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<document id="D2D5A80F29538D5680651067F9CC0391" ID-DOI="10.5281/zenodo.5333042" ID-ISSN="2345-7600" ID-Zenodo-Dep="5333042" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630173599483" checkinUser="diego" docAuthor="Chen, Hong-Wei, Toda, Masanori J., Lakim, Maklarin B. &amp; Mohamed, Maryati B." docDate="2007" docId="03CC87B2981F0065B80E6B283854FEED" docLanguage="en" docName="55rbz23-41.pdf" docOrigin="Raffles Bulletin of Zoology 55 (1)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Phortica (Phortica) epsilon Chen &amp; Toda 2007, new species" docType="treatment" docVersion="3" lastPageNumber="32" masterDocId="FFF5FFCA9818006CBB3668493A62FF95" masterDocTitle="The Phortica Sensu Stricto (Insecta: Diptera: Drosophilidae) From Malaysia" masterLastPageNumber="41" masterPageNumber="23" pageNumber="30" updateTime="1699173972972" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="924F03B048504A8C78EA4DF090EFCA3B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="18F24CF2C1BBC0FA06DE21AF099428CA">
<mods:title id="1F20C5A785570DEA14EBC91AEC9836B9">The Phortica Sensu Stricto (Insecta: Diptera: Drosophilidae) From Malaysia</mods:title>
</mods:titleInfo>
<mods:name id="5A2B9A76B35C7CB716E44B95097C0142" type="personal">
<mods:role id="5998E30669360454E6C8F6C6F0D03A15">
<mods:roleTerm id="1075E6AEEDCC1C5D58DEE7B4DE9990EF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BB02CA65C8FEEAD3C0A11E97530D8725">Chen, Hong-Wei</mods:namePart>
</mods:name>
<mods:name id="DE1BF0D0416A29AD41BFC35D3B588E88" type="personal">
<mods:role id="932AC1B952C709D851593147574C4F43">
<mods:roleTerm id="309242486E5FC7533D4718127C32029D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A3D5E3C53E669873E81C81A77018EAB0">Toda, Masanori J.</mods:namePart>
</mods:name>
<mods:name id="7FE332C9345220932A15D33F5C54118D" type="personal">
<mods:role id="D7589D10E15B94AB7044F15E4115A45B">
<mods:roleTerm id="77A15867F7D24E924494863D67407160">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FF38915E5AF6A1F662870A6588F24A10">Lakim, Maklarin B.</mods:namePart>
</mods:name>
<mods:name id="967F6CB2D26F643EDDB497984C5B4C87" type="personal">
<mods:role id="C7E812ADCDA9BEE0A87EBE56B7D8620B">
<mods:roleTerm id="116FEFD5229091EA7A7C8E937975927F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4E9692BADA97E2C8AF85C46F374626E7">Mohamed, Maryati B.</mods:namePart>
</mods:name>
<mods:typeOfResource id="076012B9EFEF917B397091FAA13EF757">text</mods:typeOfResource>
<mods:relatedItem id="060CB8015450D4CD27E2CF66A868AAFE" type="host">
<mods:titleInfo id="5281B12EB9D9CC469AC7732D12B9231D">
<mods:title id="D7A6FF4C5DC5CCD82D4E13FA9DDBC421">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="842865158416EB90087EDE79069AD22B">
<mods:date id="AD5D12BC6A6AAFCBD5C9D94895EAA0BC">2007</mods:date>
<mods:detail id="D845FB92378CC4ACBB2F89B0479BE75C" type="pubDate">
<mods:number id="D1E23F5FD494F7D963C60BE73A3C275B">2007-02-28</mods:number>
</mods:detail>
<mods:detail id="2D64D53DAB90D1C4D8D07A01DAC5B121" type="volume">
<mods:number id="D3D5B242FF0A99F998A90CC729FA8191">55</mods:number>
</mods:detail>
<mods:detail id="4E53FA5AECF2A841EA34BA208CB1E27E" type="issue">
<mods:number id="D904660B36BE7987E49934D1D3647345">1</mods:number>
</mods:detail>
<mods:extent id="1A45EC1697B71DDB046D19A588B500E9" unit="page">
<mods:start id="8ECAEA1AC9CF3AFF70A9C65221DC5F66">23</mods:start>
<mods:end id="58A37BC558880D4A42FACEB45F0D480A">41</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:classification id="247A6C2508AA86E54052452105661AFE">journal article</mods:classification>
<mods:identifier id="455A3127EADBEB9C6FCD12F01EEF4465" type="DOI">10.5281/zenodo.5333042</mods:identifier>
<mods:identifier id="FCE4AD62266B0DCCE7416E8F189EEFAB" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="22CB08AE28C79F2FF62114A4699EA5E7" type="Zenodo-Dep">5333042</mods:identifier>
</mods:mods>
<treatment id="03CC87B2981F0065B80E6B283854FEED" LSID="urn:lsid:plazi:treatment:03CC87B2981F0065B80E6B283854FEED" httpUri="http://treatment.plazi.org/id/03CC87B2981F0065B80E6B283854FEED" lastPageId="9" lastPageNumber="32" pageId="7" pageNumber="30">
<subSubSection id="C37F652F981F006BB80E6B283FC2FCED" box="[824,1440,864,889]" pageId="7" pageNumber="30" type="nomenclature">
<paragraph id="8BDA36A4981F006BB80E6B283FC2FCED" blockId="7.[824,1440,864,920]" box="[824,1440,864,889]" pageId="7" pageNumber="30">
<heading id="D09281C8981F006BB80E6B283FC2FCED" bold="true" box="[824,1440,864,889]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="30" reason="2">
<emphasis id="B911EAB6981F006BB80E6B283FC2FCED" bold="true" box="[824,1440,864,889]" pageId="7" pageNumber="30">
<taxonomicName id="4C654D27981F006BB80E6B283F6CFCED" ID-CoL="8DBVN" authority="Chen &amp; Toda" authorityName="Chen &amp; Toda" authorityYear="2007" box="[824,1294,864,889]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="30" phylum="Arthropoda" rank="species" species="epsilon" status="sp. nov." subGenus="Phortica">
<emphasis id="B911EAB6981F006BB80E6B2839F5FCEC" bold="true" box="[824,919,865,889]" italics="true" pageId="7" pageNumber="30">Phortica</emphasis>
(
<emphasis id="B911EAB6981F006BB89E6B283E65FCEC" bold="true" box="[936,1031,865,889]" italics="true" pageId="7" pageNumber="30">Phortica</emphasis>
)
<emphasis id="B911EAB6981F006BBF2E6B283E05FCEC" bold="true" box="[1048,1127,865,889]" italics="true" pageId="7" pageNumber="30">epsilon</emphasis>
Chen &amp; Toda
</taxonomicName>
,
<taxonomicNameLabel id="A22257CD981F006BBE2D6B293FC2FCED" box="[1307,1440,864,888]" pageId="7" pageNumber="30" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37F652F981F006BBF156BC83ED6FC0D" box="[1059,1204,897,920]" pageId="7" pageNumber="30" type="description">
<paragraph id="8BDA36A4981F006BBF156BC83ED6FC0D" blockId="7.[824,1440,864,920]" box="[1059,1204,897,920]" pageId="7" pageNumber="30">
(
<figureCitation id="135E2A21981F006BBF1D6BC83ECFFC0D" box="[1067,1197,897,920]" captionStart="Figs" captionStartId="8.[136,180,2035,2056]" captionTargetBox="[142,1479,1066,1985]" captionTargetId="figure-8@8.[124,1512,1005,2022]" captionTargetPageId="8" captionText="Figs. 20–25. Phortica (Phortica) epsilon Chen &amp; Toda, new species, male: 20, sixth tergite in ventral view; 21, epandrium and cercus in lateral view; 22, surstylus in frontal view; 23, tenth sternite and additional plate in frontal view; 24, hypandrium, gonopod, paramere and aedeagus in lateral view; 25, tip of aedeagal median rod in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5333054" httpUri="https://zenodo.org/record/5333054/files/figure.png" pageId="7" pageNumber="30">Figs. 20–25</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37F652F981F0064B81A6B893965FEB9" lastPageId="8" lastPageNumber="31" pageId="7" pageNumber="30" type="diagnosis">
<paragraph id="8BDA36A4981F0064B81A6B893965FEB9" blockId="7.[812,1451,960,1039]" lastBlockId="8.[136,775,192,300]" lastPageId="8" lastPageNumber="31" pageId="7" pageNumber="30">
<emphasis id="B911EAB6981F006BB81A6B8939C7FC40" bold="true" box="[812,933,960,981]" pageId="7" pageNumber="30">
<emphasis id="B911EAB6981F006BB81A6B8939F7FC40" bold="true" box="[812,917,960,981]" italics="true" pageId="7" pageNumber="30">Diagnosis.</emphasis>
–
</emphasis>
Paramere almost lacking pubescence, subapically with 1 acute projection (
<figureCitation id="135E2A21981F006BBF356B943E37FC67" box="[1027,1109,989,1010]" captionStart="Figs" captionStartId="8.[136,180,2035,2056]" captionTargetBox="[142,1479,1066,1985]" captionTargetId="figure-8@8.[124,1512,1005,2022]" captionTargetPageId="8" captionText="Figs. 20–25. Phortica (Phortica) epsilon Chen &amp; Toda, new species, male: 20, sixth tergite in ventral view; 21, epandrium and cercus in lateral view; 22, surstylus in frontal view; 23, tenth sternite and additional plate in frontal view; 24, hypandrium, gonopod, paramere and aedeagus in lateral view; 25, tip of aedeagal median rod in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5333054" httpUri="https://zenodo.org/record/5333054/files/figure.png" pageId="7" pageNumber="30">Fig. 24</figureCitation>
); aedeagal median rod apically asymmetrically bifurcated, subapically expanded and with minute warts on surface (
<figureCitation id="135E2A2198100064BA0968893BDFFF40" box="[319,445,192,213]" captionStart="Figs" captionStartId="8.[136,180,2035,2056]" captionTargetBox="[142,1479,1066,1985]" captionTargetId="figure-8@8.[124,1512,1005,2022]" captionTargetPageId="8" captionText="Figs. 20–25. Phortica (Phortica) epsilon Chen &amp; Toda, new species, male: 20, sixth tergite in ventral view; 21, epandrium and cercus in lateral view; 22, surstylus in frontal view; 23, tenth sternite and additional plate in frontal view; 24, hypandrium, gonopod, paramere and aedeagus in lateral view; 25, tip of aedeagal median rod in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5333054" httpUri="https://zenodo.org/record/5333054/files/figure.png" pageId="8" pageNumber="31">Figs. 24, 25</figureCitation>
); aedeagal basal bridge strongly sclerotized, tripartite: anterior lobe with 2 triangular teeth apically and many spinules on lateral surface; median lobe widely clavate, with 1 triangular tooth apically; posterior process slender (
<figureCitation id="135E2A2198100064B982695E3899FEB9" box="[692,763,279,300]" captionStart="Figs" captionStartId="8.[136,180,2035,2056]" captionTargetBox="[142,1479,1066,1985]" captionTargetId="figure-8@8.[124,1512,1005,2022]" captionTargetPageId="8" captionText="Figs. 20–25. Phortica (Phortica) epsilon Chen &amp; Toda, new species, male: 20, sixth tergite in ventral view; 21, epandrium and cercus in lateral view; 22, surstylus in frontal view; 23, tenth sternite and additional plate in frontal view; 24, hypandrium, gonopod, paramere and aedeagus in lateral view; 25, tip of aedeagal median rod in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5333054" httpUri="https://zenodo.org/record/5333054/files/figure.png" pageId="8" pageNumber="31">Fig. 24</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF1A662C981F006BBB47605B3F22F7D6" ID-DOI="http://doi.org/10.5281/zenodo.5333052" ID-Zenodo-Dep="5333052" httpUri="https://zenodo.org/record/5333052/files/figure.png" pageId="7" pageNumber="30" startId="7.[113,157,2066,2087]" targetBox="[119,1450,1107,2013]" targetPageId="7">
<paragraph id="8BDA36A4981F006BBB47605B3F22F7D6" blockId="7.[113,1451,2066,2115]" pageId="7" pageNumber="30">
Figs. 16–19.
<taxonomicName id="4C654D27981F006BBBC6605B3810F7B2" authority="Chen &amp; Toda" authorityName="Chen &amp; Toda" authorityYear="2007" box="[240,626,2066,2087]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="30" phylum="Arthropoda" rank="species" species="liewi" status="sp. nov." subGenus="Phortica">
<emphasis id="B911EAB6981F006BBBC6605B3B26F7B2" box="[240,324,2066,2087]" italics="true" pageId="7" pageNumber="30">Phortica</emphasis>
(
<emphasis id="B911EAB6981F006BBA65605B3BC5F7B2" box="[339,423,2066,2087]" italics="true" pageId="7" pageNumber="30">Phortica</emphasis>
)
<emphasis id="B911EAB6981F006BBA83605B3B86F7B2" box="[437,484,2066,2087]" italics="true" pageId="7" pageNumber="30">liewi</emphasis>
Chen &amp; Toda
</taxonomicName>
,
<taxonomicNameLabel id="A22257CD981F006BB94A605B3891F7B2" box="[636,755,2066,2087]" pageId="7" pageNumber="30" rank="species">new species</taxonomicNameLabel>
, male: 16, epandrium and cercus in lateral view; 17, surstylus in frontal view; 18, tenth sternite and additional plate in frontal view; 19, hypandrium, gonopod, paramere and aedeagus in lateral view.
</paragraph>
</caption>
<subSubSection id="C37F652F98100064BBBE69193BF9FE0A" pageId="8" pageNumber="31" type="materials_examined">
<paragraph id="8BDA36A498100064BBBE69193BF9FE0A" blockId="8.[136,775,336,415]" pageId="8" pageNumber="31">
<emphasis id="B911EAB698100064BBBE69193B0DFEF0" bold="true" box="[136,367,336,357]" pageId="8" pageNumber="31">
<emphasis id="B911EAB698100064BBBE69193B30FEF0" bold="true" box="[136,338,336,357]" italics="true" pageId="8" pageNumber="31">Material examined</emphasis>
. –
</emphasis>
<materialsCitation id="3B0D3CF998100064BA4F69193BF9FE0A" collectingDate="1999-03-11" collectionCode="KPSP" collectorName="M. J. Toda" country="Malaysia" location="Mt. Kinabalu" pageId="8" pageNumber="31" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus id="54DE880698100064BA4F69193BBAFEF0" box="[377,472,336,357]" pageId="8" pageNumber="31" type="holotype">Holotype</typeStatus>
male (terminalia dissected),
<collectingCountry id="F372763498100064BBBE69243B69FE17" box="[136,267,365,386]" name="Malaysia" pageId="8" pageNumber="31">MALAYSIA</collectingCountry>
: Park Headquarters,
<location id="8EBA607F98100064BAE469243836FE17" LSID="urn:lsid:plazi:treatment:03CC87B2981F0065B80E6B283854FEED:8EBA607F98100064BAE469243836FE17" box="[466,596,365,386]" country="Malaysia" name="Mt. Kinabalu" pageId="8" pageNumber="31" stateProvince="Sabah">Mt. Kinabalu</location>
,
<collectingRegion id="49A1F84698100064B968692438F9FE17" box="[606,667,365,386]" country="Malaysia" name="Sabah" pageId="8" pageNumber="31">Sabah</collectingRegion>
, coll.
<collectorName id="2690537298100064B9E369243ADEFE0A" pageId="8" pageNumber="31">M. J. Toda</collectorName>
,
<date id="FFDB106498100064BBF069C33B21FE0A" box="[198,323,394,415]" pageId="8" pageNumber="31" value="1999-03-11">
<collectingDate id="EF9FE98C98100064BBF069C33B21FE0A" box="[198,323,394,415]" pageId="8" pageNumber="31" value="1999-03-11">11 Mar.1999</collectingDate>
</date>
(
<collectionCode id="ED74AE6198100064BA6769C33BF2FE0A" box="[337,400,394,415]" pageId="8" pageNumber="31">KPSP</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37F652F98100064BBBE698E3E89FC8D" pageId="8" pageNumber="31" type="description">
<paragraph id="8BDA36A498100064BBBE698E3B22FE6B" blockId="8.[136,774,455,510]" pageId="8" pageNumber="31">
<emphasis id="B911EAB698100064BBBE698E3B4AFE4A" bold="true" box="[136,296,455,479]" pageId="8" pageNumber="31">
<emphasis id="B911EAB698100064BBBE698E3B72FE4A" bold="true" box="[136,272,455,479]" italics="true" pageId="8" pageNumber="31">Description.</emphasis>
–
</emphasis>
Male. Head: Frons brown. Arista without ventral branches.
</paragraph>
<paragraph id="8BDA36A498100064BBBE6A6E38FBFDCB" blockId="8.[136,775,551,606]" pageId="8" pageNumber="31">Thorax: Orange brown. Scutellum with trifurcate dark brown patch, pale on tip. Anepisternum lacking setulae.</paragraph>
<paragraph id="8BDA36A498100064BBBE6ACE382CFD0B" blockId="8.[136,590,647,670]" box="[136,590,647,670]" pageId="8" pageNumber="31">Wing: Slightly smoky on anterior margin.</paragraph>
<paragraph id="8BDA36A498100064BBBE6A8E38AEFD6B" blockId="8.[136,774,711,766]" pageId="8" pageNumber="31">Legs: All femora brown except apical portions. Midleg tibia subapically with 3–4 longer setae on anterior surface.</paragraph>
<paragraph id="8BDA36A498100064BBBE6B6E3B86FC4B" blockId="8.[136,775,807,990]" pageId="8" pageNumber="31">
Abdomen: First to fifth tergites yellow; second tergite with black patches submedially; third to fifth tergites each with medially slightly protruded, sublaterally interrupted, black band on posterior margin; sixth tergite nearly entirely dark, pointed anterolaterally, with a few long strong setae on posterolateral margins (
<figureCitation id="135E2A2198100064BAB16B8E3BBAFC4B" box="[391,472,967,990]" captionStart="Figs" captionStartId="8.[136,180,2035,2056]" captionTargetBox="[142,1479,1066,1985]" captionTargetId="figure-8@8.[124,1512,1005,2022]" captionTargetPageId="8" captionText="Figs. 20–25. Phortica (Phortica) epsilon Chen &amp; Toda, new species, male: 20, sixth tergite in ventral view; 21, epandrium and cercus in lateral view; 22, surstylus in frontal view; 23, tenth sternite and additional plate in frontal view; 24, hypandrium, gonopod, paramere and aedeagus in lateral view; 25, tip of aedeagal median rod in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5333054" httpUri="https://zenodo.org/record/5333054/files/figure.png" pageId="8" pageNumber="31">Fig. 20</figureCitation>
).
</paragraph>
<paragraph id="8BDA36A498100064B87568883F5DFE4D" blockId="8.[835,1474,193,472]" pageId="8" pageNumber="31">
Male terminalia: Epandrium pubescent anteroventrally and dorsomedially, with 12 setae on dorsal to posterolateral portion per side (
<figureCitation id="135E2A2198100064B8C969483E33FE8D" box="[1023,1105,257,280]" captionStart="Figs" captionStartId="8.[136,180,2035,2056]" captionTargetBox="[142,1479,1066,1985]" captionTargetId="figure-8@8.[124,1512,1005,2022]" captionTargetPageId="8" captionText="Figs. 20–25. Phortica (Phortica) epsilon Chen &amp; Toda, new species, male: 20, sixth tergite in ventral view; 21, epandrium and cercus in lateral view; 22, surstylus in frontal view; 23, tenth sternite and additional plate in frontal view; 24, hypandrium, gonopod, paramere and aedeagus in lateral view; 25, tip of aedeagal median rod in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5333054" httpUri="https://zenodo.org/record/5333054/files/figure.png" pageId="8" pageNumber="31">Fig. 21</figureCitation>
). Surstylus pubescent, with dense setae medially and apically rounded prensisetae on inner margin (
<figureCitation id="135E2A2198100064B89669083991FECD" box="[928,1011,321,344]" captionStart="Figs" captionStartId="8.[136,180,2035,2056]" captionTargetBox="[142,1479,1066,1985]" captionTargetId="figure-8@8.[124,1512,1005,2022]" captionTargetPageId="8" captionText="Figs. 20–25. Phortica (Phortica) epsilon Chen &amp; Toda, new species, male: 20, sixth tergite in ventral view; 21, epandrium and cercus in lateral view; 22, surstylus in frontal view; 23, tenth sternite and additional plate in frontal view; 24, hypandrium, gonopod, paramere and aedeagus in lateral view; 25, tip of aedeagal median rod in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5333054" httpUri="https://zenodo.org/record/5333054/files/figure.png" pageId="8" pageNumber="31">Fig. 22</figureCitation>
). Additional plate between cerci and 10th sternite pubescent, medially connected to 10th sternite (
<figureCitation id="135E2A2198100064BEA169283906FE0D" captionStart="Figs" captionStartId="8.[136,180,2035,2056]" captionTargetBox="[142,1479,1066,1985]" captionTargetId="figure-8@8.[124,1512,1005,2022]" captionTargetPageId="8" captionText="Figs. 20–25. Phortica (Phortica) epsilon Chen &amp; Toda, new species, male: 20, sixth tergite in ventral view; 21, epandrium and cercus in lateral view; 22, surstylus in frontal view; 23, tenth sternite and additional plate in frontal view; 24, hypandrium, gonopod, paramere and aedeagus in lateral view; 25, tip of aedeagal median rod in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5333054" httpUri="https://zenodo.org/record/5333054/files/figure.png" pageId="8" pageNumber="31">Fig. 23</figureCitation>
). Posterolateral lobe of hypandrium undeveloped. Paramere with 2 sensilla and 2 pointed teeth apically (
<figureCitation id="135E2A2198100064BEA169E83903FE4D" captionStart="Figs" captionStartId="8.[136,180,2035,2056]" captionTargetBox="[142,1479,1066,1985]" captionTargetId="figure-8@8.[124,1512,1005,2022]" captionTargetPageId="8" captionText="Figs. 20–25. Phortica (Phortica) epsilon Chen &amp; Toda, new species, male: 20, sixth tergite in ventral view; 21, epandrium and cercus in lateral view; 22, surstylus in frontal view; 23, tenth sternite and additional plate in frontal view; 24, hypandrium, gonopod, paramere and aedeagus in lateral view; 25, tip of aedeagal median rod in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5333054" httpUri="https://zenodo.org/record/5333054/files/figure.png" pageId="8" pageNumber="31">Fig. 24</figureCitation>
). Aedeagus with 1 pair of bridges (
<figureCitation id="135E2A2198100064BFD769883F50FE4D" box="[1249,1330,449,472]" captionStart="Figs" captionStartId="8.[136,180,2035,2056]" captionTargetBox="[142,1479,1066,1985]" captionTargetId="figure-8@8.[124,1512,1005,2022]" captionTargetPageId="8" captionText="Figs. 20–25. Phortica (Phortica) epsilon Chen &amp; Toda, new species, male: 20, sixth tergite in ventral view; 21, epandrium and cercus in lateral view; 22, surstylus in frontal view; 23, tenth sternite and additional plate in frontal view; 24, hypandrium, gonopod, paramere and aedeagus in lateral view; 25, tip of aedeagal median rod in ventral view." figureDoi="http://doi.org/10.5281/zenodo.5333054" httpUri="https://zenodo.org/record/5333054/files/figure.png" pageId="8" pageNumber="31">Fig. 24</figureCitation>
).
</paragraph>
<paragraph id="8BDA36A498100064B8756A483E70FD8D" blockId="8.[835,1042,513,536]" box="[835,1042,513,536]" pageId="8" pageNumber="31">Female: Unknown.</paragraph>
<paragraph id="8BDA36A498100064B8756A083ED6FDED" blockId="8.[835,1474,577,792]" pageId="8" pageNumber="31">
Measurements: BL = 4.00 mm in
<typeStatus id="54DE880698100064BF9E6A083F6BFDCD" box="[1192,1289,577,600]" pageId="8" pageNumber="31" type="holotype">holotype</typeStatus>
; ThL =
<quantity id="4C9D9B4198100064BE6A6A083FDCFDCD" box="[1372,1470,577,600]" metricMagnitude="-3" metricUnit="m" metricValue="1.28" pageId="8" pageNumber="31" unit="mm" value="1.28">1.28 mm</quantity>
; WL =
<quantity id="4C9D9B4198100064B8BD6A28398DFDED" box="[907,1007,609,632]" metricMagnitude="-3" metricUnit="m" metricValue="3.18" pageId="8" pageNumber="31" unit="mm" value="3.18">3.18 mm</quantity>
; WW =
<quantity id="4C9D9B4198100064BF7A6A283ED2FDED" box="[1100,1200,609,632]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="31" unit="mm" value="1.4">1.40 mm</quantity>
.
</paragraph>
<paragraph id="8BDA36A498100064B8756AC83E89FC8D" blockId="8.[835,1474,577,792]" pageId="8" pageNumber="31">Indices: arb = 3/0, adf = 1.80, flw = 1.30, FW/HW = 0.45, ch/o = 0.07, prorb = 1.10, rcorb = 0.45, vb = 0.40, dcl = 0.60, presctl = 0.80, sctl = 1.15, sterno = 0.90, orbito = 1.20, dcp = 0.23, sctlp = 1.00, C = 2.05, 4c = 1.47, 4v = 2.33, 5x = 0.70, ac = 3.67, M = 0.47, C3F = 0.73.</paragraph>
</subSubSection>
<subSubSection id="C37F652F98100064B8756B083F66FCED" pageId="8" pageNumber="31" type="etymology">
<paragraph id="8BDA36A498100064B8756B083F66FCED" blockId="8.[835,1474,832,888]" pageId="8" pageNumber="31">
<emphasis id="B911EAB698100064B8756B0839ACFCCD" bold="true" box="[835,974,832,857]" pageId="8" pageNumber="31">
<emphasis id="B911EAB698100064B8756B0839DEFCCC" bold="true" box="[835,956,833,857]" italics="true" pageId="8" pageNumber="31">Etymology.</emphasis>
–
</emphasis>
In reference to the aedeagal basal bridge looking like Greek letter ‘epsilon’ in lateral view.
</paragraph>
</subSubSection>
<subSubSection id="C37F652F98100064B8756BE83ED2FC2D" box="[835,1200,928,953]" pageId="8" pageNumber="31" type="distribution">
<paragraph id="8BDA36A498100064B8756BE83ED2FC2D" blockId="8.[835,1200,928,953]" box="[835,1200,928,953]" pageId="8" pageNumber="31">
<emphasis id="B911EAB698100064B8756BE83986FC2D" bold="true" box="[835,996,928,953]" pageId="8" pageNumber="31">
<emphasis id="B911EAB698100064B8756BE839ADFC2C" bold="true" box="[835,975,929,953]" italics="true" pageId="8" pageNumber="31">Distribution.</emphasis>
–
</emphasis>
<collectingCountry id="F372763498100064B8DA6BE83E2CFC2D" box="[1004,1102,929,952]" name="Malaysia" pageId="8" pageNumber="31">Malaysia</collectingCountry>
(
<collectingRegion id="49A1F84698100064BF686BE83EC7FC2D" box="[1118,1189,929,952]" country="Malaysia" name="Sabah" pageId="8" pageNumber="31">Sabah</collectingRegion>
).
</paragraph>
</subSubSection>
<caption id="DF1A662C98100064BBBE6FBA392EF7D7" ID-DOI="http://doi.org/10.5281/zenodo.5333054" ID-Zenodo-Dep="5333054" httpUri="https://zenodo.org/record/5333054/files/figure.png" pageId="8" pageNumber="31" startId="8.[136,180,2035,2056]" targetBox="[142,1479,1066,1985]" targetPageId="8">
<paragraph id="8BDA36A498100064BBBE6FBA392EF7D7" blockId="8.[136,1476,2035,2114]" pageId="8" pageNumber="31">
Figs. 20–25.
<taxonomicName id="4C654D2798100064BA3F6FBD38C6F79D" authority="Chen &amp; Toda" authorityName="Chen &amp; Toda" authorityYear="2007" box="[265,676,2035,2057]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="epsilon" status="sp. nov." subGenus="Phortica">
<emphasis id="B911EAB698100064BA3F6FBD3B3FF79C" box="[265,349,2036,2057]" italics="true" pageId="8" pageNumber="31">Phortica</emphasis>
(
<emphasis id="B911EAB698100064BA5A6FBD3BA2F79C" box="[364,448,2036,2057]" italics="true" pageId="8" pageNumber="31">Phortica</emphasis>
)
<emphasis id="B911EAB698100064BAF96FBD3876F79C" box="[463,532,2036,2057]" italics="true" pageId="8" pageNumber="31">epsilon</emphasis>
Chen &amp; Toda
</taxonomicName>
,
<taxonomicNameLabel id="A22257CD98100064B9996FBA3944F79D" box="[687,806,2035,2056]" pageId="8" pageNumber="31" rank="species">new species</taxonomicNameLabel>
, male: 20, sixth tergite in ventral view; 21, epandrium and cercus in lateral view; 22, surstylus in frontal view; 23, tenth sternite and additional plate in frontal view; 24, hypandrium, gonopod, paramere and aedeagus in lateral view; 25, tip of aedeagal median rod in ventral view.
</paragraph>
</caption>
<subSubSection id="C37F652F98110065BB4768883854FEED" pageId="9" pageNumber="32" type="discussion">
<paragraph id="8BDA36A498110065BB4768883854FEED" blockId="9.[113,752,192,376]" pageId="9" pageNumber="32">
<emphasis id="B911EAB698110065BB4768883AB8FF4D" box="[113,218,193,217]" italics="true" pageId="9" pageNumber="32">
<emphasis id="B911EAB698110065BB4768883AB1FF4C" bold="true" box="[113,211,193,217]" italics="true" pageId="9" pageNumber="32">Remarks</emphasis>
.
</emphasis>
<emphasis id="B911EAB698110065BBD568893A92FF4D" bold="true" box="[227,240,192,216]" pageId="9" pageNumber="32">–</emphasis>
This species is very similar to
<emphasis id="B911EAB698110065B96768883803FF4D" box="[593,609,193,216]" italics="true" pageId="9" pageNumber="32">P</emphasis>
.
<taxonomicName id="4C654D2798110065B94468883B6FFF6D" authority="(Okada, 1977)" baseAuthorityName="Okada" baseAuthorityYear="1977" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="cardua" subGenus="Phortica">
(
<emphasis id="B911EAB698110065B94D688838E9FF4D" box="[635,651,193,216]" italics="true" pageId="9" pageNumber="32">P</emphasis>
.)
<emphasis id="B911EAB698110065B99268883892FF4D" box="[676,752,193,216]" italics="true" pageId="9" pageNumber="32">cardua</emphasis>
(
<bibRefCitation id="EFF44B5598110065BB4E68A83B64FF6D" author="Okada, T" box="[120,262,225,248]" pageId="9" pageNumber="32" pagination="17 - 31" refId="ref13323" refString="Okada, T., 1977. The subgenus Phortica Schiner of the genus Amiota Loew of Japan and the Oriental Region, with reference to anti- Burla's rule (Diptera, Drosophilidae). Bulletin of the Biogeographical Society of Japan, 32: 17 - 31." type="journal article" year="1977">Okada, 1977</bibRefCitation>
)
</taxonomicName>
in that the arista lacks ventral branches and the anepisternum lacking setulae, but can be distinguished from it by the aedeagal median rod and the paramere (in
<taxonomicName id="4C654D2798110065B9EF69683AA7FECD" baseAuthorityName="Okada" baseAuthorityYear="1977" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="cardua">
<emphasis id="B911EAB698110065B9EF69683AA7FECD" italics="true" pageId="9" pageNumber="32">P. cardua</emphasis>
</taxonomicName>
: aedeagal median rod slender, unexpanded subapically; paramere pubescent distally).
</paragraph>
</subSubSection>
</treatment>
</document>
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