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<document id="93C784DC32F966BD2D2B04247D151985" ID-DOI="10.11646/phytotaxa.489.1.6" ID-ISSN="1179-3163" ID-Zenodo-Dep="5757631" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630385297280" checkinUser="marcus" docAuthor="Johnson, David M., Chaowasku, Tanawat, Murray, Nancy A. &amp; Chalermglin, Piya" docDate="2021" docId="03DB7B3F037BF010C7EBFF74FBB601BE" docLanguage="en" docName="phytotaxa.489.1.6.pdf" docOrigin="Phytotaxa 489 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.489.1.6" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Trivalvaria ochroleuca Chaowasku &amp; D. M. Johnson 2021, sp. nov." docType="treatment" docVersion="4" lastPageNumber="80" masterDocId="FFE20347037AF011C763FF80FFCA040C" masterDocTitle="Three new species of Trivalvaria (Annonaceae) from Thailand, with a key to the Thai species" masterLastPageNumber="86" masterPageNumber="79" pageNumber="80" updateTime="1699255033702" updateUser="plazi" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="1A1F4F4ADE5F5EE0C51B92C0664EFCDD" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="96DF758AB3D875604D08E79AF13FB75B">Three new species of Trivalvaria (Annonaceae) from Thailand, with a key to the Thai species</mods:title>
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<mods:namePart id="C41B18869CB7269425394E73D2CB72DB">Johnson, David M.</mods:namePart>
<mods:nameIdentifier id="405EBD35C3C373E7407084F43C9269CD" type="ORCID">0000-0003-2896-7419</mods:nameIdentifier>
<mods:affiliation id="C3F74086AC31B5F3F6639232C6109084">dmjohnso @ owu. edu; https: // orcid. org / 0000 - 0003 - 2896 - 7419</mods:affiliation>
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<mods:namePart id="C0A209EA0D8BF791AF35C628A229303C">Chaowasku, Tanawat</mods:namePart>
<mods:nameIdentifier id="CD028371F3E5728A623E092B85DB395A" type="ORCID">0000-0002-1602-8468</mods:nameIdentifier>
<mods:affiliation id="714AB46627F75819B96A8CB7D1D49959">Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai 50200, Thailand &amp; tanawat. chaowasku @ cmu. ac. th; https: // orcid. org / 0000 - 0002 - 1602 - 8468</mods:affiliation>
<mods:nameIdentifier id="E3EA5DD0B6D68B3BACC3FD5479C17006" type="email">tanawat.chaowasku@cmu.ac.th</mods:nameIdentifier>
</mods:name>
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<mods:role id="3D40A5E0F81519DE6718DFF76C8B005A">
<mods:roleTerm id="1FA7C3F74B1F7DBF41F154D648EFD1AA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6EE5EBBC74C1C2D308EAF128A312CFFF">Murray, Nancy A.</mods:namePart>
<mods:nameIdentifier id="A7612B22D963A6ABEA7DD50F68AD7D0E" type="ORCID">0000-0003-4076-6169</mods:nameIdentifier>
<mods:affiliation id="185B3FC7B00A3AAF3F8B4A659680265C">namurray @ owu. edu; https: // orcid. org / 0000 - 0003 - 4076 - 6169</mods:affiliation>
<mods:nameIdentifier id="C73BC765BEA384F2847B52C71C1411B4" type="email">namurray@owu.edu</mods:nameIdentifier>
</mods:name>
<mods:name id="2CA84EEBC24B11A8815C939199D97F88" type="personal">
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<mods:roleTerm id="A6CD3EA44D5974F5921820F8350B406F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FAD30797BAFF56714124D4256844B88F">Chalermglin, Piya</mods:namePart>
<mods:nameIdentifier id="99ECE5E06F1D4B23F90EE7739E810130" type="ORCID">0000-0002-9098-0916</mods:nameIdentifier>
<mods:affiliation id="B242D59A46FFC6421AAA2B938281F38E">Agricultural Technology Department, Thailand Institute of Scientific and Technological Research, 35 Mu 3, Khlong Ha, Khlong Luang District, Pathum Thani Province 12120, Thailand &amp; piya. chalerm @ gmail. com; https: // orcid. org / 0000 - 0002 - 9098 - 0916</mods:affiliation>
<mods:nameIdentifier id="CE0D3FB4CB715060AA30F394FC3362A5" type="email">piya.chalerm@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:title id="EF12194846E5163F7CD38E785D216E95">Phytotaxa</mods:title>
</mods:titleInfo>
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<mods:date id="BC7279EA9DE41D4A33C74CE3811B8958">2021</mods:date>
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<mods:number id="2DD5192065C4E871A6378437E1946EEA">2021-03-03</mods:number>
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<mods:url id="E5503C028D2CEADA5BE83FDFB42DB69E">http://dx.doi.org/10.11646/phytotaxa.489.1.6</mods:url>
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<mods:identifier id="CAFEF0B58C5F38F51B4FBCDA32B3EB6D" type="ISSN">1179-3163</mods:identifier>
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<treatment id="03DB7B3F037BF010C7EBFF74FBB601BE" ID-DOI="http://doi.org/10.5281/zenodo.5757749" ID-Zenodo-Dep="5757749" LSID="urn:lsid:plazi:treatment:03DB7B3F037BF010C7EBFF74FBB601BE" httpUri="http://treatment.plazi.org/id/03DB7B3F037BF010C7EBFF74FBB601BE" lastPageNumber="80" pageId="1" pageNumber="80">
<subSubSection id="C36899A2037BF010C7EBFF74FC310502" box="[136,1019,244,271]" pageId="1" pageNumber="80" type="nomenclature">
<paragraph id="8BCDCA29037BF010C7EBFF74FC310502" blockId="1.[136,1019,244,271]" box="[136,1019,244,271]" pageId="1" pageNumber="80">
<heading id="D0857D45037BF010C7EBFF74FC310502" box="[136,1019,244,271]" fontSize="11" level="2" pageId="1" pageNumber="80" reason="3">
<taxonomicName id="4C72B1AA037BF010C7EBFF74FD1D0502" authority="Chaowasku &amp; D.M.Johnson" authorityName="Chaowasku &amp; D. M. Johnson" authorityYear="2021" box="[136,727,244,271]" class="Magnoliopsida" family="Annonaceae" genus="Trivalvaria" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="80" phylum="Tracheophyta" rank="species" species="ochroleuca" status="sp. nov.">
<emphasis id="B906163B037BF010C7EBFF74FE5A0503" bold="true" box="[136,400,244,271]" italics="true" pageId="1" pageNumber="80">Trivalvaria ochroleuca</emphasis>
Chaowasku &amp; D.M.Johnson
</taxonomicName>
,
<taxonomicNameLabel id="A235AB40037BF010C580FF75FCF30503" box="[739,825,245,271]" pageId="1" pageNumber="80" rank="species">
<emphasis id="B906163B037BF010C580FF75FCFB0502" box="[739,817,245,270]" italics="true" pageId="1" pageNumber="80">sp. nov</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="1349D6AC037BF010C42BFF74FC0F0502" box="[840,965,244,270]" captionStart="FIGURE 1" captionStartId="2.[136,229,1876,1898]" captionTargetBox="[247,1339,297,1846]" captionTargetId="figure-61@2.[247,1339,296,1848]" captionTargetPageId="2" captionText="FIGURE 1. Trivalvaria ochroleuca, T. recurva, and T.stenopetala.A–B.Trivalvaria ochroleuca.A. Habit, showing inflorescence position. B. Side view of bisexual flower from which the petals and stamens have fallen, showing the cylindrical receptacle. C–E. Trivalvaria recurva. C. Staminate flower, viewed from above. D. Bisexual flower, side view. E. Stamen, abaxial view. F–H. Trivalvaria stenopetala. F. Flower, side view. G. Leaf. H. Monocarp.A–B based on van Beusekom &amp; Phengklai 193 (L), C–E based on spirit collection of Nakorn- Thiemchan NTC 13 (OWU), F–G based on Middleton et al. 3915 (A), H based on Geesink et al. 7641 (L). Illustration by D.M. Johnson." figureDoi="http://doi.org/10.5281/zenodo.5757633" httpUri="https://zenodo.org/record/5757633/files/figure.png" pageId="1" pageNumber="80">Figs. 1A,B</figureCitation>
,
<figureCitation id="1349D6AC037BF010C4B2FF74FC390502" box="[977,1011,244,270]" captionStart="FIGURE 2" captionStartId="4.[136,229,1645,1667]" captionTargetBox="[172,1416,815,1617]" captionTargetId="figure-289@4.[170,1416,814,1620]" captionTargetPageId="4" captionText="FIGURE 2. Photographs of new Trivalvaria species. A. Flower of Trivalvaria stenopetala, Phangnga Province, Thailand. B. Flower of Trivalvaria ochroleuca, Tak Province, Thailand. C. Flower of Trivalvaria recurva, Kanchanaburi Province, Thailand. D. Fruit of Trivalvaria stenopetala, Phangnga Province, Thailand. E. Fruit of probable Trivalvaria ochroleuca from Prachuap Khiri Khan Province, Thailand. F. Fruit of Trivalvaria recurva from Kanchanaburi Province, Thailand.A and D by Simon Gardner from Gardner et al. ST 1944, B by Tanawat Chaowasku, C and F by Piya Chalermglin, E by Thammanoon Temchai." figureDoi="http://doi.org/10.5281/zenodo.5757635" httpUri="https://zenodo.org/record/5757635/files/figure.png" pageId="1" pageNumber="80">2B</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36899A2037BF010C7EBFEC0FDBB077E" pageId="1" pageNumber="80" type="materials_examined">
<paragraph id="8BCDCA29037BF010C7EBFEC0FBDB0576" blockId="1.[136,1452,320,450]" pageId="1" pageNumber="80">
<materialsCitation id="3B1AC074037BF010C7EBFEC0FBC70576" collectingDate="1968-03-29" collectionCode="E, L" collectorName="van Beusekom &amp; Phengklai" country="Thailand" county="Type" elevation="1100" location="Khao Yai" municipality="Southwestern District" pageId="1" pageNumber="80" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="holotype">
<collectingCounty id="62ACB2A5037BF010C7EBFEC0FF700559" box="[136,186,320,341]" pageId="1" pageNumber="80">Type</collectingCounty>
:—
<collectingCountry id="F3658AB9037BF010C7B5FEC0FE9C0559" box="[214,342,320,341]" name="Thailand" pageId="1" pageNumber="80">THAILAND</collectingCountry>
.
<collectingRegion id="49B604CB037BF010C602FEC0FE21055A" box="[353,491,320,342]" country="Thailand" name="Kanchanaburi" pageId="1" pageNumber="80">Kanchanaburi</collectingRegion>
:
<collectingMunicipality id="6BA95053037BF010C694FEC0FD04055A" box="[503,718,320,342]" pageId="1" pageNumber="80">Southwestern District</collectingMunicipality>
,
<location id="8EAD9CF2037BF010C5B9FEC0FCF3055A" LSID="urn:lsid:plazi:treatment:03DB7B3F037BF010C7EBFF74FBB601BE:8EAD9CF2037BF010C5B9FEC0FCF3055A" box="[730,825,320,342]" country="Thailand" county="Type" municipality="Southwestern District" name="Khao Yai" pageId="1" pageNumber="80" stateProvince="Kanchanaburi">Khao Yai</location>
,
<collectionCode id="ED6352EC037BF010C426FEC0FC9E0559" box="[837,852,320,341]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="1" pageNumber="80" type="Herbarium">E</collectionCode>
of
<location id="8EAD9CF2037BF010C41AFEC0FC1F055A" LSID="urn:lsid:plazi:treatment:03DB7B3F037BF010C7EBFF74FBB601BE:8EAD9CF2037BF010C41AFEC0FC1F055A" box="[889,981,320,342]" country="Thailand" county="Type" municipality="Southwestern District" name="Sangkhla" pageId="1" pageNumber="80" stateProvince="Kanchanaburi">Sangkhla</location>
, elev. c.
<quantity id="4C8A67CC037BF010C353FEC0FBB70559" box="[1072,1149,320,341]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="80" unit="m" value="1100.0">
<elevation id="005F2D1A037BF010C353FEC0FBB70559" box="[1072,1149,320,341]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="80" unit="m" value="1100.0">1100 m</elevation>
</quantity>
,
<date id="FFCCECE9037BF010C3E9FEC0FAE8055A" box="[1162,1314,320,342]" pageId="1" pageNumber="80" value="1968-03-29">
<collectingDate id="EF881501037BF010C3E9FEC0FAE8055A" box="[1162,1314,320,342]" pageId="1" pageNumber="80" value="1968-03-29">29 March 1968</collectingDate>
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(flower),
<emphasis id="B906163B037BF010C2EAFEC0FE030575" italics="true" pageId="1" pageNumber="80">
<collectorName id="2687AFFF037BF010C2EAFEC0FED40575" pageId="1" pageNumber="80">van Beusekom</collectorName>
&amp;
<collectorName id="2687AFFF037BF010C65EFEE4FE540575" box="[317,414,356,377]" pageId="1" pageNumber="80">Phengklai</collectorName>
193
</emphasis>
(
<typeStatus id="54C9748B037BF010C6B6FEE4FDE40576" box="[469,558,356,378]" pageId="1" pageNumber="80" type="holotype">holotype</typeStatus>
: BKF-4190!; isotypes:
<collectionCode id="ED6352EC037BF010C46DFEE4FCD40576" box="[782,798,356,378]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="1" pageNumber="80" type="Herbarium">L</collectionCode>
.2066282! P-00268392!)
</materialsCitation>
.
</paragraph>
<paragraph id="8BCDCA29037BF010C7EBFE08FB0C05CE" blockId="1.[136,1452,320,450]" pageId="1" pageNumber="80">
This new species is distinguishable from
<taxonomicName id="4C72B1AA037BF010C57BFE08FDBF0591" authorityName="Hooker &amp; Thomson" authorityYear="1855" box="[536,629,392,413]" class="Magnoliopsida" family="Annonaceae" genus="Guatteria" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="80" phylum="Tracheophyta" rank="species" species="costata">
<emphasis id="B906163B037BF010C57BFE08FDBF0591" box="[536,629,392,413]" italics="true" pageId="1" pageNumber="80">T. costata</emphasis>
</taxonomicName>
by rounded (versus angular and acute) inner petal apices, greater number of stamens (ca. 180 versus 30–50), and blunt leaf acumen
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long (versus acute and up to
<quantity id="4C8A67CC037BF010C484FE2CFBE005CD" box="[999,1066,428,450]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="80" unit="mm" value="26.0">26 mm</quantity>
long) (
<tableCitation id="C6F0FF92037BF010C313FE2CFB7D05CE" box="[1136,1207,428,450]" captionStart="TABLE 1" captionStartId="3.[136,224,735,759]" captionText="TABLE 1. A morphological comparison of the three new Trivalvaria species (names in bold) to the most similar congener of each. Data for T. casseabriae from Yang et al. (2018)." pageId="1" pageNumber="80">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8BCDCA29037BF010C7EBFE70FDBB077E" blockId="1.[136,1452,496,1459]" pageId="1" pageNumber="80">
Shrubs up to
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<quantity id="4C8A67CC037BF010C646FE70FE9C0607" box="[293,342,496,523]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="80" unit="m" value="3.0">3 m</quantity>
.
</specimenCount>
Twigs densely but finely brown-tomentose, eventually glabrate. Leaves elliptic to oblanceolate, 15.7–18.2
<emphasis id="B906163B037BF010C663FD96FED20621" bold="true" box="[256,280,534,557]" pageId="1" pageNumber="80"></emphasis>
5.9–6.0 cm, base cuneate to nearly rounded, apex blunt-acuminate, the acumen
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long, glabrous except for short coarse hairs along midrib above, with short coarse hairs along midrib and secondary veins but otherwise glabrous below, secondary veins eucamptodromous, 8–9 per side, relatively straight but strongly ascending; petiole
<quantity id="4C8A67CC037BF010C2ECFDDDFF030697" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="1" pageNumber="80" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">7– 8 mm</quantity>
long. Inflorescences sub-opposite to leaves, 1–2-flowered, peduncles 1.5–3.0 mm long, pedicels ca.
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long, bearing 2 bracts ca.
<quantity id="4C8A67CC037BF010C607FD24FE6E06B3" box="[356,420,676,703]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="80" unit="mm" value="2.0">2 mm</quantity>
long; buds not seen. Flowers bisexual. Sepals triangular, 2.5–3.0 mm long, obtuse, sericeous. Petals spreading at anthesis, white to dirty pale yellow, elliptic-oblong, ca. 5.5
<emphasis id="B906163B037BF010C499FD4AFBD806ED" bold="true" box="[1018,1042,714,737]" pageId="1" pageNumber="80"></emphasis>
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, rounded, glabrous on inner surface, sericeous on outer surface. Stamens ca. 180, forming a globose boss ca.
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in diameter, oblong, ca.
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long, anther connective apex truncate, glabrous. Carpels ca. 14, ovaries ca. 1.0 mm long, pubescent, stigmas globose, ca.
<quantity id="4C8A67CC037BF010C676FCB4FEA60743" box="[277,364,820,847]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="80" unit="mm" value="0.3">0.3 mm</quantity>
long, glabrous. Receptacle short-cylindrical, ca.
<quantity id="4C8A67CC037BF010C4F7FCB4FC210743" box="[916,1003,820,847]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="80" unit="mm" value="2.5">2.5 mm</quantity>
high, ca. 2.0 mm in diameter, glabrous. Fruit unknown (but see below and
<figureCitation id="1349D6AC037BF010C56DFCD8FDAF077E" box="[526,613,856,882]" captionStart="FIGURE 2" captionStartId="4.[136,229,1645,1667]" captionTargetBox="[172,1416,815,1617]" captionTargetId="figure-289@4.[170,1416,814,1620]" captionTargetPageId="4" captionText="FIGURE 2. Photographs of new Trivalvaria species. A. Flower of Trivalvaria stenopetala, Phangnga Province, Thailand. B. Flower of Trivalvaria ochroleuca, Tak Province, Thailand. C. Flower of Trivalvaria recurva, Kanchanaburi Province, Thailand. D. Fruit of Trivalvaria stenopetala, Phangnga Province, Thailand. E. Fruit of probable Trivalvaria ochroleuca from Prachuap Khiri Khan Province, Thailand. F. Fruit of Trivalvaria recurva from Kanchanaburi Province, Thailand.A and D by Simon Gardner from Gardner et al. ST 1944, B by Tanawat Chaowasku, C and F by Piya Chalermglin, E by Thammanoon Temchai." figureDoi="http://doi.org/10.5281/zenodo.5757635" httpUri="https://zenodo.org/record/5757635/files/figure.png" pageId="1" pageNumber="80">Fig. 2E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36899A2037BF010C7DEFCFCFD7207B6" pageId="1" pageNumber="80" type="distribution">
<paragraph id="8BCDCA29037BF010C7DEFCFCFB2A079B" blockId="1.[136,1452,496,1459]" box="[189,1248,892,919]" pageId="1" pageNumber="80">
<emphasis id="B906163B037BF010C7DEFCFCFE39079A" bold="true" box="[189,499,892,918]" pageId="1" pageNumber="80">Distribution and Ecology:</emphasis>
—Southwestern
<collectingCountry id="F3658AB9037BF010C5CDFCFCFCDE079A" box="[686,788,892,918]" name="Thailand" pageId="1" pageNumber="80">Thailand</collectingCountry>
, in dry evergreen forest at
<quantity id="4C8A67CC037BF010C35CFCFCFB16079B" box="[1087,1244,892,919]" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="1" pageNumber="80" unit="m" value="1150.0" valueMax="1200.0" valueMin="1100.0">1100–1200 m</quantity>
.
</paragraph>
<paragraph id="8BCDCA29037BF010C7DEFC20FD7207B6" blockId="1.[136,1452,496,1459]" box="[189,696,928,955]" pageId="1" pageNumber="80">
<emphasis id="B906163B037BF010C7DEFC20FE9507B6" bold="true" box="[189,351,928,954]" pageId="1" pageNumber="80">Phenology:—</emphasis>
Flowering January and March.
</paragraph>
</subSubSection>
<subSubSection id="C36899A2037BF010C7DEFC44FBB601BE" pageId="1" pageNumber="80" type="etymology">
<paragraph id="8BCDCA29037BF010C7DEFC44FBD807D2" blockId="1.[136,1452,496,1459]" box="[189,1042,964,991]" pageId="1" pageNumber="80">
<emphasis id="B906163B037BF010C7DEFC44FEAE07D2" bold="true" box="[189,356,964,990]" pageId="1" pageNumber="80">Etymology:—</emphasis>
Named for the flower colours, dull yellow and creamy white.
</paragraph>
<paragraph id="8BCDCA29037BF010C7DEFC68FCBE000F" blockId="1.[136,1452,496,1459]" box="[189,884,1000,1032]" pageId="1" pageNumber="80">
<emphasis id="B906163B037BF010C7DEFC68FE7B000E" bold="true" box="[189,433,1000,1026]" pageId="1" pageNumber="80">Vernacular name:—</emphasis>
<taxonomicName id="4C72B1AA037BF010C6D2FC69FD9B000E" authority="khao" box="[433,593,1000,1026]" class="Magnoliopsida" family="Annonaceae" genus="Marsypopetalum" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="80" phylum="Tracheophyta" rank="subSpecies" species="khao" subSpecies="khao">
<emphasis id="B906163B037BF010C6D2FC69FD9B000E" box="[433,593,1000,1026]" italics="true" pageId="1" pageNumber="80">Ma khao khao</emphasis>
</taxonomicName>
(มะขาวเขา) (
<collectingRegion id="49B604CB037BF010C5A7FC68FCA2000F" box="[708,872,1000,1027]" country="Thailand" name="Kanchanaburi" pageId="1" pageNumber="80">Kanchanaburi</collectingRegion>
).
</paragraph>
<paragraph id="8BCDCA29037BF010C7DEFB8CFBF900BA" blockId="1.[136,1452,496,1459]" pageId="1" pageNumber="80">
<emphasis id="B906163B037BF010C7DEFB8CFE04002A" bold="true" box="[189,462,1036,1062]" pageId="1" pageNumber="80">Conservation status:—</emphasis>
<taxonomicName id="4C72B1AA037BF010C6ACFB8CFD1A002A" authorityName="Chaowasku &amp; D. M. Johnson" authorityYear="2021" box="[463,720,1036,1062]" class="Magnoliopsida" family="Annonaceae" genus="Trivalvaria" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="80" phylum="Tracheophyta" rank="species" species="ochroleuca">
<emphasis id="B906163B037BF010C6ACFB8CFD1A002A" box="[463,720,1036,1062]" italics="true" pageId="1" pageNumber="80">Trivalvaria ochroleuca</emphasis>
</taxonomicName>
<materialsCitation id="3B1AC074037BF010C5B5FB8CFBE500BA" collectionCode="A" county="Nothing" location="Kerr" municipality="According" pageId="1" pageNumber="80" specimenCount="1" stateProvince="Kanchanaburi">
is known from a single locality, where it has been collected twice, 42 years apart, most recently in 1968.
<collectingCounty id="62ACB2A5037BF010C532FBB0FD670046" box="[593,685,1072,1098]" pageId="1" pageNumber="80">Nothing</collectingCounty>
is known of its relative abundance, or whether it is still present.
<collectingMunicipality id="6BA95053037BF010C7EBFBD4FF340062" box="[136,254,1108,1134]" pageId="1" pageNumber="80">According</collectingMunicipality>
to the
<location id="8EAD9CF2037BF010C62BFBD5FEB10063" LSID="urn:lsid:plazi:treatment:03DB7B3F037BF010C7EBFF74FBB601BE:8EAD9CF2037BF010C62BFBD5FEB10063" box="[328,379,1109,1135]" county="Nothing" municipality="According" name="Kerr" pageId="1" pageNumber="80" stateProvince="Kanchanaburi">Kerr</location>
itinerary given by
<bibRefCitation id="EFE3B7D8037BF010C528FBD4FD230063" author="Jacobs, M." box="[587,745,1108,1135]" pageId="1" pageNumber="80" pagination="427 - 493" refId="ref5262" refString="Jacobs, M. (1962) Reliquiae Kerrianae. Blumea 11: 427 - 493." type="journal article" year="1962">Jacobs (1962)</bibRefCitation>
, this locality is
<quantity id="4C8A67CC037BF010C4F7FBD4FC100062" box="[916,986,1108,1134]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="80" unit="km" value="20.0">20 km</quantity>
east of
<location id="8EAD9CF2037BF010C34EFBD4FB420062" LSID="urn:lsid:plazi:treatment:03DB7B3F037BF010C7EBFF74FBB601BE:8EAD9CF2037BF010C34EFBD4FB420062" box="[1069,1160,1108,1134]" county="Nothing" municipality="According" name="Wangka" pageId="1" pageNumber="80" stateProvince="Kanchanaburi">Wangka</location>
in northern
<collectingRegion id="49B604CB037BF010C26DFBD4FF25009E" country="Thailand" name="Kanchanaburi" pageId="1" pageNumber="80">Kanchanaburi Province</collectingRegion>
.
<collectionCode id="ED6352EC037BF010C79BFBF8FEC1009E" box="[248,267,1144,1170]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="1" pageNumber="80" type="Herbarium">A</collectionCode>
more recent collection from
<collectingRegion id="49B604CB037BF010C531FBF8FD2D009E" box="[594,743,1144,1170]" country="Thailand" name="Tak" pageId="1" pageNumber="80">Tak Province</collectingRegion>
(see below) has been lost. In light of its narrow distribution,
<taxonomicName id="4C72B1AA037BF010C2F4FBF9FECF00BA" authorityName="Chaowasku &amp; D. M. Johnson" authorityYear="2021" class="Magnoliopsida" family="Annonaceae" genus="Trivalvaria" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="80" phylum="Tracheophyta" rank="species" species="ochroleuca">
<emphasis id="B906163B037BF010C2F4FBF9FECF00BA" italics="true" pageId="1" pageNumber="80">T. ochroleuca</emphasis>
</taxonomicName>
is given a preliminary conservation assessment of critically endangered
</materialsCitation>
.
</paragraph>
<paragraph id="8BCDCA29037BF010C7DEFB40FD8900F3" blockId="1.[136,1452,496,1459]" pageId="1" pageNumber="80">
<emphasis id="B906163B037BF010C7DEFB40FD9700D6" bold="true" box="[189,605,1216,1242]" pageId="1" pageNumber="80">Additional specimen examined:—</emphasis>
<materialsCitation id="3B1AC074037BF010C53EFB40FDF500F3" collectingDate="1926-01-31" collectionCode="BK, L" country="Thailand" elevation="1200" location="Kao Ri Yai" pageId="1" pageNumber="80" specimenCount="1" stateProvince="Kanchanaburi">
<collectingCountry id="F3658AB9037BF010C53EFB40FD3B00D6" box="[605,753,1216,1242]" name="Thailand" pageId="1" pageNumber="80">THAILAND</collectingCountry>
.
<collectingRegion id="49B604CB037BF010C462FB40FC5400D6" box="[769,926,1216,1242]" country="Thailand" name="Kanchanaburi" pageId="1" pageNumber="80">Kanchanaburi</collectingRegion>
:
<location id="8EAD9CF2037BF010C4D7FB41FBF700D6" LSID="urn:lsid:plazi:treatment:03DB7B3F037BF010C7EBFF74FBB601BE:8EAD9CF2037BF010C4D7FB41FBF700D6" box="[948,1085,1216,1243]" country="Thailand" name="Kao Ri Yai" pageId="1" pageNumber="80" stateProvince="Kanchanaburi">Kao Ri Yai</location>
, ca.
<quantity id="4C8A67CC037BF010C319FB40FB1C00D7" box="[1146,1238,1216,1243]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="80" unit="m" value="1200.0">
<elevation id="005F2D1A037BF010C319FB40FB1C00D7" box="[1146,1238,1216,1243]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="80" unit="m" value="1200.0">1200 m</elevation>
</quantity>
,
<date id="FFCCECE9037BF010C384FB40FA6100D6" box="[1255,1451,1216,1243]" pageId="1" pageNumber="80" value="1926-01-31">
<collectingDate id="EF881501037BF010C384FB40FA6100D6" box="[1255,1451,1216,1243]" pageId="1" pageNumber="80" value="1926-01-31">31 January 1926</collectingDate>
</date>
(flower),
<emphasis id="B906163B037BF010C791FB65FEBE00F2" box="[242,372,1252,1278]" italics="true" pageId="1" pageNumber="80">Kerr 10368</emphasis>
(
<collectionCode id="ED6352EC037BF010C6E0FB64FE6700F2" box="[387,429,1252,1278]" country="Thailand" lsid="urn:lsid:biocol.org:col:12928" name="Department of Agriculture" pageId="1" pageNumber="80" type="Herbarium">BK</collectionCode>
,
<collectionCode id="ED6352EC037BF010C6DAFB64FE0100F3" box="[441,459,1252,1279]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="1" pageNumber="80" type="Herbarium">L</collectionCode>
.4311236)
</materialsCitation>
.
</paragraph>
<paragraph id="8BCDCA29037BF010C7DEFA88FBB601BE" blockId="1.[136,1452,496,1459]" pageId="1" pageNumber="80">
<emphasis id="B906163B037BF010C7DEFA88FEEC012E" bold="true" box="[189,294,1288,1314]" pageId="1" pageNumber="80">Notes:—</emphasis>
An additional specimen from nearby
<collectingRegion id="49B604CB037BF010C5A3FA88FC99012E" box="[704,851,1288,1314]" country="Thailand" name="Tak" pageId="1" pageNumber="80">Tak Province</collectingRegion>
in
<collectingCountry id="F3658AB9037BF010C417FA88FC12012E" box="[884,984,1288,1314]" name="Thailand" pageId="1" pageNumber="80">Thailand</collectingCountry>
was unfortunately lost, but the flower was photographed (
<figureCitation id="1349D6AC037BF010C650FAACFE47014A" box="[307,397,1324,1350]" captionStart="FIGURE 2" captionStartId="4.[136,229,1645,1667]" captionTargetBox="[172,1416,815,1617]" captionTargetId="figure-289@4.[170,1416,814,1620]" captionTargetPageId="4" captionText="FIGURE 2. Photographs of new Trivalvaria species. A. Flower of Trivalvaria stenopetala, Phangnga Province, Thailand. B. Flower of Trivalvaria ochroleuca, Tak Province, Thailand. C. Flower of Trivalvaria recurva, Kanchanaburi Province, Thailand. D. Fruit of Trivalvaria stenopetala, Phangnga Province, Thailand. E. Fruit of probable Trivalvaria ochroleuca from Prachuap Khiri Khan Province, Thailand. F. Fruit of Trivalvaria recurva from Kanchanaburi Province, Thailand.A and D by Simon Gardner from Gardner et al. ST 1944, B by Tanawat Chaowasku, C and F by Piya Chalermglin, E by Thammanoon Temchai." figureDoi="http://doi.org/10.5281/zenodo.5757635" httpUri="https://zenodo.org/record/5757635/files/figure.png" pageId="1" pageNumber="80">Fig. 2B</figureCitation>
). A DNA sequence was obtained for phylogenetic analysis (Chaowasku, unpubl.). Similarly, the fruit (
<figureCitation id="1349D6AC037BF010C797FAD0FE800166" box="[244,330,1360,1386]" captionStart="FIGURE 2" captionStartId="4.[136,229,1645,1667]" captionTargetBox="[172,1416,815,1617]" captionTargetId="figure-289@4.[170,1416,814,1620]" captionTargetPageId="4" captionText="FIGURE 2. Photographs of new Trivalvaria species. A. Flower of Trivalvaria stenopetala, Phangnga Province, Thailand. B. Flower of Trivalvaria ochroleuca, Tak Province, Thailand. C. Flower of Trivalvaria recurva, Kanchanaburi Province, Thailand. D. Fruit of Trivalvaria stenopetala, Phangnga Province, Thailand. E. Fruit of probable Trivalvaria ochroleuca from Prachuap Khiri Khan Province, Thailand. F. Fruit of Trivalvaria recurva from Kanchanaburi Province, Thailand.A and D by Simon Gardner from Gardner et al. ST 1944, B by Tanawat Chaowasku, C and F by Piya Chalermglin, E by Thammanoon Temchai." figureDoi="http://doi.org/10.5281/zenodo.5757635" httpUri="https://zenodo.org/record/5757635/files/figure.png" pageId="1" pageNumber="80">Fig. 2E</figureCitation>
) of a
<taxonomicName id="4C72B1AA037BF010C6F2FAD0FDC50166" authorityName="Miquel" authorityYear="1865" baseAuthorityName="Miquel" baseAuthorityYear="1860" box="[401,527,1360,1386]" class="Magnoliopsida" family="Annonaceae" genus="Trivalvaria" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="80" phylum="Tracheophyta" rank="genus">
<emphasis id="B906163B037BF010C6F2FAD0FDC50166" box="[401,527,1360,1386]" italics="true" pageId="1" pageNumber="80">Trivalvaria</emphasis>
</taxonomicName>
plant photographed in
<collectingRegion id="49B604CB037BF010C479FAD0FBB20166" box="[794,1144,1360,1386]" country="Thailand" name="Prachuap Khiri Khan" pageId="1" pageNumber="80">Prachuap Khiri Khan Province</collectingRegion>
is probably this species. A specimen collected by Gallatly from a slightly higher elevation in southern
<collectingCountry id="F3658AB9037BF010C4B9FAF5FB8D0183" box="[986,1095,1397,1423]" name="Myanmar" pageId="1" pageNumber="80">Myanmar</collectingCountry>
(Molyet,
<quantity id="4C8A67CC037BF010C3D5FAF4FADA0182" box="[1206,1296,1396,1423]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="1" pageNumber="80" unit="ft" value="5000.0">5000 ft.</quantity>
,
<date id="FFCCECE9037BF010C27FFAF4FA6D0182" box="[1308,1447,1396,1423]" pageId="1" pageNumber="80" value="1877-01-19">19 Jan 1877</date>
, K), cited by
<bibRefCitation id="EFE3B7D8037BF010C677FA18FE0101BE" author="King, G." box="[276,459,1432,1458]" pageId="1" pageNumber="80" pagination="1 - 130" refId="ref5279" refString="King, G. (1892) Materials for a flora of the Malay Peninsula, no. 4. Journal of the Asiatic Society of Bengal, Part 2, Natural History 61: 1 - 130." type="journal article" year="1892">King (1892: 27)</bibRefCitation>
, under
<taxonomicName id="4C72B1AA037BF010C57EFA18FCDA01BE" authority="King" authorityName="King" authorityYear="1892" baseAuthorityName="Hook.f. &amp; Thomson" box="[541,784,1432,1458]" class="Magnoliopsida" family="Annonaceae" genus="Ellipeia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="80" phylum="Tracheophyta" rank="species" species="costata">
<emphasis id="B906163B037BF010C57EFA18FD0401BE" box="[541,718,1432,1458]" italics="true" pageId="1" pageNumber="80">Ellipeia costata</emphasis>
King
</taxonomicName>
, may also represent this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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