239 lines
32 KiB
XML
239 lines
32 KiB
XML
<document id="BE59CBC3F7E652C28F6D72D380F20A76" ID-DOI="10.5281/zenodo.5323171" ID-ISSN="0374-1036" ID-Zenodo-Dep="5323171" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630138057912" checkinUser="felipe" docAuthor="Hromádka, Lubomír" docDate="2009" docId="03FA87F7FFC7FFC1FE64BD3EFE5AAAD1" docLanguage="en" docName="ActaEntMusNatPra.2009.49.2.671-682.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 49 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Philonthus phacochoerus Hromádka 2009, sp. nov." docType="treatment" docVersion="4" lastPageNumber="680" masterDocId="FFC3FF8FFFC1FFC8FFA0B804FF9FAE3C" masterDocTitle="Five new Philonthus species (Coleoptera: Staphylinidae: Philonthina) from the Afrotropical region" masterLastPageNumber="682" masterPageNumber="671" pageNumber="677" updateTime="1699163750759" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
|
||
<mods:mods id="BFF5012A38E181A49BBCC794BD36DCB8" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="0A06BA74D2B052DFAB23B22A5285AFF1">
|
||
<mods:title id="948FD1611D017A2B7E825D47EEA04D0C">Five new Philonthus species (Coleoptera: Staphylinidae: Philonthina) from the Afrotropical region</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="31A7F0A89FC5EB440C46193F27DB09B9" type="personal">
|
||
<mods:role id="C94CB2B05181330C223F4CD2BA134FEF">
|
||
<mods:roleTerm id="1493774AC26C33993C6D4B78FC99FEB4">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="E7B5236A51679C6DDDD5413134AC0F93">Hromádka, Lubomír</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="CFACAA0A5F199E6F81C5943573753658">text</mods:typeOfResource>
|
||
<mods:relatedItem id="E55831AAF2B50947ED0F1516E1AD82FE" type="host">
|
||
<mods:titleInfo id="3D72E6246B3A3642402B161BB65567E7">
|
||
<mods:title id="49BA0AB9A72FE2DFAC450AF89931E8CD">Acta Entomologica Musei Nationalis Pragae</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="D521ACA1843063A2169DA222DCE32101">
|
||
<mods:date id="01E86A72B6CE65DD46D46160CFDEC1E0">2009</mods:date>
|
||
<mods:detail id="D5E92125C289CD7C75F8AF655E94A4A5" type="pubDate">
|
||
<mods:number id="6E92F3AD48F61A0D356DDC8064850CEA">2009-12-15</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="EB370E38FE5D248FAE122CBF5525C390" type="volume">
|
||
<mods:number id="79C5BAB3514CF5C129331E0C58B6C3E0">49</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="B711D81B39F2EAFCAE27251F9489CF05" type="issue">
|
||
<mods:number id="C12D8AAEB5505DD340B3E453194ADC56">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="EF8E071B27818900F5926FABDD4906F3" unit="page">
|
||
<mods:start id="21F3AF233FA67356A3AB89A53BE04BB6">671</mods:start>
|
||
<mods:end id="FC52526CFF423FCF71E5E4A0AE632483">682</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="332F4A68360F5FDCE2A55CF2D64FD061">journal article</mods:classification>
|
||
<mods:identifier id="413254881A98CB94474ECFF97B8EE208" type="DOI">10.5281/zenodo.5323171</mods:identifier>
|
||
<mods:identifier id="7BE6822DA71A916FB76CACB7D50D80B1" type="ISSN">0374-1036</mods:identifier>
|
||
<mods:identifier id="82DB520E0983C599C0509ADA4CFEA159" type="Zenodo-Dep">5323171</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03FA87F7FFC7FFC1FE64BD3EFE5AAAD1" ID-DOI="http://doi.org/10.5281/zenodo.5343569" ID-Zenodo-Dep="5343569" LSID="urn:lsid:plazi:treatment:03FA87F7FFC7FFC1FE64BD3EFE5AAAD1" httpUri="http://treatment.plazi.org/id/03FA87F7FFC7FFC1FE64BD3EFE5AAAD1" lastPageId="9" lastPageNumber="680" pageId="6" pageNumber="677">
|
||
<subSubSection id="C349656AFFC7FFCEFE64BD3EFCA5AB68" box="[452,826,1338,1364]" pageId="6" pageNumber="677" type="nomenclature">
|
||
<paragraph id="8BEC36E1FFC7FFCEFE64BD3EFCA5AB68" blockId="6.[452,826,1338,1364]" box="[452,826,1338,1364]" pageId="6" pageNumber="677">
|
||
