179 lines
20 KiB
XML
179 lines
20 KiB
XML
<document id="74423A6D01DFF610B7CCE3201614968E" ID-CLB-Dataset="27922" ID-DOI="10.11646/zootaxa.4524.3.4" ID-GBIF-Dataset="eb75e8f5-6d30-4886-a0ee-a08d99880b1b" ID-ISSN="1175-5326" ID-Zenodo-Dep="2610584" ID-ZooBank="C1975EB8-F799-4360-BA89-90990C97EEB3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553625673737" checkinUser="plazi" docAuthor="Laudee, Pongsak & Malicky, Hans" docDate="2018" docId="A02E8941F22D7656FF459E3EFB7CEA45" docLanguage="en" docName="zootaxa.4524.3.4.pdf" docOrigin="Zootaxa 4524 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptocerus suwannarati Laudee & Malicky 2018, n. sp." docType="treatment" docVersion="8" lastPageNumber="356" masterDocId="5C17F139F2297653FFD2980DFFB9E81A" masterDocTitle="Five new species of Caddisflies (Trichoptera: Insecta) from Upper Tenasserim Range, Thailand and Myanmar" masterLastPageNumber="358" masterPageNumber="351" pageNumber="355" updateTime="1698714027903" updateUser="ExternalLinkService">
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<mods:mods id="C76B7C9DBD518B3DCAE274C56470EB7D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="CC7F5E24ECA46D32AA84FEE029C3D9E9">
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<mods:title id="14EBB40376D541C20FCE81A19892844D">Five new species of Caddisflies (Trichoptera: Insecta) from Upper Tenasserim Range, Thailand and Myanmar</mods:title>
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<mods:name id="B48461700C5CDA5EA081C49A76D034E4" type="personal">
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<mods:roleTerm id="19E37DF79E67D837DC2948E9B9CD3D9C">Author</mods:roleTerm>
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<mods:namePart id="C4767354CF135DDE512B587C6CCD544C">Laudee, Pongsak</mods:namePart>
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<mods:name id="3E0C93483CE2D6B8ED25266DF6D1FBE2" type="personal">
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<mods:roleTerm id="86D90F2CC417CD8A3E3BFA81B235415E">Author</mods:roleTerm>
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<mods:namePart id="61746AA8F4A6F810755C8703255D0B2F">Malicky, Hans</mods:namePart>
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<mods:typeOfResource id="C40AD9E3B7FF6EDAF348C01916A78671">text</mods:typeOfResource>
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<mods:title id="5B2BDF91A3A69A9816E2C5FA8457AA95">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A7B24DC3C6045B0E03BADBB84ED20918">
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<mods:date id="F0092295E56E02151CC4D3AF8E14B186">2018</mods:date>
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<mods:detail id="79EA844B056073CE5947674F8CD3B3B2" type="pubDate">
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<mods:number id="CA3CC7773DD5B723F4CB649D505CE0F1">2018-11-22</mods:number>
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<mods:number id="CB2CE242B97955DC36D68CD0134DBD83">4524</mods:number>
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<mods:number id="9FD29877C6DD2778AA7C05989A73A35D">3</mods:number>
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<mods:classification id="B1CA33478583EF30147C0510F8E71867">journal article</mods:classification>
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<mods:identifier id="A7E014634E7F9907D16AF49F94AC2CA0" type="CLB-Dataset">27922</mods:identifier>
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<mods:identifier id="1032CD3D3C62B482CAF7D04783B9269A" type="DOI">10.11646/zootaxa.4524.3.4</mods:identifier>
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<treatment id="A02E8941F22D7656FF459E3EFB7CEA45" ID-DOI="http://doi.org/10.5281/zenodo.5957891" ID-GBIF-Taxon="154499618" ID-Zenodo-Dep="5957891" LSID="urn:lsid:plazi:treatment:A02E8941F22D7656FF459E3EFB7CEA45" httpUri="http://treatment.plazi.org/id/A02E8941F22D7656FF459E3EFB7CEA45" lastPageId="5" lastPageNumber="356" pageId="4" pageNumber="355">
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<subSubSection id="609D6BDCF22D7657FF459E3EFCB4EE57" box="[151,781,1587,1613]" pageId="4" pageNumber="355" type="nomenclature">
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<paragraph id="28383857F22D7657FF459E3EFCB4EE57" blockId="4.[151,781,1587,1647]" box="[151,781,1587,1613]" pageId="4" pageNumber="355">
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<heading id="73708F3BF22D7657FF459E3EFCB4EE57" bold="true" box="[151,781,1587,1613]" fontSize="11" level="1" pageId="4" pageNumber="355" reason="1">
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<emphasis id="1AF3E445F22D7657FF459E3EFCB4EE57" bold="true" box="[151,781,1587,1613]" pageId="4" pageNumber="355">
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<taxonomicName id="EF8743D4F22D7657FF459E3EFD7AEE57" authority="Laudee and Malicky" authorityName="Laudee & Malicky" authorityYear="2018" box="[151,707,1587,1613]" class="Insecta" family="Leptoceridae" genus="Leptocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="355" phylum="Arthropoda" rank="species" species="suwannarati" status="sp. nov.">
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<emphasis id="1AF3E445F22D7657FF459E3EFE02EE57" bold="true" box="[151,443,1587,1613]" italics="true" pageId="4" pageNumber="355">Leptocerus suwannarati</emphasis>
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Laudee and Malicky
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</taxonomicName>
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<taxonomicNameLabel id="01C0593EF22D7657FD199E3EFCB4EE57" box="[715,781,1587,1613]" pageId="4" pageNumber="355" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="609D6BDCF22D7657FF459E5BFEA5EE75" box="[151,284,1622,1647]" pageId="4" pageNumber="355" type="description">
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<paragraph id="28383857F22D7657FF459E5BFEA5EE75" blockId="4.[151,781,1587,1647]" box="[151,284,1622,1647]" pageId="4" pageNumber="355">
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<figureCitation id="B0BC24D2F22D7657FF459E5BFEA5EE75" box="[151,284,1622,1647]" captionStart="FIGURES 17–21" captionStartId="5.[151,264,1559,1582]" captionTargetBox="[151,1436,656,1506]" captionTargetId="figure@5.[151,1437,625,1508]" captionTargetPageId="5" captionText="FIGURES 17–21. Male genitalia of Leptocerus suwannarati n. sp. 17, left lateral; 18, dorsal; 19, ventral; 20, phallus, left lateral; 21, phallus, ventral. Inf = inferior appendage (paired), IX = segment IX, X = tergum X." httpUri="https://zenodo.org/record/2610592/files/figure.png" pageId="4" pageNumber="355">Figs. 17–21</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="609D6BDCF22D7657FF459E93FBC9EEC1" pageId="4" pageNumber="355" type="materials_examined">
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<paragraph id="28383857F22D7657FF459E93FBC9EEC1" blockId="4.[151,1436,1694,2007]" pageId="4" pageNumber="355">
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<emphasis id="1AF3E445F22D7657FF459E93FEFCEEAD" bold="true" box="[151,325,1694,1719]" pageId="4" pageNumber="355">Type material.</emphasis>
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<materialsCitation id="98EF320AF22D7657FE9E9E93FBC9EEC1" ID-GBIF-Occurrence="2028521987" collectingDate="2016-02-02" collectionCode="PSUNHM" collectorName="Pongsak Laudee." country="Thailand" elevation="150" latitude="12.081111" location="Kui Buri" longLatPrecision="21" longitude="99.62862" municipality="Kuiburi National Park" pageId="4" pageNumber="355" specimenCount="1" stateProvince="Prachuap Khiri Khan" typeStatus="holotype">
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<typeStatus id="F73C86F5F22D7657FE9E9E93FE0DEEAD" box="[332,436,1694,1719]" pageId="4" pageNumber="355" type="holotype">Holotype</typeStatus>
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male (
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<collectionCode id="4E96A092F22D7657FDD39E92FDC4EEAD" box="[513,637,1695,1719]" pageId="4" pageNumber="355">PSUNHM</collectionCode>
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).
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<collectingCountry id="509078C7F22D7657FD5D9E93FD4EEEAD" box="[655,759,1694,1719]" name="Thailand" pageId="4" pageNumber="355">Thailand</collectingCountry>
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:
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<collectingRegion id="EA43F6B5F22D7657FCD09E93FBE6EEAD" box="[770,1119,1694,1719]" country="Thailand" name="Prachuap Khiri Khan" pageId="4" pageNumber="355">Prachuap Khiri Khan Province</collectingRegion>
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,
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<collectingMunicipality id="C85CA22DF22D7657FBB89E93FADEEEAD" box="[1130,1383,1694,1719]" pageId="4" pageNumber="355">Kuiburi National Park</collectingMunicipality>
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,
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<location id="2D586E8CF22D7657FAA09E93FF75EEC1" LSID="urn:lsid:plazi:treatment:A02E8941F22D7656FF459E3EFB7CEA45:2D586E8CF22D7657FAA09E93FF75EEC1" country="Thailand" latitude="12.081111" longLatPrecision="21" longitude="99.62862" municipality="Kuiburi National Park" name="Kui Buri" pageId="4" pageNumber="355" stateProvince="Prachuap Khiri Khan">Kui Buri</location>
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,
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<geoCoordinate id="4DB35E90F22D7657FF059ECFFEDBEEC1" box="[215,354,1730,1755]" direction="north" orientation="latitude" pageId="4" pageNumber="355" precision="15" value="12.081111">12°04’52”N</geoCoordinate>
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,
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<geoCoordinate id="4DB35E90F22D7657FEBC9ECFFE4FEEC0" box="[366,502,1730,1754]" direction="east" orientation="longitude" pageId="4" pageNumber="355" precision="15" value="99.62862">99°37’43”E</geoCoordinate>
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, ca.
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<elevation id="A3AADF64F22D7657FDFB9ECFFD15EEC1" box="[553,684,1730,1755]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="355" unit="m" value="150.0">
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<quantity id="EF7F95B2F22D7657FDFB9ECFFDC9EEC0" box="[553,624,1730,1755]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="355" unit="m" value="150.0">150 m</quantity>
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a.s.l.
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</elevation>
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,
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<date id="5C391E97F22D7657FD6B9ECFFCCCEEC1" box="[697,885,1730,1755]" pageId="4" pageNumber="355" value="2016-02-02">
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<collectingDate id="4C7DE77FF22D7657FD6B9ECFFCCCEEC1" box="[697,885,1730,1755]" pageId="4" pageNumber="355" value="2016-02-02">2 February 2016</collectingDate>
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</date>
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, leg.
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<collectorName id="85725D81F22D7657FC629ECFFBC9EEC1" box="[944,1136,1730,1755]" pageId="4" pageNumber="355">Pongsak Laudee.</collectorName>
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="609D6BDCF22D7657FF159EEBFE60EF38" pageId="4" pageNumber="355" type="etymology">
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<paragraph id="28383857F22D7657FF159EEBFE60EF38" blockId="4.[151,1436,1694,2007]" pageId="4" pageNumber="355">
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<emphasis id="1AF3E445F22D7657FF159EEBFEF7EEE5" bold="true" box="[199,334,1766,1791]" pageId="4" pageNumber="355">Etymology.</emphasis>
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Named for Asst. Prof. Wasin Suwannarat who is the Vice President of Prince of Songkla University, Hat Yai Campus.
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</paragraph>
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</subSubSection>
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<subSubSection id="609D6BDCF22D7656FF159F23FE67E9B7" lastPageId="5" lastPageNumber="356" pageId="4" pageNumber="355" type="description">
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<paragraph id="28383857F22D7657FF159F23FDA7EF71" blockId="4.[151,1436,1694,2007]" pageId="4" pageNumber="355">
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<emphasis id="1AF3E445F22D7657FF159F23FEE0EF5D" bold="true" box="[199,345,1838,1863]" pageId="4" pageNumber="355">Description.</emphasis>
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Length of each male forewing 6.0 mm; specimens in alcohol with head, thorax, abdomen, and legs light brown. Forewings white.
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</paragraph>
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<paragraph id="28383857F22D7656FF159F7BFE67E9B7" blockId="4.[151,1436,1694,2007]" lastBlockId="5.[151,1437,151,608]" lastPageId="5" lastPageNumber="356" pageId="4" pageNumber="355">
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Male genitalia (
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<figureCitation id="B0BC24D2F22D7657FEAA9F7BFDBBEF95" box="[376,514,1910,1935]" captionStart="FIGURES 17–21" captionStartId="5.[151,264,1559,1582]" captionTargetBox="[151,1436,656,1506]" captionTargetId="figure@5.[151,1437,625,1508]" captionTargetPageId="5" captionText="FIGURES 17–21. Male genitalia of Leptocerus suwannarati n. sp. 17, left lateral; 18, dorsal; 19, ventral; 20, phallus, left lateral; 21, phallus, ventral. Inf = inferior appendage (paired), IX = segment IX, X = tergum X." httpUri="https://zenodo.org/record/2610592/files/figure.png" pageId="4" pageNumber="355">Figs. 17–21</figureCitation>
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). Segment IX in lateral view flask-like, longitudinally short (narrow) dorsally and long (expanded) ventrally, rounded basoventrally, projected apicoventrally; in dorsal view with small sharp projection anteriorly and rounded excision posteriorly; in ventral view transverse anterior margin nearly straight, posteriorly with deep V-shaped incision nearly dividing venter into two parts. Preanal appendages reduced to pair of setal patches on posterodorsal submesal margins of segment IX. Segment X divided into pair of long, slender rods, slightly diverging through most of their length and then curved ventrad and slightly asymmetrically mesad apically. Inferior appendages stout, protruding upward, fused basally with U-shaped space between them in ventral view; each with middle portion broader and with short setae on mesal margin especially in ventral view, distal portion rounded apically in lateral view, in ventral view narrow and curved slightly mesad subapically, blunt apically. Phallus tubular with long slender acute spine dorsally in lateral view, torpedo-shaped with very long slender spine in ventral view.
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</paragraph>
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</subSubSection>
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<subSubSection id="609D6BDCF22C7656FF1599BAFB7CEA45" pageId="5" pageNumber="356" type="diagnosis">
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<paragraph id="28383857F22C7656FF1599BAFB7CEA45" blockId="5.[151,1437,151,608]" pageId="5" pageNumber="356">
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<emphasis id="1AF3E445F22C7656FF1599BAFEF8E9CA" bold="true" box="[199,321,439,464]" pageId="5" pageNumber="356">Diagnosis.</emphasis>
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The male genitalia of the new species are very similar in lateral aspect to those of
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<taxonomicName id="EF8743D4F22C7656FB0499B4FE77E9EF" authority="Malicky and Chantaramongkol 1996" authorityName="Malicky and Chantaramongkol" authorityYear="1996" class="Insecta" family="Leptoceridae" genus="Leptocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="356" phylum="Arthropoda" rank="species" species="agaue">
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<emphasis id="1AF3E445F22C7656FB0499B4FB5CE9CA" box="[1238,1253,441,464]" italics="true" pageId="5" pageNumber="356">L</emphasis>
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.
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<emphasis id="1AF3E445F22C7656FB2699B4FA80E9CA" box="[1268,1337,441,464]" italics="true" pageId="5" pageNumber="356">agaue</emphasis>
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<bibRefCitation id="4C1645A6F22C7656FAED99BAFE77E9EF" author="Malicky, H. & Chantaramongkol, P." pageId="5" pageNumber="356" refId="ref3721" refString="Malicky, H. & Chantaramongkol, P. (1996) Neue Kocherfliegen aus Thailand (Trichoptera)." type="book" year="1996">Malicky and Chantaramongkol 1996</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="EF8743D4F22C7656FDD699D0FBF8E9EE" authority="Malicky and Chantaramongkol 2002" authorityName="Malicky and Chantaramongkol" authorityYear="2002" box="[516,1089,476,501]" class="Insecta" family="Leptoceridae" genus="Leptocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="356" phylum="Arthropoda" rank="species" species="ganymedes">
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<emphasis id="1AF3E445F22C7656FDD699D0FDAAE9EE" box="[516,531,477,500]" italics="true" pageId="5" pageNumber="356">L</emphasis>
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||
.
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<emphasis id="1AF3E445F22C7656FDF099D1FD26E9EE" box="[546,671,476,500]" italics="true" pageId="5" pageNumber="356">ganymedes</emphasis>
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<bibRefCitation id="4C1645A6F22C7656FD7499D1FBF8E9EE" author="Malicky, H. & Chantaramongkol, P. & Saengpradab, N & Prachuab & Chaibu, P. & Thani, I. & Changthong N & Cheunbarn, S. & Laudee, P. & Prommi, T. & Sompong, S." box="[678,1089,476,501]" pageId="5" pageNumber="356" pagination="13 - 15" refId="ref3808" refString="Malicky, H., Chantaramongkol, P., Saengpradab, N Prachuab, Chaibu, P., Thani, I., Changthong N, Cheunbarn, S., Laudee, P., Prommi, T. & Sompong, S. (2002) Neue asiatische Leptoceridae (Trichoptera) (Zugleich Arbeit Nr. 33 uber thailandische Kocherfliegen). Braueria, 29, 13 - 15. [in German]" type="journal article" year="2002">Malicky and Chantaramongkol 2002</bibRefCitation>
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||
</taxonomicName>
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(in
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||
<bibRefCitation id="4C1645A6F22C7656FBBD99D1FAF5E9EE" author="Malicky, H. & Chantaramongkol, P. & Saengpradab, N & Prachuab & Chaibu, P. & Thani, I. & Changthong N & Cheunbarn, S. & Laudee, P. & Prommi, T. & Sompong, S." box="[1135,1356,476,501]" pageId="5" pageNumber="356" pagination="13 - 15" refId="ref3808" refString="Malicky, H., Chantaramongkol, P., Saengpradab, N Prachuab, Chaibu, P., Thani, I., Changthong N, Cheunbarn, S., Laudee, P., Prommi, T. & Sompong, S. (2002) Neue asiatische Leptoceridae (Trichoptera) (Zugleich Arbeit Nr. 33 uber thailandische Kocherfliegen). Braueria, 29, 13 - 15. [in German]" type="journal article" year="2002">
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Malicky
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<emphasis id="1AF3E445F22C7656FB0199D0FABAE9EE" box="[1235,1283,476,500]" italics="true" pageId="5" pageNumber="356">et al</emphasis>
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||
. 2002
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</bibRefCitation>
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) found in
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<collectingCountry id="509078C7F22C7656FF6199F2FEA0EA02" box="[179,281,511,536]" name="Thailand" pageId="5" pageNumber="356">Thailand</collectingCountry>
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, but can be distinguished from them by the characters of segment X. In
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<taxonomicName id="EF8743D4F22C7656FBE99A0CFB24EA02" authorityName="Malicky and Chantaramongkol" authorityYear="1996" box="[1083,1181,512,536]" class="Insecta" family="Leptoceridae" genus="Leptocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="356" phylum="Arthropoda" rank="species" species="agaue">
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<emphasis id="1AF3E445F22C7656FBE99A0CFBF3EA02" box="[1083,1098,513,536]" italics="true" pageId="5" pageNumber="356">L</emphasis>
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||
.
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<emphasis id="1AF3E445F22C7656FB8A9A0CFB24EA02" box="[1112,1181,513,536]" italics="true" pageId="5" pageNumber="356">agaue</emphasis>
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</taxonomicName>
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and
|
||
<taxonomicName id="EF8743D4F22C7656FB009A0CFAD5EA02" authorityName="Malicky and Chantaramongkol" authorityYear="2002" box="[1234,1388,512,536]" class="Insecta" family="Leptoceridae" genus="Leptocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="356" phylum="Arthropoda" rank="species" species="ganymedes">
|
||
<emphasis id="1AF3E445F22C7656FB009A0CFB58EA02" box="[1234,1249,513,536]" italics="true" pageId="5" pageNumber="356">L</emphasis>
|
||
.
|
||
<emphasis id="1AF3E445F22C7656FB229A0DFAD5EA02" box="[1264,1388,512,536]" italics="true" pageId="5" pageNumber="356">ganymedes</emphasis>
|
||
</taxonomicName>
|
||
, the long slender processes of segment X are asymmetrical, but nearly symmetrical in the new species. The phallus of
|
||
<taxonomicName id="EF8743D4F22C7656FF459A44FEF9EA45" authority="Laudee & Malicky, 2018" authorityName="Laudee & Malicky" authorityYear="2018" box="[151,320,584,608]" class="Insecta" family="Leptoceridae" genus="Leptocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="356" phylum="Arthropoda" rank="species" species="suwannarati" status="sp. nov.">
|
||
<emphasis id="1AF3E445F22C7656FF459A44FF1FEA7A" box="[151,166,585,608]" italics="true" pageId="5" pageNumber="356">L</emphasis>
|
||
.
|
||
<emphasis id="1AF3E445F22C7656FF669A45FEF9EA45" box="[180,320,584,607]" italics="true" pageId="5" pageNumber="356">suwannarati</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1AF3E445F22C7656FE959A4AFE3FEA7A" bold="true" box="[327,390,583,608]" pageId="5" pageNumber="356">
|
||
<taxonomicNameLabel id="01C0593EF22C7656FE959A4AFE3FEA7A" box="[327,390,583,608]" pageId="5" pageNumber="356" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is cylindrical, but oval in
|
||
<taxonomicName id="EF8743D4F22C7656FD7E9A44FCB7EA7A" authorityName="Malicky and Chantaramongkol" authorityYear="1996" box="[684,782,584,608]" class="Insecta" family="Leptoceridae" genus="Leptocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="356" phylum="Arthropoda" rank="species" species="agaue">
|
||
<emphasis id="1AF3E445F22C7656FD7E9A44FD02EA7A" box="[684,699,585,608]" italics="true" pageId="5" pageNumber="356">L</emphasis>
|
||
.
|
||
<emphasis id="1AF3E445F22C7656FD1B9A44FCB7EA7A" box="[713,782,585,608]" italics="true" pageId="5" pageNumber="356">agaue</emphasis>
|
||
</taxonomicName>
|
||
and stout and tubular in
|
||
<taxonomicName id="EF8743D4F22C7656FBF69A44FB04EA7A" authorityName="Malicky and Chantaramongkol" authorityYear="2002" box="[1060,1213,584,608]" class="Insecta" family="Leptoceridae" genus="Leptocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="356" phylum="Arthropoda" rank="species" species="ganymedes">
|
||
<emphasis id="1AF3E445F22C7656FBF69A44FB8AEA7A" box="[1060,1075,585,608]" italics="true" pageId="5" pageNumber="356">L</emphasis>
|
||
.
|
||
<emphasis id="1AF3E445F22C7656FB939A45FB04EA7A" box="[1089,1213,584,608]" italics="true" pageId="5" pageNumber="356">ganymedes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |