280 lines
32 KiB
XML
280 lines
32 KiB
XML
<document id="372191F4655CF08B84BC6762CE4F38C2" ID-CLB-Dataset="8165" ID-DOI="10.11646/zootaxa.4926.4.3" ID-GBIF-Dataset="07abd153-3d09-4820-82dd-6991f1d14ced" ID-ISSN="1175-5326" ID-Zenodo-Dep="4529625" ID-ZooBank="C92BA48D-4493-4388-A24D-545B7FD1C19E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612956496051" checkinUser="plazi" docAuthor="Tian, Mingyi, Huang, Xinglong & Li, Chenliang" docDate="2021" docId="255087E5FFEE492D1DE9FAB3FD0A22C0" docLanguage="en" docName="zootaxa.4926.4.3.pdf" docOrigin="Zootaxa 4926 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Guizhaphaenopsodes Tian & X. Huang 2021, n. gen." docType="treatment" docVersion="7" lastPageNumber="527" masterDocId="D969FF9DFFE8492B1D7EFA24FF822454" masterDocTitle="Contribution to the knowledge of subterranean ground beetles from eastern Wuling Mountains, China (Coleoptera: Carabidae: Trechinae)" masterLastPageNumber="534" masterPageNumber="521" pageNumber="527" updateTime="1698928636667" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="FA2C06B5DD2B96E760A68E238383665B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="D905BD63C59D0F843875F7E8B7C7D112">
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<mods:title id="258D45C9ACBF1013AEE47062123AF5BC">Contribution to the knowledge of subterranean ground beetles from eastern Wuling Mountains, China (Coleoptera: Carabidae: Trechinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="81B26BB3EEE7481E00AB5207369E6184" type="personal">
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<mods:role id="94072F6E6284C3B2ED817FA9CDF794F3">
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<mods:roleTerm id="FCA938AB8F3A0D1D7CD750B9793DECDA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5D3A0F88DBDD014B78ACF76F76CB2EBA">Tian, Mingyi</mods:namePart>
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<mods:affiliation id="CCC765BB0248C59179EC08EE1CF9029A">Department of Entomology, College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou, 510642, China</mods:affiliation>
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</mods:name>
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<mods:name id="9FA4382859109601A4EEF1CB619C3086" type="personal">
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<mods:role id="A2779760AC75338FA01A4B9823784C1F">
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<mods:roleTerm id="C71273D07134919CE1C63A52CFC394FC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C8D7E09B150402BB2DC132DA4B4536B6">Huang, Xinglong</mods:namePart>
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<mods:affiliation id="7DB6DE6511E88CC5FE3ACDD5BBCCBFF0">College of Biology and Environmental Sciences, Jishou University, Jishou 416000, China</mods:affiliation>
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</mods:name>
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<mods:name id="600DE59432A03C869EDD9499EBEC9BB6" type="personal">
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<mods:role id="5CF915BF27E64996F4610807884294E8">
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<mods:roleTerm id="19D5265C273A9B3E45E67B7262B9EFAF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B301F2E0CD6CC761E0D6BFFE3D8E711E">Li, Chenliang</mods:namePart>
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<mods:affiliation id="6D37C2DAE832F4DB1B24070F5D00C185">Hubei Board Nature, Central China Normal University, 430079, Wuhan, China</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="F1E83CCE1B6B22C50B29D4A743F3F20D">text</mods:typeOfResource>
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<mods:title id="F8735D0A5561AA2A2F82526896485137">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="CDB6A5D2916755801FA9588B2C85600F">
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<mods:date id="338250C267D27E1A048299643EE0FD40">2021</mods:date>
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<mods:detail id="97677DA82B7FB8D74CEC8EF15520E763" type="pubDate">
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<mods:number id="887FF40229BEC9621F46B81431C141EE">2021-02-10</mods:number>
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</mods:detail>
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<mods:detail id="F5E5FCFEB88777BF95BC686C84267CA1" type="volume">
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<mods:number id="1B50365189FC8433AADF8557E0180BC9">4926</mods:number>
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<mods:detail id="04F5931D3E602592063337752B28CC9A" type="issue">
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<mods:number id="EAE3666876CF2E5363A362654E0664F5">4</mods:number>
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<mods:start id="135D221DA8651C486FAE8AE9D7F92075">521</mods:start>
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<mods:end id="D23DBB223C444190D939CFD1F826E9B7">534</mods:end>
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<mods:classification id="2F78476A3FA6A44795D9D4723D921D72">journal article</mods:classification>
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<mods:identifier id="65754E808E32A6AC78FBACBB55954D1C" type="CLB-Dataset">8165</mods:identifier>
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<mods:identifier id="0E117795F652A3E558187E587F2E2B88" type="DOI">10.11646/zootaxa.4926.4.3</mods:identifier>
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<mods:identifier id="DBA9F6306C39A23AA0D73E558827B29A" type="GBIF-Dataset">07abd153-3d09-4820-82dd-6991f1d14ced</mods:identifier>
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<mods:identifier id="30F5FD84DD7F178FB661F2F65AF1D916" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="7142D62742ED36C4BE13CFAEBA7CE624" type="Zenodo-Dep">4529625</mods:identifier>
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</mods:mods>
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<treatment id="255087E5FFEE492D1DE9FAB3FD0A22C0" ID-DOI="http://doi.org/10.5281/zenodo.4546217" ID-GBIF-Taxon="177422223" ID-Zenodo-Dep="4546217" LSID="urn:lsid:plazi:treatment:255087E5FFEE492D1DE9FAB3FD0A22C0" httpUri="http://treatment.plazi.org/id/255087E5FFEE492D1DE9FAB3FD0A22C0" lastPageNumber="527" pageId="6" pageNumber="527">
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<subSubSection id="E5E36578FFEE492D1DE9FAB3FD5124E5" box="[151,723,151,178]" pageId="6" pageNumber="527" type="nomenclature">
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<paragraph id="AD4636F3FFEE492D1DE9FAB3FD5124E5" blockId="6.[151,723,151,178]" box="[151,723,151,178]" pageId="6" pageNumber="527">
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<heading id="F60E819FFFEE492D1DE9FAB3FD5124E5" bold="true" box="[151,723,151,178]" fontSize="11" level="1" pageId="6" pageNumber="527" reason="1">
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<emphasis id="9F8DEAE1FFEE492D1DE9FAB3FD5124E5" bold="true" box="[151,723,151,178]" pageId="6" pageNumber="527">
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<taxonomicName id="6AF94D70FFEE492D1DE9FAB3FDF124E5" authority="Tian & X.Huang" authorityName="Tian & X. Huang" authorityYear="2021" box="[151,627,151,178]" class="Insecta" family="Carabidae" genus="Guizhaphaenopsodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="9F8DEAE1FFEE492D1DE9FAB3FE1B24E6" bold="true" box="[151,409,151,178]" italics="true" pageId="6" pageNumber="527">Guizhaphaenopsodes</emphasis>
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Tian & X.Huang
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</taxonomicName>
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,
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<taxonomicNameLabel id="84BE579AFFEE492D1FFFFAB3FD5124E5" box="[641,723,151,177]" pageId="6" pageNumber="527" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="E5E36578FFEE492D1DE9FAF8FBC72728" pageId="6" pageNumber="527" type="materials_examined">
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<paragraph id="AD4636F3FFEE492D1DE9FAF8FB4224A0" blockId="6.[151,1216,220,244]" box="[151,1216,220,244]" pageId="6" pageNumber="527">
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<materialsCitation id="1D913CAEFFEE492D1DE9FAF8FB4224A0" ID-GBIF-Occurrence="3037449304" box="[151,1216,220,244]" location="Jishou" municipality="cave Tangle Dong" pageId="6" pageNumber="527" specimenCount="1" stateProvince="Hunan" typeStatus="holotype">
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<typeStatus id="72428851FFEE492D1DE9FAF8FF4B24A0" box="[151,201,220,244]" pageId="6" pageNumber="527">Type</typeStatus>
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species:
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<taxonomicName id="6AF94D70FFEE492D1C58FAF8FD6C24A0" authority="Tian & Huang" authorityName="Tian & Huang & Li" authorityYear="2021" box="[294,750,220,244]" class="Insecta" family="Carabidae" genus="Guizhaphaenopsodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="solidior" status="sp. nov.">
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<emphasis id="9F8DEAE1FFEE492D1C58FAF8FDD624A0" box="[294,596,220,244]" italics="true" pageId="6" pageNumber="527">Guizhaphaenopsodes solidior</emphasis>
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Tian & Huang
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</taxonomicName>
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,
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<taxonomicNameLabel id="84BE579AFFEE492D1F86FAF8FCAD24A7" box="[760,815,220,243]" pageId="6" pageNumber="527" rank="species">n. sp.</taxonomicNameLabel>
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(the
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<collectingMunicipality id="4D22AC89FFEE492D1E1CFAF8FB9F24A0" box="[866,1053,220,244]" pageId="6" pageNumber="527">cave Tangle Dong</collectingMunicipality>
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,
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<location id="A8266028FFEE492D1959FAF8FBE824A0" LSID="urn:lsid:plazi:treatment:255087E5FFEE492D1DE9FAB3FD0A22C0:A8266028FFEE492D1959FAF8FBE824A0" box="[1063,1130,220,244]" municipality="cave Tangle Dong" name="Jishou" pageId="6" pageNumber="527" stateProvince="Hunan">Jishou</location>
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,
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<collectingRegion id="6F3DF811FFEE492D190AFAF8FB3B24A0" box="[1140,1209,220,244]" country="China" name="Hunan" pageId="6" pageNumber="527">Hunan</collectingRegion>
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)
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</materialsCitation>
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</paragraph>
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<paragraph id="AD4636F3FFEE492D1DE9FB05FBC72728" blockId="6.[151,1437,289,1684]" pageId="6" pageNumber="527">
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<emphasis id="9F8DEAE1FFEE492D1DE9FB05FE37256F" bold="true" box="[151,437,289,315]" pageId="6" pageNumber="527">Generic characteristics.</emphasis>
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Medium-sized beetle for cavernicolous trechines, semi-aphaenopsian, but more or less pigmented; body stout, appendages moderate long, antennae robust; dorsal surface smooth and glabrous, moderately shining. Head subquadrate, longer than wide excluding mandibles; frontal furrows well-defined, only posterior pair of supra-orbital present; right mandible bidentate; labial suture completely disappeared; mentum bisetose, base largely and deeply concave; submentum with a row of 12 setae which are very long; the 2
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<superScript id="5A8C9BBBFFEE492D190FFB95FB0325EB" attach="left" box="[1137,1153,433,447]" fontSize="6" pageId="6" pageNumber="527">nd</superScript>
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labial palpomere without additional setae apart from the two ones on inner margin; antennae moderately long, but much stouter for cavernicolous trechines, scape as long as pedicel. Prothorax not tumid at sides and so invisible from above; pronotum quadrate, evidently longer than wide, only a pair of anterior latero-marginal setae present, hind angles broadly reflexed. Elytra stout though distinctly longer than fore body including mandibles, slightly expanded at sides, widest at about middle, strongly convex, shoulders distinct, humeral angles widely rounded, lateral margins ciliate throughout, apices almost rounded; elytral striae more or less obliterated but still notable; two dorsal pores present on the 3
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<superScript id="5A8C9BBBFFEE492D182EF8ADFADC26C3" attach="left" box="[1360,1374,649,663]" fontSize="6" pageId="6" pageNumber="527">rd</superScript>
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stria, the preapical dorsal pores absent; apical striole moderately defined, ended at the apex of the 5
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<superScript id="5A8C9BBBFFEE492D19D9F889FB3626EF" attach="left" box="[1191,1204,685,699]" fontSize="6" pageId="6" pageNumber="527">th</superScript>
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stria; humeral group (the 1
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<superScript id="5A8C9BBBFFEE492D1DA9F8F6FF6026B4" attach="left" box="[215,226,722,736]" fontSize="6" pageId="6" pageNumber="527">st</superScript>
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to 4
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<superScript id="5A8C9BBBFFEE492D1C6CF8F5FE9D268B" attach="left" box="[274,287,721,735]" fontSize="6" pageId="6" pageNumber="527">th</superScript>
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pores) of marginal umbilicate series not aggregated, 1
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<superScript id="5A8C9BBBFFEE492D1E05F8F6FC0426B4" attach="left" box="[891,902,722,736]" fontSize="6" pageId="6" pageNumber="527">st</superScript>
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inwardly and a bit backwardly shifted, 2
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<superScript id="5A8C9BBBFFEE492D1832F8F5FADE268B" attach="left" box="[1356,1372,721,735]" fontSize="6" pageId="6" pageNumber="527">nd</superScript>
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close to the marginal gutter, median group (the 5
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<superScript id="5A8C9BBBFFEE492D1F0AF8D1FD032757" attach="left" box="[628,641,757,771]" fontSize="6" pageId="6" pageNumber="527">th</superScript>
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and 6
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<superScript id="5A8C9BBBFFEE492D1FBAF8D1FD532757" attach="left" box="[708,721,757,771]" fontSize="6" pageId="6" pageNumber="527">th</superScript>
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pores) close from each other, both far from 4
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<superScript id="5A8C9BBBFFEE492D19B4F8D1FB552757" attach="left" box="[1226,1239,757,771]" fontSize="6" pageId="6" pageNumber="527">th</superScript>
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pore. Only the 1
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<superScript id="5A8C9BBBFFEE492D18EFF8D2FA1E2750" attach="none" box="[1425,1436,758,772]" fontSize="6" pageId="6" pageNumber="527">st</superScript>
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protarsomere modified in male, widened and distinctly denticulate inwards at apex; the 1
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<superScript id="5A8C9BBBFFEE492D19F8F93EFB13277C" attach="left" box="[1158,1169,794,808]" fontSize="6" pageId="6" pageNumber="527">st</superScript>
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tarsomere shorter than, as long as, and longer than 2
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<superScript id="5A8C9BBBFFEE492D1CAAF919FE66271F" attach="left" box="[468,484,829,843]" fontSize="6" pageId="6" pageNumber="527">nd</superScript>
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to 4
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<superScript id="5A8C9BBBFFEE492D1F6BF919FDA0271F" attach="left" box="[533,546,829,843]" fontSize="6" pageId="6" pageNumber="527">th</superScript>
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ones combined in fore, middle and hind legs respectively. Ventrite VII with one pair of apical setae in male. Male genitalia very short and small, weakly sclerotized.
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</paragraph>
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</subSubSection>
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<subSubSection id="E5E36578FFEE492D1DB9F9A2FB612218" pageId="6" pageNumber="527" type="discussion">
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<paragraph id="AD4636F3FFEE492D1DB9F9A2FBCF215C" blockId="6.[151,1437,289,1684]" pageId="6" pageNumber="527">
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<emphasis id="9F8DEAE1FFEE492D1DB9F9A2FEB927F4" bold="true" box="[199,315,902,928]" pageId="6" pageNumber="527">Remarks.</emphasis>
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This new genus is probably close to the genus
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<taxonomicName id="6AF94D70FFEE492D1E1FF9A2FA9D27F4" authority="Vigna Taglianti, 1997" authorityName="Vigna Taglianti" authorityYear="1997" box="[865,1311,902,928]" class="Insecta" family="Carabidae" genus="Guizhaphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">
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<emphasis id="9F8DEAE1FFEE492D1E1FF9A2FB9F27F4" box="[865,1053,902,928]" italics="true" pageId="6" pageNumber="527">Guizhaphaenops</emphasis>
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<bibRefCitation id="C9684B02FFEE492D1958F9A2FA9D27F4" author="Vigna Taglianti, A." box="[1062,1311,902,928]" pageId="6" pageNumber="527" pagination="33 - 41" refId="ref6562" refString="Vigna Taglianti, A. (1997) A new genus and species of troglobitic Trechinae (Coleoptera, Carabidae) from southern China. International Journal of Speleology, 25, 33 - 41. https: // doi. org / 10.5038 / 1827 - 806 X. 25.1.3" type="journal article" year="1997">Vigna Taglianti, 1997</bibRefCitation>
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</taxonomicName>
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, especially the subgenus
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<taxonomicName id="6AF94D70FFEE492D1C49F98EFD012790" authority="Deuve, 1999" authorityName="Deuve" authorityYear="1999" box="[311,643,938,964]" class="Insecta" family="Carabidae" genus="Semiaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">
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<emphasis id="9F8DEAE1FFEE492D1C49F98EFE652790" box="[311,487,938,964]" italics="true" pageId="6" pageNumber="527">Semiaphaenops</emphasis>
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<bibRefCitation id="C9684B02FFEE492D1C8FF98EFD012790" author="Deuve, T." box="[497,643,938,964]" pageId="6" pageNumber="527" pagination="151 - 161" refId="ref5694" refString="Deuve, T. (1999) Nouveaux Trechidae cavernicoles chinois, decouverts dans les confins karstiques du Sichuan, du Hubei et du Yunnan (Coleoptera, Adephaga). Revue francaise d'Entomologie, New Series, 21, 151 - 161." type="journal article" year="1999">Deuve, 1999</bibRefCitation>
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</taxonomicName>
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since they share the following main characteristics (
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<bibRefCitation id="C9684B02FFEE492D1995F98EFF5027BC" author="Vigna Taglianti, A." pageId="6" pageNumber="527" pagination="33 - 41" refId="ref6562" refString="Vigna Taglianti, A. (1997) A new genus and species of troglobitic Trechinae (Coleoptera, Carabidae) from southern China. International Journal of Speleology, 25, 33 - 41. https: // doi. org / 10.5038 / 1827 - 806 X. 25.1.3" type="journal article" year="1997">Vigna Taglianti 1997</bibRefCitation>
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,
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<bibRefCitation id="C9684B02FFEE492D1DA0F9EAFEED27BD" author="Ueno, S. I." box="[222,367,974,1001]" pageId="6" pageNumber="527" pagination="247 - 264" refId="ref6438" refString="Ueno, S. I. (2000 b) Notes on Guizhaphaenops (Coleoptera, Trechinae), with descriptions of two new species. Elytra, 28, 247 - 264." type="journal article" year="2000">Uéno 2000b</bibRefCitation>
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): similar body shape with moderately elongated appendages, bidentate right mandibular tooth, completely fused mentum and submentum, base of mentum widely and deeply concave, ventrite VII bisetose and male with only the 1
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<superScript id="5A8C9BBBFFEE492D1C04FE32FE072070" attach="left" box="[378,389,1046,1060]" fontSize="6" pageId="6" pageNumber="527">st</superScript>
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protarsomere modified. However, they differ in many character states, e.g.: (1) Only posterior pair of supraorbital pores present in
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<taxonomicName id="6AF94D70FFEE492D1F54FE1EFC982000" authorityName="Tian & X. Huang" authorityYear="2021" box="[554,794,1082,1108]" class="Insecta" family="Carabidae" genus="Guizhaphaenopsodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">
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<emphasis id="9F8DEAE1FFEE492D1F54FE1EFC982000" box="[554,794,1082,1108]" italics="true" pageId="6" pageNumber="527">Guizhaphaenopsodes</emphasis>
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</taxonomicName>
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, versus two pairs present in
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<taxonomicName id="6AF94D70FFEE492D1928FE1EFA842000" authorityName="Deuve" authorityYear="1999" box="[1110,1286,1082,1108]" class="Insecta" family="Carabidae" genus="Semiaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">
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<emphasis id="9F8DEAE1FFEE492D1928FE1EFA842000" box="[1110,1286,1082,1108]" italics="true" pageId="6" pageNumber="527">Semiaphaenops</emphasis>
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</taxonomicName>
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, and only the anterior ones present in several species of
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<emphasis id="9F8DEAE1FFEE492D1F01FE7AFC16202C" box="[639,916,1118,1144]" italics="true" pageId="6" pageNumber="527">
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<taxonomicName id="6AF94D70FFEE492D1F01FE7AFCB9202C" authorityName="Vigna Taglianti" authorityYear="1997" box="[639,827,1118,1144]" class="Insecta" family="Carabidae" genus="Guizhaphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">Guizhaphaenops</taxonomicName>
|
|
(s. str.)
|
|
</emphasis>
|
|
(
|
|
<bibRefCitation id="C9684B02FFEE492D1EDAFE7AFB1A202C" author="Vigna Taglianti, A." box="[932,1176,1118,1144]" pageId="6" pageNumber="527" pagination="33 - 41" refId="ref6562" refString="Vigna Taglianti, A. (1997) A new genus and species of troglobitic Trechinae (Coleoptera, Carabidae) from southern China. International Journal of Speleology, 25, 33 - 41. https: // doi. org / 10.5038 / 1827 - 806 X. 25.1.3" type="journal article" year="1997">Vigna Taglianti 1997</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="C9684B02FFEE492D19DBFE7AFAB6202C" author="Ueno, S. I." box="[1189,1332,1118,1144]" pageId="6" pageNumber="527" pagination="247 - 264" refId="ref6438" refString="Ueno, S. I. (2000 b) Notes on Guizhaphaenops (Coleoptera, Trechinae), with descriptions of two new species. Elytra, 28, 247 - 264." type="journal article" year="2000">Uéno 2000b</bibRefCitation>
|
|
, Uéno & Ran 2004); (2) Fore body more or less pigmented in
|
|
<taxonomicName id="6AF94D70FFEE492D1FA2FEA6FC4E20C8" authorityName="Tian & X. Huang" authorityYear="2021" box="[732,972,1154,1180]" class="Insecta" family="Carabidae" genus="Guizhaphaenopsodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="9F8DEAE1FFEE492D1FA2FEA6FC4E20C8" box="[732,972,1154,1180]" italics="true" pageId="6" pageNumber="527">Guizhaphaenopsodes</emphasis>
|
|
</taxonomicName>
|
|
, versus depigmented in
|
|
<taxonomicName id="6AF94D70FFEE492D19A6FEA6FA1620C8" authorityName="Vigna Taglianti" authorityYear="1997" box="[1240,1428,1154,1180]" class="Insecta" family="Carabidae" genus="Guizhaphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="9F8DEAE1FFEE492D19A6FEA6FA1620C8" box="[1240,1428,1154,1180]" italics="true" pageId="6" pageNumber="527">Guizhaphaenops</emphasis>
|
|
</taxonomicName>
|
|
; (3) Antennae sturdier and shorter in
|
|
<taxonomicName id="6AF94D70FFEE492D1F48FE82FCA42094" authorityName="Tian & X. Huang" authorityYear="2021" box="[566,806,1190,1216]" class="Insecta" family="Carabidae" genus="Guizhaphaenopsodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="9F8DEAE1FFEE492D1F48FE82FCA42094" box="[566,806,1190,1216]" italics="true" pageId="6" pageNumber="527">Guizhaphaenopsodes</emphasis>
|
|
</taxonomicName>
|
|
, whereas thinner and slenderer in
|
|
<taxonomicName id="6AF94D70FFEE492D19D1FE82FAE92094" authorityName="Vigna Taglianti" authorityYear="1997" box="[1199,1387,1190,1216]" class="Insecta" family="Carabidae" genus="Guizhaphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="9F8DEAE1FFEE492D19D1FE82FAE92094" box="[1199,1387,1190,1216]" italics="true" pageId="6" pageNumber="527">Guizhaphaenops</emphasis>
|
|
</taxonomicName>
|
|
; (4) Pronotum more elongated in
|
|
<taxonomicName id="6AF94D70FFEE492D1CA6FEEEFD4A20B0" authorityName="Tian & X. Huang" authorityYear="2021" box="[472,712,1226,1252]" class="Insecta" family="Carabidae" genus="Guizhaphaenopsodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="9F8DEAE1FFEE492D1CA6FEEEFD4A20B0" box="[472,712,1226,1252]" italics="true" pageId="6" pageNumber="527">Guizhaphaenopsodes</emphasis>
|
|
</taxonomicName>
|
|
than in the latter; and, in particular, (5) Male genitalia shorter and stouter in
|
|
<taxonomicName id="6AF94D70FFEE492D1C79FECAFE75215C" authorityName="Tian & X. Huang" authorityYear="2021" box="[263,503,1262,1288]" class="Insecta" family="Carabidae" genus="Guizhaphaenopsodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="9F8DEAE1FFEE492D1C79FECAFE75215C" box="[263,503,1262,1288]" italics="true" pageId="6" pageNumber="527">Guizhaphaenopsodes</emphasis>
|
|
</taxonomicName>
|
|
, versus much longer and thinner in
|
|
<taxonomicName id="6AF94D70FFEE492D1EF4FECAFBC4215C" authorityName="Vigna Taglianti" authorityYear="1997" box="[906,1094,1262,1288]" class="Insecta" family="Carabidae" genus="Guizhaphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="9F8DEAE1FFEE492D1EF4FECAFBC4215C" box="[906,1094,1262,1288]" italics="true" pageId="6" pageNumber="527">Guizhaphaenops</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="AD4636F3FFEE492D1DB9FF36FC3221CC" blockId="6.[151,1437,289,1684]" pageId="6" pageNumber="527">
|
|
Furthermore, there is a large gap in distribution between the two genera
|
|
<emphasis id="9F8DEAE1FFEE492D1E90FF36FB592178" box="[1006,1243,1298,1324]" italics="true" pageId="6" pageNumber="527">
|
|
<taxonomicName id="6AF94D70FFEE492D1E90FF36FB282178" authorityName="Vigna Taglianti" authorityYear="1997" box="[1006,1194,1298,1324]" class="Insecta" family="Carabidae" genus="Guizhaphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">Guizhaphaenops</taxonomicName>
|
|
and
|
|
</emphasis>
|
|
<taxonomicName id="6AF94D70FFEE492D199CFF36FE902104" authority=": The" authorityName="The" class="Insecta" family="Carabidae" genus="Guizhaphaenopsodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="9F8DEAE1FFEE492D199CFF36FF5C2104" italics="true" pageId="6" pageNumber="527">Guizhaphaenopsodes:</emphasis>
|
|
The
|
|
</taxonomicName>
|
|
cave Tangle Dong, locality of
|
|
<taxonomicName id="6AF94D70FFEE492D1F0CFF12FCE02104" box="[626,866,1334,1360]" class="Insecta" family="Carabidae" genus="Guizhaphaenopsodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="9F8DEAE1FFEE492D1F0CFF12FCE02104" box="[626,866,1334,1360]" italics="true" pageId="6" pageNumber="527">Guizhaphaenopsodes</emphasis>
|
|
</taxonomicName>
|
|
, is about
|
|
<quantity id="6A019B16FFEE492D1EACFF12FBAA2104" box="[978,1064,1334,1360]" metricMagnitude="5" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="527" unit="km" value="400.0">400 km</quantity>
|
|
in linear distance from Longyan Dong in Zunyi,
|
|
<collectingRegion id="6F3DF811FFEE492D1C39FF7EFE282120" box="[327,426,1370,1396]" country="China" name="Guizhou" pageId="6" pageNumber="527">Guizhou</collectingRegion>
|
|
, the
|
|
<typeStatus id="72428851FFEE492D1CA3FF7EFD8F2120" box="[477,525,1370,1396]" pageId="6" pageNumber="527">type</typeStatus>
|
|
locality of
|
|
<taxonomicName id="6AF94D70FFEE492D1FF4FF7EFA932120" authority="Deuve & Tian, 2018" authorityName="Deuve & Tian" authorityYear="2018" box="[650,1297,1370,1396]" class="Insecta" family="Carabidae" genus="Guizhaphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="subSpecies" species="lipsorum" subSpecies="zunyiensis">
|
|
<emphasis id="9F8DEAE1FFEE492D1FF4FF7EFBAB2120" box="[650,1065,1370,1396]" italics="true" pageId="6" pageNumber="527">Guizhaphaenops lipsorum zunyiensis</emphasis>
|
|
<bibRefCitation id="C9684B02FFEE492D1951FF7EFA932120" author="Deuve, T. & Tian, M. Y." box="[1071,1297,1370,1396]" pageId="6" pageNumber="527" pagination="333 - 339" refId="ref5921" refString="Deuve, T. & Tian, M. Y. (2018) Nouveaux Trechini cavernicoles du Guizhou nord-occidental des genres Zhijinaphaenops et Guizhaphaenops (Coleoptera, Trechidae). Bulletin de la Societe Entomologique de France, 123 (3), 333 - 339. https: // doi. org / 10.32475 / bsef _ 2039" type="journal article" year="2018">Deuve & Tian, 2018</bibRefCitation>
|
|
</taxonomicName>
|
|
which is the easternmost locality of
|
|
<taxonomicName id="6AF94D70FFEE492D1CE3FF5AFDDB21CC" authorityName="Vigna Taglianti" authorityYear="1997" box="[413,601,1406,1432]" class="Insecta" family="Carabidae" genus="Guizhaphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="9F8DEAE1FFEE492D1CE3FF5AFDDB21CC" box="[413,601,1406,1432]" italics="true" pageId="6" pageNumber="527">Guizhaphaenops</emphasis>
|
|
</taxonomicName>
|
|
species (
|
|
<bibRefCitation id="C9684B02FFEE492D1FBEFF5AFC2621CC" author="Deuve, T. & Tian, M. Y." box="[704,932,1406,1432]" pageId="6" pageNumber="527" pagination="333 - 339" refId="ref5921" refString="Deuve, T. & Tian, M. Y. (2018) Nouveaux Trechini cavernicoles du Guizhou nord-occidental des genres Zhijinaphaenops et Guizhaphaenops (Coleoptera, Trechidae). Bulletin de la Societe Entomologique de France, 123 (3), 333 - 339. https: // doi. org / 10.32475 / bsef _ 2039" type="journal article" year="2018">Deuve & Tian 2018</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="AD4636F3FFEE492D1DB9FF86FB612218" blockId="6.[151,1437,289,1684]" pageId="6" pageNumber="527">
|
|
<taxonomicName id="6AF94D70FFEE492D1DB9FF86FE3521E8" box="[199,439,1442,1468]" class="Insecta" family="Carabidae" genus="Guizhaphaenopsodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="9F8DEAE1FFEE492D1DB9FF86FE3521E8" box="[199,439,1442,1468]" italics="true" pageId="6" pageNumber="527">Guizhaphaenopsodes</emphasis>
|
|
</taxonomicName>
|
|
is also easily separated from
|
|
<taxonomicName id="6AF94D70FFEE492D1E7DFF86FBC321E8" authority="Ueno, 2000" authorityName="Ueno" authorityYear="2000" box="[771,1089,1442,1468]" class="Insecta" family="Carabidae" genus="Qianaphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="9F8DEAE1FFEE492D1E7DFF86FC3621E8" box="[771,948,1442,1468]" italics="true" pageId="6" pageNumber="527">Qianaphaenops</emphasis>
|
|
Uéno, 2000
|
|
</taxonomicName>
|
|
, which is distributed in eastern
|
|
<collectingRegion id="6F3DF811FFEE492D1DE9FFE2FF7521B4" box="[151,247,1478,1504]" country="China" name="Guizhou" pageId="6" pageNumber="527">Guizhou</collectingRegion>
|
|
and might be relative to this new genus, by the absence of labial suture (present in the latter genus), the bidentate right mandibular tooth (tridentate in
|
|
<taxonomicName id="6AF94D70FFEE492D1FD6FFCEFCDB2250" authorityName="Ueno" authorityYear="2000" box="[680,857,1514,1540]" class="Insecta" family="Carabidae" genus="Qianaphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="9F8DEAE1FFEE492D1FD6FFCEFCDB2250" box="[680,857,1514,1540]" italics="true" pageId="6" pageNumber="527">Qianaphaenops</emphasis>
|
|
</taxonomicName>
|
|
), the absence of anterior pair of supraorbital pores (present in
|
|
<taxonomicName id="6AF94D70FFEE492D1C68FC2AFE45227C" authorityName="Ueno" authorityYear="2000" box="[278,455,1550,1576]" class="Insecta" family="Carabidae" genus="Qianaphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="9F8DEAE1FFEE492D1C68FC2AFE45227C" box="[278,455,1550,1576]" italics="true" pageId="6" pageNumber="527">Qianaphaenops</emphasis>
|
|
</taxonomicName>
|
|
), only the 1
|
|
<superScript id="5A8C9BBBFFEE492D1F2FFC2AFDDE2248" attach="left" box="[593,604,1550,1564]" fontSize="6" pageId="6" pageNumber="527">st</superScript>
|
|
protarsomere modified in male (1
|
|
<superScript id="5A8C9BBBFFEE492D1EA0FC2AFC6B2248" attach="left" box="[990,1001,1550,1564]" fontSize="6" pageId="6" pageNumber="527">st</superScript>
|
|
and 2
|
|
<superScript id="5A8C9BBBFFEE492D1951FC29FBBD224F" attach="left" box="[1071,1087,1549,1563]" fontSize="6" pageId="6" pageNumber="527">nd</superScript>
|
|
protarsomeres modified in
|
|
<taxonomicName id="6AF94D70FFEE492D1809FC2AFEAE2218" authorityName="Ueno" authorityYear="2000" class="Insecta" family="Carabidae" genus="Qianaphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="9F8DEAE1FFEE492D1809FC2AFEAE2218" italics="true" pageId="6" pageNumber="527">Qianaphaenops</emphasis>
|
|
</taxonomicName>
|
|
) and the absence of the preapical dorsal pores on elytra (present in
|
|
<taxonomicName id="6AF94D70FFEE492D195FFC16FB502218" authorityName="Ueno" authorityYear="2000" box="[1057,1234,1586,1612]" class="Insecta" family="Carabidae" genus="Qianaphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="9F8DEAE1FFEE492D195FFC16FB502218" box="[1057,1234,1586,1612]" italics="true" pageId="6" pageNumber="527">Qianaphaenops</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="E5E36578FFEE492D1DB9FC72FD0A22C0" pageId="6" pageNumber="527" type="etymology">
|
|
<paragraph id="AD4636F3FFEE492D1DB9FC72FB7B2224" blockId="6.[151,1437,289,1684]" box="[199,1273,1622,1648]" pageId="6" pageNumber="527">
|
|
<emphasis id="9F8DEAE1FFEE492D1DB9FC72FECC2224" bold="true" box="[199,334,1622,1648]" pageId="6" pageNumber="527">Etymology.</emphasis>
|
|
To indicate the similarity of this new genus to
|
|
<taxonomicName id="6AF94D70FFEE492D1E23FC72FB9B2224" authorityName="Vigna Taglianti" authorityYear="1997" box="[861,1049,1622,1648]" class="Insecta" family="Carabidae" genus="Guizhaphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="9F8DEAE1FFEE492D1E23FC72FB9B2224" box="[861,1049,1622,1648]" italics="true" pageId="6" pageNumber="527">Guizhaphaenops</emphasis>
|
|
</taxonomicName>
|
|
. Gender masculine.
|
|
</paragraph>
|
|
<paragraph id="AD4636F3FFEE492D1DB9FC5DFD0A22C0" blockId="6.[151,1437,289,1684]" box="[199,648,1657,1684]" pageId="6" pageNumber="527">
|
|
<emphasis id="9F8DEAE1FFEE492D1DB9FC5DFEFA22C0" bold="true" box="[199,376,1657,1684]" pageId="6" pageNumber="527">Generic range.</emphasis>
|
|
<collectingCountry id="D5EE7663FFEE492D1C01FC5EFE4022C0" box="[383,450,1658,1684]" name="China" pageId="6" pageNumber="527">China</collectingCountry>
|
|
(
|
|
<collectingRegion id="6F3DF811FFEE492D1CAFFC5EFD9D22C0" box="[465,543,1658,1684]" country="China" name="Hunan" pageId="6" pageNumber="527">Hunan</collectingRegion>
|
|
) (
|
|
<figureCitation id="35C22A76FFEE492D1F48FC5EFDF522C0" box="[566,631,1658,1684]" captionStart="FIGURE 1" captionStartId="2.[151,250,1343,1367]" captionTargetBox="[210,1378,181,1320]" captionTargetId="figure-23@2.[210,1378,181,1320]" captionTargetPageId="2" captionText="FIGURE 1. A distribution map showing the locality caves of the three new species described in the present paper (Tangle Dong: Xiangxius jiangi Tian & X. Huang, n. gen., n. sp. and Guizhaphaenopsodes solidior Tian & X. Huang, n. gen., n. sp.; Qizimei Dong: Sinotroglodytes hefengensis Tian & Li, n. sp.)." figureDoi="http://doi.org/10.5281/zenodo.4529629" httpUri="https://zenodo.org/record/4529629/files/figure.png" pageId="6" pageNumber="527">Fig. 1</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |