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<document id="6ED96E68FB13D706E5A87725D812255A" ID-DOI="10.11646/zootaxa.5315.6.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="8142492" ID-ZooBank="DCD0C570-9808-4198-9998-BBAFD4764982" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1689237537405" checkinUser="plazi" docAuthor="Hernando, Carles" docDate="2023" docId="03CCB05101484210FF6AC2884236FEE4" docLanguage="en" docName="zootaxa.5315.6.3.pdf" docOrigin="Zootaxa 5315 (6)" docSource="http://dx.doi.org/10.11646/zootaxa.5315.6.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Austramastodus Hernando 2023, gen. nov." docType="treatment" docVersion="2" lastPageNumber="561" masterDocId="FFF5C82901494213FFFDC21F432DFFA5" masterDocTitle="A new flightless genus of Thaumastodinae from Australia (Coleoptera: Limnichidae)" masterLastPageNumber="566" masterPageNumber="559" pageNumber="560" updateTime="1689237937808" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="AD1AE14145290E210C1AE16441CC086F">A new flightless genus of Thaumastodinae from Australia (Coleoptera: Limnichidae)</mods:title>
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<mods:namePart id="17C55DC34BF2CE5A262B5C1528263CC3">Hernando, Carles</mods:namePart>
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<mods:date id="7B6ECF3BBF405AAD3E4EBB9A2A12EADD">2023</mods:date>
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<mods:number id="D2EC54CF85C509678EE995A1CF18529A">2023-07-13</mods:number>
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<treatment id="03CCB05101484210FF6AC2884236FEE4" ID-DOI="http://doi.org/10.5281/zenodo.8142488" ID-Zenodo-Dep="8142488" LSID="urn:lsid:plazi:treatment:03CCB05101484210FF6AC2884236FEE4" httpUri="http://treatment.plazi.org/id/03CCB05101484210FF6AC2884236FEE4" lastPageId="3" lastPageNumber="561" pageId="1" pageNumber="560">
<subSubSection id="C37F52CC01484212FF6AC28842FDFF14" box="[151,464,151,178]" pageId="1" pageNumber="560" type="nomenclature">
<paragraph id="8BDA014701484212FF6AC28842FDFF14" blockId="1.[151,464,151,178]" box="[151,464,151,178]" pageId="1" pageNumber="560">
<heading id="D092B62B01484212FF6AC28842FDFF14" bold="true" box="[151,464,151,178]" fontSize="11" level="1" pageId="1" pageNumber="560" reason="1">
<emphasis id="B911DD5501484212FF6AC28842FDFF14" bold="true" box="[151,464,151,178]" pageId="1" pageNumber="560">
<taxonomicName id="4C657AC401484212FF6AC2884276FF17" authority="Hernando, 2023" authorityName="Hernando" authorityYear="2023" box="[151,347,151,178]" class="Insecta" family="Limnichidae" genus="Austramastodus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="560" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B911DD5501484212FF6AC2884276FF17" bold="true" box="[151,347,151,178]" italics="true" pageId="1" pageNumber="560">Austramastodus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A222602E01484212FE9FC28842FDFF14" box="[354,464,151,177]" pageId="1" pageNumber="560" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37F52CC01484212FF6AC2C3415FFF51" box="[151,626,220,244]" pageId="1" pageNumber="560" type="materials_examined">
<paragraph id="8BDA014701484212FF6AC2C3415FFF51" blockId="1.[151,626,220,244]" box="[151,626,220,244]" pageId="1" pageNumber="560">
<typeStatus id="54DEBFE501484212FF6AC2C343E4FF51" box="[151,201,220,244]" pageId="1" pageNumber="560">Type</typeStatus>
species.
<taxonomicName id="4C657AC401484212FEDBC2C34136FF56" authority="Hernando, 2023" authorityName="Hernando" authorityYear="2023" box="[294,539,220,244]" class="Insecta" family="Limnichidae" genus="Austramastodus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="560" phylum="Arthropoda" rank="species" species="apterus" status="sp. nov.">
<emphasis id="B911DD5501484212FEDBC2C34136FF56" box="[294,539,220,244]" italics="true" pageId="1" pageNumber="560">Austramastodus apterus</emphasis>
</taxonomicName>
<emphasis id="B911DD5501484212FDDFC2C3415FFF51" bold="true" box="[546,626,220,244]" pageId="1" pageNumber="560">
<taxonomicNameLabel id="A222602E01484212FDDFC2C3415FFF51" box="[546,626,220,244]" pageId="1" pageNumber="560" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C37F52CC01484212FF6AC30247EEFE25" pageId="1" pageNumber="560" type="diagnosis">
<paragraph id="8BDA014701484212FF6AC30247EEFE25" blockId="1.[151,1437,285,2040]" pageId="1" pageNumber="560">
<emphasis id="B911DD5501484212FF6AC302423FFE92" bold="true" box="[151,274,285,311]" pageId="1" pageNumber="560">Diagnosis.</emphasis>
<taxonomicName id="4C657AC401484212FEE7C30142E3FE9D" authority="Hernando, 2023" authorityName="Hernando" authorityYear="2023" box="[282,462,286,312]" class="Insecta" family="Limnichidae" genus="Austramastodus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="560" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B911DD5501484212FEE7C30142E3FE9D" box="[282,462,286,312]" italics="true" pageId="1" pageNumber="560">Austramastodus</emphasis>
</taxonomicName>
<emphasis id="B911DD5501484212FE2AC301416FFE9D" bold="true" box="[471,578,286,312]" pageId="1" pageNumber="560">
<taxonomicNameLabel id="A222602E01484212FE2AC301416FFE9D" box="[471,578,286,312]" pageId="1" pageNumber="560" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
differs from the other members of
<taxonomicName id="4C657AC401484212FC2AC30147BDFE9D" authorityName="Champion" authorityYear="1924" box="[983,1168,286,312]" pageId="1" pageNumber="563" rank="subFamily" subFamily="Thaumastodinae">Thaumastodinae</taxonomicName>
with the tarsal formula 444, in the narrow elongate body shape, transverse head with the frons not projecting forward, widely separated eyes (space between them being larger than an eye diameter) and the weakly sinuate pronotum.
