230 lines
19 KiB
XML
230 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1123.87863" ID-Pensoft-Pub="1313-2970-1123-83" ID-Pensoft-UUID="2FC5EA8BC9C455009A3BA653CA7BCE26" ID-ZooBank="0C665F99B6814C6693369F65E33B0995" ModsDocID="1313-2970-1123-83" checkinTime="1664944209266" checkinUser="pensoft" docAuthor="Gao, Cuiqing, Xiao, Shiya & Kondorosy, Elod" docDate="2022" docId="CC807A6FD1A8572DA5156F13768E517C" docLanguage="en" docName="ZooKeys 1123: 83-97" docOrigin="ZooKeys 1123" docPubDate="2022-10-04" docSource="http://dx.doi.org/10.3897/zookeys.1123.87863" docTitle="Urvaschia recta Gao & Xiao & Kondorosy 2022, sp. nov." docType="treatment" docUuid="E6FF0F07-0BAB-4014-B183-858C6A2FE4F8" docUuidSource="ZooBank" docVersion="1" id="2FC5EA8BC9C455009A3BA653CA7BCE26" lastPageNumber="83" masterDocId="2FC5EA8BC9C455009A3BA653CA7BCE26" masterDocTitle="Revision of the genus Urvaschia Hopp (Hemiptera, Lygaeoidea, Oxycarenidae), with descriptions of two new species from China and Nepal" masterLastPageNumber="97" masterPageNumber="83" pageNumber="83" updateTime="1664944209266" updateUser="pensoft">
|
|
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo>
|
|
<mods:title>Revision of the genus Urvaschia Hopp (Hemiptera, Lygaeoidea, Oxycarenidae), with descriptions of two new species from China and Nepal</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Gao, Cuiqing</mods:namePart>
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0177-5161</mods:nameIdentifier>
|
|
<mods:affiliation>Co-Innovation Center for Sustainable Forestry in Southern China, College of Forestry, Nanjing Forestry University, Long Pan Road, Nanjing, Jiangsu 210037, China</mods:affiliation>
|
|
<mods:nameIdentifier type="email">cqgao@njfu.edu.cn</mods:nameIdentifier>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Xiao, Shiya</mods:namePart>
|
|
<mods:affiliation>Co-Innovation Center for Sustainable Forestry in Southern China, College of Forestry, Nanjing Forestry University, Long Pan Road, Nanjing, Jiangsu 210037, China</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Kondorosy, Elod</mods:namePart>
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7162-0862</mods:nameIdentifier>
|
|
<mods:affiliation>Department of Conservation Biology, Georgikon Campus, Hungarian University of Agriculture and Life Sciences, H- 8360 Keszthely, Hungary</mods:affiliation>
|
|
<mods:nameIdentifier type="email">kondorosy.ee@gmail.com</mods:nameIdentifier>
|
|
</mods:name>
|
|
<mods:typeOfResource>text</mods:typeOfResource>
|
|
<mods:relatedItem type="host">
|
|
<mods:titleInfo>
|
|
<mods:title>ZooKeys</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part>
|
|
<mods:date>2022</mods:date>
|
|
<mods:detail type="pubDate">
|
|
<mods:number>2022-10-04</mods:number>
|
|
</mods:detail>
|
|
<mods:detail type="volume">
|
|
<mods:number>1123</mods:number>
|
|
</mods:detail>
|
|
<mods:extent unit="page">
|
|
<mods:start>83</mods:start>
|
|
<mods:end>97</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location>
|
|
<mods:url>http://dx.doi.org/10.3897/zookeys.1123.87863</mods:url>
|
|
</mods:location>
|
|
<mods:classification>journal article</mods:classification>
|
|
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1123.87863</mods:identifier>
|
|
<mods:identifier type="Pensoft-Pub">1313-2970-1123-83</mods:identifier>
|
|
<mods:identifier type="ZooBank">0C665F99B6814C6693369F65E33B0995</mods:identifier>
|
|
<mods:identifier type="Pensoft-UUID">2FC5EA8BC9C455009A3BA653CA7BCE26</mods:identifier>
|
|
</mods:mods>
|
|
<treatment LSID="urn:lsid:zoobank.org:act:E6FF0F07-0BAB-4014-B183-858C6A2FE4F8" httpUri="http://treatment.plazi.org/id/CC807A6FD1A8572DA5156F13768E517C" lastPageNumber="83" pageId="0" pageNumber="83">
|
|
<subSubSection pageId="0" pageNumber="83" type="nomenclature">
|
|
<paragraph pageId="0" pageNumber="83">
|
|
<taxonomicName LSID="https://zoobank.org/E6FF0F07-0BAB-4014-B183-858C6A2FE4F8" authority="Gao & Xiao & Kondorosy, 2022" authorityName="Gao & Xiao & Kondorosy" authorityYear="2022" class="Insecta" family="Oxycarenidae" genus="Urvaschia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Urvaschia recta" order="Hemiptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="recta" status="sp. nov.">Urvaschia recta</taxonomicName>
|
|
<taxonomicNameLabel pageId="0" pageNumber="83">sp. nov.</taxonomicNameLabel>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="83" type="description">
|
|
<paragraph pageId="0" pageNumber="83">
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal and ventral views a, d Urvaschia obscuripennis comb. nov. b, e Urvaschia convexa sp. nov., holotype c, f Urvaschia recta sp. nov., holotype." figureDoi="10.3897/zookeys.1123.87863.figure2" httpUri="https://binary.pensoft.net/fig/752638" pageId="0" pageNumber="83">Figs 2c, f</figureCitation>
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pygophore (dorsal views, parameres removed) and left paramere (dorsal and lateral views): a-c Urvaschia obscuripennis comb. nov. d-f Urvaschia convexa sp. nov., paratype g-i Urvaschia recta sp. nov., holotype." figureDoi="10.3897/zookeys.1123.87863.figure3" httpUri="https://binary.pensoft.net/fig/752639" pageId="0" pageNumber="83">, 3g-i</figureCitation>
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution of the species of Urvaschia." figureDoi="10.3897/zookeys.1123.87863.figure4" httpUri="https://binary.pensoft.net/fig/752640" pageId="0" pageNumber="83">, 4</figureCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="83" type="type material">
|
|
<paragraph pageId="0" pageNumber="83">Type material examined.</paragraph>
|
|
<paragraph pageId="0" pageNumber="83">
|
|
<emphasis bold="true" pageId="0" pageNumber="83">
|
|
<emphasis italics="true" pageId="0" pageNumber="83">Holotype</emphasis>
|
|
.
|
|
</emphasis>
|
|
China • ♂; Lijiang [hw], Yunnan [pr] / 11.viii. [hw]1979 [pr] / leg. Jianxin Cui [pr] (NKUM).
|
|
<emphasis bold="true" pageId="0" pageNumber="83">
|
|
<emphasis italics="true" pageId="0" pageNumber="83">Paratypes</emphasis>
|
|
.
|
|
</emphasis>
|
|
China • 1♀, Bayi town, Xizang / 6.viii.2003 / leg. Huaijun Xue, Xinpu Wang [all pr] (NKUM); 1♀, Xiaonanchuan Forestry Centre, Erlonghe, Liupanshan, Ningxia / 28.vi.2008 / 1900m. leg. Gengping Zhu [all pr] (NKUM); 1♀, Zhongreniao, Xiangcheng [all hw], Sichuan [pr] / 3950m // 1982.VII.4 [all hw] / leg. Huaicheng Chai [hw] (IZAS).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="83" type="materials_examined">
|
|
<paragraph pageId="0" pageNumber="83">Other material examined.</paragraph>
|
|
<paragraph pageId="0" pageNumber="83">
|
|
<materialsCitation collectingDate="2002-08-02" collectorName="G. Melika, Umg. Alm Darghari, Maharigaon, Gebiet von Jumla, Westnepal, H. Franz, Innsbruck, Austria" country="NEPAL" county="Linshan Mt." elevation="1749" location="China" longitude="115.5" municipality="Liyan Ling" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
|
|
<location LSID="urn:lsid:plazi:treatment:CC807A6FD1A8572DA5156F13768E517C:466B37A4E8FFA639BE3D6F51E4F41D29" country="NEPAL" county="Linshan Mt." longitude="115.5" municipality="Liyan Ling" name="China">China</location>
|
|
• pr.
|
|
<location LSID="urn:lsid:plazi:treatment:CC807A6FD1A8572DA5156F13768E517C:2A0289C0E8347CB8A4DD5237D73026DF" country="NEPAL" county="Linshan Mt." longitude="115.5" municipality="Liyan Ling" name="Beijing">Beijing</location>
|
|
/
|
|
<location LSID="urn:lsid:plazi:treatment:CC807A6FD1A8572DA5156F13768E517C:E0D3345C723F3B427D1428499FABB83A" country="NEPAL" county="Linshan Mt." longitude="115.5" municipality="Liyan Ling" name="Mentougou Dist.">Mentougou Dist.</location>
|
|
/
|
|
<location LSID="urn:lsid:plazi:treatment:CC807A6FD1A8572DA5156F13768E517C:C3A64107894D1F920015D5DC775765AF" country="NEPAL" county="Linshan Mt." longitude="115.5" municipality="Liyan Ling" name="Beijing">Beijing</location>
|
|
<quantity metricMagnitude="5" metricUnit="m" metricValue="1.3" unit="km" value="130.0">130 km</quantity>
|
|
NW /
|
|
<collectingMunicipality>Liyan Ling</collectingMunicipality>
|
|
//
|
|
<collectingCounty>Linshan Mt.</collectingCounty>
|
|
/
|
|
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.749" unit="m" value="1749.0">
|
|
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.749" unit="m" value="1749.0">1749 m</elevation>
|
|
</quantity>
|
|
,
|
|
<geoCoordinate degrees="115" direction="east" minutes="30" orientation="longitude" precision="925" value="115.5">115°30'E</geoCoordinate>
|
|
/ 40°00',
|
|
<collectingDate value="2002-08-02">2.VIII.2002</collectingDate>
|
|
// leg.
|
|
<collectorName>G. Melika</collectorName>
|
|
(HNHM);
|
|
<collectingCountry name="Nepal">NEPAL</collectingCountry>
|
|
•
|
|
<specimenCount type="male">1♂</specimenCount>
|
|
<specimenCount type="female">1♀</specimenCount>
|
|
<collectorName>Umg. Alm Darghari</collectorName>
|
|
/ b.
|
|
<collectorName>Maharigaon</collectorName>
|
|
,
|
|
<quantity metricMagnitude="3" metricUnit="m" metricValue="4.0" unit="m" value="4000.0">
|
|
<elevation metricMagnitude="3" metricUnit="m" metricValue="4.0" unit="m" value="4000.0">4000m</elevation>
|
|
</quantity>
|
|
//
|
|
<collectorName>Gebiet von Jumla</collectorName>
|
|
/
|
|
<collectorName>Westnepal</collectorName>
|
|
, lg.
|
|
<collectorName>H. Franz</collectorName>
|
|
// COLLECTION / ERNST HEISS /
|
|
<collectorName>Innsbruck</collectorName>
|
|
-
|
|
<collectorName>Austria</collectorName>
|
|
(CEHI in TLMF)
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="83" type="description">
|
|
<paragraph pageId="0" pageNumber="83">Description.</paragraph>
|
|
<paragraph pageId="0" pageNumber="83">
|
|
<emphasis bold="true" pageId="0" pageNumber="83">
|
|
<emphasis italics="true" pageId="0" pageNumber="83">Colouration</emphasis>
|
|
.
|
|
</emphasis>
|
|
Head blackish brown. Antennae dark blackish brown. Bucculae and labium dark brown. Pronotum with a yellowish white midline except area of calli. Anterior lobe of pronotum blackish brown, posterior lobe dark brown. Scutellum blackish brown. Hemelytra pale yellowish brown, with dense dark brown spots between brown veins covering exocorium as well; distal margin of corium dark brown; apical angle of corium with a blackish brown spot. Thoracal sterna blackish brown. Supracoxal lobes yellowish white to yellow. Mesopleuron black; inner 1/2 of ostiolar peritreme of metathoracic scent gland yellowish white, outer 1/2 of ostiolar peritreme brown. Posterior 1/2 of metapleura broadly yellowish white. Femora blackish brown; tibiae and tarsi ochraceous. Abdominal sterna dark reddish brown.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="83">
|
|
<emphasis bold="true" pageId="0" pageNumber="83">
|
|
<emphasis italics="true" pageId="0" pageNumber="83">Structure</emphasis>
|
|
.
|
|
</emphasis>
|
|
Head slightly declined, covered with large deep punctures and erect white setae. Eyes slightly protruding laterally. Distance between posterior margin of eyes and anterior margin of pronotum 1/2 diameter of eyes. Bucculae high, almost parallel to labium, visible laterad of clypeus from dorsal view. Antennae covered with dense oblique setae, apical 1/3 of segment I surpassing clypeus. First segment of labium surpassing bucculae, segment II surpassing base of head, labium reaching middle of mesocoxae. Venter of head flat, covered with punctures and dense white decumbent setae.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="83">
|
|
Pronotum trapezoid, flat, covered with large dense punctures and long white erect setae, slightly leaning posteriad; calli slightly emergent. Anterior margin of pronotum straight; posterior margin of pronotum straight with posterolateral pronotal angles slightly protruding posteriad. Base of scutellum slightly sunken; each margin covered with dense punctures, smaller than on pronotum and sparse setae, central area with sparse punctures and inconspicuous median carina. Hemelytra flat, sparsely covered with white and short setae; clavus with inner and outer rows of punctures, scattered with irregular one or two rows of punctures in middle. Corium with several punctures between vein Cu and clavus, and a row of punctures along inner margin of exocorium (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal and ventral views a, d Urvaschia obscuripennis comb. nov. b, e Urvaschia convexa sp. nov., holotype c, f Urvaschia recta sp. nov., holotype." figureDoi="10.3897/zookeys.1123.87863.figure2" httpUri="https://binary.pensoft.net/fig/752638" pageId="0" pageNumber="83">2c</figureCitation>
|
|
); cubital vein inconspicuous. Apical margin of corium straight, costal margin evenly arched; body broadest near apex of clavus; corium longer than 2/3 of hemelytra. Membrane comparatively broad, almost fully overlapping each other (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal and ventral views a, d Urvaschia obscuripennis comb. nov. b, e Urvaschia convexa sp. nov., holotype c, f Urvaschia recta sp. nov., holotype." figureDoi="10.3897/zookeys.1123.87863.figure2" httpUri="https://binary.pensoft.net/fig/752638" pageId="0" pageNumber="83">2c</figureCitation>
|
|
); membranal veins thick and conspicuous; apex of membrane surpassing abdomen. Femora slightly thickened, profemora sometimes with one small spine (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal and ventral views a, d Urvaschia obscuripennis comb. nov. b, e Urvaschia convexa sp. nov., holotype c, f Urvaschia recta sp. nov., holotype." figureDoi="10.3897/zookeys.1123.87863.figure2" httpUri="https://binary.pensoft.net/fig/752638" pageId="0" pageNumber="83">2f</figureCitation>
|
|
). Abdominal connexivum not exposed. Abdominal sternum impunctate, covered with sparse setae.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="83">
|
|
Pygophore (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pygophore (dorsal views, parameres removed) and left paramere (dorsal and lateral views): a-c Urvaschia obscuripennis comb. nov. d-f Urvaschia convexa sp. nov., paratype g-i Urvaschia recta sp. nov., holotype." figureDoi="10.3897/zookeys.1123.87863.figure3" httpUri="https://binary.pensoft.net/fig/752639" pageId="0" pageNumber="83">3g</figureCitation>
|
|
): Posterior margin of pygophore and cup-like sclerite fused. Parameres (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pygophore (dorsal views, parameres removed) and left paramere (dorsal and lateral views): a-c Urvaschia obscuripennis comb. nov. d-f Urvaschia convexa sp. nov., paratype g-i Urvaschia recta sp. nov., holotype." figureDoi="10.3897/zookeys.1123.87863.figure3" httpUri="https://binary.pensoft.net/fig/752639" pageId="0" pageNumber="83">3h, i</figureCitation>
|
|
): outer projection large, triangular; inner projection inconspicuous; blade bent rectangularly with shank of paramere from lateral view.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="83">
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="83">Measurements</emphasis>
|
|
(mm,
|
|
<emphasis italics="true" pageId="0" pageNumber="83">N</emphasis>
|
|
= 3).
|
|
<emphasis bold="true" pageId="0" pageNumber="83">
|
|
<emphasis italics="true" pageId="0" pageNumber="83">Holotype</emphasis>
|
|
.
|
|
</emphasis>
|
|
♂ (
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="83">Paratypes</emphasis>
|
|
. 2♀♀); Body length 3.49 (3.72-3.81). Head length 0.46 (0.50-0.51), width across eyes 0.77 (0.73-0.76); antennal segments I-IV length: 0.22: 0.46: 0.33: 0.40 (I-IV: 0.18-0.25: 0.54: 0.35: 0.45); labium length 1.34, first segment length 0.32. Pronotum length 0.73 (0.73-0.76), width of anterior margin 0.60 (0.64), width of posterior margin 1.04 (1.12-1.15); scutellum length 0.51 (0.48-0.50), width 0.56 (0.67-0.71). Distance apex clavus- apex corium 1.08 (1.08-1.18); distance apex corium-apex membrane 0.78 (0.98-1.05).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="83" type="etymology">
|
|
<paragraph pageId="0" pageNumber="83">Etymology.</paragraph>
|
|
<paragraph pageId="0" pageNumber="83">
|
|
The species epithet
|
|
<emphasis italics="true" pageId="0" pageNumber="83">recta</emphasis>
|
|
, derived from Latin adjective
|
|
<emphasis italics="true" pageId="0" pageNumber="83">rectus</emphasis>
|
|
(= straight), alludes to the straight apical margin of the corium.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="83" type="distribution">
|
|
<paragraph pageId="0" pageNumber="83">Distribution.</paragraph>
|
|
<paragraph pageId="0" pageNumber="83">
|
|
China (Beijing, Ningxia, Sichuan, Xizang, Yunnan), Nepal (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution of the species of Urvaschia." figureDoi="10.3897/zookeys.1123.87863.figure4" httpUri="https://binary.pensoft.net/fig/752640" pageId="0" pageNumber="83">4</figureCitation>
|
|
). The locality of the Nepalese specimen is very near to the type locality of
|
|
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="U. pterosticta" order="Hemiptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="pterosticta">
|
|
<emphasis italics="true" pageId="0" pageNumber="83">U. pterosticta</emphasis>
|
|
</taxonomicName>
|
|
; therefore, it cannot be seen separately on Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution of the species of Urvaschia." figureDoi="10.3897/zookeys.1123.87863.figure4" httpUri="https://binary.pensoft.net/fig/752640" pageId="0" pageNumber="83">4</figureCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="83" type="differential diagnosis">
|
|
<paragraph pageId="0" pageNumber="83">Differential diagnosis.</paragraph>
|
|
<paragraph pageId="0" pageNumber="83">
|
|
The new species is similar to
|
|
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="U. convexa" order="Hemiptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="convexa">
|
|
<emphasis italics="true" pageId="0" pageNumber="83">U. convexa</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. in having brown spots on the hemelytra and oval body shape, but the corium is not elongated, with the length of the corium almost twice the length of the membrane from the apical angle of the corium to the apex, and its apex is almost straight (vs. corium conspicuously elongated, with the length of the corium almost three times the length of the membrane from the apical angle of the corium to the apex, and the apex of the corium is conspicuously convex in
|
|
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="U. convexa" order="Hemiptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="convexa">
|
|
<emphasis italics="true" pageId="0" pageNumber="83">U. convexa</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.); membrane almost fully overlapping (vs. membrane only overlapping on the inner edge in
|
|
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="U. convexa" order="Hemiptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="convexa">
|
|
<emphasis italics="true" pageId="0" pageNumber="83">U. convexa</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |