351 lines
32 KiB
XML
351 lines
32 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.974.54794" ID-GBIF-Dataset="5f0b0760-7a15-49c0-9e80-fc6747f8046e" ID-PMC="PMC7557337" ID-Pensoft-Pub="1313-2970-974-31" ID-Pensoft-UUID="20CAFA6BA01359369DA3F27F90E22660" ID-PubMed="33110378" ID-ZooBank="9B8888660F074DECAE7B88DFB0A4621C" ModsDocID="1313-2970-974-31" checkinTime="1602130124706" checkinUser="pensoft" docAuthor="Wood, Thomas James, Michez, Denis, Cejas, Diego, Lhomme, Patrick & Rasmont, Pierre" docDate="2020" docId="E9D30E8D5F435192A6E97BAD7E7EA872" docLanguage="en" docName="ZooKeys 974: 31-92" docOrigin="ZooKeys 974" docSource="http://dx.doi.org/10.3897/zookeys.974.54794" docTitle="Andrena (Carandrena) hebescens Wood 2020, sp. nov." docType="treatment" docUuid="BECF6642-E2D6-479D-A132-18F49B9725B6" docUuidSource="ZooBank" docVersion="4" id="20CAFA6BA01359369DA3F27F90E22660" lastPageNumber="31" masterDocId="20CAFA6BA01359369DA3F27F90E22660" masterDocTitle="An update and revision of the Andrena fauna of Morocco (Hymenoptera, Apoidea, Andrenidae) with the description of eleven new North African species" masterLastPageNumber="92" masterPageNumber="31" pageNumber="31" updateTime="1668169591898" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>An update and revision of the Andrena fauna of Morocco (Hymenoptera, Apoidea, Andrenidae) with the description of eleven new North African species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wood, Thomas James</mods:namePart>
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<mods:affiliation>Laboratoire de Zoologie, Universite de Mons, 7000, Mons, Belgium</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5653-224X</mods:nameIdentifier>
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<mods:nameIdentifier type="email">thomasjames.wood@umons.ac.be</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Michez, Denis</mods:namePart>
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<mods:affiliation>Laboratoire de Zoologie, Universite de Mons, 7000, Mons, Belgium</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-8880-1838</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cejas, Diego</mods:namePart>
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<mods:affiliation>Laboratoire de Zoologie, Universite de Mons, 7000, Mons, Belgium</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lhomme, Patrick</mods:namePart>
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<mods:affiliation>Laboratoire de Zoologie, Universite de Mons, 7000, Mons, Belgium & International Center of Agricultural Research in the Dry Areas, Rabat, Morocco</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-6735-9104</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rasmont, Pierre</mods:namePart>
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<mods:affiliation>Laboratoire de Zoologie, Universite de Mons, 7000, Mons, Belgium</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>974</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>31</mods:start>
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<mods:end>92</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.974.54794</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.974.54794</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-974-31</mods:identifier>
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<mods:identifier type="ZooBank">9B8888660F074DECAE7B88DFB0A4621C</mods:identifier>
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<mods:identifier type="Pensoft-UUID">20CAFA6BA01359369DA3F27F90E22660</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="168243701" LSID="urn:lsid:zoobank.org:act:BECF6642-E2D6-479D-A132-18F49B9725B6" httpUri="http://treatment.plazi.org/id/E9D30E8D5F435192A6E97BAD7E7EA872" lastPageNumber="31" pageId="0" pageNumber="31">
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<subSubSection pageId="0" pageNumber="31" type="nomenclature">
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<paragraph pageId="0" pageNumber="31">
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<taxonomicName LSID="E9D30E8D-5F43-5192-A6E9-7BAD7E7EA872" authority="Wood" authorityName="Wood" authorityYear="2020" class="Insecta" family="Andrenidae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena (Carandrena) hebescens" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="hebescens" status="sp. nov." subGenus="Carandrena">Andrena (Carandrena) hebescens Wood</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="31">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figures 29–34" captionStartId="F5" captionText="Figures 29 - 34. Andrena (Carandrena) hebescens sp. nov. 29 female profile 30 female face 31 female dorsum 32 female tergites. Andrena (Carandrena) euzona Perez, 1895 33 female dorsum 34 female tergites." figureDoi="10.3897/zookeys.974.54794.figures29-34" httpUri="https://binary.pensoft.net/fig/460646" pageId="0" pageNumber="31">Figures 29-34</figureCitation>
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<figureCitation captionStart="Figures 35–40" captionStartId="F6" captionText="Figures 35 - 40. Andrena (Carandrena) hebescens sp. nov. 35 male profile 36 male face 37 male gena 38 male dorsum 39 male tergites 40 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures35-40" httpUri="https://binary.pensoft.net/fig/460647" pageId="0" pageNumber="31">, 35-40</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="31" type="materials_examined">
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<paragraph pageId="0" pageNumber="31">Material.</paragraph>
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<paragraph pageId="0" pageNumber="31">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Holotype</emphasis>
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: Morocco: Guelmim-Oued Noun, 15-16.iv.1995, 1♀, leg. Ma. Halada. Deposited in the
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<normalizedToken originalValue="OÖLM">OOELM</normalizedToken>
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.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Paratypes</emphasis>
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: Morocco: Guelmim-Oued Noun, 15-16.iv.1995, 4♂, 6♀, leg. Ma. Halada,
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<normalizedToken originalValue="OÖLM">OOELM</normalizedToken>
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; Souss-Massa, Tassademt, 50 km NE, Agadir, 19.iv.1996, 1♀, leg. M. Schwarz,
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<normalizedToken originalValue="OÖLM">OOELM</normalizedToken>
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; Souss-Massa, Aoulouz-Taliouine, 19.iii.1988, 1♂, leg. H. Teunissen. Paratypes are deposited at the
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<normalizedToken originalValue="OÖLM">OOELM</normalizedToken>
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and NMNL, with a male and female retained in the personal collection of TJW.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="31" type="diagnosis">
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<paragraph pageId="0" pageNumber="31">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="31">
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The female of
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<taxonomicName lsidName="A. hebescens" pageId="0" pageNumber="31" rank="species" species="hebescens">
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<emphasis italics="true" pageId="0" pageNumber="31">A. hebescens</emphasis>
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</taxonomicName>
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can be placed in the
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<taxonomicName authorityName="Warncke" authorityYear="1968" class="Insecta" family="Andrenidae" genus="Carandrena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Carandrena" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="31">Carandrena</emphasis>
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</taxonomicName>
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because the dorsolateral angle of the pronotum has a transverse ridge, the propodeal triangle is shagreened and weakly rugose at the base, and there is almost no punctation on the metasoma, and the head has a typical
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<taxonomicName authorityName="Warncke" authorityYear="1968" class="Insecta" family="Andrenidae" genus="Carandrena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Carandrena" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="31">Carandrena</emphasis>
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</taxonomicName>
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shape, broader than long, with the inner eye margins slightly converging below (Fig.
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<figureCitation captionStart="Figures 29–34" captionStartId="F5" captionText="Figures 29 - 34. Andrena (Carandrena) hebescens sp. nov. 29 female profile 30 female face 31 female dorsum 32 female tergites. Andrena (Carandrena) euzona Perez, 1895 33 female dorsum 34 female tergites." figureDoi="10.3897/zookeys.974.54794.figures29-34" httpUri="https://binary.pensoft.net/fig/460646" pageId="0" pageNumber="31">30</figureCitation>
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). It is most similar to
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<taxonomicName lsidName="A. euzona" pageId="0" pageNumber="31" rank="species" species="euzona">
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<emphasis italics="true" pageId="0" pageNumber="31">A. euzona</emphasis>
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</taxonomicName>
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<normalizedToken originalValue="Pérez">Perez</normalizedToken>
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, 1895 and
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<taxonomicName lsidName="A. microthorax" pageId="0" pageNumber="31" rank="species" species="microthorax">
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<emphasis italics="true" pageId="0" pageNumber="31">A. microthorax</emphasis>
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</taxonomicName>
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<normalizedToken originalValue="Pérez">Perez</normalizedToken>
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, 1895 given its dark, non-metallic appearance (
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<taxonomicName lsidName="A. aerinifrons" pageId="0" pageNumber="31" rank="species" species="aerinifrons">
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<emphasis italics="true" pageId="0" pageNumber="31">A. aerinifrons</emphasis>
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</taxonomicName>
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Dours, 1873,
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<taxonomicName lsidName="A. bellidis" pageId="0" pageNumber="31" rank="species" species="bellidis">
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<emphasis italics="true" pageId="0" pageNumber="31">A. bellidis</emphasis>
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</taxonomicName>
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<normalizedToken originalValue="Pérez">Perez</normalizedToken>
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, 1895,
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<taxonomicName lsidName="A. daphanea" pageId="0" pageNumber="31" rank="species" species="daphanea">
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<emphasis italics="true" pageId="0" pageNumber="31">A. daphanea</emphasis>
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</taxonomicName>
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Warncke, 1974,
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<taxonomicName lsidName="A. deserta" pageId="0" pageNumber="31" rank="species" species="deserta">
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<emphasis italics="true" pageId="0" pageNumber="31">A. deserta</emphasis>
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</taxonomicName>
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Warncke, 1974,
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<taxonomicName lsidName="A. nigroviridiula" pageId="0" pageNumber="31" rank="species" species="nigroviridiula">
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<emphasis italics="true" pageId="0" pageNumber="31">A. nigroviridiula</emphasis>
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</taxonomicName>
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Dours, 1873, and
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<taxonomicName lsidName="A. reperta" pageId="0" pageNumber="31" rank="species" species="reperta">
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<emphasis italics="true" pageId="0" pageNumber="31">A. reperta</emphasis>
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</taxonomicName>
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, Warncke, 1974 with metallic green or blue integument,
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<taxonomicName lsidName="A. binominata" pageId="0" pageNumber="31" rank="species" species="binominata">
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<emphasis italics="true" pageId="0" pageNumber="31">A. binominata</emphasis>
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</taxonomicName>
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Smith, 1853,
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<taxonomicName lsidName="A. eremobia" pageId="0" pageNumber="31" rank="species" species="eremobia">
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<emphasis italics="true" pageId="0" pageNumber="31">A. eremobia</emphasis>
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</taxonomicName>
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Guiglia, 1933, and
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<taxonomicName lsidName="A. leucophaea" pageId="0" pageNumber="31" rank="species" species="leucophaea">
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<emphasis italics="true" pageId="0" pageNumber="31">A. leucophaea</emphasis>
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</taxonomicName>
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Lepeletier, 1841 with partially red metasoma), dull domed clypeus, and white hair bands on the tergites (
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<taxonomicName lsidName="A. eddaensis" pageId="0" pageNumber="31" rank="species" species="eddaensis">
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<emphasis italics="true" pageId="0" pageNumber="31">A. eddaensis</emphasis>
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</taxonomicName>
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Gusenleitner, 1998 and
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<taxonomicName lsidName="A. decaocta" pageId="0" pageNumber="31" rank="species" species="decaocta">
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<emphasis italics="true" pageId="0" pageNumber="31">A. decaocta</emphasis>
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</taxonomicName>
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Warncke, 1967 with felt-like hair on the tergal discs). However, the scutum is shagreened and only weakly shining (Fig.
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<figureCitation captionStart="Figures 29–34" captionStartId="F5" captionText="Figures 29 - 34. Andrena (Carandrena) hebescens sp. nov. 29 female profile 30 female face 31 female dorsum 32 female tergites. Andrena (Carandrena) euzona Perez, 1895 33 female dorsum 34 female tergites." figureDoi="10.3897/zookeys.974.54794.figures29-34" httpUri="https://binary.pensoft.net/fig/460646" pageId="0" pageNumber="31">31</figureCitation>
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) whereas in
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<taxonomicName lsidName="A. euzona" pageId="0" pageNumber="31" rank="species" species="euzona">
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<emphasis italics="true" pageId="0" pageNumber="31">A. euzona</emphasis>
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</taxonomicName>
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it is extensively smooth and shiny (Fig.
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<figureCitation captionStart="Figures 29–34" captionStartId="F5" captionText="Figures 29 - 34. Andrena (Carandrena) hebescens sp. nov. 29 female profile 30 female face 31 female dorsum 32 female tergites. Andrena (Carandrena) euzona Perez, 1895 33 female dorsum 34 female tergites." figureDoi="10.3897/zookeys.974.54794.figures29-34" httpUri="https://binary.pensoft.net/fig/460646" pageId="0" pageNumber="31">33</figureCitation>
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), and the hair bands are narrower (Fig.
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<figureCitation captionStart="Figures 29–34" captionStartId="F5" captionText="Figures 29 - 34. Andrena (Carandrena) hebescens sp. nov. 29 female profile 30 female face 31 female dorsum 32 female tergites. Andrena (Carandrena) euzona Perez, 1895 33 female dorsum 34 female tergites." figureDoi="10.3897/zookeys.974.54794.figures29-34" httpUri="https://binary.pensoft.net/fig/460646" pageId="0" pageNumber="31">32</figureCitation>
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) whereas in
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<taxonomicName lsidName="A. euzona" pageId="0" pageNumber="31" rank="species" species="euzona">
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<emphasis italics="true" pageId="0" pageNumber="31">A. euzona</emphasis>
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</taxonomicName>
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they are wider (Fig.
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<figureCitation captionStart="Figures 29–34" captionStartId="F5" captionText="Figures 29 - 34. Andrena (Carandrena) hebescens sp. nov. 29 female profile 30 female face 31 female dorsum 32 female tergites. Andrena (Carandrena) euzona Perez, 1895 33 female dorsum 34 female tergites." figureDoi="10.3897/zookeys.974.54794.figures29-34" httpUri="https://binary.pensoft.net/fig/460646" pageId="0" pageNumber="31">34</figureCitation>
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). The clypeus is also evenly punctured and shagreened, weakly shining with only a subtle impunctate mid-line whereas in
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<taxonomicName lsidName="A. microthorax" pageId="0" pageNumber="31" rank="species" species="microthorax">
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<emphasis italics="true" pageId="0" pageNumber="31">A. microthorax</emphasis>
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</taxonomicName>
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it is shiny and strongly punctured with a conspicuous impunctate mid-line, particularly at the fore margin of the clypeus where it forms a broad impunctate triangle.
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</paragraph>
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<caption doi="10.3897/zookeys.974.54794.figures29-34" httpUri="https://binary.pensoft.net/fig/460646" pageId="0" pageNumber="31" start="Figures 29–34" startId="F5">
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<paragraph pageId="0" pageNumber="31">
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<emphasis bold="true" pageId="0" pageNumber="31">Figures 29-34.</emphasis>
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<taxonomicName authorityName="Wood" authorityYear="2020" class="Insecta" family="Andrenidae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena (Carandrena) hebescens" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="hebescens" subGenus="Carandrena">Andrena (Carandrena) hebescens</taxonomicName>
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sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="31">29</emphasis>
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female profile
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<emphasis bold="true" pageId="0" pageNumber="31">30</emphasis>
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female face
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<emphasis bold="true" pageId="0" pageNumber="31">31</emphasis>
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female dorsum
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<emphasis bold="true" pageId="0" pageNumber="31">32</emphasis>
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female tergites.
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<taxonomicName class="Insecta" family="Andrenidae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena (Carandrena) euzona" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="euzona" subGenus="Carandrena">Andrena (Carandrena) euzona</taxonomicName>
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<normalizedToken originalValue="Pérez">Perez</normalizedToken>
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, 1895
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<emphasis bold="true" pageId="0" pageNumber="31">33</emphasis>
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female dorsum
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<emphasis bold="true" pageId="0" pageNumber="31">34</emphasis>
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female tergites.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="31">
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The male of
|
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<taxonomicName lsidName="A. hebescens" pageId="0" pageNumber="31" rank="species" species="hebescens">
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<emphasis italics="true" pageId="0" pageNumber="31">A. hebescens</emphasis>
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</taxonomicName>
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has a yellow marking on the clypeus, but this is reduced in size and does not cover the entire clypeal surface, with two black markings that extend in from the clypeal margins giving the marking an inverted
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<normalizedToken originalValue="‘T’">'T'</normalizedToken>
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shape (Fig.
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<figureCitation captionStart="Figures 35–40" captionStartId="F6" captionText="Figures 35 - 40. Andrena (Carandrena) hebescens sp. nov. 35 male profile 36 male face 37 male gena 38 male dorsum 39 male tergites 40 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures35-40" httpUri="https://binary.pensoft.net/fig/460647" pageId="0" pageNumber="31">36</figureCitation>
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). The male is therefore superficially similar to
|
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<taxonomicName lsidName="A. semiadesus" pageId="0" pageNumber="31" rank="species" species="semiadesus">
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<emphasis italics="true" pageId="0" pageNumber="31">A. semiadesus</emphasis>
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</taxonomicName>
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, but can be separated by the typical
|
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<taxonomicName authorityName="Warncke" authorityYear="1968" class="Insecta" family="Andrenidae" genus="Carandrena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Carandrena" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="31">Carandrena</emphasis>
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</taxonomicName>
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head which is clearly wider (more than 1.5 times) than the width of a compound eye in
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<taxonomicName lsidName="A. hebescens" pageId="0" pageNumber="31" rank="species" species="hebescens">
|
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<emphasis italics="true" pageId="0" pageNumber="31">A. hebescens</emphasis>
|
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</taxonomicName>
|
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(Figs
|
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<figureCitation captionStart="Figures 35–40" captionStartId="F6" captionText="Figures 35 - 40. Andrena (Carandrena) hebescens sp. nov. 35 male profile 36 male face 37 male gena 38 male dorsum 39 male tergites 40 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures35-40" httpUri="https://binary.pensoft.net/fig/460647" pageId="0" pageNumber="31">37</figureCitation>
|
||
,
|
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<figureCitation captionStart="Figures 35–40" captionStartId="F6" captionText="Figures 35 - 40. Andrena (Carandrena) hebescens sp. nov. 35 male profile 36 male face 37 male gena 38 male dorsum 39 male tergites 40 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures35-40" httpUri="https://binary.pensoft.net/fig/460647" pageId="0" pageNumber="31">38</figureCitation>
|
||
) but only slightly wider in
|
||
<taxonomicName lsidName="A. semiadesus" pageId="0" pageNumber="31" rank="species" species="semiadesus">
|
||
<emphasis italics="true" pageId="0" pageNumber="31">A. semiadesus</emphasis>
|
||
</taxonomicName>
|
||
and lacking an angulate hind corner. The male of
|
||
<taxonomicName lsidName="A. microthorax" pageId="0" pageNumber="31" rank="species" species="microthorax">
|
||
<emphasis italics="true" pageId="0" pageNumber="31">A. microthorax</emphasis>
|
||
</taxonomicName>
|
||
can have a yellow clypeal marking, but the scutum and tergites of this species are shiny, whereas in
|
||
<taxonomicName lsidName="A. hebescens" pageId="0" pageNumber="31" rank="species" species="hebescens">
|
||
<emphasis italics="true" pageId="0" pageNumber="31">A. hebescens</emphasis>
|
||
</taxonomicName>
|
||
they are shagreened and at most, weakly shining (Fig.
|
||
<figureCitation captionStart="Figures 35–40" captionStartId="F6" captionText="Figures 35 - 40. Andrena (Carandrena) hebescens sp. nov. 35 male profile 36 male face 37 male gena 38 male dorsum 39 male tergites 40 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures35-40" httpUri="https://binary.pensoft.net/fig/460647" pageId="0" pageNumber="31">39</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="31" type="description">
|
||
<paragraph pageId="0" pageNumber="31">Description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="31">
|
||
<emphasis bold="true" pageId="0" pageNumber="31">Female</emphasis>
|
||
: Body length 8 mm (Fig.
|
||
<figureCitation captionStart="Figures 29–34" captionStartId="F5" captionText="Figures 29 - 34. Andrena (Carandrena) hebescens sp. nov. 29 female profile 30 female face 31 female dorsum 32 female tergites. Andrena (Carandrena) euzona Perez, 1895 33 female dorsum 34 female tergites." figureDoi="10.3897/zookeys.974.54794.figures29-34" httpUri="https://binary.pensoft.net/fig/460646" pageId="0" pageNumber="31">29</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Head</emphasis>
|
||
: Black, clearly wider than long (Fig.
|
||
<figureCitation captionStart="Figures 29–34" captionStartId="F5" captionText="Figures 29 - 34. Andrena (Carandrena) hebescens sp. nov. 29 female profile 30 female face 31 female dorsum 32 female tergites. Andrena (Carandrena) euzona Perez, 1895 33 female dorsum 34 female tergites." figureDoi="10.3897/zookeys.974.54794.figures29-34" httpUri="https://binary.pensoft.net/fig/460646" pageId="0" pageNumber="31">30</figureCitation>
|
||
). Clypeus arched, shagreened, weakly shining. Clypeus evenly punctured with exception of a subtle, impunctate midline, punctures otherwise separated by 0.5-1 puncture diameters. Process of labrum trapezoidal, fore margin very weakly emarginate. Gena as wide as width of compound eye. Gena, face, and scape with moderately dense white hairs, the longest not achieving length of the scape. Vertex with whitish brown hairs of a similar length. Foveae normal, occupying half the distance between the compound eye and a lateral ocellus. Antennae dark, scape black, A2-4 apically lightened to orange, A5-12 predominantly orange ventrally, A3 exceeding A4+5, shorter than A4+5+6. Ocelloccipital distance short, less than 1/3 width of lateral ocellus.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Mesosoma</emphasis>
|
||
: Scutum dark, evenly and shallowly punctured, punctures separated by 1-2 puncture diameters, underlying surface shagreened, weakly shining (Fig.
|
||
<figureCitation captionStart="Figures 29–34" captionStartId="F5" captionText="Figures 29 - 34. Andrena (Carandrena) hebescens sp. nov. 29 female profile 30 female face 31 female dorsum 32 female tergites. Andrena (Carandrena) euzona Perez, 1895 33 female dorsum 34 female tergites." figureDoi="10.3897/zookeys.974.54794.figures29-34" httpUri="https://binary.pensoft.net/fig/460646" pageId="0" pageNumber="31">31</figureCitation>
|
||
). Scutellum less densely punctured, punctures separated by 3-4 puncture diameters, shagreenation weaker, more strongly shining. Episternum and propodeum microreticulate, dull, propodeal triangle slightly more finely shagreened, weakly shining, distinct. Scutum and scutellum with faded light brownish hairs, episternum and propodeum with longer white hairs, the longest achieving the length of the scape. Legs dark, tarsi lightened brown, pubescence white, femoral and tibial scopa white, hairs simple. Wings hyaline, venation and stigma golden brown. Nervulus interstitial to slightly antefurcal.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Metasoma</emphasis>
|
||
: Tergites dark with wide lightened margins, apically translucent, basally yellowish (Fig.
|
||
<figureCitation captionStart="Figures 29–34" captionStartId="F5" captionText="Figures 29 - 34. Andrena (Carandrena) hebescens sp. nov. 29 female profile 30 female face 31 female dorsum 32 female tergites. Andrena (Carandrena) euzona Perez, 1895 33 female dorsum 34 female tergites." figureDoi="10.3897/zookeys.974.54794.figures29-34" httpUri="https://binary.pensoft.net/fig/460646" pageId="0" pageNumber="31">32</figureCitation>
|
||
). Tergal discs microreticulate, weakly shining. T1 shallowly punctured, punctures separated by two puncture diameters. Following tergites weakly and obscurely punctured, punctures hidden by microreticulation. Tergal margins with dense white hairbands, on T1 widely interrupted, on T2-4 complete. T5+6 centrally with golden hairs flanking pygidial plate, laterally with white hairs. Sternites with plumose white hairs, forming loose fringes apically on hind margins.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="31">
|
||
<emphasis bold="true" pageId="0" pageNumber="31">Male.</emphasis>
|
||
Body length 8 mm (Fig.
|
||
<figureCitation captionStart="Figures 35–40" captionStartId="F6" captionText="Figures 35 - 40. Andrena (Carandrena) hebescens sp. nov. 35 male profile 36 male face 37 male gena 38 male dorsum 39 male tergites 40 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures35-40" httpUri="https://binary.pensoft.net/fig/460647" pageId="0" pageNumber="31">35</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Head</emphasis>
|
||
: Similar to female, clypeus slightly arched, evenly and shallowly punctured, punctures separated by one puncture diameter, no impunctate central line. Clypeus centrally with yellow mark, this not reaching the lateral or basal margin of the clypeus, laterally invaded by two black marks therefore forming a broad inverted T-shape (Fig.
|
||
<figureCitation captionStart="Figures 35–40" captionStartId="F6" captionText="Figures 35 - 40. Andrena (Carandrena) hebescens sp. nov. 35 male profile 36 male face 37 male gena 38 male dorsum 39 male tergites 40 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures35-40" httpUri="https://binary.pensoft.net/fig/460647" pageId="0" pageNumber="31">36</figureCitation>
|
||
). Underlying surface weakly shagreened, moderately shiny. Process of labrum trapezoidal, fore margin inflated, slightly bulbous, very weakly emarginate. Antennae dark, A4-13 slightly lightened dark brown, A3 exceeding A4, shorter than A4+5. Gena enlarged, most 1.2 times wider than compound eye, non-carinate, with weakly angulate hind corner (Fig.
|
||
<figureCitation captionStart="Figures 35–40" captionStartId="F6" captionText="Figures 35 - 40. Andrena (Carandrena) hebescens sp. nov. 35 male profile 36 male face 37 male gena 38 male dorsum 39 male tergites 40 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures35-40" httpUri="https://binary.pensoft.net/fig/460647" pageId="0" pageNumber="31">37</figureCitation>
|
||
). Gena, vertex, and face below the level of the antennal insertions with long white hair equalling length of the scape. Scape, frons, and inner margin of compound eyes with mixture of black and white hairs. Ocelloccipital distance short, 2/3 width of lateral ocellus.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Mesosoma</emphasis>
|
||
: Similar to female but scutum with stronger shagreenation, dull except for central shining line (Fig.
|
||
<figureCitation captionStart="Figures 35–40" captionStartId="F6" captionText="Figures 35 - 40. Andrena (Carandrena) hebescens sp. nov. 35 male profile 36 male face 37 male gena 38 male dorsum 39 male tergites 40 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures35-40" httpUri="https://binary.pensoft.net/fig/460647" pageId="0" pageNumber="31">38</figureCitation>
|
||
). Scutellum centrally weakly shining, contrasting with the scutum. Episternum, propodeum, and mesosomal pubescence as in the female. Legs dark, tarsi lightened brown, with white pubescence. Wings hyaline, venation dark brown, stigma centrally light brown. Nervulus interstitial to slightly antefurcal.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Metasoma</emphasis>
|
||
: Similar to the female. Tergites more clearly punctured, punctures visible against the microreticulation with slightly raised margins giving the overall surface an uneven impression (Fig.
|
||
<figureCitation captionStart="Figures 35–40" captionStartId="F6" captionText="Figures 35 - 40. Andrena (Carandrena) hebescens sp. nov. 35 male profile 36 male face 37 male gena 38 male dorsum 39 male tergites 40 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures35-40" httpUri="https://binary.pensoft.net/fig/460647" pageId="0" pageNumber="31">39</figureCitation>
|
||
). T1-5 consistently punctured, punctures separated by 3-4 puncture diameters. T2-4 laterally with weak fringes of white hair, T5-6 with complete fringes of whitish to golden hairs. Sternites forming loose white hair bands apically. Genitalia simple, of a typical
|
||
<taxonomicName authorityName="Warncke" authorityYear="1968" class="Insecta" family="Andrenidae" genus="Carandrena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Carandrena" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="31">Carandrena</emphasis>
|
||
</taxonomicName>
|
||
form, gonocoxites apically forming weak points, rounded, diverging apically (Fig.
|
||
<figureCitation captionStart="Figures 35–40" captionStartId="F6" captionText="Figures 35 - 40. Andrena (Carandrena) hebescens sp. nov. 35 male profile 36 male face 37 male gena 38 male dorsum 39 male tergites 40 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures35-40" httpUri="https://binary.pensoft.net/fig/460647" pageId="0" pageNumber="31">40</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.974.54794.figures35-40" httpUri="https://binary.pensoft.net/fig/460647" pageId="0" pageNumber="31" start="Figures 35–40" startId="F6">
|
||
<paragraph pageId="0" pageNumber="31">
|
||
<emphasis bold="true" pageId="0" pageNumber="31">Figures 35-40.</emphasis>
|
||
<taxonomicName authorityName="Wood" authorityYear="2020" class="Insecta" family="Andrenidae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena (Carandrena) hebescens" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="hebescens" subGenus="Carandrena">Andrena (Carandrena) hebescens</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="31">35</emphasis>
|
||
male profile
|
||
<emphasis bold="true" pageId="0" pageNumber="31">36</emphasis>
|
||
male face
|
||
<emphasis bold="true" pageId="0" pageNumber="31">37</emphasis>
|
||
male gena
|
||
<emphasis bold="true" pageId="0" pageNumber="31">38</emphasis>
|
||
male dorsum
|
||
<emphasis bold="true" pageId="0" pageNumber="31">39</emphasis>
|
||
male tergites
|
||
<emphasis bold="true" pageId="0" pageNumber="31">40</emphasis>
|
||
male genitalia.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="31" type="distribution">
|
||
<paragraph pageId="0" pageNumber="31">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="31">
|
||
South-western Morocco in the Souss valley (Fig.
|
||
<figureCitation captionStart="Figure 145" captionStartId="F24" captionText="Figure 145. Distribution of sampling locations of newly described species A Andrena hebescens (circles), Andrena semiadesus (squares) B Andrena tenebricorpus (diamond), Andrena niveofacies (circle), Andrena triangulivalvis (triangle) C Andrena sparsipunctata (circles), Andrena breviceps (diamonds) D Andrena acutidentis (squares), Andrena farinosoides (triangles), Andrena nigriclypeus (circle). Relief is indicated by shading, measured in meters above sea level." figureDoi="10.3897/zookeys.974.54794.figure145" httpUri="https://binary.pensoft.net/fig/460665" pageId="0" pageNumber="31">145a</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="31" type="floral preferences">
|
||
<paragraph pageId="0" pageNumber="31">Floral preferences.</paragraph>
|
||
<paragraph pageId="0" pageNumber="31">None recorded.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="31" type="etymology">
|
||
<paragraph pageId="0" pageNumber="31">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="31">
|
||
The name
|
||
<taxonomicName authorityName="Wood" authorityYear="2020" class="Insecta" family="Andrenidae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="hebescens" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="hebescens">
|
||
<emphasis italics="true" pageId="0" pageNumber="31">hebescens</emphasis>
|
||
</taxonomicName>
|
||
was chosen because this member of the
|
||
<taxonomicName authorityName="Warncke" authorityYear="1968" class="Insecta" family="Andrenidae" genus="Carandrena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Carandrena" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="31">Carandrena</emphasis>
|
||
</taxonomicName>
|
||
, though morphologically similar to several species with metallic green colouration, is completely dark, therefore
|
||
<emphasis italics="true" pageId="0" pageNumber="31">heb</emphasis>
|
||
- (dull or blunt) +
|
||
<emphasis italics="true" pageId="0" pageNumber="31">escens</emphasis>
|
||
(becoming).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |