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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1151.97514" ID-Pensoft-Pub="1313-2970-1151-67" ID-Pensoft-UUID="0B3CEF4A954955478878AD15C022DCD5" ID-ZooBank="5FEDE55DC9AF47D591259F1758AE2A18" ModsDocID="1313-2970-1151-67" checkinTime="1677749880156" checkinUser="pensoft" docAuthor="Willemse, Luc, Tilmans, Jos, Kotitsa, Nefeli, Trichas, Apostolos, Heller, Klaus-Gerhard, Chobanov, Dragan &amp; Ode, Baudewijn" docDate="2023" docId="C2D55B1A55A35B54ACB4992941DA6D4A" docLanguage="en" docName="ZooKeys 1151: 67-158" docOrigin="ZooKeys 1151" docPubDate="2023-03-01" docSource="http://dx.doi.org/10.3897/zookeys.1151.97514" docTitle="Eupholidoptera mariannae Willemse &amp; Heller 2001" docType="treatment" docVersion="2" id="0B3CEF4A954955478878AD15C022DCD5" lastPageNumber="67" masterDocId="0B3CEF4A954955478878AD15C022DCD5" masterDocTitle="A review of Eupholidoptera (Orthoptera, Tettigoniidae) from Crete, Gavdos, Gavdopoula, and Andikithira" masterLastPageNumber="158" masterPageNumber="67" pageNumber="67" updateTime="1677750187672" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A review of Eupholidoptera (Orthoptera, Tettigoniidae) from Crete, Gavdos, Gavdopoula, and Andikithira</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Willemse, Luc</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0517-9778</mods:nameIdentifier>
<mods:affiliation>Naturalis, PO Box 9517, 2300, RA Leiden, Netherlands</mods:affiliation>
<mods:nameIdentifier type="email">luc.willemse@naturalis.nl</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tilmans, Jos</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0127-6191</mods:nameIdentifier>
<mods:affiliation>Herkenboscher Strasse 33, 41849, Wassenberg-Rothenbach, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kotitsa, Nefeli</mods:namePart>
<mods:affiliation>Natural History Museum Crete, University of Crete, P. O. Box 2208, 71409, Heraklion, Crete, Greece &amp; Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel blvd., 1000, Sofia, Bulgaria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Trichas, Apostolos</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7917-5262</mods:nameIdentifier>
<mods:affiliation>Natural History Museum Crete, University of Crete, P. O. Box 2208, 71409, Heraklion, Crete, Greece</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Heller, Klaus-Gerhard</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3331-3228</mods:nameIdentifier>
<mods:affiliation>Triesdorf Bahnhof 8, 91732, Merkendorf, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chobanov, Dragan</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1642-0363</mods:nameIdentifier>
<mods:affiliation>Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel blvd., 1000, Sofia, Bulgaria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ode, Baudewijn</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8929-2737</mods:nameIdentifier>
<mods:affiliation>Riethorsterweg 12, 6586, AC Plasmolen, Netherlands</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-01</mods:number>
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<mods:detail type="volume">
<mods:number>1151</mods:number>
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<mods:start>67</mods:start>
<mods:end>158</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1151.97514</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1151.97514</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1151-67</mods:identifier>
<mods:identifier type="ZooBank">5FEDE55DC9AF47D591259F1758AE2A18</mods:identifier>
<mods:identifier type="Pensoft-UUID">0B3CEF4A954955478878AD15C022DCD5</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:C2D55B1A55A35B54ACB4992941DA6D4A" httpUri="http://treatment.plazi.org/id/C2D55B1A55A35B54ACB4992941DA6D4A" lastPageNumber="67" pageId="0" pageNumber="67">
<subSubSection pageId="0" pageNumber="67" type="nomenclature">
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<taxonomicName ID-CoL="3CMKV" LSID="C2D55B1A-55A3-5B54-ACB4-992941DA6D4A" authority="Willemse &amp; Heller, 2001" authorityName="Willemse &amp; Heller" authorityYear="2001" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupholidoptera mariannae" order="Orthoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="mariannae">Eupholidoptera mariannae Willemse &amp; Heller, 2001</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67" type="description">
<paragraph pageId="0" pageNumber="67">
<figureCitation captionStart="Figures 11–24" captionStartId="F5" captionText="Figures 11 - 24. Colour pattern of head, frontal view 11 Eupholidoptera smyrnensis ♀ Makrigiannis RMNH 5014918 12 Eupholidoptera gemellata ♂ Mt. Idhi RMNH. INS 1141843 13 Eupholidoptera pallipes ♂ paratype Mt. Lefka RMNH. INS 1105313 14 Eupholidoptera cretica ♂ Mt. Lefka, Omalos RMNH. INS 114838 15 Eupholidoptera latens ♀ Mt. Lefka, Kalergi CT 1987.047.04 16 Eupholidoptera giuliae ♂ Argoules CT 1995.011.18 17 Eupholidoptera francisae sp. nov. ♀ allotype Andikithira CT 2002.004.11 18 Eupholidoptera annamariae ♀ Kato Zakros CT 1995.020.03 19 Eupholidoptera astyla ♀ Ano Viannos CT 1987.024.03 20 Eupholidoptera feri ♂ holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 21 Eupholidoptera mariannae ♀ Ag. Ioannis RMNH. 5014906 22 Eupholidoptera forcipata ♂ Mt. Idhi CT 1987.044.03 23 Eupholidoptera marietheresae sp. nov. ♂ paratype Mt. Dikti, FC 1606 CT 2000.096.01 24 Eupholidoptera jacquelinae ♂ holotype Gavdos CT 2000.005.02. Scale bars: 5 mm." figureDoi="10.3897/zookeys.1151.97514.figures11-24" httpUri="https://binary.pensoft.net/fig/817935" pageId="0" pageNumber="67">Figs 21</figureCitation>
<figureCitation captionStart="Figures 25–38" captionStartId="F6" captionText="Figures 25 - 38. Colour pattern of pronotum, dorsal view 25 Eupholidoptera smyrnensis ♂ Makrigiannis RMNH. INS 152909 26 Eupholidoptera gemellata ♂ Mt. Idhi, Amariou RMNH. INS 1141843 27 Eupholidoptera pallipes ♂ paratype Mt. Lefka RMNH. INS 1105313 28 Eupholidoptera cretica ♂ Mt. Lefka, Omalos RMNH. INS 1141838 29 Eupholidoptera latens ♂ Rhodopos CH 8236 30 Eupholidoptera giuliae Chora Sfakion ♂ CT 2000.014.10 31 Eupholidoptera francisae sp. nov. ♂ holotype Andikithira CT 2002.004.04 32 Eupholidoptera annamariae ♂ Kato Zakros CT 2000.030.04 33 Eupholidoptera astyla ♂ Krotos RMNH. INS 1141819 34 Eupholidoptera feri ♂ holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 35 Eupholidoptera mariannae ♀ Kalavros RMNH. 5086974 36 Eupholidoptera forcipata ♂ Mt. Idhi CT 1987.044.03 37 Eupholidoptera marietheresae sp. nov. ♂ holotype Mt. Dikti RMNH. INS 1141850 38 Eupholidoptera jacquelinae ♂ holotype Gavdos CT 2000.005.02. Scale bars: 5 mm." figureDoi="10.3897/zookeys.1151.97514.figures25-38" httpUri="https://binary.pensoft.net/fig/817936" pageId="0" pageNumber="67">, 35</figureCitation>
<figureCitation captionStart="Figures 39–52" captionStartId="F7" captionText="Figures 39 - 52. Female subgenital plate in ventral view 39 Eupholidoptera smyrnensis Makrigiannis RMNH. 5014918 40 Eupholidoptera gemellata Mt. Idhi, FC 1602 CT 2000.095.02 41 Eupholidoptera pallipes allotype Mt. Lefka RMNH. INS 1105312 42 Eupholidoptera cretica Mt. Lefka, Omalos RMNH. INS 1141837 43 Eupholidoptera latens Mt. Lefka, Kalergi CT 1987.047.04 44 Eupholidoptera giuliae Argoules CT 1995.011.03 45 Eupholidoptera francisae sp. nov. allotype Andikithira CT 2002.004.11 46 Eupholidoptera annamariae Kato Zakros CT 1995.020.03 47 Eupholidoptera astyla Krotos RMNH. INS 1141820 48 Eupholidoptera feri allotype Mt. Dikti, Katharo plain RMNH. INS 1105298 49 Eupholidoptera mariannae Kalavros RMNH 5014912 50 Eupholidoptera forcipata Mt. Idhi, FC 1602 RMNH. INS 1141845 51 Eupholidoptera marietheresae sp. nov. allotype Mt. Dikti RMNH. INS 1141849 52 Eupholidoptera jacquelinae Gavdos allotype CT 2001.004.13. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures39-52" httpUri="https://binary.pensoft.net/fig/817937" pageId="0" pageNumber="67">, 49</figureCitation>
<figureCitation captionStart="Figures 53–66" captionStartId="F8" captionText="Figures 53 - 66. Female subgenital plate in lateral view 53 Eupholidoptera smyrnensis Makrigiannis RMNH. 5014918 54 Eupholidoptera gemellata Mt. Idhi, FC 1602 CT 2000.095.02 55 Eupholidoptera pallipes allotype Mt. Lefka RMNH. INS 1105312 56 Eupholidoptera cretica Mt. Lefka, Omalos RMNH. INS 1141837 57 Eupholidoptera latens Mt. Lefka, Kalergi CT 1987.047.04 58 Eupholidoptera giuliae Argoules CT 1995.011.03 59 Eupholidoptera francisae sp. nov. allotype Andikithira CT 2002.004.11 60 Eupholidoptera annamariae Kato Zakros CT 1995.020.03 61 Eupholidoptera astyla Krotos RMNH. INS 1141820 62 Eupholidoptera feri allotype Mt. Dikti, Katharo plain RMNH. INS 1105298 63 Eupholidoptera mariannae Kalavros RMNH 5014912 64 Eupholidoptera forcipata Mt. Idhi, FC 1602 RMNH. INS 1141845 65 Eupholidoptera marietheresae sp. nov. allotype Mt. Dikti RMNH. INS 1141849 66 Eupholidoptera jacquelinae allotype Gavdos CT 2001.004.13. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures53-66" httpUri="https://binary.pensoft.net/fig/817938" pageId="0" pageNumber="67">, 63</figureCitation>
<figureCitation captionStart="Figures 69–82" captionStartId="F10" captionText="Figures 69 - 82. Male anal tergite in dorsal view 69 Eupholidoptera smyrnensis Makrigiannis RMNH. INS 152909 70 Eupholidoptera gemellata holotype Mt. Idhi RMNH. INS 1105300 71 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105313 72 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 73 Eupholidoptera latens Mt. Lefka, Kalergi CT 2003.009.01 74 Eupholidoptera giuliae Chora Sfakion CT 2000.024.04 75 Eupholidoptera francisae sp. nov. holotype Andikithira CT 2002.004.04 76 Eupholidoptera annamariae Kato Zakros CT 2000.030.04 77 Eupholidoptera astyla Krotos RMNH. INS 1141819 78 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 79 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 80 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 81 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 82 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures69-82" httpUri="https://binary.pensoft.net/fig/817940" pageId="0" pageNumber="67">, 79</figureCitation>
<figureCitation captionStart="Figures 83–96" captionStartId="F11" captionText="Figures 83 - 96. Male anal tergite in caudal view 83 Eupholidoptera smyrnensis Makrigiannis RMNH. INS 152909 84 Eupholidoptera gemellata Mt. Idhi CT 2000.095.01 85 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 86 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 87 Eupholidoptera latens Mt. Lefka, Kalergi CT 2003.009.01 88 Eupholidoptera giuliae Chora Sfakion CT 2000.014.01 89 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.02 90 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 91 Eupholidoptera astyla Krotos RMNH. INS 1141819 92 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 93 Eupholidoptera mariannae Prina FC 17798 RMNH. INS 1141840 94 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 95 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 96 Eupholidoptera jacquelinae paratype Gavdos NHMC 2001.004.12. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures83-96" httpUri="https://binary.pensoft.net/fig/817941" pageId="0" pageNumber="67">, 93</figureCitation>
<figureCitation captionStart="Figures 97–110" captionStartId="F12" captionText="Figures 97 - 110. Male anal tergite in lateral view 97 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 98 Eupholidoptera gemellata Mt. Idhi CT 2000.095.01 99 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 100 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 101 Eupholidoptera latens Mt. Lefka, Kalergi CT 1987.047.01 102 Eupholidoptera giuliae Argoules CT 1995.011.11 103 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.02 104 Eupholidoptera annamariae Kato Zakros CT 2000.030.03 105 Eupholidoptera astyla Ano Viannos CT 1995.024.01 106 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 107 Eupholidoptera mariannae Kalavros CT 2017.029.01 108 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 109 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 110 Eupholidoptera jacquelinae Gavdopoula CT 1996.019.01. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures97-110" httpUri="https://binary.pensoft.net/fig/817942" pageId="0" pageNumber="67">, 107</figureCitation>
<figureCitation captionStart="Figures 111–125" captionStartId="F13" captionText="Figures 111 - 125. Male cercus in dorsal view 111 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 112 Eupholidoptera gemellata Mt. Idhi FC 1651 RMNH. INS 1141843 113 Eupholidoptera pallipes Mt. Lefka paratype RMNH. INS 1105314 114 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 115 Eupholidoptera latens Xiloskalo IBER DC-Ort 000558 116 Eupholidoptera giuliae Chora Sfakion CT 2000.014.02 117 Eupholidoptera francisae sp. nov. holotype Andikithira CT 2002.004.04 118 Eupholidoptera annamariae Kato Zakros CT 2000.030.03 119 Eupholidoptera annamariae Xerokampos, male with anomalous cercus, IBER DC-Ort 000565 120 Eupholidoptera astyla Ierapetra MfN s. n. 121 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 122 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 123 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 124 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 125 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures111-125" httpUri="https://binary.pensoft.net/fig/817943" pageId="0" pageNumber="67">, 122</figureCitation>
<figureCitation captionStart="Figures 126–139" captionStartId="F14" captionText="Figures 126 - 139. Male cercus in lateral view 126 Eupholidoptera smyrnensis Makrigiannis RMNH. 5087053 127 Eupholidoptera gemellata Mt. Idhi FC 1651 RMNH. INS 1141843 128 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 129 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 130 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 131 Eupholidoptera giuliae Chora Sfakion CT 2000.024.04 132 Eupholidoptera francisae paratype sp. nov. Andikithira CT 2002.004.08 133 Eupholidoptera annamariae Kato Zakros CT 1995.020.02 134 Eupholidoptera astyla paratype Ierapetra MfN s. n. 135 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 136 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 137 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 138 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 139 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures126-139" httpUri="https://binary.pensoft.net/fig/817944" pageId="0" pageNumber="67">, 136</figureCitation>
<figureCitation captionStart="Figures 140–153" captionStartId="F15" captionText="Figures 140 - 153. Male subgenital plate in ventral view 140 Eupholidoptera smyrnensis Lagkada RMNH 5106270 141 Eupholidoptera gemellata Mt. Idhi FC 1651 RMNH. INS 1141843 142 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105313 143 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 144 Eupholidoptera latens Mt. Lefka Kalergi CT 1987.047.02 145 Eupholidoptera giuliae Skaloti IBER DC-Ort 000564 146 Eupholidoptera francisae sp. nov. holotype Andikithira CT 2002.004.04 147 Eupholidoptera annamariae Kato Zakros CT 2000.030.04 148 Eupholidoptera astyla Krotos RMNH. INS 1141819 149 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 150 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 151 Eupholidoptera forcipata Mt. Idhi CT 1987.044.03 152 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 153 Eupholidoptera jacquelinae holotype Gavdos 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures140-153" httpUri="https://binary.pensoft.net/fig/817945" pageId="0" pageNumber="67">, 150</figureCitation>
<figureCitation captionStart="Figures 154–167" captionStartId="F16" captionText="Figures 154 - 167. Male subgenital plate in lateral view 154 Eupholidoptera smyrnensis Lagkada RMNH. INS 152909 155 Eupholidoptera gemellata holotype Mt. Idhi RMNH. INS 1105300 156 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105313 157 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 (RMNH. INS 1141838 158 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 159 Eupholidoptera giuliae Skaloti IBER DC-Ort 000564 160 Eupholidoptera francisae sp. nov. holotype Andikithira CT 2002.004.04 161 Eupholidoptera annamariae Kato Zakros CT 2000.030.04 162 Eupholidoptera astyla Krotos RMNH 5086992 163 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 164 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 165 Eupholidoptera forcipata Mt. Idhi CT 1987.044.03 166 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 167 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures154-167" httpUri="https://binary.pensoft.net/fig/817946" pageId="0" pageNumber="67">, 164</figureCitation>
<figureCitation captionStart="Figures 168–181" captionStartId="F17" captionText="Figures 168 - 181. Male stylus in lateral view 168 Eupholidoptera smyrnensis Makrigiannis RMNH. INS 152909 169 Eupholidoptera gemellata holotype Mt. Idhi RMNH. INS 1105300 170 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105313 171 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 172 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 173 Eupholidoptera giuliae Skaloti IBER DC-Ort 000564 174 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 175 Eupholidoptera annamariae Kato Zakros CT 2000.030.04 176 Eupholidoptera astyla Krotos RMNH. INS 1141819 177 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 178 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 179 Eupholidoptera forcipata Mt. Idhi CT 1987.044.03 180 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 181 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures168-181" httpUri="https://binary.pensoft.net/fig/817947" pageId="0" pageNumber="67">, 178</figureCitation>
<figureCitation captionStart="Figures 182–197" captionStartId="F18" captionText="Figures 182 - 197. Male titillator in ventral view 182 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 183 Eupholidoptera gemellata Mt. Idhi Amariou FC 1651 RMNH. INS 1141843 184 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 185 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 186 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 187 Eupholidoptera latens Rhodopos CH 8236 188 Eupholidoptera giuliae Argoules CT 1995.011.11 189 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 190 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 191 Eupholidoptera astyla Krotos RMNH. INS 1141819 192 Eupholidoptera astyla Kofinas FC 460 RMNH. INS 1141836 193 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 194 Eupholidoptera mariannae paratype Malles CH 2906 A 195 Eupholidoptera forcipata Mt. Idhi CT 2019.047.01 196 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 197 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures182-197" httpUri="https://binary.pensoft.net/fig/817948" pageId="0" pageNumber="67">, 194</figureCitation>
<figureCitation captionStart="Figures 198–212" captionStartId="F19" captionText="Figures 198 - 212. Male titillator in lateral view 198 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 199 Eupholidoptera gemellata Mt. Idhi Amariou FC 1651 RMNH. INS 1141843 200 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 201 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 202 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 203 Eupholidoptera latens Rhodopos CH 8236 204 Eupholidoptera giuliae Argoules CT 1995.011.11 205 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 206 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 207 Eupholidoptera astyla Krotos RMNH. INS 1141819 208 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 209 Eupholidoptera mariannae Kalavros CT 2017.029.01 210 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 211 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 212 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures198-212" httpUri="https://binary.pensoft.net/fig/817949" pageId="0" pageNumber="67">, 209</figureCitation>
<figureCitation captionStart="Figures 213–215" captionStartId="F20" captionText="Figures 213 - 215. Eupholidoptera astyla paratypes 213 titillator in dorsal view Eupholidoptera astyla ♂ paratype Ierapetra s. n. 214 subgenital plate in ventral view Eupholidoptera mariannae ♀ Anadoli s. n. [paratype of Eupholidoptera astyla] 215 Eupholidoptera mariannae ♀ Kato Chorion s. n. [paratype of Eupholidoptera astyla]." figureDoi="10.3897/zookeys.1151.97514.figures213-215" httpUri="https://binary.pensoft.net/fig/817950" pageId="0" pageNumber="67">, 214</figureCitation>
<figureCitation captionStart="Figures 213–215" captionStartId="F20" captionText="Figures 213 - 215. Eupholidoptera astyla paratypes 213 titillator in dorsal view Eupholidoptera astyla ♂ paratype Ierapetra s. n. 214 subgenital plate in ventral view Eupholidoptera mariannae ♀ Anadoli s. n. [paratype of Eupholidoptera astyla] 215 Eupholidoptera mariannae ♀ Kato Chorion s. n. [paratype of Eupholidoptera astyla]." figureDoi="10.3897/zookeys.1151.97514.figures213-215" httpUri="https://binary.pensoft.net/fig/817950" pageId="0" pageNumber="67">, 215</figureCitation>
<figureCitation captionStart="Figures 224–235" captionStartId="F24" captionText="Figures 224 - 235. Habitus Eupholidoptera spp. in dorsal and lateral view 224, 225 Eupholidoptera astyla ♀ Mt. Idhi CT 1987.046.03 226, 227 Eupholidoptera cretica ♀ Mt. Lefka Omalos FC 17807 RMNH. INS 1141837 228, 229 Eupholidoptera francisae sp. nov. ♂ holotype Andikithira CT 2002.004.04 230, 231 Eupholidoptera francisae sp. nov. ♀ allotype Andikithira CT 2002.004.11 232, 233 Eupholidoptera gemellata ♀ Mt. Idhi FC 1651 RMNH. INS 1141844 234, 235 Eupholidoptera mariannae ♀ Ag. Ioannis RMNH 5014906 1844. Scale bars: 10 mm." figureDoi="10.3897/zookeys.1151.97514.figures224-235" httpUri="https://binary.pensoft.net/fig/817954" pageId="0" pageNumber="67">, 234</figureCitation>
<figureCitation captionStart="Figures 224–235" captionStartId="F24" captionText="Figures 224 - 235. Habitus Eupholidoptera spp. in dorsal and lateral view 224, 225 Eupholidoptera astyla ♀ Mt. Idhi CT 1987.046.03 226, 227 Eupholidoptera cretica ♀ Mt. Lefka Omalos FC 17807 RMNH. INS 1141837 228, 229 Eupholidoptera francisae sp. nov. ♂ holotype Andikithira CT 2002.004.04 230, 231 Eupholidoptera francisae sp. nov. ♀ allotype Andikithira CT 2002.004.11 232, 233 Eupholidoptera gemellata ♀ Mt. Idhi FC 1651 RMNH. INS 1141844 234, 235 Eupholidoptera mariannae ♀ Ag. Ioannis RMNH 5014906 1844. Scale bars: 10 mm." figureDoi="10.3897/zookeys.1151.97514.figures224-235" httpUri="https://binary.pensoft.net/fig/817954" pageId="0" pageNumber="67">, 235</figureCitation>
<figureCitation captionStart="Figures 247–253" captionStartId="F27" captionText="Figures 247 - 253. Field images Eupholidoptera spp. Crete 247 Eupholidoptera giuliae ♂ Prasies RMNH 5087051 248 Eupholidoptera giuliae ♀ Rethymnon RMNH 5014921 249 Eupholidoptera latens ♂ Kolympari RMNH 553681 250 Eupholidoptera mariannae ♂ Kalavros RMNH 5014907 251 Eupholidoptera mariannae ♀ Ag. Ioannis RMNH 5014906 252 Eupholidoptera pallipes ♀ below Pakhnes 253 Eupholidoptera smyrnensis ♂ Makrigiannis RMNH 5087053." figureDoi="10.3897/zookeys.1151.97514.figures247-253" httpUri="https://binary.pensoft.net/fig/817957" pageId="0" pageNumber="67">, 250</figureCitation>
<figureCitation captionStart="Figures 247–253" captionStartId="F27" captionText="Figures 247 - 253. Field images Eupholidoptera spp. Crete 247 Eupholidoptera giuliae ♂ Prasies RMNH 5087051 248 Eupholidoptera giuliae ♀ Rethymnon RMNH 5014921 249 Eupholidoptera latens ♂ Kolympari RMNH 553681 250 Eupholidoptera mariannae ♂ Kalavros RMNH 5014907 251 Eupholidoptera mariannae ♀ Ag. Ioannis RMNH 5014906 252 Eupholidoptera pallipes ♀ below Pakhnes 253 Eupholidoptera smyrnensis ♂ Makrigiannis RMNH 5087053." figureDoi="10.3897/zookeys.1151.97514.figures247-253" httpUri="https://binary.pensoft.net/fig/817957" pageId="0" pageNumber="67">, 251</figureCitation>
<figureCitation captionStart="Figures 254, 255" captionStartId="F28" captionText="Figures 254, 255. Distribution maps Eupholidoptera Crete and adjacent islands 254 lowland species 255 highland species." figureDoi="10.3897/zookeys.1151.97514.figures254-255" httpUri="https://binary.pensoft.net/fig/817958" pageId="0" pageNumber="67">, 254</figureCitation>
<figureCitation captionStart="Figure 256" captionStartId="F29" captionText="Figure 256. Bayesian inference phylogenetic tree of Eupholidoptera of Crete, Andikithira and Gavdos, including two outgroups and congenerics from the Greek and Turkish mainlands, based on a 1684 bp concatenated alignment of the NADH 2 + ITS fragments. Node values at branches show node support with BI posterior probabilities. Different colours correspond to the different morphological species. See Suppl. material 4 for details of the specimens and Suppl. material 5 for details of NADH 2 and IT IS sequences." figureDoi="10.3897/zookeys.1151.97514.figure256" httpUri="https://binary.pensoft.net/fig/817959" pageId="0" pageNumber="67">, 256</figureCitation>
<figureCitation captionStart="Figure 259" captionStartId="F32" captionText="Figure 259. Mean syllable duration in recorded species from Crete (incl. Gavdos and Andikithira). See Suppl. material 3 for measurements per recorded specimen." figureDoi="10.3897/zookeys.1151.97514.figure259" httpUri="https://binary.pensoft.net/fig/817962" pageId="0" pageNumber="67">, 259</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67" type="reference_group">
<paragraph pageId="0" pageNumber="67">
<taxonomicName authorityName="Willemse &amp; Heller" authorityYear="2001" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupholidoptera mariannae" order="Orthoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="mariannae">Eupholidoptera mariannae</taxonomicName>
Willemse &amp; Heller, 2001:
<bibRefCitation author="Willemse, L" journalOrPublisher="EIS Kenniscentrum Insecten and Naturalis Biodiversity Center, Leiden" pageId="0" pageNumber="67" refId="B80" refString="Willemse, L, Kleukers, R, Ode, B, 2018. The Grasshoppers of Greece. EIS Kenniscentrum Insecten and Naturalis Biodiversity Center, Leiden" title="The Grasshoppers of Greece." year="2018">Willemse et al. 2018</bibRefCitation>
: figs 946, 947.
</paragraph>
<paragraph pageId="0" pageNumber="67">
<taxonomicName authorityName="Willemse &amp; Heller" authorityYear="2001" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupholidoptera mariannae" order="Orthoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="mariannae">Eupholidoptera mariannae</taxonomicName>
Morphological description.
<bibRefCitation DOI="https://doi.org/10.1163/22119434-900000092" author="Willemse, F" journalOrPublisher="Tijdschrift voor Entomologie" pageId="0" pageNumber="67" pagination="329 - 343" refId="B76" refString="Willemse, F, Heller, K-G, 2001. Two new species of Eupholidoptera Maran (Orthoptera, Tettigoniidae) from Crete with a checklist and key to the species. Tijdschrift voor Entomologie 144 (2): 329 - 343, DOI: https://doi.org/10.1163/22119434-900000092" title="Two new species of Eupholidoptera Maran (Orthoptera, Tettigoniidae) from Crete with a checklist and key to the species." url="https://doi.org/10.1163/22119434-900000092" volume="144" year="2001">Willemse and Heller 2001</bibRefCitation>
: 329-331.
</paragraph>
<paragraph pageId="0" pageNumber="67">
<taxonomicName authorityName="Willemse &amp; Heller" authorityYear="2001" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupholidoptera mariannae" order="Orthoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="mariannae">Eupholidoptera mariannae</taxonomicName>
Bioacoustics.
<bibRefCitation DOI="https://doi.org/10.1163/22119434-900000092" author="Willemse, F" journalOrPublisher="Tijdschrift voor Entomologie" pageId="0" pageNumber="67" pagination="329 - 343" refId="B76" refString="Willemse, F, Heller, K-G, 2001. Two new species of Eupholidoptera Maran (Orthoptera, Tettigoniidae) from Crete with a checklist and key to the species. Tijdschrift voor Entomologie 144 (2): 329 - 343, DOI: https://doi.org/10.1163/22119434-900000092" title="Two new species of Eupholidoptera Maran (Orthoptera, Tettigoniidae) from Crete with a checklist and key to the species." url="https://doi.org/10.1163/22119434-900000092" volume="144" year="2001">Willemse and Heller 2001</bibRefCitation>
: 331;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2156.1.1" author="Ciplak, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="67" pagination="1 - 77" refId="B12" refString="Ciplak, B, Heller, K-G, Willemse, F, 2009. Review of the genus Eupholidoptera (Orthoptera, Tettigoniidae): Different genitalia, uniform song. Zootaxa 2156 (1): 1 - 77, DOI: https://doi.org/10.11646/zootaxa.2156.1.1" title="Review of the genus Eupholidoptera (Orthoptera, Tettigoniidae): Different genitalia, uniform song." url="https://doi.org/10.11646/zootaxa.2156.1.1" volume="2156" year="2009">
<normalizedToken originalValue="Çiplak">Ciplak</normalizedToken>
et al. 2009
</bibRefCitation>
: 27, 54, 55.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67" type="examined specimens">
<paragraph pageId="0" pageNumber="67">Examined specimens.</paragraph>
<paragraph pageId="0" pageNumber="67">
<emphasis bold="true" pageId="0" pageNumber="67">Holotype</emphasis>
, 1 ♂ (
<emphasis bold="true" pageId="0" pageNumber="67">paratype</emphasis>
); 8 ♂, 15 ♀ (for details see Suppl. material 2).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67" type="diagnostic features">
<paragraph pageId="0" pageNumber="67">Diagnostic features.</paragraph>
<paragraph pageId="0" pageNumber="67">
Frons (Fig.
<figureCitation captionStart="Figures 11–24" captionStartId="F5" captionText="Figures 11 - 24. Colour pattern of head, frontal view 11 Eupholidoptera smyrnensis ♀ Makrigiannis RMNH 5014918 12 Eupholidoptera gemellata ♂ Mt. Idhi RMNH. INS 1141843 13 Eupholidoptera pallipes ♂ paratype Mt. Lefka RMNH. INS 1105313 14 Eupholidoptera cretica ♂ Mt. Lefka, Omalos RMNH. INS 114838 15 Eupholidoptera latens ♀ Mt. Lefka, Kalergi CT 1987.047.04 16 Eupholidoptera giuliae ♂ Argoules CT 1995.011.18 17 Eupholidoptera francisae sp. nov. ♀ allotype Andikithira CT 2002.004.11 18 Eupholidoptera annamariae ♀ Kato Zakros CT 1995.020.03 19 Eupholidoptera astyla ♀ Ano Viannos CT 1987.024.03 20 Eupholidoptera feri ♂ holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 21 Eupholidoptera mariannae ♀ Ag. Ioannis RMNH. 5014906 22 Eupholidoptera forcipata ♂ Mt. Idhi CT 1987.044.03 23 Eupholidoptera marietheresae sp. nov. ♂ paratype Mt. Dikti, FC 1606 CT 2000.096.01 24 Eupholidoptera jacquelinae ♂ holotype Gavdos CT 2000.005.02. Scale bars: 5 mm." figureDoi="10.3897/zookeys.1151.97514.figures11-24" httpUri="https://binary.pensoft.net/fig/817935" pageId="0" pageNumber="67">21</figureCitation>
) pale with black dots; frontal half of pronotal disc (Fig.
<figureCitation captionStart="Figures 25–38" captionStartId="F6" captionText="Figures 25 - 38. Colour pattern of pronotum, dorsal view 25 Eupholidoptera smyrnensis ♂ Makrigiannis RMNH. INS 152909 26 Eupholidoptera gemellata ♂ Mt. Idhi, Amariou RMNH. INS 1141843 27 Eupholidoptera pallipes ♂ paratype Mt. Lefka RMNH. INS 1105313 28 Eupholidoptera cretica ♂ Mt. Lefka, Omalos RMNH. INS 1141838 29 Eupholidoptera latens ♂ Rhodopos CH 8236 30 Eupholidoptera giuliae Chora Sfakion ♂ CT 2000.014.10 31 Eupholidoptera francisae sp. nov. ♂ holotype Andikithira CT 2002.004.04 32 Eupholidoptera annamariae ♂ Kato Zakros CT 2000.030.04 33 Eupholidoptera astyla ♂ Krotos RMNH. INS 1141819 34 Eupholidoptera feri ♂ holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 35 Eupholidoptera mariannae ♀ Kalavros RMNH. 5086974 36 Eupholidoptera forcipata ♂ Mt. Idhi CT 1987.044.03 37 Eupholidoptera marietheresae sp. nov. ♂ holotype Mt. Dikti RMNH. INS 1141850 38 Eupholidoptera jacquelinae ♂ holotype Gavdos CT 2000.005.02. Scale bars: 5 mm." figureDoi="10.3897/zookeys.1151.97514.figures25-38" httpUri="https://binary.pensoft.net/fig/817936" pageId="0" pageNumber="67">35</figureCitation>
) predominantly black sharply with transverse rarely a V-shaped border with pale rear half; elytra black, veins and cross-veins more or less extensively yellow. Male (Fig.
<figureCitation captionStart="Figures 247–253" captionStartId="F27" captionText="Figures 247 - 253. Field images Eupholidoptera spp. Crete 247 Eupholidoptera giuliae ♂ Prasies RMNH 5087051 248 Eupholidoptera giuliae ♀ Rethymnon RMNH 5014921 249 Eupholidoptera latens ♂ Kolympari RMNH 553681 250 Eupholidoptera mariannae ♂ Kalavros RMNH 5014907 251 Eupholidoptera mariannae ♀ Ag. Ioannis RMNH 5014906 252 Eupholidoptera pallipes ♀ below Pakhnes 253 Eupholidoptera smyrnensis ♂ Makrigiannis RMNH 5087053." figureDoi="10.3897/zookeys.1151.97514.figures247-253" httpUri="https://binary.pensoft.net/fig/817957" pageId="0" pageNumber="67">250</figureCitation>
) - stridulatory file with 89-105 teeth (90 in
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2156.1.1" author="Ciplak, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="67" pagination="1 - 77" refId="B12" refString="Ciplak, B, Heller, K-G, Willemse, F, 2009. Review of the genus Eupholidoptera (Orthoptera, Tettigoniidae): Different genitalia, uniform song. Zootaxa 2156 (1): 1 - 77, DOI: https://doi.org/10.11646/zootaxa.2156.1.1" title="Review of the genus Eupholidoptera (Orthoptera, Tettigoniidae): Different genitalia, uniform song." url="https://doi.org/10.11646/zootaxa.2156.1.1" volume="2156" year="2009">
<normalizedToken originalValue="Çiplak">Ciplak</normalizedToken>
et al. 2009
</bibRefCitation>
) (including proximal and distal ones), density of teeth in middle two thirds of the file 18-23 teeth per mm (18-20 in
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2156.1.1" author="Ciplak, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="67" pagination="1 - 77" refId="B12" refString="Ciplak, B, Heller, K-G, Willemse, F, 2009. Review of the genus Eupholidoptera (Orthoptera, Tettigoniidae): Different genitalia, uniform song. Zootaxa 2156 (1): 1 - 77, DOI: https://doi.org/10.11646/zootaxa.2156.1.1" title="Review of the genus Eupholidoptera (Orthoptera, Tettigoniidae): Different genitalia, uniform song." url="https://doi.org/10.11646/zootaxa.2156.1.1" volume="2156" year="2009">
<normalizedToken originalValue="Çiplak">Ciplak</normalizedToken>
et al. 2009
</bibRefCitation>
); anal tergite (Figs
<figureCitation captionStart="Figures 69–82" captionStartId="F10" captionText="Figures 69 - 82. Male anal tergite in dorsal view 69 Eupholidoptera smyrnensis Makrigiannis RMNH. INS 152909 70 Eupholidoptera gemellata holotype Mt. Idhi RMNH. INS 1105300 71 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105313 72 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 73 Eupholidoptera latens Mt. Lefka, Kalergi CT 2003.009.01 74 Eupholidoptera giuliae Chora Sfakion CT 2000.024.04 75 Eupholidoptera francisae sp. nov. holotype Andikithira CT 2002.004.04 76 Eupholidoptera annamariae Kato Zakros CT 2000.030.04 77 Eupholidoptera astyla Krotos RMNH. INS 1141819 78 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 79 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 80 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 81 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 82 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures69-82" httpUri="https://binary.pensoft.net/fig/817940" pageId="0" pageNumber="67">79</figureCitation>
,
<figureCitation captionStart="Figures 83–96" captionStartId="F11" captionText="Figures 83 - 96. Male anal tergite in caudal view 83 Eupholidoptera smyrnensis Makrigiannis RMNH. INS 152909 84 Eupholidoptera gemellata Mt. Idhi CT 2000.095.01 85 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 86 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 87 Eupholidoptera latens Mt. Lefka, Kalergi CT 2003.009.01 88 Eupholidoptera giuliae Chora Sfakion CT 2000.014.01 89 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.02 90 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 91 Eupholidoptera astyla Krotos RMNH. INS 1141819 92 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 93 Eupholidoptera mariannae Prina FC 17798 RMNH. INS 1141840 94 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 95 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 96 Eupholidoptera jacquelinae paratype Gavdos NHMC 2001.004.12. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures83-96" httpUri="https://binary.pensoft.net/fig/817941" pageId="0" pageNumber="67">93</figureCitation>
,
<figureCitation captionStart="Figures 97–110" captionStartId="F12" captionText="Figures 97 - 110. Male anal tergite in lateral view 97 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 98 Eupholidoptera gemellata Mt. Idhi CT 2000.095.01 99 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 100 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 101 Eupholidoptera latens Mt. Lefka, Kalergi CT 1987.047.01 102 Eupholidoptera giuliae Argoules CT 1995.011.11 103 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.02 104 Eupholidoptera annamariae Kato Zakros CT 2000.030.03 105 Eupholidoptera astyla Ano Viannos CT 1995.024.01 106 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 107 Eupholidoptera mariannae Kalavros CT 2017.029.01 108 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 109 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 110 Eupholidoptera jacquelinae Gavdopoula CT 1996.019.01. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures97-110" httpUri="https://binary.pensoft.net/fig/817942" pageId="0" pageNumber="67">107</figureCitation>
) very wide, bilobed, lobes separated by groove, distally bend downward, centrally forming two teeth pointing forward separated by narrow, densely haired pits; cerci (Figs
<figureCitation captionStart="Figures 111–125" captionStartId="F13" captionText="Figures 111 - 125. Male cercus in dorsal view 111 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 112 Eupholidoptera gemellata Mt. Idhi FC 1651 RMNH. INS 1141843 113 Eupholidoptera pallipes Mt. Lefka paratype RMNH. INS 1105314 114 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 115 Eupholidoptera latens Xiloskalo IBER DC-Ort 000558 116 Eupholidoptera giuliae Chora Sfakion CT 2000.014.02 117 Eupholidoptera francisae sp. nov. holotype Andikithira CT 2002.004.04 118 Eupholidoptera annamariae Kato Zakros CT 2000.030.03 119 Eupholidoptera annamariae Xerokampos, male with anomalous cercus, IBER DC-Ort 000565 120 Eupholidoptera astyla Ierapetra MfN s. n. 121 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 122 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 123 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 124 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 125 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures111-125" httpUri="https://binary.pensoft.net/fig/817943" pageId="0" pageNumber="67">122</figureCitation>
,
<figureCitation captionStart="Figures 126–139" captionStartId="F14" captionText="Figures 126 - 139. Male cercus in lateral view 126 Eupholidoptera smyrnensis Makrigiannis RMNH. 5087053 127 Eupholidoptera gemellata Mt. Idhi FC 1651 RMNH. INS 1141843 128 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 129 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 130 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 131 Eupholidoptera giuliae Chora Sfakion CT 2000.024.04 132 Eupholidoptera francisae paratype sp. nov. Andikithira CT 2002.004.08 133 Eupholidoptera annamariae Kato Zakros CT 1995.020.02 134 Eupholidoptera astyla paratype Ierapetra MfN s. n. 135 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 136 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 137 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 138 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 139 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures126-139" httpUri="https://binary.pensoft.net/fig/817944" pageId="0" pageNumber="67">136</figureCitation>
) 3
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, proximally wide, flattened, strongly narrowing in second third, apical third cylindrical, pointing outward, in profile bent upward, armed with short, strong inner side-tooth at ca. one fifth of length; subgenital plate (Figs
<figureCitation captionStart="Figures 140–153" captionStartId="F15" captionText="Figures 140 - 153. Male subgenital plate in ventral view 140 Eupholidoptera smyrnensis Lagkada RMNH 5106270 141 Eupholidoptera gemellata Mt. Idhi FC 1651 RMNH. INS 1141843 142 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105313 143 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 144 Eupholidoptera latens Mt. Lefka Kalergi CT 1987.047.02 145 Eupholidoptera giuliae Skaloti IBER DC-Ort 000564 146 Eupholidoptera francisae sp. nov. holotype Andikithira CT 2002.004.04 147 Eupholidoptera annamariae Kato Zakros CT 2000.030.04 148 Eupholidoptera astyla Krotos RMNH. INS 1141819 149 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 150 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 151 Eupholidoptera forcipata Mt. Idhi CT 1987.044.03 152 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 153 Eupholidoptera jacquelinae holotype Gavdos 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures140-153" httpUri="https://binary.pensoft.net/fig/817945" pageId="0" pageNumber="67">150</figureCitation>
,
<figureCitation captionStart="Figures 154–167" captionStartId="F16" captionText="Figures 154 - 167. Male subgenital plate in lateral view 154 Eupholidoptera smyrnensis Lagkada RMNH. INS 152909 155 Eupholidoptera gemellata holotype Mt. Idhi RMNH. INS 1105300 156 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105313 157 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 (RMNH. INS 1141838 158 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 159 Eupholidoptera giuliae Skaloti IBER DC-Ort 000564 160 Eupholidoptera francisae sp. nov. holotype Andikithira CT 2002.004.04 161 Eupholidoptera annamariae Kato Zakros CT 2000.030.04 162 Eupholidoptera astyla Krotos RMNH 5086992 163 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 164 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 165 Eupholidoptera forcipata Mt. Idhi CT 1987.044.03 166 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 167 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures154-167" httpUri="https://binary.pensoft.net/fig/817946" pageId="0" pageNumber="67">164</figureCitation>
) as wide as long, widest in proximal third, sides rimmed, in profile pointing upward and backward, tip apical lobes rounded, spineless, with slit-like excision along one fifth of length; styli (Fig.
<figureCitation captionStart="Figures 168–181" captionStartId="F17" captionText="Figures 168 - 181. Male stylus in lateral view 168 Eupholidoptera smyrnensis Makrigiannis RMNH. INS 152909 169 Eupholidoptera gemellata holotype Mt. Idhi RMNH. INS 1105300 170 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105313 171 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 172 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 173 Eupholidoptera giuliae Skaloti IBER DC-Ort 000564 174 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 175 Eupholidoptera annamariae Kato Zakros CT 2000.030.04 176 Eupholidoptera astyla Krotos RMNH. INS 1141819 177 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 178 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 179 Eupholidoptera forcipata Mt. Idhi CT 1987.044.03 180 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 181 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures168-181" httpUri="https://binary.pensoft.net/fig/817947" pageId="0" pageNumber="67">178</figureCitation>
) minute, flat or depressed, inserted at ventral side of apical lobes, proximal of tip, pointing downward; titillator (Fig.
<figureCitation captionStart="Figures 182–197" captionStartId="F18" captionText="Figures 182 - 197. Male titillator in ventral view 182 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 183 Eupholidoptera gemellata Mt. Idhi Amariou FC 1651 RMNH. INS 1141843 184 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 185 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 186 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 187 Eupholidoptera latens Rhodopos CH 8236 188 Eupholidoptera giuliae Argoules CT 1995.011.11 189 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 190 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 191 Eupholidoptera astyla Krotos RMNH. INS 1141819 192 Eupholidoptera astyla Kofinas FC 460 RMNH. INS 1141836 193 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 194 Eupholidoptera mariannae paratype Malles CH 2906 A 195 Eupholidoptera forcipata Mt. Idhi CT 2019.047.01 196 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 197 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures182-197" httpUri="https://binary.pensoft.net/fig/817948" pageId="0" pageNumber="67">194</figureCitation>
,
<figureCitation captionStart="Figures 198–212" captionStartId="F19" captionText="Figures 198 - 212. Male titillator in lateral view 198 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 199 Eupholidoptera gemellata Mt. Idhi Amariou FC 1651 RMNH. INS 1141843 200 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 201 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 202 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 203 Eupholidoptera latens Rhodopos CH 8236 204 Eupholidoptera giuliae Argoules CT 1995.011.11 205 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 206 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 207 Eupholidoptera astyla Krotos RMNH. INS 1141819 208 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 209 Eupholidoptera mariannae Kalavros CT 2017.029.01 210 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 211 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 212 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures198-212" httpUri="https://binary.pensoft.net/fig/817949" pageId="0" pageNumber="67">209</figureCitation>
) slightly asymmetrical, apical arms strongly sclerotised, narrow, except for very tip, fused along entire length, smooth, needle shaped tip pointing somewhat laterad, in profile narrow, middle third slightly wider evenly upward curved.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67" type="description">
<paragraph pageId="0" pageNumber="67">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="67">
See Tables
<tableCitation captionStart="Table 6" captionStartId="T6" captionText="Table 6. Measurements (in mm), ratios, and biometrics of male Eupholidoptera." httpUri="http://table.plazi.org/id/8B3AFDE00956AD4F11FF9ECA2A2FA78F" pageId="0" pageNumber="67" tableUuid="8B3AFDE00956AD4F11FF9ECA2A2FA78F">6</tableCitation>
,
<tableCitation captionStart="Table 7" captionStartId="T7" captionText="Table 7. Measurements (in mm), ratios, and biometrics of female Eupholidoptera." httpUri="http://table.plazi.org/id/3ABD8C0FD5A9E22855CC92BF15EE30E0" pageId="0" pageNumber="67" tableUuid="3ABD8C0FD5A9E22855CC92BF15EE30E0">7</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67" type="bioacoustics">
<paragraph pageId="0" pageNumber="67">Bioacoustics.</paragraph>
<paragraph pageId="0" pageNumber="67">
Based upon the sound recordings of two specimens (20 syllables measured), the song of
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. mariannae" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="mariannae">
<emphasis italics="true" pageId="0" pageNumber="67">E. mariannae</emphasis>
</taxonomicName>
, as in all species of
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupholidoptera" order="Orthoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="67">Eupholidoptera</emphasis>
</taxonomicName>
, consists of isolated syllables produced in long series with the opening hemisyllable much shorter and weaker than the closing hemisyllable. In
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. mariannae" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="mariannae">
<emphasis italics="true" pageId="0" pageNumber="67">E. mariannae</emphasis>
</taxonomicName>
, the syllable duration is ~ 172 ms. In the present recordings, the syllable repetition rate is very low. Published records (
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2156.1.1" author="Ciplak, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="67" pagination="1 - 77" refId="B12" refString="Ciplak, B, Heller, K-G, Willemse, F, 2009. Review of the genus Eupholidoptera (Orthoptera, Tettigoniidae): Different genitalia, uniform song. Zootaxa 2156 (1): 1 - 77, DOI: https://doi.org/10.11646/zootaxa.2156.1.1" title="Review of the genus Eupholidoptera (Orthoptera, Tettigoniidae): Different genitalia, uniform song." url="https://doi.org/10.11646/zootaxa.2156.1.1" volume="2156" year="2009">
<normalizedToken originalValue="Çiplak">Ciplak</normalizedToken>
et al. 2009
</bibRefCitation>
) show a syllable duration of ~ 123 ms and a syllable repetition rate of 1/s at maximum. The song may most likely be confused with the other species of
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupholidoptera" order="Orthoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="67">Eupholidoptera</emphasis>
</taxonomicName>
in Crete, except
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. smyrnensis" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="smyrnensis">
<emphasis italics="true" pageId="0" pageNumber="67">E. smyrnensis</emphasis>
</taxonomicName>
and
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. forcipata" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="forcipata">
<emphasis italics="true" pageId="0" pageNumber="67">E. forcipata</emphasis>
</taxonomicName>
. For details of sound recordings of
<taxonomicName authorityName="Willemse &amp; Heller" authorityYear="2001" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupholidoptera mariannae" order="Orthoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="mariannae">
<emphasis italics="true" pageId="0" pageNumber="67">Eupholidoptera mariannae</emphasis>
</taxonomicName>
see Suppl. material 3.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67" type="variation">
<paragraph pageId="0" pageNumber="67">Variation.</paragraph>
<paragraph pageId="0" pageNumber="67">Apical arms of the titillator in most specimens are fused, in some they become apically somewhat separated. The central pits in the anal tergite may be more or less well developed and more or less densely haired.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67" type="description">
<paragraph pageId="0" pageNumber="67">Description of female.</paragraph>
<paragraph pageId="0" pageNumber="67">
Examined specimens. 15 ♀: LASITHI: Agios Ioannis - RMNH.5014906 (RMNH); Anatoli - s.n. [paratype of
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. astyla" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="astyla">
<emphasis italics="true" pageId="0" pageNumber="67">E. astyla</emphasis>
</taxonomicName>
<bibRefCitation author="Ramme, W" journalOrPublisher="Eos" pageId="0" pageNumber="67" pagination="111 - 200" refId="B51" refString="Ramme, W, 1927. Die Dermapteren und Orthopteren Siziliens und Kretas. Eos 3: 111 - 200" title="Die Dermapteren und Orthopteren Siziliens und Kretas." volume="3" year="1927">Ramme 1927</bibRefCitation>
] (MfNB); Kalavros -2017.029.02 (CT) RMNH.5014912, RMNH. 5086974 (RMNH); Katharo plain, 1 km NE FC17787 - RMNH.INS1124470, RMNH.INS1124471 (RMNH); Kato Chorion s.n. [paratype of
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. astyla" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="astyla">
<emphasis italics="true" pageId="0" pageNumber="67">E. astyla</emphasis>
</taxonomicName>
<bibRefCitation author="Ramme, W" journalOrPublisher="Eos" pageId="0" pageNumber="67" pagination="111 - 200" refId="B51" refString="Ramme, W, 1927. Die Dermapteren und Orthopteren Siziliens und Kretas. Eos 3: 111 - 200" title="Die Dermapteren und Orthopteren Siziliens und Kretas." volume="3" year="1927">Ramme 1927</bibRefCitation>
] (MfNB); Kavousi - 1999.029.02 (CT), RMNH.INS1141830 (RMNH); Koutsouras - 2002.007.05 (CT); Prina, 0.5 km N FC17798 - 2019.061.02 (CT), RMNH.INS1141839 (RMNH); Mt. Thrypti - 2019.032.01 (CT), FC25104 RMNH.INS1124469 (RMNH). (For details see Suppl. material 2).
</paragraph>
<paragraph pageId="0" pageNumber="67">
General appearance and colouration as male (Figs
<figureCitation captionStart="Figures 224–235" captionStartId="F24" captionText="Figures 224 - 235. Habitus Eupholidoptera spp. in dorsal and lateral view 224, 225 Eupholidoptera astyla ♀ Mt. Idhi CT 1987.046.03 226, 227 Eupholidoptera cretica ♀ Mt. Lefka Omalos FC 17807 RMNH. INS 1141837 228, 229 Eupholidoptera francisae sp. nov. ♂ holotype Andikithira CT 2002.004.04 230, 231 Eupholidoptera francisae sp. nov. ♀ allotype Andikithira CT 2002.004.11 232, 233 Eupholidoptera gemellata ♀ Mt. Idhi FC 1651 RMNH. INS 1141844 234, 235 Eupholidoptera mariannae ♀ Ag. Ioannis RMNH 5014906 1844. Scale bars: 10 mm." figureDoi="10.3897/zookeys.1151.97514.figures224-235" httpUri="https://binary.pensoft.net/fig/817954" pageId="0" pageNumber="67">234</figureCitation>
,
<figureCitation captionStart="Figures 224–235" captionStartId="F24" captionText="Figures 224 - 235. Habitus Eupholidoptera spp. in dorsal and lateral view 224, 225 Eupholidoptera astyla ♀ Mt. Idhi CT 1987.046.03 226, 227 Eupholidoptera cretica ♀ Mt. Lefka Omalos FC 17807 RMNH. INS 1141837 228, 229 Eupholidoptera francisae sp. nov. ♂ holotype Andikithira CT 2002.004.04 230, 231 Eupholidoptera francisae sp. nov. ♀ allotype Andikithira CT 2002.004.11 232, 233 Eupholidoptera gemellata ♀ Mt. Idhi FC 1651 RMNH. INS 1141844 234, 235 Eupholidoptera mariannae ♀ Ag. Ioannis RMNH 5014906 1844. Scale bars: 10 mm." figureDoi="10.3897/zookeys.1151.97514.figures224-235" httpUri="https://binary.pensoft.net/fig/817954" pageId="0" pageNumber="67">235</figureCitation>
,
<figureCitation captionStart="Figures 247–253" captionStartId="F27" captionText="Figures 247 - 253. Field images Eupholidoptera spp. Crete 247 Eupholidoptera giuliae ♂ Prasies RMNH 5087051 248 Eupholidoptera giuliae ♀ Rethymnon RMNH 5014921 249 Eupholidoptera latens ♂ Kolympari RMNH 553681 250 Eupholidoptera mariannae ♂ Kalavros RMNH 5014907 251 Eupholidoptera mariannae ♀ Ag. Ioannis RMNH 5014906 252 Eupholidoptera pallipes ♀ below Pakhnes 253 Eupholidoptera smyrnensis ♂ Makrigiannis RMNH 5087053." figureDoi="10.3897/zookeys.1151.97514.figures247-253" httpUri="https://binary.pensoft.net/fig/817957" pageId="0" pageNumber="67">251</figureCitation>
). First abdominal segment dorsally darkened, in one female black; second segment may also be somewhat darkened. Fore wings covered by pronotum, in profile barely protruding, pale coloured.
</paragraph>
<paragraph pageId="0" pageNumber="67">Cercus short, conical hardly tapering but for apical third which is distinctly narrower, tip pointed, slightly upturned in profile, straight in dorsal view, covered with pale short and long hairs.</paragraph>
<paragraph pageId="0" pageNumber="67">
Subgenital plate (Figs
<figureCitation captionStart="Figures 39–52" captionStartId="F7" captionText="Figures 39 - 52. Female subgenital plate in ventral view 39 Eupholidoptera smyrnensis Makrigiannis RMNH. 5014918 40 Eupholidoptera gemellata Mt. Idhi, FC 1602 CT 2000.095.02 41 Eupholidoptera pallipes allotype Mt. Lefka RMNH. INS 1105312 42 Eupholidoptera cretica Mt. Lefka, Omalos RMNH. INS 1141837 43 Eupholidoptera latens Mt. Lefka, Kalergi CT 1987.047.04 44 Eupholidoptera giuliae Argoules CT 1995.011.03 45 Eupholidoptera francisae sp. nov. allotype Andikithira CT 2002.004.11 46 Eupholidoptera annamariae Kato Zakros CT 1995.020.03 47 Eupholidoptera astyla Krotos RMNH. INS 1141820 48 Eupholidoptera feri allotype Mt. Dikti, Katharo plain RMNH. INS 1105298 49 Eupholidoptera mariannae Kalavros RMNH 5014912 50 Eupholidoptera forcipata Mt. Idhi, FC 1602 RMNH. INS 1141845 51 Eupholidoptera marietheresae sp. nov. allotype Mt. Dikti RMNH. INS 1141849 52 Eupholidoptera jacquelinae Gavdos allotype CT 2001.004.13. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures39-52" httpUri="https://binary.pensoft.net/fig/817937" pageId="0" pageNumber="67">49</figureCitation>
,
<figureCitation captionStart="Figures 53–66" captionStartId="F8" captionText="Figures 53 - 66. Female subgenital plate in lateral view 53 Eupholidoptera smyrnensis Makrigiannis RMNH. 5014918 54 Eupholidoptera gemellata Mt. Idhi, FC 1602 CT 2000.095.02 55 Eupholidoptera pallipes allotype Mt. Lefka RMNH. INS 1105312 56 Eupholidoptera cretica Mt. Lefka, Omalos RMNH. INS 1141837 57 Eupholidoptera latens Mt. Lefka, Kalergi CT 1987.047.04 58 Eupholidoptera giuliae Argoules CT 1995.011.03 59 Eupholidoptera francisae sp. nov. allotype Andikithira CT 2002.004.11 60 Eupholidoptera annamariae Kato Zakros CT 1995.020.03 61 Eupholidoptera astyla Krotos RMNH. INS 1141820 62 Eupholidoptera feri allotype Mt. Dikti, Katharo plain RMNH. INS 1105298 63 Eupholidoptera mariannae Kalavros RMNH 5014912 64 Eupholidoptera forcipata Mt. Idhi, FC 1602 RMNH. INS 1141845 65 Eupholidoptera marietheresae sp. nov. allotype Mt. Dikti RMNH. INS 1141849 66 Eupholidoptera jacquelinae allotype Gavdos CT 2001.004.13. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures53-66" httpUri="https://binary.pensoft.net/fig/817938" pageId="0" pageNumber="67">63</figureCitation>
) longer than wide, mitre-shaped, in profile triangular; hind margin toward middle forming two distinct pointed apical lobes, separated by a deep and wide excision along one third of length; dorsal margin usually at least partly visible in the apical half as a protruding edge or bulge, in profile the apical half straight, the basal half straight except for the proximal part which is concave; ventral side proximally with two dark coloured concavities separated by a more or less distinct keel, the apical half flattened to shallowly depressed with a median keel, surface smooth without wrinkles, with dispersed hairs.
</paragraph>
<paragraph pageId="0" pageNumber="67">
Ovipositor almost straight, apically slightly upcurved, 1.4-2.2
<normalizedToken originalValue="×">x</normalizedToken>
longer than pronotum.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67" type="differential diagnosis">
<paragraph pageId="0" pageNumber="67">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="67">
Males differ from congenerics in the stout, upturned, proximally flattened cercus, pointing outward (Figs
<figureCitation captionStart="Figures 111–125" captionStartId="F13" captionText="Figures 111 - 125. Male cercus in dorsal view 111 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 112 Eupholidoptera gemellata Mt. Idhi FC 1651 RMNH. INS 1141843 113 Eupholidoptera pallipes Mt. Lefka paratype RMNH. INS 1105314 114 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 115 Eupholidoptera latens Xiloskalo IBER DC-Ort 000558 116 Eupholidoptera giuliae Chora Sfakion CT 2000.014.02 117 Eupholidoptera francisae sp. nov. holotype Andikithira CT 2002.004.04 118 Eupholidoptera annamariae Kato Zakros CT 2000.030.03 119 Eupholidoptera annamariae Xerokampos, male with anomalous cercus, IBER DC-Ort 000565 120 Eupholidoptera astyla Ierapetra MfN s. n. 121 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 122 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 123 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 124 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 125 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures111-125" httpUri="https://binary.pensoft.net/fig/817943" pageId="0" pageNumber="67">122</figureCitation>
,
<figureCitation captionStart="Figures 126–139" captionStartId="F14" captionText="Figures 126 - 139. Male cercus in lateral view 126 Eupholidoptera smyrnensis Makrigiannis RMNH. 5087053 127 Eupholidoptera gemellata Mt. Idhi FC 1651 RMNH. INS 1141843 128 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 129 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 130 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 131 Eupholidoptera giuliae Chora Sfakion CT 2000.024.04 132 Eupholidoptera francisae paratype sp. nov. Andikithira CT 2002.004.08 133 Eupholidoptera annamariae Kato Zakros CT 1995.020.02 134 Eupholidoptera astyla paratype Ierapetra MfN s. n. 135 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 136 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 137 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 138 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 139 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures126-139" httpUri="https://binary.pensoft.net/fig/817944" pageId="0" pageNumber="67">136</figureCitation>
), sub-basally armed with short strong inner tooth, in the anal tergite (Figs
<figureCitation captionStart="Figures 69–82" captionStartId="F10" captionText="Figures 69 - 82. Male anal tergite in dorsal view 69 Eupholidoptera smyrnensis Makrigiannis RMNH. INS 152909 70 Eupholidoptera gemellata holotype Mt. Idhi RMNH. INS 1105300 71 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105313 72 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 73 Eupholidoptera latens Mt. Lefka, Kalergi CT 2003.009.01 74 Eupholidoptera giuliae Chora Sfakion CT 2000.024.04 75 Eupholidoptera francisae sp. nov. holotype Andikithira CT 2002.004.04 76 Eupholidoptera annamariae Kato Zakros CT 2000.030.04 77 Eupholidoptera astyla Krotos RMNH. INS 1141819 78 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 79 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 80 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 81 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 82 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures69-82" httpUri="https://binary.pensoft.net/fig/817940" pageId="0" pageNumber="67">79</figureCitation>
,
<figureCitation captionStart="Figures 83–96" captionStartId="F11" captionText="Figures 83 - 96. Male anal tergite in caudal view 83 Eupholidoptera smyrnensis Makrigiannis RMNH. INS 152909 84 Eupholidoptera gemellata Mt. Idhi CT 2000.095.01 85 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 86 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 87 Eupholidoptera latens Mt. Lefka, Kalergi CT 2003.009.01 88 Eupholidoptera giuliae Chora Sfakion CT 2000.014.01 89 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.02 90 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 91 Eupholidoptera astyla Krotos RMNH. INS 1141819 92 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 93 Eupholidoptera mariannae Prina FC 17798 RMNH. INS 1141840 94 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 95 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 96 Eupholidoptera jacquelinae paratype Gavdos NHMC 2001.004.12. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures83-96" httpUri="https://binary.pensoft.net/fig/817941" pageId="0" pageNumber="67">93</figureCitation>
,
<figureCitation captionStart="Figures 97–110" captionStartId="F12" captionText="Figures 97 - 110. Male anal tergite in lateral view 97 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 98 Eupholidoptera gemellata Mt. Idhi CT 2000.095.01 99 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 100 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 101 Eupholidoptera latens Mt. Lefka, Kalergi CT 1987.047.01 102 Eupholidoptera giuliae Argoules CT 1995.011.11 103 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.02 104 Eupholidoptera annamariae Kato Zakros CT 2000.030.03 105 Eupholidoptera astyla Ano Viannos CT 1995.024.01 106 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 107 Eupholidoptera mariannae Kalavros CT 2017.029.01 108 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 109 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 110 Eupholidoptera jacquelinae Gavdopoula CT 1996.019.01. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures97-110" httpUri="https://binary.pensoft.net/fig/817942" pageId="0" pageNumber="67">107</figureCitation>
) medially not extended, bent downward with very narrow V-shaped excision, tips pointing forward, in the wide, upturned, spineless subgenital plate (Figs
<figureCitation captionStart="Figures 140–153" captionStartId="F15" captionText="Figures 140 - 153. Male subgenital plate in ventral view 140 Eupholidoptera smyrnensis Lagkada RMNH 5106270 141 Eupholidoptera gemellata Mt. Idhi FC 1651 RMNH. INS 1141843 142 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105313 143 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 144 Eupholidoptera latens Mt. Lefka Kalergi CT 1987.047.02 145 Eupholidoptera giuliae Skaloti IBER DC-Ort 000564 146 Eupholidoptera francisae sp. nov. holotype Andikithira CT 2002.004.04 147 Eupholidoptera annamariae Kato Zakros CT 2000.030.04 148 Eupholidoptera astyla Krotos RMNH. INS 1141819 149 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 150 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 151 Eupholidoptera forcipata Mt. Idhi CT 1987.044.03 152 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 153 Eupholidoptera jacquelinae holotype Gavdos 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures140-153" httpUri="https://binary.pensoft.net/fig/817945" pageId="0" pageNumber="67">150</figureCitation>
,
<figureCitation captionStart="Figures 154–167" captionStartId="F16" captionText="Figures 154 - 167. Male subgenital plate in lateral view 154 Eupholidoptera smyrnensis Lagkada RMNH. INS 152909 155 Eupholidoptera gemellata holotype Mt. Idhi RMNH. INS 1105300 156 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105313 157 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 (RMNH. INS 1141838 158 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 159 Eupholidoptera giuliae Skaloti IBER DC-Ort 000564 160 Eupholidoptera francisae sp. nov. holotype Andikithira CT 2002.004.04 161 Eupholidoptera annamariae Kato Zakros CT 2000.030.04 162 Eupholidoptera astyla Krotos RMNH 5086992 163 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 164 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 165 Eupholidoptera forcipata Mt. Idhi CT 1987.044.03 166 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 167 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures154-167" httpUri="https://binary.pensoft.net/fig/817946" pageId="0" pageNumber="67">164</figureCitation>
) with minute, pre-apically inserted styli (Fig.
<figureCitation captionStart="Figures 168–181" captionStartId="F17" captionText="Figures 168 - 181. Male stylus in lateral view 168 Eupholidoptera smyrnensis Makrigiannis RMNH. INS 152909 169 Eupholidoptera gemellata holotype Mt. Idhi RMNH. INS 1105300 170 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105313 171 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 172 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 173 Eupholidoptera giuliae Skaloti IBER DC-Ort 000564 174 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 175 Eupholidoptera annamariae Kato Zakros CT 2000.030.04 176 Eupholidoptera astyla Krotos RMNH. INS 1141819 177 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 178 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 179 Eupholidoptera forcipata Mt. Idhi CT 1987.044.03 180 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 181 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures168-181" httpUri="https://binary.pensoft.net/fig/817947" pageId="0" pageNumber="67">178</figureCitation>
) pointing downward and in the barely asymmetrical, narrow, fused apical arms of the titillator (Figs
<figureCitation captionStart="Figures 182–197" captionStartId="F18" captionText="Figures 182 - 197. Male titillator in ventral view 182 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 183 Eupholidoptera gemellata Mt. Idhi Amariou FC 1651 RMNH. INS 1141843 184 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 185 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 186 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 187 Eupholidoptera latens Rhodopos CH 8236 188 Eupholidoptera giuliae Argoules CT 1995.011.11 189 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 190 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 191 Eupholidoptera astyla Krotos RMNH. INS 1141819 192 Eupholidoptera astyla Kofinas FC 460 RMNH. INS 1141836 193 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 194 Eupholidoptera mariannae paratype Malles CH 2906 A 195 Eupholidoptera forcipata Mt. Idhi CT 2019.047.01 196 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 197 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures182-197" httpUri="https://binary.pensoft.net/fig/817948" pageId="0" pageNumber="67">194</figureCitation>
,
<figureCitation captionStart="Figures 198–212" captionStartId="F19" captionText="Figures 198 - 212. Male titillator in lateral view 198 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 199 Eupholidoptera gemellata Mt. Idhi Amariou FC 1651 RMNH. INS 1141843 200 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 201 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 202 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 203 Eupholidoptera latens Rhodopos CH 8236 204 Eupholidoptera giuliae Argoules CT 1995.011.11 205 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 206 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 207 Eupholidoptera astyla Krotos RMNH. INS 1141819 208 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 209 Eupholidoptera mariannae Kalavros CT 2017.029.01 210 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 211 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 212 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures198-212" httpUri="https://binary.pensoft.net/fig/817949" pageId="0" pageNumber="67">209</figureCitation>
). Females differ from congenerics in the elongated subgenital plate (Figs
<figureCitation captionStart="Figures 39–52" captionStartId="F7" captionText="Figures 39 - 52. Female subgenital plate in ventral view 39 Eupholidoptera smyrnensis Makrigiannis RMNH. 5014918 40 Eupholidoptera gemellata Mt. Idhi, FC 1602 CT 2000.095.02 41 Eupholidoptera pallipes allotype Mt. Lefka RMNH. INS 1105312 42 Eupholidoptera cretica Mt. Lefka, Omalos RMNH. INS 1141837 43 Eupholidoptera latens Mt. Lefka, Kalergi CT 1987.047.04 44 Eupholidoptera giuliae Argoules CT 1995.011.03 45 Eupholidoptera francisae sp. nov. allotype Andikithira CT 2002.004.11 46 Eupholidoptera annamariae Kato Zakros CT 1995.020.03 47 Eupholidoptera astyla Krotos RMNH. INS 1141820 48 Eupholidoptera feri allotype Mt. Dikti, Katharo plain RMNH. INS 1105298 49 Eupholidoptera mariannae Kalavros RMNH 5014912 50 Eupholidoptera forcipata Mt. Idhi, FC 1602 RMNH. INS 1141845 51 Eupholidoptera marietheresae sp. nov. allotype Mt. Dikti RMNH. INS 1141849 52 Eupholidoptera jacquelinae Gavdos allotype CT 2001.004.13. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures39-52" httpUri="https://binary.pensoft.net/fig/817937" pageId="0" pageNumber="67">49</figureCitation>
,
<figureCitation captionStart="Figures 53–66" captionStartId="F8" captionText="Figures 53 - 66. Female subgenital plate in lateral view 53 Eupholidoptera smyrnensis Makrigiannis RMNH. 5014918 54 Eupholidoptera gemellata Mt. Idhi, FC 1602 CT 2000.095.02 55 Eupholidoptera pallipes allotype Mt. Lefka RMNH. INS 1105312 56 Eupholidoptera cretica Mt. Lefka, Omalos RMNH. INS 1141837 57 Eupholidoptera latens Mt. Lefka, Kalergi CT 1987.047.04 58 Eupholidoptera giuliae Argoules CT 1995.011.03 59 Eupholidoptera francisae sp. nov. allotype Andikithira CT 2002.004.11 60 Eupholidoptera annamariae Kato Zakros CT 1995.020.03 61 Eupholidoptera astyla Krotos RMNH. INS 1141820 62 Eupholidoptera feri allotype Mt. Dikti, Katharo plain RMNH. INS 1105298 63 Eupholidoptera mariannae Kalavros RMNH 5014912 64 Eupholidoptera forcipata Mt. Idhi, FC 1602 RMNH. INS 1141845 65 Eupholidoptera marietheresae sp. nov. allotype Mt. Dikti RMNH. INS 1141849 66 Eupholidoptera jacquelinae allotype Gavdos CT 2001.004.13. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures53-66" httpUri="https://binary.pensoft.net/fig/817938" pageId="0" pageNumber="67">63</figureCitation>
) proximally with two concavities, apical lobes pointed, separated by wide excision, as deep as one third of the length. In colouration, the amount of black shown,
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. mariannae" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="mariannae">
<emphasis italics="true" pageId="0" pageNumber="67">E. mariannae</emphasis>
</taxonomicName>
together with
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. annamariae" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="annamariae">
<emphasis italics="true" pageId="0" pageNumber="67">E. annamariae</emphasis>
</taxonomicName>
,
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. astyla" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="astyla">
<emphasis italics="true" pageId="0" pageNumber="67">E. astyla</emphasis>
</taxonomicName>
,
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. feri" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="feri">
<emphasis italics="true" pageId="0" pageNumber="67">E. feri</emphasis>
</taxonomicName>
, and
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. francisae" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="francisae">
<emphasis italics="true" pageId="0" pageNumber="67">E. francisae</emphasis>
</taxonomicName>
sp. nov. belongs to the darkest coloured species. For more details differentiating
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. mariannae" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="mariannae">
<emphasis italics="true" pageId="0" pageNumber="67">E. mariannae</emphasis>
</taxonomicName>
from other Cretan
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupholidoptera" order="Orthoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="67">Eupholidoptera</emphasis>
</taxonomicName>
see Table
<tableCitation captionStart="Table 5" captionStartId="T5" captionText="Table 5. Diagnostic characters and character-states for Eupholidoptera species from Crete, Andikithira, Gavdos, and Gavdopoula." httpUri="http://table.plazi.org/id/DEC408AC569AD02CE9FF184126BFC8DE" pageId="0" pageNumber="67" tableUuid="DEC408AC569AD02CE9FF184126BFC8DE">5</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67" type="distribution">
<paragraph pageId="0" pageNumber="67">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="67">
The species was described from southwestern Lasithi. Recent findings indicate its distribution area also includes western central Lasithi from the eastern slopes of Mt. Dikti eastward up to Kalavros in the north and Koutsouras in the south (Fig.
<figureCitation captionStart="Figures 254, 255" captionStartId="F28" captionText="Figures 254, 255. Distribution maps Eupholidoptera Crete and adjacent islands 254 lowland species 255 highland species." figureDoi="10.3897/zookeys.1151.97514.figures254-255" httpUri="https://binary.pensoft.net/fig/817958" pageId="0" pageNumber="67">254</figureCitation>
). In the east there seems to be no overlap with
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. annamariae" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="annamariae">
<emphasis italics="true" pageId="0" pageNumber="67">E. annamariae</emphasis>
</taxonomicName>
although near Kalavros and Xerokampos both species were found within 5 km of each other.
<taxonomicName authorityName="Willemse &amp; Heller" authorityYear="2001" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupholidoptera mariannae" order="Orthoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="mariannae">
<emphasis italics="true" pageId="0" pageNumber="67">Eupholidoptera mariannae</emphasis>
</taxonomicName>
and
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. astyla" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="astyla">
<emphasis italics="true" pageId="0" pageNumber="67">E. astyla</emphasis>
</taxonomicName>
may overlap in the Ierapetra area but up to now co-occurrence could not be confirmed. In the northwest,
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. mariannae" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="mariannae">
<emphasis italics="true" pageId="0" pageNumber="67">E. mariannae</emphasis>
</taxonomicName>
was also found in the Katharo plain where
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. feri" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="feri">
<emphasis italics="true" pageId="0" pageNumber="67">E. feri</emphasis>
</taxonomicName>
was discovered. For a complete list of localities, specimens and repositories see Suppl. material 1.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67" type="habitat">
<paragraph pageId="0" pageNumber="67">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="67">
The altitudinal range of
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. mariannae" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="mariannae">
<emphasis italics="true" pageId="0" pageNumber="67">E. mariannae</emphasis>
</taxonomicName>
is considerable. It ranges from sea level where it was found near Koutsouras, to 1475 m, the highest altitude at which it was found on Mt. Thrypti. The type series was found inside open pine forest and groups of planted olive trees. Near Kalavros the species was found on open hill slopes with quite a dense vegetation of small shrubs interspersed with taller shrubs. Near Kavousi (Pacheia Amos) the species was trapped on a hill covered by phrygana. Around Malles and Anadoli individual males were heard singing on the branches of olive trees a few meters above the ground.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67" type="phenology">
<paragraph pageId="0" pageNumber="67">Phenology.</paragraph>
<paragraph pageId="0" pageNumber="67">At low altitudes adults can be found already in early May, at higher altitudes starting from 300-700 m up to 1475 m, adults appear in June or July and have been caught until the end of September and mid-October.</paragraph>
</subSubSection>
</treatment>
</document>
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