944 lines
161 KiB
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944 lines
161 KiB
XML
<document id="B95C4E13985EAD7C2D26C6F56808267C" ID-DOI="10.5852/ejt.2024.947.2625" ID-ISSN="2118-9773" ID-Zenodo-Dep="13310019" ID-ZooBank="CBC9DA40-EC6D-4280-8BBC-6826D72A291E" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="julia" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1723540771803" checkinUser="plazi" docAuthor="Humara-Gil, Karla J., Granja-Fernández, Rebeca, Bautista-Guerrero, Eric, Solís-Marín, Francisco A. & Rodríguez-Troncoso, Alma P." docDate="2024" docId="03F3204BFF91820155CDB21C612DFA7B" docLanguage="en" docName="EJT.2024.947.130-174.pdf" docOrigin="European Journal of Taxonomy 947" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2625/12067" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ophioderma aija Humara-Gil, Granja-Fernández, Bautista-Guerrero, Solís-Marín & Rodríguez-Troncoso, 2024, sp. nov." docType="treatment" docVersion="3" lastPageNumber="159" masterDocId="FFCA5833FF89821C574EB3146569FFB7" masterDocTitle="Delimitation of Ophioderma teres (Lyman, 1860) and Ophioderma unicolor H. L. Clark, 1940 stat. nov. (Echinodermata: Ophiuroidea), including the description of two new species" masterLastPageNumber="174" masterPageNumber="130" pageNumber="154" updateTime="1725019940605" updateUser="julia" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F844480D5C36BB73C4AB3B8371B46961">Delimitation of Ophioderma teres (Lyman, 1860) and Ophioderma unicolor H. L. Clark, 1940 stat. nov. (Echinodermata: Ophiuroidea), including the description of two new species</mods:title>
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<mods:namePart id="9E40FB6ACDB4D3A01E6E60BF4DE9581F">Humara-Gil, Karla J.</mods:namePart>
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<mods:affiliation id="99E5586FA16A6E850B5A869B7191C846">Laboratorio de Ecología Marina, Centro Universitario de la Costa, Universidad de Guadalajara, Puerto Vallarta, Mexico.</mods:affiliation>
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<mods:nameIdentifier id="CD2DF928D54DCB27336C757CE2B7E22D" type="email">k.humaragil@gmail.com</mods:nameIdentifier>
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<mods:namePart id="AB84A96D789712E868A4D0C6126CD7D2">Granja-Fernández, Rebeca</mods:namePart>
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<mods:affiliation id="5B19D3DAE55EA0EE375F463A3ED71C83">Postdoctoral researcher (CONAHCYT) associated with Programa de Maestría en Biosistemática y Manejo de Recursos Naturales y Agrícolas (BIMARENA) / Laboratorio de Ecología Molecular, Microbiología y Taxonomía (LEMITAX). Centro Universitario de Ciencias Biológicas y Agropecuarias, Universidad de Guadalajara, Zapopan, Mexico.</mods:affiliation>
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<mods:nameIdentifier id="2B15F23ED5BE03FCB87EE061DC66D0D8" type="email">beckygranja@gmail.com</mods:nameIdentifier>
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<mods:namePart id="4CFF44ACEC9C4CBCF4CCAF02DFF689A7">Bautista-Guerrero, Eric</mods:namePart>
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<mods:affiliation id="4F90ACB969C81D5F3E03C1042CE1D162">Laboratorio de Ecología Marina, Centro Universitario de la Costa, Universidad de Guadalajara, Puerto Vallarta, Mexico.</mods:affiliation>
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<mods:namePart id="7B70BD64B42D8C2AD8DAA5BA230D03AF">Solís-Marín, Francisco A.</mods:namePart>
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<mods:affiliation id="B6CE7DD8985E2F94E88F30BB1964FD33">Colección Nacional de Equinodermos “ Dra. Ma. Elena Caso Muñoz ”, Laboratorio de Sistemática y Ecología de Equinodermos, Instituto de Ciencias del Mar y Limnología (ICML), Universidad Nacional Autónoma de México (UNAM), Mexico City, Mexico.</mods:affiliation>
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<mods:nameIdentifier id="2B373A8BCB27C2008429067EAF70B1FD" type="email">fasolis@cmarl.unam.mx</mods:nameIdentifier>
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<mods:namePart id="0D3E37A210D9C4323EAAA6C9FBBABC32">Rodríguez-Troncoso, Alma P.</mods:namePart>
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<mods:affiliation id="7D9D69004AB2E4EC78A61B3968A76C79">Laboratorio de Ecología Marina, Centro Universitario de la Costa, Universidad de Guadalajara, Puerto Vallarta, Mexico.</mods:affiliation>
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<mods:nameIdentifier id="91800A1B15C3E714338E242F93C56B70" type="email">alma.rtroncoso@academicos.udg.mx</mods:nameIdentifier>
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<mods:title id="F22B654857F0642FD81BF8D054C3F358">European Journal of Taxonomy</mods:title>
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<mods:date id="967244B6BF4E8BF869143D4CF6FA289F">2024</mods:date>
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<taxonomicName id="4C5AEADEFF91820455CDB21C6639FE94" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" box="[643,848,264,291]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="24" pageNumber="154" phylum="Echinodermata" rank="species" species="aija" status="sp. nov.">
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<emphasis id="B92E4D4FFF91820455CDB21C6639FE94" bold="true" box="[643,848,264,291]" italics="true" pageId="24" pageNumber="154">Ophioderma aija</emphasis>
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<paragraph id="8BE5915DFF918204551AB25B66B6FEDE" blockId="24.[370,1217,264,362]" box="[596,991,335,362]" pageId="24" pageNumber="154">
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<figureCitation id="13618DD8FF918204551AB25B67D5FEDE" box="[596,700,335,361]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="8.[189,232,1892,1918]" captionStartId-1="9.[189,232,1928,1954]" captionTargetBox-0="[224,1365,769,1835]" captionTargetBox-1="[222,1365,1013,1864]" captionTargetId-0="figure-204@8.[189,1398,747,1851]" captionTargetId-1="figure-223@9.[189,1398,1000,1886]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Fig. 1. Bayesian inference (BI) tree of COI sequences. Numbers on branches indicate posterior probability/bootstrap values obtained with BI and maximum likelihood (ML), respectively. A dash (–) indicates the branches not recovered in the ML analysis. Sequences obtained in this study are indicated in bold (see Supp. file 5)." captionText-1="Fig. 2. Bayesian inference (BI) tree of 16Ssequences. Numberson branches indicate posterior probability/ bootstrap values obtained with BI and maximum likelihood, respectively. Sequences obtained in this study are indicated in bold (see Supp. file 5)." figureDoi-0="http://doi.org/10.5281/zenodo.13310021" figureDoi-1="http://doi.org/10.5281/zenodo.13310023" httpUri-0="https://zenodo.org/record/13310021/files/figure.png" httpUri-1="https://zenodo.org/record/13310023/files/figure.png" pageId="24" pageNumber="154">Figs 1–2</figureCitation>
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,
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<figureCitation id="13618DD8FF9182045586B25B67B1FEDD" box="[712,728,335,362]" captionStart="Fig" captionStartId="17.[189,232,1892,1918]" captionTargetBox="[258,1329,281,1836]" captionTargetId="figure-18@17.[239,1348,265,1851]" captionTargetPageId="17" captionText="Fig. 5. Distribution of Ophioderma teres (Lyman, 1860), Ophioderma unicolor H.L. Clark, 1940 stat. nov., Ophioderma aija sp. nov., and Ophioderma bichi sp. nov. in the eastern Pacific. A. Gulf of California. B. Revillagigedo Islands. C. Mexican Tropical Pacific. D. Central America and Colombia. E. Galapagos Islands." figureDoi="http://doi.org/10.5281/zenodo.13310029" httpUri="https://zenodo.org/record/13310029/files/figure.png" pageId="24" pageNumber="154">5</figureCitation>
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,
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<figureCitation id="13618DD8FF91820455A8B25B667DFEDE" box="[742,788,335,361]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="26.[189,232,1787,1813]" captionStartId-1="28.[189,232,1716,1742]" captionTargetBox-0="[319,1268,263,1746]" captionTargetBox-1="[344,1244,265,1671]" captionTargetId-0="figure-12@26.[319,1268,263,1746]" captionTargetId-1="figure-12@28.[344,1244,265,1671]" captionTargetPageId-0="26" captionTargetPageId-1="28" captionText-0="Fig. 8. Ophioderma aija sp. nov., holotype (ICML-UNAM 18466, DD = 26.7 mm). A. Dorsal disc. B. Ventral disc (inset: section between the arm and the distal genital slit). C. Close-up of a jaw. D. Proximalmost dorsal arm. E. Median dorsal arm. F. Distalmost dorsal arm. G. Proximalmost ventral arm. H. Median ventral arm. I. Distalmost ventral arm. J. Lateral arm plates and arm spines. K. Dorsal view. Abbreviations: 2° AdShSp = secondary adoral shield spine; AdShSp = adoral shield spine; IPa = infradental papilla; LOPas = lateral oral papillae; LyOs = Lyman’s ossicle; OPRSp = oral plate ridge spine; RS = radial shield; TPa = tooth papilla; vT = ventralmost tooth." captionText-1="Fig. 9. Ophioderma aija sp. nov., non-type specimen (ICML-UNAM 18457, DD = 26.2 mm), SEM images of the ossicles. A. Radial shield, external face. The arrowhead indicates the naked section observed in the intact animal. B. Radial shield, internal face. C. Dental plate. D. Adradial genital plate, abradial face. E. Adradial genital plate, adradial face. F. Oral plate, abradial face. G. Oral plate, adradial face. Black arrows point to oral papillae sockets, white arrows point to granule sockets. H. Arm vertebra, proximal face. I. Arm vertebra, distal face. J. Ventral arm plate, external face. K. Ventral arm plate, internal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. N. Lateral arm plate, lateral face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; SA = spine articulation; Sp = spur; vent = ventral." figureDoi-0="http://doi.org/10.5281/zenodo.13310037" figureDoi-1="http://doi.org/10.5281/zenodo.13310039" httpUri-0="https://zenodo.org/record/13310037/files/figure.png" httpUri-1="https://zenodo.org/record/13310039/files/figure.png" pageId="24" pageNumber="154">8–9</figureCitation>
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,
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<figureCitation id="13618DD8FF918204546EB25B663CFEDE" box="[800,853,335,361]" captionStart="Fig" captionStartId="35.[189,232,1892,1918]" captionTargetBox="[287,1301,265,1849]" captionTargetId="figure-17@35.[287,1301,265,1849]" captionTargetPageId="35" captionText="Fig. 12. Photographs of the species in situ. A. Ophioderma teres (Lyman, 1860) (Bajo Rojo, Costa Rica) (photograph: Steven Lara-Angulo). B. Ophioderma aija sp. nov. (Estacahuite, Mexico) (photograph: Francisco Benítez-Villalobos). C. Ophioderma bichi sp. nov. (Puerto Peñasco, Mexico) (photograph: Paloma Valdivia)." figureDoi="http://doi.org/10.5281/zenodo.13310047" httpUri="https://zenodo.org/record/13310047/files/figure.png" pageId="24" pageNumber="154">12B</figureCitation>
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;
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<tableCitation id="C6D8A4E6FF918204542EB25B66B6FEDE" box="[864,991,335,361]" captionStart-0="Table 1" captionStart-1="Table 2" captionStart-2="Table 3" captionStart-3="Table 4" captionStartId-0="5.[185,255,655,683]" captionStartId-1="6.[186,256,582,610]" captionStartId-2="9.[189,259,265,291]" captionStartId-3="10.[185,255,645,673]" captionTargetBox-1="[197,1897,838,1702]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="Table 1. Main external and internal morphological characters that distinguish the studied species/morphotypes of Ophioderma Müller & Troschel, 1840. The number of DAP pieces is given as an interval, followed by the mean in parentheses. A dash (–) indicates no data are available due to lack of material. Abbreviations: AL = arm length; DAP = dorsal arm plate; DD = disc diameter." captionText-1="Table 2. Morphometric data with the highest contribution to the differentiation of the studied species/morphotypes of Ophioderma Müller & Troschel, 1840. The ratios and counts are indicated as intervals, followed by the mean in parentheses.Abbreviations: 1VAP_W:L = first ventral arm plate (VAP) width:length ratio; AS_prox_3, 5 = number of arm spines in the 3rd or 5th proximal arm segment; DAPs_med_1, 2, 3 = number of DAP pieces in the 1st, 2nd or 3rd median arm segment; DAPs_prox_3, 4, 5 = number of dorsal arm plate (DAP) pieces in the 3rd, 4th or 5th proximal arm segment; DD = disc diameter; LOPas = lateral oral papillae; OSh_W:L = oral shield width:length ratio; VAP_L:W_med = VAP length:width ratio (median arm section); VAP_pores = number of arm segments with pores present between the proximalmost VAPs." captionText-2="Table 3. Mean genetic distances (%) between the analyzed species/morphotypes of Ophioderma Müller & Troschel, 1840 and Ophiopeza Peters, 1851 (COI, lower left; 16S, upper right). A dash (–) indicates no data are available due to a lack of sequences." captionText-3="Table 4. Summary of the comparisons of the analyzed evidence by pairs of species/morphotypes of Ophioderma Müller & Troschel, 1840. A dash (–) indicates no data are available for one or both species/morphotypes compared." pageId="24" pageNumber="154">Tables 1–4</tableCitation>
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<taxonomicName id="4C5AEADEFF91820457F3B28564FFFE1C" ID-CoL="49WYH" baseAuthorityName="H. L. Clark" baseAuthorityYear="1940" box="[189,406,401,427]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="24" pageNumber="154" phylum="Echinodermata" rank="species" species="teres">
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<emphasis id="B92E4D4FFF91820457F3B28564FFFE1C" box="[189,406,401,427]" italics="true" pageId="24" pageNumber="154">Ophioderma teres</emphasis>
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–
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<bibRefCitation id="EFCBECACFF91820456F3B28567FFFE1B" author="Lyman" box="[445,662,401,428]" firstAuthor="Lyman" page="258" pageId="24" pageNumber="154" pagination="193 - 204" refId="ref25910" refString="Lyman T. 1860. Descriptions of new Ophiuridae, belonging to the Smithsonian Institution and to the Museum of Comparative Zoology at Cambridge. Proceedings of the Boston Society of Natural History 7: 193 - 204, 252 - 262. https: // doi. org / 10.5962 / bhl. part. 4822" type="journal article" year="1860">Lyman 1860: 258</bibRefCitation>
|
||
</treatmentCitation>
|
||
(partim, non
|
||
<bibRefCitation id="EFCBECACFF918204540AB2856680FE1C" author="Lyman" box="[836,1001,401,427]" firstAuthor="Lyman" pageId="24" pageNumber="154" pagination="193 - 204" refId="ref25910" refString="Lyman T. 1860. Descriptions of new Ophiuridae, belonging to the Smithsonian Institution and to the Museum of Comparative Zoology at Cambridge. Proceedings of the Boston Society of Natural History 7: 193 - 204, 252 - 262. https: // doi. org / 10.5962 / bhl. part. 4822" type="journal article" year="1860">Lyman, 1860</bibRefCitation>
|
||
); 1865: 38 (partim, non
|
||
<bibRefCitation id="EFCBECACFF9182045250B2856440FE79" author="Lyman" firstAuthor="Lyman" pageId="24" pageNumber="154" pagination="193 - 204" refId="ref25910" refString="Lyman T. 1860. Descriptions of new Ophiuridae, belonging to the Smithsonian Institution and to the Museum of Comparative Zoology at Cambridge. Proceedings of the Boston Society of Natural History 7: 193 - 204, 252 - 262. https: // doi. org / 10.5962 / bhl. part. 4822" type="journal article" year="1860">Lyman, 1860</bibRefCitation>
|
||
). —
|
||
<treatmentCitation id="0AFBB74CFF918204562EB2A06664FE79" author="Steinbeck J. & Ricketts E. F." box="[352,781,436,462]" page="391" pageId="24" pageNumber="154" year="1941">
|
||
<bibRefCitation id="EFCBECACFF918204562EB2A06664FE79" author="Steinbeck & Ricketts" box="[352,781,436,462]" firstAuthor="Steinbeck" page="391 - 392" pageId="24" pageNumber="154" refId="ref26778" refString="Steinbeck J. & Ricketts E. F. 1941. Sea of Cortez. A Leisurely Journal of Travel and Research. Viking Press, New York." type="book" year="1941">Steinbeck & Ricketts 1941: 391–392</bibRefCitation>
|
||
</treatmentCitation>
|
||
(partim, non
|
||
<bibRefCitation id="EFCBECACFF91820454E4B2A16121FE79" author="Lyman" box="[938,1096,436,463]" firstAuthor="Lyman" pageId="24" pageNumber="154" pagination="193 - 204" refId="ref25910" refString="Lyman T. 1860. Descriptions of new Ophiuridae, belonging to the Smithsonian Institution and to the Museum of Comparative Zoology at Cambridge. Proceedings of the Boston Society of Natural History 7: 193 - 204, 252 - 262. https: // doi. org / 10.5962 / bhl. part. 4822" type="journal article" year="1860">Lyman, 1860</bibRefCitation>
|
||
). —
|
||
<treatmentCitation id="0AFBB74CFF9182045330B2A0601BFE78" author="Caso M. E." box="[1150,1394,436,463]" page="258" pageId="24" pageNumber="154" year="1951">
|
||
<bibRefCitation id="EFCBECACFF9182045330B2A0601BFE78" author="Caso" box="[1150,1394,436,463]" firstAuthor="Caso" page="258 - 265" pageId="24" pageNumber="154" pagination="219 - 312" refId="ref24399" refString="Caso M. E. 1951. Contribucion al conocimiento de los ofiuroideos de Mexico. I. Algunas especies de ofiuroideos litorales. Anales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico 22 (1): 219 - 312." type="journal article" year="1951">Caso 1951: 258–265</bibRefCitation>
|
||
</treatmentCitation>
|
||
, figs 21–24 (partim, non
|
||
<bibRefCitation id="EFCBECACFF9182045548B2CC67CCFE46" author="Lyman" box="[518,677,471,498]" firstAuthor="Lyman" pageId="24" pageNumber="154" pagination="193 - 204" refId="ref25910" refString="Lyman T. 1860. Descriptions of new Ophiuridae, belonging to the Smithsonian Institution and to the Museum of Comparative Zoology at Cambridge. Proceedings of the Boston Society of Natural History 7: 193 - 204, 252 - 262. https: // doi. org / 10.5962 / bhl. part. 4822" type="journal article" year="1860">Lyman, 1860</bibRefCitation>
|
||
). —
|
||
<treatmentCitation id="0AFBB74CFF9182045590B2C36142FE46" author="Ziesenhenne F. C." box="[734,1067,471,498]" page="190" pageId="24" pageNumber="154" year="1955">
|
||
<bibRefCitation id="EFCBECACFF9182045590B2C36142FE46" author="Ziesenhenne" box="[734,1067,471,498]" firstAuthor="Ziesenhenne" page="190 - 191" pageId="24" pageNumber="154" pagination="185 - 201" refId="ref27266" refString="Ziesenhenne F. C. 1955. A review of the genus Ophioderma Muller and Troschel. In: Allan Hancock Foundation for Scientific Research (eds) Essays in the Natural Sciences in Honor of Captain Allan Hancock: 185 - 201. University of Southern California Press, Los Angeles. https: // doi. org / 10.5962 / bhl. title. 608" type="book chapter" year="1955">Ziesenhenne 1955: 190–191</bibRefCitation>
|
||
</treatmentCitation>
|
||
(partim, non
|
||
<bibRefCitation id="EFCBECACFF9182045384B2CC6003FE46" author="Lyman" box="[1226,1386,471,498]" firstAuthor="Lyman" pageId="24" pageNumber="154" pagination="193 - 204" refId="ref25910" refString="Lyman T. 1860. Descriptions of new Ophiuridae, belonging to the Smithsonian Institution and to the Museum of Comparative Zoology at Cambridge. Proceedings of the Boston Society of Natural History 7: 193 - 204, 252 - 262. https: // doi. org / 10.5962 / bhl. part. 4822" type="journal article" year="1860">Lyman, 1860</bibRefCitation>
|
||
). —
|
||
<treatmentCitation id="0AFBB74CFF918204565FB2EE6612FDA2" author="Granja-Fernandez M. R. & Lopez-Perez R. A." box="[273,891,506,533]" page="1321" pageId="24" pageNumber="154" year="2011">
|
||
<bibRefCitation id="EFCBECACFF918204565FB2EE6612FDA2" author="Granja-Fernandez & Lopez-Perez" box="[273,891,506,533]" firstAuthor="Granja-Fernandez" page="1321 - 1322" pageId="24" pageNumber="154" pagination="1320 - 1323" refId="ref24835" refString="Granja-Fernandez M. R. & Lopez-Perez R. A. 2011. Nuevos registros de ofiuroideos (Echinodermata: Ophiuroidea) para localidades de Zihuatanejo (Guerrero) y Puerto Escondido (Oaxaca), Pacifico Mexicano. Revista Mexicana de Biodiversidad 82: 1320 - 1323. https: // doi. org / 10.22201 / ib. 20078706 e. 2011.4.675" type="journal article" year="2011">Granja-Fernández & López-Pérez 2011: 1321–1322</bibRefCitation>
|
||
</treatmentCitation>
|
||
(non
|
||
<bibRefCitation id="EFCBECACFF918204548AB2EF610FFDA3" author="Lyman" box="[964,1126,506,533]" firstAuthor="Lyman" pageId="24" pageNumber="154" pagination="193 - 204" refId="ref25910" refString="Lyman T. 1860. Descriptions of new Ophiuridae, belonging to the Smithsonian Institution and to the Museum of Comparative Zoology at Cambridge. Proceedings of the Boston Society of Natural History 7: 193 - 204, 252 - 262. https: // doi. org / 10.5962 / bhl. part. 4822" type="journal article" year="1860">Lyman, 1860</bibRefCitation>
|
||
). —
|
||
<treatmentCitation id="0AFBB74CFF91820453EDB2EE6489FD8F" author="Granja-Fernandez R. & Herrero-Perezrul M. D. & Lopez-Perez R. A. & Hernandez L. & Rodriguez-Zaragoza F. A. & Jones R. W. & Pineda-Lopez R." httpUri="http://treatment.plazi.org/id/3DFFEE5952EEC94C5A3EA87EA42B6F71" page="134" pageId="24" pageNumber="154" year="2014">
|
||
<bibRefCitation id="EFCBECACFF91820453EDB2EE6489FD8F" author="Granja-Fernandez" etAl="et al." firstAuthor="Granja-Fernandez" page="134 - 135" pageId="24" pageNumber="154" pagination="101 - 145" refId="ref24943" refString="Granja-Fernandez R., Herrero-Perezrul M. D., Lopez-Perez R. A., Hernandez L., Rodriguez-Zaragoza F. A., Jones R. W. & Pineda-Lopez R. 2014. Ophiuroidea (Echinodermata) from coral reefs in the Mexican Pacific. ZooKeys 406: 101 - 145. https: // doi. org / 10.3897 / zookeys. 406.6306" type="journal article" year="2014">
|
||
Granja-Fernández
|
||
<emphasis id="B92E4D4FFF91820457A2B10A644EFD8F" box="[236,295,542,568]" italics="true" pageId="24" pageNumber="154">et al.</emphasis>
|
||
2014: 134–135
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, fig.
|
||
<quantity id="4CA23CB8FF9182045557B109675CFD80" box="[537,565,541,567]" metricMagnitude="-3" metricUnit="kg" metricValue="6.0" pageId="24" pageNumber="154" unit="g" value="6.0">6g</quantity>
|
||
–l (non
|
||
<bibRefCitation id="EFCBECACFF91820455DCB10A6658FD80" author="Lyman" box="[658,817,541,568]" firstAuthor="Lyman" pageId="24" pageNumber="154" pagination="193 - 204" refId="ref25910" refString="Lyman T. 1860. Descriptions of new Ophiuridae, belonging to the Smithsonian Institution and to the Museum of Comparative Zoology at Cambridge. Proceedings of the Boston Society of Natural History 7: 193 - 204, 252 - 262. https: // doi. org / 10.5962 / bhl. part. 4822" type="journal article" year="1860">Lyman, 1860</bibRefCitation>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFF91820457F3B155607FFD16" blockId="24.[189,1399,401,815]" pageId="24" pageNumber="154">
|
||
<treatmentCitationGroup id="AB4AB673FF91820457F3B155607FFD16" pageId="24" pageNumber="154">
|
||
<taxonomicName id="4C5AEADEFF91820457F3B155675EFDEC" authorityName="H. L. Clark" authorityYear="1940" box="[189,567,577,603]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="24" pageNumber="154" phylum="Echinodermata" rank="variety" species="teres" variety="unicolor">
|
||
<emphasis id="B92E4D4FFF91820457F3B15564FAFDEC" box="[189,403,577,603]" italics="true" pageId="24" pageNumber="154">Ophioderma teres</emphasis>
|
||
var.
|
||
<emphasis id="B92E4D4FFF918204569AB155675EFDEC" box="[468,567,577,603]" italics="true" pageId="24" pageNumber="154">unicolor</emphasis>
|
||
</taxonomicName>
|
||
– H.L.
|
||
<treatmentCitation id="0AFBB74CFF91820455D7B1556634FDEC" author="Clark H. L." box="[665,861,577,603]" page="342" pageId="24" pageNumber="154" year="1940">
|
||
<bibRefCitation id="EFCBECACFF91820455D7B1556634FDEC" author="Clark" box="[665,861,577,603]" firstAuthor="Clark" page="342" pageId="24" pageNumber="154" pagination="331 - 352" refId="ref24502" refString="Clark H. L. 1940. Eastern Pacific Expeditions of the New York Zoological Society. XXI. Notes on Echinoderms from the west coast of Central America. Zoologica 25 (3): 331 - 352. https: // doi. org / 10.5962 / p. 184708" type="journal article" year="1940">Clark 1940: 342</bibRefCitation>
|
||
</treatmentCitation>
|
||
(partim, non H.L.
|
||
<bibRefCitation id="EFCBECACFF918204530AB15561BCFDEC" author="Clark" box="[1092,1237,577,603]" firstAuthor="Clark" pageId="24" pageNumber="154" pagination="331 - 352" refId="ref24502" refString="Clark H. L. 1940. Eastern Pacific Expeditions of the New York Zoological Society. XXI. Notes on Echinoderms from the west coast of Central America. Zoologica 25 (3): 331 - 352. https: // doi. org / 10.5962 / p. 184708" type="journal article" year="1940">Clark, 1940</bibRefCitation>
|
||
). —
|
||
<treatmentCitation id="0AFBB74CFF918204525DB1556408FDC8" author="Downey M. E." page="115" pageId="24" pageNumber="154" year="1969">
|
||
<bibRefCitation id="EFCBECACFF918204525DB1556408FDC8" author="Downey" firstAuthor="Downey" page="115" pageId="24" pageNumber="154" pagination="1 - 239" refId="ref24648" refString="Downey M. E. 1969. Catalog of recent ophiuroid type specimens in major collections in the United States. United States National Museum Bulletin 293: 1 - 239. https: // doi. org / 10.5479 / si. 03629236.293" type="journal article" year="1969">Downey 1969: 115</bibRefCitation>
|
||
</treatmentCitation>
|
||
(partim, non H.L.
|
||
<bibRefCitation id="EFCBECACFF918204550DB17067BAFDC9" author="Clark" box="[579,723,612,638]" firstAuthor="Clark" pageId="24" pageNumber="154" pagination="331 - 352" refId="ref24502" refString="Clark H. L. 1940. Eastern Pacific Expeditions of the New York Zoological Society. XXI. Notes on Echinoderms from the west coast of Central America. Zoologica 25 (3): 331 - 352. https: // doi. org / 10.5962 / p. 184708" type="journal article" year="1940">Clark, 1940</bibRefCitation>
|
||
). —
|
||
<treatmentCitation id="0AFBB74CFF9182045440B17061F4FDC9" author="Granja-Fernandez M. R." box="[782,1181,612,638]" page="277" pageId="24" pageNumber="154" year="2019">
|
||
<bibRefCitation id="EFCBECACFF9182045440B17061F4FDC9" author="Granja-Fernandez" box="[782,1181,612,638]" firstAuthor="Granja-Fernandez" page="277 - 279" pageId="24" pageNumber="154" refId="ref24810" refString="Granja-Fernandez M. R. 2019. Ofiuroideos (Echinodermata) del Pacifico oriental tropical. PhD thesis, Universidad Autonoma Metropolitana, Mexico." type="book" year="2019">Granja-Fernández 2019: 277–279</bibRefCitation>
|
||
</treatmentCitation>
|
||
(partim, non H.L.
|
||
<bibRefCitation id="EFCBECACFF91820457A2B1936412FD16" author="Clark" box="[236,379,647,673]" firstAuthor="Clark" pageId="24" pageNumber="154" pagination="331 - 352" refId="ref24502" refString="Clark H. L. 1940. Eastern Pacific Expeditions of the New York Zoological Society. XXI. Notes on Echinoderms from the west coast of Central America. Zoologica 25 (3): 331 - 352. https: // doi. org / 10.5962 / p. 184708" type="journal article" year="1940">Clark, 1940</bibRefCitation>
|
||
). —
|
||
<treatmentCitation id="0AFBB74CFF91820456FDB19366E2FD16" author="Mireles-Velazquez D. & Rosales-Contreras G. I. & Conejeros-Vargas C. A. & Solis-Marin F. A. & Granja-Fernandez R." box="[435,907,647,674]" page="317" pageId="24" pageNumber="154" year="2021">
|
||
<bibRefCitation id="EFCBECACFF91820456FDB19366E2FD16" author="Mireles-Velazquez" box="[435,907,647,674]" etAl="et al." firstAuthor="Mireles-Velazquez" page="317 - 319" pageId="24" pageNumber="154" pagination="312 - 333" refId="ref26061" refString="Mireles-Velazquez D., Rosales-Contreras G. I., Conejeros-Vargas C. A., Solis-Marin F. A. & Granja-Fernandez R. 2021. Los ofiuroideos (Echinodermata: Ophiuroidea) de la bahia de Chamela, Jalisco, Mexico. Revista de Biologia Tropical 69 (S 1): 312 - 333. https: // doi. org / 10.15517 / rbt. v 69 iSuppl. 1.46364" type="journal article" year="2021">
|
||
Mireles-Velázquez
|
||
<emphasis id="B92E4D4FFF91820455DBB19C67A0FD16" box="[661,713,647,673]" italics="true" pageId="24" pageNumber="154">et al</emphasis>
|
||
. 2021: 317–319
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, fig. 2e–f (non H.L.
|
||
<bibRefCitation id="EFCBECACFF9182045334B1936060FD16" author="Clark" box="[1146,1289,647,673]" firstAuthor="Clark" pageId="24" pageNumber="154" pagination="331 - 352" refId="ref24502" refString="Clark H. L. 1940. Eastern Pacific Expeditions of the New York Zoological Society. XXI. Notes on Echinoderms from the west coast of Central America. Zoologica 25 (3): 331 - 352. https: // doi. org / 10.5962 / p. 184708" type="journal article" year="1940">Clark, 1940</bibRefCitation>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFF91820457F3B1BE6497FD50" blockId="24.[189,1399,401,815]" pageId="24" pageNumber="154">
|
||
<treatmentCitationGroup id="AB4AB673FF91820457F3B1BE6497FD50" pageId="24" pageNumber="154">
|
||
<taxonomicName id="4C5AEADEFF91820457F3B1BE674AFD73" authorityName="H. L. Clark" authorityYear="1940" box="[189,547,682,708]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" isUncertain="true" kingdom="Animalia" order="Ophiacanthida" pageId="24" pageNumber="154" phylum="Echinodermata" rank="variety" species="teres" variety="unicolor">
|
||
<emphasis id="B92E4D4FFF91820457F3B1BE6438FD73" box="[189,337,682,708]" italics="true" pageId="24" pageNumber="154">Ophioderma</emphasis>
|
||
cf.
|
||
<emphasis id="B92E4D4FFF91820456CEB1BF674AFD73" box="[384,547,682,708]" italics="true" pageId="24" pageNumber="154">teres unicolor</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AFBB74CFF918204550DB1BE66FCFD73" author="Humara-Gil K. J. & Granja-Fernandez R. & Bautista-Guerrero E. & Rodriguez-Troncoso A. P." box="[579,917,682,709]" page="367" pageId="24" pageNumber="154" year="2022">
|
||
<bibRefCitation id="EFCBECACFF918204550DB1BE66FCFD73" author="Humara-Gil" box="[579,917,682,709]" etAl="et al." firstAuthor="Humara-Gil" page="367" pageId="24" pageNumber="154" pagination="365 - 384" refId="ref25718" refString="Humara-Gil K. J., Granja-Fernandez R., Bautista-Guerrero E. & Rodriguez-Troncoso A. P. 2022. Overlooked for over a century: Ophioderma occultum sp. nov. (Echinodermata), a new species of brittle star from the Eastern Pacific. Journal of Natural History 56 (5 - 8): 365 - 384. https: // doi. org / 10.1080 / 00222933.2022.2071179" type="journal article" year="2022">
|
||
Humara-Gil
|
||
<emphasis id="B92E4D4FFF9182045596B1BF667AFD73" box="[728,787,682,708]" italics="true" pageId="24" pageNumber="154">et al.</emphasis>
|
||
2022: 367
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 379–380, fig. 5 (partial DNA sequences of genes COI and 16S).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFF91820457F3B1E5665BFCBC" blockId="24.[189,1399,401,815]" box="[189,818,753,780]" pageId="24" pageNumber="154">
|
||
<treatmentCitationGroup id="AB4AB673FF91820457F3B1E5665BFCBC" box="[189,818,753,780]" pageId="24" pageNumber="154">
|
||
<taxonomicName id="4C5AEADEFF91820457F3B1E56413FCBC" box="[189,378,753,779]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="24" pageNumber="154" phylum="Echinodermata" rank="species" species="undetermined">
|
||
<emphasis id="B92E4D4FFF91820457F3B1E56438FCBC" box="[189,337,753,779]" italics="true" pageId="24" pageNumber="154">Ophioderma</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AFBB74CFF91820456D9B1E56666FCBB" author="Granja-Fernandez R. & Rodriguez-Zaragoza F. A. & Lopez-Perez A. & Lopez-Lopez D. A." box="[407,783,753,780]" page="5" pageId="24" pageNumber="154" year="2022">
|
||
<bibRefCitation id="EFCBECACFF91820456D9B1E56666FCBB" author="Granja-Fernandez" box="[407,783,753,780]" etAl="et al." firstAuthor="Granja-Fernandez" page="5" pageId="24" pageNumber="154" refId="ref25144" refString="Granja-Fernandez R., Rodriguez-Zaragoza F. A., Lopez-Perez A. & Lopez-Lopez D. A. 2022. Coral reef ecosystem - associated echinoderms (Echinodermata) at the Southern Mexican Pacific: species richness, distribution, and composition. Marine Biodiversity 52: 65. https: // doi. org / 10.1007 / s 12526 - 022 - 01305 - 0" type="journal volume" year="2022">
|
||
Granja-Fernández
|
||
<emphasis id="B92E4D4FFF918204553CB1E667C4FCBC" box="[626,685,753,779]" italics="true" pageId="24" pageNumber="154">et al.</emphasis>
|
||
2022: 5
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 9.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFF91820457F3B0006616FC99" blockId="24.[189,1399,401,815]" box="[189,895,788,815]" pageId="24" pageNumber="154">
|
||
<taxonomicName id="4C5AEADEFF91820457F3B00064FDFC99" box="[189,404,788,815]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="24" pageNumber="154" phylum="Echinodermata" rank="species" species="undefined-A">
|
||
<emphasis id="B92E4D4FFF91820457F3B0006438FC99" box="[189,337,788,814]" italics="true" pageId="24" pageNumber="154">Ophioderma</emphasis>
|
||
sp. A
|
||
</taxonomicName>
|
||
– This study: 3–12,
|
||
<figureCitation id="13618DD8FF91820455CAB0006785FC99" box="[644,748,788,814]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="8.[189,232,1892,1918]" captionStartId-1="9.[189,232,1928,1954]" captionTargetBox-0="[224,1365,769,1835]" captionTargetBox-1="[222,1365,1013,1864]" captionTargetId-0="figure-204@8.[189,1398,747,1851]" captionTargetId-1="figure-223@9.[189,1398,1000,1886]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Fig. 1. Bayesian inference (BI) tree of COI sequences. Numbers on branches indicate posterior probability/bootstrap values obtained with BI and maximum likelihood (ML), respectively. A dash (–) indicates the branches not recovered in the ML analysis. Sequences obtained in this study are indicated in bold (see Supp. file 5)." captionText-1="Fig. 2. Bayesian inference (BI) tree of 16Ssequences. Numberson branches indicate posterior probability/ bootstrap values obtained with BI and maximum likelihood, respectively. Sequences obtained in this study are indicated in bold (see Supp. file 5)." figureDoi-0="http://doi.org/10.5281/zenodo.13310021" figureDoi-1="http://doi.org/10.5281/zenodo.13310023" httpUri-0="https://zenodo.org/record/13310021/files/figure.png" httpUri-1="https://zenodo.org/record/13310023/files/figure.png" pageId="24" pageNumber="154">Figs 1–2</figureCitation>
|
||
,
|
||
<tableCitation id="C6D8A4E6FF91820455B6B0006613FC99" box="[760,890,788,814]" captionStart-0="Table 1" captionStart-1="Table 2" captionStart-2="Table 3" captionStart-3="Table 4" captionStartId-0="5.[185,255,655,683]" captionStartId-1="6.[186,256,582,610]" captionStartId-2="9.[189,259,265,291]" captionStartId-3="10.[185,255,645,673]" captionTargetBox-1="[197,1897,838,1702]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="Table 1. Main external and internal morphological characters that distinguish the studied species/morphotypes of Ophioderma Müller & Troschel, 1840. The number of DAP pieces is given as an interval, followed by the mean in parentheses. A dash (–) indicates no data are available due to lack of material. Abbreviations: AL = arm length; DAP = dorsal arm plate; DD = disc diameter." captionText-1="Table 2. Morphometric data with the highest contribution to the differentiation of the studied species/morphotypes of Ophioderma Müller & Troschel, 1840. The ratios and counts are indicated as intervals, followed by the mean in parentheses.Abbreviations: 1VAP_W:L = first ventral arm plate (VAP) width:length ratio; AS_prox_3, 5 = number of arm spines in the 3rd or 5th proximal arm segment; DAPs_med_1, 2, 3 = number of DAP pieces in the 1st, 2nd or 3rd median arm segment; DAPs_prox_3, 4, 5 = number of dorsal arm plate (DAP) pieces in the 3rd, 4th or 5th proximal arm segment; DD = disc diameter; LOPas = lateral oral papillae; OSh_W:L = oral shield width:length ratio; VAP_L:W_med = VAP length:width ratio (median arm section); VAP_pores = number of arm segments with pores present between the proximalmost VAPs." captionText-2="Table 3. Mean genetic distances (%) between the analyzed species/morphotypes of Ophioderma Müller & Troschel, 1840 and Ophiopeza Peters, 1851 (COI, lower left; 16S, upper right). A dash (–) indicates no data are available due to a lack of sequences." captionText-3="Table 4. Summary of the comparisons of the analyzed evidence by pairs of species/morphotypes of Ophioderma Müller & Troschel, 1840. A dash (–) indicates no data are available for one or both species/morphotypes compared." pageId="24" pageNumber="154">Tables 1–4</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C340C2D6FF91820457F3B0416470FBB0" pageId="24" pageNumber="154" type="diagnosis">
|
||
<paragraph id="8BE5915DFF91820457F3B041645FFCD8" blockId="24.[189,310,853,879]" box="[189,310,853,879]" pageId="24" pageNumber="154">
|
||
<heading id="D0AD2631FF91820457F3B041645FFCD8" bold="true" box="[189,310,853,879]" fontSize="11" level="3" pageId="24" pageNumber="154" reason="3">
|
||
<emphasis id="B92E4D4FFF91820457F3B041645FFCD8" bold="true" box="[189,310,853,879]" pageId="24" pageNumber="154">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFF91820457F3B0906470FBB0" blockId="24.[189,1400,900,1031]" pageId="24" pageNumber="154">Radial shields naked. Arms short (mean AL:DD = 2.7). DAPs divided into multiple pieces (mean = 3, maximum = 9). Coloration dark brown; dorsal and ventral disc with dark, sinuous rings; dorsal arms brown; ventral arms cream (preserved specimens) or yellow (in vivo specimens) proximally, darkening distally.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C340C2D6FF91820457F3B73B6117FB2C" pageId="24" pageNumber="154" type="etymology">
|
||
<paragraph id="8BE5915DFF91820457F3B73B642DFBFE" blockId="24.[189,324,1071,1097]" box="[189,324,1071,1097]" pageId="24" pageNumber="154">
|
||
<heading id="D0AD2631FF91820457F3B73B642DFBFE" bold="true" box="[189,324,1071,1097]" fontSize="11" level="3" pageId="24" pageNumber="154" reason="3">
|
||
<emphasis id="B92E4D4FFF91820457F3B73B642DFBFE" bold="true" box="[189,324,1071,1097]" pageId="24" pageNumber="154">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFF91820457F3B7496117FB2C" blockId="24.[189,1398,1117,1179]" pageId="24" pageNumber="154">
|
||
The specific epithet
|
||
<taxonomicName id="4C5AEADEFF91820456E1B74A64B4FBC0" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" box="[431,477,1118,1143]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="24" pageNumber="154" phylum="Echinodermata" rank="species" species="aija">
|
||
<emphasis id="B92E4D4FFF91820456E1B74A64B4FBC0" box="[431,477,1118,1143]" italics="true" pageId="24" pageNumber="154">aija</emphasis>
|
||
</taxonomicName>
|
||
combines the first two letters of the given names of Aida Janet Gil-Rabadán, mother of the first author. Born under a lucky star, she deserves her own on Earth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C340C2D6FF91820457F3B7D66415F97C" pageId="24" pageNumber="154" type="materials_examined">
|
||
<paragraph id="8BE5915DFF91820457F3B7D664C3FB6B" blockId="24.[189,426,1218,1244]" box="[189,426,1218,1244]" pageId="24" pageNumber="154">
|
||
<heading id="D0AD2631FF91820457F3B7D664C3FB6B" bold="true" box="[189,426,1218,1244]" fontSize="11" level="3" pageId="24" pageNumber="154" reason="3">
|
||
<emphasis id="B92E4D4FFF91820457F3B7D664C3FB6B" bold="true" box="[189,426,1218,1244]" pageId="24" pageNumber="154">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<materialsCitation id="3B329B00FF91820457A4B7E564ABFAE6" collectingDate="2008-05" collectorName="La Mina" country="Mexico" latitude="15.673833" location="La Mina" longLatPrecision="13" longitude="-96.47667" pageId="24" pageNumber="154" specimenCode="ICML-UNAM 18466" specimenCount="1" stateProvince="Oaxaca" typeStatus="holotype">
|
||
<paragraph id="8BE5915DFF91820457A4B7E56435FABC" blockId="24.[189,1399,1265,1361]" box="[234,348,1265,1291]" pageId="24" pageNumber="154">
|
||
<emphasis id="B92E4D4FFF91820457A4B7E56435FABC" bold="true" box="[234,348,1265,1291]" pageId="24" pageNumber="154">
|
||
<typeStatus id="54E12FFFFF91820457A4B7E56435FABC" box="[234,348,1265,1291]" pageId="24" pageNumber="154" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFF91820457F3B60064ABFAE6" blockId="24.[189,1399,1265,1361]" pageId="24" pageNumber="154">
|
||
<collectingCountry id="F34DD1CDFF91820457F3B6006458FA99" box="[189,305,1300,1326]" name="Mexico" pageId="24" pageNumber="154">MEXICO</collectingCountry>
|
||
• spec. preserved in 70% ethanol;
|
||
<collectingRegion id="499E5FBFFF918204559FB6006645FA99" box="[721,812,1300,1326]" country="Mexico" name="Oaxaca" pageId="24" pageNumber="154">Oaxaca</collectingRegion>
|
||
,
|
||
<location id="8E85C786FF9182045477B60066CAFA99" LSID="urn:lsid:plazi:treatment:03F3204BFF91820155CDB21C612DFA7B:8E85C786FF9182045477B60066CAFA99" box="[825,931,1300,1326]" country="Mexico" latitude="15.673833" longLatPrecision="13" longitude="-96.47667" name="La Mina" pageId="24" pageNumber="154" stateProvince="Oaxaca">La Mina</location>
|
||
;
|
||
<geoCoordinate id="EE6EF79AFF91820454FEB606612BFA99" box="[944,1090,1298,1326]" degrees="15" direction="north" minutes="40.43" orientation="latitude" pageId="24" pageNumber="154" precision="9" value="15.673833">15°40.43′ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE6EF79AFF918204531EB606618EFA99" box="[1104,1255,1298,1326]" degrees="96" direction="west" minutes="28.60" orientation="longitude" pageId="24" pageNumber="154" precision="9" value="-96.47667">96°28.60′ W</geoCoordinate>
|
||
;
|
||
<date id="FFE4B79DFF91820453BBB600601BFA99" box="[1269,1394,1300,1326]" pageId="24" pageNumber="154" value="2008-05">
|
||
<collectingDate id="EFA04E75FF91820453BBB600601BFA99" box="[1269,1394,1300,1326]" pageId="24" pageNumber="154" value="2008-05">May 2008</collectingDate>
|
||
</date>
|
||
;
|
||
<specimenCode id="DBFC3926FF91820457F3B62364D7FAE6" box="[189,446,1335,1361]" pageId="24" pageNumber="154">ICML-UNAM 18466</specimenCode>
|
||
.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B329B00FF91820457A4B66D640EFA4B" collectingDate="2009-03-03" collectorName="D. Arellanes-Garcia & F. Benitez-Villalobos" country="Mexico" elevation="0" location="Estacahuite" pageId="24" pageNumber="154" specimenCode="ICML-UNAM 18467" specimenCount="4" stateProvince="Oaxaca" typeStatus="paratype">
|
||
<paragraph id="8BE5915DFF91820457A4B66D640EFA24" blockId="24.[189,1399,1401,1639]" box="[234,359,1401,1427]" pageId="24" pageNumber="154">
|
||
<emphasis id="B92E4D4FFF91820457A4B66D640EFA24" bold="true" box="[234,359,1401,1427]" pageId="24" pageNumber="154">
|
||
<typeStatus id="54E12FFFFF91820457A4B66D640EFA24" box="[234,359,1401,1427]" pageId="24" pageNumber="154" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFF91820457F3B688607CF9D1" blockId="24.[189,1399,1401,1639]" pageId="24" pageNumber="154">
|
||
<collectingCountry id="F34DD1CDFF91820457F3B6886458FA01" box="[189,305,1436,1462]" name="Mexico" pageId="24" pageNumber="154">MEXICO</collectingCountry>
|
||
–
|
||
<emphasis id="B92E4D4FFF9182045619B68F64DFFA02" bold="true" box="[343,438,1435,1461]" pageId="24" pageNumber="154">
|
||
<collectingRegion id="499E5FBFFF9182045619B68F64DFFA02" box="[343,438,1435,1461]" country="Mexico" name="Oaxaca" pageId="24" pageNumber="154">Oaxaca</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D5C5AD4FF9182045696B688675AFA01" box="[472,563,1436,1462]" pageId="24" pageNumber="154">4 specs</specimenCount>
|
||
(preserved in 70% ethanol);
|
||
<location id="8E85C786FF91820454D1B6886143FA01" LSID="urn:lsid:plazi:treatment:03F3204BFF91820155CDB21C612DFA7B:8E85C786FF91820454D1B6886143FA01" box="[927,1066,1436,1462]" country="Mexico" name="Estacahuite" pageId="24" pageNumber="154" stateProvince="Oaxaca">Estacahuite</location>
|
||
;
|
||
<elevation id="0077766EFF9182045374B688607CFA01" box="[1082,1301,1436,1462]" pageId="24" pageNumber="154">
|
||
3.0–
|
||
<quantity id="4CA23CB8FF9182045321B68861ADFA01" box="[1135,1220,1436,1462]" metricMagnitude="1" metricUnit="m" metricValue="1.22" pageId="24" pageNumber="154" unit="m" value="12.2">12.2 m</quantity>
|
||
depth
|
||
</elevation>
|
||
;
|
||
<date id="FFE4B79DFF918204526BB6886592FA6E" pageId="24" pageNumber="154" value="2009-03-03">
|
||
<collectingDate id="EFA04E75FF918204526BB6886592FA6E" pageId="24" pageNumber="154" value="2009-03-03">3 Mar. 2009</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26AFF48BFF9182045645B6AB6493FA6E" box="[267,506,1471,1497]" pageId="24" pageNumber="154">D. Arellanes-García</collectorName>
|
||
and
|
||
<collectorName id="26AFF48BFF9182045575B6AB6651FA6E" box="[571,824,1471,1497]" pageId="24" pageNumber="154">F. Benítez-Villalobos</collectorName>
|
||
leg.; sand under dead coral and rocks;
|
||
<specimenCode id="DBFC3926FF918204526DB6AB640EFA4B" pageId="24" pageNumber="154">ICML-UNAM 18467</specimenCode>
|
||
•
|
||
<materialsCitation id="3B329B00FF9182045631B6F66687F9A8" collectingDate="2020-02-20" collectionCode="R" collectorName="R. Granja-Fernandez" country="Mexico" elevation="9" latitude="15.668334" location="Estacahuite" longLatPrecision="21" longitude="-96.48111" pageId="24" pageNumber="154" specimenCode="ICML-UNAM 18468" specimenCount="5" stateProvince="Oaxaca" typeStatus="paratype">
|
||
<specimenCount id="9D5C5AD4FF9182045631B6F664BCFA4A" box="[383,469,1506,1533]" pageId="24" pageNumber="154">5 specs</specimenCount>
|
||
(preserved in 96% ethanol);
|
||
<location id="8E85C786FF9182045467B6F666DDFA4B" LSID="urn:lsid:plazi:treatment:03F3204BFF91820155CDB21C612DFA7B:8E85C786FF9182045467B6F666DDFA4B" box="[809,948,1506,1532]" country="Mexico" latitude="15.668334" longLatPrecision="21" longitude="-96.48111" name="Estacahuite" pageId="24" pageNumber="154" stateProvince="Oaxaca">Estacahuite</location>
|
||
;
|
||
<geoCoordinate id="EE6EF79AFF91820454F1B6F5612EFA4A" box="[959,1095,1505,1533]" degrees="15" direction="north" minutes="40" orientation="latitude" pageId="24" pageNumber="154" precision="15" seconds="6" value="15.668334">15°40′6′′ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE6EF79AFF918204531CB6F56187FA4A" box="[1106,1262,1505,1533]" degrees="96" direction="west" minutes="28" orientation="longitude" pageId="24" pageNumber="154" precision="15" seconds="52" value="-96.48111">96°28′52′′ W</geoCoordinate>
|
||
;
|
||
<elevation id="0077766EFF91820453B7B6F6601BFA4B" box="[1273,1394,1506,1533]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="24" pageNumber="154" unit="m" value="9.0">
|
||
<quantity id="4CA23CB8FF91820453B7B6F6604FFA4A" box="[1273,1318,1506,1533]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="24" pageNumber="154" unit="m" value="9.0">9 m</quantity>
|
||
depth
|
||
</elevation>
|
||
;
|
||
<date id="FFE4B79DFF91820457F3B5116432F9A8" box="[189,347,1541,1567]" pageId="24" pageNumber="154" value="2020-02-20">
|
||
<collectingDate id="EFA04E75FF91820457F3B5116432F9A8" box="[189,347,1541,1567]" pageId="24" pageNumber="154" value="2020-02-20">20 Feb. 2020</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26AFF48BFF9182045626B5126737F9A8" box="[360,606,1541,1568]" pageId="24" pageNumber="154">R. Granja-Fernández</collectorName>
|
||
leg.; rocks;
|
||
<specimenCode id="DBFC3926FF91820455BEB5116687F9A8" box="[752,1006,1541,1567]" pageId="24" pageNumber="154">ICML-UNAM 18468</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B329B00FF9182045346B5116078F9D1" collectingDate="2020-02-21" collectionCode="R, COI" collectorName="R. Granja-Fernandez" country="Mexico" elevation="10" latitude="15.6875" location="Tijera" longLatPrecision="21" longitude="-96.44195" pageId="24" pageNumber="154" specimenCode="ICML-UNAM 18469" specimenCount="2" stateProvince="Oaxaca" typeStatus="paratype">
|
||
<specimenCount id="9D5C5AD4FF9182045346B5116137F997" box="[1032,1118,1541,1568]" pageId="24" pageNumber="154">2 specs</specimenCount>
|
||
(preserved dry);
|
||
<location id="8E85C786FF9182045267B511601BF9A8" LSID="urn:lsid:plazi:treatment:03F3204BFF91820155CDB21C612DFA7B:8E85C786FF9182045267B511601BF9A8" box="[1321,1394,1541,1567]" country="Mexico" latitude="15.6875" longLatPrecision="21" longitude="-96.44195" name="Tijera" pageId="24" pageNumber="154" stateProvince="Oaxaca">Tijera</location>
|
||
;
|
||
<geoCoordinate id="EE6EF79AFF91820457F3B533643DF9F4" box="[189,340,1575,1603]" degrees="15" direction="north" minutes="41" orientation="latitude" pageId="24" pageNumber="154" precision="15" seconds="15" value="15.6875">15°41′15′′ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE6EF79AFF918204562EB5336495F9F4" box="[352,508,1575,1603]" degrees="96" direction="west" minutes="26" orientation="longitude" pageId="24" pageNumber="154" precision="15" seconds="31" value="-96.44195">96°26′31′′ W</geoCoordinate>
|
||
;
|
||
<elevation id="0077766EFF9182045547B53D67FBF9F5" box="[521,658,1576,1603]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="24" pageNumber="154" unit="m" value="10.0">
|
||
<quantity id="4CA23CB8FF9182045547B53D672CF9F4" box="[521,581,1577,1603]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="24" pageNumber="154" unit="m" value="10.0">10 m</quantity>
|
||
depth
|
||
</elevation>
|
||
;
|
||
<date id="FFE4B79DFF91820455D0B53D6655F9F4" box="[670,828,1576,1603]" pageId="24" pageNumber="154" value="2020-02-21">
|
||
<collectingDate id="EFA04E75FF91820455D0B53D6655F9F4" box="[670,828,1576,1603]" pageId="24" pageNumber="154" value="2020-02-21">21 Feb. 2020</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26AFF48BFF9182045407B53D6129F9F5" box="[841,1088,1576,1603]" pageId="24" pageNumber="154">R. Granja-Fernández</collectorName>
|
||
leg.; rocks; DNA voucher spec. labelled as “OJ5”;
|
||
<accessionNumber id="94090CBEFF9182045693B55867B2F9D0" box="[477,731,1612,1639]" pageId="24" pageNumber="154">GenBank: OR789590</accessionNumber>
|
||
<accessionNumber id="94090CBEFF91820455ADB558616DF9D1" box="[739,1028,1612,1638]" pageId="24" pageNumber="154">(COI), OR800000 (16S)</accessionNumber>
|
||
;
|
||
<specimenCode id="DBFC3926FF918204535EB5586078F9D1" box="[1040,1297,1612,1638]" pageId="24" pageNumber="154">ICML-UNAM 18469</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<paragraph id="8BE5915DFF91820457A4B5996415F97C" blockId="24.[189,422,1677,1739]" pageId="24" pageNumber="154">
|
||
<emphasis id="B92E4D4FFF91820457A4B59964CFF910" bold="true" box="[234,422,1677,1703]" pageId="24" pageNumber="154">Other material</emphasis>
|
||
See Supp. file 1.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C340C2D6FF91820757F3B5E6675FF8E3" lastPageId="27" lastPageNumber="157" pageId="24" pageNumber="154" type="description">
|
||
<paragraph id="8BE5915DFF91820457F3B5E66427F8BB" blockId="24.[189,334,1778,1804]" box="[189,334,1778,1804]" pageId="24" pageNumber="154">
|
||
<heading id="D0AD2631FF91820457F3B5E66427F8BB" bold="true" box="[189,334,1778,1804]" fontSize="11" level="3" pageId="24" pageNumber="154" reason="3">
|
||
<emphasis id="B92E4D4FFF91820457F3B5E66427F8BB" bold="true" box="[189,334,1778,1804]" pageId="24" pageNumber="154">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFF91820457A4B4346435F88D" blockId="24.[188,1399,1824,1992]" box="[234,348,1824,1850]" pageId="24" pageNumber="154">
|
||
<emphasis id="B92E4D4FFF91820457A4B4346435F88D" bold="true" box="[234,348,1824,1850]" pageId="24" pageNumber="154">
|
||
<typeStatus id="54E12FFFFF91820457A4B4346435F88D" box="[234,348,1824,1850]" pageId="24" pageNumber="154" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFF91820557F3B45064E6FE07" blockId="24.[188,1399,1824,1992]" lastBlockId="25.[188,1399,265,432]" lastPageId="25" lastPageNumber="155" pageId="24" pageNumber="154">
|
||
DD =
|
||
<quantity id="4CA23CB8FF9182045641B4576417F8E9" box="[271,382,1859,1886]" metricMagnitude="-2" metricUnit="m" metricValue="2.67" pageId="24" pageNumber="154" unit="mm" value="26.7">26.7 mm</quantity>
|
||
, AL =
|
||
<quantity id="4CA23CB8FF9182045697B4506720F8E9" box="[473,585,1860,1886]" metricMagnitude="-2" metricUnit="m" metricValue="7.89" pageId="24" pageNumber="154" unit="mm" value="78.9">78.9 mm</quantity>
|
||
, AL:DD = 3. Disc rounded, covered by rounded granules, slightly separated from each other. Granule size increasing from center to periphery. Dorsal disc granule density 60 per mm
|
||
<superScript id="7C2F3C15FF918204560EB49D6420F820" attach="right" box="[320,329,1929,1943]" fontSize="6" pageId="24" pageNumber="154">2</superScript>
|
||
. Radial shields naked; visible section approximately 1.5× as long as wide, oval; distance between shields about 4× width of shield (
|
||
<figureCitation id="13618DD8FF91820455E1B4B96664F870" box="[687,781,1965,1991]" captionStart="Fig" captionStartId="26.[189,232,1787,1813]" captionTargetBox="[319,1268,263,1746]" captionTargetId="figure-12@26.[319,1268,263,1746]" captionTargetPageId="26" captionText="Fig. 8. Ophioderma aija sp. nov., holotype (ICML-UNAM 18466, DD = 26.7 mm). A. Dorsal disc. B. Ventral disc (inset: section between the arm and the distal genital slit). C. Close-up of a jaw. D. Proximalmost dorsal arm. E. Median dorsal arm. F. Distalmost dorsal arm. G. Proximalmost ventral arm. H. Median ventral arm. I. Distalmost ventral arm. J. Lateral arm plates and arm spines. K. Dorsal view. Abbreviations: 2° AdShSp = secondary adoral shield spine; AdShSp = adoral shield spine; IPa = infradental papilla; LOPas = lateral oral papillae; LyOs = Lyman’s ossicle; OPRSp = oral plate ridge spine; RS = radial shield; TPa = tooth papilla; vT = ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.13310037" httpUri="https://zenodo.org/record/13310037/files/figure.png" pageId="24" pageNumber="154">Fig. 8A</figureCitation>
|
||
). Ventral interradii covered with granules of uniform size, slightly separated from each other. Four genital slits per interradius. Proximal genital slits oval, separated from distal section of oral shields by two rows of granules, but in contact with 1
|
||
<superScript id="7C2F3C15FF908205526EB2386042FE8D" attach="left" box="[1312,1323,300,314]" fontSize="6" pageId="25" pageNumber="155">st</superScript>
|
||
LAP; reaching up to proximal section of 2
|
||
<superScript id="7C2F3C15FF9082055539B25B67E1FEEA" attach="left" box="[631,648,335,349]" fontSize="6" pageId="25" pageNumber="155">nd</superScript>
|
||
VAP. Distal genital slits oval, 1.5× as long as proximal ones, placed between 7
|
||
<superScript id="7C2F3C15FF90820556C9B26664FDFE37" attach="left" box="[391,404,370,384]" fontSize="6" pageId="25" pageNumber="155">th</superScript>
|
||
and 9
|
||
<superScript id="7C2F3C15FF9082055695B2666481FE37" attach="left" box="[475,488,370,384]" fontSize="6" pageId="25" pageNumber="155">th</superScript>
|
||
arm segments; surrounded by granule-bearing scales and naked scales next to the arm (
|
||
<figureCitation id="13618DD8FF908205566BB28264E8FE07" box="[293,385,406,432]" captionStart="Fig" captionStartId="26.[189,232,1787,1813]" captionTargetBox="[319,1268,263,1746]" captionTargetId="figure-12@26.[319,1268,263,1746]" captionTargetPageId="26" captionText="Fig. 8. Ophioderma aija sp. nov., holotype (ICML-UNAM 18466, DD = 26.7 mm). A. Dorsal disc. B. Ventral disc (inset: section between the arm and the distal genital slit). C. Close-up of a jaw. D. Proximalmost dorsal arm. E. Median dorsal arm. F. Distalmost dorsal arm. G. Proximalmost ventral arm. H. Median ventral arm. I. Distalmost ventral arm. J. Lateral arm plates and arm spines. K. Dorsal view. Abbreviations: 2° AdShSp = secondary adoral shield spine; AdShSp = adoral shield spine; IPa = infradental papilla; LOPas = lateral oral papillae; LyOs = Lyman’s ossicle; OPRSp = oral plate ridge spine; RS = radial shield; TPa = tooth papilla; vT = ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.13310037" httpUri="https://zenodo.org/record/13310037/files/figure.png" pageId="25" pageNumber="155">Fig. 8B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFF90820557F3B2C36728FCBC" blockId="25.[188,1400,471,779]" pageId="25" pageNumber="155">
|
||
Oral shields 1.6× as wide as long, rounded triangular; proximal edge convex forming a rounded apex; lateral edges rounded; distal edge straight. Madreporite rounded trapezoidal, with a central depression deviated towards distal section; distal edge convex. Adoral shields covered by small granules. Jaws with 8–9 oral papillae: LyOs 3.5× as long as wide, angled upwards; AdShSp the largest, trapezoidal; 2°AdShSp smaller than AdShSp, rounded rectangular; LOPas 3–5, rounded rectangular to conical; IPa similar to LOPas, larger; TPa two at jaw apex, triangular to rectangular, robust. Teeth 4–5: vT triangular, slightly flattened; median and dorsalmost teeth quadrangular. One OPRSp on each side of the jaw, conspicuous. Oral plates covered with granules larger than those covering adoral shields, decreasing in size towards periphery (
|
||
<figureCitation id="13618DD8FF9082055699B1E5675AFCBC" box="[471,563,753,779]" captionStart="Fig" captionStartId="26.[189,232,1787,1813]" captionTargetBox="[319,1268,263,1746]" captionTargetId="figure-12@26.[319,1268,263,1746]" captionTargetPageId="26" captionText="Fig. 8. Ophioderma aija sp. nov., holotype (ICML-UNAM 18466, DD = 26.7 mm). A. Dorsal disc. B. Ventral disc (inset: section between the arm and the distal genital slit). C. Close-up of a jaw. D. Proximalmost dorsal arm. E. Median dorsal arm. F. Distalmost dorsal arm. G. Proximalmost ventral arm. H. Median ventral arm. I. Distalmost ventral arm. J. Lateral arm plates and arm spines. K. Dorsal view. Abbreviations: 2° AdShSp = secondary adoral shield spine; AdShSp = adoral shield spine; IPa = infradental papilla; LOPas = lateral oral papillae; LyOs = Lyman’s ossicle; OPRSp = oral plate ridge spine; RS = radial shield; TPa = tooth papilla; vT = ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.13310037" httpUri="https://zenodo.org/record/13310037/files/figure.png" pageId="25" pageNumber="155">Fig. 8C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFF90820557F3B02661C0FA37" blockId="25.[188,1399,818,1408]" pageId="25" pageNumber="155">
|
||
Five arms rounded, tapering distally: all incomplete, three regenerating, the remainder without the distalmost segments (
|
||
<figureCitation id="13618DD8FF90820556F1B0416749FCC7" box="[447,544,853,880]" captionStart="Fig" captionStartId="26.[189,232,1787,1813]" captionTargetBox="[319,1268,263,1746]" captionTargetId="figure-12@26.[319,1268,263,1746]" captionTargetPageId="26" captionText="Fig. 8. Ophioderma aija sp. nov., holotype (ICML-UNAM 18466, DD = 26.7 mm). A. Dorsal disc. B. Ventral disc (inset: section between the arm and the distal genital slit). C. Close-up of a jaw. D. Proximalmost dorsal arm. E. Median dorsal arm. F. Distalmost dorsal arm. G. Proximalmost ventral arm. H. Median ventral arm. I. Distalmost ventral arm. J. Lateral arm plates and arm spines. K. Dorsal view. Abbreviations: 2° AdShSp = secondary adoral shield spine; AdShSp = adoral shield spine; IPa = infradental papilla; LOPas = lateral oral papillae; LyOs = Lyman’s ossicle; OPRSp = oral plate ridge spine; RS = radial shield; TPa = tooth papilla; vT = ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.13310037" httpUri="https://zenodo.org/record/13310037/files/figure.png" pageId="25" pageNumber="155">Fig. 8K</figureCitation>
|
||
). Dorsal arm base with multiple scales extending to 1–2 arm segments (
|
||
<figureCitation id="13618DD8FF9082055788B06D644DFC24" box="[198,292,889,915]" captionStart="Fig" captionStartId="26.[189,232,1787,1813]" captionTargetBox="[319,1268,263,1746]" captionTargetId="figure-12@26.[319,1268,263,1746]" captionTargetPageId="26" captionText="Fig. 8. Ophioderma aija sp. nov., holotype (ICML-UNAM 18466, DD = 26.7 mm). A. Dorsal disc. B. Ventral disc (inset: section between the arm and the distal genital slit). C. Close-up of a jaw. D. Proximalmost dorsal arm. E. Median dorsal arm. F. Distalmost dorsal arm. G. Proximalmost ventral arm. H. Median ventral arm. I. Distalmost ventral arm. J. Lateral arm plates and arm spines. K. Dorsal view. Abbreviations: 2° AdShSp = secondary adoral shield spine; AdShSp = adoral shield spine; IPa = infradental papilla; LOPas = lateral oral papillae; LyOs = Lyman’s ossicle; OPRSp = oral plate ridge spine; RS = radial shield; TPa = tooth papilla; vT = ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.13310037" httpUri="https://zenodo.org/record/13310037/files/figure.png" pageId="25" pageNumber="155">Fig. 8D</figureCitation>
|
||
). DAPs wider than long, typically divided into two and up to five irregular pieces (
|
||
<figureCitation id="13618DD8FF90820553A4B06D6003FC24" box="[1258,1386,889,915]" captionStart="Fig" captionStartId="26.[189,232,1787,1813]" captionTargetBox="[319,1268,263,1746]" captionTargetId="figure-12@26.[319,1268,263,1746]" captionTargetPageId="26" captionText="Fig. 8. Ophioderma aija sp. nov., holotype (ICML-UNAM 18466, DD = 26.7 mm). A. Dorsal disc. B. Ventral disc (inset: section between the arm and the distal genital slit). C. Close-up of a jaw. D. Proximalmost dorsal arm. E. Median dorsal arm. F. Distalmost dorsal arm. G. Proximalmost ventral arm. H. Median ventral arm. I. Distalmost ventral arm. J. Lateral arm plates and arm spines. K. Dorsal view. Abbreviations: 2° AdShSp = secondary adoral shield spine; AdShSp = adoral shield spine; IPa = infradental papilla; LOPas = lateral oral papillae; LyOs = Lyman’s ossicle; OPRSp = oral plate ridge spine; RS = radial shield; TPa = tooth papilla; vT = ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.13310037" httpUri="https://zenodo.org/record/13310037/files/figure.png" pageId="25" pageNumber="155">Fig. 8D–E</figureCitation>
|
||
). DAP pieces sequence of the longest arm: first ten segments, 3, 3, 4, 5, 5, 5, 4, 4, 5, 5; 11
|
||
<superScript id="7C2F3C15FF90820553A5B08F6191FC1E" attach="none" box="[1259,1272,923,937]" fontSize="6" pageId="25" pageNumber="155">th</superScript>
|
||
–20
|
||
<superScript id="7C2F3C15FF908205526AB08F6058FC1E" attach="none" box="[1316,1329,923,937]" fontSize="6" pageId="25" pageNumber="155">th</superScript>
|
||
, 3–4; 21
|
||
<superScript id="7C2F3C15FF9082055794B0AB658CFC7A" attach="none" box="[218,229,959,973]" fontSize="6" pageId="25" pageNumber="155">st</superScript>
|
||
–30
|
||
<superScript id="7C2F3C15FF908205565CB0AA6476FC7B" attach="none" box="[274,287,958,972]" fontSize="6" pageId="25" pageNumber="155">th</superScript>
|
||
, 2–4; 31
|
||
<superScript id="7C2F3C15FF90820556C9B0AB64FBFC7A" attach="right" box="[391,402,959,973]" fontSize="6" pageId="25" pageNumber="155">st</superScript>
|
||
–40
|
||
<superScript id="7C2F3C15FF90820556F0B0AA64A2FC7B" attach="none" box="[446,459,958,972]" fontSize="6" pageId="25" pageNumber="155">th</superScript>
|
||
, 2–5; 41
|
||
<superScript id="7C2F3C15FF908205557DB0AB6757FC7A" attach="none" box="[563,574,959,973]" fontSize="6" pageId="25" pageNumber="155">st</superScript>
|
||
–50
|
||
<superScript id="7C2F3C15FF9082055525B0AA6711FC7B" attach="none" box="[619,632,958,972]" fontSize="6" pageId="25" pageNumber="155">th</superScript>
|
||
, 1–3; 51
|
||
<superScript id="7C2F3C15FF90820555AEB0AB6782FC7A" attach="right" box="[736,747,959,973]" fontSize="6" pageId="25" pageNumber="155">st</superScript>
|
||
–60
|
||
<superScript id="7C2F3C15FF9082055459B0AA664DFC7B" attach="none" box="[791,804,958,972]" fontSize="6" pageId="25" pageNumber="155">th</superScript>
|
||
, 1–2; 61
|
||
<superScript id="7C2F3C15FF90820554C2B0AB66FEFC7A" attach="none" box="[908,919,959,973]" fontSize="6" pageId="25" pageNumber="155">st</superScript>
|
||
–70
|
||
<superScript id="7C2F3C15FF908205548AB0AA66B8FC7B" attach="none" box="[964,977,958,972]" fontSize="6" pageId="25" pageNumber="155">th</superScript>
|
||
, 1–2; 71
|
||
<superScript id="7C2F3C15FF9082055377B0AB612DFC7A" attach="right" box="[1081,1092,959,973]" fontSize="6" pageId="25" pageNumber="155">st</superScript>
|
||
–74
|
||
<superScript id="7C2F3C15FF908205533EB0AA6114FC7B" attach="none" box="[1136,1149,958,972]" fontSize="6" pageId="25" pageNumber="155">th</superScript>
|
||
, 1. Distalmost DAPs trapezoidal to triangular, entire (
|
||
<figureCitation id="13618DD8FF908205550BB0F667CBFC4B" box="[581,674,994,1020]" captionStart="Fig" captionStartId="26.[189,232,1787,1813]" captionTargetBox="[319,1268,263,1746]" captionTargetId="figure-12@26.[319,1268,263,1746]" captionTargetPageId="26" captionText="Fig. 8. Ophioderma aija sp. nov., holotype (ICML-UNAM 18466, DD = 26.7 mm). A. Dorsal disc. B. Ventral disc (inset: section between the arm and the distal genital slit). C. Close-up of a jaw. D. Proximalmost dorsal arm. E. Median dorsal arm. F. Distalmost dorsal arm. G. Proximalmost ventral arm. H. Median ventral arm. I. Distalmost ventral arm. J. Lateral arm plates and arm spines. K. Dorsal view. Abbreviations: 2° AdShSp = secondary adoral shield spine; AdShSp = adoral shield spine; IPa = infradental papilla; LOPas = lateral oral papillae; LyOs = Lyman’s ossicle; OPRSp = oral plate ridge spine; RS = radial shield; TPa = tooth papilla; vT = ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.13310037" httpUri="https://zenodo.org/record/13310037/files/figure.png" pageId="25" pageNumber="155">Fig. 8F</figureCitation>
|
||
). First VAP small, 2× as wide as long, with rounded edges (
|
||
<figureCitation id="13618DD8FF9082055788B7116448FB97" box="[198,289,1029,1056]" captionStart="Fig" captionStartId="26.[189,232,1787,1813]" captionTargetBox="[319,1268,263,1746]" captionTargetId="figure-12@26.[319,1268,263,1746]" captionTargetPageId="26" captionText="Fig. 8. Ophioderma aija sp. nov., holotype (ICML-UNAM 18466, DD = 26.7 mm). A. Dorsal disc. B. Ventral disc (inset: section between the arm and the distal genital slit). C. Close-up of a jaw. D. Proximalmost dorsal arm. E. Median dorsal arm. F. Distalmost dorsal arm. G. Proximalmost ventral arm. H. Median ventral arm. I. Distalmost ventral arm. J. Lateral arm plates and arm spines. K. Dorsal view. Abbreviations: 2° AdShSp = secondary adoral shield spine; AdShSp = adoral shield spine; IPa = infradental papilla; LOPas = lateral oral papillae; LyOs = Lyman’s ossicle; OPRSp = oral plate ridge spine; RS = radial shield; TPa = tooth papilla; vT = ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.13310037" httpUri="https://zenodo.org/record/13310037/files/figure.png" pageId="25" pageNumber="155">Fig. 8B</figureCitation>
|
||
). Subsequent VAPs quadrangular, longer than wide; proximal edge straight, distal edge convex (
|
||
<figureCitation id="13618DD8FF908205578BB73D6423FBF4" box="[197,330,1065,1091]" captionStart="Fig" captionStartId="26.[189,232,1787,1813]" captionTargetBox="[319,1268,263,1746]" captionTargetId="figure-12@26.[319,1268,263,1746]" captionTargetPageId="26" captionText="Fig. 8. Ophioderma aija sp. nov., holotype (ICML-UNAM 18466, DD = 26.7 mm). A. Dorsal disc. B. Ventral disc (inset: section between the arm and the distal genital slit). C. Close-up of a jaw. D. Proximalmost dorsal arm. E. Median dorsal arm. F. Distalmost dorsal arm. G. Proximalmost ventral arm. H. Median ventral arm. I. Distalmost ventral arm. J. Lateral arm plates and arm spines. K. Dorsal view. Abbreviations: 2° AdShSp = secondary adoral shield spine; AdShSp = adoral shield spine; IPa = infradental papilla; LOPas = lateral oral papillae; LyOs = Lyman’s ossicle; OPRSp = oral plate ridge spine; RS = radial shield; TPa = tooth papilla; vT = ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.13310037" httpUri="https://zenodo.org/record/13310037/files/figure.png" pageId="25" pageNumber="155">Fig. 8G–H</figureCitation>
|
||
). Distalmost VAPs triangular, rounded, wider than long (
|
||
<figureCitation id="13618DD8FF90820554A1B73D612AFBF4" box="[1007,1091,1065,1091]" captionStart="Fig" captionStartId="26.[189,232,1787,1813]" captionTargetBox="[319,1268,263,1746]" captionTargetId="figure-12@26.[319,1268,263,1746]" captionTargetPageId="26" captionText="Fig. 8. Ophioderma aija sp. nov., holotype (ICML-UNAM 18466, DD = 26.7 mm). A. Dorsal disc. B. Ventral disc (inset: section between the arm and the distal genital slit). C. Close-up of a jaw. D. Proximalmost dorsal arm. E. Median dorsal arm. F. Distalmost dorsal arm. G. Proximalmost ventral arm. H. Median ventral arm. I. Distalmost ventral arm. J. Lateral arm plates and arm spines. K. Dorsal view. Abbreviations: 2° AdShSp = secondary adoral shield spine; AdShSp = adoral shield spine; IPa = infradental papilla; LOPas = lateral oral papillae; LyOs = Lyman’s ossicle; OPRSp = oral plate ridge spine; RS = radial shield; TPa = tooth papilla; vT = ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.13310037" httpUri="https://zenodo.org/record/13310037/files/figure.png" pageId="25" pageNumber="155">Fig. 8I</figureCitation>
|
||
). A pair of pores between the two proximalmost VAPs in all five arms (
|
||
<figureCitation id="13618DD8FF9082055599B758665BFBD1" box="[727,818,1100,1126]" captionStart="Fig" captionStartId="26.[189,232,1787,1813]" captionTargetBox="[319,1268,263,1746]" captionTargetId="figure-12@26.[319,1268,263,1746]" captionTargetPageId="26" captionText="Fig. 8. Ophioderma aija sp. nov., holotype (ICML-UNAM 18466, DD = 26.7 mm). A. Dorsal disc. B. Ventral disc (inset: section between the arm and the distal genital slit). C. Close-up of a jaw. D. Proximalmost dorsal arm. E. Median dorsal arm. F. Distalmost dorsal arm. G. Proximalmost ventral arm. H. Median ventral arm. I. Distalmost ventral arm. J. Lateral arm plates and arm spines. K. Dorsal view. Abbreviations: 2° AdShSp = secondary adoral shield spine; AdShSp = adoral shield spine; IPa = infradental papilla; LOPas = lateral oral papillae; LyOs = Lyman’s ossicle; OPRSp = oral plate ridge spine; RS = radial shield; TPa = tooth papilla; vT = ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.13310037" httpUri="https://zenodo.org/record/13310037/files/figure.png" pageId="25" pageNumber="155">Fig. 8B</figureCitation>
|
||
). LAPs conspicuous, wider than long, with up to ten arm spines. Arm spine sequence of the longest arm (right side, including arm spine bearing segments within the disc): first ten segments, 3, 3, 3, 4, 5, 5, 6, 7, 8, 9; 11
|
||
<superScript id="7C2F3C15FF90820554E1B78566D5FB28" attach="none" box="[943,956,1169,1183]" fontSize="6" pageId="25" pageNumber="155">th</superScript>
|
||
–20
|
||
<superScript id="7C2F3C15FF90820554A7B785669FFB28" attach="none" box="[1001,1014,1169,1183]" fontSize="6" pageId="25" pageNumber="155">th</superScript>
|
||
, 9–10; 21
|
||
<superScript id="7C2F3C15FF9082055320B7866110FB17" attach="right" box="[1134,1145,1170,1184]" fontSize="6" pageId="25" pageNumber="155">st</superScript>
|
||
–30
|
||
<superScript id="7C2F3C15FF90820553EBB78561DBFB28" attach="none" box="[1189,1202,1169,1183]" fontSize="6" pageId="25" pageNumber="155">th</superScript>
|
||
, 9–10; 31
|
||
<superScript id="7C2F3C15FF9082055264B786605CFB17" attach="none" box="[1322,1333,1170,1184]" fontSize="6" pageId="25" pageNumber="155">st</superScript>
|
||
–40
|
||
<superScript id="7C2F3C15FF908205522CB7856006FB28" attach="none" box="[1378,1391,1169,1183]" fontSize="6" pageId="25" pageNumber="155">th</superScript>
|
||
, 8–9; 41
|
||
<superScript id="7C2F3C15FF908205565BB7A16449FB74" attach="none" box="[277,288,1205,1219]" fontSize="6" pageId="25" pageNumber="155">st</superScript>
|
||
–50
|
||
<superScript id="7C2F3C15FF9082055603B7A16433FB74" attach="none" box="[333,346,1205,1219]" fontSize="6" pageId="25" pageNumber="155">th</superScript>
|
||
, 8; 51
|
||
<superScript id="7C2F3C15FF90820556EAB7A164C6FB74" attach="right" box="[420,431,1205,1219]" fontSize="6" pageId="25" pageNumber="155">st</superScript>
|
||
–60
|
||
<superScript id="7C2F3C15FF9082055695B7A16481FB74" attach="none" box="[475,488,1205,1219]" fontSize="6" pageId="25" pageNumber="155">th</superScript>
|
||
, 6–8; 61
|
||
<superScript id="7C2F3C15FF9082055501B7A16733FB74" attach="none" box="[591,602,1205,1219]" fontSize="6" pageId="25" pageNumber="155">st</superScript>
|
||
–70
|
||
<superScript id="7C2F3C15FF90820555C9B7A167FDFB74" attach="none" box="[647,660,1205,1219]" fontSize="6" pageId="25" pageNumber="155">th</superScript>
|
||
, 6–7; 71
|
||
<superScript id="7C2F3C15FF90820555B5B7A1666FFB74" attach="none" box="[763,774,1205,1219]" fontSize="6" pageId="25" pageNumber="155">st</superScript>
|
||
–80
|
||
<superScript id="7C2F3C15FF908205547CB7A16656FB74" attach="none" box="[818,831,1205,1219]" fontSize="6" pageId="25" pageNumber="155">th</superScript>
|
||
, 4–6; 81
|
||
<superScript id="7C2F3C15FF90820554E9B7A166DBFB74" attach="right" box="[935,946,1205,1219]" fontSize="6" pageId="25" pageNumber="155">st</superScript>
|
||
–83
|
||
<superScript id="7C2F3C15FF9082055490B7A16685FB74" attach="none" box="[990,1004,1205,1219]" fontSize="6" pageId="25" pageNumber="155">rd</superScript>
|
||
, 4. Arm spines conical with blunt tips, slightly flattened, ⅔ LAP length. Dorsalmost arm spine the shortest; ventralmost the longest and more robust, in contact with tentacle scales of the following segment (
|
||
<figureCitation id="13618DD8FF9082055358B7E86104FAA1" box="[1046,1133,1276,1302]" captionStart="Fig" captionStartId="26.[189,232,1787,1813]" captionTargetBox="[319,1268,263,1746]" captionTargetId="figure-12@26.[319,1268,263,1746]" captionTargetPageId="26" captionText="Fig. 8. Ophioderma aija sp. nov., holotype (ICML-UNAM 18466, DD = 26.7 mm). A. Dorsal disc. B. Ventral disc (inset: section between the arm and the distal genital slit). C. Close-up of a jaw. D. Proximalmost dorsal arm. E. Median dorsal arm. F. Distalmost dorsal arm. G. Proximalmost ventral arm. H. Median ventral arm. I. Distalmost ventral arm. J. Lateral arm plates and arm spines. K. Dorsal view. Abbreviations: 2° AdShSp = secondary adoral shield spine; AdShSp = adoral shield spine; IPa = infradental papilla; LOPas = lateral oral papillae; LyOs = Lyman’s ossicle; OPRSp = oral plate ridge spine; RS = radial shield; TPa = tooth papilla; vT = ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.13310037" httpUri="https://zenodo.org/record/13310037/files/figure.png" pageId="25" pageNumber="155">Fig. 8J</figureCitation>
|
||
). Two tentacle scales; adradial tentacle scale oval, elongated, just over ⅔ VAP length; abradial tentacle scale shorter and wider, ¾ adradial scale length, triangular (
|
||
<figureCitation id="13618DD8FF908205552FB656678CFAEB" box="[609,741,1346,1372]" captionStart="Fig" captionStartId="26.[189,232,1787,1813]" captionTargetBox="[319,1268,263,1746]" captionTargetId="figure-12@26.[319,1268,263,1746]" captionTargetPageId="26" captionText="Fig. 8. Ophioderma aija sp. nov., holotype (ICML-UNAM 18466, DD = 26.7 mm). A. Dorsal disc. B. Ventral disc (inset: section between the arm and the distal genital slit). C. Close-up of a jaw. D. Proximalmost dorsal arm. E. Median dorsal arm. F. Distalmost dorsal arm. G. Proximalmost ventral arm. H. Median ventral arm. I. Distalmost ventral arm. J. Lateral arm plates and arm spines. K. Dorsal view. Abbreviations: 2° AdShSp = secondary adoral shield spine; AdShSp = adoral shield spine; IPa = infradental papilla; LOPas = lateral oral papillae; LyOs = Lyman’s ossicle; OPRSp = oral plate ridge spine; RS = radial shield; TPa = tooth papilla; vT = ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.13310037" httpUri="https://zenodo.org/record/13310037/files/figure.png" pageId="25" pageNumber="155">Fig. 8G–H</figureCitation>
|
||
). In the distalmost arm section, tentacle scales oval and elongated, adradial being the longest; last arm segments with only one scale (
|
||
<figureCitation id="13618DD8FF9082055307B67161F2FA37" box="[1097,1179,1381,1408]" captionStart="Fig" captionStartId="26.[189,232,1787,1813]" captionTargetBox="[319,1268,263,1746]" captionTargetId="figure-12@26.[319,1268,263,1746]" captionTargetPageId="26" captionText="Fig. 8. Ophioderma aija sp. nov., holotype (ICML-UNAM 18466, DD = 26.7 mm). A. Dorsal disc. B. Ventral disc (inset: section between the arm and the distal genital slit). C. Close-up of a jaw. D. Proximalmost dorsal arm. E. Median dorsal arm. F. Distalmost dorsal arm. G. Proximalmost ventral arm. H. Median ventral arm. I. Distalmost ventral arm. J. Lateral arm plates and arm spines. K. Dorsal view. Abbreviations: 2° AdShSp = secondary adoral shield spine; AdShSp = adoral shield spine; IPa = infradental papilla; LOPas = lateral oral papillae; LyOs = Lyman’s ossicle; OPRSp = oral plate ridge spine; RS = radial shield; TPa = tooth papilla; vT = ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.13310037" httpUri="https://zenodo.org/record/13310037/files/figure.png" pageId="25" pageNumber="155">Fig. 8I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFF90820557F3B6B36176F9C6" blockId="25.[188,1399,1447,1649]" pageId="25" pageNumber="155">
|
||
General coloration dark brown (specimen in ethanol) (
|
||
<figureCitation id="13618DD8FF9082055417B6B366D2FA76" box="[857,955,1447,1473]" captionStart="Fig" captionStartId="26.[189,232,1787,1813]" captionTargetBox="[319,1268,263,1746]" captionTargetId="figure-12@26.[319,1268,263,1746]" captionTargetPageId="26" captionText="Fig. 8. Ophioderma aija sp. nov., holotype (ICML-UNAM 18466, DD = 26.7 mm). A. Dorsal disc. B. Ventral disc (inset: section between the arm and the distal genital slit). C. Close-up of a jaw. D. Proximalmost dorsal arm. E. Median dorsal arm. F. Distalmost dorsal arm. G. Proximalmost ventral arm. H. Median ventral arm. I. Distalmost ventral arm. J. Lateral arm plates and arm spines. K. Dorsal view. Abbreviations: 2° AdShSp = secondary adoral shield spine; AdShSp = adoral shield spine; IPa = infradental papilla; LOPas = lateral oral papillae; LyOs = Lyman’s ossicle; OPRSp = oral plate ridge spine; RS = radial shield; TPa = tooth papilla; vT = ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.13310037" httpUri="https://zenodo.org/record/13310037/files/figure.png" pageId="25" pageNumber="155">Fig. 8K</figureCitation>
|
||
). Dorsal side: disc dark brown with darker, sinuous rings; these may or may not continue over radial shields. Radial shields lighter brown (
|
||
<figureCitation id="13618DD8FF9082055789B6F9644DF9B0" box="[199,292,1517,1543]" captionStart="Fig" captionStartId="26.[189,232,1787,1813]" captionTargetBox="[319,1268,263,1746]" captionTargetId="figure-12@26.[319,1268,263,1746]" captionTargetPageId="26" captionText="Fig. 8. Ophioderma aija sp. nov., holotype (ICML-UNAM 18466, DD = 26.7 mm). A. Dorsal disc. B. Ventral disc (inset: section between the arm and the distal genital slit). C. Close-up of a jaw. D. Proximalmost dorsal arm. E. Median dorsal arm. F. Distalmost dorsal arm. G. Proximalmost ventral arm. H. Median ventral arm. I. Distalmost ventral arm. J. Lateral arm plates and arm spines. K. Dorsal view. Abbreviations: 2° AdShSp = secondary adoral shield spine; AdShSp = adoral shield spine; IPa = infradental papilla; LOPas = lateral oral papillae; LyOs = Lyman’s ossicle; OPRSp = oral plate ridge spine; RS = radial shield; TPa = tooth papilla; vT = ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.13310037" httpUri="https://zenodo.org/record/13310037/files/figure.png" pageId="25" pageNumber="155">Fig. 8A</figureCitation>
|
||
). Arms brown, lighter than disc (
|
||
<figureCitation id="13618DD8FF90820555EBB6F9662AF9B0" box="[677,835,1517,1543]" captionStart="Fig" captionStartId="26.[189,232,1787,1813]" captionTargetBox="[319,1268,263,1746]" captionTargetId="figure-12@26.[319,1268,263,1746]" captionTargetPageId="26" captionText="Fig. 8. Ophioderma aija sp. nov., holotype (ICML-UNAM 18466, DD = 26.7 mm). A. Dorsal disc. B. Ventral disc (inset: section between the arm and the distal genital slit). C. Close-up of a jaw. D. Proximalmost dorsal arm. E. Median dorsal arm. F. Distalmost dorsal arm. G. Proximalmost ventral arm. H. Median ventral arm. I. Distalmost ventral arm. J. Lateral arm plates and arm spines. K. Dorsal view. Abbreviations: 2° AdShSp = secondary adoral shield spine; AdShSp = adoral shield spine; IPa = infradental papilla; LOPas = lateral oral papillae; LyOs = Lyman’s ossicle; OPRSp = oral plate ridge spine; RS = radial shield; TPa = tooth papilla; vT = ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.13310037" httpUri="https://zenodo.org/record/13310037/files/figure.png" pageId="25" pageNumber="155">Fig. 8D–F, K</figureCitation>
|
||
). Ventral side: interradii lighter than dorsal disc, with dark, sinuous rings (
|
||
<figureCitation id="13618DD8FF90820556ACB5046754F99C" box="[482,573,1552,1579]" captionStart="Fig" captionStartId="26.[189,232,1787,1813]" captionTargetBox="[319,1268,263,1746]" captionTargetId="figure-12@26.[319,1268,263,1746]" captionTargetPageId="26" captionText="Fig. 8. Ophioderma aija sp. nov., holotype (ICML-UNAM 18466, DD = 26.7 mm). A. Dorsal disc. B. Ventral disc (inset: section between the arm and the distal genital slit). C. Close-up of a jaw. D. Proximalmost dorsal arm. E. Median dorsal arm. F. Distalmost dorsal arm. G. Proximalmost ventral arm. H. Median ventral arm. I. Distalmost ventral arm. J. Lateral arm plates and arm spines. K. Dorsal view. Abbreviations: 2° AdShSp = secondary adoral shield spine; AdShSp = adoral shield spine; IPa = infradental papilla; LOPas = lateral oral papillae; LyOs = Lyman’s ossicle; OPRSp = oral plate ridge spine; RS = radial shield; TPa = tooth papilla; vT = ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.13310037" httpUri="https://zenodo.org/record/13310037/files/figure.png" pageId="25" pageNumber="155">Fig. 8B</figureCitation>
|
||
). Oral shields light brown, two with small, darker specks; oral papillae, teeth, and arms cream (
|
||
<figureCitation id="13618DD8FF908205569DB520673AF9F9" box="[467,595,1588,1614]" captionStart="Fig" captionStartId="26.[189,232,1787,1813]" captionTargetBox="[319,1268,263,1746]" captionTargetId="figure-12@26.[319,1268,263,1746]" captionTargetPageId="26" captionText="Fig. 8. Ophioderma aija sp. nov., holotype (ICML-UNAM 18466, DD = 26.7 mm). A. Dorsal disc. B. Ventral disc (inset: section between the arm and the distal genital slit). C. Close-up of a jaw. D. Proximalmost dorsal arm. E. Median dorsal arm. F. Distalmost dorsal arm. G. Proximalmost ventral arm. H. Median ventral arm. I. Distalmost ventral arm. J. Lateral arm plates and arm spines. K. Dorsal view. Abbreviations: 2° AdShSp = secondary adoral shield spine; AdShSp = adoral shield spine; IPa = infradental papilla; LOPas = lateral oral papillae; LyOs = Lyman’s ossicle; OPRSp = oral plate ridge spine; RS = radial shield; TPa = tooth papilla; vT = ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.13310037" httpUri="https://zenodo.org/record/13310037/files/figure.png" pageId="25" pageNumber="155">Fig. 8B–C</figureCitation>
|
||
). Ventral arms darkening towards the distalmost arm section. LAPs light brown. Arm spines light brown; the ventralmost the lighter (
|
||
<figureCitation id="13618DD8FF90820554F0B543617BF9C6" box="[958,1042,1623,1649]" captionStart="Fig" captionStartId="26.[189,232,1787,1813]" captionTargetBox="[319,1268,263,1746]" captionTargetId="figure-12@26.[319,1268,263,1746]" captionTargetPageId="26" captionText="Fig. 8. Ophioderma aija sp. nov., holotype (ICML-UNAM 18466, DD = 26.7 mm). A. Dorsal disc. B. Ventral disc (inset: section between the arm and the distal genital slit). C. Close-up of a jaw. D. Proximalmost dorsal arm. E. Median dorsal arm. F. Distalmost dorsal arm. G. Proximalmost ventral arm. H. Median ventral arm. I. Distalmost ventral arm. J. Lateral arm plates and arm spines. K. Dorsal view. Abbreviations: 2° AdShSp = secondary adoral shield spine; AdShSp = adoral shield spine; IPa = infradental papilla; LOPas = lateral oral papillae; LyOs = Lyman’s ossicle; OPRSp = oral plate ridge spine; RS = radial shield; TPa = tooth papilla; vT = ventralmost tooth." figureDoi="http://doi.org/10.5281/zenodo.13310037" httpUri="https://zenodo.org/record/13310037/files/figure.png" pageId="25" pageNumber="155">Fig. 8J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFF90820557F3B58C64B8F905" blockId="25.[189,465,1688,1714]" box="[189,465,1688,1714]" pageId="25" pageNumber="155">
|
||
<heading id="D0AD2631FF90820557F3B58C64B8F905" bold="true" box="[189,465,1688,1714]" fontSize="11" level="3" pageId="25" pageNumber="155" reason="3">
|
||
<emphasis id="B92E4D4FFF90820557F3B58C64B8F905" bold="true" box="[189,465,1688,1714]" pageId="25" pageNumber="155">Disarticulated ossicles</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFF90820757F3B5D36407FC2A" blockId="25.[189,1400,1735,2008]" lastBlockId="27.[188,1401,265,925]" lastPageId="27" lastPageNumber="157" pageId="25" pageNumber="155">
|
||
Non-type specimen, ICML-UNAM 18457 (DD =
|
||
<quantity id="4CA23CB8FF908205546EB5D366F9F956" box="[800,912,1735,1761]" metricMagnitude="-2" metricUnit="m" metricValue="2.62" pageId="25" pageNumber="155" unit="mm" value="26.2">26.2 mm</quantity>
|
||
, AL =
|
||
<quantity id="4CA23CB8FF90820554A5B5D36133F956" box="[1003,1114,1735,1761]" metricMagnitude="-2" metricUnit="m" metricValue="8.19" pageId="25" pageNumber="155" unit="mm" value="81.9">81.9 mm</quantity>
|
||
, AL:DD = 3.1). Radial shields irregularly triangular, naked in the intact animal; proximal edge convex; distal edge unequally convex; adradial edge irregular, slightly convex in the middle; abradial edge with two well-defined processes, distal prominent (
|
||
<figureCitation id="13618DD8FF9082055542B42467E7F8FC" box="[524,654,1840,1867]" captionStart="Fig" captionStartId="28.[189,232,1716,1742]" captionTargetBox="[344,1244,265,1671]" captionTargetId="figure-12@28.[344,1244,265,1671]" captionTargetPageId="28" captionText="Fig. 9. Ophioderma aija sp. nov., non-type specimen (ICML-UNAM 18457, DD = 26.2 mm), SEM images of the ossicles. A. Radial shield, external face. The arrowhead indicates the naked section observed in the intact animal. B. Radial shield, internal face. C. Dental plate. D. Adradial genital plate, abradial face. E. Adradial genital plate, adradial face. F. Oral plate, abradial face. G. Oral plate, adradial face. Black arrows point to oral papillae sockets, white arrows point to granule sockets. H. Arm vertebra, proximal face. I. Arm vertebra, distal face. J. Ventral arm plate, external face. K. Ventral arm plate, internal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. N. Lateral arm plate, lateral face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; SA = spine articulation; Sp = spur; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.13310039" httpUri="https://zenodo.org/record/13310039/files/figure.png" pageId="25" pageNumber="155">Fig. 9A–B</figureCitation>
|
||
). Externally, distal half swollen, with a raised oval in the center corresponding to the naked section in the intact animal; oval surface surrounded by numerous small pores; scattered larger pores placed near edges and proximal section (
|
||
<figureCitation id="13618DD8FF9082055348B463610EF826" box="[1030,1127,1911,1937]" captionStart="Fig" captionStartId="28.[189,232,1716,1742]" captionTargetBox="[344,1244,265,1671]" captionTargetId="figure-12@28.[344,1244,265,1671]" captionTargetPageId="28" captionText="Fig. 9. Ophioderma aija sp. nov., non-type specimen (ICML-UNAM 18457, DD = 26.2 mm), SEM images of the ossicles. A. Radial shield, external face. The arrowhead indicates the naked section observed in the intact animal. B. Radial shield, internal face. C. Dental plate. D. Adradial genital plate, abradial face. E. Adradial genital plate, adradial face. F. Oral plate, abradial face. G. Oral plate, adradial face. Black arrows point to oral papillae sockets, white arrows point to granule sockets. H. Arm vertebra, proximal face. I. Arm vertebra, distal face. J. Ventral arm plate, external face. K. Ventral arm plate, internal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. N. Lateral arm plate, lateral face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; SA = spine articulation; Sp = spur; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.13310039" httpUri="https://zenodo.org/record/13310039/files/figure.png" pageId="25" pageNumber="155">Fig. 9A</figureCitation>
|
||
). Internally, distal half center with three median pores; close to distal edge, two rounded truncated bulbs slightly separated, adradial one larger, followed by a furrow (
|
||
<figureCitation id="13618DD8FF90820555FBB4A9667AF860" box="[693,787,1981,2007]" captionStart="Fig" captionStartId="28.[189,232,1716,1742]" captionTargetBox="[344,1244,265,1671]" captionTargetId="figure-12@28.[344,1244,265,1671]" captionTargetPageId="28" captionText="Fig. 9. Ophioderma aija sp. nov., non-type specimen (ICML-UNAM 18457, DD = 26.2 mm), SEM images of the ossicles. A. Radial shield, external face. The arrowhead indicates the naked section observed in the intact animal. B. Radial shield, internal face. C. Dental plate. D. Adradial genital plate, abradial face. E. Adradial genital plate, adradial face. F. Oral plate, abradial face. G. Oral plate, adradial face. Black arrows point to oral papillae sockets, white arrows point to granule sockets. H. Arm vertebra, proximal face. I. Arm vertebra, distal face. J. Ventral arm plate, external face. K. Ventral arm plate, internal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. N. Lateral arm plate, lateral face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; SA = spine articulation; Sp = spur; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.13310039" httpUri="https://zenodo.org/record/13310039/files/figure.png" pageId="25" pageNumber="155">Fig. 9B</figureCitation>
|
||
). Dental plate fragmented into several pieces (up to five), bearing one or two teeth in oval or round non-penetrating sockets; ventralmost piece also with round sockets for TPa (
|
||
<figureCitation id="13618DD8FF92820756CBB238648AFEF1" box="[389,483,300,326]" captionStart="Fig" captionStartId="28.[189,232,1716,1742]" captionTargetBox="[344,1244,265,1671]" captionTargetId="figure-12@28.[344,1244,265,1671]" captionTargetPageId="28" captionText="Fig. 9. Ophioderma aija sp. nov., non-type specimen (ICML-UNAM 18457, DD = 26.2 mm), SEM images of the ossicles. A. Radial shield, external face. The arrowhead indicates the naked section observed in the intact animal. B. Radial shield, internal face. C. Dental plate. D. Adradial genital plate, abradial face. E. Adradial genital plate, adradial face. F. Oral plate, abradial face. G. Oral plate, adradial face. Black arrows point to oral papillae sockets, white arrows point to granule sockets. H. Arm vertebra, proximal face. I. Arm vertebra, distal face. J. Ventral arm plate, external face. K. Ventral arm plate, internal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. N. Lateral arm plate, lateral face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; SA = spine articulation; Sp = spur; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.13310039" httpUri="https://zenodo.org/record/13310039/files/figure.png" pageId="27" pageNumber="157">Fig. 9C</figureCitation>
|
||
). Adradial genital plate falcate, elongated. Abradial face slightly depressed in the middle, longitudinally, with three pores close to distal edge; distal edge with two truncated knobs and a distal depression, noticed from the other side (
|
||
<figureCitation id="13618DD8FF9282075469B26666EFFE3A" box="[807,902,370,397]" captionStart="Fig" captionStartId="28.[189,232,1716,1742]" captionTargetBox="[344,1244,265,1671]" captionTargetId="figure-12@28.[344,1244,265,1671]" captionTargetPageId="28" captionText="Fig. 9. Ophioderma aija sp. nov., non-type specimen (ICML-UNAM 18457, DD = 26.2 mm), SEM images of the ossicles. A. Radial shield, external face. The arrowhead indicates the naked section observed in the intact animal. B. Radial shield, internal face. C. Dental plate. D. Adradial genital plate, abradial face. E. Adradial genital plate, adradial face. F. Oral plate, abradial face. G. Oral plate, adradial face. Black arrows point to oral papillae sockets, white arrows point to granule sockets. H. Arm vertebra, proximal face. I. Arm vertebra, distal face. J. Ventral arm plate, external face. K. Ventral arm plate, internal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. N. Lateral arm plate, lateral face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; SA = spine articulation; Sp = spur; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.13310039" httpUri="https://zenodo.org/record/13310039/files/figure.png" pageId="27" pageNumber="157">Fig. 9D</figureCitation>
|
||
). Adradial face with a longitudinal groove and a notorious pore close to distal section. Distal edge rounded, with a lateral depression followed by a knob (
|
||
<figureCitation id="13618DD8FF928207566CB2AD64E9FE64" box="[290,384,441,467]" captionStart="Fig" captionStartId="28.[189,232,1716,1742]" captionTargetBox="[344,1244,265,1671]" captionTargetId="figure-12@28.[344,1244,265,1671]" captionTargetPageId="28" captionText="Fig. 9. Ophioderma aija sp. nov., non-type specimen (ICML-UNAM 18457, DD = 26.2 mm), SEM images of the ossicles. A. Radial shield, external face. The arrowhead indicates the naked section observed in the intact animal. B. Radial shield, internal face. C. Dental plate. D. Adradial genital plate, abradial face. E. Adradial genital plate, adradial face. F. Oral plate, abradial face. G. Oral plate, adradial face. Black arrows point to oral papillae sockets, white arrows point to granule sockets. H. Arm vertebra, proximal face. I. Arm vertebra, distal face. J. Ventral arm plate, external face. K. Ventral arm plate, internal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. N. Lateral arm plate, lateral face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; SA = spine articulation; Sp = spur; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.13310039" httpUri="https://zenodo.org/record/13310039/files/figure.png" pageId="27" pageNumber="157">Fig. 9E</figureCitation>
|
||
). Oral plates longer than high, middle section slightly lower than ends (
|
||
<figureCitation id="13618DD8FF92820753A7B2AD6003FE64" box="[1257,1386,441,467]" captionStart="Fig" captionStartId="28.[189,232,1716,1742]" captionTargetBox="[344,1244,265,1671]" captionTargetId="figure-12@28.[344,1244,265,1671]" captionTargetPageId="28" captionText="Fig. 9. Ophioderma aija sp. nov., non-type specimen (ICML-UNAM 18457, DD = 26.2 mm), SEM images of the ossicles. A. Radial shield, external face. The arrowhead indicates the naked section observed in the intact animal. B. Radial shield, internal face. C. Dental plate. D. Adradial genital plate, abradial face. E. Adradial genital plate, adradial face. F. Oral plate, abradial face. G. Oral plate, adradial face. Black arrows point to oral papillae sockets, white arrows point to granule sockets. H. Arm vertebra, proximal face. I. Arm vertebra, distal face. J. Ventral arm plate, external face. K. Ventral arm plate, internal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. N. Lateral arm plate, lateral face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; SA = spine articulation; Sp = spur; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.13310039" httpUri="https://zenodo.org/record/13310039/files/figure.png" pageId="27" pageNumber="157">Fig. 9F–G</figureCitation>
|
||
); abradial face with muscle fossa irregularly oval, elongated (
|
||
<figureCitation id="13618DD8FF92820754CFB2C866B2FE41" box="[897,987,476,502]" captionStart="Fig" captionStartId="28.[189,232,1716,1742]" captionTargetBox="[344,1244,265,1671]" captionTargetId="figure-12@28.[344,1244,265,1671]" captionTargetPageId="28" captionText="Fig. 9. Ophioderma aija sp. nov., non-type specimen (ICML-UNAM 18457, DD = 26.2 mm), SEM images of the ossicles. A. Radial shield, external face. The arrowhead indicates the naked section observed in the intact animal. B. Radial shield, internal face. C. Dental plate. D. Adradial genital plate, abradial face. E. Adradial genital plate, adradial face. F. Oral plate, abradial face. G. Oral plate, adradial face. Black arrows point to oral papillae sockets, white arrows point to granule sockets. H. Arm vertebra, proximal face. I. Arm vertebra, distal face. J. Ventral arm plate, external face. K. Ventral arm plate, internal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. N. Lateral arm plate, lateral face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; SA = spine articulation; Sp = spur; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.13310039" httpUri="https://zenodo.org/record/13310039/files/figure.png" pageId="27" pageNumber="157">Fig. 9F</figureCitation>
|
||
); adradial face with multiple pores at proximoventral edge of plate corresponding to oral papillae (lateral) and granules (ventral) sockets (
|
||
<figureCitation id="13618DD8FF9282075786B136644CFD8A" box="[200,293,546,573]" captionStart="Fig" captionStartId="28.[189,232,1716,1742]" captionTargetBox="[344,1244,265,1671]" captionTargetId="figure-12@28.[344,1244,265,1671]" captionTargetPageId="28" captionText="Fig. 9. Ophioderma aija sp. nov., non-type specimen (ICML-UNAM 18457, DD = 26.2 mm), SEM images of the ossicles. A. Radial shield, external face. The arrowhead indicates the naked section observed in the intact animal. B. Radial shield, internal face. C. Dental plate. D. Adradial genital plate, abradial face. E. Adradial genital plate, adradial face. F. Oral plate, abradial face. G. Oral plate, adradial face. Black arrows point to oral papillae sockets, white arrows point to granule sockets. H. Arm vertebra, proximal face. I. Arm vertebra, distal face. J. Ventral arm plate, external face. K. Ventral arm plate, internal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. N. Lateral arm plate, lateral face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; SA = spine articulation; Sp = spur; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.13310039" httpUri="https://zenodo.org/record/13310039/files/figure.png" pageId="27" pageNumber="157">Fig. 9G</figureCitation>
|
||
). Vertebrae zygospondylus (
|
||
<figureCitation id="13618DD8FF9282075523B136678AFD8A" box="[621,739,546,573]" captionStart="Fig" captionStartId="28.[189,232,1716,1742]" captionTargetBox="[344,1244,265,1671]" captionTargetId="figure-12@28.[344,1244,265,1671]" captionTargetPageId="28" captionText="Fig. 9. Ophioderma aija sp. nov., non-type specimen (ICML-UNAM 18457, DD = 26.2 mm), SEM images of the ossicles. A. Radial shield, external face. The arrowhead indicates the naked section observed in the intact animal. B. Radial shield, internal face. C. Dental plate. D. Adradial genital plate, abradial face. E. Adradial genital plate, adradial face. F. Oral plate, abradial face. G. Oral plate, adradial face. Black arrows point to oral papillae sockets, white arrows point to granule sockets. H. Arm vertebra, proximal face. I. Arm vertebra, distal face. J. Ventral arm plate, external face. K. Ventral arm plate, internal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. N. Lateral arm plate, lateral face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; SA = spine articulation; Sp = spur; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.13310039" httpUri="https://zenodo.org/record/13310039/files/figure.png" pageId="27" pageNumber="157">Fig. 9H–I</figureCitation>
|
||
). Proximal vertebrae wider than long, with dorsal muscle fossae larger than ventral ones (
|
||
<figureCitation id="13618DD8FF928207557DB15267F8FDD7" box="[563,657,582,608]" captionStart="Fig" captionStartId="28.[189,232,1716,1742]" captionTargetBox="[344,1244,265,1671]" captionTargetId="figure-12@28.[344,1244,265,1671]" captionTargetPageId="28" captionText="Fig. 9. Ophioderma aija sp. nov., non-type specimen (ICML-UNAM 18457, DD = 26.2 mm), SEM images of the ossicles. A. Radial shield, external face. The arrowhead indicates the naked section observed in the intact animal. B. Radial shield, internal face. C. Dental plate. D. Adradial genital plate, abradial face. E. Adradial genital plate, adradial face. F. Oral plate, abradial face. G. Oral plate, adradial face. Black arrows point to oral papillae sockets, white arrows point to granule sockets. H. Arm vertebra, proximal face. I. Arm vertebra, distal face. J. Ventral arm plate, external face. K. Ventral arm plate, internal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. N. Lateral arm plate, lateral face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; SA = spine articulation; Sp = spur; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.13310039" httpUri="https://zenodo.org/record/13310039/files/figure.png" pageId="27" pageNumber="157">Fig. 9H</figureCitation>
|
||
). VAPs (from proximal arm section) quadrangular, longer than wide; proximal edge with three points, the median one corresponding to a spur; lateral edges with two points forming concave areas; distal edge convex (
|
||
<figureCitation id="13618DD8FF9282075444B1986634FD11" box="[778,861,652,678]" captionStart="Fig" captionStartId="28.[189,232,1716,1742]" captionTargetBox="[344,1244,265,1671]" captionTargetId="figure-12@28.[344,1244,265,1671]" captionTargetPageId="28" captionText="Fig. 9. Ophioderma aija sp. nov., non-type specimen (ICML-UNAM 18457, DD = 26.2 mm), SEM images of the ossicles. A. Radial shield, external face. The arrowhead indicates the naked section observed in the intact animal. B. Radial shield, internal face. C. Dental plate. D. Adradial genital plate, abradial face. E. Adradial genital plate, adradial face. F. Oral plate, abradial face. G. Oral plate, adradial face. Black arrows point to oral papillae sockets, white arrows point to granule sockets. H. Arm vertebra, proximal face. I. Arm vertebra, distal face. J. Ventral arm plate, external face. K. Ventral arm plate, internal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. N. Lateral arm plate, lateral face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; SA = spine articulation; Sp = spur; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.13310039" httpUri="https://zenodo.org/record/13310039/files/figure.png" pageId="27" pageNumber="157">Fig. 9J</figureCitation>
|
||
). Internal face with three spurs, two elongated and lateral, and one smaller, oval in the middle (
|
||
<figureCitation id="13618DD8FF92820755A2B1BB6622FD7E" box="[748,843,687,713]" captionStart="Fig" captionStartId="28.[189,232,1716,1742]" captionTargetBox="[344,1244,265,1671]" captionTargetId="figure-12@28.[344,1244,265,1671]" captionTargetPageId="28" captionText="Fig. 9. Ophioderma aija sp. nov., non-type specimen (ICML-UNAM 18457, DD = 26.2 mm), SEM images of the ossicles. A. Radial shield, external face. The arrowhead indicates the naked section observed in the intact animal. B. Radial shield, internal face. C. Dental plate. D. Adradial genital plate, abradial face. E. Adradial genital plate, adradial face. F. Oral plate, abradial face. G. Oral plate, adradial face. Black arrows point to oral papillae sockets, white arrows point to granule sockets. H. Arm vertebra, proximal face. I. Arm vertebra, distal face. J. Ventral arm plate, external face. K. Ventral arm plate, internal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. N. Lateral arm plate, lateral face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; SA = spine articulation; Sp = spur; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.13310039" httpUri="https://zenodo.org/record/13310039/files/figure.png" pageId="27" pageNumber="157">Fig. 9K</figureCitation>
|
||
). LAPs curved, 2× as high as wide; dorsal edge straight; ventral edge convex, with a small, rounded condyle developing from internal side; proximal edge concave; distal edge convex (
|
||
<figureCitation id="13618DD8FF9282075524B1E26787FCA7" box="[618,750,758,784]" captionStart="Fig" captionStartId="28.[189,232,1716,1742]" captionTargetBox="[344,1244,265,1671]" captionTargetId="figure-12@28.[344,1244,265,1671]" captionTargetPageId="28" captionText="Fig. 9. Ophioderma aija sp. nov., non-type specimen (ICML-UNAM 18457, DD = 26.2 mm), SEM images of the ossicles. A. Radial shield, external face. The arrowhead indicates the naked section observed in the intact animal. B. Radial shield, internal face. C. Dental plate. D. Adradial genital plate, abradial face. E. Adradial genital plate, adradial face. F. Oral plate, abradial face. G. Oral plate, adradial face. Black arrows point to oral papillae sockets, white arrows point to granule sockets. H. Arm vertebra, proximal face. I. Arm vertebra, distal face. J. Ventral arm plate, external face. K. Ventral arm plate, internal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. N. Lateral arm plate, lateral face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; SA = spine articulation; Sp = spur; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.13310039" httpUri="https://zenodo.org/record/13310039/files/figure.png" pageId="27" pageNumber="157">Fig. 9L–N</figureCitation>
|
||
). Proximal external LAP edge with two conspicuous, elongated, and triangular spurs in the middle (
|
||
<figureCitation id="13618DD8FF92820755AFB00D6654FC84" box="[737,829,793,819]" captionStart="Fig" captionStartId="28.[189,232,1716,1742]" captionTargetBox="[344,1244,265,1671]" captionTargetId="figure-12@28.[344,1244,265,1671]" captionTargetPageId="28" captionText="Fig. 9. Ophioderma aija sp. nov., non-type specimen (ICML-UNAM 18457, DD = 26.2 mm), SEM images of the ossicles. A. Radial shield, external face. The arrowhead indicates the naked section observed in the intact animal. B. Radial shield, internal face. C. Dental plate. D. Adradial genital plate, abradial face. E. Adradial genital plate, adradial face. F. Oral plate, abradial face. G. Oral plate, adradial face. Black arrows point to oral papillae sockets, white arrows point to granule sockets. H. Arm vertebra, proximal face. I. Arm vertebra, distal face. J. Ventral arm plate, external face. K. Ventral arm plate, internal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. N. Lateral arm plate, lateral face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; SA = spine articulation; Sp = spur; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.13310039" httpUri="https://zenodo.org/record/13310039/files/figure.png" pageId="27" pageNumber="157">Fig. 9L</figureCitation>
|
||
), having their counterparts internally (
|
||
<figureCitation id="13618DD8FF928207524BB00D6003FC84" box="[1285,1386,793,819]" captionStart="Fig" captionStartId="28.[189,232,1716,1742]" captionTargetBox="[344,1244,265,1671]" captionTargetId="figure-12@28.[344,1244,265,1671]" captionTargetPageId="28" captionText="Fig. 9. Ophioderma aija sp. nov., non-type specimen (ICML-UNAM 18457, DD = 26.2 mm), SEM images of the ossicles. A. Radial shield, external face. The arrowhead indicates the naked section observed in the intact animal. B. Radial shield, internal face. C. Dental plate. D. Adradial genital plate, abradial face. E. Adradial genital plate, adradial face. F. Oral plate, abradial face. G. Oral plate, adradial face. Black arrows point to oral papillae sockets, white arrows point to granule sockets. H. Arm vertebra, proximal face. I. Arm vertebra, distal face. J. Ventral arm plate, external face. K. Ventral arm plate, internal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. N. Lateral arm plate, lateral face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; SA = spine articulation; Sp = spur; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.13310039" httpUri="https://zenodo.org/record/13310039/files/figure.png" pageId="27" pageNumber="157">Fig. 9M</figureCitation>
|
||
). Internal side with four pores near center, concave proximal ridge, two separated bulbs near ventral edge, ventralmost protruding from plate. Ten spine articulations on distal edge, each surrounded by a thick lobe (
|
||
<figureCitation id="13618DD8FF928207564CB0966408FC2A" box="[258,353,898,925]" captionStart="Fig" captionStartId="28.[189,232,1716,1742]" captionTargetBox="[344,1244,265,1671]" captionTargetId="figure-12@28.[344,1244,265,1671]" captionTargetPageId="28" captionText="Fig. 9. Ophioderma aija sp. nov., non-type specimen (ICML-UNAM 18457, DD = 26.2 mm), SEM images of the ossicles. A. Radial shield, external face. The arrowhead indicates the naked section observed in the intact animal. B. Radial shield, internal face. C. Dental plate. D. Adradial genital plate, abradial face. E. Adradial genital plate, adradial face. F. Oral plate, abradial face. G. Oral plate, adradial face. Black arrows point to oral papillae sockets, white arrows point to granule sockets. H. Arm vertebra, proximal face. I. Arm vertebra, distal face. J. Ventral arm plate, external face. K. Ventral arm plate, internal face. L. Lateral arm plate, external face. M. Lateral arm plate, internal face. N. Lateral arm plate, lateral face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; SA = spine articulation; Sp = spur; vent = ventral." figureDoi="http://doi.org/10.5281/zenodo.13310039" httpUri="https://zenodo.org/record/13310039/files/figure.png" pageId="27" pageNumber="157">Fig. 9N</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF25C1D5FF93820657F3B5EF669CF85F" ID-DOI="http://doi.org/10.5281/zenodo.13310037" ID-Zenodo-Dep="13310037" httpUri="https://zenodo.org/record/13310037/files/figure.png" pageId="26" pageNumber="156" startId="26.[189,232,1787,1813]" targetBox="[319,1268,263,1746]" targetPageId="26">
|
||
<paragraph id="8BE5915DFF93820657F3B5EF669CF85F" blockId="26.[189,1399,1787,2025]" pageId="26" pageNumber="156">
|
||
<emphasis id="B92E4D4FFF93820657F3B5EF6467F8A2" bold="true" box="[189,270,1787,1813]" pageId="26" pageNumber="156">Fig. 8.</emphasis>
|
||
<taxonomicName id="4C5AEADEFF9382065657B5EF648CF8A2" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" box="[281,485,1787,1813]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="26" pageNumber="156" phylum="Echinodermata" rank="species" species="aija" status="sp. nov.">
|
||
<emphasis id="B92E4D4FFF9382065657B5EF648CF8A2" box="[281,485,1787,1813]" italics="true" pageId="26" pageNumber="156">Ophioderma aija</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21DF034FF93820656A1B5E86726F8A1" box="[495,591,1788,1814]" pageId="26" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (ICML-UNAM 18466, DD = 26.7 mm).
|
||
<emphasis id="B92E4D4FFF938206538FB5EF61BFF8A2" bold="true" box="[1217,1238,1787,1813]" pageId="26" pageNumber="156">A</emphasis>
|
||
. Dorsal disc.
|
||
<emphasis id="B92E4D4FFF93820657F3B40A65B8F88F" bold="true" box="[189,209,1822,1848]" pageId="26" pageNumber="156">B</emphasis>
|
||
. Ventral disc (inset: section between the arm and the distal genital slit).
|
||
<emphasis id="B92E4D4FFF9382065329B40A6115F88F" bold="true" box="[1127,1148,1822,1848]" pageId="26" pageNumber="156">C</emphasis>
|
||
. Close-up of a jaw.
|
||
<emphasis id="B92E4D4FFF93820657F3B45665BBF8EB" bold="true" box="[189,210,1858,1884]" pageId="26" pageNumber="156">D</emphasis>
|
||
. Proximalmost dorsal arm.
|
||
<emphasis id="B92E4D4FFF9382065558B4566743F8EB" bold="true" box="[534,554,1858,1884]" pageId="26" pageNumber="156">E</emphasis>
|
||
. Median dorsal arm.
|
||
<emphasis id="B92E4D4FFF938206546DB456665CF8EB" bold="true" box="[803,821,1858,1884]" pageId="26" pageNumber="156">F</emphasis>
|
||
. Distalmost dorsal arm.
|
||
<emphasis id="B92E4D4FFF938206531AB4556102F8EC" bold="true" box="[1108,1131,1857,1883]" pageId="26" pageNumber="156">G</emphasis>
|
||
. Proximalmost ventral arm.
|
||
<emphasis id="B92E4D4FFF93820657B7B4716479F8C8" bold="true" box="[249,272,1893,1919]" pageId="26" pageNumber="156">H</emphasis>
|
||
. Median ventral arm.
|
||
<emphasis id="B92E4D4FFF938206555BB4716749F8C8" bold="true" box="[533,544,1893,1919]" pageId="26" pageNumber="156">I</emphasis>
|
||
. Distalmost ventral arm.
|
||
<emphasis id="B92E4D4FFF9382065405B4716633F8C8" bold="true" box="[843,858,1893,1919]" pageId="26" pageNumber="156">J</emphasis>
|
||
. Lateral arm plates and arm spines.
|
||
<emphasis id="B92E4D4FFF938206524DB4716073F8C8" bold="true" box="[1283,1306,1893,1919]" pageId="26" pageNumber="156">K</emphasis>
|
||
. Dorsal view. Abbreviations: 2° AdShSp = secondary adoral shield spine; AdShSp = adoral shield spine; IPa = infradental papilla; LOPas = lateral oral papillae; LyOs = Lyman’s ossicle; OPRSp = oral plate ridge spine; RS = radial shield; TPa = tooth papilla; vT = ventralmost tooth.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE5915DFF92820757F3B0D06709FC69" blockId="27.[189,608,964,990]" box="[189,608,964,990]" pageId="27" pageNumber="157">
|
||
<heading id="D0AD2631FF92820757F3B0D06709FC69" bold="true" box="[189,608,964,990]" fontSize="11" level="3" pageId="27" pageNumber="157" reason="3">
|
||
<emphasis id="B92E4D4FFF92820757F3B0D06709FC69" bold="true" box="[189,608,964,990]" pageId="27" pageNumber="157">
|
||
<typeStatus id="54E12FFFFF92820757F3B0D06446FC69" box="[189,303,964,990]" pageId="27" pageNumber="157" type="paratype">Paratype</typeStatus>
|
||
and non-type variations
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFF92820757F3B0E7645AFA61" blockId="27.[188,1401,1010,1494]" pageId="27" pageNumber="157">
|
||
<typeStatus id="54E12FFFFF92820757F3B0E76446FBBA" box="[189,303,1011,1037]" pageId="27" pageNumber="157" type="paratype">Paratypes</typeStatus>
|
||
varied in size from
|
||
<quantity id="4CA23CB8FF9282075558B0E667AEFBBA" box="[534,711,1010,1037]" metricMagnitude="-2" metricUnit="m" metricValue="1.365" metricValueMax="2.37" metricValueMin="0.36" pageId="27" pageNumber="157" unit="mm" value="13.65" valueMax="23.7" valueMin="3.6">3.6 to 23.7 mm</quantity>
|
||
(DD). The two smallest specimens (DD = 3.6 and
|
||
<quantity id="4CA23CB8FF928207525AB0E76006FBBA" box="[1300,1391,1011,1037]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="27" pageNumber="157" unit="mm" value="4.8">4.8 mm</quantity>
|
||
) showed radial shields, jaws (including oral shields), DAPs, VAPs, and LAPs covered with granules, as well as the arm base and the section between the arm and distal genital slit. The arm spines in these specimens were up to five, short and pointed. Larger specimens (DD =
|
||
<quantity id="4CA23CB8FF9282075342B74861C3FBC1" box="[1036,1194,1116,1142]" metricMagnitude="-2" metricUnit="m" metricValue="1.655" metricValueMax="2.37" metricValueMin="0.94" pageId="27" pageNumber="157" unit="mm" value="16.55" valueMax="23.7" valueMin="9.4">9.4–23.7 mm</quantity>
|
||
) resembled more the
|
||
<typeStatus id="54E12FFFFF92820757A6B76B643BFB2E" box="[232,338,1151,1177]" pageId="27" pageNumber="157" type="holotype">holotype</typeStatus>
|
||
, as they had naked radial shields, oral shields and arms without granules, and scales on the arm base and distal genital slit. These presented exposed DAPs, divided into a maximum of three pieces in the smallest specimen (DD =
|
||
<quantity id="4CA23CB8FF9282075574B7D267FFFB57" box="[570,662,1222,1248]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="27" pageNumber="157" unit="mm" value="9.4">9.4 mm</quantity>
|
||
) and seven in larger ones (DD>
|
||
<quantity id="4CA23CB8FF9282075351B7D261E5FB57" box="[1055,1164,1222,1248]" metricMagnitude="-2" metricUnit="m" metricValue="2.2399999999999998" pageId="27" pageNumber="157" unit="mm" value="22.4">22.4 mm</quantity>
|
||
), varying according to size. The maximum number of arm spines also varied with size, ranging from seven (DD =
|
||
<quantity id="4CA23CB8FF928207525DB7FD6019FAB4" box="[1299,1392,1257,1283]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="27" pageNumber="157" unit="mm" value="9.4">9.4 mm</quantity>
|
||
) to nine (DD =
|
||
<quantity id="4CA23CB8FF9282075638B6186740FA91" box="[374,553,1292,1318]" metricMagnitude="-2" metricUnit="m" metricValue="2.045" metricValueMax="2.37" metricValueMin="1.72" pageId="27" pageNumber="157" unit="mm" value="20.45" valueMax="23.7" valueMin="17.2">17.2–23.7 mm</quantity>
|
||
). As for coloration, all
|
||
<typeStatus id="54E12FFFFF9282075406B61966D1FA90" box="[840,952,1293,1319]" pageId="27" pageNumber="157" type="paratype">paratypes</typeStatus>
|
||
exhibited dark, sinuous rings on the disc. The number and color intensity of these rings varied within (dorsal/ventral sides) and between specimens. The smallest specimens showed the faintest rings, which were quite susceptible to fading with preservation.
|
||
<typeStatus id="54E12FFFFF92820756D0B6626779FA27" box="[414,528,1398,1424]" pageId="27" pageNumber="157" type="paratype">Paratypes</typeStatus>
|
||
presented light brown, dark brown, or cream-colored oral shields with a brown center. On the ventral side, the interradii were as dark as the disc or light, darkening towards the disc edge.
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFF92820757F3B6E9675FF8E3" blockId="27.[188,1400,1533,1877]" pageId="27" pageNumber="157">
|
||
Other variations were observed in non-type specimens (DD = 6.0–
|
||
<quantity id="4CA23CB8FF92820754B8B6E96101F9AF" box="[1014,1128,1533,1560]" metricMagnitude="-2" metricUnit="m" metricValue="2.6399999999999997" pageId="27" pageNumber="157" unit="mm" value="26.4">26.4 mm</quantity>
|
||
). As in the
|
||
<typeStatus id="54E12FFFFF92820753B1B6EA601BF9AF" box="[1279,1394,1534,1560]" pageId="27" pageNumber="157" type="paratype">paratypes</typeStatus>
|
||
, the smallest specimens (DD <
|
||
<quantity id="4CA23CB8FF9282075565B53567E3F98C" box="[555,650,1569,1596]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="27" pageNumber="157" unit="mm" value="8.5">8.5 mm</quantity>
|
||
) had radial shields covered with granules; however, two large specimens (DD = 18.8 and
|
||
<quantity id="4CA23CB8FF92820756B6B550670BF9E9" box="[504,610,1604,1630]" metricMagnitude="-2" metricUnit="m" metricValue="2.36" pageId="27" pageNumber="157" unit="mm" value="23.6">23.6 mm</quantity>
|
||
) also presented covered radial shields.
|
||
<specimenCount id="9D5C5AD4FF9282075368B55161B4F9E9" box="[1062,1245,1604,1631]" count="4" pageId="27" pageNumber="157" type="generic">Four specimens</specimenCount>
|
||
(DD = 10.6– 19.0 mm) had one to several small, rounded plates on the disc. A few specimens showed trilobed, rather than rounded triangular, oral shields. Specimens with DD from
|
||
<quantity id="4CA23CB8FF92820754EBB59E6696F912" box="[933,1023,1674,1701]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="27" pageNumber="157" unit="mm" value="5.6">5.6 mm</quantity>
|
||
began to show some uncovered DAPs in the proximal and median arm sections, mostly entire or divided into up to two pieces, while their VAPs, LAPs, and oral shields remained covered. On the other hand, those with DD from
|
||
<quantity id="4CA23CB8FF9282075250B5C56011F95C" box="[1310,1400,1745,1771]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="27" pageNumber="157" unit="mm" value="7.7">7.7 mm</quantity>
|
||
had all their DAPs, VAPs, LAPs, and oral shields without granules. In terms of coloration,
|
||
<specimenCount id="9D5C5AD4FF9282075396B5E1601EF8B9" box="[1240,1399,1780,1807]" count="1" pageId="27" pageNumber="157" type="generic">one specimen</specimenCount>
|
||
presented a uniform light brown ventral side, and another had a general cream coloration with dark, sinuous rings on the dorsal disc.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C340C2D6FF92820157F3B46866C9FE07" lastPageId="29" lastPageNumber="159" pageId="27" pageNumber="157" type="distribution">
|
||
<paragraph id="8BE5915DFF92820757F3B4686487F821" blockId="27.[189,494,1916,1942]" box="[189,494,1916,1942]" pageId="27" pageNumber="157">
|
||
<heading id="D0AD2631FF92820757F3B4686487F821" bold="true" box="[189,494,1916,1942]" fontSize="11" level="3" pageId="27" pageNumber="157" reason="3">
|
||
<emphasis id="B92E4D4FFF92820757F3B4686487F821" bold="true" box="[189,494,1916,1942]" pageId="27" pageNumber="157">Distribution and habitat</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFF92820157F3B4BE66C9FE07" blockId="27.[187,1397,1962,2024]" lastBlockId="29.[188,1399,265,432]" lastPageId="29" lastPageNumber="159" pageId="27" pageNumber="157">
|
||
<taxonomicName id="4C5AEADEFF92820757F3B4BE64ECF873" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" box="[189,389,1962,1988]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="27" pageNumber="157" phylum="Echinodermata" rank="species" species="aija" status="sp. nov.">
|
||
<emphasis id="B92E4D4FFF92820757F3B4BE64ECF873" box="[189,389,1962,1988]" italics="true" pageId="27" pageNumber="157">Ophioderma aija</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21DF034FF92820756C7B4BF648BF872" box="[393,482,1963,1989]" pageId="27" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has been collected in
|
||
<collectingCountry id="F34DD1CDFF9282075593B4BE665EF873" box="[733,823,1962,1988]" name="Mexico" pageId="27" pageNumber="157">Mexico</collectingCountry>
|
||
in the Gulf of California (
|
||
<collectingRegion id="499E5FBFFF928207532FB4BE61D0F873" box="[1121,1209,1962,1988]" country="Mexico" name="Sonora" pageId="27" pageNumber="157">Sonora</collectingRegion>
|
||
,
|
||
<collectingRegion id="499E5FBFFF928207538CB4BE6581F85F" country="Mexico" name="Baja California Sur" pageId="27" pageNumber="157">Baja California Sur</collectingRegion>
|
||
),
|
||
<collectingRegion id="499E5FBFFF92820757B2B4D96426F850" box="[252,335,1997,2023]" country="Mexico" name="Jalisco" pageId="27" pageNumber="157">Jalisco</collectingRegion>
|
||
,
|
||
<collectingRegion id="499E5FBFFF9282075614B4D964DAF850" box="[346,435,1997,2023]" country="Mexico" name="Colima" pageId="27" pageNumber="157">Colima</collectingRegion>
|
||
,
|
||
<collectingRegion id="499E5FBFFF92820756F1B4D9672DF850" box="[447,580,1997,2023]" country="Mexico" name="Michoacan de Ocampo" pageId="27" pageNumber="157">Michoacán</collectingRegion>
|
||
,
|
||
<collectingRegion id="499E5FBFFF928207551EB4DA67D4F85F" box="[592,701,1998,2024]" country="Mexico" name="Guerrero" pageId="27" pageNumber="157">Guerrero</collectingRegion>
|
||
, and
|
||
<collectingRegion id="499E5FBFFF92820755B7B4DA663DF85F" box="[761,852,1998,2024]" country="Mexico" name="Oaxaca" pageId="27" pageNumber="157">Oaxaca</collectingRegion>
|
||
, as well as in
|
||
<collectingCountry id="F34DD1CDFF92820754BBB4D96118F850" box="[1013,1137,1997,2023]" name="Nicaragua" pageId="27" pageNumber="157">Nicaragua</collectingCountry>
|
||
, in Cardon Island (see Supp. file 1). Its northernmost record is from Guaymas,
|
||
<collectingRegion id="499E5FBFFF948201541EB21D66CFFE94" box="[848,934,265,291]" country="Mexico" name="Sonora" pageId="29" pageNumber="159">Sonora</collectingRegion>
|
||
,
|
||
<collectingCountry id="F34DD1CDFF94820154FCB21D6165FE94" box="[946,1036,265,291]" name="Mexico" pageId="29" pageNumber="159">Mexico</collectingCountry>
|
||
(
|
||
<geoCoordinate id="EE6EF79AFF9482015352B21D610FFE94" box="[1052,1126,265,291]" degrees="27" direction="north" orientation="latitude" pageId="29" pageNumber="159" precision="55555" value="27.0">27° N</geoCoordinate>
|
||
), and the southernmost from Cardon Island, Corinto,
|
||
<collectingCountry id="F34DD1CDFF9482015560B23867CEFEF1" box="[558,679,300,326]" name="Nicaragua" pageId="29" pageNumber="159">Nicaragua</collectingCountry>
|
||
(
|
||
<geoCoordinate id="EE6EF79AFF94820155F2B2386660FEF1" box="[700,777,300,326]" degrees="12" direction="north" orientation="latitude" pageId="29" pageNumber="159" precision="55555" value="12.0">12° N</geoCoordinate>
|
||
) (
|
||
<figureCitation id="13618DD8FF9482015469B238661DFEF0" box="[807,884,300,327]" captionStart="Fig" captionStartId="17.[189,232,1892,1918]" captionTargetBox="[258,1329,281,1836]" captionTargetId="figure-18@17.[239,1348,265,1851]" captionTargetPageId="17" captionText="Fig. 5. Distribution of Ophioderma teres (Lyman, 1860), Ophioderma unicolor H.L. Clark, 1940 stat. nov., Ophioderma aija sp. nov., and Ophioderma bichi sp. nov. in the eastern Pacific. A. Gulf of California. B. Revillagigedo Islands. C. Mexican Tropical Pacific. D. Central America and Colombia. E. Galapagos Islands." figureDoi="http://doi.org/10.5281/zenodo.13310029" httpUri="https://zenodo.org/record/13310029/files/figure.png" pageId="29" pageNumber="159">Fig. 5</figureCitation>
|
||
). The species may also occur in
|
||
<collectingCountry id="F34DD1CDFF948201525BB238601EFEF1" box="[1301,1399,300,326]" name="Ecuador" pageId="29" pageNumber="159">Ecuador</collectingCountry>
|
||
(
|
||
<collectingRegion id="499E5FBFFF948201578BB25B64F4FEDE" box="[197,413,335,361]" country="Ecuador" name="Galapagos" pageId="29" pageNumber="159">Galapagos Islands</collectingRegion>
|
||
and La Plata), according to
|
||
<bibRefCitation id="EFCBECACFF94820155A2B25B617CFEDE" author="Ziesenhenne" box="[748,1045,335,362]" firstAuthor="Ziesenhenne" page="191" pageId="29" pageNumber="159" pagination="185 - 201" refId="ref27266" refString="Ziesenhenne F. C. 1955. A review of the genus Ophioderma Muller and Troschel. In: Allan Hancock Foundation for Scientific Research (eds) Essays in the Natural Sciences in Honor of Captain Allan Hancock: 185 - 201. University of Southern California Press, Los Angeles. https: // doi. org / 10.5962 / bhl. title. 608" type="book chapter" year="1955">Ziesenhenne (1955: 191)</bibRefCitation>
|
||
. Revising material from these areas would help to confirm this.
|
||
<taxonomicName id="4C5AEADEFF948201550EB2666661FE3B" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" box="[576,776,370,396]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="aija">
|
||
<emphasis id="B92E4D4FFF948201550EB2666661FE3B" box="[576,776,370,396]" italics="true" pageId="29" pageNumber="159">Ophioderma aija</emphasis>
|
||
</taxonomicName>
|
||
is found typically on rocks, but also on shingle, sand, and live and dead coral, from the intertidal zone to
|
||
<quantity id="4CA23CB8FF948201545AB2826639FE07" box="[788,848,406,432]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="29" pageNumber="159" unit="m" value="27.0">27 m</quantity>
|
||
depth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF25C1D5FF95820057F3B5A06480F85F" ID-DOI="http://doi.org/10.5281/zenodo.13310039" ID-Zenodo-Dep="13310039" httpUri="https://zenodo.org/record/13310039/files/figure.png" pageId="28" pageNumber="158" startId="28.[189,232,1716,1742]" targetBox="[344,1244,265,1671]" targetPageId="28">
|
||
<paragraph id="8BE5915DFF95820057F3B5A06480F85F" blockId="28.[187,1399,1716,2025]" pageId="28" pageNumber="158">
|
||
<emphasis id="B92E4D4FFF95820057F3B5A06467F978" bold="true" box="[189,270,1716,1743]" pageId="28" pageNumber="158">Fig. 9.</emphasis>
|
||
<taxonomicName id="4C5AEADEFF9582005656B5A1648CF978" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" box="[280,485,1717,1743]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="28" pageNumber="158" phylum="Echinodermata" rank="species" species="aija" status="sp. nov.">
|
||
<emphasis id="B92E4D4FFF9582005656B5A1648CF978" box="[280,485,1717,1743]" italics="true" pageId="28" pageNumber="158">Ophioderma aija</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21DF034FF95820056A1B5A16726F978" box="[495,591,1717,1743]" pageId="28" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, non-type specimen (ICML-UNAM 18457, DD = 26.2 mm), SEM images of the ossicles.
|
||
<emphasis id="B92E4D4FFF95820056AEB5CC649CF945" bold="true" box="[480,501,1752,1778]" pageId="28" pageNumber="158">A</emphasis>
|
||
. Radial shield, external face. The arrowhead indicates the naked section observed in the intact animal.
|
||
<emphasis id="B92E4D4FFF9582005551B5EF675BF8A2" bold="true" box="[543,562,1787,1813]" pageId="28" pageNumber="158">B</emphasis>
|
||
. Radial shield, internal face.
|
||
<emphasis id="B92E4D4FFF95820054C5B5EF66C9F8A2" bold="true" box="[907,928,1787,1813]" pageId="28" pageNumber="158">C</emphasis>
|
||
. Dental plate.
|
||
<emphasis id="B92E4D4FFF9582005303B5EF610BF8A2" bold="true" box="[1101,1122,1787,1813]" pageId="28" pageNumber="158">D</emphasis>
|
||
. Adradial genital plate, abradial face.
|
||
<emphasis id="B92E4D4FFF958200562EB40A641DF88F" bold="true" box="[352,372,1822,1848]" pageId="28" pageNumber="158">E</emphasis>
|
||
. Adradial genital plate, adradial face.
|
||
<emphasis id="B92E4D4FFF9582005463B40A6656F88F" bold="true" box="[813,831,1822,1848]" pageId="28" pageNumber="158">F</emphasis>
|
||
. Oral plate, abradial face.
|
||
<emphasis id="B92E4D4FFF958200533FB40A61E1F88F" bold="true" box="[1137,1160,1822,1848]" pageId="28" pageNumber="158">G</emphasis>
|
||
. Oral plate, adradial face. Black arrows point to oral papillae sockets, white arrows point to granule sockets.
|
||
<emphasis id="B92E4D4FFF95820053E0B45661ACF8EB" bold="true" box="[1198,1221,1858,1884]" pageId="28" pageNumber="158">H</emphasis>
|
||
. Arm vertebra, proximal face.
|
||
<emphasis id="B92E4D4FFF958200563DB4716417F8C8" bold="true" box="[371,382,1893,1919]" pageId="28" pageNumber="158">I</emphasis>
|
||
. Arm vertebra, distal face.
|
||
<emphasis id="B92E4D4FFF9582005583B47167B5F8C8" bold="true" box="[717,732,1893,1919]" pageId="28" pageNumber="158">J</emphasis>
|
||
. Ventral arm plate, external face.
|
||
<emphasis id="B92E4D4FFF9582005336B47161E6F8C8" bold="true" box="[1144,1167,1893,1919]" pageId="28" pageNumber="158">K</emphasis>
|
||
. Ventral arm plate, internal face.
|
||
<emphasis id="B92E4D4FFF958200562DB49C641EF815" bold="true" box="[355,375,1928,1954]" pageId="28" pageNumber="158">L</emphasis>
|
||
. Lateral arm plate, external face.
|
||
<emphasis id="B92E4D4FFF958200545CB49C6647F815" bold="true" box="[786,814,1928,1954]" pageId="28" pageNumber="158">M</emphasis>
|
||
. Lateral arm plate, internal face.
|
||
<emphasis id="B92E4D4FFF958200538BB49C61B3F815" bold="true" box="[1221,1242,1928,1954]" pageId="28" pageNumber="158">N</emphasis>
|
||
. Lateral arm plate, lateral face. Abbreviations: dist = distal; dors = dorsal; prox = proximal; SA = spine articulation; Sp = spur; vent = ventral.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C340C2D6FF94820157F3B2C3612DFA7B" pageId="29" pageNumber="159" type="discussion">
|
||
<paragraph id="8BE5915DFF94820157F3B2C36446FE46" blockId="29.[189,303,471,497]" box="[189,303,471,497]" pageId="29" pageNumber="159">
|
||
<heading id="D0AD2631FF94820157F3B2C36446FE46" bold="true" box="[189,303,471,497]" fontSize="11" level="3" pageId="29" pageNumber="159" reason="3">
|
||
<emphasis id="B92E4D4FFF94820157F3B2C36446FE46" bold="true" box="[189,303,471,497]" pageId="29" pageNumber="159">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFF94820157F3B1126123FC71" blockId="29.[186,1399,518,967]" pageId="29" pageNumber="159">
|
||
Despite being recorded since 1860 (
|
||
<bibRefCitation id="EFCBECACFF948201553AB112667BFD97" author="Lyman" box="[628,786,518,544]" firstAuthor="Lyman" pageId="29" pageNumber="159" pagination="193 - 204" refId="ref25910" refString="Lyman T. 1860. Descriptions of new Ophiuridae, belonging to the Smithsonian Institution and to the Museum of Comparative Zoology at Cambridge. Proceedings of the Boston Society of Natural History 7: 193 - 204, 252 - 262. https: // doi. org / 10.5962 / bhl. part. 4822" type="journal article" year="1860">Lyman 1860</bibRefCitation>
|
||
),
|
||
<taxonomicName id="4C5AEADEFF9482015467B1126616FD97" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" box="[809,895,518,544]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="aija" status="sp. nov.">
|
||
<emphasis id="B92E4D4FFF9482015467B1126616FD97" box="[809,895,518,544]" italics="true" pageId="29" pageNumber="159">O. aija</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21DF034FF94820154C4B1126681FD97" box="[906,1000,518,544]" pageId="29" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
|
||
remained undescribed due to its misidentification as either
|
||
<taxonomicName id="4C5AEADEFF94820156A2B13D672CFDF4" baseAuthorityName="H. L. Clark" baseAuthorityYear="1940" box="[492,581,553,579]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="teres">
|
||
<emphasis id="B92E4D4FFF94820156A2B13D672CFDF4" box="[492,581,553,579]" italics="true" pageId="29" pageNumber="159">O. teres</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C5AEADEFF9482015529B13D6782FDF4" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" baseAuthorityName="H. L. Clark" baseAuthorityYear="1940" box="[615,747,553,579]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="unicolor" status="stat. nov.">
|
||
<emphasis id="B92E4D4FFF9482015529B13D6782FDF4" box="[615,747,553,579]" italics="true" pageId="29" pageNumber="159">O. unicolor</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21DF034FF94820155A0B13D663CFDF4" box="[750,853,553,579]" pageId="29" pageNumber="159">stat. nov.</taxonomicNameLabel>
|
||
Confusion with
|
||
<taxonomicName id="4C5AEADEFF948201535FB13D6102FDF4" baseAuthorityName="H. L. Clark" baseAuthorityYear="1940" box="[1041,1131,553,579]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="teres">
|
||
<emphasis id="B92E4D4FFF948201535FB13D6102FDF4" box="[1041,1131,553,579]" italics="true" pageId="29" pageNumber="159">O. teres</emphasis>
|
||
</taxonomicName>
|
||
arose from the original description of this species, where
|
||
<bibRefCitation id="EFCBECACFF9482015507B1596645FDD0" author="Lyman" box="[585,812,588,615]" firstAuthor="Lyman" page="258" pageId="29" pageNumber="159" pagination="193 - 204" refId="ref25910" refString="Lyman T. 1860. Descriptions of new Ophiuridae, belonging to the Smithsonian Institution and to the Museum of Comparative Zoology at Cambridge. Proceedings of the Boston Society of Natural History 7: 193 - 204, 252 - 262. https: // doi. org / 10.5962 / bhl. part. 4822" type="journal article" year="1860">Lyman (1860: 258)</bibRefCitation>
|
||
included a specimen of
|
||
<taxonomicName id="4C5AEADEFF9482015303B15961C9FDD1" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" box="[1101,1184,589,615]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="aija">
|
||
<emphasis id="B92E4D4FFF9482015303B15961C9FDD1" box="[1101,1184,589,615]" italics="true" pageId="29" pageNumber="159">O. aija</emphasis>
|
||
</taxonomicName>
|
||
(DD =
|
||
<quantity id="4CA23CB8FF94820153B4B1586026FDD1" box="[1274,1359,588,614]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="29" pageNumber="159" unit="mm" value="14.0">14 mm</quantity>
|
||
) as additional material, describing its characteristic color pattern: “…very dark umber, with fine, sinuous, black lines on the disc”. Since then,
|
||
<taxonomicName id="4C5AEADEFF9482015528B18767D1FD1B" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" box="[614,696,659,685]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="aija">
|
||
<emphasis id="B92E4D4FFF9482015528B18767D1FD1B" box="[614,696,659,685]" italics="true" pageId="29" pageNumber="159">O. aija</emphasis>
|
||
</taxonomicName>
|
||
has been recorded as
|
||
<taxonomicName id="4C5AEADEFF94820154F4B187617CFD1B" baseAuthorityName="H. L. Clark" baseAuthorityYear="1940" box="[954,1045,659,685]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="teres">
|
||
<emphasis id="B92E4D4FFF94820154F4B187617CFD1B" box="[954,1045,659,685]" italics="true" pageId="29" pageNumber="159">O. teres</emphasis>
|
||
</taxonomicName>
|
||
by other authors (e.g.,
|
||
<bibRefCitation id="EFCBECACFF948201526BB1876592FD66" author="Lyman" firstAuthor="Lyman" pageId="29" pageNumber="159" pagination="1 - 200" refId="ref25967" refString="Lyman T. 1865. Ophiuridae and Astrophytidae. Illustrated Catalogue of the Museum of Comparative Zoology at Harvard College 1: 1 - 200. https: // doi. org / 10.5962 / bhl. title. 40077" type="journal article" year="1865">Lyman 1865</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFCBECACFF9482015644B1A26738FD67" author="Steinbeck & Ricketts" box="[266,593,694,720]" firstAuthor="Steinbeck" pageId="29" pageNumber="159" refId="ref26778" refString="Steinbeck J. & Ricketts E. F. 1941. Sea of Cortez. A Leisurely Journal of Travel and Research. Viking Press, New York." type="book" year="1941">Steinbeck & Ricketts 1941</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFCBECACFF948201552EB1A2678AFD66" author="Caso" box="[608,739,694,721]" firstAuthor="Caso" pageId="29" pageNumber="159" pagination="219 - 312" refId="ref24399" refString="Caso M. E. 1951. Contribucion al conocimiento de los ofiuroideos de Mexico. I. Algunas especies de ofiuroideos litorales. Anales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico 22 (1): 219 - 312." type="journal article" year="1951">Caso 1951</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFCBECACFF94820155BCB1A261A4FD67" author="Granja-Fernandez & Lopez-Perez" box="[754,1229,694,720]" firstAuthor="Granja-Fernandez" pageId="29" pageNumber="159" pagination="1320 - 1323" refId="ref24835" refString="Granja-Fernandez M. R. & Lopez-Perez R. A. 2011. Nuevos registros de ofiuroideos (Echinodermata: Ophiuroidea) para localidades de Zihuatanejo (Guerrero) y Puerto Escondido (Oaxaca), Pacifico Mexicano. Revista Mexicana de Biodiversidad 82: 1320 - 1323. https: // doi. org / 10.22201 / ib. 20078706 e. 2011.4.675" type="journal article" year="2011">Granja-Fernández & López-Pérez 2011</bibRefCitation>
|
||
), occasionally attributing their morphological differences to distribution, habitat (
|
||
<bibRefCitation id="EFCBECACFF9482015489B1CD61CBFD43" author="Ziesenhenne" box="[967,1186,729,756]" firstAuthor="Ziesenhenne" pageId="29" pageNumber="159" pagination="185 - 201" refId="ref27266" refString="Ziesenhenne F. C. 1955. A review of the genus Ophioderma Muller and Troschel. In: Allan Hancock Foundation for Scientific Research (eds) Essays in the Natural Sciences in Honor of Captain Allan Hancock: 185 - 201. University of Southern California Press, Los Angeles. https: // doi. org / 10.5962 / bhl. title. 608" type="book chapter" year="1955">Ziesenhenne 1955</bibRefCitation>
|
||
), or a juvenile life stage (
|
||
<bibRefCitation id="EFCBECACFF9482015644B1E8671BFCA1" author="Granja-Fernandez" box="[266,626,764,791]" etAl="et al." firstAuthor="Granja-Fernandez" pageId="29" pageNumber="159" pagination="101 - 145" refId="ref24943" refString="Granja-Fernandez R., Herrero-Perezrul M. D., Lopez-Perez R. A., Hernandez L., Rodriguez-Zaragoza F. A., Jones R. W. & Pineda-Lopez R. 2014. Ophiuroidea (Echinodermata) from coral reefs in the Mexican Pacific. ZooKeys 406: 101 - 145. https: // doi. org / 10.3897 / zookeys. 406.6306" type="journal article" year="2014">
|
||
Granja-Fernández
|
||
<emphasis id="B92E4D4FFF94820156A5B1E96748FCA1" box="[491,545,764,790]" italics="true" pageId="29" pageNumber="159">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
). On the other hand, the misidentification as
|
||
<taxonomicName id="4C5AEADEFF94820153DAB1E96077FCA1" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" baseAuthorityName="H. L. Clark" baseAuthorityYear="1940" box="[1172,1310,764,791]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="unicolor">
|
||
<emphasis id="B92E4D4FFF94820153DAB1E96077FCA1" box="[1172,1310,764,791]" italics="true" pageId="29" pageNumber="159">O. unicolor</emphasis>
|
||
</taxonomicName>
|
||
started due to a mixture of
|
||
<specimenCount id="9D5C5AD4FF94820156ECB034672BFC8E" box="[418,578,799,826]" count="1" pageId="29" pageNumber="159" type="generic">one specimen</specimenCount>
|
||
of
|
||
<taxonomicName id="4C5AEADEFF9482015526B03467D3FC8E" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" box="[616,698,800,826]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="aija">
|
||
<emphasis id="B92E4D4FFF9482015526B03467D3FC8E" box="[616,698,800,826]" italics="true" pageId="29" pageNumber="159">O. aija</emphasis>
|
||
</taxonomicName>
|
||
in the type series of
|
||
<taxonomicName id="4C5AEADEFF94820154E2B034615AFC8E" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" baseAuthorityName="H. L. Clark" baseAuthorityYear="1940" box="[940,1075,799,826]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="unicolor">
|
||
<emphasis id="B92E4D4FFF94820154E2B034615AFC8E" box="[940,1075,799,826]" italics="true" pageId="29" pageNumber="159">O. unicolor</emphasis>
|
||
</taxonomicName>
|
||
(H.L.
|
||
<bibRefCitation id="EFCBECACFF94820153CFB00B606EFC8E" author="Clark" box="[1153,1287,799,825]" firstAuthor="Clark" pageId="29" pageNumber="159" pagination="331 - 352" refId="ref24502" refString="Clark H. L. 1940. Eastern Pacific Expeditions of the New York Zoological Society. XXI. Notes on Echinoderms from the west coast of Central America. Zoologica 25 (3): 331 - 352. https: // doi. org / 10.5962 / p. 184708" type="journal article" year="1940">Clark 1940</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFCBECACFF948201525DB0346595FCEB" author="Downey" firstAuthor="Downey" pageId="29" pageNumber="159" pagination="1 - 239" refId="ref24648" refString="Downey M. E. 1969. Catalog of recent ophiuroid type specimens in major collections in the United States. United States National Museum Bulletin 293: 1 - 239. https: // doi. org / 10.5479 / si. 03629236.293" type="journal article" year="1969">Downey 1969</bibRefCitation>
|
||
). Revision of the former
|
||
<typeStatus id="54E12FFFFF948201556AB05767E0FCEA" box="[548,649,835,861]" pageId="29" pageNumber="159" type="paratype">paratype</typeStatus>
|
||
(MCZ IZ OPH-167471) led some authors to identify
|
||
<taxonomicName id="4C5AEADEFF948201524BB057603EFCEB" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" box="[1285,1367,835,861]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="aija">
|
||
<emphasis id="B92E4D4FFF948201524BB057603EFCEB" box="[1285,1367,835,861]" italics="true" pageId="29" pageNumber="159">O. aija</emphasis>
|
||
</taxonomicName>
|
||
as
|
||
<taxonomicName id="4C5AEADEFF94820157F3B07264D1FC37" authorityName="H. L. Clark" authorityYear="1940" box="[189,440,870,896]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="variety" species="teres" variety="unicolor">
|
||
<emphasis id="B92E4D4FFF94820157F3B0726473FC37" box="[189,282,870,896]" italics="true" pageId="29" pageNumber="159">O. teres</emphasis>
|
||
var.
|
||
<emphasis id="B92E4D4FFF948201561BB07264D1FC37" box="[341,440,870,896]" italics="true" pageId="29" pageNumber="159">unicolor</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFCBECACFF9482015687B072678CFC37" author="Granja-Fernandez" box="[457,741,870,896]" firstAuthor="Granja-Fernandez" pageId="29" pageNumber="159" refId="ref24810" refString="Granja-Fernandez M. R. 2019. Ofiuroideos (Echinodermata) del Pacifico oriental tropical. PhD thesis, Universidad Autonoma Metropolitana, Mexico." type="book" year="2019">Granja-Fernández 2019</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFCBECACFF94820155BCB072613EFC37" author="Mireles-Velazquez" box="[754,1111,870,896]" etAl="et al." firstAuthor="Mireles-Velazquez" pageId="29" pageNumber="159" pagination="312 - 333" refId="ref26061" refString="Mireles-Velazquez D., Rosales-Contreras G. I., Conejeros-Vargas C. A., Solis-Marin F. A. & Granja-Fernandez R. 2021. Los ofiuroideos (Echinodermata: Ophiuroidea) de la bahia de Chamela, Jalisco, Mexico. Revista de Biologia Tropical 69 (S 1): 312 - 333. https: // doi. org / 10.15517 / rbt. v 69 iSuppl. 1.46364" type="journal article" year="2021">
|
||
Mireles-Velázquez
|
||
<emphasis id="B92E4D4FFF9482015499B073617BFC37" box="[983,1042,870,896]" italics="true" pageId="29" pageNumber="159">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
) or
|
||
<taxonomicName id="4C5AEADEFF94820153C7B0726484FC14" authority="(Humara-Gil et al. 2022)" baseAuthorityName="Humara-Gil" baseAuthorityYear="2022" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" isUncertain="true" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="variety" species="teres" variety="unicolor">
|
||
<emphasis id="B92E4D4FFF94820153C7B07261CCFC37" box="[1161,1189,870,896]" italics="true" pageId="29" pageNumber="159">O.</emphasis>
|
||
cf.
|
||
<emphasis id="B92E4D4FFF948201539DB073601FFC37" box="[1235,1398,870,896]" italics="true" pageId="29" pageNumber="159">teres unicolor</emphasis>
|
||
(
|
||
<bibRefCitation id="EFCBECACFF948201578BB09D648FFC14" author="Humara-Gil" box="[197,486,905,931]" etAl="et al." firstAuthor="Humara-Gil" pageId="29" pageNumber="159" pagination="365 - 384" refId="ref25718" refString="Humara-Gil K. J., Granja-Fernandez R., Bautista-Guerrero E. & Rodriguez-Troncoso A. P. 2022. Overlooked for over a century: Ophioderma occultum sp. nov. (Echinodermata), a new species of brittle star from the Eastern Pacific. Journal of Natural History 56 (5 - 8): 365 - 384. https: // doi. org / 10.1080 / 00222933.2022.2071179" type="journal article" year="2022">
|
||
Humara-Gil
|
||
<emphasis id="B92E4D4FFF9482015610B09E64F5FC14" box="[350,412,905,931]" italics="true" pageId="29" pageNumber="159">et al.</emphasis>
|
||
2022
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. In this work, the comprehensive revision of the type series of
|
||
<taxonomicName id="4C5AEADEFF94820153A3B09D601FFC14" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" baseAuthorityName="H. L. Clark" baseAuthorityYear="1940" box="[1261,1398,905,931]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="unicolor">
|
||
<emphasis id="B92E4D4FFF94820153A3B09D601FFC14" box="[1261,1398,905,931]" italics="true" pageId="29" pageNumber="159">O. unicolor</emphasis>
|
||
</taxonomicName>
|
||
resulted in the reclassification of the specimen of
|
||
<taxonomicName id="4C5AEADEFF948201544DB0B9663DFC71" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" box="[771,852,941,967]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="aija">
|
||
<emphasis id="B92E4D4FFF948201544DB0B9663DFC71" box="[771,852,941,967]" italics="true" pageId="29" pageNumber="159">O. aija</emphasis>
|
||
</taxonomicName>
|
||
as a distinct species.
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFF94820157F3B0F96422FB48" blockId="29.[189,1400,1005,1279]" pageId="29" pageNumber="159">
|
||
<taxonomicName id="4C5AEADEFF94820157F3B0F964E3FBB0" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" box="[189,394,1005,1031]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="aija" status="sp. nov.">
|
||
<emphasis id="B92E4D4FFF94820157F3B0F964E3FBB0" box="[189,394,1005,1031]" italics="true" pageId="29" pageNumber="159">Ophioderma aija</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21DF034FF94820156DAB0FA649BFBBF" box="[404,498,1006,1032]" pageId="29" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
|
||
shares with
|
||
<taxonomicName id="4C5AEADEFF94820155DEB0FA6786FBB0" baseAuthorityName="H. L. Clark" baseAuthorityYear="1940" box="[656,751,1006,1032]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="teres">
|
||
<emphasis id="B92E4D4FFF94820155DEB0FA6786FBB0" box="[656,751,1006,1032]" italics="true" pageId="29" pageNumber="159">O. teres</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5AEADEFF9482015461B0FA66D0FBB0" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" baseAuthorityName="H. L. Clark" baseAuthorityYear="1940" box="[815,953,1005,1032]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="unicolor" status="stat. nov.">
|
||
<emphasis id="B92E4D4FFF9482015461B0FA66D0FBB0" box="[815,953,1005,1032]" italics="true" pageId="29" pageNumber="159">O. unicolor</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21DF034FF948201548AB0FA6158FBBF" box="[964,1073,1006,1032]" pageId="29" pageNumber="159">stat. nov.</taxonomicNameLabel>
|
||
the covered adoral shields and divided DAPs. However, they differ as follows: 1) radial shields typically naked in
|
||
<taxonomicName id="4C5AEADEFF94820153A3B705602BFB9C" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" box="[1261,1346,1041,1067]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="aija">
|
||
<emphasis id="B92E4D4FFF94820153A3B705602BFB9C" box="[1261,1346,1041,1067]" italics="true" pageId="29" pageNumber="159">O. aija</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5AEADEFF94820157F3B720642CFBF9" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" baseAuthorityName="H. L. Clark" baseAuthorityYear="1940" box="[189,325,1076,1102]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="unicolor">
|
||
<emphasis id="B92E4D4FFF94820157F3B720642CFBF9" box="[189,325,1076,1102]" italics="true" pageId="29" pageNumber="159">O. unicolor</emphasis>
|
||
</taxonomicName>
|
||
versus radial shields either covered or naked in
|
||
<taxonomicName id="4C5AEADEFF94820154C7B720668CFBF9" baseAuthorityName="H. L. Clark" baseAuthorityYear="1940" box="[905,997,1076,1102]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="teres">
|
||
<emphasis id="B92E4D4FFF94820154C7B720668CFBF9" box="[905,997,1076,1102]" italics="true" pageId="29" pageNumber="159">O. teres</emphasis>
|
||
</taxonomicName>
|
||
; 2) shorter arms in
|
||
<taxonomicName id="4C5AEADEFF948201539DB720604FFBF9" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" box="[1235,1318,1076,1102]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="aija">
|
||
<emphasis id="B92E4D4FFF948201539DB720604FFBF9" box="[1235,1318,1076,1102]" italics="true" pageId="29" pageNumber="159">O. aija</emphasis>
|
||
</taxonomicName>
|
||
(mean AL:DD = 2.7) than in
|
||
<taxonomicName id="4C5AEADEFF948201569EB74C6747FBC6" baseAuthorityName="H. L. Clark" baseAuthorityYear="1940" box="[464,558,1112,1138]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="teres">
|
||
<emphasis id="B92E4D4FFF948201569EB74C6747FBC6" box="[464,558,1112,1138]" italics="true" pageId="29" pageNumber="159">O. teres</emphasis>
|
||
</taxonomicName>
|
||
(mean AL:DD = 4.1) and
|
||
<taxonomicName id="4C5AEADEFF948201543EB74C6690FBC6" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" baseAuthorityName="H. L. Clark" baseAuthorityYear="1940" box="[880,1017,1111,1138]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="unicolor">
|
||
<emphasis id="B92E4D4FFF948201543EB74C6690FBC6" box="[880,1017,1111,1138]" italics="true" pageId="29" pageNumber="159">O. unicolor</emphasis>
|
||
</taxonomicName>
|
||
(mean AL:DD = 3.6); 3) DAPs divided into more pieces in
|
||
<taxonomicName id="4C5AEADEFF948201554BB76F6731FB23" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" box="[517,600,1147,1173]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="aija">
|
||
<emphasis id="B92E4D4FFF948201554BB76F6731FB23" box="[517,600,1147,1173]" italics="true" pageId="29" pageNumber="159">O. aija</emphasis>
|
||
</taxonomicName>
|
||
(mean = 3, maximum = 9) and
|
||
<taxonomicName id="4C5AEADEFF9482015480B76F6142FB23" baseAuthorityName="H. L. Clark" baseAuthorityYear="1940" box="[974,1067,1147,1173]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="teres">
|
||
<emphasis id="B92E4D4FFF9482015480B76F6142FB23" box="[974,1067,1147,1173]" italics="true" pageId="29" pageNumber="159">O. teres</emphasis>
|
||
</taxonomicName>
|
||
(mean = 3, maximum = 13) than in
|
||
<taxonomicName id="4C5AEADEFF9482015658B78A64F4FB00" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" baseAuthorityName="H. L. Clark" baseAuthorityYear="1940" box="[278,413,1181,1208]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="unicolor">
|
||
<emphasis id="B92E4D4FFF9482015658B78A64F4FB00" box="[278,413,1181,1208]" italics="true" pageId="29" pageNumber="159">O. unicolor</emphasis>
|
||
</taxonomicName>
|
||
(mean = 2, maximum = 6), and 4) color pattern with dark, sinuous rings in the disc in
|
||
<taxonomicName id="4C5AEADEFF9482015792B7D56447FB6C" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" box="[220,302,1217,1243]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="aija">
|
||
<emphasis id="B92E4D4FFF9482015792B7D56447FB6C" box="[220,302,1217,1243]" italics="true" pageId="29" pageNumber="159">O. aija</emphasis>
|
||
</taxonomicName>
|
||
, rounded cream specks in the disc and arms in
|
||
<taxonomicName id="4C5AEADEFF9482015411B7D566D3FB6C" baseAuthorityName="H. L. Clark" baseAuthorityYear="1940" box="[863,954,1217,1243]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="teres">
|
||
<emphasis id="B92E4D4FFF9482015411B7D566D3FB6C" box="[863,954,1217,1243]" italics="true" pageId="29" pageNumber="159">O. teres</emphasis>
|
||
</taxonomicName>
|
||
, and uniform brown disc and arms in
|
||
<taxonomicName id="4C5AEADEFF94820157F0B7F0642DFB49" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" baseAuthorityName="H. L. Clark" baseAuthorityYear="1940" box="[190,324,1252,1278]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="unicolor">
|
||
<emphasis id="B92E4D4FFF94820157F0B7F0642DFB49" box="[190,324,1252,1278]" italics="true" pageId="29" pageNumber="159">O. unicolor</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE5915DFF94820157F3B632612DFA7B" blockId="29.[189,1400,1317,1485]" pageId="29" pageNumber="159">
|
||
As mentioned previously, one of the most distinctive characters of
|
||
<taxonomicName id="4C5AEADEFF9482015494B6326147FA88" authorityName="Humara-Gil & Granja-Fernández & Bautista-Guerrero & Solís-Marín & Rodríguez-Troncoso" authorityYear="2024" box="[986,1070,1318,1344]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiacanthida" pageId="29" pageNumber="159" phylum="Echinodermata" rank="species" species="aija" status="sp. nov.">
|
||
<emphasis id="B92E4D4FFF9482015494B6326147FA88" box="[986,1070,1318,1344]" italics="true" pageId="29" pageNumber="159">O. aija</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21DF034FF9482015378B63261FBFAF7" box="[1078,1170,1318,1344]" pageId="29" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is its color pattern. Although the rings remain visible in preserved specimens as old as 163 years (MZC IZ OPH-112), they may fade due to fixation. It is advisable to photograph or label specimens appropriately in the field to avoid confusion with other similar species in case the rings fade. This is particularly important for juvenile specimens whose other diagnostic characters may not be developed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |