treatments-xml/data/03/F7/D8/03F7D871AF795D5FFF0FABF375A1DC4F.xml
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<document id="5153DFDF8433155D2BECB5E282447F8F" ID-DOI="10.5962/bhl.part.117802" ID-ISSN="0035-418X" ID-Zenodo-Dep="6311706" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1645021050174" checkinUser="carolina" docAuthor="Cuccodoro, Giulio" docDate="2011" docId="03F7D871AF795D5FFF0FABF375A1DC4F" docLanguage="en" docName="RevSuisseZool.118.107-148.pdf.imf" docOrigin="Revue suisse de Zoologie 118 (1)" docStyle="DocumentStyle:0846B485C0C87F6EB7648DEEBF34B77F.2:RevSuisseZool.2011-2014.journal_article" docStyleId="0846B485C0C87F6EB7648DEEBF34B77F" docStyleName="RevSuisseZool.2011-2014.journal_article" docStyleVersion="2" docTitle="Megarthrus machu Cuccodoro &amp; C. &amp; P. &amp; Ch- 2011, sp. n." docType="treatment" docVersion="3" lastPageNumber="115" masterDocId="FFCEA009AF7E5D57FF98AE43766CDA37" masterDocTitle="Revision of the Neotropical types of Megarthrus Curtis, 1829 and description of two new species from Costa Rica and Peru (Coleoptera, Staphylinidae, Proteininae)" masterLastPageNumber="147" masterPageNumber="107" pageNumber="114" updateTime="1699296264638" updateUser="plazi">
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<mods:title id="B1C647CC569C38903B2C4923E609F44B">Revision of the Neotropical types of Megarthrus Curtis, 1829 and description of two new species from Costa Rica and Peru (Coleoptera, Staphylinidae, Proteininae)</mods:title>
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<mods:namePart id="1664F062F205C9EB8BF10BE05A56667E">Cuccodoro, Giulio</mods:namePart>
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<mods:typeOfResource id="3416A9DB7057DE92E7609D17F1CC1226">text</mods:typeOfResource>
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<mods:title id="F226E5DD7366DCD5EC276A77A1946D1E">Revue suisse de Zoologie</mods:title>
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<mods:part id="3A23C4238C89452EEC2ECBCDAC78A78B">
<mods:date id="FE1934B63495C8C93F0EFF60BEABBD9E">2011</mods:date>
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<mods:number id="6E97F39A8CFA1C50B0589DA11D3A9B5F">2011-03-31</mods:number>
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<mods:identifier id="F657CC325E4735A126A2FCE29BB574BC" type="DOI">10.5962/bhl.part.117802</mods:identifier>
<mods:identifier id="B334A14C1A36603D20B8B0BD5B170DF0" type="ISSN">0035-418X</mods:identifier>
<mods:identifier id="826B0D5A6C3C84D85B9F9090BE17557D" type="Zenodo-Dep">6311706</mods:identifier>
</mods:mods>
<treatment id="03F7D871AF795D5FFF0FABF375A1DC4F" LSID="urn:lsid:plazi:treatment:03F7D871AF795D5FFF0FABF375A1DC4F" httpUri="http://treatment.plazi.org/id/03F7D871AF795D5FFF0FABF375A1DC4F" lastPageId="8" lastPageNumber="115" pageId="7" pageNumber="114">
<subSubSection id="C3443AECAF795D50FF0FABF3725CDFFF" box="[151,1072,1456,1480]" pageId="7" pageNumber="114" type="nomenclature">
<paragraph id="8BE16967AF795D50FF0FABF3725CDFFF" blockId="7.[151,1074,1456,1589]" box="[151,1072,1456,1480]" pageId="7" pageNumber="114">
<heading id="D0A9DE0BAF795D50FF0FABF3725CDFFF" box="[151,1072,1456,1480]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="114" reason="2">
<emphasis id="B92AB575AF795D50FF0FABF377DCDFFF" bold="true" box="[151,432,1456,1480]" pageId="7" pageNumber="114">
<taxonomicName id="4C5E12E4AF795D50FF0FABF37701DFFF" ID-CoL="6R7VD" authority="Cuccodoro &amp; C. &amp; P. &amp; Ch-, 2011" authorityName="Cuccodoro &amp; C. &amp; P. &amp; Ch-" authorityYear="2011" box="[151,365,1456,1480]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="114" phylum="Arthropoda" rank="species" species="machu" status="sp. nov.">
<emphasis id="B92AB575AF795D50FF0FABF37701DFFF" bold="true" box="[151,365,1456,1480]" italics="true" pageId="7" pageNumber="114">Megarthrus machu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A219080EAF795D50FEECABF377DCDFFF" box="[372,432,1456,1480]" pageId="7" pageNumber="114" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Figs 11, 87-104
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3443AECAF795D50FF7BAB9774B7DC02" pageId="7" pageNumber="114" type="materials_examined">
<paragraph id="8BE16967AF795D50FF7BAB9774B7DC02" blockId="7.[151,1074,1456,1589]" pageId="7" pageNumber="114">
<collectionCode id="ED4FF1A2AF795D50FF7BAB97771BDFDF" box="[227,375,1492,1513]" pageId="7" pageNumber="114">TYPE MATERIAL</collectionCode>
:
<materialsCitation id="3B36633AAF795D50FE1EAB97757CDC35" collectingDate="1982-09-27" collectionCode="SEMC" collectorName="Watrous &amp; Mazurek" country="Peru" county="Dept." location="Manu Rd." municipality="Pillahuata" pageId="7" pageNumber="114" specimenCount="1" stateProvince="Cusco" typeStatus="holotype">
<typeStatus id="54E5D7C5AF795D50FE1EAB9777B3DFDE" box="[390,479,1492,1513]" pageId="7" pageNumber="114" type="holotype">Holotype</typeStatus>
(Ƌ, in
<collectionCode id="ED4FF1A2AF795D50FDB5AB97741FDFDE" box="[557,627,1492,1513]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="7" pageNumber="114" type="Museum">SEMC</collectionCode>
):
<collectingCountry id="F34929F7AF795D50FD1EAB9774AADFDE" box="[646,710,1492,1513]" name="Peru" pageId="7" pageNumber="114">PERU</collectingCountry>
,
<collectingRegion id="499AA785AF795D50FD4AAB977563DFDE" box="[722,783,1492,1513]" country="Peru" name="Cusco" pageId="7" pageNumber="114">Cuzco</collectingRegion>
<collectingCounty id="628011EBAF795D50FC8FAB977523DFDE" box="[791,847,1492,1513]" pageId="7" pageNumber="114">Dept.</collectingCounty>
,
<collectingMunicipality id="6B85F31DAF795D50FCC2AB9775D1DFDE" box="[858,957,1492,1513]" pageId="7" pageNumber="114">Pillahuata</collectingMunicipality>
,
<location id="8E813FBCAF795D50FC51AB977242DFDE" LSID="urn:lsid:plazi:treatment:03F7D871AF795D5FFF0FABF375A1DC4F:8E813FBCAF795D50FC51AB977242DFDE" box="[969,1070,1492,1513]" country="Peru" county="Dept." municipality="Pillahuata" name="Manu Rd." pageId="7" pageNumber="114" stateProvince="Cusco">Manu Rd.</location>
,
<location id="8E813FBCAF795D50FF0FABAE76D7DC35" LSID="urn:lsid:plazi:treatment:03F7D871AF795D5FFF0FABF375A1DC4F:8E813FBCAF795D50FF0FABAE76D7DC35" box="[151,187,1517,1538]" country="Peru" county="Dept." municipality="Pillahuata" name="Km" pageId="7" pageNumber="114" stateProvince="Cusco">Km</location>
128,
<date id="FFE04FA7AF795D50FF6AABAE7734DC35" box="[242,344,1517,1538]" pageId="7" pageNumber="114" value="1982-09-27">
<collectingDate id="EFA4B64FAF795D50FF6AABAE7734DC35" box="[242,344,1517,1538]" pageId="7" pageNumber="114" value="1982-09-27">27.ix.1982</collectingDate>
</date>
(
<collectorName id="26AB0CB1AF795D50FEFDABAE77DADC35" box="[357,438,1517,1538]" pageId="7" pageNumber="114">Watrous</collectorName>
&amp;
<collectorName id="26AB0CB1AF795D50FE4DABAE7440DC35" box="[469,556,1517,1538]" pageId="7" pageNumber="114">Mazurek</collectorName>
) #82-308, ex vine litter
</materialsCitation>
.
<materialsCitation id="3B36633AAF795D50FCB3ABAE77E3DC2C" collectingDate="1982-09-27" collectionCode="FMNH" collectorName="Watrous &amp; Mazurek" country="Peru" county="Dept." location="Manu Rd." municipality="Pillahuata" pageId="7" pageNumber="114" specimenCount="1" specimenCount-female="1" stateProvince="Cusco" typeStatus="paratype">
<typeStatus id="54E5D7C5AF795D50FCB3ABAE75E4DC35" box="[811,904,1517,1538]" pageId="7" pageNumber="114" type="paratype">Paratypes</typeStatus>
(4): same data as holotype,
<specimenCount id="9D58A2EEAF795D50FF62A845774FDC2C" box="[250,291,1541,1563]" pageId="7" pageNumber="114" type="female">1 ♀</specimenCount>
in
<collectionCode id="ED4FF1A2AF795D50FEDFA84577E3DC2C" box="[327,399,1542,1563]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="7" pageNumber="114" type="Museum">FMNH</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B36633AAF795D50FE03A8457241DC2C" box="[411,1069,1541,1563]" collectingDate="1982-09-26" collectionCode="FMNH, MHNG" collectorName="Watrous &amp; Mazurek" country="Peru" county="Dept." location="Manu Rd." municipality="Pillahuata" pageId="7" pageNumber="114" specimenCount="2" specimenCount-female="2" stateProvince="Cusco" typeStatus="paratype">
same data, but
<date id="FFE04FA7AF795D50FDABA84574F0DC2C" box="[563,668,1542,1563]" pageId="7" pageNumber="114" value="1982-09-26">
<collectingDate id="EFA4B64FAF795D50FDABA84574F0DC2C" box="[563,668,1542,1563]" pageId="7" pageNumber="114" value="1982-09-26">26.ix.1982</collectingDate>
</date>
, #82-301,
<specimenCount id="9D58A2EEAF795D50FC9FA8457528DC2C" box="[775,836,1541,1563]" pageId="7" pageNumber="114" type="female">2 ♀♀</specimenCount>
in
<collectionCode id="ED4FF1A2AF795D50FCF0A84575C1DC2C" box="[872,941,1542,1563]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="7" pageNumber="114" type="Museum">FMNH</collectionCode>
and
<collectionCode id="ED4FF1A2AF795D50FC79A8457241DC2C" box="[993,1069,1542,1563]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="7" pageNumber="114" type="Museum">MHNG</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B36633AAF795D50FF0FA86374BBDC02" box="[151,727,1566,1589]" collectingDate="1982-09-25" collectionCode="FMNH" collectorName="Watrous &amp; Mazurek" country="Peru" county="Dept." location="Manu Rd." municipality="Pillahuata" pageId="7" pageNumber="114" specimenCount="1" specimenCount-female="1" stateProvince="Cusco" typeStatus="paratype">
same data, but
<date id="FFE04FA7AF795D50FEB1A86377FDDC02" box="[297,401,1568,1589]" pageId="7" pageNumber="114" value="1982-09-25">
<collectingDate id="EFA4B64FAF795D50FEB1A86377FDDC02" box="[297,401,1568,1589]" pageId="7" pageNumber="114" value="1982-09-25">25.ix.1982</collectingDate>
</date>
, #82-290, on tent,
<specimenCount id="9D58A2EEAF795D50FDD0A8637403DC03" box="[584,623,1566,1589]" pageId="7" pageNumber="114" type="female">1 ♀</specimenCount>
in
<collectionCode id="ED4FF1A2AF795D50FD16A86374BBDC02" box="[654,727,1568,1589]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="7" pageNumber="114" type="Museum">FMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3443AECAF795D5FFF7BA8067454DF50" lastPageId="8" lastPageNumber="115" pageId="7" pageNumber="114" type="description">
<paragraph id="8BE16967AF795D50FF7BA806725DDC6B" blockId="7.[151,1073,1605,1660]" box="[227,1073,1605,1628]" pageId="7" pageNumber="114">DESCRIPTION: Habitus as in Fig. 11. Combined length of pronotum and elytra =</paragraph>
<paragraph id="8BE16967AF795D5FFF0FA82674E6DBC1" blockId="7.[151,1073,1605,1660]" lastBlockId="8.[136,1059,227,1383]" lastPageId="8" lastPageNumber="115" pageId="7" pageNumber="114">
<quantity id="4CA6C482AF795D50FF0FA826777ADC4B" box="[151,278,1637,1660]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" metricValueMax="2.3" metricValueMin="2.1" pageId="7" pageNumber="114" unit="mm" value="2.2" valueMax="2.3" valueMin="2.1">2.1-2.3 mm</quantity>
; maximal pronotal width =
<quantity id="4CA6C482AF795D50FDA1A82674DBDC4B" box="[569,695,1637,1660]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="7" pageNumber="114" unit="mm" value="1.55" valueMax="1.6" valueMin="1.5">1.5-1.6 mm</quantity>
. Body dark brown with appendages and elytra slightly paler; antennomeres 10-11 paler than antennomeres 1-9. Dorsal pubescence fairly uniform, denser on head and pronotum than on elytral disc; frontal setae directed forward; elytral and pronotal setae fairly straight, recumbent; metasternal pubescence fairly uniform, shorter than prosternal; pubescence on abdominal tergites parallel, becoming denser near posterior margin of tergite VII. Frons, humeral callus of elytra and anterior portion of prohypomera granulate; frontal granulation conspicuous, with granula about as high as their diameter, or higher; pronotum granulofossulate; metasternum with granulofossulation becoming denser and finer posteriomedially; elytral disc coarsely punctate.
</paragraph>
<paragraph id="8BE16967AF765D5FFF4CAFBC724FD843" blockId="8.[136,1059,227,1383]" pageId="8" pageNumber="115">Frons forming above clypeus a sharp ridge. the latter finely evenly carinate, weakly arcuate in middle and laterally oblique in dorsal view; mesal portion of disc strongly evenly convex lateral view; U-shaped frontal impression deep. Temples strongly convex in dorsal view. Antennae (Fig. 98) 2.3-2.6 times longer than pronotum.</paragraph>
<paragraph id="8BE16967AF765D5FFF4CAC3F741CD906" blockId="8.[136,1059,227,1383]" pageId="8" pageNumber="115">Pronotum (Fig. 97) with center strongly convex in frontal view; disc deeply depressed near middle of lateral edge, shallowly depressed along anterior and posterior margins; medial groove slightly arcuate in lateral view, deep, parallel-sided; hypomera ridged from anterior margin to laterobasal angle, disc without pit. Prosternal medial ridge entire. Scutellum with anterior margin angulate in middle and posterior margin slighly arcuate toward obtusely angular apex.</paragraph>
<paragraph id="8BE16967AF765D5FFF4CAD7A7547D9B8" blockId="8.[136,1059,227,1383]" pageId="8" pageNumber="115">Elytra abruptly widened subbasally (Fig. 11); humeral callus low, moderately convex; disc with low swellings, deeply depressed along lateral edge; the latter finely carinate, markedly denticulate, strongly arcuate in dorsal view.</paragraph>
<paragraph id="8BE16967AF765D5FFF4CADDB74C5D9F9" blockId="8.[136,1059,227,1383]" pageId="8" pageNumber="115">Abdominal sternites II and III with medial processes as in Fig. 96, posterior portion of process of sternite III widened, or trifid.</paragraph>
<paragraph id="8BE16967AF765D5FFF4CAD947492DEDE" blockId="8.[136,1059,227,1383]" pageId="8" pageNumber="115">Male: Frontoclypeal area not modified. Protarsomeres 1 lacking tenent setae. Mesofemora (Fig. 90) as long as metafemora (Fig. 89). Mesotibiae (Fig. 94) shorter than metatibiae (Fig. 95). Metatarsomeres 1 about as long as combined length of metatarsomeres 2-4. Peg-like setae arranged in a single row on mesotrochanters (Fig. 90), grouped in a field on mesotibiae and metatibiae, and absent from protrochanters, profemora, protibiae, mesofemora, metatrochanters (Fig. 89) and metafemora. Pubescence on sternites IV-VII becoming denser posteriomedially. Apex of abdominal tergite VIII as in Figs 91, 93. Sternite VIII as in Fig. 92. Sternite IX lacking subbasal protuberance. Aedeagus as in Figs 87-88.</paragraph>
<paragraph id="8BE16967AF765D5FFF4CAAB17454DF50" blockId="8.[136,1059,227,1383]" pageId="8" pageNumber="115">Female: Pubescence on sternites IV-VII uniform, except for a pair of subapical macrosetae on each sternite. Abdominal tergite VIII (Figs 102-103) without medioapical projection. Sternite VIII as in Fig. 104. Genital segment as in Figs 99-101; gonocoxal plate bearing a mediodorsal ridge.</paragraph>
</subSubSection>
<subSubSection id="C3443AECAF765D5FFF4CAB39755DDFC7" pageId="8" pageNumber="115" type="distribution">
<paragraph id="8BE16967AF765D5FFF4CAB39755DDFC7" blockId="8.[136,1059,1402,1520]" pageId="8" pageNumber="115">
<materialsCitation id="3B36633AAF765D5FFF4CAB397541DFC7" country="Peru" elevation="2500" latitude="-3.1166666" location="Peru" longLatPrecision="1307" longitude="-71.416664" pageId="8" pageNumber="115" specimenCount="1">
DISTRIBUTION AND NATURAL HISTORY:
<taxonomicName id="4C5E12E4AF765D5FFDFBAB387543DFA5" authorityName="Cuccodoro &amp; C. &amp; P. &amp; Ch-" authorityYear="2011" box="[611,815,1403,1426]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="115" phylum="Arthropoda" rank="species" species="machu">
<emphasis id="B92AB575AF765D5FFDFBAB387543DFA5" box="[611,815,1403,1426]" italics="true" pageId="8" pageNumber="115">Megarthrus machu</emphasis>
</taxonomicName>
is know only from the
<typeStatus id="54E5D7C5AF765D5FFF10ABD976DADF86" box="[136,182,1434,1457]" pageId="8" pageNumber="115">type</typeStatus>
locality in
<collectingCountry id="F34929F7AF765D5FFEB5ABD97732DF86" box="[301,350,1434,1457]" name="Peru" pageId="8" pageNumber="115">Peru</collectingCountry>
(Pillahuata &lt;
<geoCoordinate id="EE6A0FA0AF765D5FFE77ABD97459DF86" box="[495,565,1434,1457]" degrees="3" direction="south" minutes="7" orientation="latitude" pageId="8" pageNumber="115" precision="925" value="-3.1166666">3° 7' S</geoCoordinate>
;
<geoCoordinate id="EE6A0FA0AF765D5FFDD8ABD974C1DF86" box="[576,685,1434,1457]" degrees="71" direction="west" minutes="25" orientation="longitude" pageId="8" pageNumber="115" precision="925" value="-71.416664">71° 25' W</geoCoordinate>
&gt;), which is situated at an elevation of nearly
<quantity id="4CA6C482AF765D5FFF6EABFA7726DFE7" box="[246,330,1465,1488]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="115" unit="m" value="2500.0">
<elevation id="00738E54AF765D5FFF6EABFA7726DFE7" box="[246,330,1465,1488]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="115" unit="m" value="2500.0">2500 m</elevation>
</quantity>
just below a patch of forest. It was found in vine leaf litter, and showed good ability to fly (
<specimenCount id="9D58A2EEAF765D5FFE29AB9A742ADFC7" box="[433,582,1497,1520]" pageId="8" pageNumber="115" type="generic">one specimen</specimenCount>
was found on a tent)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3443AECAF765D5FFF4CA84075A1DC4F" pageId="8" pageNumber="115" type="discussion">
<paragraph id="8BE16967AF765D5FFF4CA84075A1DC4F" blockId="8.[136,1059,1539,1656]" pageId="8" pageNumber="115">
COMMENTS: Within Neotropical
<taxonomicName id="4C5E12E4AF765D5FFDA9A84074C3DC2D" authorityName="Curtis" authorityYear="1829" box="[561,687,1539,1562]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis id="B92AB575AF765D5FFDA9A84074C3DC2D" box="[561,687,1539,1562]" italics="true" pageId="8" pageNumber="115">Megarthrus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5E12E4AF765D5FFD26A8407546DC2D" authorityName="Cuccodoro &amp; C. &amp; P. &amp; Ch-" authorityYear="2011" box="[702,810,1539,1562]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="115" phylum="Arthropoda" rank="species" species="machu">
<emphasis id="B92AB575AF765D5FFD26A8407546DC2D" box="[702,810,1539,1562]" italics="true" pageId="8" pageNumber="115">M. machu</emphasis>
</taxonomicName>
is easily distinguished by its abruptly and broadly expanded elytra. Exceeding
<quantity id="4CA6C482AF765D5FFC80A8617531DC0E" box="[792,861,1570,1593]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="115" unit="mm" value="4.0">4 mm</quantity>
in length (when measured from the middle of the frontal margin to the tip of the abdomen), it is the largest species of
<taxonomicName id="4C5E12E4AF765D5FFED0A82277ADDC4F" authorityName="Erichson" authorityYear="1839" box="[328,449,1633,1656]" pageId="8" pageNumber="107" rank="subFamily" subFamily="Proteininae">Proteininae</taxonomicName>
of the world. See comments under
<taxonomicName id="4C5E12E4AF765D5FFCD9A82275A9DC4F" authorityName="Bierig" authorityYear="2011" box="[833,965,1633,1656]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="115" phylum="Arthropoda" rank="species" species="adelphus">
<emphasis id="B92AB575AF765D5FFCD9A82275A9DC4F" box="[833,965,1633,1656]" italics="true" pageId="8" pageNumber="115">M. adelphus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>