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<mods:title id="11AD5E557F91A71B39ADFFD1CDC40A56">Sabellid worms from the Patagonian Shelf and Humboldt Current System (Annelida, Sabellidae): Phyllis Knight-Jones and José María Orensanzs collections</mods:title>
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<paragraph id="662836D0FF85A362FF7FD97AFDAA5828" blockId="46.[151,584,1556,1582]" box="[151,584,1556,1582]" pageId="46" pageNumber="47">
<heading id="3D6081BCFF85A362FF7FD97AFDAA5828" bold="true" box="[151,584,1556,1582]" fontSize="11" level="1" pageId="46" pageNumber="47" reason="1">
<emphasis id="54E3EAC2FF85A362FF7FD97AFDAA5828" bold="true" box="[151,584,1556,1582]" pageId="46" pageNumber="47">
Genus
<taxonomicName id="A1974D53FF85A362FF06D97AFDAA5828" ID-CoL="84SYS" authority="Bush, 1905" authorityName="Bush" authorityYear="1905" box="[238,584,1556,1582]" class="Polychaeta" family="Sabellidae" genus="Pseudopotamilla" kingdom="Animalia" order="Sabellida" pageId="46" pageNumber="47" phylum="Annelida" rank="genus">
<emphasis id="54E3EAC2FF85A362FF06D97AFE5A5828" bold="true" box="[238,440,1556,1582]" italics="true" pageId="46" pageNumber="47">Pseudopotamilla</emphasis>
<bibRefCitation id="02064B21FF85A362FE57D97AFDAA5828" author="Bush" box="[447,584,1556,1582]" pageId="46" pageNumber="47" refString="Bush, K. J. (1905) Tubicolous annelids of the tribes Sabellides and Serpulides from the Pacific Ocean. Harriman Alaska Expedition, 12, 169 - 346." type="journal article" year="1905">Bush, 1905</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2E8D655BFF85A363FF7FD93BFA635ED2" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="discussion">
<paragraph id="662836D0FF85A362FF7FD93BFCA058F8" blockId="46.[151,1436,1621,2009]" pageId="46" pageNumber="47">
<materialsCitation id="D6FF3C8DFF85A362FF7FD93BFCA058F8" ID-GBIF-Occurrence="1572324646" country="Iceland" county="North Pacific" location="Caribbean Sea" municipality="Arctic" pageId="46" pageNumber="47" specimenCount="1" typeStatus="holotype">
<emphasis id="54E3EAC2FF85A362FF7FD93BFEE95868" bold="true" box="[151,267,1621,1646]" pageId="46" pageNumber="47">Remarks.</emphasis>
<taxonomicName id="A1974D53FF85A362FEFCD93BFE305868" box="[276,466,1621,1646]" class="Polychaeta" family="Sabellidae" genus="Pseudopotamilla" kingdom="Animalia" order="Sabellida" pageId="46" pageNumber="47" phylum="Annelida" rank="genus">
<emphasis id="54E3EAC2FF85A362FEFCD93BFE305868" box="[276,466,1621,1646]" italics="true" pageId="46" pageNumber="47">Pseudopotamilla</emphasis>
</taxonomicName>
is represented by some boring species in rocks and dead coral, and characterized by the presence of unpaired proximal compound radiolar eyes on the outer margins of radioles. The
<typeStatus id="B92C8872FF85A362FAE1D914FADB5894" box="[1289,1337,1658,1682]" pageId="46" pageNumber="47">type</typeStatus>
species,
<taxonomicName id="A1974D53FF85A362FF7FD9F3FD4458B0" authority="Bruguiere, 1789" authorityName="Bruguiere" authorityYear="1789" box="[151,678,1693,1718]" class="Polychaeta" family="Sabellidae" genus="Pseudopotamilla" kingdom="Animalia" order="Sabellida" pageId="46" pageNumber="47" phylum="Annelida" rank="species" species="reniformis">
<emphasis id="54E3EAC2FF85A362FF7FD9F3FE3358B0" box="[151,465,1693,1718]" italics="true" pageId="46" pageNumber="47">Pseudopotamilla reniformis</emphasis>
(
<bibRefCitation id="02064B21FF85A362FE0BD9F3FD7D58B0" author="Bruguiere" box="[483,671,1693,1718]" pageId="46" pageNumber="47" refString="Bruguiere, L. G. (1789) Encyclopedie Methodique. Vol. 1. Part 1. Histoire naturelle des vers, Paris, 344 pp." type="book" year="1789">Bruguière, 1789</bibRefCitation>
)
</taxonomicName>
, was originally described from
<collectingCountry id="1E807640FF85A362FBF9D9F3FB8658B0" box="[1041,1124,1693,1718]" name="Iceland" pageId="46" pageNumber="47">Iceland</collectingCountry>
and has been subsequently reported from the
<location id="6348600BFF85A362FE86D9AFFDF558DC" LSID="urn:lsid:plazi:treatment:EE3E87C6FF85A363FF7FD97AFB7D5FAB:6348600BFF85A362FE86D9AFFDF558DC" box="[366,535,1729,1754]" country="Iceland" county="North Pacific" municipality="Arctic" name="Caribbean Sea" pageId="46" pageNumber="47">Caribbean Sea</location>
,
<location id="6348600BFF85A362FDCED9ACFD7358DF" LSID="urn:lsid:plazi:treatment:EE3E87C6FF85A363FF7FD97AFB7D5FAB:6348600BFF85A362FDCED9ACFD7358DF" box="[550,657,1730,1753]" country="Iceland" county="North Pacific" municipality="Arctic" name="European" pageId="46" pageNumber="47">European</location>
waters,
<location id="6348600BFF85A362FD10D9AFFC8458DC" LSID="urn:lsid:plazi:treatment:EE3E87C6FF85A363FF7FD97AFB7D5FAB:6348600BFF85A362FD10D9AFFC8458DC" box="[760,870,1729,1754]" country="Iceland" county="North Pacific" municipality="Arctic" name="American" pageId="46" pageNumber="47">American</location>
coasts of North and
<location id="6348600BFF85A362FB8BD9AFFAEF58DC" LSID="urn:lsid:plazi:treatment:EE3E87C6FF85A363FF7FD97AFB7D5FAB:6348600BFF85A362FB8BD9AFFAEF58DC" box="[1123,1293,1729,1754]" country="Iceland" county="North Pacific" municipality="Arctic" name="South Atlantic" pageId="46" pageNumber="47">South Atlantic</location>
,
<collectingMunicipality id="864CACAAFF85A362FAF4D9AFFA8758DC" box="[1308,1381,1729,1754]" pageId="46" pageNumber="47">Arctic</collectingMunicipality>
, and
<collectingCounty id="8F494E5CFF85A362FF7FD98BFECF58F8" box="[151,301,1765,1790]" pageId="46" pageNumber="47">North Pacific</collectingCounty>
(
<bibRefCitation id="02064B21FF85A362FED3D98BFDD158F8" author="Kolbasova" box="[315,563,1765,1790]" pageId="46" pageNumber="47" refString="Kolbasova, G. D., Tzetlin, A. B. &amp; Kupriyanova, E. K. (2013) Biology of Pseudopotamilla reniformis (Muller, 1771) in the White Sea, with description of asexual reproduction. Invertebrate Reproduction &amp; Development, 57, 264 - 275. https: // doi. org / 10.1080 / 07924259.2012.759164" type="journal article" year="2013">
Kolbasova
<emphasis id="54E3EAC2FF85A362FE54D988FE1658F8" box="[444,500,1765,1790]" italics="true" pageId="46" pageNumber="47">et al.</emphasis>
2013
</bibRefCitation>
and references therein).
</materialsCitation>
</paragraph>
<paragraph id="662836D0FF85A363FF2FD862FA635ED2" blockId="46.[151,1436,1621,2009]" lastBlockId="47.[151,1436,151,429]" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
As shown in her poster presented during the 8th International Polychaeta Conference in
<collectingRegion id="A453F832FF85A362FB77D862FB165923" box="[1183,1268,1804,1829]" country="Spain" name="Madrid" pageId="46" pageNumber="47">Madrid</collectingRegion>
, Knight-Jones, Darbyshire and Petersen were working on the designation of a
<typeStatus id="B92C8872FF85A362FC9DD85FFC2C594F" box="[885,974,1841,1865]" pageId="46" pageNumber="47" type="neotype">neotype</typeStatus>
for
<taxonomicName id="A1974D53FF85A362FBEAD85EFADC594E" box="[1026,1342,1840,1865]" class="Polychaeta" family="Sabellidae" genus="Pseudopotamilla" kingdom="Animalia" order="Sabellida" pageId="46" pageNumber="47" phylum="Annelida" rank="species" species="reniformis">
<emphasis id="54E3EAC2FF85A362FBEAD85EFADC594E" box="[1026,1342,1840,1865]" italics="true" pageId="46" pageNumber="47">Pseudopotamilla reniformis</emphasis>
</taxonomicName>
and the resurrection of two species previously synonymized with
<taxonomicName id="A1974D53FF85A362FCAFD83BFAC6596A" authority="Knight-Jones et al. 2004" authorityName="Knight-Jones et al." authorityYear="2004" box="[839,1316,1876,1901]" class="Polychaeta" family="Sabellidae" genus="Pseudopotamilla" kingdom="Animalia" order="Sabellida" pageId="46" pageNumber="47" phylum="Annelida" rank="species" species="reniformis">
<emphasis id="54E3EAC2FF85A362FCAFD83BFC3E596A" box="[839,988,1876,1900]" italics="true" pageId="46" pageNumber="47">P. reniformis</emphasis>
(
<bibRefCitation id="02064B21FF85A362FC18D83AFAFF596A" author="Knight-Jones" box="[1008,1309,1876,1901]" pageId="46" pageNumber="47" refString="Knight-Jones, P., Darbyshire, T. &amp; Petersen, M. E. (2004) What is Pseudopotamilla reniformis (Sabellidae)? Comparisons of populations from Britain, Iceland and Nova Scotia. Poster presented during the VIII International Polychaeta Conference, Madrid, Spain. [poster]" type="book" year="2004">
Knight-Jones
<emphasis id="54E3EAC2FF85A362FB7CD83BFB31596B" box="[1172,1235,1876,1901]" italics="true" pageId="46" pageNumber="47">et al.</emphasis>
2004
</bibRefCitation>
)
</taxonomicName>
; however, Knight-Jones and Petersen died before this work was completed, but the designation of a
<typeStatus id="B92C8872FF85A362FB4CD817FB1F5997" box="[1188,1277,1913,1937]" pageId="46" pageNumber="47" type="neotype">neotype</typeStatus>
is now under revision (
<bibRefCitation id="02064B21FF85A362FEE8D8F2FDFB59B2" author="Knight-Jones" box="[256,537,1948,1973]" pageId="46" pageNumber="47" refString="Knight-Jones, P., Darbyshire, T., Petersen, M. E. &amp; Tovar-Hernandez, M. A. (2017) What is Pseudopotamilla reniformis (Sabellidae)? Comparisons of populations from Britain, Iceland and Canada with comments on Eudistylia and Schizobranchia. Zootaxa, 4254 (2), 201 - 220." type="journal article" year="2017">
Knight-Jones
<emphasis id="54E3EAC2FF85A362FE75D8F3FE3459B3" box="[413,470,1948,1973]" italics="true" pageId="46" pageNumber="47">et al.</emphasis>
2017
</bibRefCitation>
). In addition, records of
<taxonomicName id="A1974D53FF85A362FCC4D8F3FC5959B2" box="[812,955,1948,1972]" class="Polychaeta" family="Sabellidae" genus="Pseudopotamilla" kingdom="Animalia" order="Sabellida" pageId="46" pageNumber="47" phylum="Annelida" rank="species" species="reniformis">
<emphasis id="54E3EAC2FF85A362FCC4D8F3FC5959B2" box="[812,955,1948,1972]" italics="true" pageId="46" pageNumber="47">P. reniformis</emphasis>
</taxonomicName>
from the Caribbean area were corrected as
<taxonomicName id="A1974D53FF85A362FF7FD8AFFCAA59DF" authority="Tovar-Hernandez &amp; Salazar-Vallejo, 2006" authorityName="Tovar-Hernandez &amp; Salazar-Vallejo" authorityYear="2006" box="[151,840,1984,2009]" class="Polychaeta" family="Sabellidae" genus="Pseudopotamilla" kingdom="Animalia" order="Sabellida" pageId="46" pageNumber="47" phylum="Annelida" rank="species" species="fitzhughi">
<emphasis id="54E3EAC2FF85A362FF7FD8AFFEC359DF" box="[151,289,1984,2009]" italics="true" pageId="46" pageNumber="47">P. fitzhughi</emphasis>
(
<bibRefCitation id="02064B21FF85A362FED4D8AEFCA259DF" author="Tovar-Hernandez" box="[316,832,1984,2009]" pageId="46" pageNumber="47" refString="Tovar-Hernandez, M. A. &amp; Sosa-Rodriguez, T. (2006) Redescription of Chone infundibuliformis Kroyer, 1856 (Polychaeta: Sabellidae) and histology of the branchial crown appendages, collar and glandular ridge. Zootaxa, 1115, 31 - 59." type="journal article" year="2006">Tovar-Hernández &amp; Salazar-Vallejo, 2006</bibRefCitation>
)
</taxonomicName>
, and those from
<collectingCountry id="1E807640FF85A362FBDDD8AEFB7E59DF" box="[1077,1180,1984,2009]" name="Australia" pageId="46" pageNumber="47">Australia</collectingCountry>
were reassigned as
<taxonomicName id="A1974D53FF84A363FF7FDFF9FEB75EB6" box="[151,341,151,176]" class="Polychaeta" family="Sabellidae" genus="Pseudopotamilla" kingdom="Animalia" order="Sabellida" pageId="47" pageNumber="48" phylum="Annelida" rank="genus">
<emphasis id="54E3EAC2FF84A363FF7FDFF9FEB75EB6" box="[151,341,151,176]" italics="true" pageId="47" pageNumber="48">Pseudopotamilla</emphasis>
</taxonomicName>
sp. A and sp. B by
<bibRefCitation id="02064B21FF84A363FDC4DFF6FD5B5EB6" author="Capa" box="[556,697,152,176]" pageId="47" pageNumber="48" refString="Capa, M. (2007) Taxonomic revision and phylogenetic relationships of apomorphic sabellids (Sabellidae: Polychaeta) from Australia. Invertebrate Systematics, 21, 537 - 567." type="journal article" year="2007">Capa (2007)</bibRefCitation>
, showing that some misidentifications of
<taxonomicName id="A1974D53FF84A363FB6FDFF9FAA75EB6" box="[1159,1349,151,176]" class="Polychaeta" family="Sabellidae" genus="Pseudopotamilla" kingdom="Animalia" order="Sabellida" pageId="47" pageNumber="48" phylum="Annelida" rank="genus">
<emphasis id="54E3EAC2FF84A363FB6FDFF9FAA75EB6" box="[1159,1349,151,176]" italics="true" pageId="47" pageNumber="48">Pseudopotamilla</emphasis>
</taxonomicName>
species have occurred. A full revision of
<taxonomicName id="A1974D53FF84A363FDE3DFD2FD2B5ED3" box="[523,713,188,213]" class="Polychaeta" family="Sabellidae" genus="Pseudopotamilla" kingdom="Animalia" order="Sabellida" pageId="47" pageNumber="48" phylum="Annelida" rank="genus">
<emphasis id="54E3EAC2FF84A363FDE3DFD2FD2B5ED3" box="[523,713,188,213]" italics="true" pageId="47" pageNumber="48">Pseudopotamilla</emphasis>
</taxonomicName>
is needed in order to confirm the distribution of
<taxonomicName id="A1974D53FF84A363FB03DFD3FA985ED2" box="[1259,1402,188,212]" class="Polychaeta" family="Sabellidae" genus="Pseudopotamilla" kingdom="Animalia" order="Sabellida" pageId="47" pageNumber="48" phylum="Annelida" rank="species" species="reniformis">
<emphasis id="54E3EAC2FF84A363FB03DFD3FA985ED2" box="[1259,1402,188,212]" italics="true" pageId="47" pageNumber="48">P. reniformis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="2E8D655BFF84A363FF2FDFB1FB7D5FAB" pageId="47" pageNumber="48" type="reference_group">
<paragraph id="662836D0FF84A363FF2FDFB1FB7D5FAB" blockId="47.[151,1436,151,429]" pageId="47" pageNumber="48">
<taxonomicName id="A1974D53FF84A363FF2FDFB1FE675EFE" ID-CoL="84SYS" box="[199,389,223,248]" class="Polychaeta" family="Sabellidae" genus="Pseudopotamilla" kingdom="Animalia" order="Sabellida" pageId="47" pageNumber="48" phylum="Annelida" rank="genus">
<emphasis id="54E3EAC2FF84A363FF2FDFB1FE675EFE" box="[199,389,223,248]" italics="true" pageId="47" pageNumber="48">Pseudopotamilla</emphasis>
</taxonomicName>
was emended by
<bibRefCitation id="02064B21FF84A363FDA3DF8EFD3B5EFE" author="Capa" box="[587,729,224,248]" pageId="47" pageNumber="48" refString="Capa, M. (2007) Taxonomic revision and phylogenetic relationships of apomorphic sabellids (Sabellidae: Polychaeta) from Australia. Invertebrate Systematics, 21, 537 - 567." type="journal article" year="2007">Capa (2007)</bibRefCitation>
. The genus includes 16 species (
<bibRefCitation id="02064B21FF84A363FBABDF8EFB1C5EFE" author="Capa" box="[1091,1278,223,248]" pageId="47" pageNumber="48" refString="Capa, M., Giangrande, A., Nogueira, J. M. M. &amp; Tovar-Hernandez, M. A. (2014) Sabellidae Latreille, 1825. In: Westheide, G. &amp; Purschke, G. (Eds.), The Handbook of Zoology. De Gruyter, Germany. Available from: http: // www. degruyter. com / view / Zoology / bp _ 029147 - 6 - 62 (accessed 23 November 2015)" type="journal article" year="2014">
Capa
<emphasis id="54E3EAC2FF84A363FB6CDF8FFB5E5EFE" box="[1156,1212,223,248]" italics="true" pageId="47" pageNumber="48">et al.</emphasis>
2014
</bibRefCitation>
), but only one was described from South America:
<taxonomicName id="A1974D53FF84A363FDAFDE6BFBA05F1B" authority="Hartmann-Schroder, 1965" authorityName="Hartmann-Schroder" authorityYear="1965" box="[583,1090,260,285]" class="Polychaeta" family="Sabellidae" genus="Pseudopotamilla" kingdom="Animalia" order="Sabellida" pageId="47" pageNumber="48" phylum="Annelida" rank="species" species="polyophthalma">
<emphasis id="54E3EAC2FF84A363FDAFDE6BFCED5F1B" box="[583,783,260,285]" italics="true" pageId="47" pageNumber="48">P. polyophthalma</emphasis>
<bibRefCitation id="02064B21FF84A363FCF2DE6AFBA05F1B" author="Hartmann-Schroder" box="[794,1090,260,285]" pageId="47" pageNumber="48" refString="Hartmann-Schroder, G. (1965) Zur Kentnnis des Sublitorals der chilenischen Kuste, unter besonderer Berucksichtigung der Polychaeten und Ostracoden. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 62, 59 - 305." type="journal article" year="1965">Hartmann-Schröder, 1965</bibRefCitation>
</taxonomicName>
from Punta Lavapie (
<collectingRegion id="A453F832FF84A363FAA0DE6AFA7E5F1B" box="[1352,1436,260,285]" country="Sri Lanka" name="Central" pageId="47" pageNumber="48">Central</collectingRegion>
<collectingCountry id="1E807640FF84A363FF7FDE49FF375F46" box="[151,213,295,320]" name="Chile" pageId="47" pageNumber="48">Chile</collectingCountry>
) to
<quantity id="A16F9B35FF84A363FEEEDE49FEA05F39" box="[262,322,295,320]" metricMagnitude="1" metricUnit="m" metricValue="5.8" pageId="47" pageNumber="48" unit="m" value="58.0">58 m</quantity>
depth. In the present paper, a new species of
<taxonomicName id="A1974D53FF84A363FCB2DE49FBFA5F46" ID-CoL="84SYS" box="[858,1048,295,320]" class="Polychaeta" family="Sabellidae" genus="Pseudopotamilla" kingdom="Animalia" order="Sabellida" pageId="47" pageNumber="48" phylum="Annelida" rank="genus">
<emphasis id="54E3EAC2FF84A363FCB2DE49FBFA5F46" box="[858,1048,295,320]" italics="true" pageId="47" pageNumber="48">Pseudopotamilla</emphasis>
</taxonomicName>
is described from
<collectingCountry id="1E807640FF84A363FB1BDE49FA875F46" box="[1267,1381,295,320]" name="Argentina" pageId="47" pageNumber="48">Argentina</collectingCountry>
, and
<taxonomicName id="A1974D53FF84A363FF7FDE22FDC75F63" ID-CoL="4M3CM" authority="Hartman, 1953" authorityName="Hartman" authorityYear="1953" box="[151,549,332,357]" class="Polychaeta" family="Sabellidae" genus="Potamilla" kingdom="Animalia" order="Sabellida" pageId="47" pageNumber="48" phylum="Annelida" rank="species" species="platensis">
<emphasis id="54E3EAC2FF84A363FF7FDE22FE935F63" box="[151,369,332,357]" italics="true" pageId="47" pageNumber="48">Potamilla platensis</emphasis>
<bibRefCitation id="02064B21FF84A363FE91DE23FDC75F63" author="Hartman" box="[377,549,332,357]" pageId="47" pageNumber="48" refString="Hartman, O. (1953) Non-pelagic polychaeta of the Swedish Antarctic Expedition 1901 - 1903. Further Zoological Results of the Swedish Antarctic Expedition 1901 - 1903, 4, 1 - 83." type="journal article" year="1953">Hartman, 1953</bibRefCitation>
</taxonomicName>
, is transferred to
<taxonomicName id="A1974D53FF84A363FD04DE22FC485F63" ID-CoL="84SYS" box="[748,938,332,357]" class="Polychaeta" family="Sabellidae" genus="Pseudopotamilla" kingdom="Animalia" order="Sabellida" pageId="47" pageNumber="48" phylum="Annelida" rank="genus">
<emphasis id="54E3EAC2FF84A363FD04DE22FC485F63" box="[748,938,332,357]" italics="true" pageId="47" pageNumber="48">Pseudopotamilla</emphasis>
</taxonomicName>
and redescribed.
<taxonomicName id="A1974D53FF84A363FB9ADE22FA7E5F63" ID-CoL="78GQF" box="[1138,1436,332,357]" class="Polychaeta" family="Sabellidae" genus="Pseudopotamilla" kingdom="Animalia" order="Sabellida" pageId="47" pageNumber="48" phylum="Annelida" rank="species" species="platensis">
<emphasis id="54E3EAC2FF84A363FB9ADE22FA7E5F63" box="[1138,1436,332,357]" italics="true" pageId="47" pageNumber="48">Pseudopotamilla platensis</emphasis>
</taxonomicName>
is commonly found in translucent, chitinous tubes with an external layer of sand grains, from intertidal to
<quantity id="A16F9B35FF84A363FABCDE1EFA7E5F81" box="[1364,1436,368,392]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="47" pageNumber="48" unit="m" value="200.0">200 m</quantity>
depth, whereas
<taxonomicName id="A1974D53FF84A363FEAEDEFBFE1F5FAB" ID-CoL="4P8TP" box="[326,509,404,429]" class="Polychaeta" family="Sabellidae" genus="Pseudopotamilla" kingdom="Animalia" order="Sabellida" pageId="47" pageNumber="48" phylum="Annelida" rank="species" species="knightjonesae" status="sp. nov.">
<emphasis id="54E3EAC2FF84A363FEAEDEFBFE1F5FAB" box="[326,509,404,429]" italics="true" pageId="47" pageNumber="48">P. knightjonesae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FD057B9FF84A363FDECDEFAFDBF5FAB" box="[516,605,404,429]" pageId="47" pageNumber="48" rank="species">
<emphasis id="54E3EAC2FF84A363FDECDEFAFDBF5FAB" bold="true" box="[516,605,404,429]" pageId="47" pageNumber="48">sp. nov.</emphasis>
</taxonomicNameLabel>
, was collected intertidally in hard sediment matrix.
</paragraph>
</subSubSection>
</treatment>
</document>