</paragraph>
</subSubSection>
<subSubSection id="C37F52CC01484212FF3AC39647B9F910" pageId="1" pageNumber="560" type="description">
<paragraph id="8BDA014701484212FF3AC396420DFE49" blockId="1.[151,1437,285,2040]" pageId="1" pageNumber="560">
<emphasis id="B911DD5501484212FF3AC3964274FE06" bold="true" box="[199,345,393,419]" pageId="1" pageNumber="560">Description.</emphasis>
Length:
<quantity id="4C9DACA201484212FE39C3954144FE01" box="[452,617,394,421]" metricMagnitude="-3" metricUnit="m" metricValue="2.33" metricValueMax="2.5" metricValueMin="2.16" pageId="1" pageNumber="560" unit="mm" value="2.33" valueMax="2.5" valueMin="2.16">2.162.50 mm</quantity>
(head included). Integument unicolored dark brown. Body form (
<figureCitation id="135E1DC201484212FA99C39543ECFE6D" captionStart="FIGURES 17" captionStartId="2.[151,264,1757,1781]" captionTargetBox="[183,1405,232,1733]" captionTargetId="figure-22@2.[183,1405,181,1733]" captionTargetPageId="2" captionText="FIGURES 17. Austramastodus apterus sp. nov. 1. Male habitus, dorsal view. 2. Male habitus, ventral view. 3. Male habitus, lateral view. 4. Head and pronotum, fronto-dorsal view. 5. Head, fronto-lateral view and male fore tarsi. 6. Male last abdominal sternite. 7. Female last abdominal sternite. The arrow points to the only two large lateral spiniform setae on the female." figureDoi="http://doi.org/10.5281/zenodo.8142494" httpUri="https://zenodo.org/record/8142494/files/figure.png" pageId="1" pageNumber="560">Figs. 13</figureCitation>
) elongate-oval, dorsal surface covered with short and recumbent silvery pubescence and spots of golden pubescence.
</paragraph>
<paragraph id="8BDA014701484212FF3AC3E940CCFD4D" blockId="1.[151,1437,285,2040]" pageId="1" pageNumber="560">
<emphasis id="B911DD5501484212FF3AC3E9423DFDB5" bold="true" box="[199,272,502,528]" pageId="1" pageNumber="560">Head:</emphasis>
Strongly hypognathous (
<figureCitation id="135E1DC201484212FDC1C3E941E7FDB4" box="[572,714,502,529]" captionStart="FIGURES 17" captionStartId="2.[151,264,1757,1781]" captionTargetBox="[183,1405,232,1733]" captionTargetId="figure-22@2.[183,1405,181,1733]" captionTargetPageId="2" captionText="FIGURES 17. Austramastodus apterus sp. nov. 1. Male habitus, dorsal view. 2. Male habitus, ventral view. 3. Male habitus, lateral view. 4. Head and pronotum, fronto-dorsal view. 5. Head, fronto-lateral view and male fore tarsi. 6. Male last abdominal sternite. 7. Female last abdominal sternite. The arrow points to the only two large lateral spiniform setae on the female." figureDoi="http://doi.org/10.5281/zenodo.8142494" httpUri="https://zenodo.org/record/8142494/files/figure.png" pageId="1" pageNumber="560">Figs. 145</figureCitation>
), transverse; frons broad, not projecting forward; eyes large, subcircular and slightly convex (
<figureCitation id="135E1DC201484212FDE8C00541AAFD90" box="[533,647,538,565]" captionStart="FIGURES 17" captionStartId="2.[151,264,1757,1781]" captionTargetBox="[183,1405,232,1733]" captionTargetId="figure-22@2.[183,1405,181,1733]" captionTargetPageId="2" captionText="FIGURES 17. Austramastodus apterus sp. nov. 1. Male habitus, dorsal view. 2. Male habitus, ventral view. 3. Male habitus, lateral view. 4. Head and pronotum, fronto-dorsal view. 5. Head, fronto-lateral view and male fore tarsi. 6. Male last abdominal sternite. 7. Female last abdominal sternite. The arrow points to the only two large lateral spiniform setae on the female." figureDoi="http://doi.org/10.5281/zenodo.8142494" httpUri="https://zenodo.org/record/8142494/files/figure.png" pageId="1" pageNumber="560">Figs. 45</figureCitation>
); frontoclypeal suture straight, shallow and inconspicuous; antennal insertion exposed, laterally contiguous the frontoclypeal suture (
<figureCitation id="135E1DC201484212FC75C02140FCFDFC" box="[904,977,574,601]" captionStart="FIGURES 17" captionStartId="2.[151,264,1757,1781]" captionTargetBox="[183,1405,232,1733]" captionTargetId="figure-22@2.[183,1405,181,1733]" captionTargetPageId="2" captionText="FIGURES 17. Austramastodus apterus sp. nov. 1. Male habitus, dorsal view. 2. Male habitus, ventral view. 3. Male habitus, lateral view. 4. Head and pronotum, fronto-dorsal view. 5. Head, fronto-lateral view and male fore tarsi. 6. Male last abdominal sternite. 7. Female last abdominal sternite. The arrow points to the only two large lateral spiniform setae on the female." figureDoi="http://doi.org/10.5281/zenodo.8142494" httpUri="https://zenodo.org/record/8142494/files/figure.png" pageId="1" pageNumber="560">Fig. 5</figureCitation>
); antennal groove present and shallow, where only the first six antennomeres fit (