<heading id="D0A4818DFFC7FFCEFE64BD3EFCA5AB68" bold="true" box="[452,826,1338,1364]" centered="true" fontSize="8" level="2" pageId="6" pageNumber="677" reason="3">
|
||
<emphasis id="B927EAF3FFC7FFCEFE64BD3EFCA5AB68" bold="true" box="[452,826,1338,1364]" pageId="6" pageNumber="677">
|
||
<taxonomicName id="4C534D62FFC7FFCEFE64BD3EFD41AB6F" ID-CoL="4GBHY" authority="Hromádka, 2009" authorityName="Hromádka" authorityYear="2009" box="[452,734,1338,1363]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="677" phylum="Arthropoda" rank="species" species="phacochoerus" status="sp. nov.">
|
||
<emphasis id="B927EAF3FFC7FFCEFE64BD3EFD41AB6F" bold="true" box="[452,734,1338,1363]" italics="true" pageId="6" pageNumber="677">Philonthus phacochoerus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2145788FFC7FFCEFD45BD38FCA5AB68" box="[741,826,1340,1364]" pageId="6" pageNumber="677" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C349656AFFC7FFCEFDE5BD5EFD27AB51" box="[581,696,1370,1389]" pageId="6" pageNumber="677" type="description">
|
||
<paragraph id="8BEC36E1FFC7FFCEFDE5BD5EFD27AB51" blockId="6.[581,696,1370,1389]" box="[581,696,1370,1389]" pageId="6" pageNumber="677">
|
||
(
|
||
<figureCitation id="13682A64FFC7FFCEFDECBD5EFD2DAB51" box="[588,690,1370,1389]" captionStart="Figs 20–31. 20–26" captionStartId="8.[152,189,1461,1480]" captionTargetBox="[245,1049,218,1428]" captionTargetId="figure-10@8.[196,1082,195,1445]" captionTargetPageId="8" captionText="Figs 20–31. 20–26 – Philonthus phacochoerus sp. nov. 20 – aedeagus, ventral view; 21 – aedeagus, lateral view; 22 – paramere; 23 – apical portion of male sternite VIII; 24 – male sternite IX; 25 – female tergite X; 26 – gonocoxite of female genital segment. 27–31 – Philonthus tragelaphus sp. nov. 27 – aedeagus, ventral view; 28 – aedeagus, lateral view; 29 – apex of paramere with sensory peg setae; 30 – apical portion of male sternite VIII; 31 – male sternite IX." figureDoi="http://doi.org/10.5281/zenodo.5323179" httpUri="https://zenodo.org/record/5323179/files/figure.png" pageId="6" pageNumber="677">Figs. 20–26</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C349656AFFC7FFCEFF37BD80FEF6A835" pageId="6" pageNumber="677" type="materials_examined">
|
||
<paragraph id="8BEC36E1FFC7FFCEFF37BD80FDBCABAB" blockId="6.[151,1127,1411,1545]" box="[151,547,1411,1431]" pageId="6" pageNumber="677">
|
||
<emphasis id="B927EAF3FFC7FFCEFF37BD80FE8FABAB" bold="true" box="[151,272,1411,1431]" pageId="6" pageNumber="677">
|
||
<typeStatus id="54E88843FFC7FFCEFF37BD80FF5BABAB" box="[151,196,1412,1431]" pageId="6" pageNumber="677">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="F3447671FFC7FFCEFEB5BD80FECAABAB" box="[277,341,1412,1431]" name="Liberia" pageId="6" pageNumber="677">Liberia</collectingCountry>
|
||
, Mt.
|
||
<collectingRegion id="4997F803FFC7FFCEFE21BD80FE22ABAB" box="[385,445,1412,1431]" country="Liberia" name="Nimba" pageId="6" pageNumber="677">Nimba</collectingRegion>
|
||
, Grassfield.
|
||
</paragraph>
|
||
<paragraph id="8BEC36E1FFC7FFCEFF37BDA2FEF6A835" blockId="6.[151,1127,1411,1545]" pageId="6" pageNumber="677">
|
||
<emphasis id="B927EAF3FFC7FFCEFF37BDA2FE81AB85" bold="true" box="[151,286,1445,1465]" pageId="6" pageNumber="677">Type material.</emphasis>
|
||
<materialsCitation id="3B3B3CBCFFC7FFCEFE85BDA2FE4DABE8" collectingDate="1979-09-16" collectingDateMax="1979-09-25" collectingDateMin="1979-09-16" collectorName="I. Hanski" country="Liberia" elevation="500" location="Grassfield" pageId="6" pageNumber="677" specimenCount="1" stateProvince="Nimba" typeStatus="holotype">
|
||
<typeStatus id="54E88843FFC7FFCEFE85BDA2FEE6AB84" box="[293,377,1446,1465]" pageId="6" pageNumber="677" type="holotype">HOLOTYPE</typeStatus>
|
||
: J, ‛
|
||
<collectingCountry id="F3447671FFC7FFCEFE0ABDA2FD9FAB85" box="[426,512,1446,1465]" name="Liberia" pageId="6" pageNumber="677">LIBERIA</collectingCountry>
|
||
, Mt.
|
||
<collectingRegion id="4997F803FFC7FFCEFD91BDA2FDF2AB85" box="[561,621,1446,1465]" country="Liberia" name="Nimba" pageId="6" pageNumber="677">Nimba</collectingRegion>
|
||
,
|
||
<location id="8E8C603AFFC7FFCEFDD8BDA2FD4EAB85" LSID="urn:lsid:plazi:treatment:03FA87F7FFC7FFC1FE64BD3EFE5AAAD1:8E8C603AFFC7FFCEFDD8BDA2FD4EAB85" box="[632,721,1446,1465]" country="Liberia" name="Grassfield" pageId="6" pageNumber="677" stateProvince="Nimba">Grassfield</location>
|
||
,
|
||
<date id="FFED1021FFC7FFCEFD7CBDA2FCC1AB85" box="[732,862,1446,1465]" pageId="6" pageNumber="677" value="1979-09-16" valueMax="1979-09-25" valueMin="1979-09-16">
|
||
<collectingDate id="EFA9E9C9FFC7FFCEFD7CBDA2FCC1AB85" box="[732,862,1446,1465]" pageId="6" pageNumber="677" value="1979-09-16" valueMax="1979-09-25" valueMin="1979-09-16">16.–25.ix.1979</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26A65337FFC7FFCEFCC9BDA2FC23AB85" box="[873,956,1446,1465]" pageId="6" pageNumber="677">I. Hanski</collectorName>
|
||
, B. M. 1980-85, in human faeces, lowland forest
|
||
<quantity id="4CAB9B04FFC7FFCEFE31BDC4FE5DABEF" box="[401,450,1472,1491]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="677" unit="m" value="500.0">
|
||
<elevation id="007ED1D2FFC7FFCEFE31BDC4FE5DABEF" box="[401,450,1472,1491]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="677" unit="m" value="500.0">500m</elevation>
|
||
</quantity>
|
||
//
|
||
</materialsCitation>
|
||
<materialsCitation id="3B3B3CBCFFC7FFCEFE76BDC4FEF9ABD2" collectionCode="FMNH" country="Liberia" determinerName="Hromadka" pageId="6" pageNumber="677" specimenCount="1" stateProvince="Nimba" typeStatus="holotype">
|
||
<typeStatus id="54E88843FFC7FFCEFE76BDC4FDB7ABEF" box="[470,552,1472,1491]" pageId="6" pageNumber="677" type="holotype">Holotype</typeStatus>
|
||
,
|
||
<taxonomicName id="4C534D62FFC7FFCEFD8FBDC6FC98ABE8" authority="Hromádka, 2009" authorityName="Hromádka" authorityYear="2009" box="[559,775,1474,1492]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="677" phylum="Arthropoda" rank="species" species="phacochoerus" status="sp. nov.">
|
||
<emphasis id="B927EAF3FFC7FFCEFD8FBDC6FC98ABE8" box="[559,775,1474,1492]" italics="true" pageId="6" pageNumber="677">Philonthus phacochoerus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2145788FFC7FFCEFCABBDC5FCD2ABE8" box="[779,845,1473,1492]" pageId="6" pageNumber="677" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<determinerName id="7E30DA25FFC7FFCEFCEFBDC4FC36ABEF" box="[847,937,1472,1491]" pageId="6" pageNumber="677">Hromádka</determinerName>
|
||
det., 2008 [red oblong printed label]’ (
|
||
<collectionCode id="ED42AE24FFC7FFCEFEBDBDDFFEFFABD2" box="[285,352,1499,1518]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="6" pageNumber="677" type="Museum">FMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<typeStatus id="54E88843FFC7FFCEFECFBDD8FE58ABD1" box="[367,455,1500,1519]" pageId="6" pageNumber="677" type="paratype">PARATYPES</typeStatus>
|
||
:
|
||
<specimenCount id="9D55FD68FFC7FFCEFE73BDDFFD82ABD3" box="[467,541,1499,1519]" pageId="6" pageNumber="677" type="generic">10 spec.</specimenCount>
|
||
, same label data as in
|
||
<typeStatus id="54E88843FFC7FFCEFD43BDDFFCB1ABD2" box="[739,814,1499,1518]" pageId="6" pageNumber="677" type="holotype">holotype</typeStatus>
|
||
[all
|
||
<typeStatus id="54E88843FFC7FFCEFCF8BDD8FC35ABD3" box="[856,938,1500,1519]" pageId="6" pageNumber="677" type="paratype">paratypes</typeStatus>
|
||
with red printed type labels] (
|
||
<collectionCode id="ED42AE24FFC7FFCEFF7BBDF2FE83A835" box="[219,284,1526,1545]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="6" pageNumber="677" type="Museum">FMNH</collectionCode>
|
||
,
|
||
<collectionCode id="ED42AE24FFC7FFCEFE85BDF2FEC0A835" box="[293,351,1526,1545]" pageId="6" pageNumber="677">LHPC</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C349656AFFC6FFCFFFD1B8DFFCDBA80F" pageId="7" pageNumber="678" type="description">
|
||
<paragraph id="8BEC36E1FFC6FFCFFFD1B8DFFBAFAECF" blockId="7.[113,1089,219,1588]" box="[113,1072,219,244]" pageId="7" pageNumber="678">
|
||
<emphasis id="B927EAF3FFC6FFCFFFD1B8DFFF63AECF" bold="true" box="[113,252,219,243]" pageId="7" pageNumber="678">Description.</emphasis>
|
||
Body length
|
||
<quantity id="4CAB9B04FFC6FFCFFE2CB8DFFD90AECF" box="[396,527,219,243]" metricMagnitude="-3" metricUnit="m" metricValue="7.75" metricValueMax="8.9" metricValueMin="6.6" pageId="7" pageNumber="678" unit="mm" value="7.75" valueMax="8.9" valueMin="6.6">6.6–8.9 mm</quantity>
|
||
, length of fore body (to end of elytra)
|
||
<quantity id="4CAB9B04FFC6FFCFFC08B8D8FBB3AECF" box="[936,1068,220,244]" metricMagnitude="-3" metricUnit="m" metricValue="3.8499999999999996" metricValueMax="4.3" metricValueMin="3.4" pageId="7" pageNumber="678" unit="mm" value="3.8499999999999996" valueMax="4.3" valueMin="3.4">3.4–4.3 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEC36E1FFC6FFCFFF30B8FFFEEFAFEF" blockId="7.[113,1089,219,1588]" pageId="7" pageNumber="678">Colouration. Head black, maxillary and labial palpi yellow-brown, terminal maxillary palpomere with paler apex, mandibles dark brown, antennal sockets, labrum, ventral side of antennomere 1 yellow-brown, dorsal side of antennomere 1 and remaining antennomeres black-brown, pronotum black-brown with sides and posterior margins slightly golden iridescent, scutellum brown-black, elytra yellow-brown, suture narrowly darker, posterior margin of all abdominal tergites broadly yellow-brown, paratergites, femora and tarsi yellow-brown, tibiae somewhat darker.</paragraph>
|
||
<paragraph id="8BEC36E1FFC6FFCFFF30B9DFFE8DAC8F" blockId="7.[113,1089,219,1588]" pageId="7" pageNumber="678">Head with square outline with obtusely rounded posterior angles, wider than long (ratio 36: 28). Eyes as long as temples, four punctures situated between eyes, distance between medial interocular punctures almost three times as long as distance between medial and lateral interocular puncture, posterior margin of eyes with two coarse punctures, temporal area almost impunctate, posterior angles with several coarse punctures and with one long black bristle, surface with very fine and dense microsculpture consisting of transverse and oblique waves.</paragraph>
|
||
<paragraph id="8BEC36E1FFC6FFCFFF30BAB8FDF6ACCF" blockId="7.[113,1089,219,1588]" pageId="7" pageNumber="678">Antennae long, reaching posterior sixth of pronotum when reclined, relative lengths of antennomeres: 1–11: 11-6-6-4.5-4-4-4-4-4-4-5.</paragraph>
|
||
<paragraph id="8BEC36E1FFC6FFCFFF30BAF8FEF8ADAF" blockId="7.[113,1089,219,1588]" pageId="7" pageNumber="678">Pronotum longer than wide (ratio 40: 38), very slightly narrowed anteriad, posterior angles conspicuously rounded, each dorsal row with four coarse equidistant punctures, each sublateral row with two fine punctures, puncture 2 situated behind level of puncture 3 of dorsal row. Anterior angles and sides with several bristles of unequal length, microsculpture similar to that on head.</paragraph>
|
||
<paragraph id="8BEC36E1FFC6FFCFFF30BB9FFF44ADCF" blockId="7.[113,1089,219,1588]" pageId="7" pageNumber="678">Scutellum coarsely punctate, punctures as large as eye facets, separated by 1.5–2 puncture diameters in transverse direction, surface with fine, mostly transverse and undulating microsculpture.</paragraph>
|
||
<paragraph id="8BEC36E1FFC6FFCFFF30BBFFFDBFAAAF" blockId="7.[113,1089,219,1588]" pageId="7" pageNumber="678">Elytra wider than long (ratio 50: 45), widened posteriad, punctation fine and sparse, punctures of two sizes, smaller punctures somewhat smaller than eye facets, separated by 2–3 puncture diameters, larger punctures distinctly larger than eye facets, irregularly arranged, mostly separated by three puncture diameters. Sides with several long black bristles, surface without microsculpture; setation yellow.</paragraph>
|
||
<paragraph id="8BEC36E1FFC6FFCFFF30BC98FE09AAE8" blockId="7.[113,1089,219,1588]" pageId="7" pageNumber="678">Legs. Metatibia slightly shorter than metatarsus (ratio 28: 30). Relative lengths of metatarsomeres: 1–5: 9-4-4-4-7.</paragraph>
|
||
<paragraph id="8BEC36E1FFC6FFCFFF30BCDFFCEAAB4F" blockId="7.[113,1089,219,1588]" pageId="7" pageNumber="678">Abdomen wide, slightly narrowed posteriad from fifth visible tergite, first three visible tergites with two basal lines, elevated area between basal lines impunctate. Punctation of visible tergites very fine and dense, punctures smaller than eye facets, separated by distance equal to diameter of puncture or slightly smaller. Lateral margins with several long brown bristles, setation long, yellowish brown, surface without microsculpture.</paragraph>
|
||
<paragraph id="8BEC36E1FFC6FFCFFF30BD7FFD5FABE8" blockId="7.[113,1089,219,1588]" pageId="7" pageNumber="678">
|
||
Male. Protarsomeres 1–3 conspicuously dilated and sub-bilobed, covered with modified pale setae ventrally, protarsomere 4 distinctly narrower than preceding ones. Sternite VIII (
|
||
<figureCitation id="13682A64FFC6FFCFFFD9BDBFFF56ABE8" box="[121,201,1467,1492]" captionStart="Figs 20–31. 20–26" captionStartId="8.[152,189,1461,1480]" captionTargetBox="[245,1049,218,1428]" captionTargetId="figure-10@8.[196,1082,195,1445]" captionTargetPageId="8" captionText="Figs 20–31. 20–26 – Philonthus phacochoerus sp. nov. 20 – aedeagus, ventral view; 21 – aedeagus, lateral view; 22 – paramere; 23 – apical portion of male sternite VIII; 24 – male sternite IX; 25 – female tergite X; 26 – gonocoxite of female genital segment. 27–31 – Philonthus tragelaphus sp. nov. 27 – aedeagus, ventral view; 28 – aedeagus, lateral view; 29 – apex of paramere with sensory peg setae; 30 – apical portion of male sternite VIII; 31 – male sternite IX." figureDoi="http://doi.org/10.5281/zenodo.5323179" httpUri="https://zenodo.org/record/5323179/files/figure.png" pageId="7" pageNumber="678">Fig. 23</figureCitation>
|
||
); sternite IX (
|
||
<figureCitation id="13682A64FFC6FFCFFEFDBDBFFE32ABE8" box="[349,429,1467,1492]" captionStart="Figs 20–31. 20–26" captionStartId="8.[152,189,1461,1480]" captionTargetBox="[245,1049,218,1428]" captionTargetId="figure-10@8.[196,1082,195,1445]" captionTargetPageId="8" captionText="Figs 20–31. 20–26 – Philonthus phacochoerus sp. nov. 20 – aedeagus, ventral view; 21 – aedeagus, lateral view; 22 – paramere; 23 – apical portion of male sternite VIII; 24 – male sternite IX; 25 – female tergite X; 26 – gonocoxite of female genital segment. 27–31 – Philonthus tragelaphus sp. nov. 27 – aedeagus, ventral view; 28 – aedeagus, lateral view; 29 – apex of paramere with sensory peg setae; 30 – apical portion of male sternite VIII; 31 – male sternite IX." figureDoi="http://doi.org/10.5281/zenodo.5323179" httpUri="https://zenodo.org/record/5323179/files/figure.png" pageId="7" pageNumber="678">Fig. 24</figureCitation>
|
||
); aedeagus (
|
||
<figureCitation id="13682A64FFC6FFCFFD90BDBFFD2BABE8" box="[560,692,1467,1492]" captionStart="Figs 20–31. 20–26" captionStartId="8.[152,189,1461,1480]" captionTargetBox="[245,1049,218,1428]" captionTargetId="figure-10@8.[196,1082,195,1445]" captionTargetPageId="8" captionText="Figs 20–31. 20–26 – Philonthus phacochoerus sp. nov. 20 – aedeagus, ventral view; 21 – aedeagus, lateral view; 22 – paramere; 23 – apical portion of male sternite VIII; 24 – male sternite IX; 25 – female tergite X; 26 – gonocoxite of female genital segment. 27–31 – Philonthus tragelaphus sp. nov. 27 – aedeagus, ventral view; 28 – aedeagus, lateral view; 29 – apex of paramere with sensory peg setae; 30 – apical portion of male sternite VIII; 31 – male sternite IX." figureDoi="http://doi.org/10.5281/zenodo.5323179" httpUri="https://zenodo.org/record/5323179/files/figure.png" pageId="7" pageNumber="678">Figs. 20–22</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEC36E1FFC6FFCFFF30BDDFFCDBA80F" blockId="7.[113,1089,219,1588]" pageId="7" pageNumber="678">
|
||
Female. Protarsomeres 1–3 only slightly dilated, not sub-bilobed, covered with only few modified pale setae ventrally, protarsomere 4 only slightly narrower than preceding ones. Tergite X (
|
||
<figureCitation id="13682A64FFC6FFCFFF45BE1FFEABA80F" box="[229,308,1563,1587]" captionStart="Figs 20–31. 20–26" captionStartId="8.[152,189,1461,1480]" captionTargetBox="[245,1049,218,1428]" captionTargetId="figure-10@8.[196,1082,195,1445]" captionTargetPageId="8" captionText="Figs 20–31. 20–26 – Philonthus phacochoerus sp. nov. 20 – aedeagus, ventral view; 21 – aedeagus, lateral view; 22 – paramere; 23 – apical portion of male sternite VIII; 24 – male sternite IX; 25 – female tergite X; 26 – gonocoxite of female genital segment. 27–31 – Philonthus tragelaphus sp. nov. 27 – aedeagus, ventral view; 28 – aedeagus, lateral view; 29 – apex of paramere with sensory peg setae; 30 – apical portion of male sternite VIII; 31 – male sternite IX." figureDoi="http://doi.org/10.5281/zenodo.5323179" httpUri="https://zenodo.org/record/5323179/files/figure.png" pageId="7" pageNumber="678">Fig. 25</figureCitation>
|
||
); gonocoxite of female genital segment (
|
||
<figureCitation id="13682A64FFC6FFCFFD48BE1FFCA7A80F" box="[744,824,1563,1587]" captionStart="Figs 20–31. 20–26" captionStartId="8.[152,189,1461,1480]" captionTargetBox="[245,1049,218,1428]" captionTargetId="figure-10@8.[196,1082,195,1445]" captionTargetPageId="8" captionText="Figs 20–31. 20–26 – Philonthus phacochoerus sp. nov. 20 – aedeagus, ventral view; 21 – aedeagus, lateral view; 22 – paramere; 23 – apical portion of male sternite VIII; 24 – male sternite IX; 25 – female tergite X; 26 – gonocoxite of female genital segment. 27–31 – Philonthus tragelaphus sp. nov. 27 – aedeagus, ventral view; 28 – aedeagus, lateral view; 29 – apex of paramere with sensory peg setae; 30 – apical portion of male sternite VIII; 31 – male sternite IX." figureDoi="http://doi.org/10.5281/zenodo.5323179" httpUri="https://zenodo.org/record/5323179/files/figure.png" pageId="7" pageNumber="678">Fig. 26</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF2C6669FFC9FFC0FF38BDB1FECDA80F" ID-DOI="http://doi.org/10.5281/zenodo.5323179" ID-Zenodo-Dep="5323179" httpUri="https://zenodo.org/record/5323179/files/figure.png" pageId="8" pageNumber="679" startId="8.[152,189,1461,1480]" targetBox="[245,1049,218,1428]" targetPageId="8">
|
||
<paragraph id="8BEC36E1FFC9FFC0FF38BDB1FECDA80F" blockId="8.[152,1128,1461,1587]" pageId="8" pageNumber="679">
|
||
Figs 20–31. 20–26 –
|
||
<taxonomicName id="4C534D62FFC9FFC0FEF0BDB2FDB4ABF4" authority="Hromádka, 2009" authorityName="Hromádka" authorityYear="2009" box="[336,555,1462,1480]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="679" phylum="Arthropoda" rank="species" species="phacochoerus" status="sp. nov.">
|
||
<emphasis id="B927EAF3FFC9FFC0FEF0BDB2FDB4ABF4" box="[336,555,1462,1480]" italics="true" pageId="8" pageNumber="679">Philonthus phacochoerus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2145788FFC9FFC0FD90BDB1FDEBABF4" box="[560,628,1461,1480]" pageId="8" pageNumber="679" rank="species">sp. nov.</taxonomicNameLabel>
|
||
20 – aedeagus, ventral view; 21 – aedeagus, lateral view; 22 – paramere; 23 – apical portion of male sternite VIII; 24 – male sternite IX; 25 – female tergite X; 26 – gonocoxite of female genital segment. 27–31 –
|
||
<taxonomicName id="4C534D62FFC9FFC0FDFFBDEFFCB8ABC1" authority="Hromádka, 2009" authorityName="Hromádka" authorityYear="2009" box="[607,807,1515,1533]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="679" phylum="Arthropoda" rank="species" species="tragelaphus" status="sp. nov.">
|
||
<emphasis id="B927EAF3FFC9FFC0FDFFBDEFFCB8ABC1" box="[607,807,1515,1533]" italics="true" pageId="8" pageNumber="679">Philonthus tragelaphus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2145788FFC9FFC0FC8CBDEFFCF0ABC2" box="[812,879,1515,1534]" pageId="8" pageNumber="679" rank="species">sp. nov.</taxonomicNameLabel>
|
||
27 – aedeagus, ventral view; 28 – aedeagus, lateral view; 29 – apex of paramere with sensory peg setae; 30 – apical portion of male sternite VIII; 31 – male sternite IX.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2C6669FFC8FFC1FFD1BB44FE96AD9F" ID-DOI="http://doi.org/10.5281/zenodo.5323181" ID-Zenodo-Dep="5323181" httpUri="https://zenodo.org/record/5323181/files/figure.png" pageId="9" pageNumber="680" startId="9.[113,152,832,851]" targetBox="[119,1083,238,799]" targetPageId="9">
|
||
<paragraph id="8BEC36E1FFC8FFC1FFD1BB44FE96AD9F" blockId="9.[113,1090,832,931]" pageId="9" pageNumber="680">
|
||
Figs. 32–36. 32–33 –
|
||
<taxonomicName id="4C534D62FFC8FFC1FE93BB45FE62AD6F" authority="Hromádka, 2009" authorityName="Hromádka" authorityYear="2009" box="[307,509,833,851]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="680" phylum="Arthropoda" rank="species" species="tragelaphus" status="sp. nov.">
|
||
<emphasis id="B927EAF3FFC8FFC1FE93BB45FE62AD6F" box="[307,509,833,851]" italics="true" pageId="9" pageNumber="680">Philonthus tragelaphus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2145788FFC8FFC1FDA4BB45FDD6AD68" box="[516,585,833,852]" pageId="9" pageNumber="680" rank="species">sp. nov.</taxonomicNameLabel>
|
||
32 – female tergite X; 33 – gonocoxite of female genital segment. 34 –
|
||
<taxonomicName id="4C534D62FFC8FFC1FF56BB58FD85AD52" authority="Tottenham, 1949" authorityName="Tottenham" authorityYear="1949" box="[246,538,859,878]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="680" phylum="Arthropoda" rank="species" species="smaragdinus">
|
||
<emphasis id="B927EAF3FFC8FFC1FF56BB58FEE2AD52" box="[246,381,860,878]" italics="true" pageId="9" pageNumber="680">P. smaragdinus</emphasis>
|
||
<bibRefCitation id="EFC24B10FFC8FFC1FE25BB5FFD85AD52" author="TOTTENHAM C. E." box="[389,538,859,878]" pageId="9" pageNumber="680" pagination="291 - 362" refId="ref5163" refString="TOTTENHAM C. E. 1949: Studies in the genus Philonthus Stephens (Coleoptera). Transactions of the Royal Entomological Society of London 100: 291 - 362." type="journal article" year="1949">Tottenham, 1949</bibRefCitation>
|
||
</taxonomicName>
|
||
, aedeagus, ventral view (Democratic Republic of the Congo). 35 –
|
||
<taxonomicName id="4C534D62FFC8FFC1FF3DBB73FEE9ADB5" authority="Boheman, 1848" authorityName="Boheman" authorityYear="1848" box="[157,374,886,905]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="680" phylum="Arthropoda" rank="species" species="morio">
|
||
<emphasis id="B927EAF3FFC8FFC1FF3DBB73FF79ADB5" box="[157,230,887,905]" italics="true" pageId="9" pageNumber="680">P. morio</emphasis>
|
||
Boheman, 1848
|
||
</taxonomicName>
|
||
, aedeagus, ventral view (Malawi); 36 –
|
||
<taxonomicName id="4C534D62FFC8FFC1FD70BB73FC3CADB5" authority="Tottenham, 1955" authorityName="Tottenham" authorityYear="1955" box="[720,931,886,905]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="680" phylum="Arthropoda" rank="species" species="lasti">
|
||
<emphasis id="B927EAF3FFC8FFC1FD70BB73FC95ADB5" box="[720,778,887,905]" italics="true" pageId="9" pageNumber="680">P. lasti</emphasis>
|
||
Tottenham, 1955
|
||
</taxonomicName>
|
||
, aedeagus, ventral view (Botswana).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C349656AFFC8FFC1FFD1BBF1FD6BAA71" pageId="9" pageNumber="680" type="diagnosis">
|
||
<paragraph id="8BEC36E1FFC8FFC1FFD1BBF1FD6BAA71" blockId="9.[113,1089,1013,1261]" pageId="9" pageNumber="680">
|
||
<emphasis id="B927EAF3FFC8FFC1FFD1BBF1FEF1AA31" bold="true" box="[113,366,1013,1037]" pageId="9" pageNumber="680">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C534D62FFC8FFC1FED7BBF2FD15AA31" authority="Hromádka, 2009" authorityName="Hromádka" authorityYear="2009" box="[375,650,1014,1037]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="680" phylum="Arthropoda" rank="species" species="phacochaerus" status="sp. nov.">
|
||
<emphasis id="B927EAF3FFC8FFC1FED7BBF2FD15AA31" box="[375,650,1014,1037]" italics="true" pageId="9" pageNumber="680">Philonthus phacochaerus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2145788FFC8FFC1FD33BBF2FD76AA31" box="[659,745,1014,1037]" pageId="9" pageNumber="680" rank="species">sp. nov.</taxonomicNameLabel>
|
||
may be distinguished from the similar
|
||
<taxonomicName id="4C534D62FFC8FFC1FF61BC12FE20AA11" authority="Tottenham, 1955" authorityName="Tottenham" authorityYear="1955" box="[193,447,1045,1069]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="680" phylum="Arthropoda" rank="species" species="lasti">
|
||
<emphasis id="B927EAF3FFC8FFC1FF61BC12FE97AA11" box="[193,264,1046,1069]" italics="true" pageId="9" pageNumber="680">P. lasti</emphasis>
|
||
Tottenham, 1955
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13682A64FFC8FFC1FE6DBC11FD85AA11" box="[461,538,1045,1069]" captionStart="Figs" captionStartId="9.[113,152,832,851]" captionTargetBox="[119,1083,238,799]" captionTargetId="figure-183@9.[114,1089,219,810]" captionTargetPageId="9" captionText="Figs. 32–36. 32–33 – Philonthus tragelaphus sp. nov. 32 – female tergite X; 33 – gonocoxite of female genital segment. 34 – P. smaragdinus Tottenham, 1949, aedeagus, ventral view (Democratic Republic of the Congo). 35 – P. morio Boheman, 1848, aedeagus, ventral view (Malawi); 36 – P. lasti Tottenham, 1955, aedeagus, ventral view (Botswana)." figureDoi="http://doi.org/10.5281/zenodo.5323181" httpUri="https://zenodo.org/record/5323181/files/figure.png" pageId="9" pageNumber="680">Fig. 36</figureCitation>
|
||
) by a quadrate head, shorter eyes, uniformly red and sparsely punctate elytra and different shape of the aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C349656AFFC8FFC1FFD1BC51FE85AAF1" pageId="9" pageNumber="680" type="etymology">
|
||
<paragraph id="8BEC36E1FFC8FFC1FFD1BC51FD7BAAB1" blockId="9.[113,1089,1013,1261]" pageId="9" pageNumber="680">
|
||
<emphasis id="B927EAF3FFC8FFC1FFD1BC51FF6DAA51" bold="true" box="[113,242,1109,1133]" pageId="9" pageNumber="680">Etymology.</emphasis>
|
||
The name of this species, a noun in apposition, is the Latin generic name of the African warthog
|
||
<emphasis id="B927EAF3FFC8FFC1FE87BC72FDDDAAB1" box="[295,578,1142,1165]" italics="true" pageId="9" pageNumber="680">
|
||
<taxonomicName id="4C534D62FFC8FFC1FE87BC72FE5FAAB1" authorityName="Hromádka" authorityYear="2009" box="[295,448,1142,1165]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="680" phylum="Arthropoda" rank="species" species="phacochoerus">Phacochoerus</taxonomicName>
|
||
aethiopicus
|
||
</emphasis>
|
||
(Pallas, 1767).
|
||
</paragraph>
|
||
<paragraph id="8BEC36E1FFC8FFC1FFD1BC91FE85AAF1" blockId="9.[113,1089,1013,1261]" pageId="9" pageNumber="680">
|
||
<emphasis id="B927EAF3FFC8FFC1FFD1BC91FF70AA91" bold="true" box="[113,239,1173,1197]" pageId="9" pageNumber="680">Bionomics.</emphasis>
|
||
All specimens of
|
||
<taxonomicName id="4C534D62FFC8FFC1FE17BC92FDF6AA91" authority="Hromádka, 2009" authorityName="Hromádka" authorityYear="2009" box="[439,617,1174,1197]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="680" phylum="Arthropoda" rank="species" species="phacochoerus" status="sp. nov.">
|
||
<emphasis id="B927EAF3FFC8FFC1FE17BC92FDF6AA91" box="[439,617,1174,1197]" italics="true" pageId="9" pageNumber="680">P. phacochoerus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2145788FFC8FFC1FDD1BC92FD5AAA91" box="[625,709,1174,1197]" pageId="9" pageNumber="680" rank="species">sp. nov.</taxonomicNameLabel>
|
||
were collected in human faeces in mid September.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C349656AFFC8FFC1FFD1BCD1FE5AAAD1" box="[113,453,1237,1261]" pageId="9" pageNumber="680" type="distribution">
|
||
<paragraph id="8BEC36E1FFC8FFC1FFD1BCD1FE5AAAD1" blockId="9.[113,1089,1013,1261]" box="[113,453,1237,1261]" pageId="9" pageNumber="680">
|
||
<emphasis id="B927EAF3FFC8FFC1FFD1BCD1FE9BAAD1" bold="true" box="[113,260,1237,1261]" pageId="9" pageNumber="680">Distribution.</emphasis>
|
||
Northern
|
||
<collectingCountry id="F3447671FFC8FFC1FED1BCD1FE5DAAD1" box="[369,450,1237,1261]" name="Liberia" pageId="9" pageNumber="680">Liberia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |