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<mods:title id="A4876BFCE2D79851AE42E53F037DC82D">Revision of the subgenus Nessus of the genus Hypocaccus from Central Europe, with description of a new species (Coleoptera: Histeridae)</mods:title>
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<mods:namePart id="96EBEB960AC0133176B897100D4A6E63">Lackner, Tomáš</mods:namePart>
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<taxonomicName id="4C44B1F2FFED470AFC9CFF04CA13FF39" ID-CoL="9TMC5" authority="(Rosenhauer, 1847)" authorityName="Lackner &amp; Seres" authorityYear="2018" baseAuthorityName="Rosenhauer" baseAuthorityYear="1847" box="[864,1399,189,213]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="curtus" status="comb. nov." subGenus="Nessus">
<emphasis id="B9301663FFED470AFC9CFF04CC81FF38" bold="true" box="[864,997,189,212]" italics="true" pageId="13" pageNumber="432">Hypocaccus</emphasis>
(
<emphasis id="B9301663FFED470AFC09FF04CB26FF38" bold="true" box="[1013,1090,189,212]" italics="true" pageId="13" pageNumber="432">Nessus</emphasis>
) 
<emphasis id="B9301663FFED470AFBADFF04CBF3FF38" bold="true" box="[1105,1175,189,212]" italics="true" pageId="13" pageNumber="432">curtus</emphasis>
(
<bibRefCitation id="EFD5B780FFED470AFB59FF07CA15FF39" author="ROSENHAUER W. G." box="[1189,1393,190,213]" pageId="13" pageNumber="432" refId="ref21812" refString="ROSENHAUER W. G. 1847: Beitrage zur Insekten-Fauna Europas. Erstes Bandchen; enthalt die Beschreibung von sechzig neuen Kafern aus Bayern, Tyrol, Ungarn, etc., so wie die Kafer Tyrols, nach dem Ergebnisse von vier Reisen zusammengestellt. Mit einer Tafel Abbildungen. Th. Blasing, Erlangen, x + 159 pp." type="book" year="1847">Rosenhauer, 1847</bibRefCitation>
)
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<taxonomicNameLabel id="A203AB18FFED470AFBD3FF67CBCDFF19" box="[1071,1193,222,245]" pageId="13" pageNumber="432" rank="species">comb. nov.</taxonomicNameLabel>
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<subSubSection id="C35E99FAFFED470AFBF9FF45CBB7FEE3" box="[1029,1235,252,271]" pageId="13" pageNumber="432" type="description">
<paragraph id="8BFBCA71FFED470AFBF9FF45CBB7FEE3" blockId="13.[828,1438,252,998]" box="[1029,1235,252,271]" pageId="13" pageNumber="432">
(
<figureCitation id="137FD6F4FFED470AFBF7FF45CB29FEE3" box="[1035,1101,252,271]" captionStart="Figs 31–32" captionStartId="11.[151,188,1619,1638]" captionTargetBox="[172,742,1091,1585]" captionTargetId="figure-518@11.[151,760,1066,1603]" captionTargetPageId="11" captionText="Figs 31–32. Protibiae, dorsal view. 31 – Hypocaccus (Nessus) rubripes (Erichson, 1834); 32 – H. (N.) curtus (Rosenhauer, 1847) (both re-drawn from VIENNA 1980)." figureDoi="http://doi.org/10.5281/zenodo.4504938" httpUri="https://zenodo.org/record/4504938/files/figure.png" pageId="13" pageNumber="432">Figs 32</figureCitation>
, 
<figureCitation id="137FD6F4FFED470AFBAAFF45CBEAFEE3" box="[1110,1166,252,271]" captionStart-0="Figs 40–43" captionStart-1="Figs 44–45" captionStartId-0="15.[151,188,2006,2025]" captionStartId-1="16.[151,188,2008,2027]" captionTargetBox-0="[212,1382,479,1970]" captionTargetBox-1="[156,1428,1206,1975]" captionTargetId-0="figure-17@15.[201,1387,467,1981]" captionTargetId-1="figure-13@16.[151,1436,1200,1986]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="Figs 40–43. Hypocaccus (Nessus) curtus (Rosenhauer, 1847), dorsal habitus.40 – Saprinus curtus Rosenhauer, 1847, lectotype.41 – Saprinus puncticollis Küster, 1849, lectotype. 42 – Saprinus cribellaticollis Jacquelin Du Val, 1858, lectotype. 43 – Saprinus sicanus Marseul, 1862, lectotype." captionText-1="Figs 44–45.Hypocaccus (Nessus) curtus (Rosenhauer, 1847), dorsal habitus. 44 – Saprinus revisus Marseul, 1876, lectotype. 45 – Hypocacculus (Nessus) controversus G. Müller, 1937, lectotype." figureDoi-0="http://doi.org/10.5281/zenodo.4504944" figureDoi-1="http://doi.org/10.5281/zenodo.4504946" httpUri-0="https://zenodo.org/record/4504944/files/figure.png" httpUri-1="https://zenodo.org/record/4504946/files/figure.png" pageId="13" pageNumber="432">40–44</figureCitation>
, 
<figureCitation id="137FD6F4FFED470AFB6AFF45CBA8FEE3" box="[1174,1228,252,271]" captionStart="Figs 46–54" captionStartId="17.[151,188,1365,1384]" captionTargetBox="[271,1310,177,1331]" captionTargetId="figure-374@17.[252,1324,170,1345]" captionTargetPageId="17" captionText="Figs 46–54. Male genitalia of Hypocaccus (Nessus) curtus (Rosenhauer, 1847). 45 – VIII sternite and tergite, ventral view; 46 – ditto, dorsal view; 47 – ditto, lateral view; 48 – IX and X tergites, dorsal view; 49 – ditto, lateral view; 50 – IX sternite (spiculum gastrale), ventral view; 51 – ditto, lateral view; 52 – aedeagus, dorsal view; 53 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4504952" httpUri="https://zenodo.org/record/4504952/files/figure.png" pageId="13" pageNumber="432">46–54</figureCitation>
)
</paragraph>
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<subSubSection id="C35E99FAFFED470AFCC0FEA7CBB1FC0A" pageId="13" pageNumber="432" type="reference_group">
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<taxonomicName id="4C44B1F2FFED470AFCC0FEA7CBE2FEDD" authority="Rosenhauer, 1847: 26" authorityName="Rosenhauer" authorityPageNumber="26" authorityYear="1847" box="[828,1158,286,305]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="curtus">
<emphasis id="B9301663FFED470AFCC0FEA7CCA7FEDD" box="[828,963,286,305]" italics="true" pageId="13" pageNumber="432">Saprinus curtus</emphasis>
<bibRefCitation id="EFD5B780FFED470AFC36FEA7CBE2FEDD" author="ROSENHAUER W. G." box="[970,1158,286,305]" pageId="13" pageNumber="432" refId="ref21812" refString="ROSENHAUER W. G. 1847: Beitrage zur Insekten-Fauna Europas. Erstes Bandchen; enthalt die Beschreibung von sechzig neuen Kafern aus Bayern, Tyrol, Ungarn, etc., so wie die Kafer Tyrols, nach dem Ergebnisse von vier Reisen zusammengestellt. Mit einer Tafel Abbildungen. Th. Blasing, Erlangen, x + 159 pp." type="book" year="1847">Rosenhauer, 1847: 26</bibRefCitation>
</taxonomicName>
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(original description). 
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<bibRefCitation id="EFD5B780FFED470AFAB3FEA7CCF2FEA0" author="MARSEUL S. A." pageId="13" pageNumber="432" pagination="677 - 758" refId="ref21234" refString="MARSEUL S. A. 1855: Essai monographique sur la familie des Histerides. (Suite). Annales de Societe Entomologique de France, Troiseme Serie 3: 83 - 165, 327 - 506, 677 - 758." type="journal article" year="1855">MARSEUL (1855)</bibRefCitation>
: 751
</treatmentCitation>
(redescription).
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</paragraph>
<paragraph id="8BFBCA71FFED470AFCC0FEEACAFAFE8A" blockId="13.[828,1438,252,998]" box="[828,1438,339,358]" pageId="13" pageNumber="432">
<treatmentCitationGroup id="AB54ED5FFFED470AFCC0FEEACAFAFE8A" box="[828,1438,339,358]" pageId="13" pageNumber="432">
<treatmentCitation id="0AE5EC60FFED470AFCC0FEEACA72FE8A" author="GANGLBAUER L." box="[828,1302,339,358]" page="389" pageId="13" pageNumber="432" year="1899">
<taxonomicName id="4C44B1F2FFED470AFCC0FEEACB8BFE8A" authority=": GANGLBAUER (1899)" authorityName=": GANGLBAUER" authorityYear="1899" box="[828,1263,339,358]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="curtus" subGenus="Hypocaccus">
<emphasis id="B9301663FFED470AFCC0FEEACCECFE8A" box="[828,904,339,358]" italics="true" pageId="13" pageNumber="432">Saprinus</emphasis>
(
<emphasis id="B9301663FFED470AFC6EFEEACC9EFE8A" box="[914,1018,339,358]" italics="true" pageId="13" pageNumber="432">Hypocaccus</emphasis>
) 
<emphasis id="B9301663FFED470AFBF9FEEACB5EFE8A" box="[1029,1082,339,358]" italics="true" pageId="13" pageNumber="432">curtus</emphasis>
: 
<bibRefCitation id="EFD5B780FFED470AFBB8FEEACB8BFE8A" author="GANGLBAUER L." box="[1092,1263,339,358]" pageId="13" pageNumber="432" refId="ref20673" refString="GANGLBAUER L. 1899: Die Kafer von Mitteleuropa. Die Kafer der osterreichisch-ungarischen Monarchie, Deutschland, der Schweiz, sowie des franzosischen und italienischen Alpengebiets. III, 1. Familienreihe Staphylinoidea. 2 Theil: Scydmaenidae, Silphidae, Clambidae, Leptinidae, Platypsyllidae, Corylophidae, Sphaeritidae, Trichopterigydae, Hydroscaphidae, Scaphidiidae, Histeridae. Carl Gerolds Sohn, Wien, 408 pp." type="book" year="1899">GANGLBAUER (1899)</bibRefCitation>
</taxonomicName>
: 389
</treatmentCitation>
(redescription).
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</paragraph>
<paragraph id="8BFBCA71FFED470AFCC0FED7CB38FE5A" blockId="13.[828,1438,252,998]" pageId="13" pageNumber="432">
<treatmentCitationGroup id="AB54ED5FFFED470AFCC0FED7CB38FE5A" pageId="13" pageNumber="432">
<treatmentCitation id="0AE5EC60FFED470AFCC0FED7CA68FE6D" author="REICHARDT A." box="[828,1292,366,385]" page="49" pageId="13" pageNumber="432" year="1932">
<taxonomicName id="4C44B1F2FFED470AFCC0FED7CB8FFE6D" ID-CoL="6N25R" authority=": REICHARDT (1932)" authorityName="REICHARDT" authorityYear="1932" box="[828,1259,366,385]" class="Insecta" family="Histeridae" genus="Hypocacculus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="curtus" subGenus="Nessus">
<emphasis id="B9301663FFED470AFCC0FED7CCD1FE6D" box="[828,949,366,385]" italics="true" pageId="13" pageNumber="432">Hypocacculus</emphasis>
(
<emphasis id="B9301663FFED470AFC3FFED7CC9AFE6D" box="[963,1022,366,385]" italics="true" pageId="13" pageNumber="432">Nessus</emphasis>
) 
<emphasis id="B9301663FFED470AFBF0FED7CB25FE6D" box="[1036,1089,366,385]" italics="true" pageId="13" pageNumber="432">curtus</emphasis>
: 
<bibRefCitation id="EFD5B780FFED470AFBB2FED7CB8FFE6D" author="REICHARDT A." box="[1102,1259,366,385]" pageId="13" pageNumber="432" pagination="1 - 164" refId="ref21636" refString="REICHARDT A. 1932: Beitrage zur einer Monographie der Saprininae (Coleoptera, Histeridae). Mitteilungen aus dem Zoologischen Museum in Berlin 18: 1 - 164." type="journal article" year="1932">REICHARDT (1932)</bibRefCitation>
</taxonomicName>
: 49
</treatmentCitation>
, 122 (keyed, redescription, incl. pl. IV, fig. 9); 
<treatmentCitation id="0AE5EC60FFED470AFB94FE30CA4AFE70" author="REICHARDT A." box="[1128,1326,393,412]" page="285" pageId="13" pageNumber="432" year="1941">
<bibRefCitation id="EFD5B780FFED470AFB94FE30CA60FE70" author="REICHARDT A." box="[1128,1284,393,412]" pageId="13" pageNumber="432" refId="ref21666" refString="REICHARDT A. 1941: Semeystva Sphaeritidae i Histeridae (Vol. 1). [Families Sphaeritidae and Histeridae]. Fauna SSSR, Nasekomye Zhestokrylye, V, 3. Nauka, Moskva-Leningrad, xiii + 419 pp." type="book" year="1941">REICHARDT (1941)</bibRefCitation>
: 285
</treatmentCitation>
, 300 (keyed, redescription, incl. fig. 147C).
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</paragraph>
<paragraph id="8BFBCA71FFED470AFCC0FE07CA1EFE00" blockId="13.[828,1438,252,998]" pageId="13" pageNumber="432">
<treatmentCitationGroup id="AB54ED5FFFED470AFCC0FE07CA1EFE00" pageId="13" pageNumber="432">
<treatmentCitation id="0AE5EC60FFED470AFCC0FE07CBE3FE3D" author="KUSTER H. C." box="[828,1159,446,465]" page="30" pageId="13" pageNumber="432" year="1849">
<taxonomicName id="4C44B1F2FFED470AFCC0FE07CBE3FE3D" ID-CoL="9N3B7" authority="Kuster, 1849: 30" authorityName="Kuster" authorityPageNumber="30" authorityYear="1849" box="[828,1159,446,465]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="B9301663FFED470AFCC0FE07CC96FE3D" box="[828,1010,446,465]" italics="true" pageId="13" pageNumber="432">Saprinus puncticollis</emphasis>
<bibRefCitation id="EFD5B780FFED470AFC04FE07CBE3FE3D" author="KUSTER H. C." box="[1016,1159,446,465]" pageId="13" pageNumber="432" refId="ref21108" refString="KUSTER H. C. 1849: Die Kafer Europa's. Nach der Natur beschreiben. XVII Heft. Bauer and Raspe, Nurnberg, 100 pls." type="book" year="1849">Küster, 1849: 30</bibRefCitation>
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(original description). 
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<bibRefCitation id="EFD5B780FFED470AFAB3FE07CCF2FE00" author="MARSEUL S. A." pageId="13" pageNumber="432" pagination="677 - 758" refId="ref21234" refString="MARSEUL S. A. 1855: Essai monographique sur la familie des Histerides. (Suite). Annales de Societe Entomologique de France, Troiseme Serie 3: 83 - 165, 327 - 506, 677 - 758." type="journal article" year="1855">MARSEUL (1855)</bibRefCitation>
: 755
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(redescription); 
<treatmentCitation id="0AE5EC60FFED470AFBB0FE60CA62FE00" author="BICKHARDT H." box="[1100,1286,473,492]" page="96" pageId="13" pageNumber="432" year="1916">
<bibRefCitation id="EFD5B780FFED470AFBB0FE60CB8CFE00" author="BICKHARDT H." box="[1100,1256,473,492]" pageId="13" pageNumber="432" refId="ref20506" refString="BICKHARDT H. 1916 - 1917: Histeridae. In: WYTSMAN P. (ed.): Genera Insectorum. Fasc. 166 a, b. La Haye, 302 pp." type="book" year="1916">BICKHARDT (1916)</bibRefCitation>
: 96
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(synonymy).
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</paragraph>
<paragraph id="8BFBCA71FFED470AFCC0FE4ACCD6FDCD" blockId="13.[828,1438,252,998]" pageId="13" pageNumber="432">
<treatmentCitationGroup id="AB54ED5FFFED470AFCC0FE4ACCD6FDCD" pageId="13" pageNumber="432">
<treatmentCitation id="0AE5EC60FFED470AFCC0FE4ACA04FDEA" author="GANGLBAUER L." box="[828,1376,499,518]" page="389" pageId="13" pageNumber="432" year="1899">
<taxonomicName id="4C44B1F2FFED470AFCC0FE4ACA52FDEA" ID-CoL="9N3B7" authority=": GANGLBAUER (1899)" authorityName=": GANGLBAUER" authorityYear="1899" box="[828,1334,499,518]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="puncticollis" subGenus="Hypocaccus">
<emphasis id="B9301663FFED470AFCC0FE4ACCEDFDEA" box="[828,905,499,518]" italics="true" pageId="13" pageNumber="432">Saprinus</emphasis>
(
<emphasis id="B9301663FFED470AFC64FE4ACB65FDEA" box="[920,1025,499,518]" italics="true" pageId="13" pageNumber="432">Hypocaccus</emphasis>
) 
<emphasis id="B9301663FFED470AFBEDFE4ACB12FDEA" box="[1041,1142,499,518]" italics="true" pageId="13" pageNumber="432">puncticollis</emphasis>
: 
<bibRefCitation id="EFD5B780FFED470AFB78FE4ACA52FDEA" author="GANGLBAUER L." box="[1156,1334,499,518]" pageId="13" pageNumber="432" refId="ref20673" refString="GANGLBAUER L. 1899: Die Kafer von Mitteleuropa. Die Kafer der osterreichisch-ungarischen Monarchie, Deutschland, der Schweiz, sowie des franzosischen und italienischen Alpengebiets. III, 1. Familienreihe Staphylinoidea. 2 Theil: Scydmaenidae, Silphidae, Clambidae, Leptinidae, Platypsyllidae, Corylophidae, Sphaeritidae, Trichopterigydae, Hydroscaphidae, Scaphidiidae, Histeridae. Carl Gerolds Sohn, Wien, 408 pp." type="book" year="1899">GANGLBAUER (1899)</bibRefCitation>
</taxonomicName>
: 389
</treatmentCitation>
(redescription).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFBCA71FFED470AFCC0FD90CA77FD2D" blockId="13.[828,1438,252,998]" pageId="13" pageNumber="432">
<treatmentCitationGroup id="AB54ED5FFFED470AFCC0FD90CA77FD2D" pageId="13" pageNumber="432">
<treatmentCitation id="0AE5EC60FFED470AFCC0FD90CC18FDBA" author="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N." page="204" pageId="13" pageNumber="432" year="1976">
<taxonomicName id="4C44B1F2FFED470AFCC0FD90CAFDFDD0" ID-CoL="7VHQH" authority=": KRYZHANOVSKIJ &amp; REICHARDT (1976)" authorityName="KRYZHANOVSKIJ &amp; REICHARDT" authorityYear="1976" box="[828,1433,553,572]" class="Insecta" family="Histeridae" genus="Hypocacculus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="puncticollis" subGenus="Nessus">
<emphasis id="B9301663FFED470AFCC0FD90CCD7FDD0" box="[828,947,553,572]" italics="true" pageId="13" pageNumber="432">Hypocacculus</emphasis>
(
<emphasis id="B9301663FFED470AFC41FD90CC93FDD0" box="[957,1015,553,572]" italics="true" pageId="13" pageNumber="432">Nessus</emphasis>
) 
<emphasis id="B9301663FFED470AFBFEFD90CB07FDD0" box="[1026,1123,553,572]" italics="true" pageId="13" pageNumber="432">puncticollis</emphasis>
: 
<bibRefCitation id="EFD5B780FFED470AFB91FD90CAFDFDD0" author="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N." box="[1133,1433,553,572]" pageId="13" pageNumber="432" refId="ref20992" refString="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N. 1976: Zhuki nadsemeystva Histeroidea (semeystva Sphaeritidae, Histeridae, Synteliidae). Fauna SSSR, Zhestokrylye, Tom V, vyp. 4. [Beetles of the superfamily Histeroidea (families Sphaeritidae, Histeridae, Syntelidae) Fauna SSSR, Coleoptera, Vol. 5, part 4.]. Nauka, Leningrad, 434 pp (in Russian)." type="book" year="1976">KRYZHANOVSKIJ &amp; REICHARDT (1976)</bibRefCitation>
</taxonomicName>
: 204
</treatmentCitation>
, 213 (keyed, redescription); 
<treatmentCitation id="0AE5EC60FFED470AFB87FDFACA4DFDBA" author="VIENNA P." box="[1147,1321,579,598]" page="179" pageId="13" pageNumber="432" year="1980">
<bibRefCitation id="EFD5B780FFED470AFB87FDFACB98FDBA" author="VIENNA P." box="[1147,1276,579,598]" pageId="13" pageNumber="432" refId="ref21978" refString="VIENNA P. 1980: Fauna d'Italia. Vol. XVI. Coleoptera Histeridae. Bologna, Calderini Press, ix + 386 pp." type="book" year="1980">
<collectingRegion id="49800493FFED470AFB87FDFACBDCFDB9" box="[1147,1208,579,598]" country="Austria" name="Wien" pageId="13" pageNumber="432">VIENNA</collectingRegion>
(1980)
</bibRefCitation>
: 179
</treatmentCitation>
, 181 (keyed, redescription, incl. fig. 64b); 
<treatmentCitation id="0AE5EC60FFED470AFBA8FDE7CB8BFD9D" author="MAZUR S." box="[1108,1263,606,625]" page="89" pageId="13" pageNumber="432" year="1984">
<bibRefCitation id="EFD5B780FFED470AFBA8FDE7CBB5FD9D" author="MAZUR S." box="[1108,1233,606,625]" pageId="13" pageNumber="432" pagination="1 - 376" refId="ref21359" refString="MAZUR S. 1984: A world catalogue of Histeridae. Polskie Pismo Entomologiczne 54: 1 - 376." type="journal article" year="1984">MAZUR (1984)</bibRefCitation>
: 89
</treatmentCitation>
(catalogue); 
<treatmentCitation id="0AE5EC60FFED470AFA9DFDE7CCD9FD60" author="MAZUR S." page="254" pageId="13" pageNumber="432" year="1997">
<bibRefCitation id="EFD5B780FFED470AFA9DFDE7CCF2FD60" author="MAZUR S." pageId="13" pageNumber="432" pagination="1 - 373" refId="ref21379" refString="MAZUR S. 1997: A world catalogue of the Histeridae (Coleoptera: Histeridae). Genus Supplement 7: 1 - 373." type="journal article" year="1997">MAZUR (1997)</bibRefCitation>
: 254
</treatmentCitation>
(catalogue); 
<treatmentCitation id="0AE5EC60FFED470AFBD7FDC0CBBBFD60" author="YELAMOS T." box="[1067,1247,633,652]" page="320" pageId="13" pageNumber="432" year="2002">
<bibRefCitation id="EFD5B780FFED470AFBD7FDC0CBDCFD60" author="YELAMOS T." box="[1067,1208,633,652]" pageId="13" pageNumber="432" refId="ref22068" refString="YELAMOS T. 2002: Coleoptera, Histeridae. In: RAMOS M. A., TERCE- DOR J. A, BELLES-ROS X., GOSALBEZ-NOGUERA J., SIERRA A. G., MAYOL E. M., PIERA F. M., MARINO J. S. &amp; GONZALEZ J. T. (eds): Fauna Iberica. Vol. 17. Museo Nacional de Ciencias Naturales, CSCI, Madrid, 411 pp." type="book" year="2002">YÉLAMOS (2002)</bibRefCitation>
: 320
</treatmentCitation>
(keyed, redescription, incl. fig. 157f); 
<treatmentCitation id="0AE5EC60FFED470AFC18FD2ACBE5FD4A" author="MAZUR S." box="[996,1153,659,678]" page="94" pageId="13" pageNumber="432" year="2004">
<bibRefCitation id="EFD5B780FFED470AFC18FD2ACB06FD4A" author="MAZUR S." box="[996,1122,659,678]" pageId="13" pageNumber="432" refId="ref21404" refString="MAZUR S. 2004: Family Histeridae. Pp. 68 - 102. In: LOBL I. &amp; SMETANA A. (eds): Catalogue of Palaearctic Coleoptera, Vol. 2, Hydrophiloidea-Histeroidea-Staphylinidea. Apollo Books, Stenstrup, 942 pp." type="book" year="2004">MAZUR (2004)</bibRefCitation>
: 94
</treatmentCitation>
(catalogue); 
<treatmentCitation id="0AE5EC60FFED470AFB09FD2ACAF8FD4A" author="MAZUR S." box="[1269,1436,659,678]" page="209" pageId="13" pageNumber="432" year="2011">
<bibRefCitation id="EFD5B780FFED470AFB09FD2ACA16FD4A" author="MAZUR S." box="[1269,1394,659,678]" pageId="13" pageNumber="432" refId="ref21450" refString="MAZUR S. 2011: A concise catalogue of the Histeridae (Insecta: Coleoptera). Warsaw Univeristy of Life Sciences - SGGW Press, Warsaw, 332 pp." type="book" year="2011">MAZUR (2011)</bibRefCitation>
: 209
</treatmentCitation>
(catalogue); 
<treatmentCitation id="0AE5EC60FFED470AFC39FD17CBC3FD2D" author="LACKNER T. &amp; MAZUR S. &amp; NEWTON A. F." box="[965,1191,686,705]" page="118" pageId="13" pageNumber="432" year="2015">
<bibRefCitation id="EFD5B780FFED470AFC39FD17CB1BFD2D" author="LACKNER T. &amp; MAZUR S. &amp; NEWTON A. F." box="[965,1151,686,705]" pageId="13" pageNumber="432" refId="ref21168" refString="LACKNER T., MAZUR S. &amp; NEWTON A. F. 2015: Family Histeridae. Pp. 76 - 130. In: LOBL I. &amp; LOBL D. (eds): Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Staphylinoidea. Volume 1. Brill Publishers, Leiden, Boston, xxiii + 1 - 900." type="book" year="2015">LACKNER et al. (2015)</bibRefCitation>
: 118
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFBCA71FFED470AFCC0FD70CBFEFCFD" blockId="13.[828,1438,252,998]" pageId="13" pageNumber="432">
<treatmentCitationGroup id="AB54ED5FFFED470AFCC0FD70CBFEFCFD" pageId="13" pageNumber="432">
<taxonomicName id="4C44B1F2FFED470AFCC0FD70CB86FD30" ID-CoL="9N38S" authority="Jacquelin du Val, 1858: 99" authorityName="Jacquelin du Val" authorityPageNumber="99" authorityYear="1858" box="[828,1250,713,732]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="cribellaticollis">
<emphasis id="B9301663FFED470AFCC0FD70CB61FD30" box="[828,1029,713,732]" italics="true" pageId="13" pageNumber="432">Saprinus cribellaticollis</emphasis>
<bibRefCitation id="EFD5B780FFED470AFBF5FD70CB86FD30" author="JACQUELIN DU VAL M." box="[1033,1250,713,732]" pageId="13" pageNumber="432" pagination="97 - 108" refId="ref20919" refString="JACQUELIN DU VAL M. 1858: Genere des Coleopteres d'Europe comprenant leur classification et famillies naturelles, la description de tous les genres, des tableaux synoptiques destines a faciliter l'etude, le catalogue de toutes les especes de nombreux dessins au trait ce caracteres. II. A. Deyrolle, Paris, pp. 1 - 168. Cat. pp. 53 - 83, 97 - 108." type="journal article" year="1858">Jacquelin du Val, 1858:99</bibRefCitation>
</taxonomicName>
(original description). Fauvel in 
<treatmentCitation id="0AE5EC60FFED470AFC4CFD5ACB2FFD1A" author="GOZIS M." box="[944,1099,739,758]" page="202" pageId="13" pageNumber="432" year="1886">
<bibRefCitation id="EFD5B780FFED470AFC4CFD5ACB46FD1A" author="GOZIS M." box="[944,1058,739,758]" pageId="13" pageNumber="432" pagination="152 - 213" refId="ref20776" refString="GOZIS M. 1886: Les Histeridae Gallo-Rhenans. Tableaux traduits et abreges de l'allemand de Joh. Schmidt. Avec catalogue supplementaire par Albert Fauvel. Revue d'Entomologie 5: 152 - 213." type="journal article" year="1886">GOZIS (1886)</bibRefCitation>
: 202
</treatmentCitation>
(as synonym of 
<taxonomicName id="4C44B1F2FFED470AFB27FD5ACAF4FD1A" ID-CoL="9N3B7" authorityName=": GANGLBAUER" authorityYear="1899" box="[1243,1424,739,758]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="B9301663FFED470AFB27FD5ACAF4FD1A" box="[1243,1424,739,758]" italics="true" pageId="13" pageNumber="432">Saprinus puncticollis</emphasis>
</taxonomicName>
). 
<treatmentCitation id="0AE5EC60FFED470AFCA6FD47CB74FCFD" author="MARSEUL S. A." box="[858,1040,766,785]" page="509" pageId="13" pageNumber="432" year="1862">
<bibRefCitation id="EFD5B780FFED470AFCA6FD47CC8CFCFD" author="MARSEUL S. A." box="[858,1000,766,785]" pageId="13" pageNumber="432" pagination="677 - 758" refId="ref21276" refString="MARSEUL S. A. 1862: Supplement a la monographie des Histerides. (Suite). Annales de la Societe Entomologique de France, Quartieme Serie 3: 83 - 165, 327 - 506, 677 - 758." type="journal article" year="1862">MARSEUL (1862)</bibRefCitation>
: 509
</treatmentCitation>
(redescription).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFBCA71FFED470AFCC0FCA0CAEAFCC0" blockId="13.[828,1438,252,998]" box="[828,1422,793,812]" pageId="13" pageNumber="432">
<treatmentCitationGroup id="AB54ED5FFFED470AFCC0FCA0CAEAFCC0" box="[828,1422,793,812]" pageId="13" pageNumber="432">
<treatmentCitation id="0AE5EC60FFED470AFCC0FCA0CA25FCC0" author="SCHMIDT J." box="[828,1345,793,812]" page="312" pageId="13" pageNumber="432" year="1885">
<taxonomicName id="4C44B1F2FFED470AFCC0FCA0CA7DFCC0" ID-CoL="9N38S" authority=": SCHMIDT (1885)" authorityName="SCHMIDT" authorityYear="1885" box="[828,1305,793,812]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="cribellaticollis" subGenus="Hypocaccus">
<emphasis id="B9301663FFED470AFCC0FCA0CCECFCC0" box="[828,904,793,812]" italics="true" pageId="13" pageNumber="432">Saprinus</emphasis>
(
<emphasis id="B9301663FFED470AFC68FCA0CC98FCC0" box="[916,1020,793,812]" italics="true" pageId="13" pageNumber="432">Hypocaccus</emphasis>
) 
<emphasis id="B9301663FFED470AFBF5FCA0CBE1FCC0" box="[1033,1157,793,812]" italics="true" pageId="13" pageNumber="432">cribellaticollis</emphasis>
: 
<bibRefCitation id="EFD5B780FFED470AFB6CFCA0CA7DFCC0" author="SCHMIDT J." box="[1168,1305,793,812]" pageId="13" pageNumber="432" pagination="279 - 330" refId="ref21874" refString="SCHMIDT J. 1885: Bestimmungs-Tabellen der europaischen Coleopteren. XIV. Histeridae. Berliner Entomologische Zeitschrift 29: 279 - 330." type="journal article" year="1885">SCHMIDT (1885)</bibRefCitation>
</taxonomicName>
: 312
</treatmentCitation>
(keyed).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFBCA71FFED470AFCC0FC8ACCAEFC90" blockId="13.[828,1438,252,998]" pageId="13" pageNumber="432">
<treatmentCitationGroup id="AB54ED5FFFED470AFCC0FC8ACCAEFC90" pageId="13" pageNumber="432">
<treatmentCitation id="0AE5EC60FFED470AFCC0FC8ACBEFFCAA" author="MARSEUL S. A." box="[828,1163,819,838]" page="490" pageId="13" pageNumber="432" year="1862">
<taxonomicName id="4C44B1F2FFED470AFCC0FC8ACBEFFCAA" ID-CoL="9N3C4" authority="Marseul, 1862: 490" authorityName="Marseul" authorityPageNumber="490" authorityYear="1862" box="[828,1163,819,838]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="sicanus">
<emphasis id="B9301663FFED470AFCC0FC8ACCB7FCAA" box="[828,979,819,838]" italics="true" pageId="13" pageNumber="432">Saprinus sicanus</emphasis>
<bibRefCitation id="EFD5B780FFED470AFC20FC8ACBEFFCAA" author="MARSEUL S. A." box="[988,1163,819,838]" pageId="13" pageNumber="432" pagination="677 - 758" refId="ref21276" refString="MARSEUL S. A. 1862: Supplement a la monographie des Histerides. (Suite). Annales de la Societe Entomologique de France, Quartieme Serie 3: 83 - 165, 327 - 506, 677 - 758." type="journal article" year="1862">Marseul, 1862: 490</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description, incl. pl. XVII, fig. 47). 
<treatmentCitation id="0AE5EC60FFED470AFC26FCF7CBA6FC8D" author="BAUDI DI SELVE F." box="[986,1218,846,865]" page="233" pageId="13" pageNumber="432" year="1864">
<bibRefCitation id="EFD5B780FFED470AFC26FCF7CBFDFC8D" author="BAUDI DI SELVE F." box="[986,1177,846,865]" pageId="13" pageNumber="432" pagination="195 - 233" refId="ref20435" refString="BAUDI DI SELVE F. 1864: Coleopterorum messis in insula Cypro et Asia minore ab Eugenio Truqui congregatae recensito: de Europaeis notis quibusdam additis. Pars prima. Berliner Entomologische Zeitschrift 8: 195 - 233." type="journal article" year="1864">BAUDI DI SELVE (1864)</bibRefCitation>
: 233
</treatmentCitation>
(as synonym of 
<taxonomicName id="4C44B1F2FFED470AFAACFCF7CCDAFC90" ID-CoL="9N3B7" authorityName=": GANGLBAUER" authorityYear="1899" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="B9301663FFED470AFAACFCF7CCDAFC90" italics="true" pageId="13" pageNumber="432">Saprinus puncticollis</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFBCA71FFED470AFCC0FC3ACBC1FC5D" blockId="13.[828,1438,252,998]" pageId="13" pageNumber="432">
<treatmentCitationGroup id="AB54ED5FFFED470AFCC0FC3ACBC1FC5D" pageId="13" pageNumber="432">
<treatmentCitation id="0AE5EC60FFED470AFCC0FC3ACB16FC7A" author="MARSEUL S. A." box="[828,1138,899,918]" page="715" pageId="13" pageNumber="432" year="1862">
<taxonomicName id="4C44B1F2FFED470AFCC0FC3ACB16FC7A" authority="Marseul, 1862: 715" authorityName="Marseul" authorityPageNumber="715" authorityYear="1862" box="[828,1138,899,918]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="kuesteri">
<emphasis id="B9301663FFED470AFCC0FC3ACCA8FC7A" box="[828,972,899,918]" italics="true" pageId="13" pageNumber="432">Saprinus kuesteri</emphasis>
<bibRefCitation id="EFD5B780FFED470AFC2CFC3ACB16FC7A" author="MARSEUL S. A." box="[976,1138,899,918]" pageId="13" pageNumber="432" pagination="677 - 758" refId="ref21276" refString="MARSEUL S. A. 1862: Supplement a la monographie des Histerides. (Suite). Annales de la Societe Entomologique de France, Quartieme Serie 3: 83 - 165, 327 - 506, 677 - 758." type="journal article" year="1862">Marseul, 1862: 715</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(catalogue; unecessary replacement name for 
<taxonomicName id="4C44B1F2FFED470AFC50FC27CBFFFC5D" ID-CoL="9N3B7" authority="Kuster, 1849" authorityName="Kuster" authorityYear="1849" box="[940,1179,926,945]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="B9301663FFED470AFC50FC27CB41FC5D" box="[940,1061,926,945]" italics="true" pageId="13" pageNumber="432">S. puncticollis</emphasis>
<bibRefCitation id="EFD5B780FFED470AFBD6FC27CBFFFC5D" author="KUSTER H. C." box="[1066,1179,926,945]" pageId="13" pageNumber="432" refId="ref21108" refString="KUSTER H. C. 1849: Die Kafer Europa's. Nach der Natur beschreiben. XVII Heft. Bauer and Raspe, Nurnberg, 100 pls." type="book" year="1849">Küster, 1849</bibRefCitation>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFBCA71FFED470AFCC0FC00CBB1FC0A" blockId="13.[828,1438,252,998]" pageId="13" pageNumber="432">
<treatmentCitationGroup id="AB54ED5FFFED470AFCC0FC00CBB1FC0A" pageId="13" pageNumber="432">
<taxonomicName id="4C44B1F2FFED470AFCC0FC00CB10FC20" ID-CoL="9N3BJ" authority="Marseul, 1876: 39" authorityName="Marseul" authorityPageNumber="39" authorityYear="1876" box="[828,1140,953,972]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="revisus">
<emphasis id="B9301663FFED470AFCC0FC00CCAFFC20" box="[828,971,953,972]" italics="true" pageId="13" pageNumber="432">Saprinus revisus</emphasis>
Marseul, 1876: 39
</taxonomicName>
(original description). 
<treatmentCitation id="0AE5EC60FFED470AFABDFC00CCD7FC0A" author="BICKHARDT H." page="97" pageId="13" pageNumber="432" year="1916">
<bibRefCitation id="EFD5B780FFED470AFABDFC00CCF1FC0A" author="BICKHARDT H." pageId="13" pageNumber="432" refId="ref20506" refString="BICKHARDT H. 1916 - 1917: Histeridae. In: WYTSMAN P. (ed.): Genera Insectorum. Fasc. 166 a, b. La Haye, 302 pp." type="book" year="1916">BICKHARDT (1916)</bibRefCitation>
: 97
</treatmentCitation>
(as synonym of 
<taxonomicName id="4C44B1F2FFED470AFBBEFC6ACBACFC0A" authorityName=": GANGLBAUER" authorityYear="1899" box="[1090,1224,979,998]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="curtus">
<emphasis id="B9301663FFED470AFBBEFC6ACBACFC0A" box="[1090,1224,979,998]" italics="true" pageId="13" pageNumber="432">Saprinus curtus</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35E99FAFFED4709FCC0FC44CD15FBE0" lastPageId="14" lastPageNumber="433" pageId="13" pageNumber="432" type="materials_examined">
<paragraph id="8BFBCA71FFED470AFCC0FC44CBC6FA87" blockId="13.[826,1438,1021,2080]" pageId="13" pageNumber="432">
<emphasis id="B9301663FFED470AFCC0FC44CB7DFBFC" bold="true" box="[828,1049,1021,1040]" pageId="13" pageNumber="432">Type material examined.</emphasis>
<taxonomicName id="4C44B1F2FFED470AFBE0FC44CA59FBFC" authority="Rosenhauer, 1847" authorityName="Rosenhauer" authorityYear="1847" box="[1052,1341,1021,1040]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="curtus">
<emphasis id="B9301663FFED470AFBE0FC44CBC4FBFC" box="[1052,1184,1021,1040]" italics="true" pageId="13" pageNumber="432">Saprinus curtus</emphasis>
<bibRefCitation id="EFD5B780FFED470AFB58FC44CA59FBFC" author="ROSENHAUER W. G." box="[1188,1341,1021,1040]" pageId="13" pageNumber="432" refId="ref21812" refString="ROSENHAUER W. G. 1847: Beitrage zur Insekten-Fauna Europas. Erstes Bandchen; enthalt die Beschreibung von sechzig neuen Kafern aus Bayern, Tyrol, Ungarn, etc., so wie die Kafer Tyrols, nach dem Ergebnisse von vier Reisen zusammengestellt. Mit einer Tafel Abbildungen. Th. Blasing, Erlangen, x + 159 pp." type="book" year="1847">Rosenhauer, 1847</bibRefCitation>
</taxonomicName>
. 
<typeStatus id="54FF74D3FFED470AFAB8FC44CAF8FBE2" box="[1348,1436,1021,1040]" pageId="13" pageNumber="432" type="lectotype">LECTOTYPE</typeStatus>
(present designation): ♁ (
<figureCitation id="137FD6F4FFED470AFBE9FBA1CB31FBC7" box="[1045,1109,1048,1067]" captionStart="Figs 40–43" captionStartId="15.[151,188,2006,2025]" captionTargetBox="[212,1382,479,1970]" captionTargetId="figure-17@15.[201,1387,467,1981]" captionTargetPageId="15" captionText="Figs 40–43. Hypocaccus (Nessus) curtus (Rosenhauer, 1847), dorsal habitus.40 – Saprinus curtus Rosenhauer, 1847, lectotype.41 – Saprinus puncticollis Küster, 1849, lectotype. 42 – Saprinus cribellaticollis Jacquelin Du Val, 1858, lectotype. 43 – Saprinus sicanus Marseul, 1862, lectotype." figureDoi="http://doi.org/10.5281/zenodo.4504944" httpUri="https://zenodo.org/record/4504944/files/figure.png" pageId="13" pageNumber="432">Fig. 40</figureCitation>
), originally pinned with pin-hole in its right elytron, mounted on a rectangular mounting card, right antennal funicle and left mesotarsus missing, genitalia extracted and disarticulated, glued to the same mounting card as the specimen, ‘curtus / Rosenh. [written] // Hungaria [written] // herbeus Mars. [written] // Ex Musaeo / 
<emphasis id="B9301663FFED470AFCBBFB24CCD0FB5C" bold="true" box="[839,948,1181,1200]" pageId="13" pageNumber="432">Rosenhauer</emphasis>
[black-margined, printed label] // pas synonime / d’Herbeus Mars. / Dr. Auzat 1917 [written-printed] // 
<collectingCountry id="F3538AE1FFED470AFB1DFB01CA43FB27" box="[1249,1319,1208,1227]" name="Hungary" pageId="13" pageNumber="432">Hongrie</collectingCountry>
/ Ex-Musaeo / ROSENHAUER [printed] // 
<taxonomicName id="4C44B1F2FFED470AFBBAFB6BCBDBFB09" authorityName="Bickhardt" authorityYear="1914" box="[1094,1215,1234,1253]" class="Insecta" family="Histeridae" genus="Hypocacculus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="genus">
<emphasis id="B9301663FFED470AFBBAFB6BCBDBFB09" box="[1094,1215,1234,1253]" italics="true" pageId="13" pageNumber="432">Hypocacculus</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C44B1F2FFED470AFB28FB6BCAF8FB09" authorityName="Rosenhauer" authorityYear="1847" box="[1236,1436,1234,1253]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="curtus" subGenus="Nannolepidius">
(
<emphasis id="B9301663FFED470AFB20FB6BCA3CFB09" box="[1244,1368,1234,1253]" italics="true" pageId="13" pageNumber="432">Nannolepidius</emphasis>
) 
<emphasis id="B9301663FFED470AFA9BFB6BCAF8FB09" box="[1383,1436,1234,1253]" italics="true" pageId="13" pageNumber="432">curtus</emphasis>
</taxonomicName>
/ (
<bibRefCitation id="EFD5B780FFED470AFCACFB54CC95FAEC" author="ROSENHAUER W. G." box="[848,1009,1261,1280]" pageId="13" pageNumber="432" refId="ref21812" refString="ROSENHAUER W. G. 1847: Beitrage zur Insekten-Fauna Europas. Erstes Bandchen; enthalt die Beschreibung von sechzig neuen Kafern aus Bayern, Tyrol, Ungarn, etc., so wie die Kafer Tyrols, nach dem Ergebnisse von vier Reisen zusammengestellt. Mit einer Tafel Abbildungen. Th. Blasing, Erlangen, x + 159 pp." type="book" year="1847">Rosenhauer, 1847</bibRefCitation>
) / Dr. Auzat Dét. 1917 [printed] // Exemplaire provenant de la / collection Vauloger de Beaupré / Marcel (1862-1904) et inclus dans / la collection S. Risser en 2011 [black-margined, printed label] // 
<taxonomicName id="4C44B1F2FFED470AFC7AFA84CB6AFABC" authorityName="Rosenhauer" authorityYear="1847" box="[902,1038,1341,1360]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="curtus">
<emphasis id="B9301663FFED470AFC7AFA84CB6AFABC" box="[902,1038,1341,1360]" italics="true" pageId="13" pageNumber="432">Saprinus curtus</emphasis>
</taxonomicName>
/ 
<bibRefCitation id="EFD5B780FFED470AFBDEFA84CBD9FABC" author="ROSENHAUER W. G." box="[1058,1213,1341,1360]" pageId="13" pageNumber="432" refId="ref21812" refString="ROSENHAUER W. G. 1847: Beitrage zur Insekten-Fauna Europas. Erstes Bandchen; enthalt die Beschreibung von sechzig neuen Kafern aus Bayern, Tyrol, Ungarn, etc., so wie die Kafer Tyrols, nach dem Ergebnisse von vier Reisen zusammengestellt. Mit einer Tafel Abbildungen. Th. Blasing, Erlangen, x + 159 pp." type="book" year="1847">Rosenhauer, 1847</bibRefCitation>
/ 
<typeStatus id="54FF74D3FFED470AFB2DFA84CA2FFABC" box="[1233,1355,1341,1360]" pageId="13" pageNumber="432" type="lectotype">LECTOTYPE</typeStatus>
/ Des. T. Lackner 2017 [red label, printed]’ (
<collectionCode id="ED5552B4FFED470AFB9BFAE1CBF3FA87" box="[1127,1175,1368,1387]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="13" pageNumber="432" type="Museum">ZSM</collectionCode>
).
</paragraph>
<paragraph id="8BFBCA71FFED470AFCA6FACBCA4DF9C9" blockId="13.[826,1438,1021,2080]" pageId="13" pageNumber="432">
<taxonomicName id="4C44B1F2FFED470AFCA6FACBCBE4FA69" authority="Kuster, 1849" authorityName="Kuster" authorityYear="1849" box="[858,1152,1394,1413]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="B9301663FFED470AFCA6FACBCB6AFA69" box="[858,1038,1394,1413]" italics="true" pageId="13" pageNumber="432">Saprinus puncticollis</emphasis>
<bibRefCitation id="EFD5B780FFED470AFBEDFACBCBE4FA69" author="KUSTER H. C." box="[1041,1152,1394,1413]" pageId="13" pageNumber="432" refId="ref21108" refString="KUSTER H. C. 1849: Die Kafer Europa's. Nach der Natur beschreiben. XVII Heft. Bauer and Raspe, Nurnberg, 100 pls." type="book" year="1849">Küster, 1849</bibRefCitation>
</taxonomicName>
. 
<typeStatus id="54FF74D3FFED470AFB7BFACBCB84FA68" box="[1159,1248,1394,1413]" pageId="13" pageNumber="432" type="lectotype">LECTOTYPE</typeStatus>
(present designation): ♁ (
<figureCitation id="137FD6F4FFED470AFCABFA34CCFCFA4C" box="[855,920,1421,1440]" captionStart="Figs 40–43" captionStartId="15.[151,188,2006,2025]" captionTargetBox="[212,1382,479,1970]" captionTargetId="figure-17@15.[201,1387,467,1981]" captionTargetPageId="15" captionText="Figs 40–43. Hypocaccus (Nessus) curtus (Rosenhauer, 1847), dorsal habitus.40 – Saprinus curtus Rosenhauer, 1847, lectotype.41 – Saprinus puncticollis Küster, 1849, lectotype. 42 – Saprinus cribellaticollis Jacquelin Du Val, 1858, lectotype. 43 – Saprinus sicanus Marseul, 1862, lectotype." figureDoi="http://doi.org/10.5281/zenodo.4504944" httpUri="https://zenodo.org/record/4504944/files/figure.png" pageId="13" pageNumber="432">Fig. 42</figureCitation>
), glued onto a rectangular mounting card, two left and three right mesotarsomeres missing, genitalia extracted, disarticulated and glued to the same mounting card as the specimen, ‘Typ! [written] // Cagliari / Dr. Küster [written] // puncticollis / Küst. [written] // 
<taxonomicName id="4C44B1F2FFED470AFAB4FA64CAF6FA1C" authorityName="Erichson" authorityYear="1834" box="[1352,1426,1501,1520]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
/ 
<taxonomicName id="4C44B1F2FFED470AFCC0FA41CCDDF9E7" authority="Rosenh." authorityName="Rosenhauer" authorityYear="1847" box="[828,953,1528,1547]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="curtus">curtus Rosenh.</taxonomicName>
[written] // 
<taxonomicName id="4C44B1F2FFED470AFBE5FA41CBAFF9E7" authorityName="Kuster" authorityYear="1849" box="[1049,1227,1528,1547]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="B9301663FFED470AFBE5FA41CBAFF9E7" box="[1049,1227,1528,1547]" italics="true" pageId="13" pageNumber="432">Saprinus puncticollis</emphasis>
</taxonomicName>
/ 
<bibRefCitation id="EFD5B780FFED470AFB24FA41CA27F9E7" author="KUSTER H. C." box="[1240,1347,1528,1547]" pageId="13" pageNumber="432" refId="ref21108" refString="KUSTER H. C. 1849: Die Kafer Europa's. Nach der Natur beschreiben. XVII Heft. Bauer and Raspe, Nurnberg, 100 pls." type="book" year="1849">Küster, 1849</bibRefCitation>
/ LECTOTYPE / Des. T. Lackner 2017 [red label, printed]’ (
<collectionCode id="ED5552B4FFED470AFB13F9ABCA7BF9C9" box="[1263,1311,1554,1573]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="13" pageNumber="432" type="Museum">ZSM</collectionCode>
).
</paragraph>
<paragraph id="8BFBCA71FFED470AFCA6F994CAF4F917" blockId="13.[826,1438,1021,2080]" pageId="13" pageNumber="432">
<taxonomicName id="4C44B1F2FFED470AFCA6F994CB95F9AC" authority="Jacquelin du Val, 1858" authorityName="Jacquelin du Val" authorityYear="1858" box="[858,1265,1581,1600]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="cribellaticollis">
<emphasis id="B9301663FFED470AFCA6F994CB43F9AC" box="[858,1063,1581,1600]" italics="true" pageId="13" pageNumber="432">Saprinus cribellaticollis</emphasis>
<bibRefCitation id="EFD5B780FFED470AFBD1F994CB95F9AC" author="JACQUELIN DU VAL M." box="[1069,1265,1581,1600]" pageId="13" pageNumber="432" pagination="97 - 108" refId="ref20919" refString="JACQUELIN DU VAL M. 1858: Genere des Coleopteres d'Europe comprenant leur classification et famillies naturelles, la description de tous les genres, des tableaux synoptiques destines a faciliter l'etude, le catalogue de toutes les especes de nombreux dessins au trait ce caracteres. II. A. Deyrolle, Paris, pp. 1 - 168. Cat. pp. 53 - 83, 97 - 108." type="journal article" year="1858">Jacquelin du Val, 1858</bibRefCitation>
</taxonomicName>
. 
<typeStatus id="54FF74D3FFED470AFB05F994CA36F9D2" box="[1273,1362,1581,1600]" pageId="13" pageNumber="432" type="lectotype">LECTOTYPE</typeStatus>
(present designation): 
<specimenCount id="9D4201F8FFED470AFC53F9FDCCDBF9B7" box="[943,959,1604,1627]" pageId="13" pageNumber="432" type="female">♀</specimenCount>
(
<figureCitation id="137FD6F4FFED470AFC35F9F1CB62F9B7" box="[969,1030,1608,1627]" captionStart="Figs 40–43" captionStartId="15.[151,188,2006,2025]" captionTargetBox="[212,1382,479,1970]" captionTargetId="figure-17@15.[201,1387,467,1981]" captionTargetPageId="15" captionText="Figs 40–43. Hypocaccus (Nessus) curtus (Rosenhauer, 1847), dorsal habitus.40 – Saprinus curtus Rosenhauer, 1847, lectotype.41 – Saprinus puncticollis Küster, 1849, lectotype. 42 – Saprinus cribellaticollis Jacquelin Du Val, 1858, lectotype. 43 – Saprinus sicanus Marseul, 1862, lectotype." figureDoi="http://doi.org/10.5281/zenodo.4504944" httpUri="https://zenodo.org/record/4504944/files/figure.png" pageId="13" pageNumber="432">Fig.41</figureCitation>
), glued on a rectangular mounting card, both antennal funicles broken off; legs: except for right foreleg and left foretibia, all tibiae broken off; with the following labels:tiny, green rectangular label that is glued onto much larger translucent plastic mounting card (original mounting card of J. du Val) and tiny, red, quadrate label, followed by, ‘ 
<taxonomicName id="4C44B1F2FFED470AFCBFF974CB6BF90C" authorityName="Jacquelin du Val" authorityYear="1858" box="[835,1039,1741,1760]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="cribellaticollis">
<emphasis id="B9301663FFED470AFCBFF974CB6BF90C" box="[835,1039,1741,1760]" italics="true" pageId="13" pageNumber="432">Saprinus cribellaticollis</emphasis>
</taxonomicName>
/ 
<bibRefCitation id="EFD5B780FFED470AFBE3F974CBBAF90C" author="JACQUELIN DU VAL M." box="[1055,1246,1741,1760]" pageId="13" pageNumber="432" pagination="97 - 108" refId="ref20919" refString="JACQUELIN DU VAL M. 1858: Genere des Coleopteres d'Europe comprenant leur classification et famillies naturelles, la description de tous les genres, des tableaux synoptiques destines a faciliter l'etude, le catalogue de toutes les especes de nombreux dessins au trait ce caracteres. II. A. Deyrolle, Paris, pp. 1 - 168. Cat. pp. 53 - 83, 97 - 108." type="journal article" year="1858">Jacquelin du Val, 1858</bibRefCitation>
/ 
<typeStatus id="54FF74D3FFED470AFB11F974CA03F90C" box="[1261,1383,1741,1760]" pageId="13" pageNumber="432" type="lectotype">LECTOTYPE</typeStatus>
/ Des. T. Lackner 2017 [red label, printed]’ (
<collectionCode id="ED5552B4FFED470AFB81F951CBA6F917" box="[1149,1218,1768,1787]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="432">MNHN</collectionCode>
; coll. Jacquelin du Val).
</paragraph>
<paragraph id="8BFBCA71FFED470AFCA6F8BBCAEAF7CC" blockId="13.[826,1438,1021,2080]" pageId="13" pageNumber="432">
<taxonomicName id="4C44B1F2FFED470AFCA6F8BBCB0FF8F9" authority="Marseul, 1862" authorityName="Marseul" authorityYear="1862" box="[858,1131,1794,1813]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="sicanus">
<emphasis id="B9301663FFED470AFCA6F8BBCC8EF8F9" box="[858,1002,1794,1813]" italics="true" pageId="13" pageNumber="432">Saprinus sicanus</emphasis>
<bibRefCitation id="EFD5B780FFED470AFC12F8BBCB0FF8F9" author="MARSEUL S. A." box="[1006,1131,1794,1813]" pageId="13" pageNumber="432" pagination="677 - 758" refId="ref21276" refString="MARSEUL S. A. 1862: Supplement a la monographie des Histerides. (Suite). Annales de la Societe Entomologique de France, Quartieme Serie 3: 83 - 165, 327 - 506, 677 - 758." type="journal article" year="1862">Marseul, 1862</bibRefCitation>
</taxonomicName>
. 
<typeStatus id="54FF74D3FFED470AFB8EF8BBCBAFF8F8" box="[1138,1227,1794,1813]" pageId="13" pageNumber="432" type="lectotype">LECTOTYPE</typeStatus>
(present designation):♁ (
<figureCitation id="137FD6F4FFED470AFCBEF8A4CCE5F8DC" box="[834,897,1821,1840]" captionStart="Figs 40–43" captionStartId="15.[151,188,2006,2025]" captionTargetBox="[212,1382,479,1970]" captionTargetId="figure-17@15.[201,1387,467,1981]" captionTargetPageId="15" captionText="Figs 40–43. Hypocaccus (Nessus) curtus (Rosenhauer, 1847), dorsal habitus.40 – Saprinus curtus Rosenhauer, 1847, lectotype.41 – Saprinus puncticollis Küster, 1849, lectotype. 42 – Saprinus cribellaticollis Jacquelin Du Val, 1858, lectotype. 43 – Saprinus sicanus Marseul, 1862, lectotype." figureDoi="http://doi.org/10.5281/zenodo.4504944" httpUri="https://zenodo.org/record/4504944/files/figure.png" pageId="13" pageNumber="432">Fig. 43</figureCitation>
), glued onto a rectangular mounting card, right antennal funicle, both protarsi, two segments of right mesotarsus, as well as both metatibiae missing, male genitalia extracted, disarticulated and glued onto the same mounting card as the specimen, with the following labels: small, square-shaped blue label, followed by, ‘ 
<taxonomicName id="4C44B1F2FFED470AFB71F831CBB8F877" authorityName="Erichson" authorityYear="1834" box="[1165,1244,1928,1947]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
/ sicanus m. / Schaum ‘59 [round label, written] // 129c / 
<taxonomicName id="4C44B1F2FFED470AFB98F81BCBCBF859" authorityName="Erichson" authorityYear="1834" box="[1124,1199,1954,1973]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
/ sicanus m. / 
<collectingRegion id="49800493FFED470AFAD6F81BCA3FF859" box="[1322,1371,1954,1973]" country="Italy" name="Sicilia" pageId="13" pageNumber="432">Sicile</collectingRegion>
/ Schm 679 [round label, written] // 47 (129c) Saprin / sicanus m60 / Sicil. [written] // MUSEUM PARIS / Coll. De Marseul / 2842-90 [printed] // TYPE [red-printed label; followed by: “ 
<taxonomicName id="4C44B1F2FFED470AFB7BF84BCA71F7E9" authorityName="Marseul" authorityYear="1862" box="[1159,1301,2034,2053]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="sicanus">
<emphasis id="B9301663FFED470AFB7BF84BCA71F7E9" box="[1159,1301,2034,2053]" italics="true" pageId="13" pageNumber="432">Saprinus sicanus</emphasis>
</taxonomicName>
/ 
<bibRefCitation id="EFD5B780FFED470AFADFF84BCAF8F7E9" author="MARSEUL S. A." box="[1315,1436,2034,2053]" pageId="13" pageNumber="432" pagination="677 - 758" refId="ref21276" refString="MARSEUL S. A. 1862: Supplement a la monographie des Histerides. (Suite). Annales de la Societe Entomologique de France, Quartieme Serie 3: 83 - 165, 327 - 506, 677 - 758." type="journal article" year="1862">Marseul, 1862</bibRefCitation>
/ 
<typeStatus id="54FF74D3FFED470AFCBBF7B4CCA5F7CC" box="[839,961,2061,2080]" pageId="13" pageNumber="432" type="lectotype">LECTOTYPE</typeStatus>
/ Des. T. Lackner 2017 [red label, printed]’ (
<collectionCode id="ED5552B4FFED470AFAC2F7B4CAE0F7CC" box="[1342,1412,2061,2080]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="432">MNHN</collectionCode>
).
</paragraph>
<paragraph id="8BFBCA71FFEE4709FF49FF07CEF3FE60" blockId="14.[150,761,190,1036]" pageId="14" pageNumber="433">
<taxonomicName id="4C44B1F2FFEE4709FF49FF07CEAAFF3D" authority="Marseul, 1876" authorityName="Marseul" authorityYear="1876" box="[181,462,190,209]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="433" phylum="Arthropoda" rank="species" species="revisus">
<emphasis id="B9301663FFEE4709FF49FF07CE22FF3D" box="[181,326,190,209]" italics="true" pageId="14" pageNumber="433">Saprinus revisus</emphasis>
Marseul, 1876
</taxonomicName>
. 
<typeStatus id="54FF74D3FFEE4709FE25FF07CD56FF23" box="[473,562,190,209]" pageId="14" pageNumber="433" type="lectotype">LECTOTYPE</typeStatus>
(present designation): 
<specimenCount id="9D4201F8FFEE4709FF6BFF6CCFC3FF00" box="[151,167,213,236]" pageId="14" pageNumber="433" type="female">♀</specimenCount>
(
<figureCitation id="137FD6F4FFEE4709FF4EFF60CF95FF00" box="[178,241,217,236]" captionStart="Figs 44–45" captionStartId="16.[151,188,2008,2027]" captionTargetBox="[156,1428,1206,1975]" captionTargetId="figure-13@16.[151,1436,1200,1986]" captionTargetPageId="16" captionText="Figs 44–45.Hypocaccus (Nessus) curtus (Rosenhauer, 1847), dorsal habitus. 44 – Saprinus revisus Marseul, 1876, lectotype. 45 – Hypocacculus (Nessus) controversus G. Müller, 1937, lectotype." figureDoi="http://doi.org/10.5281/zenodo.4504946" httpUri="https://zenodo.org/record/4504946/files/figure.png" pageId="14" pageNumber="433">Fig. 44</figureCitation>
), left antennal funicle, left protarsus, and left metatarsus missing, glued onto a rectangular mounting card, female genitalia extracted, glued to the same card as the specimen, ‘ 
<taxonomicName id="4C44B1F2FFEE4709FE17FEB7CD5EFECD" authorityName="Erichson" authorityYear="1834" box="[491,570,270,289]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="433" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
/ revisus / rest of label illegible [round, blue label, written] // MUSEUM PARIS / Coll. / De Marseul 1890 [light-green label, printed] // TYPE [red-printed label] // 
<taxonomicName id="4C44B1F2FFEE4709FF6BFEE7CE47FE9D" authorityName="Marseul" authorityYear="1876" box="[151,291,350,369]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="433" phylum="Arthropoda" rank="species" species="revisus">
<emphasis id="B9301663FFEE4709FF6BFEE7CE47FE9D" box="[151,291,350,369]" italics="true" pageId="14" pageNumber="433">Saprinus revisus</emphasis>
</taxonomicName>
/ Marseul, 1876 / 
<typeStatus id="54FF74D3FFEE4709FE47FEE7CD51FE9D" box="[443,565,350,369]" pageId="14" pageNumber="433" type="lectotype">LECTOTYPE</typeStatus>
/ Des. T. Lackner 2017 [red label, printed]’ (
<collectionCode id="ED5552B4FFEE4709FEBBFEC0CEE9FE60" box="[327,397,377,396]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="433">MNHN</collectionCode>
).
</paragraph>
<paragraph id="8BFBCA71FFEE4709FF6BFE2ACD15FBE0" blockId="14.[150,761,190,1036]" pageId="14" pageNumber="433">
<emphasis id="B9301663FFEE4709FF6BFE2ACD10FE4A" bold="true" box="[151,628,403,422]" pageId="14" pageNumber="433">
Additional material examined. 
<collectingCountry id="F3538AE1FFEE4709FE4BFE2ACD7AFE4A" box="[439,542,403,422]" name="Algeria" pageId="14" pageNumber="433">ALGERIA</collectingCountry>
: 
<collectingRegion id="49800493FFEE4709FDD4FE2ACD09FE48" box="[552,621,403,422]" country="Algeria" name="Annaba" pageId="14" pageNumber="433">ANNABA</collectingRegion>
:
</emphasis>
Bône [= 
<collectingRegion id="49800493FFEE4709FD3CFE2ACFCBFE2D" country="Algeria" name="Annaba" pageId="14" pageNumber="433">Annaba</collectingRegion>
], 
<specimenCount id="9D4201F8FFEE4709FF42FE17CFBAFE2D" box="[190,222,426,449]" pageId="14" pageNumber="433" type="female">1 ♀</specimenCount>
, coll. Dr. Buysson (
<collectionCode id="ED5552B4FFEE4709FE71FE17CEB6FE2D" box="[397,466,430,449]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="433">MNHN</collectionCode>
; coll. Thérond); Bône [= 
<collectingRegion id="49800493FFEE4709FD5BFE17CD94FE2D" box="[679,752,430,449]" country="Algeria" name="Annaba" pageId="14" pageNumber="433">Annaba</collectingRegion>
], 1 ♁, Desbr. (
<collectionCode id="ED5552B4FFEE4709FEFFFE70CE20FE30" box="[259,324,457,476]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="14" pageNumber="433">MFNB</collectionCode>
). 
<emphasis id="B9301663FFEE4709FEADFE70CEC5FE30" bold="true" box="[337,417,457,476]" pageId="14" pageNumber="433">
<collectingCountry id="F3538AE1FFEE4709FEADFE70CEF8FE30" box="[337,412,457,476]" name="Egypt" pageId="14" pageNumber="433">EGYPT</collectingCountry>
:
</emphasis>
Egypt, no further data, 
<specimenCount id="9D4201F8FFEE4709FD96FE70CDEDFE30" box="[618,649,453,476]" pageId="14" pageNumber="433" type="female">1 ♀</specimenCount>
, coll.Ancey, (
<collectionCode id="ED5552B4FFEE4709FF61FE5ACF86FE1A" box="[157,226,483,502]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="433">MNHN</collectionCode>
; coll. Thérond). 
<emphasis id="B9301663FFEE4709FE8BFE5ACDF7FE1A" bold="true" box="[375,659,483,502]" pageId="14" pageNumber="433">
<collectingCountry id="F3538AE1FFEE4709FE8BFE5ACEB7FE1A" box="[375,467,483,502]" name="France" pageId="14" pageNumber="433">FRANCE</collectingCountry>
: BOUCHES- DU- RHÔNE:
</emphasis>
Camargue, 2 ♁♁, L. Puel lgt., Auzat coll. (
<collectionCode id="ED5552B4FFEE4709FE47FE47CD65FDFD" box="[443,513,510,529]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="433">MNHN</collectionCode>
; coll. Thérond); Camargue, Vaccares, no date, 1 ♁, 
<date id="FFFAECB1FFEE4709FE95FDA0CEDBFDC0" box="[361,447,537,556]" pageId="14" pageNumber="433" value="1937-05-29">29.v.1937</date>
, 1 ♁, J. Thérond lgt. (
<collectionCode id="ED5552B4FFEE4709FD7FFDA0CDACFDC0" box="[643,712,537,556]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="433">MNHN</collectionCode>
; coll. Thérond); Camargue, La Sauvage, 
<date id="FFFAECB1FFEE4709FE38FD8ACD6BFDAA" box="[452,527,563,582]" pageId="14" pageNumber="433" value="1928-05-01">1.v.1928</date>
, 1 ♁, L. Puel lgt. (
<collectionCode id="ED5552B4FFEE4709FD53FD8ACD90FDAA" box="[687,756,563,582]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="433">MNHN</collectionCode>
; coll. Thérond); St. Maries de la Mer, 
<date id="FFFAECB1FFEE4709FE27FDF7CD5AFD8D" box="[475,574,590,609]" pageId="14" pageNumber="433" value="1922-07-18">18.vii.1922</date>
, 
<specimenCount id="9D4201F8FFEE4709FDB4FDF7CD0CFD8D" box="[584,616,586,609]" pageId="14" pageNumber="433" type="female">1 ♀</specimenCount>
, Dr. A. Chobaut lgt., coll.Dr.Auzat (
<collectionCode id="ED5552B4FFEE4709FEC3FDD0CEE0FD90" box="[319,388,617,636]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="433">MNHN</collectionCode>
; coll.Thérond). 
<emphasis id="B9301663FFEE4709FDF1FDD0CDCEFD90" bold="true" box="[525,682,617,636]" pageId="14" pageNumber="433">
<collectingCountry id="F3538AE1FFEE4709FDF1FDD0CD2AFD90" box="[525,590,617,636]" name="Italy" pageId="14" pageNumber="433">ITALY</collectingCountry>
: 
<collectingRegion id="49800493FFEE4709FDAAFDD0CDC6FD96" box="[598,674,617,636]" country="Italy" name="Sardegna" pageId="14" pageNumber="433">SARDINIA</collectingRegion>
:
</emphasis>
Cagliari, Saline di Stato, 
<date id="FFFAECB1FFEE4709FEDCFD3ACE12FD7A" box="[288,374,643,662]" pageId="14" pageNumber="433" value="1989-05-10">10.v.1989</date>
, 1 ♁, 
<specimenCount id="9D4201F8FFEE4709FE57FD3ACEBFFD7B" box="[427,475,640,663]" pageId="14" pageNumber="433" type="female">3 ♀♀</specimenCount>
, C. Meloni lgt. (1 ♁ in 
<collectionCode id="ED5552B4FFEE4709FD56FD3ACD80FD7A" box="[682,740,643,662]" pageId="14" pageNumber="433">CTLA</collectionCode>
, 
<specimenCount id="9D4201F8FFEE4709FD11FD3ACFD3FD5D" pageId="14" pageNumber="433" type="female">3 ♀♀</specimenCount>
in 
<collectionCode id="ED5552B4FFEE4709FF2BFD27CE7DFD5D" box="[215,281,670,689]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="14" pageNumber="433">MSNG</collectionCode>
); Stagno di Molentargius, 
<date id="FFFAECB1FFEE4709FDF9FD27CD07FD5D" box="[517,611,670,689]" pageId="14" pageNumber="433" value="1979-03-27">27.iii.1979</date>
, 1 ♁, C. Meloni lgt. (
<collectionCode id="ED5552B4FFEE4709FF3CFD00CF99FD20" box="[192,253,697,716]" pageId="14" pageNumber="433">CPVV</collectionCode>
), 
<date id="FFFAECB1FFEE4709FEF2FD00CE00FD20" box="[270,356,697,716]" pageId="14" pageNumber="433" value="1988-05-29">29.v.1988</date>
, 1 ♁, 
<specimenCount id="9D4201F8FFEE4709FE61FD00CEDBFD20" box="[413,447,693,716]" pageId="14" pageNumber="433" type="female">1 ♀</specimenCount>
, C. Meloni lgt. (
<collectionCode id="ED5552B4FFEE4709FDA9FD00CDF3FD20" box="[597,663,697,716]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="14" pageNumber="433">MSNG</collectionCode>
); Serdiana, 
<date id="FFFAECB1FFEE4709FF6BFD6ACF8EFD0A" box="[151,234,723,742]" pageId="14" pageNumber="433" value="2003-06-08">8.vi.2003</date>
, 6 ♁♁, 
<specimenCount id="9D4201F8FFEE4709FED6FD6ACE33FD0B" box="[298,343,720,743]" pageId="14" pageNumber="433" type="female">6 ♀♀</specimenCount>
, Fancello lgt. (
<collectionCode id="ED5552B4FFEE4709FE2AFD6ACD7CFD0A" box="[470,536,723,742]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="14" pageNumber="433">MSNG</collectionCode>
); Molentargius, 
<date id="FFFAECB1FFEE4709FD5EFD6ACD90FD0A" box="[674,756,723,742]" pageId="14" pageNumber="433" value="1979-01-31">31.i.1979</date>
, 1 ♁, C. Meloni lgt. (
<collectionCode id="ED5552B4FFEE4709FEC3FD57CEE5FCED" box="[319,385,750,769]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="14" pageNumber="433">MSNG</collectionCode>
); Cagliari, Campo Santa Gilla, 
<date id="FFFAECB1FFEE4709FD7BFD57CD82FCED" box="[647,742,750,769]" pageId="14" pageNumber="433" value="1983-03-28">28.iii.1983</date>
, 
<specimenCount id="9D4201F8FFEE4709FD11FD57CFD1FCF0" pageId="14" pageNumber="433" type="female">2 ♀♀</specimenCount>
, C. Meloni lgt.(
<collectionCode id="ED5552B4FFEE4709FEC2FCB0CEE4FCF0" box="[318,384,777,796]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="14" pageNumber="433">MSNG</collectionCode>
). 
<emphasis id="B9301663FFEE4709FE71FCB0CEAEFCF0" bold="true" box="[397,458,777,796]" pageId="14" pageNumber="433">
<collectingRegion id="49800493FFEE4709FE71FCB0CEA7FCF6" box="[397,451,777,796]" country="Italy" name="Sicilia" pageId="14" pageNumber="433">SICILY</collectingRegion>
:
</emphasis>
<collectingRegion id="49800493FFEE4709FE32FCB0CD64FCF0" box="[462,512,777,796]" country="Italy" name="Sicilia" pageId="14" pageNumber="433">Sicily</collectingRegion>
, no further data, 1♁., 
<specimenCount id="9D4201F8FFEE4709FD44FCB0CD91FCF0" box="[696,757,777,796]" pageId="14" pageNumber="433" type="generic">1 spec.</specimenCount>
, Krtz. (
<collectionCode id="ED5552B4FFEE4709FF31FC9ACE77FCDA" box="[205,275,803,822]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="433">MNHN</collectionCode>
); 
<collectingRegion id="49800493FFEE4709FEDDFC9ACE3EFCDA" box="[289,346,803,822]" country="Italy" name="Sicilia" pageId="14" pageNumber="433">Sicilia</collectingRegion>
, no further data, 
<specimenCount id="9D4201F8FFEE4709FE14FC9ACD62FCDB" box="[488,518,800,823]" pageId="14" pageNumber="433" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED5552B4FFEE4709FDEDFC9ACD36FCDA" box="[529,594,803,822]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="14" pageNumber="433">MFNB</collectionCode>
). 
<emphasis id="B9301663FFEE4709FD9CFC9ACD9CFCDA" bold="true" box="[608,760,803,822]" pageId="14" pageNumber="433">
<collectingCountry id="F3538AE1FFEE4709FD9CFC9ACDC1FCDA" box="[608,677,803,822]" name="Libya" pageId="14" pageNumber="433">LIBYA</collectingCountry>
: 
<collectingRegion id="49800493FFEE4709FD51FC9ACD94FCD8" box="[685,752,803,822]" country="Libya" name="Tarabulus" pageId="14" pageNumber="433">TRIPOLI</collectingRegion>
:
</emphasis>
Tripolis, no further data, 
<specimenCount id="9D4201F8FFEE4709FE92FC87CEEAFCBD" box="[366,398,826,849]" pageId="14" pageNumber="433" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED5552B4FFEE4709FE65FC87CEBEFCBD" box="[409,474,830,849]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="14" pageNumber="433">MFNB</collectionCode>
). 
<emphasis id="B9301663FFEE4709FE15FC87CDF3FCBD" bold="true" box="[489,663,830,849]" pageId="14" pageNumber="433">
<collectingCountry id="F3538AE1FFEE4709FE15FC87CD4FFCBD" box="[489,555,830,849]" name="Spain" pageId="14" pageNumber="433">SPAIN</collectingCountry>
: 
<collectingRegion id="49800493FFEE4709FDCFFC87CDF4FCA3" box="[563,656,830,849]" country="Spain" name="Andalucia" pageId="14" pageNumber="433">ANDALUSIA</collectingRegion>
:
</emphasis>
<collectingRegion id="49800493FFEE4709FD60FC87CD91FCBD" box="[668,757,830,849]" country="Spain" name="Andalucia" pageId="14" pageNumber="433">Andalusia</collectingRegion>
, no further data, 
<specimenCount id="9D4201F8FFEE4709FEE7FCE0CE5CFC80" box="[283,312,853,876]" pageId="14" pageNumber="433" type="female">1♀</specimenCount>
(
<collectionCode id="ED5552B4FFEE4709FEBFFCE0CEE0FC80" box="[323,388,857,876]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="14" pageNumber="433">MFNB</collectionCode>
). 
<emphasis id="B9301663FFEE4709FE6DFCE0CD49FC80" bold="true" box="[401,557,857,876]" pageId="14" pageNumber="433">
<collectingCountry id="F3538AE1FFEE4709FE6DFCE0CE89FC80" box="[401,493,857,876]" name="Tunisia" pageId="14" pageNumber="433">TUNISIA</collectingCountry>
: 
<collectingCountry id="F3538AE1FFEE4709FE09FCE0CD43FC86" box="[501,551,857,876]" name="Tunisia" pageId="14" pageNumber="433">TUNIS</collectingCountry>
:
</emphasis>
<collectingRegion id="49800493FFEE4709FDCDFCE0CD07FC80" box="[561,611,857,876]" country="Tunisia" name="Tunis" pageId="14" pageNumber="433">Tunis</collectingRegion>
, 
<specimenCount id="9D4201F8FFEE4709FD96FCE0CDC3FC80" box="[618,679,857,876]" pageId="14" pageNumber="433" type="generic">1 spec.</specimenCount>
, collector unknown, Reitter coll. (
<collectionCode id="ED5552B4FFEE4709FE98FCCACEF0FC6A" box="[356,404,883,902]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="14" pageNumber="433" type="Museum">ZSM</collectionCode>
); Tint, 
<date id="FFFAECB1FFEE4709FE2EFCCACD42FC6A" box="[466,550,883,902]" pageId="14" pageNumber="433" value="1882-01" valueMax="1882-02-31" valueMin="1882-01-00">i.–ii.1882</date>
, 1 ♁, G. &amp; L. Doria lgt. (
<collectionCode id="ED5552B4FFEE4709FF61FC37CFA1FC4D" box="[157,197,910,929]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="14" pageNumber="433" type="Museum">ZIN</collectionCode>
); Carthage, 
<date id="FFFAECB1FFEE4709FED5FC37CE12FC4D" box="[297,374,910,929]" pageId="14" pageNumber="433" value="1914-07">vii. 1914</date>
, 1♁, Novak lgt. (
<collectionCode id="ED5552B4FFEE4709FDF5FC37CD55FC4D" box="[521,561,910,929]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="14" pageNumber="433" type="Museum">ZIN</collectionCode>
); 
<collectingRegion id="49800493FFEE4709FDC2FC37CD14FC4D" box="[574,624,910,929]" country="Tunisia" name="Tunis" pageId="14" pageNumber="433">Tunis</collectingRegion>
, no further data, iv.[18]83, 1 ♁ (
<collectionCode id="ED5552B4FFEE4709FEE1FC10CE3AFC50" box="[285,350,937,956]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="14" pageNumber="433">MFNB</collectionCode>
); 
<collectingRegion id="49800493FFEE4709FE92FC10CEC4FC50" box="[366,416,937,956]" country="Tunisia" name="Tunis" pageId="14" pageNumber="433">Tunis</collectingRegion>
, no further data, 6 ♁♁, 
<specimenCount id="9D4201F8FFEE4709FD8CFC10CDC6FC50" box="[624,674,933,956]" pageId="14" pageNumber="433" type="female">3 ♀♀</specimenCount>
(
<collectionCode id="ED5552B4FFEE4709FD51FC10CD8AFC50" box="[685,750,937,956]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="14" pageNumber="433">MFNB</collectionCode>
); 
<collectingRegion id="49800493FFEE4709FF6BFC7ACFADFC3A" box="[151,201,963,982]" country="Tunisia" name="Tunis" pageId="14" pageNumber="433">Tunis</collectingRegion>
, 
<date id="FFFAECB1FFEE4709FF2CFC7ACE54FC3A" box="[208,304,963,982]" pageId="14" pageNumber="433" value="1882-02" valueMax="1882-03-31" valueMin="1882-02-00">ii.–iii.1882</date>
, 1♁, G. &amp; L. Doria lgt. (
<collectionCode id="ED5552B4FFEE4709FE03FC7ACD24FC3A" box="[511,576,963,982]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="14" pageNumber="433">MFNB</collectionCode>
); Radès, 
<date id="FFFAECB1FFEE4709FD77FC7ACDA9FC3A" box="[651,717,963,982]" pageId="14" pageNumber="433" value="1933-04">iv.1933</date>
, 1♁, M. Grossclaude (
<collectionCode id="ED5552B4FFEE4709FED1FC67CE16FC1D" box="[301,370,990,1009]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="433">MNHN</collectionCode>
; coll. Thérond). 
<emphasis id="B9301663FFEE4709FDF8FC67CD23FC1D" bold="true" box="[516,583,990,1009]" pageId="14" pageNumber="433">
<collectingRegion id="49800493FFEE4709FDF8FC67CD24FC03" box="[516,576,990,1009]" country="Tunisia" name="Sousse" pageId="14" pageNumber="433">SOUSSE</collectingRegion>
:
</emphasis>
Sebkha Kelbia lake near 
<collectingRegion id="49800493FFEE4709FF3DFC40CF9AFBE0" box="[193,254,1017,1036]" country="Tunisia" name="Sousse" pageId="14" pageNumber="433">Sousse</collectingRegion>
, 
<date id="FFFAECB1FFEE4709FEFBFC40CE3CFBE0" box="[263,344,1017,1036]" pageId="14" pageNumber="433" value="1962-04-08">8.iv.1962</date>
, 1 ♁, Cl. Besuchet lgt. (
<collectionCode id="ED5552B4FFEE4709FDD9FC40CD0CFBE0" box="[549,616,1017,1036]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="14" pageNumber="433">MSNG</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C35E99FAFFEE4709FF6BFB9DCBE7F859" pageId="14" pageNumber="433" type="description">
<paragraph id="8BFBCA71FFEE4709FF6BFB9DCEF2FB37" blockId="14.[151,761,1060,2075]" pageId="14" pageNumber="433">
<emphasis id="B9301663FFEE4709FF6BFB9DCE5CFBD7" bold="true" box="[151,312,1060,1083]" pageId="14" pageNumber="433">Redescription.</emphasis>
PEL: 1.60–2.00 mm; APW: 0.75–1.00 mm; PPW: 1.40–1.60 mm; EW: 1.50–1.75 mm; EL: 1.00–1.40 mm. Body (
<figureCitation id="137FD6F4FFEE4709FEE5FBDDCE0DFB97" box="[281,361,1124,1147]" captionStart="Figs 40–43" captionStartId="15.[151,188,2006,2025]" captionTargetBox="[212,1382,479,1970]" captionTargetId="figure-17@15.[201,1387,467,1981]" captionTargetPageId="15" captionText="Figs 40–43. Hypocaccus (Nessus) curtus (Rosenhauer, 1847), dorsal habitus.40 – Saprinus curtus Rosenhauer, 1847, lectotype.41 – Saprinus puncticollis Küster, 1849, lectotype. 42 – Saprinus cribellaticollis Jacquelin Du Val, 1858, lectotype. 43 – Saprinus sicanus Marseul, 1862, lectotype." figureDoi="http://doi.org/10.5281/zenodo.4504944" httpUri="https://zenodo.org/record/4504944/files/figure.png" pageId="14" pageNumber="433">Fig. 40</figureCitation>
) oblong, oval, rather convex, cuticle dark-brown to black with faint to pronounced greenish hue; legs and antennal funicle light reddish-brown; antennal scape somewhat darker.
</paragraph>
<paragraph id="8BFBCA71FFEE4709FF49FB5DCE88F997" blockId="14.[151,761,1060,2075]" pageId="14" pageNumber="433">
<emphasis id="B9301663FFEE4709FF49FB5DCF9DFB17" box="[181,249,1252,1275]" italics="true" pageId="14" pageNumber="433">Head:</emphasis>
mandibles densely punctate dorsally; clypeus densely and coarsely punctate, almost rugose-lacunose, anterior margin slightly elevated; frontal disc with similar, if somewhat weaker punctation; occasionally this punctation is confluent and forms tiny rugae; frontal stria slightly outwardly arcuate, complete to reduced to interrupted medially, supraorbital stria well developed; eyes flattened, but visible from above. Basal third of frontal disc with irregular rounded glabrous area; occipital stria weak, but visible. Antennal scape somewhat darker than reddish antennal funicle, antennae similar to other species of the subgenus, sensory structures of the antennal club studied by 
<bibRefCitation id="EFD5B780FFEE4709FF44F9DDCE8CF997" author="DE MARZO L. &amp; VIENNA P." box="[184,488,1636,1659]" pageId="14" pageNumber="433" pagination="79 - 89" refId="ref20541" refString="DE MARZO L. &amp; VIENNA P. 1982: Osservazioni morfologiche e ultrastrutturali su particolari organi di senso delle clave antennali in Isteridi della subf. Saprininae e considerazioni sistematiche. Entomologica 17: 79 - 89." type="journal article" year="1982">
DE MARZO &amp; 
<collectingRegion id="49800493FFEE4709FEB6F9DDCEF3F995" box="[330,407,1636,1659]" country="Austria" name="Wien" pageId="14" pageNumber="433">VIENNA</collectingRegion>
(1982)
</bibRefCitation>
.
</paragraph>
<paragraph id="8BFBCA71FFEE4709FF49F93DCEE6F8B7" blockId="14.[151,761,1060,2075]" pageId="14" pageNumber="433">
<emphasis id="B9301663FFEE4709FF49F93DCE47F977" box="[181,291,1668,1691]" italics="true" pageId="14" pageNumber="433">Pronotum</emphasis>
convex, lateral sides slightly narrowing anteriorly; anterior pronotal angles obtuse, marginal pronotal stria complete, its lateral portion observable in some cases from lateral view only. Entire pronotal disc covered with punctures separated by one to several times their diameter, punctation weakens medially. Scutellum very small, triangular.
</paragraph>
<paragraph id="8BFBCA71FFEE4709FF49F8DDCBA9FD39" blockId="14.[151,761,1060,2075]" lastBlockId="14.[828,1437,190,2069]" pageId="14" pageNumber="433">
<emphasis id="B9301663FFEE4709FF49F8DDCF9BF897" box="[181,255,1892,1915]" italics="true" pageId="14" pageNumber="433">Elytra:</emphasis>
elytral epipleuron impunctate, marginal epipleural stria complete, marginal elytral stria well developed, complete, continued as apical elytral stria for short distance. Humeral elytral stria well developed, present on basal elytral third; internal subhumeral stria present as a median fragment. Dorsal elytral striae 1–4 well developed, first the longest, slightly bisinuate, reaching approximately two-thirds of elytral length apically, occasionally even slightly longer, striae 2–4 shorter, reaching approximately elytral mid-length apically, while second stria may be longer than striae 3–4; fourth stria usually the shortest, formed in most cases of beads of punctures, stopping short of elytral mid-length apically. Fourth dorsal elytral stria usually not connected (connected in specimens that belong to form ‘ 
<taxonomicName id="4C44B1F2FFEE4709FB66FE27CA55FE59" authorityName="Jacquelin du Val" authorityYear="1858" box="[1178,1329,414,437]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="433" phylum="Arthropoda" rank="species" species="cribellaticollis">
<emphasis id="B9301663FFEE4709FB66FE27CA55FE59" box="[1178,1329,414,437]" italics="true" pageId="14" pageNumber="433">cribellaticollis</emphasis>
</taxonomicName>
’) with the basal end of (in)complete sutural elytral stria, which is in punctures and can be basally shortened. Elytral punctation covers approximately apical half of elytral disc, slightly surpassing elytral mid-length basally, slightly and scatteredly entering elytral intervals in some specimens; punctation rather dense, punctures separated by approximately their own diameter. Basal elytral fifth, fourth elytral interval, elytral flanks and extreme elytral apex impunctate, or with scattered microscopic punctation only.
</paragraph>
<paragraph id="8BFBCA71FFEE4709FCA6FD67CAF9FB99" blockId="14.[828,1437,190,2069]" pageId="14" pageNumber="433">
<emphasis id="B9301663FFEE4709FCA6FD67CBF0FD19" box="[858,1172,734,757]" italics="true" pageId="14" pageNumber="433">Propygidium and pygidium</emphasis>
: propygidium covered with punctation similar to that of elytra; pygidium with much finer and sparser punctation. 
<taxonomicName id="4C44B1F2FFEE4709FB56FCA7CA4DFCD9" box="[1194,1321,798,821]" class="Insecta" family="Elateridae" genus="Prosternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="423" phylum="Arthropoda" rank="genus">Prosternum</taxonomicName>
: prosternal process slightly to moderately concave (observed from lateral view); carinal prosternal striae carinate, divergent on prosternal apophysis, running convergent to sub-parallel to almost approximate apically; from mid-length of prosternal process slightly divergent anteriorly, apically united under tiny loop; interspaces between carinal prosternal striae with scattered punctures. Lateral prosternal stria strongly carinate, convergent apically, united in front of united carinal prosternal striae; lateral sides of prosternal process densely punctate; prosternal foveae moderately large, deep.
</paragraph>
<paragraph id="8BFBCA71FFEE4709FCA6FBC7CBC8FAD9" blockId="14.[828,1437,190,2069]" pageId="14" pageNumber="433">
<emphasis id="B9301663FFEE4709FCA6FBC7CC80FB79" box="[858,996,1150,1173]" italics="true" pageId="14" pageNumber="433">Mesoventrite</emphasis>
: disc of mesoventrite approximately three times as wide as long, with scattered punctures (occasionally almost glabrous); marginal mesoventral stria complete, slightly inwardly arcuate medially; meso-metaventral stria undulate, bisinuate, in punctures, slightly distanced from meso-metaventral suture medially.
</paragraph>
<paragraph id="8BFBCA71FFEE4709FCA6FA87CA24F999" blockId="14.[828,1437,190,2069]" pageId="14" pageNumber="433">
<emphasis id="B9301663FFEE4709FCA6FA87CC8FFAB9" box="[858,1003,1342,1365]" italics="true" pageId="14" pageNumber="433">Metaventrite:</emphasis>
disc of metaventrite apart from several rows of tiny punctures situated along basal margin entirely glabrous; lateral metaventral stria almost straight, slightly bisinuate, deeply impressed, in punctures, stopping short of metacoxa; lateral disc of metaventrite depressed, with large oval deep punctures separated by less than their diameter; metepisternum with similar punctation, punctures of smaller sizes than those of lateral disc of metaventrite. First visible abdominal ventrite striate laterally, with scattered fine punctation, occasionally almost impunctate.
</paragraph>
<paragraph id="8BFBCA71FFEE4709FCA6F9C7CB64F919" blockId="14.[828,1437,190,2069]" pageId="14" pageNumber="433">
<emphasis id="B9301663FFEE4709FCA6F9C7CCF1F979" box="[858,917,1662,1685]" italics="true" pageId="14" pageNumber="433">Legs:</emphasis>
protibia (
<figureCitation id="137FD6F4FFEE4709FC05F9C7CB20F979" box="[1017,1092,1662,1685]" captionStart="Figs 31–32" captionStartId="11.[151,188,1619,1638]" captionTargetBox="[172,742,1091,1585]" captionTargetId="figure-518@11.[151,760,1066,1603]" captionTargetPageId="11" captionText="Figs 31–32. Protibiae, dorsal view. 31 – Hypocaccus (Nessus) rubripes (Erichson, 1834); 32 – H. (N.) curtus (Rosenhauer, 1847) (both re-drawn from VIENNA 1980)." figureDoi="http://doi.org/10.5281/zenodo.4504938" httpUri="https://zenodo.org/record/4504938/files/figure.png" pageId="14" pageNumber="433">Fig. 32</figureCitation>
) on outer margin with 8–11 short to moderately long denticles diminishing in size proximally, protibial groove deep; rest of leg characters similar to preceding species.
</paragraph>
<paragraph id="8BFBCA71FFEE4709FCA6F947CBE7F859" blockId="14.[828,1437,190,2069]" pageId="14" pageNumber="433">
<emphasis id="B9301663FFEE4709FCA6F947CB66F8F9" box="[858,1026,1790,1813]" italics="true" pageId="14" pageNumber="433">Male genitalia:</emphasis>
sternite VIII (
<figureCitation id="137FD6F4FFEE4709FB5DF947CA7AF8F9" box="[1185,1310,1790,1813]" captionStart="Figs 46–54" captionStartId="17.[151,188,1365,1384]" captionTargetBox="[271,1310,177,1331]" captionTargetId="figure-374@17.[252,1324,170,1345]" captionTargetPageId="17" captionText="Figs 46–54. Male genitalia of Hypocaccus (Nessus) curtus (Rosenhauer, 1847). 45 – VIII sternite and tergite, ventral view; 46 – ditto, dorsal view; 47 – ditto, lateral view; 48 – IX and X tergites, dorsal view; 49 – ditto, lateral view; 50 – IX sternite (spiculum gastrale), ventral view; 51 – ditto, lateral view; 52 – aedeagus, dorsal view; 53 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4504952" httpUri="https://zenodo.org/record/4504952/files/figure.png" pageId="14" pageNumber="433">Figs 46–47</figureCitation>
) narrowing apically; sternite VIII and tergite VIII fused laterally (
<figureCitation id="137FD6F4FFEE4709FA8DF8A7CC3EF8B9" captionStart="Figs 46–54" captionStartId="17.[151,188,1365,1384]" captionTargetBox="[271,1310,177,1331]" captionTargetId="figure-374@17.[252,1324,170,1345]" captionTargetPageId="17" captionText="Figs 46–54. Male genitalia of Hypocaccus (Nessus) curtus (Rosenhauer, 1847). 45 – VIII sternite and tergite, ventral view; 46 – ditto, dorsal view; 47 – ditto, lateral view; 48 – IX and X tergites, dorsal view; 49 – ditto, lateral view; 50 – IX sternite (spiculum gastrale), ventral view; 51 – ditto, lateral view; 52 – aedeagus, dorsal view; 53 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4504952" httpUri="https://zenodo.org/record/4504952/files/figure.png" pageId="14" pageNumber="433">Fig. 48</figureCitation>
). Tergite IX medio-laterally with tiny acute projection (
<figureCitation id="137FD6F4FFEE4709FCBFF8E7CCA6F899" box="[835,962,1886,1909]" captionStart="Figs 46–54" captionStartId="17.[151,188,1365,1384]" captionTargetBox="[271,1310,177,1331]" captionTargetId="figure-374@17.[252,1324,170,1345]" captionTargetPageId="17" captionText="Figs 46–54. Male genitalia of Hypocaccus (Nessus) curtus (Rosenhauer, 1847). 45 – VIII sternite and tergite, ventral view; 46 – ditto, dorsal view; 47 – ditto, lateral view; 48 – IX and X tergites, dorsal view; 49 – ditto, lateral view; 50 – IX sternite (spiculum gastrale), ventral view; 51 – ditto, lateral view; 52 – aedeagus, dorsal view; 53 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4504952" httpUri="https://zenodo.org/record/4504952/files/figure.png" pageId="14" pageNumber="433">Figs 49–50</figureCitation>
). Spiculum gastrale (
<figureCitation id="137FD6F4FFEE4709FB54F8E7CA41F899" box="[1192,1317,1886,1909]" captionStart="Figs 46–54" captionStartId="17.[151,188,1365,1384]" captionTargetBox="[271,1310,177,1331]" captionTargetId="figure-374@17.[252,1324,170,1345]" captionTargetPageId="17" captionText="Figs 46–54. Male genitalia of Hypocaccus (Nessus) curtus (Rosenhauer, 1847). 45 – VIII sternite and tergite, ventral view; 46 – ditto, dorsal view; 47 – ditto, lateral view; 48 – IX and X tergites, dorsal view; 49 – ditto, lateral view; 50 – IX sternite (spiculum gastrale), ventral view; 51 – ditto, lateral view; 52 – aedeagus, dorsal view; 53 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4504952" httpUri="https://zenodo.org/record/4504952/files/figure.png" pageId="14" pageNumber="433">Figs 51–52</figureCitation>
) similar to other congeners. Aedeagus (
<figureCitation id="137FD6F4FFEE4709FB88F8C7CB95F879" box="[1140,1265,1918,1941]" captionStart="Figs 46–54" captionStartId="17.[151,188,1365,1384]" captionTargetBox="[271,1310,177,1331]" captionTargetId="figure-374@17.[252,1324,170,1345]" captionTargetPageId="17" captionText="Figs 46–54. Male genitalia of Hypocaccus (Nessus) curtus (Rosenhauer, 1847). 45 – VIII sternite and tergite, ventral view; 46 – ditto, dorsal view; 47 – ditto, lateral view; 48 – IX and X tergites, dorsal view; 49 – ditto, lateral view; 50 – IX sternite (spiculum gastrale), ventral view; 51 – ditto, lateral view; 52 – aedeagus, dorsal view; 53 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4504952" httpUri="https://zenodo.org/record/4504952/files/figure.png" pageId="14" pageNumber="433">Figs 53–54</figureCitation>
) almost subparallel, bluntly pointed apically.
</paragraph>
</subSubSection>
<subSubSection id="C35E99FAFFEE4709FCC0F807CB48F7F9" pageId="14" pageNumber="433" type="distribution">
<paragraph id="8BFBCA71FFEE4709FCC0F807CB48F7F9" blockId="14.[828,1437,190,2069]" pageId="14" pageNumber="433">
<emphasis id="B9301663FFEE4709FCC0F807CCAAF839" bold="true" box="[828,974,1982,2005]" pageId="14" pageNumber="433">Distribution.</emphasis>
<collectingCountry id="F3538AE1FFEE4709FC29F807CB5DF839" box="[981,1081,1982,2005]" name="Hungary" pageId="14" pageNumber="433">Hungary</collectingCountry>
(?), 
<collectingCountry id="F3538AE1FFEE4709FBA1F807CBCCF839" box="[1117,1192,1982,2005]" name="France" pageId="14" pageNumber="433">France</collectingCountry>
, 
<collectingCountry id="F3538AE1FFEE4709FB4FF807CB8CF839" box="[1203,1256,1982,2005]" name="Italy" pageId="14" pageNumber="433">Italy</collectingCountry>
: 
<collectingRegion id="49800493FFEE4709FB0FF807CA2BF839" box="[1267,1359,1982,2005]" country="Italy" name="Sardegna" pageId="14" pageNumber="433">Sardinia</collectingRegion>
, 
<collectingRegion id="49800493FFEE4709FAA5F807CAFCF839" box="[1369,1432,1982,2005]" country="Italy" name="Sicilia" pageId="14" pageNumber="433">Sicily</collectingRegion>
, 
<collectingCountry id="F3538AE1FFEE4709FCC0F867CC1FF819" box="[828,891,2014,2037]" name="Spain" pageId="14" pageNumber="433">Spain</collectingCountry>
, 
<collectingCountry id="F3538AE1FFEE4709FC7AF867CC87F819" box="[902,995,2014,2037]" name="Portugal" pageId="14" pageNumber="433">Portugal</collectingCountry>
, 
<collectingCountry id="F3538AE1FFEE4709FC13F867CB59F819" box="[1007,1085,2014,2037]" name="Greece" pageId="14" pageNumber="433">Greece</collectingCountry>
, 
<collectingCountry id="F3538AE1FFEE4709FBB4F867CBEDF819" box="[1096,1161,2014,2037]" name="Malta" pageId="14" pageNumber="433">Malta</collectingCountry>
, 
<collectingCountry id="F3538AE1FFEE4709FB68F867CB80F819" box="[1172,1252,2014,2037]" name="Cyprus" pageId="14" pageNumber="433">Cyprus</collectingCountry>
, 
<collectingCountry id="F3538AE1FFEE4709FB13F867CA58F819" box="[1263,1340,2014,2037]" name="Turkey" pageId="14" pageNumber="433">Turkey</collectingCountry>
, 
<collectingCountry id="F3538AE1FFEE4709FABAF867CAFCF819" box="[1350,1432,2014,2037]" name="Tunisia" pageId="14" pageNumber="433">Tunisia</collectingCountry>
, 
<collectingCountry id="F3538AE1FFEE4709FCC0F847CCEBF7F9" box="[828,911,2046,2069]" name="Algeria" pageId="14" pageNumber="433">Algeria</collectingCountry>
, 
<collectingCountry id="F3538AE1FFEE4709FC65F847CCBEF7F9" box="[921,986,2046,2069]" name="Libya" pageId="14" pageNumber="433">Libya</collectingCountry>
, 
<collectingCountry id="F3538AE1FFEE4709FC18F847CB43F7F9" box="[996,1063,2046,2069]" name="Egypt" pageId="14" pageNumber="433">Egypt</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C35E99FAFFEF4708FF6BFF07CD4CFED9" pageId="15" pageNumber="434" type="biology_ecology">
<paragraph id="8BFBCA71FFEF4708FF6BFF07CD4CFED9" blockId="15.[151,760,190,437]" pageId="15" pageNumber="434">
<emphasis id="B9301663FFEF4708FF6BFF07CF97FF39" bold="true" box="[151,243,190,213]" pageId="15" pageNumber="434">Biology.</emphasis>
According to 
<bibRefCitation id="EFD5B780FFEF4708FE6CFF07CD4BFF39" author="VIENNA P." box="[400,559,190,213]" pageId="15" pageNumber="434" refId="ref21978" refString="VIENNA P. 1980: Fauna d'Italia. Vol. XVI. Coleoptera Histeridae. Bologna, Calderini Press, ix + 386 pp." type="book" year="1980">
<collectingRegion id="49800493FFEF4708FE6CFF07CEB9FF3F" box="[400,477,190,213]" country="Austria" name="Wien" pageId="15" pageNumber="434">VIENNA</collectingRegion>
(1980)
</bibRefCitation>
, who repeats THÉROND (1975), 
<taxonomicName id="4C44B1F2FFEF4708FED8FF67CED2FF19" authorityName="Rosenhauer" authorityYear="1847" box="[292,438,222,245]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="438" phylum="Arthropoda" rank="species" species="curtus" subGenus="Nannolepidius">
<emphasis id="B9301663FFEF4708FED8FF67CE53FF19" box="[292,311,222,245]" italics="true" pageId="15" pageNumber="434">H</emphasis>
. (
<emphasis id="B9301663FFEF4708FEB0FF67CE3AFF19" box="[332,350,222,245]" italics="true" pageId="15" pageNumber="434">N</emphasis>
.) 
<emphasis id="B9301663FFEF4708FE8FFF67CED2FF19" box="[371,438,222,245]" italics="true" pageId="15" pageNumber="434">curtus</emphasis>
</taxonomicName>
is found under detritus in sand near the seacoast, where it was collected from near 
<taxonomicName id="4C44B1F2FFEF4708FD55FF47CD93FEF9" box="[681,759,254,277]" class="Magnoliopsida" family="Amaranthaceae" genus="Suaeda" kingdom="Plantae" order="Caryophyllales" pageId="15" pageNumber="434" phylum="Tracheophyta" rank="genus">
<emphasis id="B9301663FFEF4708FD55FF47CD93FEF9" box="[681,759,254,277]" italics="true" pageId="15" pageNumber="434">Suaeda</emphasis>
</taxonomicName>
sp. and 
<taxonomicName id="4C44B1F2FFEF4708FF15FEA7CEE1FED9" box="[233,389,286,309]" class="Magnoliopsida" family="Plumbaginaceae" genus="Statice" kingdom="Plantae" order="Caryophyllales" pageId="15" pageNumber="434" phylum="Tracheophyta" rank="species" species="virgata">
<emphasis id="B9301663FFEF4708FF15FEA7CEE1FED9" box="[233,389,286,309]" italics="true" pageId="15" pageNumber="434">Statice virgata</emphasis>
</taxonomicName>
W. plant roots.
</paragraph>
</subSubSection>
<subSubSection id="C35E99FAFFEF4716FF6BFE87CBD7F9FE" lastPageId="17" lastPageNumber="436" pageId="15" pageNumber="434" type="discussion">
<paragraph id="8BFBCA71FFEF4717FF6BFE87CEF6FE39" blockId="15.[151,760,190,437]" lastBlockId="16.[151,760,190,1141]" lastPageId="16" lastPageNumber="435" pageId="15" pageNumber="434">
<emphasis id="B9301663FFEF4708FF6BFE87CE6CFEB9" bold="true" box="[151,264,318,341]" pageId="15" pageNumber="434">Remarks.</emphasis>
The type specimen was part of Rosenhauer’s collection, which later became partly a part of R. Oberthür’s collection (A. Taghavian, pers. comm. 2017), currently housed in MNHN. The senior author has visited MNHN multiple times and failed to locate the type specimen(s) of this species in the collections of MNHN (including R. Oberthür’s collection). Mr. Serge Risser (Pleucadeuc, 
<collectingCountry id="F3538AE1FFEF4708FC2CFEA7CB78FED9" box="[976,1052,286,309]" name="France" pageId="15" pageNumber="434">France</collectingCountry>
) recently purchased the 
<taxonomicName id="4C44B1F2FFEF4708FAD0FEA7CAF8FED9" authorityName="Gyllenhaal" authorityYear="1808" box="[1324,1436,286,309]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="434" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
collection of the late Marcel René Paul de Vauloger de Beaupré and published its contents in two separate papers (
<bibRefCitation id="EFD5B780FFEF4708FCBFFEC7CCBFFE79" author="RISSER S." box="[835,987,382,405]" pageId="15" pageNumber="434" pagination="197 - 214" refId="ref21752" refString="RISSER S. 2013 a: Les Histeridae de la collection de Vauloger de Beaupre (premiere partie) (Coleoptera). L'Entomologiste 69: 197 - 214." type="journal article" year="2013">RISSER 2013a</bibRefCitation>
,b). When reading RISSER’ S paper (2013a) we were intrigued by a specimen identified as 
<emphasis id="B9301663FFEF4717FAE0FE27CFD2FF39" italics="true" lastPageId="16" lastPageNumber="435" pageId="15" pageNumber="434">Hypocacculus</emphasis>
(
<taxonomicName id="4C44B1F2FFF04717FF38FF07CE3BFF39" authorityName="Reichardt" authorityYear="1932" box="[196,351,190,213]" class="Insecta" family="Histeridae" genus="Nannolepidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="435" phylum="Arthropoda" rank="genus">
<emphasis id="B9301663FFF04717FF38FF07CE3BFF39" box="[196,351,190,213]" italics="true" pageId="16" pageNumber="435">Nannolepidius</emphasis>
</taxonomicName>
!) 
<taxonomicName id="4C44B1F2FFF04717FE8AFF07CEDCFF39" authorityName="Rosenhauer" authorityYear="1847" box="[374,440,190,213]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="435" phylum="Arthropoda" rank="species" species="curtus">
<emphasis id="B9301663FFF04717FE8AFF07CEDCFF39" box="[374,440,190,213]" italics="true" pageId="16" pageNumber="435">curtus</emphasis>
</taxonomicName>
originating from 
<collectingCountry id="F3538AE1FFF04717FD93FF07CDA8FF39" box="[623,716,190,213]" name="Hungary" pageId="16" pageNumber="435">Hungary</collectingCountry>
and from ‘Musaeo Rosenhauer’. Mr. Risser was kind enough to send this specimen to one of us (T. L.). Having examined it as well as compared it to Rosenhauer’s original description we concluded that this is the long-lost type specimen of Rosenhauer’s species 
<taxonomicName id="4C44B1F2FFF04717FDECFEE7CDDFFE99" authorityName="Rosenhauer" authorityYear="1847" box="[528,699,350,373]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="435" phylum="Arthropoda" rank="species" species="curtus">
<emphasis id="B9301663FFF04717FDECFEE7CDDFFE99" box="[528,699,350,373]" italics="true" pageId="16" pageNumber="435">Saprinus curtus</emphasis>
</taxonomicName>
. This species was described based on an unspecified number of specimens and therefore we designate a 
<typeStatus id="54FF74D3FFF04717FD8AFE27CDBEFE59" box="[630,730,414,437]" pageId="16" pageNumber="435" type="lectotype">lectotype</typeStatus>
to fix the species identity.
</paragraph>
<caption id="DF3B9AF9FFEF4708FF6BF86FCA48F7E8" ID-DOI="http://doi.org/10.5281/zenodo.4504944" ID-Zenodo-Dep="4504944" httpUri="https://zenodo.org/record/4504944/files/figure.png" pageId="15" pageNumber="434" startId="15.[151,188,2006,2025]" targetBox="[212,1382,479,1970]" targetPageId="15">
<paragraph id="8BFBCA71FFEF4708FF6BF86FCA48F7E8" blockId="15.[151,1436,2006,2052]" pageId="15" pageNumber="434">
Figs 40–43. 
<taxonomicName id="4C44B1F2FFEF4708FF02F86FCDF2F805" authority="(Rosenhauer, 1847)" authorityName="Lackner &amp; Seres" authorityYear="2018" baseAuthorityName="Rosenhauer" baseAuthorityYear="1847" box="[254,662,2006,2025]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="434" phylum="Arthropoda" rank="species" species="curtus" subGenus="Nessus">
<emphasis id="B9301663FFEF4708FF02F86FCE02F805" box="[254,358,2006,2025]" italics="true" pageId="15" pageNumber="434">Hypocaccus</emphasis>
(
<emphasis id="B9301663FFEF4708FE8CF86FCECFF805" box="[368,427,2006,2025]" italics="true" pageId="15" pageNumber="434">Nessus</emphasis>
) 
<emphasis id="B9301663FFEF4708FE4AF86FCE8FF805" box="[438,491,2006,2025]" italics="true" pageId="15" pageNumber="434">curtus</emphasis>
(
<bibRefCitation id="EFD5B780FFEF4708FE09F86FCDF4F805" author="ROSENHAUER W. G." box="[501,656,2006,2025]" pageId="15" pageNumber="434" refId="ref21812" refString="ROSENHAUER W. G. 1847: Beitrage zur Insekten-Fauna Europas. Erstes Bandchen; enthalt die Beschreibung von sechzig neuen Kafern aus Bayern, Tyrol, Ungarn, etc., so wie die Kafer Tyrols, nach dem Ergebnisse von vier Reisen zusammengestellt. Mit einer Tafel Abbildungen. Th. Blasing, Erlangen, x + 159 pp." type="book" year="1847">Rosenhauer, 1847</bibRefCitation>
)
</taxonomicName>
, dorsal habitus.40 – 
<taxonomicName id="4C44B1F2FFEF4708FCBFF86FCB00F805" authority="Rosenhauer, 1847" authorityName="Rosenhauer" authorityYear="1847" box="[835,1124,2006,2025]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="434" phylum="Arthropoda" rank="species" species="curtus">
<emphasis id="B9301663FFEF4708FCBFF86FCCA3F805" box="[835,967,2006,2025]" italics="true" pageId="15" pageNumber="434">Saprinus curtus</emphasis>
<bibRefCitation id="EFD5B780FFEF4708FC37F86FCB00F805" author="ROSENHAUER W. G." box="[971,1124,2006,2025]" pageId="15" pageNumber="434" refId="ref21812" refString="ROSENHAUER W. G. 1847: Beitrage zur Insekten-Fauna Europas. Erstes Bandchen; enthalt die Beschreibung von sechzig neuen Kafern aus Bayern, Tyrol, Ungarn, etc., so wie die Kafer Tyrols, nach dem Ergebnisse von vier Reisen zusammengestellt. Mit einer Tafel Abbildungen. Th. Blasing, Erlangen, x + 159 pp." type="book" year="1847">Rosenhauer, 1847</bibRefCitation>
</taxonomicName>
, lectotype.41 – 
<taxonomicName id="4C44B1F2FFEF4708FB16F86FCE63F7E8" authority="Kuster, 1849" authorityName="Kuster" authorityYear="1849" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="434" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="B9301663FFEF4708FB16F86FCAF8F805" box="[1258,1436,2006,2025]" italics="true" pageId="15" pageNumber="434">Saprinus puncticollis</emphasis>
<bibRefCitation id="EFD5B780FFEF4708FF6BF848CE63F7E8" author="KUSTER H. C." box="[151,263,2033,2052]" pageId="15" pageNumber="434" refId="ref21108" refString="KUSTER H. C. 1849: Die Kafer Europa's. Nach der Natur beschreiben. XVII Heft. Bauer and Raspe, Nurnberg, 100 pls." type="book" year="1849">Küster, 1849</bibRefCitation>
</taxonomicName>
, lectotype. 42 – 
<taxonomicName id="4C44B1F2FFEF4708FE6FF848CC4BF7E8" authority="Jacquelin Du Val, 1858" authorityName="Jacquelin du Val" authorityYear="1858" box="[403,815,2033,2052]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="434" phylum="Arthropoda" rank="species" species="cribellaticollis">
<emphasis id="B9301663FFEF4708FE6FF848CD04F7E8" box="[403,608,2033,2052]" italics="true" pageId="15" pageNumber="434">Saprinus cribellaticollis</emphasis>
<bibRefCitation id="EFD5B780FFEF4708FD9AF848CC4BF7E8" author="JACQUELIN DU VAL M." box="[614,815,2033,2052]" pageId="15" pageNumber="434" pagination="97 - 108" refId="ref20919" refString="JACQUELIN DU VAL M. 1858: Genere des Coleopteres d'Europe comprenant leur classification et famillies naturelles, la description de tous les genres, des tableaux synoptiques destines a faciliter l'etude, le catalogue de toutes les especes de nombreux dessins au trait ce caracteres. II. A. Deyrolle, Paris, pp. 1 - 168. Cat. pp. 53 - 83, 97 - 108." type="journal article" year="1858">Jacquelin Du Val, 1858</bibRefCitation>
</taxonomicName>
, lectotype. 43 – 
<taxonomicName id="4C44B1F2FFEF4708FC46F848CBABF7E8" authority="Marseul, 1862" authorityName="Marseul" authorityYear="1862" box="[954,1231,2033,2052]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="434" phylum="Arthropoda" rank="species" species="sicanus">
<emphasis id="B9301663FFEF4708FC46F848CB28F7E8" box="[954,1100,2033,2052]" italics="true" pageId="15" pageNumber="434">Saprinus sicanus</emphasis>
<bibRefCitation id="EFD5B780FFEF4708FBADF848CBABF7E8" author="MARSEUL S. A." box="[1105,1231,2033,2052]" pageId="15" pageNumber="434" pagination="677 - 758" refId="ref21276" refString="MARSEUL S. A. 1862: Supplement a la monographie des Histerides. (Suite). Annales de la Societe Entomologique de France, Quartieme Serie 3: 83 - 165, 327 - 506, 677 - 758." type="journal article" year="1862">Marseul, 1862</bibRefCitation>
</taxonomicName>
, lectotype.
</paragraph>
</caption>
<paragraph id="8BFBCA71FFF04717FF49FE67CB29FDF9" blockId="16.[151,760,190,1141]" lastBlockId="16.[828,1437,190,1141]" pageId="16" pageNumber="435">
<taxonomicName id="4C44B1F2FFF04717FF49FE67CE3FFE19" authorityName="Rosenhauer" authorityYear="1847" box="[181,347,478,501]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="435" phylum="Arthropoda" rank="species" species="curtus">
<emphasis id="B9301663FFF04717FF49FE67CE3FFE19" box="[181,347,478,501]" italics="true" pageId="16" pageNumber="435">Saprinus curtus</emphasis>
</taxonomicName>
has become a mystery practically since its description, which was, however, rather detailed and served the purpose well. The reason for this was probably the fact that the 
<typeStatus id="54FF74D3FFF04717FEB3FD87CE1AFDB9" box="[335,382,574,597]" pageId="16" pageNumber="435">type</typeStatus>
specimen(s) were unavailable for comparison and perhaps also because no more specimens matching this species were ever reported from ‘Hungary’. Based on the description alone, 
<bibRefCitation id="EFD5B780FFF04717FDCDFD27CD9CFD59" author="BICKHARDT H." box="[561,760,670,693]" pageId="16" pageNumber="435" refId="ref20506" refString="BICKHARDT H. 1916 - 1917: Histeridae. In: WYTSMAN P. (ed.): Genera Insectorum. Fasc. 166 a, b. La Haye, 302 pp." type="book" year="1916">BICKHARDT (1916)</bibRefCitation>
correctly synonymized the 
<emphasis id="B9301663FFF04717FE3FFD07CEB2FD39" box="[451,470,702,725]" italics="true" pageId="16" pageNumber="435">H</emphasis>
. 
<taxonomicName id="4C44B1F2FFF04717FE19FD07CFB3FD19" authority="(Kuster, 1849)" baseAuthorityName="Kuster" baseAuthorityYear="1849" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="435" phylum="Arthropoda" rank="species" species="puncticollis" subGenus="Nessus">
(
<emphasis id="B9301663FFF04717FE12FD07CD64FD39" box="[494,512,702,725]" italics="true" pageId="16" pageNumber="435">N</emphasis>
.) 
<emphasis id="B9301663FFF04717FDE5FD07CDFCFD39" box="[537,664,702,725]" italics="true" pageId="16" pageNumber="435">puncticollis</emphasis>
(
<bibRefCitation id="EFD5B780FFF04717FD54FD07CFABFD19" author="KUSTER H. C." pageId="16" pageNumber="435" refId="ref21108" refString="KUSTER H. C. 1849: Die Kafer Europa's. Nach der Natur beschreiben. XVII Heft. Bauer and Raspe, Nurnberg, 100 pls." type="book" year="1849">Küster, 1849</bibRefCitation>
)
</taxonomicName>
with 
<taxonomicName id="4C44B1F2FFF04717FEE4FD67CED6FD19" authorityName="Rosenhauer" authorityYear="1847" box="[280,434,734,757]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="438" phylum="Arthropoda" rank="species" species="curtus" subGenus="Nannolepidius">
<emphasis id="B9301663FFF04717FEE4FD67CE4FFD19" box="[280,299,734,757]" italics="true" pageId="16" pageNumber="435">H</emphasis>
. (
<emphasis id="B9301663FFF04717FEB8FD67CE32FD19" box="[324,342,734,757]" italics="true" pageId="16" pageNumber="435">N</emphasis>
.) 
<emphasis id="B9301663FFF04717FE92FD67CED6FD19" box="[366,434,734,757]" italics="true" pageId="16" pageNumber="435">curtus</emphasis>
</taxonomicName>
, which was also followed by 
<bibRefCitation id="EFD5B780FFF04717FF6BFD47CE3EFCF9" author="REICHARDT A." box="[151,346,766,789]" pageId="16" pageNumber="435" pagination="1 - 164" refId="ref21636" refString="REICHARDT A. 1932: Beitrage zur einer Monographie der Saprininae (Coleoptera, Histeridae). Mitteilungen aus dem Zoologischen Museum in Berlin 18: 1 - 164." type="journal article" year="1932">REICHARDT (1932)</bibRefCitation>
. 
<bibRefCitation id="EFD5B780FFF04717FE9FFD47CD6CFCF9" author="MULLER G." box="[355,520,766,789]" pageId="16" pageNumber="435" pagination="97 - 112" refId="ref21522" refString="MULLER G. 1937: Histeriden-Studien. Entomologische Blatter 33: 97 - 112." type="journal article" year="1937">MÜLLER (1937)</bibRefCitation>
, however, doubted the two species are synonymous since the apical elytral stria in 
<taxonomicName id="4C44B1F2FFF04717FF4EFC87CE22FCB9" authorityName="Rosenhauer" authorityYear="1847" box="[178,326,830,853]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="438" phylum="Arthropoda" rank="species" species="curtus" subGenus="Nannolepidius">
<emphasis id="B9301663FFF04717FF4EFC87CFA1FCB9" box="[178,197,830,853]" italics="true" pageId="16" pageNumber="435">H</emphasis>
. (
<emphasis id="B9301663FFF04717FF27FC87CF89FCB9" box="[219,237,830,853]" italics="true" pageId="16" pageNumber="435">N</emphasis>
.) 
<emphasis id="B9301663FFF04717FEFEFC87CE22FCB9" box="[258,326,830,853]" italics="true" pageId="16" pageNumber="435">curtus</emphasis>
</taxonomicName>
reaches only mid-length of elytral apex, while, according to 
<bibRefCitation id="EFD5B780FFF04717FE88FCE7CD7EFC99" author="MULLER G." box="[372,538,862,885]" pageId="16" pageNumber="435" pagination="97 - 112" refId="ref21522" refString="MULLER G. 1937: Histeriden-Studien. Entomologische Blatter 33: 97 - 112." type="journal article" year="1937">MÜLLER (1937)</bibRefCitation>
it is complete in 
<emphasis id="B9301663FFF04717FD21FCE7CD94FC99" box="[733,752,862,885]" italics="true" pageId="16" pageNumber="435">H</emphasis>
. 
<taxonomicName id="4C44B1F2FFF04717FF6BFCC7CE21FC79" baseAuthorityName="Kuster" baseAuthorityYear="1849" box="[151,325,894,917]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="435" phylum="Arthropoda" rank="species" species="puncticollis" subGenus="Nessus">
(
<emphasis id="B9301663FFF04717FF5CFCC7CFD6FC79" box="[160,178,894,917]" italics="true" pageId="16" pageNumber="435">N</emphasis>
.) 
<emphasis id="B9301663FFF04717FF3BFCC7CE21FC79" box="[199,325,894,917]" italics="true" pageId="16" pageNumber="435">puncticollis</emphasis>
</taxonomicName>
. Furthermore, 
<bibRefCitation id="EFD5B780FFF04717FE18FCC7CDE1FC79" author="MULLER G." box="[484,645,894,917]" pageId="16" pageNumber="435" pagination="97 - 112" refId="ref21522" refString="MULLER G. 1937: Histeriden-Studien. Entomologische Blatter 33: 97 - 112." type="journal article" year="1937">MÜLLER (1937)</bibRefCitation>
advocated using Küster’s 
<emphasis id="B9301663FFF04717FEC4FC27CE2FFC59" box="[312,331,926,949]" italics="true" pageId="16" pageNumber="435">H</emphasis>
. 
<taxonomicName id="4C44B1F2FFF04717FEA4FC27CD63FC59" baseAuthorityName="Kuster" baseAuthorityYear="1849" box="[344,519,926,949]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="435" phylum="Arthropoda" rank="species" species="puncticollis" subGenus="Nessus">
(
<emphasis id="B9301663FFF04717FE9DFC27CE17FC59" box="[353,371,926,949]" italics="true" pageId="16" pageNumber="435">N</emphasis>
.) 
<emphasis id="B9301663FFF04717FE74FC27CD63FC59" box="[392,519,926,949]" italics="true" pageId="16" pageNumber="435">puncticollis</emphasis>
</taxonomicName>
as the valid (albeit not the earliest) name for this species and suggested, perhaps because of the incomplete description or the absence of the 
<typeStatus id="54FF74D3FFF04717FF41FC47CF8FFBF9" box="[189,235,1022,1045]" pageId="16" pageNumber="435">type</typeStatus>
material, that 
<taxonomicName id="4C44B1F2FFF04717FE7FFC47CD71FBF9" authorityName="Rosenhauer" authorityYear="1847" box="[387,533,1022,1045]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="435" phylum="Arthropoda" rank="species" species="curtus" subGenus="Nannolepidius">
<emphasis id="B9301663FFF04717FE7FFC47CEF2FBF9" box="[387,406,1022,1045]" italics="true" pageId="16" pageNumber="435">H</emphasis>
. (
<emphasis id="B9301663FFF04717FE57FC47CEA0FBF9" box="[427,452,1022,1045]" italics="true" pageId="16" pageNumber="435">N.</emphasis>
) 
<emphasis id="B9301663FFF04717FE2EFC47CD71FBF9" box="[466,533,1022,1045]" italics="true" pageId="16" pageNumber="435">curtus</emphasis>
</taxonomicName>
was a dubious taxon. In the latest treatise on the 
<taxonomicName id="4C44B1F2FFF04717FE4AFBA7CD42FBD9" authorityName="Gyllenhaal" authorityYear="1808" box="[438,550,1054,1077]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="435" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
of the USSR (KRYZHANOVSKIJ &amp; REICHARDT 1976), which in fact included almost the entire Palaearctic fauna, Kryzhanovskij upheld MÜLLER’ S (1937) opinion, and the name 
<taxonomicName id="4C44B1F2FFF04717FAE9FF07CB7BFF19" baseAuthorityName="Kuster" baseAuthorityYear="1849" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="435" phylum="Arthropoda" rank="species" species="puncticollis" subGenus="Nessus">
<emphasis id="B9301663FFF04717FAE9FF07CAF8FF39" box="[1301,1436,190,213]" italics="true" pageId="16" pageNumber="435">Hypocaccus</emphasis>
(
<emphasis id="B9301663FFF04717FCB9FF67CCF4FF19" box="[837,912,222,245]" italics="true" pageId="16" pageNumber="435">Nessus</emphasis>
) 
<emphasis id="B9301663FFF04717FC5CFF67CB7BFF19" box="[928,1055,222,245]" italics="true" pageId="16" pageNumber="435">puncticollis</emphasis>
</taxonomicName>
gained priority. This was followed by 
<bibRefCitation id="EFD5B780FFF04717FCA2FF47CC96FEF9" author="MAZUR S." box="[862,1010,254,277]" pageId="16" pageNumber="435" pagination="1 - 376" refId="ref21359" refString="MAZUR S. 1984: A world catalogue of Histeridae. Polskie Pismo Entomologiczne 54: 1 - 376." type="journal article" year="1984">MAZUR (1984</bibRefCitation>
, 
<bibRefCitation id="EFD5B780FFF04717FC01FF47CB52FEF9" author="MAZUR S." box="[1021,1078,254,277]" pageId="16" pageNumber="435" pagination="1 - 373" refId="ref21379" refString="MAZUR S. 1997: A world catalogue of the Histeridae (Coleoptera: Histeridae). Genus Supplement 7: 1 - 373." type="journal article" year="1997">1997</bibRefCitation>
, 
<bibRefCitation id="EFD5B780FFF04717FBBDFF47CB1CFEF9" author="MAZUR S." box="[1089,1144,254,277]" pageId="16" pageNumber="435" refId="ref21450" refString="MAZUR S. 2011: A concise catalogue of the Histeridae (Insecta: Coleoptera). Warsaw Univeristy of Life Sciences - SGGW Press, Warsaw, 332 pp." type="book" year="2011">2011</bibRefCitation>
) in all three editions of his world catalogue of the 
<taxonomicName id="4C44B1F2FFF04717FBC8FEA7CBC0FED9" authorityName="Gyllenhaal" authorityYear="1808" box="[1076,1188,286,309]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="435" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
as well as by the latest edition of the Palaearctic Catalogue by 
<bibRefCitation id="EFD5B780FFF04717FB00FE87CCE3FE99" author="LACKNER T. &amp; MAZUR S. &amp; NEWTON A. F." pageId="16" pageNumber="435" refId="ref21168" refString="LACKNER T., MAZUR S. &amp; NEWTON A. F. 2015: Family Histeridae. Pp. 76 - 130. In: LOBL I. &amp; LOBL D. (eds): Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Staphylinoidea. Volume 1. Brill Publishers, Leiden, Boston, xxiii + 1 - 900." type="book" year="2015">LACKNER et al. (2015)</bibRefCitation>
. Having examined both 
<typeStatus id="54FF74D3FFF04717FB77FEE7CBDEFE99" box="[1163,1210,350,373]" pageId="16" pageNumber="435">type</typeStatus>
specimens as well as numerous non-type specimens we can conclude that the two species are synonymous, and the earlier described taxon (
<taxonomicName id="4C44B1F2FFF04717FC7AFE07CB7CFE39" authorityName="Rosenhauer" authorityYear="1847" box="[902,1048,446,469]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="438" phylum="Arthropoda" rank="species" species="curtus" subGenus="Nannolepidius">
<emphasis id="B9301663FFF04717FC7AFE07CCFDFE39" box="[902,921,446,469]" italics="true" pageId="16" pageNumber="435">H</emphasis>
. (
<emphasis id="B9301663FFF04717FC52FE07CCA4FE39" box="[942,960,446,469]" italics="true" pageId="16" pageNumber="435">N</emphasis>
.) 
<emphasis id="B9301663FFF04717FC29FE07CB7CFE39" box="[981,1048,446,469]" italics="true" pageId="16" pageNumber="435">curtus</emphasis>
</taxonomicName>
) has the priority. Regarding external morphological variation of this species, see Remarks section of 
<taxonomicName id="4C44B1F2FFF04717FC52FE47CB2CFDF9" authorityName="Rosenhauer" authorityYear="1847" box="[942,1096,510,533]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="438" phylum="Arthropoda" rank="species" species="curtus" subGenus="Nannolepidius">
<emphasis id="B9301663FFF04717FC52FE47CCA5FDF9" box="[942,961,510,533]" italics="true" pageId="16" pageNumber="435">H</emphasis>
. (
<emphasis id="B9301663FFF04717FC24FE47CC8EFDF9" box="[984,1002,510,533]" italics="true" pageId="16" pageNumber="435">N</emphasis>
.) 
<emphasis id="B9301663FFF04717FBFDFE47CB2CFDF9" box="[1025,1096,510,533]" italics="true" pageId="16" pageNumber="435">curtus</emphasis>
</taxonomicName>
<emphasis id="B9301663FFF04717FBB5FE47CB29FDF9" box="[1097,1101,510,533]" italics="true" pageId="16" pageNumber="435">.</emphasis>
</paragraph>
<paragraph id="8BFBCA71FFF04717FCA6FDA7CBD9FD19" blockId="16.[828,1437,190,1141]" pageId="16" pageNumber="435">
<taxonomicName id="4C44B1F2FFF04717FCA6FDA7CB59FDD9" baseAuthorityName="Kuster" baseAuthorityYear="1849" box="[858,1085,542,565]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="435" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="B9301663FFF04717FCA6FDA7CB59FDD9" box="[858,1085,542,565]" italics="true" pageId="16" pageNumber="435">Saprinus puncticollis</emphasis>
</taxonomicName>
was described from a specimen found in Cagliari by Küster himself, as well as from specimen(s) brought by Mr. Handschuh from Cartagena (
<collectingCountry id="F3538AE1FFF04717FAAAFDE7CAF1FD99" box="[1366,1429,606,629]" name="Spain" pageId="16" pageNumber="435">Spain</collectingCountry>
) (
<bibRefCitation id="EFD5B780FFF04717FCBFFDC7CCB4FD79" author="KUSTER H. C." box="[835,976,638,661]" pageId="16" pageNumber="435" refId="ref21108" refString="KUSTER H. C. 1849: Die Kafer Europa's. Nach der Natur beschreiben. XVII Heft. Bauer and Raspe, Nurnberg, 100 pls." type="book" year="1849">KÜSTER 1849</bibRefCitation>
). The depository of the Spanish specimens is unknown and hence we designate the male specimen from Cagliari (Sardinia) as the 
<typeStatus id="54FF74D3FFF04717FBAAFD07CBDCFD39" box="[1110,1208,702,725]" pageId="16" pageNumber="435" type="lectotype">lectotype</typeStatus>
to fix the identity of this taxon for purpose of synonymy.
</paragraph>
<paragraph id="8BFBCA71FFF04717FCA6FD47CBC2FB99" blockId="16.[828,1437,190,1141]" pageId="16" pageNumber="435">
<taxonomicName id="4C44B1F2FFF04717FCA6FD47CB39FCF9" authorityName="Jacquelin du Val" authorityYear="1858" box="[858,1117,766,789]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="435" phylum="Arthropoda" rank="species" species="cribellaticollis">
<emphasis id="B9301663FFF04717FCA6FD47CB39FCF9" box="[858,1117,766,789]" italics="true" pageId="16" pageNumber="435">Saprinus cribellaticollis</emphasis>
</taxonomicName>
was described based on unknown number of specimens.A single specimen was located in the original collection of Jacquelin du Val, deposited in MNHN, under the label ‘ 
<taxonomicName id="4C44B1F2FFF04717FBC1FCE7CA53FC99" authorityName="Jacquelin du Val" authorityYear="1858" box="[1085,1335,862,885]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="435" phylum="Arthropoda" rank="species" species="cribellaticollis">
<emphasis id="B9301663FFF04717FBC1FCE7CA53FC99" box="[1085,1335,862,885]" italics="true" pageId="16" pageNumber="435">Saprinus cribellaticollis</emphasis>
</taxonomicName>
’. Jacquelin du Val did not provide his specimens with any labels, but, according to the curator of 
<taxonomicName id="4C44B1F2FFF04717FB71FC27CA66FC59" box="[1165,1282,926,949]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="16" pageNumber="435" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
in MNHN, A. Taghavian, he kept his types in his private collection. Therefore we presume that this specimen, which completely matches J. du Val’s description, is a 
<typeStatus id="54FF74D3FFF04717FB43FC47CA71FBF9" box="[1215,1301,1022,1045]" pageId="16" pageNumber="435" type="syntype">syntype</typeStatus>
. The species was described based on an unknown number of specimens and therefore we designate the 
<typeStatus id="54FF74D3FFF04717FB72FB87CB94FBB9" box="[1166,1264,1086,1109]" pageId="16" pageNumber="435" type="lectotype">lectotype</typeStatus>
to fix the taxon identity for purpose of synonymy.
</paragraph>
<caption id="DF3B9AF9FFF04717FF6BF861CE96F7E9" ID-DOI="http://doi.org/10.5281/zenodo.4504946" ID-Zenodo-Dep="4504946" httpUri="https://zenodo.org/record/4504946/files/figure.png" pageId="16" pageNumber="435" startId="16.[151,188,2008,2027]" targetBox="[156,1428,1206,1975]" targetPageId="16">
<paragraph id="8BFBCA71FFF04717FF6BF861CE96F7E9" blockId="16.[151,1436,2008,2053]" pageId="16" pageNumber="435">
Figs 44–45. 
<taxonomicName id="4C44B1F2FFF04717FF02F861CDFCF807" authority="(Rosenhauer, 1847)" authorityName="Lackner &amp; Seres" authorityYear="2018" baseAuthorityName="Rosenhauer" baseAuthorityYear="1847" box="[254,664,2008,2027]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="435" phylum="Arthropoda" rank="species" species="curtus" subGenus="Nessus">
<emphasis id="B9301663FFF04717FF02F861CE02F807" box="[254,358,2008,2027]" italics="true" pageId="16" pageNumber="435">Hypocaccus</emphasis>
(
<emphasis id="B9301663FFF04717FE8DF861CEC8F807" box="[369,428,2008,2027]" italics="true" pageId="16" pageNumber="435">Nessus</emphasis>
) 
<emphasis id="B9301663FFF04717FE4BF861CE88F807" box="[439,492,2008,2027]" italics="true" pageId="16" pageNumber="435">curtus</emphasis>
(
<bibRefCitation id="EFD5B780FFF04717FE0AF861CDF5F807" author="ROSENHAUER W. G." box="[502,657,2008,2027]" pageId="16" pageNumber="435" refId="ref21812" refString="ROSENHAUER W. G. 1847: Beitrage zur Insekten-Fauna Europas. Erstes Bandchen; enthalt die Beschreibung von sechzig neuen Kafern aus Bayern, Tyrol, Ungarn, etc., so wie die Kafer Tyrols, nach dem Ergebnisse von vier Reisen zusammengestellt. Mit einer Tafel Abbildungen. Th. Blasing, Erlangen, x + 159 pp." type="book" year="1847">Rosenhauer, 1847</bibRefCitation>
)
</taxonomicName>
, dorsal habitus. 44 – 
<taxonomicName id="4C44B1F2FFF04717FCBAF861CB35F807" authority="Marseul, 1876" authorityName="Marseul" authorityYear="1876" box="[838,1105,2008,2027]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="435" phylum="Arthropoda" rank="species" species="revisus">
<emphasis id="B9301663FFF04717FCBAF861CCB4F807" box="[838,976,2008,2027]" italics="true" pageId="16" pageNumber="435">Saprinus revisus</emphasis>
Marseul, 1876
</taxonomicName>
, lectotype. 45 – 
<taxonomicName id="4C44B1F2FFF04717FB2AF861CEF1F7E9" authority="G. Muller, 1937" authorityName="G. Muller" authorityYear="1937" class="Insecta" family="Histeridae" genus="Hypocacculus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="435" phylum="Arthropoda" rank="species" species="controversus" subGenus="Nessus">
<emphasis id="B9301663FFF04717FB2AF861CA2BF807" box="[1238,1359,2008,2027]" italics="true" pageId="16" pageNumber="435">Hypocacculus</emphasis>
(
<emphasis id="B9301663FFF04717FAA6F861CAF1F807" box="[1370,1429,2008,2027]" italics="true" pageId="16" pageNumber="435">Nessus</emphasis>
) 
<emphasis id="B9301663FFF04717FF6BF84BCE60F7E9" box="[151,260,2034,2053]" italics="true" pageId="16" pageNumber="435">controversus</emphasis>
G. 
<bibRefCitation id="EFD5B780FFF04717FED8F84BCEF1F7E9" author="MULLER G." box="[292,405,2034,2053]" pageId="16" pageNumber="435" pagination="97 - 112" refId="ref21522" refString="MULLER G. 1937: Histeriden-Studien. Entomologische Blatter 33: 97 - 112." type="journal article" year="1937">Müller, 1937</bibRefCitation>
</taxonomicName>
, lectotype.
</paragraph>
</caption>
<caption id="DF3B9AF9FFF14716FF6BFAECCDE0FA72" ID-DOI="http://doi.org/10.5281/zenodo.4504952" ID-Zenodo-Dep="4504952" httpUri="https://zenodo.org/record/4504952/files/figure.png" pageId="17" pageNumber="436" startId="17.[151,188,1365,1384]" targetBox="[271,1310,177,1331]" targetPageId="17">
<paragraph id="8BFBCA71FFF14716FF6BFAECCDE0FA72" blockId="17.[151,1436,1365,1438]" pageId="17" pageNumber="436">
Figs 46–54. Male genitalia of 
<taxonomicName id="4C44B1F2FFF14716FE66FAECCC58FA84" authority="(Rosenhauer, 1847)" authorityName="Lackner &amp; Seres" authorityYear="2018" baseAuthorityName="Rosenhauer" baseAuthorityYear="1847" box="[410,828,1365,1384]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="436" phylum="Arthropoda" rank="species" species="curtus" subGenus="Nessus">
<emphasis id="B9301663FFF14716FE66FAECCD66FA84" box="[410,514,1365,1384]" italics="true" pageId="17" pageNumber="436">Hypocaccus</emphasis>
(
<emphasis id="B9301663FFF14716FDF3FAECCD2EFA84" box="[527,586,1365,1384]" italics="true" pageId="17" pageNumber="436">Nessus</emphasis>
) 
<emphasis id="B9301663FFF14716FDABFAECCDE8FA84" box="[599,652,1365,1384]" italics="true" pageId="17" pageNumber="436">curtus</emphasis>
(
<bibRefCitation id="EFD5B780FFF14716FD64FAECCC52FA84" author="ROSENHAUER W. G." box="[664,822,1365,1384]" pageId="17" pageNumber="436" refId="ref21812" refString="ROSENHAUER W. G. 1847: Beitrage zur Insekten-Fauna Europas. Erstes Bandchen; enthalt die Beschreibung von sechzig neuen Kafern aus Bayern, Tyrol, Ungarn, etc., so wie die Kafer Tyrols, nach dem Ergebnisse von vier Reisen zusammengestellt. Mit einer Tafel Abbildungen. Th. Blasing, Erlangen, x + 159 pp." type="book" year="1847">Rosenhauer, 1847</bibRefCitation>
)
</taxonomicName>
. 45 – VIII sternite and tergite, ventral view; 46 – ditto, dorsal view; 47 – ditto, lateral view; 48 – IX and X tergites, dorsal view; 49 – ditto, lateral view; 50 – IX sternite (spiculum gastrale), ventral view; 51 – ditto, lateral view; 52 – aedeagus, dorsal view; 53 – ditto, lateral view.
</paragraph>
</caption>
<paragraph id="8BFBCA71FFF14716FF49FA50CEE0F98C" blockId="17.[151,760,1513,2080]" pageId="17" pageNumber="436">
<taxonomicName id="4C44B1F2FFF14716FF49FA50CE10F9EC" authorityName="Marseul" authorityYear="1862" box="[181,372,1513,1536]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="436" phylum="Arthropoda" rank="species" species="sicanus">
<emphasis id="B9301663FFF14716FF49FA50CE10F9EC" box="[181,372,1513,1536]" italics="true" pageId="17" pageNumber="436">Saprinus sicanus</emphasis>
</taxonomicName>
was described from 
<collectingRegion id="49800493FFF14716FD94FA50CDCCF9EC" box="[616,680,1513,1536]" country="Italy" name="Sicilia" pageId="17" pageNumber="436">Sicily</collectingRegion>
(
<collectingCountry id="F3538AE1FFF14716FD46FA50CD8BF9EC" box="[698,751,1513,1536]" name="Italy" pageId="17" pageNumber="436">Italy</collectingCountry>
) based on an unspecified number of specimens, therefore we designate the 
<typeStatus id="54FF74D3FFF14716FEA7F990CED9F9AC" box="[347,445,1577,1600]" pageId="17" pageNumber="436" type="lectotype">lectotype</typeStatus>
to fix the taxon identity for purpose of synonymy.
</paragraph>
<paragraph id="8BFBCA71FFF14716FF49F9D0CE4DF90C" blockId="17.[151,760,1513,2080]" pageId="17" pageNumber="436">
<taxonomicName id="4C44B1F2FFF14716FF49F9D0CE02F96C" authorityName="Marseul" authorityYear="1876" box="[181,358,1641,1664]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="436" phylum="Arthropoda" rank="species" species="revisus">
<emphasis id="B9301663FFF14716FF49F9D0CE02F96C" box="[181,358,1641,1664]" italics="true" pageId="17" pageNumber="436">Saprinus revisus</emphasis>
</taxonomicName>
was described from 
<collectingCountry id="F3538AE1FFF14716FDBDF9D0CDF4F96C" box="[577,656,1641,1664]" name="Algeria" pageId="17" pageNumber="436">Algiers</collectingCountry>
(
<collectingCountry id="F3538AE1FFF14716FD61F9D0CD94F96C" box="[669,752,1641,1664]" name="Algeria" pageId="17" pageNumber="436">Algeria</collectingCountry>
) based on an unknown number of specimens, therefore we designate the 
<typeStatus id="54FF74D3FFF14716FED9F910CEE2F92C" box="[293,390,1705,1728]" pageId="17" pageNumber="436" type="lectotype">lectotype</typeStatus>
to fix the taxon identity for purpose of synonymy.
</paragraph>
<paragraph id="8BFBCA71FFF14716FF49F950CEEAF7CC" blockId="17.[151,760,1513,2080]" pageId="17" pageNumber="436">
The 
<typeStatus id="54FF74D3FFF14716FF15F950CE73F8EC" box="[233,279,1769,1792]" pageId="17" pageNumber="436">type</typeStatus>
of 
<taxonomicName id="4C44B1F2FFF14716FEBCF950CEC5F8EC" authorityName="Lackner &amp; Seres" authorityYear="2018" baseAuthorityName="Rosenhauer" baseAuthorityYear="1847" box="[320,417,1769,1792]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="436" phylum="Arthropoda" rank="species" species="curtus">
<emphasis id="B9301663FFF14716FEBCF950CEC5F8EC" box="[320,417,1769,1792]" italics="true" pageId="17" pageNumber="436">S. curtus</emphasis>
</taxonomicName>
was found in mid-19 
<superScript id="7C316739FFF14716FD6EF95ECDFAF919" attach="left" box="[658,670,1767,1781]" fontSize="6" pageId="17" pageNumber="436">th</superScript>
century ‘Hungary’. This vague locality could refer to anywhere in the former Hungarian monarchy, which stretched south to the Adriatic Sea. It is possible that this species will be discovered in countries of the former 
<collectingCountry id="F3538AE1FFF14716FD9CF8D0CDBEF86C" box="[608,730,1897,1920]" name="Serbia" pageId="17" pageNumber="436">Yugoslavia</collectingCountry>
. It is a rather rare and seldom-collected species apparently spread around the Mediterranean Sea. Its rarity and slight morphological differences regarding dorsal punctation or course of carinal prosternal striae probably account for its numerous synonymies.
</paragraph>
<paragraph id="8BFBCA71FFF14716FCB4FA58CBD7F9FE" blockId="17.[828,1438,1505,1722]" pageId="17" pageNumber="436">
<taxonomicName id="4C44B1F2FFF14716FCB4FA58CAEBFA14" authority="(G. Muller, 1937)" baseAuthorityName="G. Muller" baseAuthorityYear="1937" box="[840,1423,1505,1528]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="436" phylum="Arthropoda" rank="species" species="controversus" subGenus="Nessus">
<emphasis id="B9301663FFF14716FCB4FA58CAEBFA14" bold="true" box="[840,1423,1505,1528]" pageId="17" pageNumber="436">
<emphasis id="B9301663FFF14716FCB4FA58CCA9FA14" bold="true" box="[840,973,1505,1528]" italics="true" pageId="17" pageNumber="436">Hypocaccus</emphasis>
(
<emphasis id="B9301663FFF14716FC21FA58CB4EFA14" bold="true" box="[989,1066,1505,1528]" italics="true" pageId="17" pageNumber="436">Nessus</emphasis>
) 
<emphasis id="B9301663FFF14716FBC6FA58CBA3FA14" bold="true" box="[1082,1223,1505,1528]" italics="true" pageId="17" pageNumber="436">controversus</emphasis>
(G. 
<bibRefCitation id="EFD5B780FFF14716FB04FA58CAECFA14" author="MULLER G." box="[1272,1416,1505,1528]" pageId="17" pageNumber="436" pagination="97 - 112" refId="ref21522" refString="MULLER G. 1937: Histeriden-Studien. Entomologische Blatter 33: 97 - 112." type="journal article" year="1937">Müller, 1937</bibRefCitation>
)
</emphasis>
</taxonomicName>
(
<figureCitation id="137FD6F4FFF14716FBD7FA46CB09F9FE" box="[1067,1133,1535,1554]" captionStart="Figs 44–45" captionStartId="16.[151,188,2008,2027]" captionTargetBox="[156,1428,1206,1975]" captionTargetId="figure-13@16.[151,1436,1200,1986]" captionTargetPageId="16" captionText="Figs 44–45.Hypocaccus (Nessus) curtus (Rosenhauer, 1847), dorsal habitus. 44 – Saprinus revisus Marseul, 1876, lectotype. 45 – Hypocacculus (Nessus) controversus G. Müller, 1937, lectotype." figureDoi="http://doi.org/10.5281/zenodo.4504946" httpUri="https://zenodo.org/record/4504946/files/figure.png" pageId="17" pageNumber="436">Figs 45</figureCitation>
, 
<figureCitation id="137FD6F4FFF14716FB8AFA46CBC8F9FE" box="[1142,1196,1535,1554]" captionStart="Figs 55–63" captionStartId="18.[150,187,1309,1328]" captionTargetBox="[294,1285,171,1249]" captionTargetId="figure-468@18.[280,1288,166,1252]" captionTargetPageId="18" captionText="Figs 55–63. Male genitalia of Hypocaccus (Nessus) controversus (Müller, 1937). 55 – VIII sternite and tergite, ventral view; 56 – ditto, dorsal view; 57 – ditto, lateral view; 58 – IX and X tergites, dorsal view; 59 – ditto, lateral view; 60 – IX sternite (spiculum gastrale), ventral view; 61 – ditto, lateral view; 62 – aedeagus, dorsal view; 63 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4504960" httpUri="https://zenodo.org/record/4504960/files/figure.png" pageId="17" pageNumber="436">55–63</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35E99FAFFF14716FCC0F998CBCCF956" pageId="17" pageNumber="436" type="reference_group">
<paragraph id="8BFBCA71FFF14716FCC0F998CAFAF9D8" blockId="17.[828,1438,1505,1722]" box="[828,1438,1569,1588]" pageId="17" pageNumber="436">
<treatmentCitationGroup id="AB54ED5FFFF14716FCC0F998CAFAF9D8" box="[828,1438,1569,1588]" pageId="17" pageNumber="436">
<treatmentCitation id="0AE5EC60FFF14716FCC0F998CBB9F9D8" author="MULLER G." box="[828,1245,1569,1588]" page="115" pageId="17" pageNumber="436" year="1937">
<taxonomicName id="4C44B1F2FFF14716FCC0F998CBB9F9D8" ID-CoL="7VHPG" authority="G. Muller, 1937: 115" authorityName="G. Muller" authorityPageNumber="115" authorityYear="1937" box="[828,1245,1569,1588]" class="Insecta" family="Histeridae" genus="Hypocacculus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="436" phylum="Arthropoda" rank="species" species="controversus">
<emphasis id="B9301663FFF14716FCC0F998CB42F9D8" box="[828,1062,1569,1588]" italics="true" pageId="17" pageNumber="436">Hypocacculus controversus</emphasis>
G. 
<bibRefCitation id="EFD5B780FFF14716FBB9F998CBB9F9D8" author="MULLER G." box="[1093,1245,1569,1588]" pageId="17" pageNumber="436" pagination="97 - 112" refId="ref21522" refString="MULLER G. 1937: Histeriden-Studien. Entomologische Blatter 33: 97 - 112." type="journal article" year="1937">Müller, 1937: 115</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFBCA71FFF14716FCC0F985CAFAF968" blockId="17.[828,1438,1505,1722]" pageId="17" pageNumber="436">
<treatmentCitationGroup id="AB54ED5FFFF14716FCC0F985CAFAF968" pageId="17" pageNumber="436">
<treatmentCitation id="0AE5EC60FFF14716FCC0F985CCA5F986" author="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N." page="204" pageId="17" pageNumber="436" year="1976">
<taxonomicName id="4C44B1F2FFF14716FCC0F985CCF1F986" ID-CoL="7VHPG" authority=": KRYZHANOVSKIJ &amp; REICHARDT (1976)" authorityName=": KRYZHANOVSKIJ &amp; REICHARDT" authorityYear="1976" class="Insecta" family="Histeridae" genus="Hypocacculus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="436" phylum="Arthropoda" rank="species" species="controversus" subGenus="Nessus">
<emphasis id="B9301663FFF14716FCC0F985CCDDF9A3" box="[828,953,1596,1615]" italics="true" pageId="17" pageNumber="436">Hypocacculus</emphasis>
(
<emphasis id="B9301663FFF14716FC35F985CB62F9A3" box="[969,1030,1596,1615]" italics="true" pageId="17" pageNumber="436">Nessus</emphasis>
) 
<emphasis id="B9301663FFF14716FBEBF985CBEDF9A3" box="[1047,1161,1596,1615]" italics="true" pageId="17" pageNumber="436">controversus</emphasis>
: 
<bibRefCitation id="EFD5B780FFF14716FB64F985CCF1F986" author="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N." pageId="17" pageNumber="436" refId="ref20992" refString="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N. 1976: Zhuki nadsemeystva Histeroidea (semeystva Sphaeritidae, Histeridae, Synteliidae). Fauna SSSR, Zhestokrylye, Tom V, vyp. 4. [Beetles of the superfamily Histeroidea (families Sphaeritidae, Histeridae, Syntelidae) Fauna SSSR, Coleoptera, Vol. 5, part 4.]. Nauka, Leningrad, 434 pp (in Russian)." type="book" year="1976">KRYZHANOVSKIJ &amp; REICHARDT (1976)</bibRefCitation>
</taxonomicName>
: 204
</treatmentCitation>
, 212 (keyed, redescription); 
<treatmentCitation id="0AE5EC60FFF14716FB45F9EECA31F986" author="MAZUR S." box="[1209,1365,1623,1642]" page="89" pageId="17" pageNumber="436" year="1984">
<bibRefCitation id="EFD5B780FFF14716FB45F9EECA52F986" author="MAZUR S." box="[1209,1334,1623,1642]" pageId="17" pageNumber="436" pagination="1 - 376" refId="ref21359" refString="MAZUR S. 1984: A world catalogue of Histeridae. Polskie Pismo Entomologiczne 54: 1 - 376." type="journal article" year="1984">MAZUR (1984)</bibRefCitation>
: 89
</treatmentCitation>
(catalogue); 
<treatmentCitation id="0AE5EC60FFF14716FC76F9C8CB49F968" author="MAZUR S." box="[906,1069,1649,1668]" page="252" pageId="17" pageNumber="436" year="1997">
<bibRefCitation id="EFD5B780FFF14716FC76F9C8CB62F968" author="MAZUR S." box="[906,1030,1649,1668]" pageId="17" pageNumber="436" pagination="1 - 373" refId="ref21379" refString="MAZUR S. 1997: A world catalogue of the Histeridae (Coleoptera: Histeridae). Genus Supplement 7: 1 - 373." type="journal article" year="1997">MAZUR (1997)</bibRefCitation>
: 252
</treatmentCitation>
(catalogue); 
<treatmentCitation id="0AE5EC60FFF14716FB67F9C8CA56F968" author="MAZUR S." box="[1179,1330,1649,1668]" page="93" pageId="17" pageNumber="436" year="2004">
<bibRefCitation id="EFD5B780FFF14716FB67F9C8CA72F968" author="MAZUR S." box="[1179,1302,1649,1668]" pageId="17" pageNumber="436" refId="ref21404" refString="MAZUR S. 2004: Family Histeridae. Pp. 68 - 102. In: LOBL I. &amp; SMETANA A. (eds): Catalogue of Palaearctic Coleoptera, Vol. 2, Hydrophiloidea-Histeroidea-Staphylinidea. Apollo Books, Stenstrup, 942 pp." type="book" year="2004">MAZUR (2004)</bibRefCitation>
:93
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFBCA71FFF14716FCC0F935CBCCF956" blockId="17.[828,1438,1505,1722]" pageId="17" pageNumber="436">
<treatmentCitationGroup id="AB54ED5FFFF14716FCC0F935CBCCF956" pageId="17" pageNumber="436">
<treatmentCitation id="0AE5EC60FFF14716FCC0F935CA4FF973" author="MAZUR S." box="[828,1323,1676,1695]" page="208" pageId="17" pageNumber="436" year="2011">
<taxonomicName id="4C44B1F2FFF14716FCC0F935CA64F973" ID-CoL="7VHTV" authority=": MAZUR (2011)" authorityName="MAZUR" authorityYear="2011" box="[828,1280,1676,1695]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="436" phylum="Arthropoda" rank="species" species="controversus" subGenus="Nessus">
<emphasis id="B9301663FFF14716FCC0F935CCC2F973" box="[828,934,1676,1695]" italics="true" pageId="17" pageNumber="436">Hypocaccus</emphasis>
(
<emphasis id="B9301663FFF14716FC49F935CC95F973" box="[949,1009,1676,1695]" italics="true" pageId="17" pageNumber="436">Nessus</emphasis>
) 
<emphasis id="B9301663FFF14716FBFDF935CB15F973" box="[1025,1137,1676,1695]" italics="true" pageId="17" pageNumber="436">controversus</emphasis>
: 
<bibRefCitation id="EFD5B780FFF14716FB83F935CA64F973" author="MAZUR S." box="[1151,1280,1676,1695]" pageId="17" pageNumber="436" refId="ref21450" refString="MAZUR S. 2011: A concise catalogue of the Histeridae (Insecta: Coleoptera). Warsaw Univeristy of Life Sciences - SGGW Press, Warsaw, 332 pp." type="book" year="2011">MAZUR (2011)</bibRefCitation>
</taxonomicName>
: 208
</treatmentCitation>
(catalogue); 
<treatmentCitation id="0AE5EC60FFF14716FCA6F91ECB58F956" author="LACKNER T. &amp; MAZUR S. &amp; NEWTON A. F." box="[858,1084,1703,1722]" page="117" pageId="17" pageNumber="436" year="2015">
<bibRefCitation id="EFD5B780FFF14716FCA6F91ECB70F956" author="LACKNER T. &amp; MAZUR S. &amp; NEWTON A. F." box="[858,1044,1703,1722]" pageId="17" pageNumber="436" refId="ref21168" refString="LACKNER T., MAZUR S. &amp; NEWTON A. F. 2015: Family Histeridae. Pp. 76 - 130. In: LOBL I. &amp; LOBL D. (eds): Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Staphylinoidea. Volume 1. Brill Publishers, Leiden, Boston, xxiii + 1 - 900." type="book" year="2015">LACKNER et al. (2015)</bibRefCitation>
: 117
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35E99FAFFF14714FCC0F969CE56FD39" lastPageId="19" lastPageNumber="438" pageId="17" pageNumber="436" type="materials_examined">
<paragraph id="8BFBCA71FFF14715FCC0F969CD6BFA15" blockId="17.[828,1438,1744,2084]" lastBlockId="18.[151,761,1430,1929]" lastPageId="18" lastPageNumber="437" pageId="17" pageNumber="436">
<emphasis id="B9301663FFF14716FCC0F969CB49F90F" bold="true" box="[828,1069,1744,1763]" pageId="17" pageNumber="436">Type material examined.</emphasis>
<emphasis id="B9301663FFF14716FBCAF969CA53F90F" box="[1078,1335,1744,1763]" italics="true" pageId="17" pageNumber="436">
<taxonomicName id="4C44B1F2FFF14716FBCAF969CA57F90F" authorityName=": KRYZHANOVSKIJ &amp; REICHARDT" authorityYear="1976" box="[1078,1331,1744,1763]" class="Insecta" family="Histeridae" genus="Hypocacculus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="436" phylum="Arthropoda" rank="species" species="controversus">Hypocacculus controversus</taxonomicName>
.
</emphasis>
<typeStatus id="54FF74D3FFF14716FAC3F969CAF9F90E" box="[1343,1437,1744,1763]" pageId="17" pageNumber="436" type="lectotype">LECTOTYPE</typeStatus>
(present designation): 
<specimenCount id="9D4201F8FFF14716FC03F951CB6BF913" box="[1023,1039,1768,1791]" pageId="17" pageNumber="436" type="female">♀</specimenCount>
(
<figureCitation id="137FD6F4FFF14716FBE0F952CB3AF912" box="[1052,1118,1771,1790]" captionStart="Figs 44–45" captionStartId="16.[151,188,2008,2027]" captionTargetBox="[156,1428,1206,1975]" captionTargetId="figure-13@16.[151,1436,1200,1986]" captionTargetPageId="16" captionText="Figs 44–45.Hypocaccus (Nessus) curtus (Rosenhauer, 1847), dorsal habitus. 44 – Saprinus revisus Marseul, 1876, lectotype. 45 – Hypocacculus (Nessus) controversus G. Müller, 1937, lectotype." figureDoi="http://doi.org/10.5281/zenodo.4504946" httpUri="https://zenodo.org/record/4504946/files/figure.png" pageId="17" pageNumber="436">Fig. 45</figureCitation>
), mounted on a triangular mounting card, right metatarsus missing, ‘ 
<specimenCount id="9D4201F8FFF14716FBA9F8BBCB01F8F5" box="[1109,1125,1794,1817]" pageId="17" pageNumber="436" type="female">♀</specimenCount>
[written] // Banat 1909 / Herkulesbad / leg. M. Hilf / Coll. O. Leonhardt [printed] // sbsp. / controversus [written] // TYPUS [light-ochre label, printed] // scat. / Hist. 6 [yellow label, written] // 
<taxonomicName id="4C44B1F2FFF14716FC2DF8EFCB2EF885" authorityName="Bickhardt" authorityYear="1914" box="[977,1098,1878,1897]" class="Insecta" family="Histeridae" genus="Hypocacculus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="436" phylum="Arthropoda" rank="genus">Hypocacculus</taxonomicName>
/ (
<taxonomicName id="4C44B1F2FFF14716FB99F8EFCBC0F885" authorityName="Reichardt" authorityYear="1932" box="[1125,1188,1878,1897]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="432" phylum="Arthropoda" rank="subGenus" subGenus="Nessus">Nessus</taxonomicName>
) / controversus / G. 
<bibRefCitation id="EFD5B780FFF14716FAA2F8EFCC03F86F" author="MULLER G." pageId="17" pageNumber="436" pagination="97 - 112" refId="ref21522" refString="MULLER G. 1937: Histeriden-Studien. Entomologische Blatter 33: 97 - 112." type="journal article" year="1937">Müller, 1937</bibRefCitation>
/ 
<materialsCitation id="3B2CC02CFFF14716FC8BF8C9CAFDF86F" ID-GBIF-Occurrence="3313348301" box="[887,1433,1904,1923]" collectionCode="CST" collectorName="T. Lackner" pageId="17" pageNumber="436" specimenCount="1" typeStatus="lectotype">
<typeStatus id="54FF74D3FFF14716FC8BF8C9CC95F86F" box="[887,1009,1904,1923]" pageId="17" pageNumber="436" type="lectotype">LECTOTYPE</typeStatus>
/ des. 
<collectorName id="26B1AFA7FFF14716FBDBF8C9CBE7F86F" box="[1063,1155,1904,1923]" pageId="17" pageNumber="436">T. Lackner</collectorName>
2017 [red label, written]’ (
<collectionCode id="ED5552B4FFF14716FA94F8C9CAF7F86F" box="[1384,1427,1904,1923]" pageId="17" pageNumber="436">CST</collectionCode>
)
</materialsCitation>
. 
<typeStatus id="54FF74D3FFF14716FCC0F832CCA1F870" box="[828,965,1931,1950]" pageId="17" pageNumber="436" type="paralectotype">PARALECTOTYPES</typeStatus>
: 
<specimenCount id="9D4201F8FFF14716FC33F832CC8BF873" box="[975,1007,1928,1951]" pageId="17" pageNumber="436" type="female">1 ♀</specimenCount>
, side-mounted on a triangular mounting point, left meso- and metatarsus missing, ‘Athen / Phaleron [written] // Da Scat. / 6 [yellow label, written] // 
<taxonomicName id="4C44B1F2FFF14716FBDDF879CB8EF83F" authority="" box="[1057,1258,1984,2003]" class="Insecta" family="Histeridae" genus="Hypocacculus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="436" phylum="Arthropoda" rank="subGenus" subGenus="Nessus">Hypocacculus (Nessus)</taxonomicName>
/ 
<taxonomicName id="4C44B1F2FFF14716FB07F879CAFAF83F" authority="Mull." authorityName="G. Muller" authorityYear="1937" box="[1275,1438,1984,2003]" class="Insecta" family="Histeridae" genus="Hypocacculus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="436" phylum="Arthropoda" rank="species" species="controversus">controversus Müll.</taxonomicName>
/ Det. T. Lackner 2017 [printed-written] // 
<taxonomicName id="4C44B1F2FFF14716FB4AF862CA4BF802" authorityName="Bickhardt" authorityYear="1914" box="[1206,1327,2011,2030]" class="Insecta" family="Histeridae" genus="Hypocacculus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="436" phylum="Arthropoda" rank="genus">Hypocacculus</taxonomicName>
/ (
<taxonomicName id="4C44B1F2FFF14716FAB6F862CAEDF802" authorityName="Reichardt" authorityYear="1932" box="[1354,1417,2011,2030]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="432" phylum="Arthropoda" rank="subGenus" subGenus="Nessus">Nessus</taxonomicName>
) / controversus / G. 
<bibRefCitation id="EFD5B780FFF14716FC29F84FCB21F7E5" author="MULLER G." box="[981,1093,2038,2057]" pageId="17" pageNumber="436" pagination="97 - 112" refId="ref21522" refString="MULLER G. 1937: Histeriden-Studien. Entomologische Blatter 33: 97 - 112." type="journal article" year="1937">Müller, 1937</bibRefCitation>
/ 
<typeStatus id="54FF74D3FFF14716FBAAF84FCA63F7E5" box="[1110,1287,2038,2057]" pageId="17" pageNumber="436" type="paralectotype">PARALECTOTYPE</typeStatus>
/ des. T. Lackner 2017 [red label, written]’ (
<collectionCode id="ED5552B4FFF14716FBE7F7A9CB2DF7CF" box="[1051,1097,2064,2083]" pageId="17" pageNumber="436">CST</collectionCode>
). 
<specimenCount id="9D4201F8FFF14716FBAEF7A9CB0AF7C8" box="[1106,1134,2061,2084]" pageId="17" pageNumber="436" type="female">1 ♀</specimenCount>
, ‘Saloniki / Schatzmayr [written] // Da Scat. / 6 [yellow label, written] // 
<taxonomicName id="4C44B1F2FFF24715FE21FA2FCDC3FA45" authority="" box="[477,679,1430,1449]" class="Insecta" family="Histeridae" genus="Hypocacculus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="437" phylum="Arthropoda" rank="subGenus" subGenus="Nessus">Hypocacculus (Nessus)</taxonomicName>
/ 
<taxonomicName id="4C44B1F2FFF24715FD46FA2FCE67FA2F" authority="Mull." authorityName="G. Muller" authorityYear="1937" class="Insecta" family="Histeridae" genus="Hypocacculus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="437" phylum="Arthropoda" rank="species" species="controversus">controversus Müll.</taxonomicName>
/ Det. T. Lackner 2017 [printed-written] // 
<taxonomicName id="4C44B1F2FFF24715FD8FFA09CD88FA2F" authorityName="Bickhardt" authorityYear="1914" box="[627,748,1456,1475]" class="Insecta" family="Histeridae" genus="Hypocacculus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="437" phylum="Arthropoda" rank="genus">Hypocacculus</taxonomicName>
/ (
<taxonomicName id="4C44B1F2FFF24715FF61FA72CFB8FA32" authorityName="Reichardt" authorityYear="1932" box="[157,220,1483,1502]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="432" phylum="Arthropoda" rank="subGenus" subGenus="Nessus">Nessus</taxonomicName>
) / controversus / G. 
<bibRefCitation id="EFD5B780FFF24715FE6CFA72CD64FA32" author="MULLER G." box="[400,512,1483,1502]" pageId="18" pageNumber="437" pagination="97 - 112" refId="ref21522" refString="MULLER G. 1937: Histeriden-Studien. Entomologische Blatter 33: 97 - 112." type="journal article" year="1937">Müller, 1937</bibRefCitation>
/ 
<typeStatus id="54FF74D3FFF24715FDEEFA72CDA7FA32" box="[530,707,1483,1502]" pageId="18" pageNumber="437" type="paralectotype">PARALECTOTYPE</typeStatus>
/ des. T. Lackner 2017 [red label, written]’ (
<collectionCode id="ED5552B4FFF24715FE25FA5FCD60FA15" box="[473,516,1510,1529]" pageId="18" pageNumber="437">CST</collectionCode>
).
</paragraph>
<caption id="DF3B9AF9FFF24715FF6AFAA4CDE0FA89" ID-DOI="http://doi.org/10.5281/zenodo.4504960" ID-Zenodo-Dep="4504960" httpUri="https://zenodo.org/record/4504960/files/figure.png" pageId="18" pageNumber="437" startId="18.[150,187,1309,1328]" targetBox="[294,1285,171,1249]" targetPageId="18">
<paragraph id="8BFBCA71FFF24715FF6AFAA4CDE0FA89" blockId="18.[150,1437,1309,1381]" pageId="18" pageNumber="437">
Figs 55–63. Male genitalia of 
<taxonomicName id="4C44B1F2FFF24715FE61FAA4CC2BFADC" authority="(Muller, 1937)" baseAuthorityName="G. Muller" baseAuthorityYear="1937" box="[413,847,1309,1328]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="437" phylum="Arthropoda" rank="species" species="controversus" subGenus="Nessus">
<emphasis id="B9301663FFF24715FE61FAA4CD61FADC" box="[413,517,1309,1328]" italics="true" pageId="18" pageNumber="437">Hypocaccus</emphasis>
(
<emphasis id="B9301663FFF24715FDEFFAA4CD2AFADC" box="[531,590,1309,1328]" italics="true" pageId="18" pageNumber="437">Nessus</emphasis>
) 
<emphasis id="B9301663FFF24715FDA7FAA4CDACFADC" box="[603,712,1309,1328]" italics="true" pageId="18" pageNumber="437">controversus</emphasis>
(
<bibRefCitation id="EFD5B780FFF24715FD29FAA4CC2DFADC" author="MULLER G." box="[725,841,1309,1328]" pageId="18" pageNumber="437" pagination="97 - 112" refId="ref21522" refString="MULLER G. 1937: Histeriden-Studien. Entomologische Blatter 33: 97 - 112." type="journal article" year="1937">Müller, 1937</bibRefCitation>
)
</taxonomicName>
. 55 – VIII sternite and tergite, ventral view; 56 – ditto, dorsal view; 57 – ditto, lateral view; 58 – IX and X tergites, dorsal view; 59 – ditto, lateral view; 60 – IX sternite (spiculum gastrale), ventral view; 61 – ditto, lateral view; 62 – aedeagus, dorsal view; 63 – ditto, lateral view.
</paragraph>
</caption>
<paragraph id="8BFBCA71FFF24715FF6BF9B9CE58F865" blockId="18.[151,761,1430,1929]" pageId="18" pageNumber="437">
<emphasis id="B9301663FFF24715FF6BF9B9CD7FF9FF" bold="true" box="[151,539,1536,1555]" pageId="18" pageNumber="437">
Additional material examined. 
<collectingCountry id="F3538AE1FFF24715FE41F9B9CD73F9FF" box="[445,535,1536,1555]" name="Cyprus" pageId="18" pageNumber="437">CYPRUS</collectingCountry>
:
</emphasis>
Cyprus, no further data, 
<specimenCount id="9D4201F8FFF24715FF6BF9A2CFB2F9C2" box="[151,214,1563,1582]" pageId="18" pageNumber="437" type="generic">1 spec.</specimenCount>
(probably a male, genitalia lost), Baudi, (
<collectionCode id="ED5552B4FFF24715FDB9F9A2CDE2F9C2" box="[581,646,1563,1582]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="18" pageNumber="437">MFNB</collectionCode>
). 
<emphasis id="B9301663FFF24715FD6AF9A2CD9CF9C2" bold="true" box="[662,760,1563,1582]" pageId="18" pageNumber="437">
<collectingCountry id="F3538AE1FFF24715FD6AF9A2CD97F9C2" box="[662,755,1563,1582]" name="Greece" pageId="18" pageNumber="437">GREECE</collectingCountry>
:
</emphasis>
Greece, 1 ♁ (genitalia lost, sexed by the protarsi), 
<specimenCount id="9D4201F8FFF24715FDA9F98FCD12F9A5" box="[597,630,1586,1609]" pageId="18" pageNumber="437" type="female">1 ♀</specimenCount>
, Emge lgt., C. &amp; O. Vogt coll. (
<specimenCount id="9D4201F8FFF24715FED5F9E9CE2DF988" box="[297,329,1613,1636]" pageId="18" pageNumber="437" type="female">1 ♀</specimenCount>
in 
<collectionCode id="ED5552B4FFF24715FE99F9E9CEFBF98F" box="[357,415,1616,1635]" pageId="18" pageNumber="437">CTLA</collectionCode>
, 1 ♁ in 
<collectionCode id="ED5552B4FFF24715FE18F9E9CD42F98F" box="[484,550,1616,1635]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="18" pageNumber="437">MSNG</collectionCode>
); Greece, 1 ♁, (
<collectionCode id="ED5552B4FFF24715FD52F9E9CD8BF98F" box="[686,751,1616,1635]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="18" pageNumber="437">MFNB</collectionCode>
). 
<emphasis id="B9301663FFF24715FF6BF9D2CFB9F992" bold="true" box="[151,221,1643,1662]" pageId="18" pageNumber="437">
<collectingRegion id="49800493FFF24715FF6BF9D2CFB2F990" box="[151,214,1643,1662]" country="Greece" name="Attiki" pageId="18" pageNumber="437">ATTICA</collectingRegion>
:
</emphasis>
Attica, no further data, 
<specimenCount id="9D4201F8FFF24715FE48F9D2CE81F992" box="[436,485,1639,1662]" pageId="18" pageNumber="437" type="female">2 ♀♀</specimenCount>
(
<collectionCode id="ED5552B4FFF24715FE0FF9D2CD50F992" box="[499,564,1643,1662]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="18" pageNumber="437">MFNB</collectionCode>
). 
<emphasis id="B9301663FFF24715FDB9F9D2CDE0F992" bold="true" box="[581,644,1643,1662]" pageId="18" pageNumber="437">
<collectingRegion id="49800493FFF24715FDB9F9D2CD19F990" box="[581,637,1643,1662]" country="Greece" name="Kriti" pageId="18" pageNumber="437">CRETE</collectingRegion>
:
</emphasis>
Lerapetra E, 
<quantity id="4CBC6794FFF24715FF6BF93FCFBEF975" box="[151,218,1670,1689]" metricMagnitude="1" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="18" pageNumber="437" unit="m" value="10.0" valueMax="20.0" valueMin="0.0">0–20 m</quantity>
, 
<date id="FFFAECB1FFF24715FF1FF93FCE00F975" box="[227,356,1670,1689]" pageId="18" pageNumber="437" value="2000-04-17" valueMax="2000-04-23" valueMin="2000-04-17">17.–23.iv.2000</date>
, 
<specimenCount id="9D4201F8FFF24715FE91F93FCEE9F975" box="[365,397,1666,1689]" pageId="18" pageNumber="437" type="female">1 ♀</specimenCount>
, A. Kopetz ltg. (
<collectionCode id="ED5552B4FFF24715FDE1F93FCD04F975" box="[541,608,1670,1689]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="18" pageNumber="437">MSNG</collectionCode>
). 
<emphasis id="B9301663FFF24715FD91F93FCD9CF975" bold="true" box="[621,760,1670,1689]" pageId="18" pageNumber="437">
<collectingCountry id="F3538AE1FFF24715FD91F93FCD95F97B" box="[621,753,1670,1689]" name="Greece" pageId="18" pageNumber="437">IONIAN ISLANDS</collectingCountry>
:
</emphasis>
Zante [=Zakynthos],Kalamaki,1909, 1♁, M.Hilf lgt., Coll. O. Leonhard (
<collectionCode id="ED5552B4FFF24715FF61F902CFB9F922" box="[157,221,1723,1742]" pageId="18" pageNumber="437">MNFB</collectionCode>
). 
<emphasis id="B9301663FFF24715FF16F902CEE4F922" bold="true" box="[234,384,1723,1742]" pageId="18" pageNumber="437">
<collectingCountry id="F3538AE1FFF24715FF16F902CE21F922" box="[234,325,1723,1742]" name="Jordan" pageId="18" pageNumber="437">JORDAN</collectingCountry>
: 
<collectingRegion id="49800493FFF24715FEB0F902CE1DF920" box="[332,377,1723,1742]" country="Jordan" name="Irbid" pageId="18" pageNumber="437">IRBID</collectingRegion>
:
</emphasis>
<quantity id="4CBC6794FFF24715FE78F902CEC9F922" box="[388,429,1723,1742]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="437" unit="km" value="5.0">5 km</quantity>
NE of El Karama, 
<date id="FFFAECB1FFF24715FDB4F902CDC0F922" box="[584,676,1723,1742]" pageId="18" pageNumber="437" value="1994-03-31">31.iii.1994</date>
, 
<geoCoordinate id="EE70ACB6FFF24715FD50F902CD90F922" box="[684,756,1723,1742]" degrees="31.58" direction="north" orientation="latitude" pageId="18" pageNumber="437" precision="555" value="31.58">31.58°N</geoCoordinate>
, 
<geoCoordinate id="EE70ACB6FFF24715FF6BF96FCFBBF905" box="[151,223,1750,1769]" degrees="35.36" direction="east" orientation="longitude" pageId="18" pageNumber="437" precision="555" value="35.36">35.36°E</geoCoordinate>
, 
<quantity id="4CBC6794FFF24715FF1BF96FCE44F905" box="[231,288,1750,1769]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="437" unit="m" value="200.0">200 m</quantity>
, 
<specimenCount id="9D4201F8FFF24715FED4F96FCE23F905" box="[296,327,1746,1769]" pageId="18" pageNumber="437" type="female">1 ♀</specimenCount>
, S. Bečvář jun. &amp; sen. lgt. (dubious identification) (
<collectionCode id="ED5552B4FFF24715FF61F949CFBBF8EF" box="[157,223,1776,1795]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="18" pageNumber="437">MSNG</collectionCode>
); Toten Meer [= Dead Sea], 
<date id="FFFAECB1FFF24715FE2BF949CD48F8EF" box="[471,556,1776,1795]" pageId="18" pageNumber="437" value="1963-05-10">10.v.1963</date>
, 
<specimenCount id="9D4201F8FFF24715FDCBF949CD33F8E8" box="[567,599,1773,1796]" pageId="18" pageNumber="437" type="female">1 ♀</specimenCount>
, J. Klapperich lgt. (dubious identification) (
<collectionCode id="ED5552B4FFF24715FE97F8B2CECAF8F2" box="[363,430,1803,1822]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="18" pageNumber="437">MSNG</collectionCode>
). 
<emphasis id="B9301663FFF24715FE40F8B2CD16F8F2" bold="true" box="[444,626,1803,1822]" pageId="18" pageNumber="437">
<collectingCountry id="F3538AE1FFF24715FE40F8B2CD4EF8F2" box="[444,554,1803,1822]" name="Romania" pageId="18" pageNumber="437">ROMANIA</collectingCountry>
: 
<collectingRegion id="49800493FFF24715FDCFF8B2CD0FF8F0" box="[563,619,1803,1822]" country="Romania" name="BANAT" pageId="18" pageNumber="437">BANAT</collectingRegion>
:
</emphasis>
Banat, Orșova, 1909, 
<specimenCount id="9D4201F8FFF24715FF32F89FCF8AF8D5" box="[206,238,1826,1849]" pageId="18" pageNumber="437" type="female">1 ♀</specimenCount>
, M. Hilf lgt., coll. O. Leonhard (
<collectionCode id="ED5552B4FFF24715FDF2F89FCD2BF8D5" box="[526,591,1830,1849]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="18" pageNumber="437">MFNB</collectionCode>
). 
<emphasis id="B9301663FFF24715FDA2F89FCFD7F8BF" bold="true" pageId="18" pageNumber="437">
<collectingCountry id="F3538AE1FFF24715FDA2F89FCDDEF8D5" box="[606,698,1830,1849]" name="Tunisia" pageId="18" pageNumber="437">TUNISIA</collectingCountry>
: 
<collectingRegion id="49800493FFF24715FD39F89FCFC8F8BD" country="Tunisia" name="BA" pageId="18" pageNumber="437">DJERBA</collectingRegion>
:
</emphasis>
Rass Taguernes, 
<date id="FFFAECB1FFF24715FEB0F8F9CEADF8BF" box="[332,457,1856,1875]" pageId="18" pageNumber="437" value="1997-02-10" valueMax="1997-02-20" valueMin="1997-02-10">10.–20.ii.1997</date>
, 
<specimenCount id="9D4201F8FFF24715FE28F8F9CE91F8B8" box="[468,501,1853,1876]" pageId="18" pageNumber="437" type="female">1 ♀</specimenCount>
, Egger Manfred lgt. (dubious identification) (
<collectionCode id="ED5552B4FFF24715FEEBF8E2CE3DF882" box="[279,345,1883,1902]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="18" pageNumber="437">MSNG</collectionCode>
). 
<emphasis id="B9301663FFF24715FE9AF8E2CD65F882" bold="true" box="[358,513,1883,1902]" pageId="18" pageNumber="437">
<collectingCountry id="F3538AE1FFF24715FE9AF8E2CEA6F882" box="[358,450,1883,1902]" name="Turkey" pageId="18" pageNumber="437">TURKEY</collectingCountry>
: 
<collectingRegion id="49800493FFF24715FE36F8E2CE9EF880" box="[458,506,1883,1902]" country="Turkey" name="Izmir" pageId="18" pageNumber="437">IZMIR</collectingRegion>
:
</emphasis>
Smyrna? [=Izmir], no further data, 1 ♁ (
<collectionCode id="ED5552B4FFF24715FF0CF8CFCE55F865" box="[240,305,1910,1929]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="18" pageNumber="437">MFNB</collectionCode>
).
</paragraph>
<paragraph id="8BFBCA71FFF24714FF6BF818CD9CFDF9" blockId="18.[151,760,1953,2072]" lastBlockId="19.[151,761,190,2069]" lastPageId="19" lastPageNumber="438" pageId="18" pageNumber="437">
<emphasis id="B9301663FFF24715FF6BF818CEF9F854" bold="true" box="[151,413,1953,1976]" pageId="18" pageNumber="437">Diagnostic description.</emphasis>
This species is externally rather similar to the preceding species and therefore here we provide only the diagnostic description outlining the differences between the two taxa. Body (
<figureCitation id="137FD6F4FFF24715FDD3F7B8CD19F7F4" box="[559,637,2049,2072]" captionStart="Figs 44–45" captionStartId="16.[151,188,2008,2027]" captionTargetBox="[156,1428,1206,1975]" captionTargetId="figure-13@16.[151,1436,1200,1986]" captionTargetPageId="16" captionText="Figs 44–45.Hypocaccus (Nessus) curtus (Rosenhauer, 1847), dorsal habitus. 44 – Saprinus revisus Marseul, 1876, lectotype. 45 – Hypocacculus (Nessus) controversus G. Müller, 1937, lectotype." figureDoi="http://doi.org/10.5281/zenodo.4504946" httpUri="https://zenodo.org/record/4504946/files/figure.png" pageId="18" pageNumber="437">Fig. 45</figureCitation>
) somewhat more round and more flattened, light to dark brown, with light bronze hue (never with greenish hue). PEL: 2.00–2.30 mm; APW: 1.00–1.10 mm; PPW: 1.50–1.70 mm; EW: 1.65–1.90 mm; EL: 1.25–1.50 mm. Frontal disc more finely punctate than the one of 
<taxonomicName id="4C44B1F2FFF24715FB72F9AFCA41F9C1" authorityName="Lackner &amp; Seres" authorityYear="2018" baseAuthorityName="Rosenhauer" baseAuthorityYear="1847" box="[1166,1317,1558,1581]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="438" phylum="Arthropoda" rank="species" species="curtus" subGenus="Nessus">
<emphasis id="B9301663FFF24715FB72F9AFCBC5F9C1" box="[1166,1185,1558,1581]" italics="true" pageId="18" pageNumber="437">H</emphasis>
. (
<emphasis id="B9301663FFF24715FB45F9AFCBAFF9C1" box="[1209,1227,1558,1581]" italics="true" pageId="18" pageNumber="437">N</emphasis>
.) 
<emphasis id="B9301663FFF24715FB1EF9AFCA41F9C1" box="[1250,1317,1558,1581]" italics="true" pageId="18" pageNumber="437">curtus</emphasis>
</taxonomicName>
; pronotum medially almost impunctate. The first dorsal elytral stria is only slightly longer than the second (apically both striae 1–2 surpass slightly elytral half), never reaching ¾ of the elytral length apically (in 
<taxonomicName id="4C44B1F2FFF24715FBB3F92FCBD7F941" authorityName="Lackner &amp; Seres" authorityYear="2018" baseAuthorityName="Rosenhauer" baseAuthorityYear="1847" box="[1103,1203,1686,1709]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="437" phylum="Arthropoda" rank="species" species="curtus">
<emphasis id="B9301663FFF24715FBB3F92FCBD7F941" box="[1103,1203,1686,1709]" italics="true" pageId="18" pageNumber="437">H. curtus</emphasis>
</taxonomicName>
the first dorsal elytral stria is substantially longer, occasionally surpassing ¾ of elytral length apically). Sutural elytral stria always connected basally with fourth dorsal elytral stria (in 
<taxonomicName id="4C44B1F2FFF24715FA7EF94FCC1BF8C1" authorityName="Lackner &amp; Seres" authorityYear="2018" baseAuthorityName="Rosenhauer" baseAuthorityYear="1847" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="437" phylum="Arthropoda" rank="species" species="curtus">
<emphasis id="B9301663FFF24715FA7EF94FCC1BF8C1" italics="true" pageId="18" pageNumber="437">H. curtus</emphasis>
</taxonomicName>
these two striae are joined only in specimens that belong to the ‘ 
<taxonomicName id="4C44B1F2FFF24715FC2BF88FCB17F8A1" authorityName="Jacquelin du Val" authorityYear="1858" box="[983,1139,1846,1869]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="437" phylum="Arthropoda" rank="species" species="cribellaticollis">
<emphasis id="B9301663FFF24715FC2BF88FCB17F8A1" box="[983,1139,1846,1869]" italics="true" pageId="18" pageNumber="437">cribellaticollis</emphasis>
</taxonomicName>
’ form), can occasionally be shortened apically. Carinal prosternal striae strongly convergent apically, their apices very approximate, stopping posterad of united lateral prosternal striae; their united apices not forming a ‘loop’ as in 
<taxonomicName id="4C44B1F2FFF24715FB73F80FCB95F821" authorityName="Lackner &amp; Seres" authorityYear="2018" baseAuthorityName="Rosenhauer" baseAuthorityYear="1847" box="[1167,1265,1974,1997]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="437" phylum="Arthropoda" rank="species" species="curtus">
<emphasis id="B9301663FFF24715FB73F80FCB95F821" box="[1167,1265,1974,1997]" italics="true" pageId="18" pageNumber="437">H. curtus</emphasis>
</taxonomicName>
. 
<bibRefCitation id="EFD5B780FFF24715FB00F80FCAF8F821" author="MULLER G." box="[1276,1436,1974,1997]" pageId="18" pageNumber="437" pagination="97 - 112" refId="ref21522" refString="MULLER G. 1937: Histeriden-Studien. Entomologische Blatter 33: 97 - 112." type="journal article" year="1937">MÜLLER (1937)</bibRefCitation>
mentioned another character: the mesoventral punctation is supposed to be denser and coarser in ‘ 
<taxonomicName id="4C44B1F2FFF24715FAF7F84FCAF1F7E1" baseAuthorityName="G. Muller" baseAuthorityYear="1937" box="[1291,1429,2038,2061]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="437" phylum="Arthropoda" rank="species" species="controversus">
<emphasis id="B9301663FFF24715FAF7F84FCAF1F7E1" box="[1291,1429,2038,2061]" italics="true" pageId="18" pageNumber="437">controversus</emphasis>
</taxonomicName>
’ than in ‘ 
<taxonomicName id="4C44B1F2FFF34714FF0DFF07CE09FF39" baseAuthorityName="Kuster" baseAuthorityYear="1849" box="[241,365,190,213]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="438" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="B9301663FFF34714FF0DFF07CE09FF39" box="[241,365,190,213]" italics="true" pageId="19" pageNumber="438">puncticollis</emphasis>
</taxonomicName>
’ (= 
<taxonomicName id="4C44B1F2FFF34714FE60FF07CD64FF39" authorityName="Lackner &amp; Seres" authorityYear="2018" baseAuthorityName="Rosenhauer" baseAuthorityYear="1847" box="[412,512,190,213]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="438" phylum="Arthropoda" rank="species" species="curtus">
<emphasis id="B9301663FFF34714FE60FF07CD64FF39" box="[412,512,190,213]" italics="true" pageId="19" pageNumber="438">H. curtus</emphasis>
</taxonomicName>
). According to our observations, this is a valid, but not entirely stable character, since even among the few ‘ 
<taxonomicName id="4C44B1F2FFF34714FE34FF47CD35FEF9" baseAuthorityName="G. Muller" baseAuthorityYear="1937" box="[456,593,254,277]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="438" phylum="Arthropoda" rank="species" species="controversus">
<emphasis id="B9301663FFF34714FE34FF47CD35FEF9" box="[456,593,254,277]" italics="true" pageId="19" pageNumber="438">controversus</emphasis>
</taxonomicName>
’ specimens we were able to examine we saw a specimen with only weak mesoventral punctation; the majority of specimens had their mesoventrite densely punctate. Male genitalia (
<figureCitation id="137FD6F4FFF34714FD34FEE7CFBFFE79" captionStart="Figs 55–63" captionStartId="18.[150,187,1309,1328]" captionTargetBox="[294,1285,171,1249]" captionTargetId="figure-468@18.[280,1288,166,1252]" captionTargetPageId="18" captionText="Figs 55–63. Male genitalia of Hypocaccus (Nessus) controversus (Müller, 1937). 55 – VIII sternite and tergite, ventral view; 56 – ditto, dorsal view; 57 – ditto, lateral view; 58 – IX and X tergites, dorsal view; 59 – ditto, lateral view; 60 – IX sternite (spiculum gastrale), ventral view; 61 – ditto, lateral view; 62 – aedeagus, dorsal view; 63 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4504960" httpUri="https://zenodo.org/record/4504960/files/figure.png" pageId="19" pageNumber="438">Figs 55–63</figureCitation>
) are generally similar to the preceding species, the aedeagi differ most markedly: the one of 
<taxonomicName id="4C44B1F2FFF34714FDB0FE27CD84FE59" authorityName="Lackner &amp; Seres" authorityYear="2018" baseAuthorityName="Rosenhauer" baseAuthorityYear="1847" box="[588,736,414,437]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="438" phylum="Arthropoda" rank="species" species="curtus" subGenus="Nessus">
<emphasis id="B9301663FFF34714FDB0FE27CD3BFE59" box="[588,607,414,437]" italics="true" pageId="19" pageNumber="438">H</emphasis>
. (
<emphasis id="B9301663FFF34714FD89FE27CDE3FE59" box="[629,647,414,437]" italics="true" pageId="19" pageNumber="438">N</emphasis>
.) 
<emphasis id="B9301663FFF34714FD61FE27CD84FE59" box="[669,736,414,437]" italics="true" pageId="19" pageNumber="438">curtus</emphasis>
</taxonomicName>
is sub-parallel and blunted apically, while the one of 
<emphasis id="B9301663FFF34714FD51FE07CDA4FE39" box="[685,704,446,469]" italics="true" pageId="19" pageNumber="438">H</emphasis>
. 
<taxonomicName id="4C44B1F2FFF34714FD31FE07CE44FE19" baseAuthorityName="G. Muller" baseAuthorityYear="1937" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="438" phylum="Arthropoda" rank="species" species="controversus" subGenus="Nessus">
(
<emphasis id="B9301663FFF34714FD2AFE07CD8CFE39" box="[726,744,446,469]" italics="true" pageId="19" pageNumber="438">N</emphasis>
.) 
<emphasis id="B9301663FFF34714FF6BFE67CE44FE19" box="[151,288,478,501]" italics="true" pageId="19" pageNumber="438">controversus</emphasis>
</taxonomicName>
is shorter, stouter, slightly dilated in apical third with acutely pointed apex (compare 
<figureCitation id="137FD6F4FFF34714FDB2FE47CDF8FDF9" box="[590,668,510,533]" captionStart="Figs 46–54" captionStartId="17.[151,188,1365,1384]" captionTargetBox="[271,1310,177,1331]" captionTargetId="figure-374@17.[252,1324,170,1345]" captionTargetPageId="17" captionText="Figs 46–54. Male genitalia of Hypocaccus (Nessus) curtus (Rosenhauer, 1847). 45 – VIII sternite and tergite, ventral view; 46 – ditto, dorsal view; 47 – ditto, lateral view; 48 – IX and X tergites, dorsal view; 49 – ditto, lateral view; 50 – IX sternite (spiculum gastrale), ventral view; 51 – ditto, lateral view; 52 – aedeagus, dorsal view; 53 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4504952" httpUri="https://zenodo.org/record/4504952/files/figure.png" pageId="19" pageNumber="438">Figs 53</figureCitation>
and 
<figureCitation id="137FD6F4FFF34714FD31FE47CD8FFDF9" box="[717,747,510,533]" captionStart="Figs 55–63" captionStartId="18.[150,187,1309,1328]" captionTargetBox="[294,1285,171,1249]" captionTargetId="figure-468@18.[280,1288,166,1252]" captionTargetPageId="18" captionText="Figs 55–63. Male genitalia of Hypocaccus (Nessus) controversus (Müller, 1937). 55 – VIII sternite and tergite, ventral view; 56 – ditto, dorsal view; 57 – ditto, lateral view; 58 – IX and X tergites, dorsal view; 59 – ditto, lateral view; 60 – IX sternite (spiculum gastrale), ventral view; 61 – ditto, lateral view; 62 – aedeagus, dorsal view; 63 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4504960" httpUri="https://zenodo.org/record/4504960/files/figure.png" pageId="19" pageNumber="438">62</figureCitation>
).
</paragraph>
<paragraph id="8BFBCA71FFF34714FF6BFDA7CE56FD39" blockId="19.[151,761,190,2069]" pageId="19" pageNumber="438">
<emphasis id="B9301663FFF34714FF6BFDA7CFB7FDD9" bold="true" box="[151,211,542,565]" pageId="19" pageNumber="438">Note.</emphasis>
The two female specimens from 
<collectingCountry id="F3538AE1FFF34714FDDAFDA7CD08FDD9" box="[550,620,542,565]" name="Jordan" pageId="19" pageNumber="438">Jordan</collectingCountry>
as well as the female from 
<collectingCountry id="F3538AE1FFF34714FEE7FD87CE0DFDB9" box="[283,361,574,597]" name="Tunisia" pageId="19" pageNumber="438">Tunisia</collectingCountry>
are generally somewhat narrower, and their frons is adorned with coarse elongate rugae in place of dense punctures that are present at the 
<typeStatus id="54FF74D3FFF34714FDB2FDC7CD18FD79" box="[590,636,638,661]" pageId="19" pageNumber="438">type</typeStatus>
specimens. Therefore we identified these specimens with doubts as 
<taxonomicName id="4C44B1F2FFF34714FF6BFD07CE48FD39" authorityName="Lackner &amp; Seres" authorityYear="2018" baseAuthorityName="Rosenhauer" baseAuthorityYear="1847" box="[151,300,702,725]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="438" phylum="Arthropoda" rank="species" species="curtus" subGenus="Nessus">
<emphasis id="B9301663FFF34714FF6BFD07CFD6FD39" box="[151,178,702,725]" italics="true" pageId="19" pageNumber="438">H.</emphasis>
(
<emphasis id="B9301663FFF34714FF3DFD07CFBEFD39" box="[193,218,702,725]" italics="true" pageId="19" pageNumber="438">N.</emphasis>
) 
<emphasis id="B9301663FFF34714FF15FD07CE48FD39" box="[233,300,702,725]" italics="true" pageId="19" pageNumber="438">curtus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35E99FAFFF34714FF6BFD67CDAAFCB9" pageId="19" pageNumber="438" type="distribution">
<paragraph id="8BFBCA71FFF34714FF6BFD67CDAAFCB9" blockId="19.[151,761,190,2069]" pageId="19" pageNumber="438">
<emphasis id="B9301663FFF34714FF6BFD67CE4EFD19" bold="true" box="[151,298,734,757]" pageId="19" pageNumber="438">Distribution.</emphasis>
<collectingCountry id="F3538AE1FFF34714FECCFD67CEF7FD19" box="[304,403,734,757]" name="Romania" pageId="19" pageNumber="438">Romania</collectingCountry>
, 
<collectingCountry id="F3538AE1FFF34714FE62FD67CE8DFD19" box="[414,489,734,757]" name="Greece" pageId="19" pageNumber="438">Greece</collectingCountry>
(including Zakynthos Island and 
<collectingRegion id="49800493FFF34714FF04FD47CE52FCF9" box="[248,310,766,789]" country="Greece" name="Kriti" pageId="19" pageNumber="438">Crete</collectingRegion>
), 
<collectingCountry id="F3538AE1FFF34714FEBBFD47CEA8FCF9" box="[327,460,766,789]" name="Serbia" pageId="19" pageNumber="438">Montenegro</collectingCountry>
, 
<collectingCountry id="F3538AE1FFF34714FE2AFD47CD71FCF9" box="[470,533,766,789]" name="Spain" pageId="19" pageNumber="438">Spain</collectingCountry>
, 
<collectingCountry id="F3538AE1FFF34714FDE3FD47CDE6FCF9" box="[543,642,766,789]" name="Morocco" pageId="19" pageNumber="438">Morocco</collectingCountry>
, 
<collectingCountry id="F3538AE1FFF34714FD70FD47CDBEFCF9" box="[652,730,766,789]" name="Jordan" pageId="19" pageNumber="438">Jordan</collectingCountry>
(?), 
<collectingCountry id="F3538AE1FFF34714FF6BFCA7CF80FCD9" box="[151,228,798,821]" name="Turkey" pageId="19" pageNumber="438">Turkey</collectingCountry>
, and 
<collectingCountry id="F3538AE1FFF34714FEDCFCA7CEC9FCD9" box="[288,429,798,821]" name="Saudi Arabia" pageId="19" pageNumber="438">Saudi Arabia</collectingCountry>
(
<bibRefCitation id="EFD5B780FFF34714FE41FCA7CDF9FCD9" author="LACKNER T. &amp; MAZUR S. &amp; NEWTON A. F." box="[445,669,798,821]" pageId="19" pageNumber="438" refId="ref21168" refString="LACKNER T., MAZUR S. &amp; NEWTON A. F. 2015: Family Histeridae. Pp. 76 - 130. In: LOBL I. &amp; LOBL D. (eds): Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Staphylinoidea. Volume 1. Brill Publishers, Leiden, Boston, xxiii + 1 - 900." type="book" year="2015">LACKNER et al. 2015</bibRefCitation>
). Newly reported from 
<collectingCountry id="F3538AE1FFF34714FECDFC87CE1AFCB9" box="[305,382,830,853]" name="Cyprus" pageId="19" pageNumber="438">Cyprus</collectingCountry>
and from 
<collectingCountry id="F3538AE1FFF34714FE10FC87CD5FFCB9" box="[492,571,830,853]" name="Tunisia" pageId="19" pageNumber="438">Tunisia</collectingCountry>
(with doubt).
</paragraph>
</subSubSection>
<subSubSection id="C35E99FAFFF34714FF6BFCE7CD9CFC59" pageId="19" pageNumber="438" type="biology_ecology">
<paragraph id="8BFBCA71FFF34714FF6BFCE7CD9CFC59" blockId="19.[151,761,190,2069]" pageId="19" pageNumber="438">
<emphasis id="B9301663FFF34714FF6BFCE7CF95FC99" bold="true" box="[151,241,862,885]" pageId="19" pageNumber="438">Biology.</emphasis>
According to 
<bibRefCitation id="EFD5B780FFF34714FE7EFCE7CD9CFC99" author="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N." box="[386,760,862,885]" pageId="19" pageNumber="438" refId="ref20992" refString="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N. 1976: Zhuki nadsemeystva Histeroidea (semeystva Sphaeritidae, Histeridae, Synteliidae). Fauna SSSR, Zhestokrylye, Tom V, vyp. 4. [Beetles of the superfamily Histeroidea (families Sphaeritidae, Histeridae, Syntelidae) Fauna SSSR, Coleoptera, Vol. 5, part 4.]. Nauka, Leningrad, 434 pp (in Russian)." type="book" year="1976">KRYZHANOVSKIJ &amp; REICHARDT (1976)</bibRefCitation>
this species is found on sandy banks of rivers and seas. The examined specimens did not bear any ecological data.
</paragraph>
</subSubSection>
<subSubSection id="C35E99FAFFF34714FF6BFC07CB55FCF9" pageId="19" pageNumber="438" type="discussion">
<paragraph id="8BFBCA71FFF34714FF6BFC07CB91FE19" blockId="19.[151,761,190,2069]" lastBlockId="19.[828,1437,190,789]" pageId="19" pageNumber="438">
<emphasis id="B9301663FFF34714FF6BFC07CE62FC39" bold="true" box="[151,262,958,981]" pageId="19" pageNumber="438">Remarks.</emphasis>
This species was described from the following localities, but the number of specimens from each locality was not specified: 
<collectingCountry id="F3538AE1FFF34714FE90FC47CEB5FBF9" box="[364,465,1022,1045]" name="Romania" pageId="19" pageNumber="438">Romania</collectingCountry>
, Banat: Băile Herculeane; 
<collectingCountry id="F3538AE1FFF34714FF6BFBA7CF83FBD9" box="[151,231,1054,1077]" name="Greece" pageId="19" pageNumber="438">Greece</collectingCountry>
: Thessaloniki; Phaleron near Athens; Parnass (= Mount Parnassus?), and Zakynthos Island: Kalamaki. We were able to examine the specimens from 
<collectingCountry id="F3538AE1FFF34714FDB3FBE7CDD6FB99" box="[591,690,1118,1141]" name="Romania" pageId="19" pageNumber="438">Romania</collectingCountry>
, Thessaloniki, Zakynthos Island: Kalamaki and Phaleron near Athens. We chose the best-preserved female specimen from 
<collectingCountry id="F3538AE1FFF34714FF6BFB07CF93FB39" box="[151,247,1214,1237]" name="Romania" pageId="19" pageNumber="438">Romania</collectingCountry>
as the 
<typeStatus id="54FF74D3FFF34714FEBFFB07CECCFB39" box="[323,424,1214,1237]" pageId="19" pageNumber="438" type="lectotype">lectotype</typeStatus>
, since it was the only specimen bearing a “type” label and the remaining three specimens as the 
<typeStatus id="54FF74D3FFF34714FF21FB47CE1EFAF9" box="[221,378,1278,1301]" pageId="19" pageNumber="438" type="paralectotype">paralectotypes</typeStatus>
, respectively. The remaining specimen(s) from Parnass (= Mount Parnassus?) should qualify as 
<typeStatus id="54FF74D3FFF34714FF4FFA87CE2EFAB9" box="[179,330,1342,1365]" pageId="19" pageNumber="438" type="paralectotype">paralectotype</typeStatus>
(s), but their depository is unknown to us. The specimen from Zakynthos Island: Kalamaki was labelled by the MFNB staff as ‘ 
<taxonomicName id="4C44B1F2FFF34714FDFBFAC7CFBEFA59" authority="Payk." authorityName="Payk." class="Insecta" family="Histeridae" genus="Hypocacculus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="438" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="B9301663FFF34714FDFBFAC7CD93FA79" box="[519,759,1406,1429]" italics="true" pageId="19" pageNumber="438">Hypocacculus rufipes</emphasis>
Payk.
</taxonomicName>
’, since it was placed among other specimens of 
<emphasis id="B9301663FFF34714FD22FA27CD95FA59" box="[734,753,1438,1461]" italics="true" pageId="19" pageNumber="438">H</emphasis>
. 
<taxonomicName id="4C44B1F2FFF34714FF6BFA07CE70FA39" box="[151,276,1470,1493]" class="Insecta" family="Histeridae" genus="Hypocacculus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="438" phylum="Arthropoda" rank="species" species="rufipes" subGenus="Nessus">
(
<emphasis id="B9301663FFF34714FF5CFA07CFD6FA39" box="[160,178,1470,1493]" italics="true" pageId="19" pageNumber="438">N</emphasis>
.) 
<emphasis id="B9301663FFF34714FF37FA07CE70FA39" box="[203,276,1470,1493]" italics="true" pageId="19" pageNumber="438">rufipes</emphasis>
</taxonomicName>
originating from the collections of Schmidt and Bickhardt. Although 
<bibRefCitation id="EFD5B780FFF34714FE56FA67CD2AFA19" author="MULLER G." box="[426,590,1502,1525]" pageId="19" pageNumber="438" pagination="97 - 112" refId="ref21522" refString="MULLER G. 1937: Histeriden-Studien. Entomologische Blatter 33: 97 - 112." type="journal article" year="1937">MÜLLER (1937)</bibRefCitation>
mentioned that he examined specimen(s) from Zakynthos Island, we cannot be sure that this very specimen can be attributed a 
<typeStatus id="54FF74D3FFF34714FF6BF987CE42F9B9" box="[151,294,1598,1621]" pageId="19" pageNumber="438" type="paralectotype">paralectotype</typeStatus>
for the following reasons: 
<bibRefCitation id="EFD5B780FFF34714FDC1F987CFACF999" author="REICHARDT A." pageId="19" pageNumber="438" pagination="1 - 164" refId="ref21636" refString="REICHARDT A. 1932: Beitrage zur einer Monographie der Saprininae (Coleoptera, Histeridae). Mitteilungen aus dem Zoologischen Museum in Berlin 18: 1 - 164." type="journal article" year="1932">REICHARDT (1932: 124)</bibRefCitation>
already mentioned a series of five specimens with the same label data (
<collectingCountry id="F3538AE1FFF34714FE8AF9C7CEA2F979" box="[374,454,1662,1685]" name="Greece" pageId="19" pageNumber="438">Greece</collectingCountry>
, Zante [=Zakynthos] 
<collectingCountry id="F3538AE1FFF34714FD4CF9C7CD90F979" box="[688,756,1662,1685]" name="Iceland" pageId="19" pageNumber="438">Island</collectingCountry>
, Kalamaki, 1909, Hilf lgt., coll. Leonhard); at least one of these specimens was examined also by Müller. According to 
<bibRefCitation id="EFD5B780FFF34714FF4CF967CE0EF919" author="REICHARDT A." box="[176,362,1758,1781]" pageId="19" pageNumber="438" pagination="1 - 164" refId="ref21636" refString="REICHARDT A. 1932: Beitrage zur einer Monographie der Saprininae (Coleoptera, Histeridae). Mitteilungen aus dem Zoologischen Museum in Berlin 18: 1 - 164." type="journal article" year="1932">REICHARDT (1932)</bibRefCitation>
, two of these were identified by Müller as 
<emphasis id="B9301663FFF34714FF49F947CFACF8F9" box="[181,200,1790,1813]" italics="true" pageId="19" pageNumber="438">H</emphasis>
. 
<taxonomicName id="4C44B1F2FFF34714FF2AF947CEE0F8F9" baseAuthorityName="Kuster" baseAuthorityYear="1849" box="[214,388,1790,1813]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="438" phylum="Arthropoda" rank="species" species="puncticollis" subGenus="Nessus">
(
<emphasis id="B9301663FFF34714FF23F947CF95F8F9" box="[223,241,1790,1813]" italics="true" pageId="19" pageNumber="438">N</emphasis>
.) 
<emphasis id="B9301663FFF34714FEF4F947CEE0F8F9" box="[264,388,1790,1813]" italics="true" pageId="19" pageNumber="438">puncticollis</emphasis>
</taxonomicName>
(= 
<taxonomicName id="4C44B1F2FFF34714FE57F947CD6BF8F9" authorityName="Lackner &amp; Seres" authorityYear="2018" baseAuthorityName="Rosenhauer" baseAuthorityYear="1847" box="[427,527,1790,1813]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="438" phylum="Arthropoda" rank="species" species="curtus">
<emphasis id="B9301663FFF34714FE57F947CD6BF8F9" box="[427,527,1790,1813]" italics="true" pageId="19" pageNumber="438">H. curtus</emphasis>
</taxonomicName>
); two were deposited in Schmidt’s collection and identified as 
<emphasis id="B9301663FFF34714FDA1F8A7CD14F8D9" box="[605,624,1822,1845]" italics="true" pageId="19" pageNumber="438">H</emphasis>
. 
<taxonomicName id="4C44B1F2FFF34714FD7DF8A7CEEBF8B9" authority="Schmidt" authorityName="Schmidt" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="438" phylum="Arthropoda" rank="variety" species="curtus" subGenus="Nessus" variety="aenescens">
(
<emphasis id="B9301663FFF34714FD76F8A7CDF8F8D9" box="[650,668,1822,1845]" italics="true" pageId="19" pageNumber="438">N</emphasis>
.) 
<emphasis id="B9301663FFF34714FD49F8A7CD9CF8D9" box="[693,760,1822,1845]" italics="true" pageId="19" pageNumber="438">curtus</emphasis>
var. 
<emphasis id="B9301663FFF34714FF38F887CE54F8B9" box="[196,304,1854,1877]" italics="true" pageId="19" pageNumber="438">aenescens</emphasis>
Schmidt
</taxonomicName>
in litt.; and a single specimen was deposited in Schmidt’s collection and identified as 
<emphasis id="B9301663FFF34714FD52F8E7CDA5F899" box="[686,705,1886,1909]" italics="true" pageId="19" pageNumber="438">H</emphasis>
. 
<taxonomicName id="4C44B1F2FFF34714FD31F8E7CE29F879" authority="Payk. The" authorityName="Payk. The" class="Insecta" family="Histeridae" genus="Hypocacculus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="438" phylum="Arthropoda" rank="species" species="rufipes" subGenus="Nessus">
(
<emphasis id="B9301663FFF34714FD2AF8E7CD8CF899" box="[726,744,1886,1909]" italics="true" pageId="19" pageNumber="438">N</emphasis>
.) 
<emphasis id="B9301663FFF34714FF6BF8C7CFBBF879" box="[151,223,1918,1941]" italics="true" pageId="19" pageNumber="438">rufipes</emphasis>
Payk. The
</taxonomicName>
five specimens were supposedly divided between MFNB and Deutsches Entomologisches Institut, Müncheberg, 
<collectingCountry id="F3538AE1FFF34714FEC9F807CEFAF839" box="[309,414,1982,2005]" name="Germany" pageId="19" pageNumber="438">Germany</collectingCountry>
. According to Reichardt, who examined the whole lot, the five specimens doubtlessly belonged to the same species, albeit Reichardt was not sure to which, and placed them as ‘near to 
<taxonomicName id="4C44B1F2FFF34714FB2CFF07CA7DFF39" box="[1232,1305,190,213]" class="Insecta" family="Histeridae" genus="Hypocacculus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="438" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="B9301663FFF34714FB2CFF07CA7DFF39" box="[1232,1305,190,213]" italics="true" pageId="19" pageNumber="438">rufipes</emphasis>
</taxonomicName>
or transitional forms between 
<taxonomicName id="4C44B1F2FFF34714FBF5FF67CB36FF19" box="[1033,1106,222,245]" class="Insecta" family="Histeridae" genus="Hypocacculus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="438" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="B9301663FFF34714FBF5FF67CB36FF19" box="[1033,1106,222,245]" italics="true" pageId="19" pageNumber="438">rufipes</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C44B1F2FFF34714FB7BFF67CBAEFF19" authorityName="Lackner &amp; Seres" authorityYear="2018" baseAuthorityName="Rosenhauer" baseAuthorityYear="1847" box="[1159,1226,222,245]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="438" phylum="Arthropoda" rank="species" species="curtus">
<emphasis id="B9301663FFF34714FB7BFF67CBAEFF19" box="[1159,1226,222,245]" italics="true" pageId="19" pageNumber="438">curtus</emphasis>
</taxonomicName>
, or even hybrids of the two species’. The specimen from Zakynthos we were able to examine is most likely the one that was identified as ‘ 
<taxonomicName id="4C44B1F2FFF34714FC98FE87CCC9FEB9" box="[868,941,318,341]" class="Insecta" family="Histeridae" genus="Hypocacculus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="438" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="B9301663FFF34714FC98FE87CCC9FEB9" box="[868,941,318,341]" italics="true" pageId="19" pageNumber="438">rufipes</emphasis>
</taxonomicName>
’ in Schmidt’s collection, currently housed in MFNB and therefore probably not examined by Müller, when he described 
<emphasis id="B9301663FFF34714FBF4FEC7CB7FFE79" box="[1032,1051,382,405]" italics="true" pageId="19" pageNumber="438">H</emphasis>
. 
<taxonomicName id="4C44B1F2FFF34714FBD4FEC7CB85FE79" baseAuthorityName="G. Muller" baseAuthorityYear="1937" box="[1064,1249,382,405]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="438" phylum="Arthropoda" rank="species" species="controversus" subGenus="Nessus">
(
<emphasis id="B9301663FFF34714FBCDFEC7CB27FE79" box="[1073,1091,382,405]" italics="true" pageId="19" pageNumber="438">N</emphasis>
.) 
<emphasis id="B9301663FFF34714FBA4FEC7CB85FE79" box="[1112,1249,382,405]" italics="true" pageId="19" pageNumber="438">controversus</emphasis>
</taxonomicName>
. Albeit the specimen cannot be ascribed a 
<typeStatus id="54FF74D3FFF34714FBABFE27CB83FE59" box="[1111,1255,414,437]" pageId="19" pageNumber="438" type="paralectotype">paralectotype</typeStatus>
status this is the only male specimen of 
<emphasis id="B9301663FFF34714FBCDFE07CB20FE39" box="[1073,1092,446,469]" italics="true" pageId="19" pageNumber="438">H</emphasis>
. 
<taxonomicName id="4C44B1F2FFF34714FBADFE07CA6EFE39" baseAuthorityName="G. Muller" baseAuthorityYear="1937" box="[1105,1290,446,469]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="438" phylum="Arthropoda" rank="species" species="controversus" subGenus="Nessus">
(
<emphasis id="B9301663FFF34714FBA6FE07CB08FE39" box="[1114,1132,446,469]" italics="true" pageId="19" pageNumber="438">N</emphasis>
.) 
<emphasis id="B9301663FFF34714FB7DFE07CA6EFE39" box="[1153,1290,446,469]" italics="true" pageId="19" pageNumber="438">controversus</emphasis>
</taxonomicName>
we have seen and we therefore depict its genitalia here.
</paragraph>
<paragraph id="8BFBCA71FFF34714FCA6FE47CB55FCF9" blockId="19.[828,1437,190,789]" pageId="19" pageNumber="438">
<bibRefCitation id="EFD5B780FFF34714FCA6FE47CB75FDF9" author="REICHARDT A." box="[858,1041,510,533]" pageId="19" pageNumber="438" pagination="1 - 164" refId="ref21636" refString="REICHARDT A. 1932: Beitrage zur einer Monographie der Saprininae (Coleoptera, Histeridae). Mitteilungen aus dem Zoologischen Museum in Berlin 18: 1 - 164." type="journal article" year="1932">REICHARDT (1932)</bibRefCitation>
expressed his frustration with a couple of specimens of 
<taxonomicName id="4C44B1F2FFF34714FC11FDA7CA78FDD9" authorityName="Lackner &amp; Seres" authorityYear="2018" baseAuthorityName="Rosenhauer" baseAuthorityYear="1847" box="[1005,1308,542,565]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="438" phylum="Arthropoda" rank="species" species="curtus" subGenus="Nessus">
<emphasis id="B9301663FFF34714FC11FDA7CB0BFDD9" box="[1005,1135,542,565]" italics="true" pageId="19" pageNumber="438">Hypocaccus</emphasis>
(
<emphasis id="B9301663FFF34714FB83FDA7CBADFDD9" box="[1151,1225,542,565]" italics="true" pageId="19" pageNumber="438">Nessus</emphasis>
) 
<emphasis id="B9301663FFF34714FB25FDA7CA78FDD9" box="[1241,1308,542,565]" italics="true" pageId="19" pageNumber="438">curtus</emphasis>
</taxonomicName>
from North Africa, which look externally as 
<taxonomicName id="4C44B1F2FFF34714FB4AFD87CA0BFDB9" authorityName="Marseul" authorityYear="1876" box="[1206,1391,574,597]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="438" phylum="Arthropoda" rank="species" species="revisus">
<emphasis id="B9301663FFF34714FB4AFD87CA0BFDB9" box="[1206,1391,574,597]" italics="true" pageId="19" pageNumber="438">Saprinus revisus</emphasis>
</taxonomicName>
but the aedeagus is different and similar to another species, 
<taxonomicName id="4C44B1F2FFF34714FCC0FDC7CAF8FD79" authority="(Peyerimhoff, 1917)" baseAuthorityName="Peyerimhoff" baseAuthorityYear="1917" box="[828,1436,638,661]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="438" phylum="Arthropoda" rank="species" species="emendatus" subGenus="Nessus">
<emphasis id="B9301663FFF34714FCC0FDC7CCA6FD79" box="[828,962,638,661]" italics="true" pageId="19" pageNumber="438">Hypocaccus</emphasis>
(
<emphasis id="B9301663FFF34714FC2AFDC7CB46FD79" box="[982,1058,638,661]" italics="true" pageId="19" pageNumber="438">Nessus</emphasis>
) 
<emphasis id="B9301663FFF34714FBCBFDC7CBC9FD79" box="[1079,1197,638,661]" italics="true" pageId="19" pageNumber="438">emendatus</emphasis>
(Peyerimhoff, 1917)
</taxonomicName>
occurring in 
<collectingCountry id="F3538AE1FFF34714FC3DFD27CB70FD59" box="[961,1044,670,693]" name="Algeria" pageId="19" pageNumber="438">Algeria</collectingCountry>
, 
<collectingCountry id="F3538AE1FFF34714FBE2FD27CB3BFD59" box="[1054,1119,670,693]" name="Libya" pageId="19" pageNumber="438">Libya</collectingCountry>
, 
<collectingCountry id="F3538AE1FFF34714FB94FD27CBDEFD59" box="[1128,1210,670,693]" name="Tunisia" pageId="19" pageNumber="438">Tunisia</collectingCountry>
, and 
<collectingCountry id="F3538AE1FFF34714FB0DFD27CA55FD59" box="[1265,1329,670,693]" name="Egypt" pageId="19" pageNumber="438">Egypt</collectingCountry>
(LACKNER et. al. 2015). Without examination of the 
<typeStatus id="54FF74D3FFF34714FB00FD07CA4EFD39" box="[1276,1322,702,725]" pageId="19" pageNumber="438">type</typeStatus>
of 
<emphasis id="B9301663FFF34714FAACFD07CA07FD39" box="[1360,1379,702,725]" italics="true" pageId="19" pageNumber="438">H</emphasis>
. 
<taxonomicName id="4C44B1F2FFF34714FA8EFD07CCCAFD19" baseAuthorityName="Peyerimhoff" baseAuthorityYear="1917" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="438" phylum="Arthropoda" rank="species" species="emendatus" subGenus="Nessus">
(
<emphasis id="B9301663FFF34714FA87FD07CAE9FD39" box="[1403,1421,702,725]" italics="true" pageId="19" pageNumber="438">N</emphasis>
.) 
<emphasis id="B9301663FFF34714FCC0FD67CCCAFD19" box="[828,942,734,757]" italics="true" pageId="19" pageNumber="438">emendatus</emphasis>
</taxonomicName>
, we are unable to solve this riddle and opt for keeping the status quo.
</paragraph>
</subSubSection>
</treatment>
</document>