<figureCitation id="135E1DC201484212FD84C07D4192FDD8" box="[633,703,610,637]" captionStart="FIGURES 17" captionStartId="2.[151,264,1757,1781]" captionTargetBox="[183,1405,232,1733]" captionTargetId="figure-22@2.[183,1405,181,1733]" captionTargetPageId="2" captionText="FIGURES 17. Austramastodus apterus sp. nov. 1. Male habitus, dorsal view. 2. Male habitus, ventral view. 3. Male habitus, lateral view. 4. Head and pronotum, fronto-dorsal view. 5. Head, fronto-lateral view and male fore tarsi. 6. Male last abdominal sternite. 7. Female last abdominal sternite. The arrow points to the only two large lateral spiniform setae on the female." figureDoi="http://doi.org/10.5281/zenodo.8142494" httpUri="https://zenodo.org/record/8142494/files/figure.png" pageId="1" pageNumber="560">Fig. 5</figureCitation>
). Antenna 11-segmented, short and stout, with 4-segmented and slightly asymmetrical antennal club; scape and pedicel enlarged, antennomere 3 stout, antennomeres 47 narrow, antennomeres 810 dilated and gradually shortened. Labrum subquadrate (
<figureCitation id="135E1DC201484212FC10C0B5471EFD60" box="[1005,1075,682,709]" captionStart="FIGURES 17" captionStartId="2.[151,264,1757,1781]" captionTargetBox="[183,1405,232,1733]" captionTargetId="figure-22@2.[183,1405,181,1733]" captionTargetPageId="2" captionText="FIGURES 17. Austramastodus apterus sp. nov. 1. Male habitus, dorsal view. 2. Male habitus, ventral view. 3. Male habitus, lateral view. 4. Head and pronotum, fronto-dorsal view. 5. Head, fronto-lateral view and male fore tarsi. 6. Male last abdominal sternite. 7. Female last abdominal sternite. The arrow points to the only two large lateral spiniform setae on the female." figureDoi="http://doi.org/10.5281/zenodo.8142494" httpUri="https://zenodo.org/record/8142494/files/figure.png" pageId="1" pageNumber="560">Fig. 5</figureCitation>
). Maxillary palps 4-segmented, last palpomere enlarged and apically acuminate. Labial palps 3-segmented.
</paragraph>
<paragraph id="8BDA014701484212FF3AC0ED4797FAE9" blockId="1.[151,1437,285,2040]" pageId="1" pageNumber="560">
<emphasis id="B911DD5501484212FF3AC0ED4204FCA9" bold="true" box="[199,297,754,780]" pageId="1" pageNumber="560">Thorax:</emphasis>
Pronotum (
<figureCitation id="135E1DC201484212FE49C0ED410CFCA9" box="[436,545,754,780]" captionStart="FIGURES 17" captionStartId="2.[151,264,1757,1781]" captionTargetBox="[183,1405,232,1733]" captionTargetId="figure-22@2.[183,1405,181,1733]" captionTargetPageId="2" captionText="FIGURES 17. Austramastodus apterus sp. nov. 1. Male habitus, dorsal view. 2. Male habitus, ventral view. 3. Male habitus, lateral view. 4. Head and pronotum, fronto-dorsal view. 5. Head, fronto-lateral view and male fore tarsi. 6. Male last abdominal sternite. 7. Female last abdominal sternite. The arrow points to the only two large lateral spiniform setae on the female." figureDoi="http://doi.org/10.5281/zenodo.8142494" httpUri="https://zenodo.org/record/8142494/files/figure.png" pageId="1" pageNumber="560">Figs. 14</figureCitation>
) transverse, about twice as wide as long; lateral margins practically straight, slightly narrowing medially, posterior margin slightly sinuate (
<figureCitation id="135E1DC201484212FC8FC1094091FC95" box="[882,956,790,816]" captionStart="FIGURES 17" captionStartId="2.[151,264,1757,1781]" captionTargetBox="[183,1405,232,1733]" captionTargetId="figure-22@2.[183,1405,181,1733]" captionTargetPageId="2" captionText="FIGURES 17. Austramastodus apterus sp. nov. 1. Male habitus, dorsal view. 2. Male habitus, ventral view. 3. Male habitus, lateral view. 4. Head and pronotum, fronto-dorsal view. 5. Head, fronto-lateral view and male fore tarsi. 6. Male last abdominal sternite. 7. Female last abdominal sternite. The arrow points to the only two large lateral spiniform setae on the female." figureDoi="http://doi.org/10.5281/zenodo.8142494" httpUri="https://zenodo.org/record/8142494/files/figure.png" pageId="1" pageNumber="560">Fig. 4</figureCitation>
); anterior angles broadly acute, posterior angles rounded; disc with coarse and somewhat rough punctation, covered with dense and short golden and silvery setae (
<figureCitation id="135E1DC201484212FF22C1414208FCDD" box="[223,293,862,888]" captionStart="FIGURES 17" captionStartId="2.[151,264,1757,1781]" captionTargetBox="[183,1405,232,1733]" captionTargetId="figure-22@2.[183,1405,181,1733]" captionTargetPageId="2" captionText="FIGURES 17. Austramastodus apterus sp. nov. 1. Male habitus, dorsal view. 2. Male habitus, ventral view. 3. Male habitus, lateral view. 4. Head and pronotum, fronto-dorsal view. 5. Head, fronto-lateral view and male fore tarsi. 6. Male last abdominal sternite. 7. Female last abdominal sternite. The arrow points to the only two large lateral spiniform setae on the female." figureDoi="http://doi.org/10.5281/zenodo.8142494" httpUri="https://zenodo.org/record/8142494/files/figure.png" pageId="1" pageNumber="560">Fig. 2</figureCitation>
). Hypomeron broad and slightly depressed, without suture or carina, covered with short silvery setae. Prosternum: area before procoxae about 0.7 times as long as coxal cavities; prosternal process slightly longer than wide, flat and finely bordered laterally, apically bluntly pointed. Scutellum small and triangular. Elytra very long, twice as long as wide (
<figureCitation id="135E1DC201484212FE5DC1D542C9FC41" box="[416,484,970,996]" captionStart="FIGURES 17" captionStartId="2.[151,264,1757,1781]" captionTargetBox="[183,1405,232,1733]" captionTargetId="figure-22@2.[183,1405,181,1733]" captionTargetPageId="2" captionText="FIGURES 17. Austramastodus apterus sp. nov. 1. Male habitus, dorsal view. 2. Male habitus, ventral view. 3. Male habitus, lateral view. 4. Head and pronotum, fronto-dorsal view. 5. Head, fronto-lateral view and male fore tarsi. 6. Male last abdominal sternite. 7. Female last abdominal sternite. The arrow points to the only two large lateral spiniform setae on the female." figureDoi="http://doi.org/10.5281/zenodo.8142494" httpUri="https://zenodo.org/record/8142494/files/figure.png" pageId="1" pageNumber="560">Fig. 1</figureCitation>
); lateral margins finely beaded along their entire length; apical interlocking device connecting elytra and last sternite, well developed; surface of disc shiny, densely and finely punctate, covered with recumbent, generally silvery setae, but in some areas with golden spots; epipleuron broad at base and slightly impressed to receive femoral tips, extending to apical interlocking device. Hind wings absent. Mesoventrite short and transverse, with large anterior depression to receive prosternal process. Metaventrite projecting posteriorly, covering the metacoxa. Mesocoxae slightly transverse, trochantins exposed. Metacoxae strongly transverse and oblique, subcontiguous; metacoxal plates (
<figureCitation id="135E1DC201484212FD8EC6BD419BFB19" box="[627,694,1186,1212]" captionStart="FIGURES 17" captionStartId="2.[151,264,1757,1781]" captionTargetBox="[183,1405,232,1733]" captionTargetId="figure-22@2.[183,1405,181,1733]" captionTargetPageId="2" captionText="FIGURES 17. Austramastodus apterus sp. nov. 1. Male habitus, dorsal view. 2. Male habitus, ventral view. 3. Male habitus, lateral view. 4. Head and pronotum, fronto-dorsal view. 5. Head, fronto-lateral view and male fore tarsi. 6. Male last abdominal sternite. 7. Female last abdominal sternite. The arrow points to the only two large lateral spiniform setae on the female." figureDoi="http://doi.org/10.5281/zenodo.8142494" httpUri="https://zenodo.org/record/8142494/files/figure.png" pageId="1" pageNumber="560">Fig. 2</figureCitation>
) large and strongly expanded laterally, posterior margins with two pointed projections, partly covering metatrochantins (
<figureCitation id="135E1DC201484212FD0FC6D94015FB45" box="[754,824,1222,1248]" captionStart="FIGURES 17" captionStartId="2.[151,264,1757,1781]" captionTargetBox="[183,1405,232,1733]" captionTargetId="figure-22@2.[183,1405,181,1733]" captionTargetPageId="2" captionText="FIGURES 17. Austramastodus apterus sp. nov. 1. Male habitus, dorsal view. 2. Male habitus, ventral view. 3. Male habitus, lateral view. 4. Head and pronotum, fronto-dorsal view. 5. Head, fronto-lateral view and male fore tarsi. 6. Male last abdominal sternite. 7. Female last abdominal sternite. The arrow points to the only two large lateral spiniform setae on the female." figureDoi="http://doi.org/10.5281/zenodo.8142494" httpUri="https://zenodo.org/record/8142494/files/figure.png" pageId="1" pageNumber="560">Fig. 2</figureCitation>
). Forelegs relatively short and stout, with a few apical spines; tibia clearly shorter than femur. Middle legs relatively short and stout, with a few apical spines; tibia as long as femur. Hind legs very long, with a longitudinal row of strong spurs; tibia much longer than femur. All legs sexually dimorphic, especially protarsus (
<figureCitation id="135E1DC201484212FD97C72D4181FAE8" box="[618,684,1330,1357]" captionStart="FIGURES 17" captionStartId="2.[151,264,1757,1781]" captionTargetBox="[183,1405,232,1733]" captionTargetId="figure-22@2.[183,1405,181,1733]" captionTargetPageId="2" captionText="FIGURES 17. Austramastodus apterus sp. nov. 1. Male habitus, dorsal view. 2. Male habitus, ventral view. 3. Male habitus, lateral view. 4. Head and pronotum, fronto-dorsal view. 5. Head, fronto-lateral view and male fore tarsi. 6. Male last abdominal sternite. 7. Female last abdominal sternite. The arrow points to the only two large lateral spiniform setae on the female." figureDoi="http://doi.org/10.5281/zenodo.8142494" httpUri="https://zenodo.org/record/8142494/files/figure.png" pageId="1" pageNumber="560">Fig. 5</figureCitation>
) and claws (see below). Tarsal formula 444.
</paragraph>
<paragraph id="8BDA014701484212FF3AC74A41E6F9A5" blockId="1.[151,1437,285,2040]" pageId="1" pageNumber="560">
<emphasis id="B911DD5501484212FF3AC74A4268FACA" bold="true" box="[199,325,1365,1391]" pageId="1" pageNumber="560">Abdomen:</emphasis>
(
<figureCitation id="135E1DC201484212FEA9C74942BAFAD5" box="[340,407,1366,1392]" captionStart="FIGURES 17" captionStartId="2.[151,264,1757,1781]" captionTargetBox="[183,1405,232,1733]" captionTargetId="figure-22@2.[183,1405,181,1733]" captionTargetPageId="2" captionText="FIGURES 17. Austramastodus apterus sp. nov. 1. Male habitus, dorsal view. 2. Male habitus, ventral view. 3. Male habitus, lateral view. 4. Head and pronotum, fronto-dorsal view. 5. Head, fronto-lateral view and male fore tarsi. 6. Male last abdominal sternite. 7. Female last abdominal sternite. The arrow points to the only two large lateral spiniform setae on the female." figureDoi="http://doi.org/10.5281/zenodo.8142494" httpUri="https://zenodo.org/record/8142494/files/figure.png" pageId="1" pageNumber="560">Fig. 2</figureCitation>
) with five ventrites, covered with short recumbent setae, finely punctate (not very dense and uniform), and covered by a polygonal squamiform reticulation; intercoxal process of the first abdominal ventrite very small; first three ventrites connate; ventrite 1 long; ventrites 24 subequal in length; lateral margin of ventrite 5 with a pair of subapical notches (
<figureCitation id="135E1DC201484212FDE6C7DD41ABFA79" box="[539,646,1474,1500]" captionStart="FIGURES 17" captionStartId="2.[151,264,1757,1781]" captionTargetBox="[183,1405,232,1733]" captionTargetId="figure-22@2.[183,1405,181,1733]" captionTargetPageId="2" captionText="FIGURES 17. Austramastodus apterus sp. nov. 1. Male habitus, dorsal view. 2. Male habitus, ventral view. 3. Male habitus, lateral view. 4. Head and pronotum, fronto-dorsal view. 5. Head, fronto-lateral view and male fore tarsi. 6. Male last abdominal sternite. 7. Female last abdominal sternite. The arrow points to the only two large lateral spiniform setae on the female." figureDoi="http://doi.org/10.5281/zenodo.8142494" httpUri="https://zenodo.org/record/8142494/files/figure.png" pageId="1" pageNumber="560">Figs. 67</figureCitation>
) (being part of the elytral-abdominal interlocking device); setation and apex of ventrite 5 sexually dimorphic (see below).
</paragraph>
<paragraph id="8BDA014701484212FF3AC41647ECF9ED" blockId="1.[151,1437,285,2040]" pageId="1" pageNumber="560">
<emphasis id="B911DD5501484212FF3AC416426CF986" bold="true" box="[199,321,1545,1571]" pageId="1" pageNumber="560">Aedeagus.</emphasis>
Articulated and strongly arched in lateral view (
<figureCitation id="135E1DC201484212FC8FC4154094F981" box="[882,953,1546,1572]" captionStart="FIGURES 815" captionStartId="3.[151,264,1920,1944]" captionTargetBox="[183,1405,345,1897]" captionTargetId="figure-89@3.[183,1405,345,1897]" captionTargetPageId="3" captionText="FIGURES 815. Austramastodus apterus sp. nov. 8. Aedeagus, ventral view. 9. Aedeagus, lateral view. 10. Ninth male genital segment, ventral view. 11. Male sternite VIII. 12. Gonocoxites and gonocoxal struts, ventral view. 13. Spiculum ventral. 14. gonocoxites 15. Spiculum ventral, distal plaque." figureDoi="http://doi.org/10.5281/zenodo.8142496" httpUri="https://zenodo.org/record/8142496/files/figure.png" pageId="1" pageNumber="560">Fig. 9</figureCitation>
), with the base longer than the parameres (
<figureCitation id="135E1DC201484212FF62C4314221F9ED" box="[159,268,1582,1608]" captionStart="FIGURES 815" captionStartId="3.[151,264,1920,1944]" captionTargetBox="[183,1405,345,1897]" captionTargetId="figure-89@3.[183,1405,345,1897]" captionTargetPageId="3" captionText="FIGURES 815. Austramastodus apterus sp. nov. 8. Aedeagus, ventral view. 9. Aedeagus, lateral view. 10. Ninth male genital segment, ventral view. 11. Male sternite VIII. 12. Gonocoxites and gonocoxal struts, ventral view. 13. Spiculum ventral. 14. gonocoxites 15. Spiculum ventral, distal plaque." figureDoi="http://doi.org/10.5281/zenodo.8142496" httpUri="https://zenodo.org/record/8142496/files/figure.png" pageId="1" pageNumber="560">Figs. 89</figureCitation>
); median lobe with the apex strongly curved inwards, falciform in shape (
<figureCitation id="135E1DC201484212FBBAC4314799F9ED" box="[1095,1204,1582,1608]" captionStart="FIGURES 815" captionStartId="3.[151,264,1920,1944]" captionTargetBox="[183,1405,345,1897]" captionTargetId="figure-89@3.[183,1405,345,1897]" captionTargetPageId="3" captionText="FIGURES 815. Austramastodus apterus sp. nov. 8. Aedeagus, ventral view. 9. Aedeagus, lateral view. 10. Ninth male genital segment, ventral view. 11. Male sternite VIII. 12. Gonocoxites and gonocoxal struts, ventral view. 13. Spiculum ventral. 14. gonocoxites 15. Spiculum ventral, distal plaque." figureDoi="http://doi.org/10.5281/zenodo.8142496" httpUri="https://zenodo.org/record/8142496/files/figure.png" pageId="1" pageNumber="560">Figs. 89</figureCitation>
).
</paragraph>
<paragraph id="8BDA014701484212FF3AC44D47B9F910" blockId="1.[151,1437,285,2040]" pageId="1" pageNumber="560">
<emphasis id="B911DD5501484212FF3AC44D42BEF9CE" bold="true" box="[199,403,1617,1644]" pageId="1" pageNumber="560">Female genitalia.</emphasis>
Ovopositor in ventral view as in
<figureCitation id="135E1DC201484212FCF7C44D4070F9C9" box="[778,861,1618,1644]" captionStart="FIGURES 815" captionStartId="3.[151,264,1920,1944]" captionTargetBox="[183,1405,345,1897]" captionTargetId="figure-89@3.[183,1405,345,1897]" captionTargetPageId="3" captionText="FIGURES 815. Austramastodus apterus sp. nov. 8. Aedeagus, ventral view. 9. Aedeagus, lateral view. 10. Ninth male genital segment, ventral view. 11. Male sternite VIII. 12. Gonocoxites and gonocoxal struts, ventral view. 13. Spiculum ventral. 14. gonocoxites 15. Spiculum ventral, distal plaque." figureDoi="http://doi.org/10.5281/zenodo.8142496" httpUri="https://zenodo.org/record/8142496/files/figure.png" pageId="1" pageNumber="560">Fig. 12</figureCitation>
; gonocoxal struts very long, ten times the length of gonocoxites; gonocoxites strongly acuminate (
<figureCitation id="135E1DC201484212FD63C4694009F935" box="[670,804,1654,1680]" captionStart="FIGURES 815" captionStartId="3.[151,264,1920,1944]" captionTargetBox="[183,1405,345,1897]" captionTargetId="figure-89@3.[183,1405,345,1897]" captionTargetPageId="3" captionText="FIGURES 815. Austramastodus apterus sp. nov. 8. Aedeagus, ventral view. 9. Aedeagus, lateral view. 10. Ninth male genital segment, ventral view. 11. Male sternite VIII. 12. Gonocoxites and gonocoxal struts, ventral view. 13. Spiculum ventral. 14. gonocoxites 15. Spiculum ventral, distal plaque." figureDoi="http://doi.org/10.5281/zenodo.8142496" httpUri="https://zenodo.org/record/8142496/files/figure.png" pageId="1" pageNumber="560">Figs. 1214</figureCitation>
) and articulate at apex of the gonocoxal struts. Spiculum ventrale with a long manubrium, distal part with two long lateral expansions (
<figureCitation id="135E1DC201484212FC03C48547AAF910" box="[1022,1159,1690,1717]" captionStart="FIGURES 815" captionStartId="3.[151,264,1920,1944]" captionTargetBox="[183,1405,345,1897]" captionTargetId="figure-89@3.[183,1405,345,1897]" captionTargetPageId="3" captionText="FIGURES 815. Austramastodus apterus sp. nov. 8. Aedeagus, ventral view. 9. Aedeagus, lateral view. 10. Ninth male genital segment, ventral view. 11. Male sternite VIII. 12. Gonocoxites and gonocoxal struts, ventral view. 13. Spiculum ventral. 14. gonocoxites 15. Spiculum ventral, distal plaque." figureDoi="http://doi.org/10.5281/zenodo.8142496" httpUri="https://zenodo.org/record/8142496/files/figure.png" pageId="1" pageNumber="560">Figs. 1315</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37F52CC01484210FF3AC4A24236FEE4" lastPageId="3" lastPageNumber="562" pageId="1" pageNumber="560" type="diagnosis">
<paragraph id="8BDA014701484212FF3AC4A24176F85D" blockId="1.[151,1437,285,2040]" pageId="1" pageNumber="560">
<emphasis id="B911DD5501484212FF3AC4A242E1F972" bold="true" box="[199,460,1725,1751]" pageId="1" pageNumber="560">Differential diagnosis</emphasis>
.
<taxonomicName id="4C657AC401484212FE1DC4A141B9F97D" authority="Hernando, 2023" authorityName="Hernando" authorityYear="2023" box="[480,660,1726,1752]" class="Insecta" family="Limnichidae" genus="Austramastodus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="560" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B911DD5501484212FE1DC4A141B9F97D" box="[480,660,1726,1752]" italics="true" pageId="1" pageNumber="560">Austramastodus</emphasis>
</taxonomicName>
<emphasis id="B911DD5501484212FD5FC4A1403CF97D" bold="true" box="[674,785,1726,1752]" pageId="1" pageNumber="560">
<taxonomicNameLabel id="A222602E01484212FD5FC4A1403CF97D" box="[674,785,1726,1752]" pageId="1" pageNumber="560" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
can be clearly distinguished from the other genera of
<taxonomicName id="4C657AC401484212FF6AC4FD427DF959" authorityName="Champion" authorityYear="1924" box="[151,336,1762,1788]" pageId="1" pageNumber="563" rank="subFamily" subFamily="Thaumastodinae">Thaumastodinae</taxonomicName>
(
<taxonomicName id="4C657AC401484212FE99C4FD41EFF959" authority="Champion, 1924" authorityName="Champion" authorityYear="1924" box="[356,706,1762,1788]" class="Insecta" family="Limnichidae" genus="Acontosceles" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="B911DD5501484212FE99C4FD42DBF959" box="[356,502,1762,1788]" italics="true" pageId="1" pageNumber="560">Acontosceles</emphasis>
Champion, 1924
</taxonomicName>
,
<taxonomicName id="4C657AC401484212FD2DC4FD4053F959" authorityName="Heller" authorityYear="1921" box="[720,894,1762,1788]" class="Insecta" family="Limnichidae" genus="Pseudeucinetus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="B911DD5501484212FD2DC4FD4053F959" box="[720,894,1762,1788]" italics="true" pageId="1" pageNumber="560">Pseudeucinetus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C657AC401484212FC6DC4FC40CFF959" authorityName="Spilman" authorityYear="1972" box="[912,994,1763,1788]" class="Insecta" family="Limnichidae" genus="Mexico" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="B911DD5501484212FC6DC4FC40CFF959" box="[912,994,1763,1788]" italics="true" pageId="1" pageNumber="560">Mexico</emphasis>
</taxonomicName>
and
<taxonomicName id="4C657AC401484212FBDDC4FC466CF958" authority="Spilman, 1959" authorityName="Spilman" authorityYear="1959" box="[1056,1345,1762,1789]" class="Insecta" family="Limnichidae" genus="Martinius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="B911DD5501484212FBDDC4FC47A3F959" box="[1056,1166,1763,1788]" italics="true" pageId="1" pageNumber="560">Martinius</emphasis>
Spilman, 1959
</taxonomicName>
) by the widely separated eyes, which are in lateral position and with the space between them larger than the diameter of an eye (eyes close together and in a more frontal position in the other four genera and with the space between the eyes always narrower than the maximum length of an eye). Additional distinguishing characters: tarsal formula 444 (4 5
<quantity id="4C9DACA201484212FF49C56D43F2F829" box="[180,223,1906,1933]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="1" pageNumber="560" unit="in" value="5.0">5 in</quantity>
<taxonomicName id="4C657AC401484212FF1BC56D4255F829" authorityName="Champion" authorityYear="1924" box="[230,376,1906,1932]" class="Insecta" family="Limnichidae" genus="Acontosceles" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="B911DD5501484212FF1BC56D4255F829" box="[230,376,1906,1932]" italics="true" pageId="1" pageNumber="560">Acontosceles</emphasis>
</taxonomicName>
); 11-segmented antennae (7-segmented in
<taxonomicName id="4C657AC401484212FCA0C56C40E6F829" authorityName="Spilman" authorityYear="1959" box="[861,971,1907,1932]" class="Insecta" family="Limnichidae" genus="Martinius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="B911DD5501484212FCA0C56C40E6F829" box="[861,971,1907,1932]" italics="true" pageId="1" pageNumber="560">Martinius</emphasis>
</taxonomicName>
); lateral margin of elytra smooth (serrate in
<taxonomicName id="4C657AC401484212FF4BC5884209F815" authorityName="Spilman" authorityYear="1959" box="[182,292,1943,1968]" class="Insecta" family="Limnichidae" genus="Martinius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="B911DD5501484212FF4BC5884209F815" box="[182,292,1943,1968]" italics="true" pageId="1" pageNumber="560">Martinius</emphasis>
</taxonomicName>
and
<taxonomicName id="4C657AC401484212FEA3C588429DF815" authorityName="Spilman" authorityYear="1972" box="[350,432,1943,1968]" class="Insecta" family="Limnichidae" genus="Mexico" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="B911DD5501484212FEA3C588429DF815" box="[350,432,1943,1968]" italics="true" pageId="1" pageNumber="560">Mexico</emphasis>
</taxonomicName>
); presence of antennal groove (lacking in
<taxonomicName id="4C657AC401484212FC6AC5884728F815" authorityName="Spilman" authorityYear="1959" box="[919,1029,1943,1968]" class="Insecta" family="Limnichidae" genus="Martinius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="B911DD5501484212FC6AC5884728F815" box="[919,1029,1943,1968]" italics="true" pageId="1" pageNumber="560">Martinius</emphasis>
</taxonomicName>
); remarkable sexual dimorphism in the protarsus (lacking in
<taxonomicName id="4C657AC401484212FE50C5A54112F871" authorityName="Champion" authorityYear="1924" box="[429,575,1978,2004]" class="Insecta" family="Limnichidae" genus="Acontosceles" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="B911DD5501484212FE50C5A54112F871" box="[429,575,1978,2004]" italics="true" pageId="1" pageNumber="560">Acontosceles</emphasis>
</taxonomicName>
,
<taxonomicName id="4C657AC401484212FDB0C5A44196F871" authorityName="Spilman" authorityYear="1959" box="[589,699,1979,2004]" class="Insecta" family="Limnichidae" genus="Martinius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="B911DD5501484212FDB0C5A44196F871" box="[589,699,1979,2004]" italics="true" pageId="1" pageNumber="560">Martinius</emphasis>
</taxonomicName>
and
<taxonomicName id="4C657AC401484212FD0CC5A4406EF871" authorityName="Spilman" authorityYear="1972" box="[753,835,1979,2004]" class="Insecta" family="Limnichidae" genus="Mexico" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="B911DD5501484212FD0CC5A4406EF871" box="[753,835,1979,2004]" italics="true" pageId="1" pageNumber="560">Mexico</emphasis>
</taxonomicName>
); eyes large and subcircular (narrow and elongated in
<taxonomicName id="4C657AC401484212FF6AC5C04228F85D" authorityName="Spilman" authorityYear="1959" box="[151,261,2015,2040]" class="Insecta" family="Limnichidae" genus="Martinius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="B911DD5501484212FF6AC5C04228F85D" box="[151,261,2015,2040]" italics="true" pageId="1" pageNumber="560">Martinius</emphasis>
</taxonomicName>
,
<taxonomicName id="4C657AC401484212FEE9C5C0424BF85D" authorityName="Spilman" authorityYear="1972" box="[276,358,2015,2040]" class="Insecta" family="Limnichidae" genus="Mexico" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="B911DD5501484212FEE9C5C0424BF85D" box="[276,358,2015,2040]" italics="true" pageId="1" pageNumber="560">Mexico</emphasis>
</taxonomicName>
and
<taxonomicName id="4C657AC401484212FE61C5C14167F85D" authorityName="Heller" authorityYear="1921" box="[412,586,2014,2040]" class="Insecta" family="Limnichidae" genus="Pseudeucinetus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="B911DD5501484212FE61C5C14167F85D" box="[412,586,2014,2040]" italics="true" pageId="1" pageNumber="560">Pseudeucinetus</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="DF1A51CF014B4211FF6AC4C24663F898" ID-DOI="http://doi.org/10.5281/zenodo.8142494" ID-Zenodo-Dep="8142494" httpUri="https://zenodo.org/record/8142494/files/figure.png" pageId="2" pageNumber="561" startId="2.[151,264,1757,1781]" targetBox="[183,1405,232,1733]" targetPageId="2" targetType="figure">
<paragraph id="8BDA0147014B4211FF6AC4C24663F898" blockId="2.[151,1436,1757,1853]" pageId="2" pageNumber="561">
<emphasis id="B911DD55014B4211FF6AC4C24216F950" bold="true" box="[151,315,1757,1781]" pageId="2" pageNumber="561">FIGURES 17.</emphasis>
<taxonomicName id="4C657AC4014B4211FEBFC4C24115F950" authority="Hernando, 2023" authorityName="Hernando" authorityYear="2023" box="[322,568,1757,1781]" class="Insecta" family="Limnichidae" genus="Austramastodus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="apterus" status="sp. nov.">
<emphasis id="B911DD55014B4211FEBFC4C24115F950" box="[322,568,1757,1781]" italics="true" pageId="2" pageNumber="561">Austramastodus apterus</emphasis>
</taxonomicName>
<emphasis id="B911DD55014B4211FDC2C4C24187F950" bold="true" box="[575,682,1757,1781]" pageId="2" pageNumber="561">
<taxonomicNameLabel id="A222602E014B4211FDC2C4C241BDF950" box="[575,656,1757,1781]" pageId="2" pageNumber="561" rank="species">sp. nov.</taxonomicNameLabel>
1.
</emphasis>
Male habitus, dorsal view.
<emphasis id="B911DD55014B4211FC3EC4C240FBF950" bold="true" box="[963,982,1757,1781]" pageId="2" pageNumber="561">2.</emphasis>
Male habitus, ventral view.
<emphasis id="B911DD55014B4211FB05C4C24626F950" bold="true" box="[1272,1291,1757,1781]" pageId="2" pageNumber="561">3.</emphasis>
Male habitus, lateral view.
<emphasis id="B911DD55014B4211FEE4C51E4201F8BC" bold="true" box="[281,300,1793,1817]" pageId="2" pageNumber="561">4.</emphasis>
Head and pronotum, fronto-dorsal view.
<emphasis id="B911DD55014B4211FD30C51E41CDF8BC" bold="true" box="[717,736,1793,1817]" pageId="2" pageNumber="561">5.</emphasis>
Head, fronto-lateral view and male fore tarsi.
<emphasis id="B911DD55014B4211FB48C51E47EAF8BC" bold="true" box="[1205,1223,1793,1817]" pageId="2" pageNumber="561">6.</emphasis>
Male last abdominal sternite.
<emphasis id="B911DD55014B4211FF13C53A422CF898" bold="true" box="[238,257,1829,1853]" pageId="2" pageNumber="561">7.</emphasis>
Female last abdominal sternite. The arrow points to the only two large lateral spiniform setae on the female.
</paragraph>
</caption>
<paragraph id="8BDA0147014B4210FF3AC5904236FEE4" blockId="2.[151,1436,1935,2034]" lastBlockId="3.[151,1436,151,321]" lastPageId="3" lastPageNumber="562" pageId="2" pageNumber="561">
Within the subfamily,
<taxonomicName id="4C657AC4014B4211FE3BC5904157F80C" authority="Hernando, 2023" authorityName="Hernando" authorityYear="2023" box="[454,634,1935,1961]" class="Insecta" family="Limnichidae" genus="Austramastodus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B911DD55014B4211FE3BC5904157F80C" box="[454,634,1935,1961]" italics="true" pageId="2" pageNumber="561">Austramastodus</emphasis>
</taxonomicName>
<emphasis id="B911DD55014B4211FD7EC58F41C3F80F" bold="true" box="[643,750,1936,1962]" pageId="2" pageNumber="561">
<taxonomicNameLabel id="A222602E014B4211FD7EC58F41C3F80F" box="[643,750,1936,1962]" pageId="2" pageNumber="561" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
, seems to be close to
<taxonomicName id="4C657AC4014B4211FC12C59047B0F80C" authorityName="Heller" authorityYear="1921" box="[1007,1181,1935,1961]" class="Insecta" family="Limnichidae" genus="Pseudeucinetus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus">
<emphasis id="B911DD55014B4211FC12C59047B0F80C" box="[1007,1181,1935,1961]" italics="true" pageId="2" pageNumber="561">Pseudeucinetus</emphasis>
</taxonomicName>
, from which it can be distinguished (apart from the position of the eyes) by the following characters: body form elongate-oval (
<emphasis id="B911DD55014B4211FAE5C5AB4602F868" box="[1304,1327,1972,1997]" italics="true" pageId="2" pageNumber="561">vs</emphasis>
. fusiform in
<taxonomicName id="4C657AC4014B4211FF49C5C8424FF854" authorityName="Heller" authorityYear="1921" box="[180,354,2007,2033]" class="Insecta" family="Limnichidae" genus="Pseudeucinetus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus">
<emphasis id="B911DD55014B4211FF49C5C8424FF854" box="[180,354,2007,2033]" italics="true" pageId="2" pageNumber="561">Pseudeucinetus</emphasis>
</taxonomicName>
); head transverse and regularly curved (
<emphasis id="B911DD55014B4211FCD4C5C7406DF854" box="[809,832,2008,2033]" italics="true" pageId="2" pageNumber="561">vs</emphasis>
. projecting forward and pointed); posterior margin of the pronotum weakly sinuate (
<emphasis id="B911DD55014A4210FE0FC2884124FF15" box="[498,521,151,176]" italics="true" pageId="3" pageNumber="562">vs</emphasis>
. strongly sinuate); male with mesotarsal and metatarsal claws asymmetrical (
<emphasis id="B911DD55014A4210FA83C28846B8FF15" box="[1406,1429,151,176]" italics="true" pageId="3" pageNumber="562">vs</emphasis>
. symmetrical); parameres and median lobe shorter than the phallobase, strongly curved apically (
<emphasis id="B911DD55014A4210FB14C2A4462DFF71" box="[1257,1280,187,212]" italics="true" pageId="3" pageNumber="562">vs</emphasis>
. equal length or longer, always straight); lamina of the genital segment extending beyond insertion of the parameres (
<emphasis id="B911DD55014A4210FAD7C2C0466CFF5D" box="[1322,1345,223,248]" italics="true" pageId="3" pageNumber="562">vs</emphasis>
. lamina not extending, parameres inserted at the apex of the lamina); gonocoxites separate (
<emphasis id="B911DD55014A4210FB9DC31C475AFEB9" box="[1120,1143,259,284]" italics="true" pageId="3" pageNumber="562">vs</emphasis>
. convergent and strongly acuminate).
</paragraph>
</subSubSection>
</treatment>
</document>