222 lines
34 KiB
XML
222 lines
34 KiB
XML
<document id="69015F7A5E9A7F26FA03446E9649322E" ID-DOI="10.11646/zootaxa.5539.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="14240426" ID-ZooBank="7BDB9666-CA90-40BF-8F65-B9897CDA1F2B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1732831664062" checkinUser="plazi" docAuthor="Runyon, Justin B." docDate="2024" docId="FD1A87BFA516FFDE50D3FB1DFDA6FCEA" docLanguage="en" docName="zootaxa.5539.1.1.pdf" docOrigin="Zootaxa 5539 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5539.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Calyxochaetus lacrima Runyon, 2024, sp. nov." docType="treatment" docVersion="3" lastPageNumber="60" masterDocId="0123FFC7A52CFFE55044FFD3FFFBFFD3" masterDocTitle="Revision of the genus Calyxochaetus Bigot (Diptera: Dolichopodidae: Sympycninae)" masterLastPageNumber="74" masterPageNumber="1" pageNumber="59" updateTime="1732912132906" updateUser="carolina">
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<mods:mods id="30524454A30A9CF6068BEADC35E02D6B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="6AF00E62AA264C5BBDD0139F52789D21">
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<mods:title id="02A3DD6EADFFF07735FCDF527F65FA22">Revision of the genus Calyxochaetus Bigot (Diptera: Dolichopodidae: Sympycninae)</mods:title>
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<mods:name id="F678C1D72C6D662CF2507FD0EBA73A03" type="personal">
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<mods:roleTerm id="9C304B2A4BA505E9AF9CE00DFB1BBC36">Author</mods:roleTerm>
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<mods:namePart id="6E0281721E3719B158289800F42C973C">Runyon, Justin B.</mods:namePart>
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<mods:nameIdentifier id="00B4F07A7CCAC28C5E64AEDF608312C7" type="ORCID">0000-0002-0271-0511</mods:nameIdentifier>
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<mods:affiliation id="E215DCE7BD58FEAC555152282B50CA56">Rocky Mountain Research Station, USDA Forest Service, 1648 S. 7 th Avenue, Bozeman, Montana 59717, USA</mods:affiliation>
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<mods:nameIdentifier id="6D2794EDB5042B852A6E05D2F2D95974" type="email">justin.runyon@usda.gov</mods:nameIdentifier>
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<mods:typeOfResource id="CC4834611A4D70DDAE5ACA3324451A3E">text</mods:typeOfResource>
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<mods:title id="D5122EB3495F2A34233BC2B9C58FAC5A">Zootaxa</mods:title>
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<mods:part id="DB41C912F841EB6299CBEACFA026BFD7">
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<mods:date id="B3F6F24CE2D0AC020880330A576B2DF5">2024</mods:date>
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<mods:number id="23932E7F198D5ECDA4AC8F7F947EF519">2024-11-21</mods:number>
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<mods:number id="EE100724827E169F885978C723372D5F">5539</mods:number>
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<mods:url id="BD87983A857A5990C201EEB92ABE3064">http://dx.doi.org/10.11646/zootaxa.5539.1.1</mods:url>
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<mods:classification id="ABBBB241ACBC8BA04888E2DFC003C474">journal article</mods:classification>
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<mods:identifier id="BA4476CE211562DD2BEC9FAEE3306672" type="DOI">10.11646/zootaxa.5539.1.1</mods:identifier>
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<mods:identifier id="5CCF6AA97ED30ED9188A30B88350C922" type="ISSN">1175-5334</mods:identifier>
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<treatment id="FD1A87BFA516FFDE50D3FB1DFDA6FCEA" ID-DOI="http://doi.org/10.5281/zenodo.14240398" ID-Zenodo-Dep="14240398" LSID="urn:lsid:plazi:treatment:FD1A87BFA516FFDE50D3FB1DFDA6FCEA" httpUri="http://treatment.plazi.org/id/FD1A87BFA516FFDE50D3FB1DFDA6FCEA" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">
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<subSubSection id="3DA96522A516FFDF50D3FB1DFDE9FB3A" box="[151,530,1230,1257]" pageId="58" pageNumber="59" type="nomenclature">
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<paragraph id="750C36A9A516FFDF50D3FB1DFDE9FB3A" blockId="58.[151,545,1230,1293]" box="[151,530,1230,1257]" pageId="58" pageNumber="59">
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<heading id="2E4481C5A516FFDF50D3FB1DFDE9FB3A" bold="true" box="[151,530,1230,1257]" fontSize="11" level="2" pageId="58" pageNumber="59" reason="4">
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<emphasis id="47C7EABBA516FFDF50D3FB1DFDE9FB3A" bold="true" box="[151,530,1230,1257]" pageId="58" pageNumber="59">
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<taxonomicName id="B2B34D2AA516FFDF50D3FB1DFE56FB3A" authorityName="Runyon" authorityYear="2024" box="[151,429,1230,1257]" class="Insecta" family="Dolichopodidae" genus="Calyxochaetus" kingdom="Animalia" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="lacrima" status="sp. nov.">
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<emphasis id="47C7EABBA516FFDF50D3FB1DFE56FB3A" bold="true" box="[151,429,1230,1257]" italics="true" pageId="58" pageNumber="59">Calyxochaetus lacrima</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="5CF457C0A516FFDF51F0FB1CFDE9FB3A" box="[436,530,1231,1257]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="3DA96522A516FFDF50D3FB20FDDAFADE" box="[151,545,1266,1293]" pageId="58" pageNumber="59" type="description">
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<paragraph id="750C36A9A516FFDF50D3FB20FDDAFADE" blockId="58.[151,545,1230,1293]" box="[151,545,1266,1293]" pageId="58" pageNumber="59">
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(
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<figureCitation id="ED882A2CA516FFDF50DBFB20FEC1FADE" box="[159,314,1266,1293]" captionStart="FIGURES 104–119" captionStartId="50.[151,264,1472,1496]" captionTargetBox="[151,1437,181,1448]" captionTargetId="figure-27@50.[151,1437,181,1448]" captionTargetPageId="50" captionText="FIGURES 104–119. Male antennae (lateral view) and forelegs of the Calyxochaetus frontalis species group. 104. C. binodatus (Harmston & Knowlton). 105. C. nodatus (Loew). 106. C. desertus sp. nov. 107. C. pictipes (Harmston & Knowlton). 108. C. sobrinus (Wheeler). 109. C. ornatus Parent. 110. C. remifer sp. nov. 111. C. tohono sp. nov. 112. C. frontalis (Loew). 113. C. furcatus (Van Duzee). 114. C. lamellicornis Parent (photo by Emmanuel Delfosse, MNHN). 115. C. lacrima sp. nov. 116. C. binodatus, fore tarsus, anterior view. 117. C. sobrinus, fore tarsus, anterior view. 118. C. remifer sp. nov., fore tarsus, dorsal view. 119. C. lacrima sp. nov., fore tibia and tarsus, posterior view. Scale bars = 0.5 mm; arista-like styli approximately 1 mm long." figureDoi="http://doi.org/10.5281/zenodo.14240476" httpUri="https://zenodo.org/record/14240476/files/figure.png" pageId="58" pageNumber="59">Figs 115, 119</figureCitation>
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,
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<figureCitation id="ED882A2CA516FFDF5102FB20FE89FADE" box="[326,370,1267,1293]" captionStart="FIGURES 120–131" captionStartId="52.[151,264,1711,1735]" captionTargetBox="[196,1392,181,1688]" captionTargetId="figure-27@52.[196,1392,181,1688]" captionTargetPageId="52" captionText="FIGURES 120–131. Male mid tarsi of the Calyxochaetus frontalis species group (anterior on left side). 120. C. binodatus (Harmston & Knowlton), dorsal view. 121. C. nodatus (Loew), ventral view. 122. C. bovanodatus sp. nov., dorsal view. 123. C. desertus sp. nov., mid tarsomeres 2–5, dorsal view. 124. C. sobrinus (Wheeler), mid tarsomeres 2–5, dorsal view. 125. C. pictipes (Harmston & Knowlton), dorsal view. 126. C. ornatus Parent, mid tarsomeres 2–5, ventral view. 127. C. tohono sp. nov., mid tarsomeres 2–5, dorsal view. 128. C. remifer sp. nov., dorsal view. 129. C. frontalis (Loew), dorsal view. 130. C. furcatus (Van Duzee), dorsal view. 131. C. lacrima sp. nov., dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14240480" httpUri="https://zenodo.org/record/14240480/files/figure.png" pageId="58" pageNumber="59">131</figureCitation>
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,
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<figureCitation id="ED882A2CA516FFDF513AFB20FE51FADE" box="[382,426,1267,1293]" captionStart="FIGURES 132–138" captionStartId="54.[151,264,1423,1447]" captionTargetBox="[152,1436,381,1400]" captionTargetId="figure-114@54.[152,1436,381,1400]" captionTargetPageId="54" captionText="FIGURES 132–138. Male hind legs and wings of the Calyxochaetus frontalis species group. 132. C. binodatus (Harmston & Knowlton), hind tibia, anterior view. 133. C. nodatus (Loew), hind tibia, anterior view. 134. C. ornatus Parent, hind tibia, posterior view. 135. C. frontalis (Loew), hind tibia, posterior view. 136. C. lacrima sp. nov., apical third of hind femur and basal half of hind tibia, posterior view. 137. C. nodatus, wing. 138. C. ornatus, wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14240484" httpUri="https://zenodo.org/record/14240484/files/figure.png" pageId="58" pageNumber="59">136</figureCitation>
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,
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<figureCitation id="ED882A2CA516FFDF51F2FB20FE19FADE" box="[438,482,1267,1293]" captionStart="FIGURES 139–143" captionStartId="60.[151,264,1395,1419]" captionTargetBox="[162,1432,181,1371]" captionTargetId="figure-27@60.[151,1436,181,1371]" captionTargetPageId="60" captionText="FIGURES 139–143. Holotypes of new species in the Calyxochaetus frontalis group. 139. C. bovanodatus sp. nov. 140. C. lacrima sp. nov. 141. C. desertus sp. nov. 142. C. remifer sp. nov. 143. C. tohono sp. nov. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14240486" httpUri="https://zenodo.org/record/14240486/files/figure.png" pageId="58" pageNumber="59">140</figureCitation>
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,
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<figureCitation id="ED882A2CA516FFDF51AAFB20FDE2FADE" box="[494,537,1267,1293]" captionStart="FIGURES 147–148" captionStartId="66.[151,264,1875,1899]" captionTargetBox="[274,1315,181,1851]" captionTargetId="figure-27@66.[271,1315,181,1851]" captionTargetPageId="66" captionText="FIGURES 147–148. Known geographical distributions of species in the Calyxochaetus frontalis group. 147. C. furcatus (Van Duzee), C. lacrima sp. nov. and C. lamellicornis Parent. 148. C. ornatus Parent, C. remifer sp. nov. and C. tohono sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14240492" httpUri="https://zenodo.org/record/14240492/files/figure.png" pageId="58" pageNumber="59">147</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="3DA96522A516FFDF50D3FAE8FD4CFA4E" pageId="58" pageNumber="59" type="materials_examined">
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||
<paragraph id="750C36A9A516FFDF50D3FAE8FD4CFA4E" blockId="58.[151,1437,1338,2013]" pageId="58" pageNumber="59">
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||
<emphasis id="47C7EABBA516FFDF50D3FAE8FEB1FA87" bold="true" box="[151,330,1338,1365]" pageId="58" pageNumber="59">Type material.</emphasis>
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||
<materialsCitation id="C5DB3CF4A516FFDF5112FAE9FB1EFAAA" collectingDate="1985-05-10" collectionCode="CNC" collectorName="A. Contreras & J. B. Runyon" country="Mexico" location="Nuevo Leon" pageId="58" pageNumber="59" specimenCount="1" specimenCount-male="1" stateProvince="Nuevo Leon" typeStatus="holotype">
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||
<emphasis id="47C7EABBA516FFDF5112FAE9FE0EFA87" bold="true" box="[342,501,1338,1364]" pageId="58" pageNumber="59">
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||
<typeStatus id="AA08880BA516FFDF5112FAE9FE0EFA87" box="[342,501,1338,1364]" pageId="58" pageNumber="59" type="holotype">HOLOTYPE</typeStatus>
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||
</emphasis>
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||
,
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<specimenCount id="63B5FD20A516FFDF5243FAE8FDE5FA86" box="[519,542,1339,1365]" count="1" pageId="58" pageNumber="59" type="male">♂</specimenCount>
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||
, labeled: “
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||
<collectingCountry id="0DA47639A516FFDF52D9FAE8FCE9FA86" box="[669,786,1339,1365]" name="Mexico" pageId="58" pageNumber="59">MEXICO</collectingCountry>
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||
:
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<collectingRegion id="B777F84BA516FFDF5366FAE8FC4AFA86" box="[802,945,1339,1365]" country="Mexico" name="Nuevo Leon" pageId="58" pageNumber="59">Nuevo Leon</collectingRegion>
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/ Santiago, Potrero/ Redondo.
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<date id="010D1069A516FFDF555CFAE9FA96FA86" box="[1304,1389,1338,1365]" pageId="58" pageNumber="59" value="1985-05-10">
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||
<collectingDate id="1149E981A516FFDF555CFAE9FA96FA86" box="[1304,1389,1338,1365]" pageId="58" pageNumber="59" value="1985-05-10">10.V.85</collectingDate>
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</date>
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/
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<collectorName id="D846537FA516FFDF55C5FAE8FEFFFAAA" pageId="58" pageNumber="59">A. Contreras</collectorName>
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R.”; “HOLOTYPE/
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<taxonomicName id="B2B34D2AA516FFDF51AAFA8CFD68FAAA" box="[494,659,1375,1401]" class="Insecta" family="Dolichopodidae" genus="Calyxochaetus" kingdom="Animalia" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
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<emphasis id="47C7EABBA516FFDF51AAFA8CFD68FAAA" box="[494,659,1375,1401]" italics="true" pageId="58" pageNumber="59">Calyxochaetus</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="B2B34D2AA516FFDF52E5FA8CFD03FAAA" authorityName="Runyon" authorityYear="2024" box="[673,760,1375,1401]" class="Insecta" family="Dolichopodidae" genus="Calyxochaetus" kingdom="Animalia" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="lacrima">
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<emphasis id="47C7EABBA516FFDF52E5FA8CFD03FAAA" box="[673,760,1375,1401]" italics="true" pageId="58" pageNumber="59">lacrima</emphasis>
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</taxonomicName>
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/
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<collectorName id="D846537FA516FFDF5343FA8CFC69FAAA" box="[775,914,1375,1401]" pageId="58" pageNumber="59">J.B. Runyon</collectorName>
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[red label]” (
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<collectionCode id="13A2AE6CA516FFDF5463FA8CFB9FFAAA" box="[1063,1124,1375,1401]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" pageId="58" pageNumber="59">CNC</collectionCode>
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||
) (
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<figureCitation id="ED882A2CA516FFDF543EFA8CFB27FAAA" box="[1146,1244,1375,1401]" captionStart="FIGURES 139–143" captionStartId="60.[151,264,1395,1419]" captionTargetBox="[162,1432,181,1371]" captionTargetId="figure-27@60.[151,1436,181,1371]" captionTargetPageId="60" captionText="FIGURES 139–143. Holotypes of new species in the Calyxochaetus frontalis group. 139. C. bovanodatus sp. nov. 140. C. lacrima sp. nov. 141. C. desertus sp. nov. 142. C. remifer sp. nov. 143. C. tohono sp. nov. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14240486" httpUri="https://zenodo.org/record/14240486/files/figure.png" pageId="58" pageNumber="59">Fig. 140</figureCitation>
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||
)
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||
</materialsCitation>
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||
.
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||
<materialsCitation id="C5DB3CF4A516FFDF54ABFA8DFD48FA4E" collectionCode="CNC" collectorName="A. Contreras & J. B. Runyon" country="Mexico" location="Mexico" pageId="58" pageNumber="59" specimenCount="12" specimenCount-female="7" specimenCount-male="5" stateProvince="Mexico" typeStatus="paratype">
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||
<emphasis id="47C7EABBA516FFDF54ABFA8DFF03FA4F" bold="true" pageId="58" pageNumber="59">
|
||
<typeStatus id="AA08880BA516FFDF54ABFA8DFA6CFAAB" box="[1263,1431,1374,1400]" pageId="58" pageNumber="59" type="paratype">PARATYPES</typeStatus>
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||
:
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||
<collectingRegion id="B777F84BA516FFDF50D3FA51FF09FA4F" box="[151,242,1410,1436]" country="Mexico" name="Mexico" pageId="58" pageNumber="59">Mexico</collectingRegion>
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||
:
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</emphasis>
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Same data as holotype (
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||
<specimenCount id="63B5FD20A516FFDF524EFA51FDD4FA4E" box="[522,559,1410,1437]" count="5" pageId="58" pageNumber="59" type="male">5♂</specimenCount>
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||
,
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||
<specimenCount id="63B5FD20A516FFDF527FFA50FDA4FA4E" box="[571,607,1411,1437]" count="7" pageId="58" pageNumber="59" type="female">7♀</specimenCount>
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||
,
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||
<collectionCode id="13A2AE6CA516FFDF5228FA50FD51FA4E" box="[620,682,1411,1437]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" pageId="58" pageNumber="59">CNC</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="3DA96522A516FFDF5083FA75FA67FA36" pageId="58" pageNumber="59" type="diagnosis">
|
||
<paragraph id="750C36A9A516FFDF5083FA75FA67FA36" blockId="58.[151,1437,1338,2013]" pageId="58" pageNumber="59">
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||
<emphasis id="47C7EABBA516FFDF5083FA75FEB9FA13" bold="true" box="[199,322,1446,1472]" pageId="58" pageNumber="59">Diagnosis.</emphasis>
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The very large, nearly round apical lamella on the arista-like stylus of males is diagnostic (
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<figureCitation id="ED882A2CA516FFDF5574FA74FA6BFA12" box="[1328,1424,1446,1473]" captionStart="FIGURES 104–119" captionStartId="50.[151,264,1472,1496]" captionTargetBox="[151,1437,181,1448]" captionTargetId="figure-27@50.[151,1437,181,1448]" captionTargetPageId="50" captionText="FIGURES 104–119. Male antennae (lateral view) and forelegs of the Calyxochaetus frontalis species group. 104. C. binodatus (Harmston & Knowlton). 105. C. nodatus (Loew). 106. C. desertus sp. nov. 107. C. pictipes (Harmston & Knowlton). 108. C. sobrinus (Wheeler). 109. C. ornatus Parent. 110. C. remifer sp. nov. 111. C. tohono sp. nov. 112. C. frontalis (Loew). 113. C. furcatus (Van Duzee). 114. C. lamellicornis Parent (photo by Emmanuel Delfosse, MNHN). 115. C. lacrima sp. nov. 116. C. binodatus, fore tarsus, anterior view. 117. C. sobrinus, fore tarsus, anterior view. 118. C. remifer sp. nov., fore tarsus, dorsal view. 119. C. lacrima sp. nov., fore tibia and tarsus, posterior view. Scale bars = 0.5 mm; arista-like styli approximately 1 mm long." figureDoi="http://doi.org/10.5281/zenodo.14240476" httpUri="https://zenodo.org/record/14240476/files/figure.png" pageId="58" pageNumber="59">Fig. 115</figureCitation>
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). This is the only species in the
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<taxonomicName id="B2B34D2AA516FFDF51ADFA18FD93FA36" box="[489,616,1483,1509]" class="Insecta" family="Dolichopodidae" genus="Calyxochaetus" kingdom="Animalia" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="frontalis">
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<emphasis id="47C7EABBA516FFDF51ADFA18FD93FA36" box="[489,616,1483,1509]" italics="true" pageId="58" pageNumber="59">C. frontalis</emphasis>
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</taxonomicName>
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group whose males have long ventral setae on the hind femur (
|
||
<figureCitation id="ED882A2CA516FFDF556AFA18FA6BFA36" box="[1326,1424,1483,1509]" captionStart="FIGURES 132–138" captionStartId="54.[151,264,1423,1447]" captionTargetBox="[152,1436,381,1400]" captionTargetId="figure-114@54.[152,1436,381,1400]" captionTargetPageId="54" captionText="FIGURES 132–138. Male hind legs and wings of the Calyxochaetus frontalis species group. 132. C. binodatus (Harmston & Knowlton), hind tibia, anterior view. 133. C. nodatus (Loew), hind tibia, anterior view. 134. C. ornatus Parent, hind tibia, posterior view. 135. C. frontalis (Loew), hind tibia, posterior view. 136. C. lacrima sp. nov., apical third of hind femur and basal half of hind tibia, posterior view. 137. C. nodatus, wing. 138. C. ornatus, wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14240484" httpUri="https://zenodo.org/record/14240484/files/figure.png" pageId="58" pageNumber="59">Fig. 136</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3DA96522A516FFDE5083FA3DFAC6FD7A" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59" type="description">
|
||
<paragraph id="750C36A9A516FFDE5083FA3DFBDBFE02" blockId="58.[151,1437,1338,2013]" lastBlockId="59.[151,1437,150,825]" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">
|
||
<emphasis id="47C7EABBA516FFDF5083FA3DFE53F9DA" bold="true" box="[199,424,1518,1545]" pageId="58" pageNumber="59">Description. Male:</emphasis>
|
||
Wing length
|
||
<quantity id="B24B9B4CA516FFDF5203FA3CFD2FF9DA" box="[583,724,1519,1545]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.1" metricValueMin="2.8" pageId="58" pageNumber="59" unit="mm" value="2.95" valueMax="3.1" valueMin="2.8">2.8–3.1 mm</quantity>
|
||
.
|
||
<emphasis id="47C7EABBA516FFDF52A4FA3CFCD2F9DA" bold="true" box="[736,809,1519,1545]" pageId="58" pageNumber="59">Head:</emphasis>
|
||
Frons brown pruinose with large glabrous shiny blue-violet spot on either side of ocellar tubercle. Face silver, about 2 ommatidia wide on ventral half. Antenna (
|
||
<figureCitation id="ED882A2CA516FFDF552BF9C0FF3AF982" captionStart="FIGURES 104–119" captionStartId="50.[151,264,1472,1496]" captionTargetBox="[151,1437,181,1448]" captionTargetId="figure-27@50.[151,1437,181,1448]" captionTargetPageId="50" captionText="FIGURES 104–119. Male antennae (lateral view) and forelegs of the Calyxochaetus frontalis species group. 104. C. binodatus (Harmston & Knowlton). 105. C. nodatus (Loew). 106. C. desertus sp. nov. 107. C. pictipes (Harmston & Knowlton). 108. C. sobrinus (Wheeler). 109. C. ornatus Parent. 110. C. remifer sp. nov. 111. C. tohono sp. nov. 112. C. frontalis (Loew). 113. C. furcatus (Van Duzee). 114. C. lamellicornis Parent (photo by Emmanuel Delfosse, MNHN). 115. C. lacrima sp. nov. 116. C. binodatus, fore tarsus, anterior view. 117. C. sobrinus, fore tarsus, anterior view. 118. C. remifer sp. nov., fore tarsus, dorsal view. 119. C. lacrima sp. nov., fore tibia and tarsus, posterior view. Scale bars = 0.5 mm; arista-like styli approximately 1 mm long." figureDoi="http://doi.org/10.5281/zenodo.14240476" httpUri="https://zenodo.org/record/14240476/files/figure.png" pageId="58" pageNumber="59">Fig. 115</figureCitation>
|
||
) dark brown; scape relatively short and stout, with apex narrowly yellow medially; postpedicel elongate oval, about as long as wide; arista-like stylus bare and becoming very thin before broadening into large (subequal in size to postpedicel), teardrop-shaped lamella. Palpus small, elongate oval, brown at base and broadly yellow apically, with yellow setulae.
|
||
<emphasis id="47C7EABBA516FFDF513BF970FE1AF96E" bold="true" box="[383,481,1699,1725]" pageId="58" pageNumber="59">Thorax:</emphasis>
|
||
Pleura brown with light gray pruinosity, yellowish below halter and above mid coxa; metepimeron yellow. Scutellum with distinct blue-violet reflections medially.
|
||
<emphasis id="47C7EABBA516FFDF545CF914FBA2F932" bold="true" box="[1048,1113,1735,1761]" pageId="58" pageNumber="59">Legs:</emphasis>
|
||
<emphasis id="47C7EABBA516FFDF5427F914FB40F932" box="[1123,1211,1735,1761]" italics="true" pageId="58" pageNumber="59">Foreleg</emphasis>
|
||
(
|
||
<figureCitation id="ED882A2CA516FFDF5489F914FAC9F932" box="[1229,1330,1735,1761]" captionStart="FIGURES 104–119" captionStartId="50.[151,264,1472,1496]" captionTargetBox="[151,1437,181,1448]" captionTargetId="figure-27@50.[151,1437,181,1448]" captionTargetPageId="50" captionText="FIGURES 104–119. Male antennae (lateral view) and forelegs of the Calyxochaetus frontalis species group. 104. C. binodatus (Harmston & Knowlton). 105. C. nodatus (Loew). 106. C. desertus sp. nov. 107. C. pictipes (Harmston & Knowlton). 108. C. sobrinus (Wheeler). 109. C. ornatus Parent. 110. C. remifer sp. nov. 111. C. tohono sp. nov. 112. C. frontalis (Loew). 113. C. furcatus (Van Duzee). 114. C. lamellicornis Parent (photo by Emmanuel Delfosse, MNHN). 115. C. lacrima sp. nov. 116. C. binodatus, fore tarsus, anterior view. 117. C. sobrinus, fore tarsus, anterior view. 118. C. remifer sp. nov., fore tarsus, dorsal view. 119. C. lacrima sp. nov., fore tibia and tarsus, posterior view. Scale bars = 0.5 mm; arista-like styli approximately 1 mm long." figureDoi="http://doi.org/10.5281/zenodo.14240476" httpUri="https://zenodo.org/record/14240476/files/figure.png" pageId="58" pageNumber="59">Fig. 119</figureCitation>
|
||
): Yellow, except tarsus becoming brown from near apex of tarsomere 3. Tibia slender, with small posterodorsal seta near 1/3. Tarsomere 1 very short, slightly longer than wide. Tarsomere 2 about half tibia length. Tarsomere 3 about half tarsomere 2 length, slightly broadened apically and pointed dorsoapically with a few short distally curved black dorsal setulae across apex. Tarsomere 4 about 1.5X tarsomere 3 length, slightly narrowed at base, slightly thicker near 2/3 (appearing slightly bowed, in anterior and posterior views), with row of small black dorsal setulae across apex.
|
||
<emphasis id="47C7EABBA516FFDF5163F84CFE8DF86A" box="[295,374,1951,1977]" italics="true" pageId="58" pageNumber="59">Midleg</emphasis>
|
||
: Yellow, except tarsus brown from near apex of tarsomere 1. Tarsus (
|
||
<figureCitation id="ED882A2CA516FFDF5439F84CFB26F86A" box="[1149,1245,1951,1977]" captionStart="FIGURES 120–131" captionStartId="52.[151,264,1711,1735]" captionTargetBox="[196,1392,181,1688]" captionTargetId="figure-27@52.[196,1392,181,1688]" captionTargetPageId="52" captionText="FIGURES 120–131. Male mid tarsi of the Calyxochaetus frontalis species group (anterior on left side). 120. C. binodatus (Harmston & Knowlton), dorsal view. 121. C. nodatus (Loew), ventral view. 122. C. bovanodatus sp. nov., dorsal view. 123. C. desertus sp. nov., mid tarsomeres 2–5, dorsal view. 124. C. sobrinus (Wheeler), mid tarsomeres 2–5, dorsal view. 125. C. pictipes (Harmston & Knowlton), dorsal view. 126. C. ornatus Parent, mid tarsomeres 2–5, ventral view. 127. C. tohono sp. nov., mid tarsomeres 2–5, dorsal view. 128. C. remifer sp. nov., dorsal view. 129. C. frontalis (Loew), dorsal view. 130. C. furcatus (Van Duzee), dorsal view. 131. C. lacrima sp. nov., dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14240480" httpUri="https://zenodo.org/record/14240480/files/figure.png" pageId="58" pageNumber="59">Fig. 131</figureCitation>
|
||
) length subequal to tibia length. Tarsomere 1 longer than combined length of tarsomeres 2–5. Tarsomere 2 subequal to combined length of tarsomeres 3–5, slightly broadened apically with small anteroapical projection bearing a few short pale setulae. Tarsomeres 3–4 subequal in size, short, wider than long, with short pale anterior setulae (most are hooked). Tarsomere 5 distinctly longer than combined length of tarsomeres 3–4.
|
||
<emphasis id="47C7EABBA517FFDE53FEFF0CFBEFFF2A" box="[954,1044,223,249]" italics="true" pageId="59" pageNumber="60">Hindleg</emphasis>
|
||
: Yellow, except tarsus brown from near apex of tarsomere 1. Femur (
|
||
<figureCitation id="ED882A2CA517FFDE5258FED0FD86FECE" box="[540,637,259,285]" captionStart="FIGURES 132–138" captionStartId="54.[151,264,1423,1447]" captionTargetBox="[152,1436,381,1400]" captionTargetId="figure-114@54.[152,1436,381,1400]" captionTargetPageId="54" captionText="FIGURES 132–138. Male hind legs and wings of the Calyxochaetus frontalis species group. 132. C. binodatus (Harmston & Knowlton), hind tibia, anterior view. 133. C. nodatus (Loew), hind tibia, anterior view. 134. C. ornatus Parent, hind tibia, posterior view. 135. C. frontalis (Loew), hind tibia, posterior view. 136. C. lacrima sp. nov., apical third of hind femur and basal half of hind tibia, posterior view. 137. C. nodatus, wing. 138. C. ornatus, wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14240484" httpUri="https://zenodo.org/record/14240484/files/figure.png" pageId="59" pageNumber="60">Fig. 136</figureCitation>
|
||
) with row of longer ventral setae on apical half. Tibia (
|
||
<figureCitation id="ED882A2CA517FFDE54BDFED0FAA2FECE" box="[1273,1369,259,285]" captionStart="FIGURES 132–138" captionStartId="54.[151,264,1423,1447]" captionTargetBox="[152,1436,381,1400]" captionTargetId="figure-114@54.[152,1436,381,1400]" captionTargetPageId="54" captionText="FIGURES 132–138. Male hind legs and wings of the Calyxochaetus frontalis species group. 132. C. binodatus (Harmston & Knowlton), hind tibia, anterior view. 133. C. nodatus (Loew), hind tibia, anterior view. 134. C. ornatus Parent, hind tibia, posterior view. 135. C. frontalis (Loew), hind tibia, posterior view. 136. C. lacrima sp. nov., apical third of hind femur and basal half of hind tibia, posterior view. 137. C. nodatus, wing. 138. C. ornatus, wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14240484" httpUri="https://zenodo.org/record/14240484/files/figure.png" pageId="59" pageNumber="60">Fig. 136</figureCitation>
|
||
) with posteroventral row of 4–5 longer, weakly curvate setae on basal half.
|
||
<emphasis id="47C7EABBA517FFDE53EBFEF5FC02FE93" bold="true" box="[943,1017,294,320]" pageId="59" pageNumber="60">Wing:</emphasis>
|
||
Evenly tinged light brown. Elongate oval and rather narrow, with anal area somewhat reduced. Distal section of M
|
||
<subScript id="E93734ECA517FFDE5464FE8EFBD3FEB8" attach="left" box="[1056,1064,349,363]" fontSize="6" pageId="59" pageNumber="60">4</subScript>
|
||
about 3X longer than crossvein dm-m.
|
||
<emphasis id="47C7EABBA517FFDE50AEFEBDFE93FE5B" bold="true" box="[234,360,366,392]" pageId="59" pageNumber="60">Abdomen:</emphasis>
|
||
Tergite 1 yellow-brown to yellow laterally. Tergites 2–3 almost wholly yellow. Tergite 4 (and sometimes tergites 5–6) yellow ventrally at base. Sternites yellow.
|
||
<emphasis id="47C7EABBA517FFDE53C5FE40FBEBFE7F" box="[897,1040,403,428]" italics="true" pageId="59" pageNumber="60">Hypopygium</emphasis>
|
||
: Postgonites with apical lobes very narrow, sharply pointed, translucent with narrow brown stripe along dorsal edge.
|
||
</paragraph>
|
||
<paragraph id="750C36A9A517FFDE5083FE08FAC6FD7A" blockId="59.[151,1437,150,825]" pageId="59" pageNumber="60">
|
||
<emphasis id="47C7EABBA517FFDE5083FE08FEDEFE26" bold="true" box="[199,293,475,501]" pageId="59" pageNumber="60">Female.</emphasis>
|
||
Wing length
|
||
<quantity id="B24B9B4CA517FFDE51FCFE09FDBBFE26" box="[440,576,474,501]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" metricValueMax="3.5" metricValueMin="2.9" pageId="59" pageNumber="60" unit="mm" value="3.2" valueMax="3.5" valueMin="2.9">2.9–3.5 mm</quantity>
|
||
. Similar to male except as follows:
|
||
<emphasis id="47C7EABBA517FFDE5387FE08FBF0FE26" bold="true" box="[963,1035,475,501]" pageId="59" pageNumber="60">Head:</emphasis>
|
||
Face wider (subequal to postpedicel width), nearly parallel-sided, gray-silver. Antenna with postpedicel about as wide as long, rounded but apex slightly flattened in a few specimens; arista-like stylus unmodified. Palpus much larger, triangular, narrowly rounded apically, yellow-brown.
|
||
<emphasis id="47C7EABBA517FFDE51EFFD94FDF6FDB2" bold="true" box="[427,525,583,609]" pageId="59" pageNumber="60">Thorax:</emphasis>
|
||
Proepimeron with cluster of 2–4 small white hairs.
|
||
<emphasis id="47C7EABBA517FFDE5421FD94FB5DFDB2" bold="true" box="[1125,1190,583,609]" pageId="59" pageNumber="60">Legs:</emphasis>
|
||
Unmodified, without specialized setae.
|
||
<emphasis id="47C7EABBA517FFDE5124FDB9FE51FD57" bold="true" box="[352,426,618,644]" pageId="59" pageNumber="60">Wing:</emphasis>
|
||
Anal lobe broader. Distal section of M
|
||
<subScript id="E93734ECA517FFDE5319FDAEFC9EFD58" attach="left" box="[861,869,637,651]" fontSize="6" pageId="59" pageNumber="60">4</subScript>
|
||
about 2X longer than crossvein dm-m.
|
||
<emphasis id="47C7EABBA517FFDE555AFDB9FA67FD57" bold="true" box="[1310,1436,618,644]" pageId="59" pageNumber="60">Abdomen:</emphasis>
|
||
Tergites 2–3 mostly yellow, but browner along posterior margins and sometimes narrowly brown dorsally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3DA96522A517FFDE5083FD61FDA7FD22" pageId="59" pageNumber="60" type="distribution">
|
||
<paragraph id="750C36A9A517FFDE5083FD61FDA7FD22" blockId="59.[151,1437,150,825]" pageId="59" pageNumber="60">
|
||
<emphasis id="47C7EABBA517FFDE5083FD61FD68FD1E" bold="true" box="[199,659,690,717]" pageId="59" pageNumber="60">Distribution and seasonal occurrence.</emphasis>
|
||
Known only from the village of Potrero Redondo in
|
||
<collectingRegion id="B777F84BA517FFDE5543FD60FA63FD1E" box="[1287,1432,691,717]" country="Mexico" name="Nuevo Leon" pageId="59" pageNumber="60">Nuevo León</collectingRegion>
|
||
,
|
||
<collectingCountry id="0DA47639A517FFDE50D3FD04FF16FD22" box="[151,237,727,753]" name="Mexico" pageId="59" pageNumber="60">Mexico</collectingCountry>
|
||
(
|
||
<figureCitation id="ED882A2CA517FFDE50B8FD04FEA0FD22" box="[252,347,727,753]" captionStart="FIGURES 147–148" captionStartId="66.[151,264,1875,1899]" captionTargetBox="[274,1315,181,1851]" captionTargetId="figure-27@66.[271,1315,181,1851]" captionTargetPageId="66" captionText="FIGURES 147–148. Known geographical distributions of species in the Calyxochaetus frontalis group. 147. C. furcatus (Van Duzee), C. lacrima sp. nov. and C. lamellicornis Parent. 148. C. ornatus Parent, C. remifer sp. nov. and C. tohono sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14240492" httpUri="https://zenodo.org/record/14240492/files/figure.png" pageId="59" pageNumber="60">Fig. 147</figureCitation>
|
||
) and collected in May.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3DA96522A517FFDE5083FD28FDA6FCEA" pageId="59" pageNumber="60" type="etymology">
|
||
<paragraph id="750C36A9A517FFDE5083FD28FDA6FCEA" blockId="59.[151,1437,150,825]" pageId="59" pageNumber="60">
|
||
<emphasis id="47C7EABBA517FFDE5083FD28FEB5FCC6" bold="true" box="[199,334,763,789]" pageId="59" pageNumber="60">Etymology.</emphasis>
|
||
The specific epithet is the Latin noun
|
||
<taxonomicName id="B2B34D2AA517FFDE5345FD28FCA3FCC6" authorityName="Runyon" authorityYear="2024" box="[769,856,763,789]" class="Insecta" family="Dolichopodidae" genus="Calyxochaetus" kingdom="Animalia" order="Diptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="lacrima">
|
||
<emphasis id="47C7EABBA517FFDE5345FD28FCA3FCC6" box="[769,856,763,789]" italics="true" pageId="59" pageNumber="60">lacrima</emphasis>
|
||
</taxonomicName>
|
||
(teardrop) and refers to the shape of the lamella on the arista-like stylus of males (
|
||
<figureCitation id="ED882A2CA517FFDE51ABFCCCFDABFCEA" box="[495,592,798,825]" captionStart="FIGURES 104–119" captionStartId="50.[151,264,1472,1496]" captionTargetBox="[151,1437,181,1448]" captionTargetId="figure-27@50.[151,1437,181,1448]" captionTargetPageId="50" captionText="FIGURES 104–119. Male antennae (lateral view) and forelegs of the Calyxochaetus frontalis species group. 104. C. binodatus (Harmston & Knowlton). 105. C. nodatus (Loew). 106. C. desertus sp. nov. 107. C. pictipes (Harmston & Knowlton). 108. C. sobrinus (Wheeler). 109. C. ornatus Parent. 110. C. remifer sp. nov. 111. C. tohono sp. nov. 112. C. frontalis (Loew). 113. C. furcatus (Van Duzee). 114. C. lamellicornis Parent (photo by Emmanuel Delfosse, MNHN). 115. C. lacrima sp. nov. 116. C. binodatus, fore tarsus, anterior view. 117. C. sobrinus, fore tarsus, anterior view. 118. C. remifer sp. nov., fore tarsus, dorsal view. 119. C. lacrima sp. nov., fore tibia and tarsus, posterior view. Scale bars = 0.5 mm; arista-like styli approximately 1 mm long." figureDoi="http://doi.org/10.5281/zenodo.14240476" httpUri="https://zenodo.org/record/14240476/files/figure.png" pageId="59" pageNumber="60">Fig. 115</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |