244 lines
28 KiB
XML
244 lines
28 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.863.34662" ID-GBIF-Dataset="af839023-d05c-4769-b2a1-f9f9f8561df7" ID-PMC="PMC6639351" ID-Pensoft-Pub="1313-2970-863-35" ID-PubMed="31341392" ID-ZBK="763EDE2B5F0C414D8289D37765E993E4" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-863-35" ModsDocOrigin="ZooKeys 863" ModsDocTitle="Unexpected palaeodiversity of omaliine rove beetles in Eocene Baltic amber (Coleoptera, Staphylinidae, Omaliinae)" checkinTime="1562893176132" checkinUser="pensoft" docAuthor="Shavrin, Alexey V. & Yamamoto, Shuhei" docDate="2019" docId="6D603CEA25B9DC50B3E6794424327770" docLanguage="en" docName="ZooKeys 863: 35-83" docOrigin="ZooKeys 863" docSource="http://dx.doi.org/10.3897/zookeys.863.34662" docTitle="Phyllodrepa daedali Shavrin & Yamamoto, sp. nov." docType="treatment" docUuid="BDE372B6-F773-433D-B98E-C0E272FB8FA7" docUuidSource="ZooBank" docVersion="5" lastPageNumber="35" masterDocId="FFE7E644384D885DFFF3E730B1230775" masterDocTitle="Unexpected palaeodiversity of omaliine rove beetles in Eocene Baltic amber (Coleoptera, Staphylinidae, Omaliinae)" masterLastPageNumber="83" masterPageNumber="35" pageNumber="35" updateTime="1668167561321" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Unexpected palaeodiversity of omaliine rove beetles in Eocene Baltic amber (Coleoptera, Staphylinidae, Omaliinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shavrin, Alexey V.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yamamoto, Shuhei</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>863</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>35</mods:start>
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<mods:end>83</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.863.34662</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.863.34662</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-863-35</mods:identifier>
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<mods:identifier type="ZBK">763EDE2B5F0C414D8289D37765E993E4</mods:identifier>
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<mods:identifier type="ZooBank">763EDE2B5F0C414D8289D37765E993E4</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="158605288" LSID="urn:lsid:zoobank.org:act:BDE372B6-F773-433D-B98E-C0E272FB8FA7" httpUri="http://treatment.plazi.org/id/6D603CEA25B9DC50B3E6794424327770" lastPageNumber="35" pageId="0" pageNumber="35">
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<subSubSection pageId="0" pageNumber="35" type="nomenclature">
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<paragraph pageId="0" pageNumber="35">
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†
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<taxonomicName LSID="http://zoobank.org/BDE372B6-F773-433D-B98E-C0E272FB8FA7" authority="Shavrin & Yamamoto" class="Insecta" family="Staphylinidae" genus="Phyllodrepa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodrepa daedali" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="daedali">Phyllodrepa daedali Shavrin & Yamamoto</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="35">sp. nov.</taxonomicNameLabel>
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Figures 7-9, 54-56, 57-64
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="35" type="type material">
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<paragraph pageId="0" pageNumber="35">Type materials examined.</paragraph>
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<paragraph pageId="0" pageNumber="35">
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Holotype (male), FMNHINS-3260629, complete specimen as inclusion in very small piece of light yellow Baltic amber, 9.3 mm
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<normalizedToken originalValue="×">x</normalizedToken>
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5.9 mm
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<normalizedToken originalValue="×">x</normalizedToken>
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2.9 mm in size (Figs 7-9), with the following labels: "16 [printed] 02 [handwritten] SYAC 0 [printed] 294 [handwritten] | Baltic / Burmite | Other: | Larva / Adult | Omalium sp. [handwritten] | Omaliinae [handwritten] | Yantarny, Kaliningrad [handwritten] |
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<normalizedToken originalValue="Shûhei">Shuhei</normalizedToken>
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<normalizedToken originalValue="Yamamoto’s">Yamamoto's</normalizedToken>
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| Amber Collection" <large rectangular label, printed>, "[FMNH barcode at left side of label] FMNHINS | 3260629 | AMBER [handwritten] | FIELD MUSEUM | Wet" <small rectangular label, printed>, "HOLOTYPE | Phyllodrepa | daedali sp. nov. | Shavrin A. & Yamamoto S. des. 2019" <red rectangular label, printed> (FMNH).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="35" type="preservation">
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<paragraph pageId="0" pageNumber="35">Preservation.</paragraph>
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<paragraph pageId="0" pageNumber="35">The specimen is relatively well preserved and many details are visible, from the dorsal, ventral and lateral sides (Figs 7-9). However, most body parts, except the dorsal surface of the head, are covered with cloud of milky substance, especially most of the ventral side.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="35" type="locality and horizon">
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<paragraph pageId="0" pageNumber="35">Locality and horizon.</paragraph>
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<paragraph pageId="0" pageNumber="35">
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Baltic amber from Yantarny, Kaliningrad, westernmost Russia; mid-Eocene (ca 44 Ma;
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<bibRefCitation pageId="0" pageNumber="35">Wappler 2005</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="35" type="description">
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<paragraph pageId="0" pageNumber="35">Description.</paragraph>
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<paragraph pageId="0" pageNumber="35">
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Measurements: HW: 0.32; HL: 0.22; OL: 0.11; AL: 0.52; PML
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<normalizedToken originalValue="×">x</normalizedToken>
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PMW (III, IV): III: 0.02
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<normalizedToken originalValue="×">x</normalizedToken>
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0.02, IV: 0.07
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<normalizedToken originalValue="×">x</normalizedToken>
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0.02; PL: 0.35; PW: 0.48; ESL: 0.56; EW: 0.51; MTbL: 0.31; MTrL: 0.20 (
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<normalizedToken originalValue="I–IV">I-IV</normalizedToken>
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: 0.08; V: 0.12); AW: 0.50; TL: ~1.80. Antennomeres with lengths
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<normalizedToken originalValue="×">x</normalizedToken>
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widths: 1: 0.08
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<normalizedToken originalValue="×">x</normalizedToken>
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0.03; 2: 0.06
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<normalizedToken originalValue="×">x</normalizedToken>
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0.02; 3-4: 0.05
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<normalizedToken originalValue="×">x</normalizedToken>
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0.02; 5: 0.04
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<normalizedToken originalValue="×">x</normalizedToken>
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0.02; 6: 0.04
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<normalizedToken originalValue="×">x</normalizedToken>
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0.03; 7: 0.03
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<normalizedToken originalValue="×">x</normalizedToken>
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0.03; 8-10: 0.03
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<normalizedToken originalValue="×">x</normalizedToken>
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0.04; 11: 0.08
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<normalizedToken originalValue="×">x</normalizedToken>
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0.04.
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</paragraph>
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<paragraph pageId="0" pageNumber="35">Body elongate and slightly convex, glossy (Fig. 54), reddish-brown, with darker head and abdomen; mouthparts, antennae, legs and apical margins of abdominal sclerites yellow-brown. Body lateroventrally as in Figure 55 and laterally as in Figure 56. Lateral margins of pronotum (Figs 54, 57), paratergites and abdominal tergite VIII (Fig. 64) with several long erect setae.</paragraph>
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<paragraph pageId="0" pageNumber="35">Head 1.4 times as wide as long, with slightly convex median portion and slight oval lateroapical impressions (Fig. 57), with sparse, small and moderately deep punctation, with shallow postocular carina. Eyes large and broadly convex (Figs 55-57, 60). Ocelli large and convex, situated at level of posterior margins of eyes, distance between ocelli much more than twice as long as distance between ocellus and posterior margin of eye; grooves in front of ocelli present, moderately deep and short (Fig. 57). Apical segment of maxillary palp significantly longer than small penultimate segment, from swollen middle gradually narrowing apicad (Figs 56, 57, 63). Antenna moderately short, just surpassing basal margin of pronotum, with sparse very long setae on antennomeres 5-11; basal and antennomere 2 swollen and elongate, 3 and 4 narrow and elongate, 5 ovoid, 6 and 7 slightly transverse and 8-10 distinctly transverse, apical antennomere large, strongly narrowing from about middle apicad (Figs 54-58).</paragraph>
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<caption pageId="0" pageNumber="35">
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<paragraph pageId="0" pageNumber="35">
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Figures 57-64.
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Phyllodrepa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodrepa daedali" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="daedali">Phyllodrepa daedali</taxonomicName>
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sp. nov. 57 head and pronotum, laterodorsal view 58 left antenna, ventral view 59 thorax, legs and abdomen, lateroventral view 60 head and prothorax, ventral view 61 protarsus, dorsal view 62 hind tarsi, lateral view 63 head and maxillar palpi, dorsolateral view 64 apex of abdomen, ventral view. Abbreviations: a1-a11 = antennomeres 1-11; amp = apical maxillary palpomere; ip = intercoxal process; mp2-mp4 = maxillary palpomeres 2-4; mds = modified setae; mst = mesotibia; msv = mesoventrite; mt1-mt5 = metatarsomeres 1-5; mtv = metaventrite; oc = ocellus; s8 = sternite VIII. Scale bars: 0.1 mm (58, 61-63), 0.2 mm (57, 60, 64), 0.3 mm (59).
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="35">Pronotum slightly convex, without longitudinal impressions, 1.3 times as wide as long, 1.5 times as wide as head, from middle distinctly more narrowed posterad than apicad, with widely rounded anterior and obtuse posterior angles; apical margin widely rounded, slightly shorter than somewhat straight posterior margin; lateral margins slightly sinuate posteriorly, narrowly emarginate and finely crenulate; lateroposterior portions with indistinct, moderately wide impressions (Figs 54, 57). Dorsal surface of pronotum without visible microsculpture between punctures, with dense, very large and deep punctation, markedly sparser in lateral and smaller in apical and basal portions (Figs 54, 57). Prosternum with wide procoxal fissures and moderately short prosternal process, with acute apex (Fig. 60). Scutellum moderately large, triangular, with somewhat rounded apex, without visible punctures or microsculpture (Fig. 54).</paragraph>
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<paragraph pageId="0" pageNumber="35">Elytra slightly convex, longer than wide, 1.6 times as long as pronotum, reaching basal margin of abdominal tergite IV, with somewhat parallel lateral sides and widely rounded lateroapical angles, with sutural apices truncate to very oblique (Fig. 54). Punctation as that in pronotum, but shallower and somewhat smoothed on apical portion, smaller and sparser on basal and apical portions. Surface between punctures with shallow dense isodiametric microsculpture.</paragraph>
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<paragraph pageId="0" pageNumber="35">Legs long and slender, similar in shape, with moderately wide femora; tibiae slender, gradually widened apicad, covered by elongate setae, denser and stronger on inner margins, and with a few strong spines near apex and additional spine on outer margin in apical third (Figs 55, 56, 59); tarsi long, with apical metatarsomere distinctly longer than previous tarsomeres together (Figs 59, 61, 62); protarsus as in Figure 61, with long tenent setae (probably only in males); tarsal claw simple (Figs 61, 62).</paragraph>
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<paragraph pageId="0" pageNumber="35">
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Abdomen markedly convex, slightly narrower at base than elytra; wing-folding patches in middle of tergite IV and/or V not visible; intersegmental membranes between tergites
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<normalizedToken originalValue="IV–VII">IV-VII</normalizedToken>
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with brick-wall sculpture, apical margin of tergite VII with indistinct very narrow palisade fringe (Fig. 54). Abdominal tergites without visible punctation, with large distinct transverse microsculpture.
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</paragraph>
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<paragraph pageId="0" pageNumber="35">Male. First four protarsomeres wide (Figs 55, 59, 61); ventral surface of protarsomeres 1-4 with several rows of modified tenent setae (not all details visible) consisting of internal rows formed by markedly elongate setae with leaf-shaped apical parts (Figs 55, 56, 59, 61); ventral surface of mesotarsomeres 1-4 with two rows of elongate setae with broadened apical parts as that in protarsomeres 1-4 but without additional internal rows (Figs 55, 56); metatarsi as in Figure 62. Apical margin of abdominal tergite VIII straight.</paragraph>
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<paragraph pageId="0" pageNumber="35">Female unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="35" type="etymology">
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<paragraph pageId="0" pageNumber="35">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="35">
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The specific epithet is the Latinized name of
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<taxonomicName class="Insecta" family="Tettigoniidae" genus="Daedalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Daedalus" order="Orthoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="genus">Daedalus</taxonomicName>
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, -i, m, the Greek architect of the times of Theseus and Minos, and father of Icarus.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="35" type="remarks">
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<paragraph pageId="0" pageNumber="35">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="35">
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In external characters such as proportions of the body, antennomeres, and maxillary palpomeres, and, more substantially, by the proportions of tarsi with elongate apical tarsomere, the fossil undoubtedly belongs to the tribe
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<taxonomicName lsidName="" pageId="0" pageNumber="35" rank="tribe" tribe="Omaliini">Omaliini</taxonomicName>
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. Based on the triangular and elongate apical maxillary palpomere, shape of slightly convex head and slightly transverse antennomere 7, presence of two small depressions between bases of antennae, short grooves (dorsal tentorial pits) in front of the ocelli, and shape of the moderately convex pronotum with slightly sinuate lateral margins in front of obtuse posterior angles, the new species belong to the
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Phyllodrepa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodrepa" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="genus">Phyllodrepa</taxonomicName>
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complex, specifically to the genus
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Phyllodrepa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodrepa" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="genus">Phyllodrepa</taxonomicName>
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.
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Phyllodrepa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodrepa" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="genus">Phyllodrepa</taxonomicName>
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includes about 30 species distributed in Palaearctic, Nearctic, and Neotropical regions (
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<bibRefCitation pageId="0" pageNumber="35">Newton et al. 2000</bibRefCitation>
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;
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<bibRefCitation author="Herman, LH" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="0" pageNumber="35" pagination="1 - 650" title="Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millenium. I. Introduction, history, biogeographical sketches, and Omaliine group." url="http://hdl.handle.net/2246/5826" volume="265" year="2001">Herman 2001</bibRefCitation>
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;
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<bibRefCitation pageId="0" pageNumber="35">
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<normalizedToken originalValue="Schülke">Schuelke</normalizedToken>
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and Smetana 2015
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</bibRefCitation>
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). The genus requires a worldwide revision and apparently includes some taxa that belong to other related genera (
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<bibRefCitation author="Shavrin, AV" journalOrPublisher="Zootaxa" pageId="0" pageNumber="35" pagination="1 - 94" title="A review of the genus Pycnoglypta Thomson, 1858 (Staphylinidae, Omaliinae, Omaliini) with notes on related taxa." url="https://doi.org/10.11646/zootaxa.4077.1.1" volume="4077" year="2016">Shavrin 2016</bibRefCitation>
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;
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<bibRefCitation author="Zanetti, A" journalOrPublisher="Arthropod Systematics & Phylogeny" pageId="0" pageNumber="35" pagination="53 - 64" title="Two new fossil species species of Omaliinae from Baltic amber (Coleoptera: Staphylinidae) and their significance for understanding the Eocene-Oligocene climate." url="http://www.senckenberg.de/files/content/forschung/publikationen/arthropodsystematics/asp_74_1/03_asp_74_1_zanetti_53-64.pdf" volume="74" year="2016">Zanetti et. al. 2016</bibRefCitation>
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).
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Phyllodrepa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodrepa daedali" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="daedali">Phyllodrepa daedali</taxonomicName>
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sp. nov. and
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<taxonomicName lsidName="Ph. icari" pageId="0" pageNumber="35" rank="species" species="icari">Ph. icari</taxonomicName>
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sp. nov., described below, are species with a very small and pale body that reminds of some Palaearctic species of the genus
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Dropephylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dropephylla" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="genus">Dropephylla</taxonomicName>
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Mulsant & Rey, 1880, that for a long time had been regarded as a subgenus of
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Phyllodrepa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodrepa" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="genus">Phyllodrepa</taxonomicName>
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. Nevertheless, species of
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Dropephylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dropephylla" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="genus">Dropephylla</taxonomicName>
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differ by the absence of grooves in front of ocelli and microsculpture on the elytra, wider apical maxillary palpomere, oval antennomere 4, by the presence of moderately defined short and rounded temples, shorter apical tarsomere, and other morphological characters that were considered in the revision of the Palaearctic fauna of the genus by
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<bibRefCitation author="Jaszay, T" journalOrPublisher="Entomological Problems" pageId="0" pageNumber="35" pagination="31 - 62" title="A revision of the Palaearctic species of the genus Dropephylla (Coleoptera: Staphylinidae: Omaliinae)." volume="36" year="2006">
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<normalizedToken originalValue="Jászay">Jaszay</normalizedToken>
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and
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<normalizedToken originalValue="Hlaváč">Hlavac</normalizedToken>
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(2006)
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</bibRefCitation>
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. Although faintly crenulate lateral margins of the pronotum (Fig. 57) are similar to those in
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Dialycera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dialycera" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="genus">Dialycera</taxonomicName>
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Ganglbauer and
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Hapalaraea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hapalaraea" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="genus">Hapalaraea</taxonomicName>
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Thomson (
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<bibRefCitation pageId="0" pageNumber="35">Zanetti 1987</bibRefCitation>
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,
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<bibRefCitation pageId="0" pageNumber="35">2012b</bibRefCitation>
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;
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<bibRefCitation author="Zanetti, A" journalOrPublisher="Arthropod Systematics & Phylogeny" pageId="0" pageNumber="35" pagination="53 - 64" title="Two new fossil species species of Omaliinae from Baltic amber (Coleoptera: Staphylinidae) and their significance for understanding the Eocene-Oligocene climate." url="http://www.senckenberg.de/files/content/forschung/publikationen/arthropodsystematics/asp_74_1/03_asp_74_1_zanetti_53-64.pdf" volume="74" year="2016">Zanetti et al. 2016</bibRefCitation>
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), they are also known to the first author in European
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Phyllodrepa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodrepa puberula" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="puberula">Phyllodrepa puberula</taxonomicName>
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Bernhauer, 1903 and some little-known species distributed in the eastern Palaearctic Region. Despite this, the new species can not be reliably associated with any extant species of the genus due to its unique morphological characters and the fact that most species differ only by the external structure of the aedeagus. Both new species of
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Phyllodrepa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodrepa" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="genus">Phyllodrepa</taxonomicName>
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described herein differ from the more ancient Transbaikal †
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Eophyllodrepa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eophyllodrepa" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="genus">Eophyllodrepa</taxonomicName>
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Ryvkin from the Middle-Upper Jurassic of Novospasskoe (
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<bibRefCitation author="Ryvkin, AB" editor="Rasnitsyn, AP" journalOrPublisher="Trudy Paleontologischeskogo Instituta, Akademia Nauk SSSR" pageId="0" pageNumber="35" pagination="88 - 91" title="Zhuki semeistva Staphylinidae iz Iury Zabaikal'ia [Beetles of the family Staphylinidae from the Jurassic of Transbaikal]." volume="211" volumeTitle="Jurskiye nasekomyie Sibiri i Mongolii [Jurassic insects of Siberia and Mongolia]." year="1985">Ryvkin 1985</bibRefCitation>
|
||
) and †
|
||
<taxonomicName genus="Daidromus" lsidName="Daidromus" pageId="0" pageNumber="35" rank="genus">Daidromus</taxonomicName>
|
||
Ryvkin from the Upper Jurassic of Daya (
|
||
<bibRefCitation author="Ryvkin, AB" journalOrPublisher=") Pozdne-Mesozoiskie nasekomye vostochnogo Zabaikal'ia [Upper Mesozoic insects of eastern Transbaikalia], Trudy Paleontologischskogo Instituta, Akademia Nauk SSSR" pageId="0" pageNumber="35" pagination="1 - 222" title="Semeistvo Staphylinidae Latreille, 1802 [Family Staphylinidae Latreille, 1802] [pp. 52 - 66]. In: Rasnitsyn AP (Ed." volume="239" year="1990">Ryvkin 1990</bibRefCitation>
|
||
), by the same morphological characters as in
|
||
<taxonomicName lsidName="Ph. electrica" pageId="0" pageNumber="35" rank="species" species="electrica">Ph. electrica</taxonomicName>
|
||
(see
|
||
<bibRefCitation author="Zanetti, A" journalOrPublisher="Arthropod Systematics & Phylogeny" pageId="0" pageNumber="35" pagination="53 - 64" title="Two new fossil species species of Omaliinae from Baltic amber (Coleoptera: Staphylinidae) and their significance for understanding the Eocene-Oligocene climate." url="http://www.senckenberg.de/files/content/forschung/publikationen/arthropodsystematics/asp_74_1/03_asp_74_1_zanetti_53-64.pdf" volume="74" year="2016">Zanetti et al. 2016</bibRefCitation>
|
||
). Based on the small body, shape of head, general shape of apical antennomeres, and pronotum with finely crenulate lateral margins,
|
||
<taxonomicName lsidName="Ph. daedalum" pageId="0" pageNumber="35" rank="species" species="daedalum">Ph. daedalum</taxonomicName>
|
||
sp. nov. is similar to
|
||
<taxonomicName lsidName="Ph. antiqua" pageId="0" pageNumber="35" rank="species" species="antiqua">Ph. antiqua</taxonomicName>
|
||
Zanetti, Perreau & Solodovnikov, 2016, which was recently described from Baltic amber (
|
||
<bibRefCitation author="Zanetti, A" journalOrPublisher="Arthropod Systematics & Phylogeny" pageId="0" pageNumber="35" pagination="53 - 64" title="Two new fossil species species of Omaliinae from Baltic amber (Coleoptera: Staphylinidae) and their significance for understanding the Eocene-Oligocene climate." url="http://www.senckenberg.de/files/content/forschung/publikationen/arthropodsystematics/asp_74_1/03_asp_74_1_zanetti_53-64.pdf" volume="74" year="2016">Zanetti et. al 2016</bibRefCitation>
|
||
). It is also similar to
|
||
<taxonomicName lsidName="Ph. icari" pageId="0" pageNumber="35" rank="species" species="icari">Ph. icari</taxonomicName>
|
||
sp. nov. (see below), based on the crenulate lateral pronotal margins, pale body, and large and deep punctation of the elytra. It differs from
|
||
<taxonomicName lsidName="Ph. antiqua" pageId="0" pageNumber="35" rank="species" species="antiqua">Ph. antiqua</taxonomicName>
|
||
by the smaller, paler and slightly more convex body (Figs 54, 56), coarser and deeper punctation of pronotum (Figs 54, 57) and elytra (Fig. 54), and elongate antennomeres 2-5 (Fig. 58), and from
|
||
<taxonomicName lsidName="Ph. icari" pageId="0" pageNumber="35" rank="species" species="icari">Ph. icari</taxonomicName>
|
||
sp. nov. by the darker abdomen, wider apical maxillary palpomere (Fig. 57), shape of anterior angles of the pronotum not protruded apicad (Fig. 57), denser punctation of the pronotum, less transverse head and pronotum (Fig. 57), and longer antennomeres 4-5 and 11 (Fig. 58). From both these species it differs by longer elytra, and from
|
||
<taxonomicName lsidName="Ph. antiqua" pageId="0" pageNumber="35" rank="species" species="antiqua">Ph. antiqua</taxonomicName>
|
||
by the presence of modified setae on tarsomeres 1-4 of front and middle legs of the male.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="35">
|
||
A remarkable morphological feature of
|
||
<taxonomicName lsidName="Ph. daedali" pageId="0" pageNumber="35" rank="species" species="daedali">Ph. daedali</taxonomicName>
|
||
sp. nov. is the presence of modified rows of elongate setae (Figs 55, 56, 61) with leaf-shaped apical parts on ventral surface of pro- and mesotarsomeres 1-4, described earlier as disco-setae (
|
||
<bibRefCitation author="Stork, NE" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="35" pagination="173 - 306" title="A scanning electron microscope study of tarsal adhesive setae in the Coleoptera." url="https://doi.org/10.1111/j.1096-3642.1980.tb01121.x" volume="68" year="1980">Stork 1980</bibRefCitation>
|
||
) or clavate adhesive setae (
|
||
<bibRefCitation author="Smetana, A" journalOrPublisher="Coleopterists Bulletin" pageId="0" pageNumber="35" pagination="367 - 380" title="Prosopaspis, a new omaliine genus with two new species from the Himalayas (Coleoptera: Staphylinidae)." volume="40" year="1987">Smetana 1987</bibRefCitation>
|
||
). Similar structures were observed in species of the Oriental genera
|
||
<taxonomicName class="Insecta" family="Staphylinidae" genus="Xanthonomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xanthonomus" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="genus">Xanthonomus</taxonomicName>
|
||
Bernhauer by
|
||
<bibRefCitation author="Steel, WO" journalOrPublisher="Entomologist's Monthly Magazine" pageId="0" pageNumber="35" pagination="275 - 278" title="Notes on the Omaliinae (Col., Staphylinidae). 7. The genus Xanthonomus Bernhauer." volume="91" year="1955">Steel (1955</bibRefCitation>
|
||
: fig. 6),
|
||
<taxonomicName class="Insecta" family="Staphylinidae" genus="Prosopaspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosopaspis" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="genus">Prosopaspis</taxonomicName>
|
||
Smetana (
|
||
<bibRefCitation author="Smetana, A" journalOrPublisher="Coleopterists Bulletin" pageId="0" pageNumber="35" pagination="367 - 380" title="Prosopaspis, a new omaliine genus with two new species from the Himalayas (Coleoptera: Staphylinidae)." volume="40" year="1987">Smetana 1987</bibRefCitation>
|
||
: fig. 22),
|
||
<taxonomicName class="Insecta" family="Staphylinidae" genus="Duocalcar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Duocalcar" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="genus">Duocalcar</taxonomicName>
|
||
Peris & Thayer, 2014 (at least protarsi), and
|
||
<taxonomicName class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraphloeostiba" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="genus">Paraphloeostiba</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Steel, WO" journalOrPublisher="Transactions of the Royal Entomological Society of London" pageId="0" pageNumber="35" pagination="141 - 172" title="Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson." url="https://doi.org/10.1111/j.1365-2311.1960.tb00497.x" volume="112" year="1960 a">Steel 1960a</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |