277">
|
||
<taxonomicNameLabel id="89DA57ECFFD6206AFDC5FE94FD7CA2B1" box="[590,671,1748,1772]" pageId="4" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
habitus, dorsal view. 6. Holotype, male. 7. Paratype, female (Ph. G. Nicolosi). Scale: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A0223685FFD6206BFF4CFF7FFC30A48A" blockId="4.[151,1437,1854,2025]" lastBlockId="5.[151,1437,153,827]" lastPageId="5" lastPageNumber="278" pageId="4" pageNumber="277">
|
||
Pronotum wider (
|
||
<quantity id="67659B60FFD6206AFE1BFF7EFE1BA304" box="[400,504,1854,1881]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="4" pageNumber="277" unit="mm" value="0.31">0.31 mm</quantity>
|
||
) than long (
|
||
<quantity id="67659B60FFD6206AFD0EFF7EFD1DA304" box="[645,766,1854,1881]" metricMagnitude="-4" metricUnit="m" metricValue="2.9499999999999997" pageId="4" pageNumber="277" unit="mm" value="0.295">0.295 mm</quantity>
|
||
), widest near middle, anteriorly narrowed with convergent sides, posteriorly very slightly narrowed with rounded and subparallel sides very flattened laterally. Dorsal surface shiny with some sparse and faint punctures. Two well-impressed antebasal lateral foveae linked by wide antebasal sulcus. Tegument between pronotal posterior margin and antebasal sulcus rough, makeing it difficult to see small median antebasal fovea. Metaventrite distinctly raised at middle, its surface with dense and large punctures, with median sulcus beginning from its posterior margin and extending to just posterior to mesocoxal cavities. Base of mesocoxal cavities with pubescent lateral mesosternal fovea on each side.
|
||
</paragraph>
|
||
<paragraph id="A0223685FFD7206BFF4CF8A1FD86A53A" blockId="5.[151,1437,153,827]" pageId="5" pageNumber="278">
|
||
Elytra distinctly wider (
|
||
<quantity id="67659B60FFD7206BFE58F8A0FDD9A4A6" box="[467,570,224,251]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="5" pageNumber="278" unit="mm" value="0.57">0.57 mm</quantity>
|
||
) than long (
|
||
<quantity id="67659B60FFD7206BFD48F8A0FCCDA4A6" box="[707,814,224,251]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="5" pageNumber="278" unit="mm" value="0.52">0.52 mm</quantity>
|
||
), convex, sides slightly rounded from base to the apex, widest near middle. Humeral calli strongly reduced. Dorsal surface shiny with only some superficial punctures. Each elytron with two basal foveae, subhumeral fovea well-defined. Both marginal and sutural striae reaching to about elytral apices, discal striae lacking.
|
||
</paragraph>
|
||
<paragraph id="A0223685FFD7206BFF4CF931FC6FA5D6" blockId="5.[151,1437,153,827]" box="[199,908,369,395]" pageId="5" pageNumber="278">Abdomen normally shaped without species-specific characters.</paragraph>
|
||
<paragraph id="A0223685FFD7206BFF4CF9D5FE76A646" blockId="5.[151,1437,153,827]" pageId="5" pageNumber="278">
|
||
Legs relatively long and thin. Protrochanters with 1–2 tubercles on ventral surface, protibiae slightly flattened and sinuate at distal third, protarsomere II slightly dilated. Meso- and metatrochanters simple; femora simple, mesotibiae slightly enlarged and sinuate for distal third, length of metatibiae:
|
||
<quantity id="67659B60FFD7206BFC7CF99CFB7DA5AA" box="[1015,1182,476,503]" metricMagnitude="-4" metricUnit="m" metricValue="4.55" metricValueMax="4.6" metricValueMin="4.5" pageId="5" pageNumber="278" unit="mm" value="0.455" valueMax="0.46" valueMin="0.45">0.45–0.46 mm</quantity>
|
||
, slightly enlarged and sinuate for distal third.
|
||
</paragraph>
|
||
<paragraph id="A0223685FFD7206BFF4CFA65FE03A6DA" blockId="5.[151,1437,153,827]" pageId="5" pageNumber="278">
|
||
Aedeagus (
|
||
<figureCitation id="38A62A00FFD7206BFECFFA65FE77A662" box="[324,404,549,575]" captionStart="FIGURES 12–16" captionStartId="6.[151,264,949,974]" captionTargetBox="[178,1407,187,920]" captionTargetId="figure@6.[174,1413,181,923]" captionTargetPageId="6" captionText="FIGURES 12–16. Tychobythinus inopinatus sp. nov. 12. Paratype female, left antenna. 13. Paratype female, right maxillary palpus. 14. Holotype male, aedeagus, dorsal view. 15. Paratype male, internal sac of aedeagus, dorsal view. 16. Paratype female, telisternite, dorsal view. Scale: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3833593" httpUri="https://zenodo.org/record/3833593/files/figure.png" pageId="5" pageNumber="278">Fig. 14</figureCitation>
|
||
)
|
||
<quantity id="67659B60FFD7206BFE29FA64FDB1A662" box="[418,594,548,575]" metricMagnitude="-4" metricUnit="m" metricValue="3.275" metricValueMax="3.3" metricValueMin="3.25" pageId="5" pageNumber="278" unit="mm" value="0.3275" valueMax="0.33" valueMin="0.325">0.325–0.33 mm</quantity>
|
||
long, ovoid with relatively short parameres, parameres convergent and with narrow apex, each bearing two subapical setae. Internal sac (
|
||
<figureCitation id="38A62A00FFD7206BFCC0FA09FC2FA63E" box="[843,972,584,611]" captionStart="FIGURES 12–16" captionStartId="6.[151,264,949,974]" captionTargetBox="[178,1407,187,920]" captionTargetId="figure@6.[174,1413,181,923]" captionTargetPageId="6" captionText="FIGURES 12–16. Tychobythinus inopinatus sp. nov. 12. Paratype female, left antenna. 13. Paratype female, right maxillary palpus. 14. Holotype male, aedeagus, dorsal view. 15. Paratype male, internal sac of aedeagus, dorsal view. 16. Paratype female, telisternite, dorsal view. Scale: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3833593" httpUri="https://zenodo.org/record/3833593/files/figure.png" pageId="5" pageNumber="278">Figs 14–15</figureCitation>
|
||
) with two long apophyses converging to apices with numerous spines.
|
||
</paragraph>
|
||
<paragraph id="A0223685FFD7206BFF4CFAD1FC1BA766" blockId="5.[151,1437,153,827]" pageId="5" pageNumber="278">
|
||
<emphasis id="92E9EA97FFD7206BFF4CFAD1FEF8A6F6" box="[199,283,657,683]" italics="true" pageId="5" pageNumber="278">Female</emphasis>
|
||
(
|
||
<figureCitation id="38A62A00FFD7206BFEA0FAD1FE93A6F6" box="[299,368,657,683]" captionStart="FIGURES 6–7" captionStartId="4.[151,264,1747,1772]" captionTargetBox="[170,1419,673,1714]" captionTargetId="figure@4.[161,1427,669,1722]" captionTargetPageId="4" captionText="FIGURES 6–7. Tychobythinus inopinatus sp. nov. habitus, dorsal view. 6. Holotype, male. 7. Paratype, female (Ph. G. Nicolosi). Scale: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3833589" httpUri="https://zenodo.org/record/3833589/files/figure.png" pageId="5" pageNumber="278">Fig. 7</figureCitation>
|
||
): Similar to male, length
|
||
<quantity id="67659B60FFD7206BFD06FAD0FCD1A6F6" box="[653,818,656,683]" metricMagnitude="-3" metricUnit="m" metricValue="1.325" metricValueMax="1.35" metricValueMin="1.3" pageId="5" pageNumber="278" unit="mm" value="1.325" valueMax="1.35" valueMin="1.3">1.30–1.35 mm</quantity>
|
||
, head (
|
||
<figureCitation id="38A62A00FFD7206BFC09FAD1FC26A6F6" box="[898,965,657,683]" captionStart="FIGURES 8–11" captionStartId="5.[151,264,1929,1954]" captionTargetBox="[222,1362,852,1901]" captionTargetId="figure@5.[218,1369,849,1904]" captionTargetPageId="5" captionText="FIGURES 8–11. Tychobythinus inopinatus sp. nov. 8. Male head in dorsal view, schematic drawning. 9. Female head in dorsal view, schematic drawning. 10. Paratype male, head, lateral view (Ph. G. Nicolosi). 11. Paratype male, head, ventral view (Ph. G. Nicolosi). Scale: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3833591" httpUri="https://zenodo.org/record/3833591/files/figure.png" pageId="5" pageNumber="278">Fig. 9</figureCitation>
|
||
) slightly wider (
|
||
<quantity id="67659B60FFD7206BFBF5FAD0FAC7A6F6" box="[1150,1316,656,683]" metricMagnitude="-4" metricUnit="m" metricValue="2.55" metricValueMax="2.6" metricValueMin="2.5" pageId="5" pageNumber="278" unit="mm" value="0.255" valueMax="0.26" valueMin="0.25">0.25–0.26 mm</quantity>
|
||
) than long (
|
||
<quantity id="67659B60FFD7206BFF14FAF5FEA9A692" box="[159,330,693,719]" metricMagnitude="-4" metricUnit="m" metricValue="2.3499999999999996" metricValueMax="2.4" metricValueMin="2.3" pageId="5" pageNumber="278" unit="mm" value="0.235" valueMax="0.24" valueMin="0.23">0.23–0.24 mm</quantity>
|
||
), gular region unmodified, slightly convex. Antennae (
|
||
<figureCitation id="38A62A00FFD7206BFC4BFAF5FBF7A692" box="[960,1044,693,719]" captionStart="FIGURES 12–16" captionStartId="6.[151,264,949,974]" captionTargetBox="[178,1407,187,920]" captionTargetId="figure@6.[174,1413,181,923]" captionTargetPageId="6" captionText="FIGURES 12–16. Tychobythinus inopinatus sp. nov. 12. Paratype female, left antenna. 13. Paratype female, right maxillary palpus. 14. Holotype male, aedeagus, dorsal view. 15. Paratype male, internal sac of aedeagus, dorsal view. 16. Paratype female, telisternite, dorsal view. Scale: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3833593" httpUri="https://zenodo.org/record/3833593/files/figure.png" pageId="5" pageNumber="278">Fig. 12</figureCitation>
|
||
) a little shorter (length:
|
||
<quantity id="67659B60FFD7206BFABAFAF5FF27A6AE" metricMagnitude="-4" metricUnit="m" metricValue="6.35" metricValueMax="6.4" metricValueMin="6.3" pageId="5" pageNumber="278" unit="mm" value="0.635" valueMax="0.64" valueMin="0.63">0.63-0.64 mm</quantity>
|
||
) and with sligthly thinner antennomeres than in male; surface of palpomere II covered by 10–14 tubercles, anterior sides of pronotum narrower than in male. Metaventrite lacking median impression; legs with protibiae, tarsomere II of protarsi, and metatibiae unmodified. Telisternite as in
|
||
<figureCitation id="38A62A00FFD7206BFC2AFB61FC17A766" box="[929,1012,801,827]" captionStart="FIGURES 12–16" captionStartId="6.[151,264,949,974]" captionTargetBox="[178,1407,187,920]" captionTargetId="figure@6.[174,1413,181,923]" captionTargetPageId="6" captionText="FIGURES 12–16. Tychobythinus inopinatus sp. nov. 12. Paratype female, left antenna. 13. Paratype female, right maxillary palpus. 14. Holotype male, aedeagus, dorsal view. 15. Paratype male, internal sac of aedeagus, dorsal view. 16. Paratype female, telisternite, dorsal view. Scale: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3833593" httpUri="https://zenodo.org/record/3833593/files/figure.png" pageId="5" pageNumber="278">Fig. 16</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="F4E2660DFFD7206BFF1CFFC9FE59A3B4" ID-DOI="http://doi.org/10.5281/zenodo.3833591" ID-Zenodo-Dep="3833591" httpUri="https://zenodo.org/record/3833591/files/figure.png" pageId="5" pageNumber="278" startId="5.[151,264,1929,1954]" targetBox="[222,1362,852,1901]" targetPageId="5">
|
||
<paragraph id="A0223685FFD7206BFF1CFFC9FE59A3B4" blockId="5.[151,1437,1929,2026]" pageId="5" pageNumber="278">
|
||
<emphasis id="92E9EA97FFD7206BFF1CFFC9FEDCA3FF" bold="true" box="[151,319,1929,1954]" pageId="5" pageNumber="278">FIGURES 8–11</emphasis>
|
||
.
|
||
<taxonomicName id="679D4D06FFD7206BFEC0FFCAFDADA3FF" authority="Sabella & Costanzo & Nicolosi, 2020" authorityName="Sabella & Costanzo & Nicolosi" authorityYear="2020" box="[331,590,1930,1954]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="inopinatus" status="sp. nov.">
|
||
<emphasis id="92E9EA97FFD7206BFEC0FFCAFDADA3FF" box="[331,590,1930,1954]" italics="true" pageId="5" pageNumber="278">Tychobythinus inopinatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="92E9EA97FFD7206BFDDFFFCAFD46A3FF" bold="true" box="[596,677,1930,1954]" pageId="5" pageNumber="278">
|
||
<taxonomicNameLabel id="89DA57ECFFD7206BFDDFFFCAFD46A3FF" box="[596,677,1930,1954]" pageId="5" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
8. Male head in dorsal view, schematic drawning. 9. Female head in dorsal view, schematic drawning. 10. Paratype male, head, lateral view (Ph. G. Nicolosi). 11. Paratype male, head, ventral view (Ph. G. Nicolosi). Scale: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="F4E2660DFFD42068FF1CFBF5FDCBA048" ID-DOI="http://doi.org/10.5281/zenodo.3833593" ID-Zenodo-Dep="3833593" httpUri="https://zenodo.org/record/3833593/files/figure.png" pageId="6" pageNumber="279" startId="6.[151,264,949,974]" targetBox="[178,1407,187,920]" targetPageId="6">
|
||
<paragraph id="A0223685FFD42068FF1CFBF5FDCBA048" blockId="6.[151,1437,949,1046]" pageId="6" pageNumber="279">
|
||
<emphasis id="92E9EA97FFD42068FF1CFBF5FEACA793" bold="true" box="[151,335,949,974]" pageId="6" pageNumber="279">FIGURES 12–16</emphasis>
|
||
.
|
||
<taxonomicName id="679D4D06FFD42068FED5FBF5FD81A790" authority="Sabella & Costanzo & Nicolosi, 2020" authorityName="Sabella & Costanzo & Nicolosi" authorityYear="2020" box="[350,610,949,973]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="279" phylum="Arthropoda" rank="species" species="inopinatus" status="sp. nov.">
|
||
<emphasis id="92E9EA97FFD42068FED5FBF5FD81A790" box="[350,610,949,973]" italics="true" pageId="6" pageNumber="279">Tychobythinus inopinatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="89DA57ECFFD42068FDE1FBF6FD5EA793" box="[618,701,950,974]" pageId="6" pageNumber="279" rank="species">
|
||
<emphasis id="92E9EA97FFD42068FDE1FBF6FD5EA793" bold="true" box="[618,701,950,974]" pageId="6" pageNumber="279">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
12. Paratype female, left antenna. 13. Paratype female, right maxillary palpus. 14. Holotype male, aedeagus, dorsal view. 15. Paratype male, internal sac of aedeagus, dorsal view. 16. Paratype female, telisternite, dorsal view. Scale: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="E887650EFFD42069FF4CFC03FE10A5D6" lastPageId="7" lastPageNumber="280" pageId="6" pageNumber="279" type="discussion">
|
||
<paragraph id="A0223685FFD42068FF4CFC03FC4FA123" blockId="6.[151,1437,1091,2018]" pageId="6" pageNumber="279">
|
||
<emphasis id="92E9EA97FFD42068FF4CFC03FEAFA003" bold="true" box="[199,332,1091,1118]" pageId="6" pageNumber="279">Discussion.</emphasis>
|
||
<taxonomicName id="679D4D06FFD42068FEDDFC04FD95A003" authority="Sabella & Costanzo & Nicolosi, 2020" authorityName="Sabella & Costanzo & Nicolosi" authorityYear="2020" box="[342,630,1092,1118]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="279" phylum="Arthropoda" rank="species" species="inopinatus" status="sp. nov.">
|
||
<emphasis id="92E9EA97FFD42068FEDDFC04FD95A003" box="[342,630,1092,1118]" italics="true" pageId="6" pageNumber="279">Tychobythinus inopinatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="89DA57ECFFD42068FD0BFC04FD3EA003" box="[640,733,1092,1118]" pageId="6" pageNumber="279" rank="species">
|
||
<emphasis id="92E9EA97FFD42068FD0BFC04FD34A003" bold="true" box="[640,727,1092,1118]" pageId="6" pageNumber="279">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
differs from all other congeneric species by its aedeagal and exoskeletal features. It shares affinities with
|
||
<taxonomicName id="679D4D06FFD42068FD03FC28FAF8A0DF" authority="Sabella, Amore, Nicolosi, 2019" authorityName="Sabella, Amore, Nicolosi" authorityYear="2019" box="[648,1307,1128,1154]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="279" phylum="Arthropoda" rank="species" species="villasmundi">
|
||
<emphasis id="92E9EA97FFD42068FD03FC28FC50A0DF" box="[648,947,1128,1154]" italics="true" pageId="6" pageNumber="279">Tychobythinus villasmundi</emphasis>
|
||
<bibRefCitation id="C40C4B74FFD42068FC31FC28FAF8A0DF" author="Sabella, G. & Amore, E. & Nicolosi, G." box="[954,1307,1128,1154]" pageId="6" pageNumber="279" pagination="379 - 386" refId="ref4703" refString="Sabella, G., Amore, E. & Nicolosi, G. (2019) A new troglobitic Tychobythinus from Sicily (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa, 4567 (2), 379 - 386. https: // doi. org / 10.11646 / zootaxa. 4567.2.10" type="journal article" year="2019">Sabella, Amore, Nicolosi, 2019</bibRefCitation>
|
||
</taxonomicName>
|
||
, however it differs from the latter in numerous characters: antennal scape more than 4 times longer than wide (less than 4 times longer than wide for
|
||
<taxonomicName id="679D4D06FFD42068FE0EFCF1FDC7A097" authorityName="Sabella, Amore, Nicolosi" authorityYear="2019" box="[389,548,1200,1226]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="279" phylum="Arthropoda" rank="species" species="villasmundi">
|
||
<emphasis id="92E9EA97FFD42068FE0EFCF1FDC7A097" box="[389,548,1200,1226]" italics="true" pageId="6" pageNumber="279">T. villasmundi</emphasis>
|
||
</taxonomicName>
|
||
); last palpomere about 5 times longer than wide, with lateral margin distinctly curved and sinuate at middle (about 4 times longer than wide with lateral margin slightly curved and sinuate at middle for
|
||
<taxonomicName id="679D4D06FFD42068FE9CFCB9FE54A14F" authorityName="Sabella, Amore, Nicolosi" authorityYear="2019" box="[279,439,1272,1298]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="279" phylum="Arthropoda" rank="species" species="villasmundi">
|
||
<emphasis id="92E9EA97FFD42068FE9CFCB9FE54A14F" box="[279,439,1272,1298]" italics="true" pageId="6" pageNumber="279">T. villasmundi</emphasis>
|
||
</taxonomicName>
|
||
), different shape of gular ridge of male (cfr.
|
||
<figureCitation id="38A62A00FFD42068FC37FCB8FBD9A14F" box="[956,1082,1272,1298]" captionStart="FIGURES 8–11" captionStartId="5.[151,264,1929,1954]" captionTargetBox="[222,1362,852,1901]" captionTargetId="figure@5.[218,1369,849,1904]" captionTargetPageId="5" captionText="FIGURES 8–11. Tychobythinus inopinatus sp. nov. 8. Male head in dorsal view, schematic drawning. 9. Female head in dorsal view, schematic drawning. 10. Paratype male, head, lateral view (Ph. G. Nicolosi). 11. Paratype male, head, ventral view (Ph. G. Nicolosi). Scale: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3833591" httpUri="https://zenodo.org/record/3833591/files/figure.png" pageId="6" pageNumber="279">Figs 10–11</figureCitation>
|
||
with
|
||
<figureCitation id="38A62A00FFD42068FBF6FCB8FB03A14F" box="[1149,1248,1272,1298]" captionStart="FIGURES 8–11" captionStartId="5.[151,264,1929,1954]" captionTargetBox="[222,1362,852,1901]" captionTargetId="figure@5.[218,1369,849,1904]" captionTargetPageId="5" captionText="FIGURES 8–11. Tychobythinus inopinatus sp. nov. 8. Male head in dorsal view, schematic drawning. 9. Female head in dorsal view, schematic drawning. 10. Paratype male, head, lateral view (Ph. G. Nicolosi). 11. Paratype male, head, ventral view (Ph. G. Nicolosi). Scale: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3833591" httpUri="https://zenodo.org/record/3833591/files/figure.png" pageId="6" pageNumber="279">
|
||
Figs
|
||
<quantity id="67659B60FFD42068FB3DFCB8FB03A14F" box="[1206,1248,1272,1298]" metricMagnitude="-1" metricUnit="m" metricValue="2.159" metricValueMax="2.286" metricValueMin="2.032" pageId="6" pageNumber="279" unit="in" value="8.5" valueMax="9.0" valueMin="8.0">8–9</quantity>
|
||
</figureCitation>
|
||
in
|
||
<bibRefCitation id="C40C4B74FFD42068FA8CFCB8FF30A16B" author="Sabella, G. & Amore, E. & Nicolosi, G." pageId="6" pageNumber="279" pagination="379 - 386" refId="ref4703" refString="Sabella, G., Amore, E. & Nicolosi, G. (2019) A new troglobitic Tychobythinus from Sicily (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa, 4567 (2), 379 - 386. https: // doi. org / 10.11646 / zootaxa. 4567.2.10" type="journal article" year="2019">
|
||
Sabella
|
||
<emphasis id="92E9EA97FFD42068FAE9FCB9FA7EA14F" box="[1378,1437,1272,1298]" italics="true" pageId="6" pageNumber="279">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
), aedeagus with different apical morphology of the parameres, which are not narrowed in
|
||
<taxonomicName id="679D4D06FFD42068FB55FD5DFA91A16B" authority="Sabella & Costanzo & Nicolosi, 2020" authorityName="Sabella & Costanzo & Nicolosi" authorityYear="2020" box="[1246,1394,1309,1334]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="279" phylum="Arthropoda" rank="species" species="inopinatus" status="sp. nov.">
|
||
<emphasis id="92E9EA97FFD42068FB55FD5DFA91A16B" box="[1246,1394,1309,1334]" italics="true" pageId="6" pageNumber="279">T. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="89DA57ECFFD42068FAF0FD5CFF2AA107" pageId="6" pageNumber="279" rank="species">
|
||
<emphasis id="92E9EA97FFD42068FAF0FD5CFF20A107" bold="true" pageId="6" pageNumber="279">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, and are distinctly narrowed in
|
||
<taxonomicName id="679D4D06FFD42068FDA6FD01FD2FA107" authorityName="Sabella, Amore, Nicolosi" authorityYear="2019" box="[557,716,1344,1370]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="279" phylum="Arthropoda" rank="species" species="villasmundi">
|
||
<emphasis id="92E9EA97FFD42068FDA6FD01FD2FA107" box="[557,716,1344,1370]" italics="true" pageId="6" pageNumber="279">T. villasmundi</emphasis>
|
||
</taxonomicName>
|
||
(cfr.
|
||
<figureCitation id="38A62A00FFD42068FC8CFD00FCB5A107" box="[775,854,1344,1370]" captionStart="FIGURES 12–16" captionStartId="6.[151,264,949,974]" captionTargetBox="[178,1407,187,920]" captionTargetId="figure@6.[174,1413,181,923]" captionTargetPageId="6" captionText="FIGURES 12–16. Tychobythinus inopinatus sp. nov. 12. Paratype female, left antenna. 13. Paratype female, right maxillary palpus. 14. Holotype male, aedeagus, dorsal view. 15. Paratype male, internal sac of aedeagus, dorsal view. 16. Paratype female, telisternite, dorsal view. Scale: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3833593" httpUri="https://zenodo.org/record/3833593/files/figure.png" pageId="6" pageNumber="279">Fig. 14</figureCitation>
|
||
with
|
||
<figureCitation id="38A62A00FFD42068FC1DFD00FC05A107" box="[918,998,1344,1370]" captionStart="FIGURES 12–16" captionStartId="6.[151,264,949,974]" captionTargetBox="[178,1407,187,920]" captionTargetId="figure@6.[174,1413,181,923]" captionTargetPageId="6" captionText="FIGURES 12–16. Tychobythinus inopinatus sp. nov. 12. Paratype female, left antenna. 13. Paratype female, right maxillary palpus. 14. Holotype male, aedeagus, dorsal view. 15. Paratype male, internal sac of aedeagus, dorsal view. 16. Paratype female, telisternite, dorsal view. Scale: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3833593" httpUri="https://zenodo.org/record/3833593/files/figure.png" pageId="6" pageNumber="279">
|
||
Fig.
|
||
<quantity id="67659B60FFD42068FC41FD00FC05A107" box="[970,998,1344,1370]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="6" pageNumber="279" unit="in" value="12.0">12</quantity>
|
||
</figureCitation>
|
||
in
|
||
<bibRefCitation id="C40C4B74FFD42068FB82FD00FB3DA107" author="Sabella, G. & Amore, E. & Nicolosi, G." box="[1033,1246,1344,1370]" pageId="6" pageNumber="279" pagination="379 - 386" refId="ref4703" refString="Sabella, G., Amore, E. & Nicolosi, G. (2019) A new troglobitic Tychobythinus from Sicily (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa, 4567 (2), 379 - 386. https: // doi. org / 10.11646 / zootaxa. 4567.2.10" type="journal article" year="2019">
|
||
Sabella
|
||
<emphasis id="92E9EA97FFD42068FBE8FD01FB77A107" box="[1123,1172,1344,1370]" italics="true" pageId="6" pageNumber="279">et al</emphasis>
|
||
. 2019
|
||
</bibRefCitation>
|
||
), and differences of the internal sac (cfr.
|
||
<figureCitation id="38A62A00FFD42068FE10FD24FDFBA123" box="[411,536,1379,1406]" captionStart="FIGURES 12–16" captionStartId="6.[151,264,949,974]" captionTargetBox="[178,1407,187,920]" captionTargetId="figure@6.[174,1413,181,923]" captionTargetPageId="6" captionText="FIGURES 12–16. Tychobythinus inopinatus sp. nov. 12. Paratype female, left antenna. 13. Paratype female, right maxillary palpus. 14. Holotype male, aedeagus, dorsal view. 15. Paratype male, internal sac of aedeagus, dorsal view. 16. Paratype female, telisternite, dorsal view. Scale: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3833593" httpUri="https://zenodo.org/record/3833593/files/figure.png" pageId="6" pageNumber="279">Figs 14–15</figureCitation>
|
||
with
|
||
<figureCitation id="38A62A00FFD42068FDD3FD24FD44A123" box="[600,679,1380,1406]" captionStart="FIGURES 12–16" captionStartId="6.[151,264,949,974]" captionTargetBox="[178,1407,187,920]" captionTargetId="figure@6.[174,1413,181,923]" captionTargetPageId="6" captionText="FIGURES 12–16. Tychobythinus inopinatus sp. nov. 12. Paratype female, left antenna. 13. Paratype female, right maxillary palpus. 14. Holotype male, aedeagus, dorsal view. 15. Paratype male, internal sac of aedeagus, dorsal view. 16. Paratype female, telisternite, dorsal view. Scale: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3833593" httpUri="https://zenodo.org/record/3833593/files/figure.png" pageId="6" pageNumber="279">
|
||
Fig.
|
||
<quantity id="67659B60FFD42068FD00FD24FD44A123" box="[651,679,1380,1406]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="6" pageNumber="279" unit="in" value="12.0">12</quantity>
|
||
</figureCitation>
|
||
in
|
||
<bibRefCitation id="C40C4B74FFD42068FD40FD24FC43A123" author="Sabella, G. & Amore, E. & Nicolosi, G." box="[715,928,1380,1406]" pageId="6" pageNumber="279" pagination="379 - 386" refId="ref4703" refString="Sabella, G., Amore, E. & Nicolosi, G. (2019) A new troglobitic Tychobythinus from Sicily (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa, 4567 (2), 379 - 386. https: // doi. org / 10.11646 / zootaxa. 4567.2.10" type="journal article" year="2019">
|
||
Sabella
|
||
<emphasis id="92E9EA97FFD42068FCAFFD25FCB5A123" box="[804,854,1380,1406]" italics="true" pageId="6" pageNumber="279">et al</emphasis>
|
||
. 2019
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="A0223685FFD42068FF4CFDC8FEB0A253" blockId="6.[151,1437,1091,2018]" pageId="6" pageNumber="279">
|
||
Like
|
||
<emphasis id="92E9EA97FFD42068FE89FDC9FE44A1FF" box="[258,423,1416,1442]" italics="true" pageId="6" pageNumber="279">
|
||
<taxonomicName id="679D4D06FFD42068FE89FDC9FE41A1FF" authorityName="Sabella, Amore, Nicolosi" authorityYear="2019" box="[258,418,1416,1442]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="279" phylum="Arthropoda" rank="species" species="villasmundi">T. villasmundi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicName id="679D4D06FFD42068FE25FDC9FDA1A1FF" authority="Sabella & Costanzo & Nicolosi, 2020" authorityName="Sabella & Costanzo & Nicolosi" authorityYear="2020" box="[430,578,1416,1442]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="279" phylum="Arthropoda" rank="species" species="inopinatus" status="sp. nov.">
|
||
<emphasis id="92E9EA97FFD42068FE25FDC9FE5DA1FF" box="[430,446,1417,1442]" italics="true" pageId="6" pageNumber="279">T</emphasis>
|
||
.
|
||
<emphasis id="92E9EA97FFD42068FE47FDC9FDA1A1FF" box="[460,578,1417,1442]" italics="true" pageId="6" pageNumber="279">inopinatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="89DA57ECFFD42068FDC1FDC8FD46A1FF" box="[586,677,1416,1442]" pageId="6" pageNumber="279" rank="species">
|
||
<emphasis id="92E9EA97FFD42068FDC1FDC8FD7DA1FF" bold="true" box="[586,670,1416,1442]" pageId="6" pageNumber="279">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
shares affinities to the congeneric species from North Africa which belong to two distinct groups (
|
||
<bibRefCitation id="C40C4B74FFD42068FE7EFDECFD31A19B" author="Sabella, G. & Viglianisi, F. M. & D'Urso, V. & Nouira, S." box="[501,722,1452,1478]" pageId="6" pageNumber="279" pagination="579 - 586" refId="ref4637" refString="Sabella, G., Viglianisi, F. M., D'Urso, V. & Nouira, S. (2014) A review of the Tychobythinus algiricus species group from North Africa (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa, 3893 (4), 579 - 586. https: // doi. org / 10.11646 / zootaxa. 3893.4.7" type="journal article" year="2014">
|
||
Sabella
|
||
<emphasis id="92E9EA97FFD42068FDD9FDEDFD6FA19B" box="[594,652,1452,1478]" italics="true" pageId="6" pageNumber="279">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
). However, they do not represent, with certainty, two homogeneous phyletic lineages because their external morphology is closely related to their different levels of adaptation to endogean life.
|
||
</paragraph>
|
||
<paragraph id="A0223685FFD42068FF4CFE58FE33A2C3" blockId="6.[151,1437,1091,2018]" pageId="6" pageNumber="279">
|
||
The aedeagus of
|
||
<taxonomicName id="679D4D06FFD42068FE02FE59FDFFA26F" authority="Sabella & Costanzo & Nicolosi, 2020" authorityName="Sabella & Costanzo & Nicolosi" authorityYear="2020" box="[393,540,1561,1586]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="279" phylum="Arthropoda" rank="species" species="inopinatus" status="sp. nov.">
|
||
<emphasis id="92E9EA97FFD42068FE02FE59FDFFA26F" box="[393,540,1561,1586]" italics="true" pageId="6" pageNumber="279">T. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="89DA57ECFFD42068FDAFFE58FD63A26F" box="[548,640,1560,1586]" pageId="6" pageNumber="279" rank="species">
|
||
<emphasis id="92E9EA97FFD42068FDAFFE58FD99A26F" bold="true" box="[548,634,1560,1586]" pageId="6" pageNumber="279">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is comparable to the species belonging to the
|
||
<taxonomicName id="679D4D06FFD42068FB04FE58FA7EA26F" baseAuthorityName="Raffray" baseAuthorityYear="1871" box="[1167,1437,1560,1586]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="279" phylum="Arthropoda" rank="species" species="algiricus">
|
||
<emphasis id="92E9EA97FFD42068FB04FE58FA7EA26F" box="[1167,1437,1560,1586]" italics="true" pageId="6" pageNumber="279">Tychobythinus algiricus</emphasis>
|
||
</taxonomicName>
|
||
group (
|
||
<bibRefCitation id="C40C4B74FFD42068FF6CFE7CFE5FA20B" author="Sabella, G. & Viglianisi, F. M. & D'Urso, V. & Nouira, S." box="[231,444,1596,1622]" pageId="6" pageNumber="279" pagination="579 - 586" refId="ref4637" refString="Sabella, G., Viglianisi, F. M., D'Urso, V. & Nouira, S. (2014) A review of the Tychobythinus algiricus species group from North Africa (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa, 3893 (4), 579 - 586. https: // doi. org / 10.11646 / zootaxa. 3893.4.7" type="journal article" year="2014">
|
||
Sabella
|
||
<emphasis id="92E9EA97FFD42068FECAFE7DFE9AA20B" box="[321,377,1596,1622]" italics="true" pageId="6" pageNumber="279">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
;
|
||
<taxonomicName id="679D4D06FFD42068FE4DFE7CFC89A20B" authority="sensu Jeannel 1956" authorityName="Jeannel" authorityYear="1956" box="[454,874,1595,1622]" class="Insecta" family="Staphylinidae" genus="Chiasmatobythus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="279" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="92E9EA97FFD42068FE4DFE7CFD2EA20B" box="[454,717,1596,1622]" italics="true" pageId="6" pageNumber="279">Chiasmatobythus sensu</emphasis>
|
||
<bibRefCitation id="C40C4B74FFD42068FD5FFE7CFC89A20B" author="Jeannel, R." box="[724,874,1595,1622]" pageId="6" pageNumber="279" pagination="1 - 233" refId="ref4160" refString="Jeannel, R. (1956) Les Pselaphides de l'Afrique du Nord. Essai de biogeographie berbere. Memoires du Museum National d'Histoire Naturelle, Paris, New Series, Serie A, Zoologie, 14, 1 - 233." type="journal article" year="1956">Jeannel 1956</bibRefCitation>
|
||
</taxonomicName>
|
||
), due to their troglobitic adaptations.
|
||
<taxonomicName id="679D4D06FFD42068FA80FE7DFA7FA20B" authority="Sabella & Costanzo & Nicolosi, 2020" authorityName="Sabella & Costanzo & Nicolosi" authorityYear="2020" box="[1291,1436,1597,1622]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="279" phylum="Arthropoda" rank="species" species="inopinatus" status="sp. nov.">
|
||
<emphasis id="92E9EA97FFD42068FA80FE7DFA7FA20B" box="[1291,1436,1597,1622]" italics="true" pageId="6" pageNumber="279">T. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="89DA57ECFFD42068FF1CFE20FF11A227" box="[151,242,1632,1658]" pageId="6" pageNumber="279" rank="species">
|
||
<emphasis id="92E9EA97FFD42068FF1CFE20FF0FA227" bold="true" box="[151,236,1632,1658]" pageId="6" pageNumber="279">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is externally close to the species belonging to the
|
||
<taxonomicName id="679D4D06FFD42068FCAFFE20FBEEA227" baseAuthorityName="Guillebeau" baseAuthorityYear="1893" box="[804,1037,1632,1658]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="279" phylum="Arthropoda" rank="species" species="theryi">
|
||
<emphasis id="92E9EA97FFD42068FCAFFE20FBEEA227" box="[804,1037,1632,1658]" italics="true" pageId="6" pageNumber="279">Tychobythinus theryi</emphasis>
|
||
</taxonomicName>
|
||
group (
|
||
<bibRefCitation id="C40C4B74FFD42068FBEEFE20FADAA227" author="Sabella, G. & D'Urso, V. & Nouira, S. & Souissi, R. & Besuchet, C." box="[1125,1337,1632,1658]" pageId="6" pageNumber="279" pagination="63 - 75" refId="ref4564" refString="Sabella, G., D'Urso, V., Nouira, S., Souissi, R. & Besuchet, C. (2013) A review of Tychobythinus theryi species group with description of a new species from Algeria (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa, 3701 (1), 63 - 75. https: // doi. org / 10.11646 / zootaxa. 3701.1.5" type="journal article" year="2013">
|
||
Sabella
|
||
<emphasis id="92E9EA97FFD42068FB4BFE21FB12A227" box="[1216,1265,1632,1658]" italics="true" pageId="6" pageNumber="279">et al</emphasis>
|
||
. 2013
|
||
</bibRefCitation>
|
||
;
|
||
<taxonomicName id="679D4D06FFD42068FACEFE20FE20A2C3" authority="sensu Jeannel 1956" authorityName="Jeannel" authorityYear="1956" class="Insecta" family="Staphylinidae" genus="Anopsibythus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="279" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="92E9EA97FFD42068FACEFE20FEC6A2C3" italics="true" pageId="6" pageNumber="279">Anopsibythus sensu</emphasis>
|
||
<bibRefCitation id="C40C4B74FFD42068FEA6FEC4FE20A2C3" author="Jeannel, R." box="[301,451,1667,1694]" pageId="6" pageNumber="279" pagination="1 - 233" refId="ref4160" refString="Jeannel, R. (1956) Les Pselaphides de l'Afrique du Nord. Essai de biogeographie berbere. Memoires du Museum National d'Histoire Naturelle, Paris, New Series, Serie A, Zoologie, 14, 1 - 233." type="journal article" year="1956">Jeannel 1956</bibRefCitation>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="A0223685FFD42068FF4CFEE8FA8CA357" blockId="6.[151,1437,1091,2018]" pageId="6" pageNumber="279">
|
||
<materialsCitation id="10F53CD8FFD42068FF4CFEE8FA88A357" ID-GBIF-Occurrence="2620591304" collectingDate="2019-06-06" collectionCode="DBUC, VI, VII" collectorName="G. Nicolosi & This" location="Among" municipality="Monello Cave" pageId="6" pageNumber="279" specimenCount="1">
|
||
<location id="A542605EFFD42068FF4CFEE8FEF8A29F" LSID="urn:lsid:plazi:treatment:28348793FFD12069FF1CFCAAFE10A5D6:A542605EFFD42068FF4CFEE8FEF8A29F" box="[199,283,1704,1730]" municipality="Monello Cave" name="Among" pageId="6" pageNumber="279">Among</location>
|
||
the material examined we also found a female collected in
|
||
<collectingMunicipality id="4046ACFFFFD42068FC22FEE8FBA9A29F" box="[937,1098,1704,1730]" pageId="6" pageNumber="279">Monello Cave</collectingMunicipality>
|
||
,
|
||
<date id="D4231045FFD42068FBDEFEE8FB35A29F" box="[1109,1238,1704,1730]" pageId="6" pageNumber="279" value="2019-06-06">
|
||
<collectingDate id="C467E9ADFFD42068FBDEFEE8FB35A29F" box="[1109,1238,1704,1730]" pageId="6" pageNumber="279" value="2019-06-06">06.VI.2019</collectingDate>
|
||
</date>
|
||
, trap 13,
|
||
<collectorName id="0D685353FFD42068FAB0FEE8FF5CA2BB" pageId="6" pageNumber="279">G. Nicolosi</collectorName>
|
||
leg. (
|
||
<collectionCode id="C68CAE40FFD42068FF76FE8CFEB3A2BB" box="[253,336,1740,1766]" pageId="6" pageNumber="279">DBUC</collectionCode>
|
||
), with only 10 antennomeres, and with antennomere
|
||
<collectionCode id="C68CAE40FFD42068FC13FE8CFC55A2BB" box="[920,950,1740,1766]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="6" pageNumber="279">VI</collectionCode>
|
||
distinctly longer than wide, which is probably derived from the fusion of antennomers
|
||
<collectionCode id="C68CAE40FFD42068FD0FFEB0FD41A357" box="[644,674,1776,1802]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="6" pageNumber="279">VI</collectionCode>
|
||
and
|
||
<collectionCode id="C68CAE40FFD42068FD53FEB0FCE1A357" box="[728,770,1776,1802]" pageId="6" pageNumber="279">VII</collectionCode>
|
||
.
|
||
<collectorName id="0D685353FFD42068FC87FEB0FCDDA357" box="[780,830,1776,1802]" pageId="6" pageNumber="279">This</collectorName>
|
||
specimen has been excluded from the
|
||
<typeStatus id="7F268827FFD42068FB7AFEB0FAC2A357" box="[1265,1313,1776,1802]" pageId="6" pageNumber="279">type</typeStatus>
|
||
series
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="A0223685FFD42068FF4CFF54FB79A3E3" blockId="6.[151,1437,1091,2018]" pageId="6" pageNumber="279">
|
||
Finally, it seems interesting to underline that the previous research conducted inside Monello Cave in 1991 (
|
||
<bibRefCitation id="C40C4B74FFD42068FAF9FF54FEE9A30F" author="Caruso, D." pageId="6" pageNumber="279" pagination="87 - 121" refId="ref3972" refString="Caruso, D. (1995) La fauna della Grotta Monello. Atti e Memorie dell'Ente Fauna Siciliana, 2, 87 - 121." type="journal article" year="1995">Caruso 1995</bibRefCitation>
|
||
), which used the same collection techniques, did not produce any specimens of the new species. On the contrary, our investigation that began on
|
||
<date id="D4231045FFD42068FDD4FF1CFCF5A32B" box="[607,790,1884,1910]" pageId="6" pageNumber="279" value="2019-01-11">11 January 2019</date>
|
||
and is still in progress allowed us to collect, in addition to the species relieved in the previous study,
|
||
<specimenCount id="B69BFD0CFFD42068FDE7FFC0FCE7A3C7" box="[620,772,1920,1946]" pageId="6" pageNumber="279" type="generic">18 specimens</specimenCount>
|
||
(
|
||
<specimenCount id="B69BFD0CFFD42068FC99FFC0FC8AA3C7" box="[786,873,1920,1946]" pageId="6" pageNumber="279" type="male">2 males</specimenCount>
|
||
and
|
||
<specimenCount id="B69BFD0CFFD42068FC15FFC0FBF9A3C6" box="[926,1050,1920,1947]" pageId="6" pageNumber="279" type="female">16 females</specimenCount>
|
||
) of the new species of
|
||
<taxonomicName id="679D4D06FFD42068FA92FFC0FF5DA3E3" authorityName="Ganglbauer" authorityYear="1896" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="279" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="92E9EA97FFD42068FA92FFC0FF5DA3E3" italics="true" pageId="6" pageNumber="279">Tychobythinus</emphasis>
|
||
</taxonomicName>
|
||
in various areas of the cave, but only during the period from
|
||
<date id="D4231045FFD42068FCE7FFE4FB74A3E3" box="[876,1175,1956,1982]" pageId="6" pageNumber="279" value="2019-08" valueMax="2019-12-31" valueMin="2019-08-00">August to December 2019</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="A0223685FFD42069FF4CFF88FAFDA4A6" blockId="6.[151,1437,1091,2018]" lastBlockId="7.[151,1437,153,395]" lastPageId="7" lastPageNumber="280" pageId="6" pageNumber="279">This fact could be explained considering that the ecological conditions inside the cave have improved signifi- cantly in the last thirty years, probably in relation to the implemented strict conservation measures, but the factors driving this change certainly deserve further study (with a careful comparison of the environmental data emerged from the study conducted in 1991 and in 2019), which however goes beyond the purpose of this article.</paragraph>
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<paragraph id="A0223685FFD52069FF4CF945FE10A5D6" blockId="7.[151,1437,153,395]" pageId="7" pageNumber="280">A simpler explanation can be provided by the different sampling periods for the two research programs: research in 1991 covered the period January–July, while our research program, still in progress, covered the entire year of 2019, with collection of the new species happening between August and December, a period not covered by the previous research program.</paragraph>
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</subSubSection>
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</treatment>
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</document> |