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<document id="4D0189577B6C76DBC6AC95DD04B1A661" ID-DOI="10.5281/zenodo.275727" ID-GBIF-Dataset="1e79f840-6ced-48b5-ab20-94992c6f0a88" ID-ISSN="1175-5326" ID-Zenodo-Dep="275727" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460160971924" checkinUser="plazi" docAuthor="West, Rick C. &amp; Nunn, Steven C." docDate="2010" docId="03CC87B17645FFB69293FDF7FF78D10A" docLanguage="en" docName="zt02362p043.pdf" docOrigin="Zootaxa 2362" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Lyrognathus lessunda West &amp; Nunn, 2010, sp. nov." docType="treatment" docVersion="8" lastPageNumber="31" masterDocId="FFF5FFC9765DFFA89204FFFAFF92D456" masterDocTitle="A taxonomic revision of the tarantula spider genus Lyrognathus Pocock 1895 (Araneae, Theraphosidae), with notes on the Selenocosmiinae" masterLastPageNumber="43" masterPageNumber="1" pageNumber="25" updateTime="1698235793967" updateUser="plazi">
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<mods:title id="C3B56455099D75D1FC72EE6B8639C1E4">A taxonomic revision of the tarantula spider genus Lyrognathus Pocock 1895 (Araneae, Theraphosidae), with notes on the Selenocosmiinae</mods:title>
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<mods:namePart id="CFF5E636C78313E3A884B88291D6B495">West, Rick C.</mods:namePart>
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<mods:namePart id="0012412224B454435F334B93CBC82A3C">Nunn, Steven C.</mods:namePart>
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<mods:date id="802BD8BB2089C837B4DD43E241CD400E">2010</mods:date>
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<treatment id="03CC87B17645FFB69293FDF7FF78D10A" ID-DOI="http://doi.org/10.5281/zenodo.6210156" ID-GBIF-Taxon="127654812" ID-Zenodo-Dep="6210156" LSID="urn:lsid:plazi:treatment:03CC87B17645FFB69293FDF7FF78D10A" httpUri="http://treatment.plazi.org/id/03CC87B17645FFB69293FDF7FF78D10A" lastPageId="30" lastPageNumber="31" pageId="24" pageNumber="25">
<subSubSection id="C37F652C7645FFB09293FDF7FEF5D61C" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="8BDA36A77645FFB09293FDF7FD94D671" blockId="24.[151,518,525,586]" box="[151,518,525,551]" pageId="24" pageNumber="25">
<heading id="D09281CB7645FFB09293FDF7FD94D671" bold="true" box="[151,518,525,551]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
<emphasis id="B911EAB57645FFB09293FDF7FD94D671" bold="true" box="[151,518,525,551]" pageId="24" pageNumber="25">
<taxonomicName id="4C654D247645FFB09293FDF7FE30D671" ID-CoL="3WTQM" box="[151,418,525,551]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="lessunda" status="sp. nov.">
<emphasis id="B911EAB57645FFB09293FDF7FE30D671" bold="true" box="[151,418,525,551]" italics="true" pageId="24" pageNumber="25">Lyrognathus lessunda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257CE7645FFB093ADFDF7FD94D671" box="[425,518,525,551]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA36A77645FFB09293FDCAFEF5D61C" blockId="24.[151,518,525,586]" box="[151,359,560,586]" pageId="24" pageNumber="25">
(
<figureCitation id="135E2A227645FFB0929BFDCAFEB4D61C" box="[159,294,560,586]" captionStart-0="FIGURE 63" captionStart-1="FIGURES 64 70" captionStart-2="FIGURES 71 76" captionStart-3="FIGURES 77 80" captionStart-4="FIGURES 81 90" captionStartId-0="24.[151,255,1755,1779]" captionStartId-1="25.[151,269,1653,1677]" captionStartId-2="26.[151,270,1906,1930]" captionStartId-3="27.[151,269,1142,1166]" captionStartId-4="29.[151,271,1676,1700]" captionTargetBox-0="[151,1435,853,1732]" captionTargetBox-1="[163,1423,343,1632]" captionTargetBox-2="[165,1409,415,1873]" captionTargetBox-3="[152,1434,194,1119]" captionTargetBox-4="[151,1434,412,1651]" captionTargetId-0="figure@24.[151,1436,853,1734]" captionTargetId-1="figure@25.[163,1424,343,1632]" captionTargetId-2="figure@26.[165,1409,415,1874]" captionTargetId-3="figure@27.[152,1435,194,1121]" captionTargetId-4="figure@29.[151,1436,412,1652]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionTargetPageId-2="26" captionTargetPageId-3="27" captionTargetPageId-4="29" captionText-0="FIGURE 63. Lyrognathus lessunda sp. nov., holotype, female in life. Photo: R. C. West." captionText-1="FIGURES 64 70. Lyrognathus lessunda sp. nov. holotype female: 64. Carapace, dorsal view. 65. Eyes, dorsal view. 66. Sternum, labium, maxillae and coxa, ventral view. 67. Labium, ventral view. 68. Left palp, legs I, II, ventral view. 69. Abdomen, legs IV, ventral view. 70. Spermathecae, dorsal view. Scale bar = 10 mm for 64, 66, 69; 5 mm for 68; 2 mm for 65, 67, 70. Photos: S. C. Nunn." captionText-2="FIGURES 71 76. Lyrognathus lessunda sp. nov., holotype female (except: paratype female QM S 74066; 75, 76): 71. Abdomen and legs IV, dorsal view. 72. Left leg IV, retrolateral view. 73. Left leg IV, prolateral view. 74. Left met. and tar. IV, ventral view. 75. Left maxillae, prolateral view. 76. Left maxillary lyra, prolateral view. Scale bar = 10 mm for 71 73; 5 mm for 74; 2 mm for 75; 1 mm for 76. Photos: S. C. Nunn." captionText-3="FIGURES 77 80. Lyrognathus lessunda sp. nov. paratype female QM S 74066: 77. Left chelicerae, prolateral view. 78. Left intercheliceral pegs, prolateral view. 79. Left cheliceral strikers, ectal view. 80. Left basomedial cheliceral spines, retrolateral view. Scale bar = 2 mm for 77; 1 mm for 78, 79; 0.5 mm for 80. Photos: S. C. Nunn." captionText-4="FIGURES 81 90. Lyrognathus lessunda sp. nov. paratype male QM S 74069: 81. Habitus, dorsal view. 82. Carapace, dorsal view. 83. Eyes, dorsal view. 84. Sternum, labium, maxillae and coxa, ventral view. 85. Right intercheliceral pegs, prolateral view. 86. Right cheliceral strikers, ectal view. 87. Right maxillae showing lyra, prolateral view. 88. Left bulb, retrolateral view. 89. Left bulb, prolateral view. 90. Left bulb, anterior view. Scale bar = 10 mm for 81; 5 mm for 82, 84; 2 mm for 83, 86, 87; 1 mm for 85, 88 90. Photos: S. C. Nunn." httpUri-0="https://zenodo.org/record/275742/files/figure.png" httpUri-1="https://zenodo.org/record/275743/files/figure.png" httpUri-2="https://zenodo.org/record/275744/files/figure.png" httpUri-3="https://zenodo.org/record/275745/files/figure.png" httpUri-4="https://zenodo.org/record/275746/files/figure.png" pageId="24" pageNumber="25">Figs 6383</figureCitation>
,
<figureCitation id="135E2A227645FFB09336FDCAFECCD61C" box="[306,350,560,586]" captionStart="FIGURE 114" captionStartId="38.[151,255,1297,1321]" captionTargetBox="[151,1436,194,1276]" captionTargetId="figure@38.[151,1436,194,1276]" captionTargetPageId="38" captionText="FIGURE 114. Records of Lyrognathus species in Southeast Asia. C. Senescall" httpUri="https://zenodo.org/record/275751/files/figure.png" pageId="24" pageNumber="25">114</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37F652C7645FFB09293FD80FE3CD689" pageId="24" pageNumber="25" type="description">
<paragraph id="8BDA36A77645FFB09293FD80FE3CD689" blockId="24.[151,1437,634,810]" pageId="24" pageNumber="25">
<emphasis id="B911EAB57645FFB09293FD80FF62D6C2" bold="true" box="[151,240,634,660]" pageId="24" pageNumber="25">
<typeStatus id="54DE88057645FFB09293FD80FF71D6C2" box="[151,227,634,660]" pageId="24" pageNumber="25">Types</typeStatus>
:
</emphasis>
female
<typeStatus id="54DE88057645FFB0935CFD80FE58D6C2" box="[344,458,634,660]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
,
<specimenCount id="9D63FD2E7645FFB093DCFD80FDD8D6C2" box="[472,586,634,660]" pageId="24" pageNumber="25" type="female">1 female</specimenCount>
,
<specimenCount id="9D63FD2E7645FFB0905CFD80FD3CD6C2" box="[600,686,634,660]" pageId="24" pageNumber="25" type="male">3 male</specimenCount>
<typeStatus id="54DE88057645FFB090B3FD80FCA6D6C2" box="[695,820,634,660]" pageId="24" pageNumber="25" type="paratype">paratypes</typeStatus>
, S74067, S74066,
<accessionNumber id="9436AB447645FFB09620FD80FB1AD6C2" box="[1060,1160,634,660]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S74070" pageId="24" pageNumber="25" type="EnaNcbi">S74070</accessionNumber>
, S74068 and
<accessionNumber id="9436AB447645FFB09731FD80FA0AD6C2" box="[1333,1432,634,660]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S74069" pageId="24" pageNumber="25" type="EnaNcbi">S74069</accessionNumber>
, respectively, Lombok Island (
<geoCoordinate id="EE5150607645FFB093FEFD5AFDC0D6EC" box="[506,594,672,698]" direction="south" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="-8.133333">8°08' S</geoCoordinate>
,
<geoCoordinate id="EE5150607645FFB09065FD5AFD4AD6EC" box="[609,728,672,698]" direction="east" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="117.4">117°24' E</geoCoordinate>
), Lesser Sunda Islands,
<collectingCountry id="F37276377645FFB091FEFD5AFBE2D6EC" box="[1018,1136,672,698]" name="Indonesia" pageId="24" pageNumber="25">Indonesia</collectingCountry>
, col. B. Sutrino,
<date id="FFDB10677645FFB09738FD5AFF47D689" pageId="24" pageNumber="25" value="2002-08-15">15 Aug. 2002</date>
, QM.—examined.
</paragraph>
</subSubSection>
<subSubSection id="C37F652C7645FFB092C2FD10FD2AD77C" pageId="24" pageNumber="25" type="etymology">
<paragraph id="8BDA36A77645FFB092C2FD10FD2AD77C" blockId="24.[151,1437,634,810]" pageId="24" pageNumber="25">
<emphasis id="B911EAB57645FFB092C2FD10FEC5D752" bold="true" box="[198,343,746,772]" pageId="24" pageNumber="25">Etymology:</emphasis>
The specific name is a contraction of Lesser Sunda Islands in the Malay Archipelago of
<collectingCountry id="F37276377645FFB09293FCEAFE9BD77C" box="[151,265,784,810]" name="Indonesia" pageId="24" pageNumber="25">Indonesia</collectingCountry>
and, therefore, a noun in apposition.
</paragraph>
</subSubSection>
<caption id="DF1A662F7645FFB09293F921FBDED2A4" box="[151,1100,1755,1779]" httpUri="https://zenodo.org/record/275742/files/figure.png" pageId="24" pageNumber="25" targetBox="[151,1435,853,1732]" targetPageId="24">
<paragraph id="8BDA36A77645FFB09293F921FBDED2A4" blockId="24.[151,1100,1755,1779]" box="[151,1100,1755,1779]" pageId="24" pageNumber="25">
<emphasis id="B911EAB57645FFB09293F921FEB2D2A5" bold="true" box="[151,288,1755,1779]" pageId="24" pageNumber="25">FIGURE 63</emphasis>
.
<taxonomicName id="4C654D247645FFB09329F921FD84D2A4" box="[301,534,1755,1778]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="lessunda" status="sp. nov.">
<emphasis id="B911EAB57645FFB09329F921FD84D2A4" box="[301,534,1755,1778]" italics="true" pageId="24" pageNumber="25">Lyrognathus lessunda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257CE7645FFB09018F921FDE0D2A5" box="[540,626,1755,1779]" pageId="24" pageNumber="25" rank="species">
<emphasis id="B911EAB57645FFB09018F921FDE0D2A5" bold="true" box="[540,626,1755,1779]" pageId="24" pageNumber="25">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, female in life. Photo: R. C. West.
</paragraph>
</caption>
<subSubSection id="C37F652C7645FFB192C2F8DAFF7AD574" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="8BDA36A77645FFB092C2F8DAFA0ED3A3" blockId="24.[151,1437,1824,2037]" pageId="24" pageNumber="25">
<emphasis id="B911EAB57645FFB092C2F8DAFEDAD36C" bold="true" box="[198,328,1824,1850]" pageId="24" pageNumber="25">Diagnosis:</emphasis>
Differs from all other
<taxonomicName id="4C654D247645FFB09051F8DBFD75D36C" box="[597,743,1825,1850]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B911EAB57645FFB09051F8DBFD75D36C" box="[597,743,1825,1850]" italics="true" pageId="24" pageNumber="25">Lyrognathus</emphasis>
</taxonomicName>
species in the apically swollen “mushroom cap” that form bilobular spermathecae, present on both lateral and medial lobes in females and undivided scopula on tar. IV. Differs from
<taxonomicName id="4C654D247645FFB09345F891FE5ED3D2" box="[321,460,1899,1924]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="crotalus">
<emphasis id="B911EAB57645FFB09345F891FE5ED3D2" box="[321,460,1899,1924]" italics="true" pageId="24" pageNumber="25">L. crotalus</emphasis>
</taxonomicName>
in intercheliceral pegs (not spines), bilobular spermathecae and retrolateral basomedial cheliceral spines.
<taxonomicName id="4C654D247645FFB093F1F86BFD64D3FC" box="[501,758,1937,1962]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="lessunda" status="sp. nov.">
<emphasis id="B911EAB57645FFB093F1F86BFD64D3FC" box="[501,758,1937,1962]" italics="true" pageId="24" pageNumber="25">Lyrognathus lessunda</emphasis>
</taxonomicName>
<emphasis id="B911EAB57645FFB090F9F86AFCC9D3FC" bold="true" box="[765,859,1936,1962]" pageId="24" pageNumber="25">
<taxonomicNameLabel id="A22257CE7645FFB090F9F86AFCC9D3FC" box="[765,859,1936,1962]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles
<taxonomicName id="4C654D247645FFB091DBF86BFBC8D3FC" box="[991,1114,1937,1962]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="achilles" status="sp. nov.">
<emphasis id="B911EAB57645FFB091DBF86BFBC8D3FC" box="[991,1114,1937,1962]" italics="true" pageId="24" pageNumber="25">L. achilles</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257CE7645FFB09665F86AFB2DD3FC" box="[1121,1215,1936,1962]" pageId="24" pageNumber="25" rank="species">
<emphasis id="B911EAB57645FFB09665F86AFB2DD3FC" bold="true" box="[1121,1215,1936,1962]" pageId="24" pageNumber="25">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="4C654D247645FFB096CAF86BFAA4D3FC" box="[1230,1334,1937,1962]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fuscus" status="sp. nov.">
<emphasis id="B911EAB57645FFB096CAF86BFAA4D3FC" box="[1230,1334,1937,1962]" italics="true" pageId="24" pageNumber="25">L. fuscus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257CE7645FFB0973AF86AFA09D3FC" box="[1342,1435,1936,1962]" pageId="24" pageNumber="25" rank="species">
<emphasis id="B911EAB57645FFB0973AF86AFA09D3FC" bold="true" box="[1342,1435,1936,1962]" pageId="24" pageNumber="25">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C654D247645FFB092CBF84CFEF1D399" box="[207,355,1974,1999]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B911EAB57645FFB092CBF84CFEF1D399" box="[207,355,1974,1999]" italics="true" pageId="24" pageNumber="25">L. robustus</emphasis>
</taxonomicName>
in having intercheliceral pegs (females and males) in combination with bilobular spermathecae and retrolateral basomedial cheliceral spines. Differs from
<taxonomicName id="4C654D247645FFB091F4F821FBF8D3A2" box="[1008,1130,2011,2036]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="achilles" status="sp. nov.">
<emphasis id="B911EAB57645FFB091F4F821FBF8D3A2" box="[1008,1130,2011,2036]" italics="true" pageId="24" pageNumber="25">L. achilles</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257CE7645FFB09676F821FB42D3A3" box="[1138,1232,2011,2037]" pageId="24" pageNumber="25" rank="species">
<emphasis id="B911EAB57645FFB09676F821FB42D3A3" bold="true" box="[1138,1232,2011,2037]" pageId="24" pageNumber="25">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C654D247645FFB0970DF821FAE3D3A2" box="[1289,1393,2011,2036]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fuscus" status="sp.">
<emphasis id="B911EAB57645FFB0970DF821FAE3D3A2" box="[1289,1393,2011,2036]" italics="true" pageId="24" pageNumber="25">L. fuscus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257CE7645FFB0977DF821FA0ED3A3" box="[1401,1436,2011,2037]" pageId="24" pageNumber="25">
<emphasis id="B911EAB57645FFB0977DF821FA0ED3A3" bold="true" box="[1401,1436,2011,2037]" pageId="24" pageNumber="25">sp.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BDA36A77644FFB19293FF6DFF7AD574" blockId="25.[151,1437,151,290]" pageId="25" pageNumber="26">
<emphasis id="B911EAB57644FFB19293FF6DFF5FD4E7" bold="true" box="[151,205,151,177]" pageId="25" pageNumber="26">nov.</emphasis>
in the apical “mushroom cap” form medial lobes of spermathecae. Differs from
<taxonomicName id="4C654D247644FFB196C5FF62FADFD4E7" box="[1217,1357,152,177]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B911EAB57644FFB196C5FF62FADFD4E7" box="[1217,1357,152,177]" italics="true" pageId="25" pageNumber="26">L. robustus</emphasis>
</taxonomicName>
in the undivided metatarsi IV scopula for the full length ventrally and the absence of a third claw on tarsi IV. Differs from
<taxonomicName id="4C654D247644FFB192DCFF18FEC6D4AD" box="[216,340,226,251]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="saltator">
<emphasis id="B911EAB57644FFB192DCFF18FEC6D4AD" box="[216,340,226,251]" italics="true" pageId="25" pageNumber="26">L. saltator</emphasis>
</taxonomicName>
in the undivided tarsi IV scopula, intercheliceral pegs and retrolateral basomedial cheliceral spines.
</paragraph>
</subSubSection>
<caption id="DF1A662F7644FFB19293F98FFE71D2BB" httpUri="https://zenodo.org/record/275743/files/figure.png" pageId="25" pageNumber="26" targetBox="[163,1423,343,1632]" targetPageId="25">
<paragraph id="8BDA36A77644FFB19293F98FFE71D2BB" blockId="25.[151,1436,1653,1773]" pageId="25" pageNumber="26">
<emphasis id="B911EAB57644FFB19293F98FFECFD2DB" bold="true" box="[151,349,1653,1677]" pageId="25" pageNumber="26">FIGURES 6470.</emphasis>
<taxonomicName id="4C654D247644FFB19360F98CFDDFD2DB" box="[356,589,1654,1677]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="lessunda" status="sp. nov.">
<emphasis id="B911EAB57644FFB19360F98CFDDFD2DB" box="[356,589,1654,1677]" italics="true" pageId="25" pageNumber="26">Lyrognathus lessunda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257CE7644FFB19050F98FFD3BD2DB" box="[596,681,1653,1677]" pageId="25" pageNumber="26" rank="species">
<emphasis id="B911EAB57644FFB19050F98FFD3BD2DB" bold="true" box="[596,681,1653,1677]" pageId="25" pageNumber="26">sp. nov.</emphasis>
</taxonomicNameLabel>
holotype female: 64. Carapace, dorsal view. 65. Eyes, dorsal view. 66. Sternum, labium, maxillae and coxa, ventral view. 67. Labium, ventral view. 68. Left palp, legs I, II, ventral view. 69. Abdomen, legs IV, ventral view. 70. Spermathecae, dorsal view. Scale bar = 10mm for 64, 66, 69; 5mm for 68; 2mm for 65, 67, 70. Photos: S. C. Nunn.
</paragraph>
</caption>
<subSubSection id="C37F652C7644FFB692C0F8E1FBB8D064" lastPageId="30" lastPageNumber="31" pageId="25" pageNumber="26" type="description">
<paragraph id="8BDA36A77644FFB192C0F8E1FB80D363" blockId="25.[151,1437,1819,2032]" box="[196,1042,1819,1845]" pageId="25" pageNumber="26">
<emphasis id="B911EAB57644FFB192C0F8E1FECCD363" bold="true" box="[196,350,1819,1845]" pageId="25" pageNumber="26">Description:</emphasis>
Female
<typeStatus id="54DE88057644FFB193C7F8E1FDB8D363" box="[451,554,1819,1845]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
(in life,
<figureCitation id="135E2A227644FFB1908BF8E1FD70D363" box="[655,738,1819,1845]" captionStart="FIGURE 63" captionStartId="24.[151,255,1755,1779]" captionTargetBox="[151,1435,853,1732]" captionTargetId="figure@24.[151,1436,853,1734]" captionTargetPageId="24" captionText="FIGURE 63. Lyrognathus lessunda sp. nov., holotype, female in life. Photo: R. C. West." httpUri="https://zenodo.org/record/275742/files/figure.png" pageId="25" pageNumber="26">Fig. 63</figureCitation>
) with body length: 32.91.
</paragraph>
<paragraph id="8BDA36A77644FFB192C0F8BBFD88D3D6" blockId="25.[151,1437,1819,2032]" pageId="25" pageNumber="26">Color (in life): intermolt and premolt entirely reddish brown/rust, postmolt dark brown posteriorly, shading to rust orange anteriorly.</paragraph>
<paragraph id="8BDA36A77644FFB292C0F871FE6ED53A" blockId="25.[151,1437,1819,2032]" lastBlockId="26.[151,1436,152,364]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
Carapace (
<figureCitation id="135E2A227644FFB19341F871FE30D3F3" box="[325,418,1931,1957]" captionStart="FIGURES 64 70" captionStartId="25.[151,269,1653,1677]" captionTargetBox="[163,1423,343,1632]" captionTargetId="figure@25.[163,1424,343,1632]" captionTargetPageId="25" captionText="FIGURES 64 70. Lyrognathus lessunda sp. nov. holotype female: 64. Carapace, dorsal view. 65. Eyes, dorsal view. 66. Sternum, labium, maxillae and coxa, ventral view. 67. Labium, ventral view. 68. Left palp, legs I, II, ventral view. 69. Abdomen, legs IV, ventral view. 70. Spermathecae, dorsal view. Scale bar = 10 mm for 64, 66, 69; 5 mm for 68; 2 mm for 65, 67, 70. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275743/files/figure.png" pageId="25" pageNumber="26">Fig. 64</figureCitation>
): length 13.3, width 11.7 (width of anterior edge 8.7). Fovea 2.15, procurved, deep, slightly wider than OT (
<figureCitation id="135E2A227644FFB193BEF84BFD86D39D" box="[442,532,1969,1995]" captionStart="FIGURES 20 23" captionStartId="13.[151,270,1796,1820]" captionTargetBox="[151,1436,343,1773]" captionTargetId="figure@13.[151,1436,343,1775]" captionTargetPageId="13" captionText="FIGURES 20 23. Lyrognathus achilles sp. nov., holotype female: 20. Leg IV, dorsal view. 21. Leg IV, prolateral view. 22. Tar. IV, ventral view. 23. Spermathecae, dorsal view. Scale bar = 10 mm for 20, 21; 5 mm for 22; 2 mm for 23. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275733/files/figure.png" pageId="25" pageNumber="26">Fig. 21</figureCitation>
). Carapace with 4 discernible hair
<typeStatus id="54DE88057644FFB191BDF84BFC6FD39D" box="[953,1021,1969,1995]" pageId="25" pageNumber="26">types</typeStatus>
: 4 distinct
<typeStatus id="54DE88057644FFB19684F84BFB2DD39D" box="[1152,1215,1969,1995]" pageId="25" pageNumber="26">types</typeStatus>
noted (C1 to C4):
<typeStatus id="54DE88057644FFB19293F82CFF59D3A6" box="[151,203,2006,2032]" pageId="25" pageNumber="26">type</typeStatus>
C1, long, needle-form pallids border carapace entirely (darker basally):
<typeStatus id="54DE88057644FFB1964EF82CFBECD3A6" box="[1098,1150,2006,2032]" pageId="25" pageNumber="26">type</typeStatus>
C2, short blunt tipped, basally dark gray to distally pallid, cover entire carapace evenly, but reasonably sparse:
<typeStatus id="54DE88057647FFB296B1FF62FB7AD4E4" box="[1205,1256,152,178]" pageId="26" pageNumber="27">type</typeStatus>
C3, short, thin wavy pallids, form basis of carapace mat:
<typeStatus id="54DE88057647FFB2908DFF47FD2ED481" box="[649,700,189,215]" pageId="26" pageNumber="27">type</typeStatus>
C4, emerge from bothria-like collars, long, distally spiniform, pallid, lie in uniform arrangement of (&gt;4) along anterior edge of each radial groove, (&gt;4) along anterior edge of each thoracic groove, (&gt;6) along anterior edge of each sulcal groove, (&gt;8) along anterior edge of each caput groove and line of&gt;4 along medial caput ridge (between OT and fovea).&gt;4 more located behind each side of OT, nearest median caput row.
</paragraph>
<caption id="DF1A662F7647FFB29293F888FC9CD3BF" httpUri="https://zenodo.org/record/275744/files/figure.png" pageId="26" pageNumber="27" targetBox="[165,1409,415,1873]" targetPageId="26">
<paragraph id="8BDA36A77647FFB29293F888FC9CD3BF" blockId="26.[151,1437,1906,2025]" pageId="26" pageNumber="27">
<emphasis id="B911EAB57647FFB29293F888FEF2D3DC" bold="true" box="[151,352,1906,1930]" pageId="26" pageNumber="27">FIGURES 7176.</emphasis>
<taxonomicName id="4C654D247647FFB29363F888FDC1D3DF" box="[359,595,1906,1929]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="lessunda" status="sp. nov.">
<emphasis id="B911EAB57647FFB29363F888FDC1D3DF" box="[359,595,1906,1929]" italics="true" pageId="26" pageNumber="27">Lyrognathus lessunda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257CE7647FFB2905FF888FD20D3DC" box="[603,690,1906,1930]" pageId="26" pageNumber="27" rank="species">
<emphasis id="B911EAB57647FFB2905FF888FD20D3DC" bold="true" box="[603,690,1906,1930]" pageId="26" pageNumber="27">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype female (except: paratype female QM S74066; 75, 76): 71. Abdomen and legs IV, dorsal view. 72. Left leg IV, retrolateral view. 73. Left leg IV, prolateral view. 74. Left met. and tar. IV, ventral view. 75. Left maxillae, prolateral view. 76. Left maxillary
<taxonomicName id="4C654D247647FFB29186F848FC3CD39F" box="[898,942,1970,1993]" class="Arachnida" family="Theraphosidae" genus="Selenocosmia" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="lyra">lyra</taxonomicName>
, prolateral view. Scale bar = 10mm for 7173; 5mm for 74; 2mm for 75; 1mm for 76. Photos: S. C. Nunn.
</paragraph>
</caption>
<caption id="DF1A662F7646FFB39293FB8CFB16D09B" httpUri="https://zenodo.org/record/275745/files/figure.png" pageId="27" pageNumber="28" targetBox="[152,1434,194,1119]" targetPageId="27">
<paragraph id="8BDA36A77646FFB39293FB8CFB16D09B" blockId="27.[151,1436,1142,1229]" pageId="27" pageNumber="28">
<emphasis id="B911EAB57646FFB39293FB8CFECCD0D8" bold="true" box="[151,350,1142,1166]" pageId="27" pageNumber="28">FIGURES 7780.</emphasis>
<taxonomicName id="4C654D247646FFB39361FB8DFDDCD0D8" box="[357,590,1143,1166]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="lessunda" status="sp. nov.">
<emphasis id="B911EAB57646FFB39361FB8DFDDCD0D8" box="[357,590,1143,1166]" italics="true" pageId="27" pageNumber="28">Lyrognathus lessunda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257CE7646FFB39051FB8CFD3ED0D8" box="[597,684,1142,1166]" pageId="27" pageNumber="28" rank="species">
<emphasis id="B911EAB57646FFB39051FB8CFD3ED0D8" bold="true" box="[597,684,1142,1166]" pageId="27" pageNumber="28">sp. nov.</emphasis>
</taxonomicNameLabel>
paratype female QM S74066: 77. Left chelicerae, prolateral view. 78. Left intercheliceral pegs, prolateral view. 79. Left cheliceral strikers, ectal view. 80. Left basomedial cheliceral spines, retrolateral view. Scale bar = 2mm for 77; 1mm for 78, 79; 0.5mm for 80. Photos: S. C. Nunn.
</paragraph>
</caption>
<paragraph id="8BDA36A77646FFB392C0FB06FCF3D1FD" blockId="27.[151,1437,1276,2011]" pageId="27" pageNumber="28">
Eyes (
<figureCitation id="135E2A227646FFB39314FB06FEFED140" box="[272,364,1276,1302]" captionStart="FIGURES 64 70" captionStartId="25.[151,269,1653,1677]" captionTargetBox="[163,1423,343,1632]" captionTargetId="figure@25.[163,1424,343,1632]" captionTargetPageId="25" captionText="FIGURES 64 70. Lyrognathus lessunda sp. nov. holotype female: 64. Carapace, dorsal view. 65. Eyes, dorsal view. 66. Sternum, labium, maxillae and coxa, ventral view. 67. Labium, ventral view. 68. Left palp, legs I, II, ventral view. 69. Abdomen, legs IV, ventral view. 70. Spermathecae, dorsal view. Scale bar = 10 mm for 64, 66, 69; 5 mm for 68; 2 mm for 65, 67, 70. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275743/files/figure.png" pageId="27" pageNumber="28">Fig. 65</figureCitation>
): ocular tubercle; length 2.0, width 2.35. Anterior row transverse. Posterior row slightly recurved. Eyes: AME, ALE, PLE = PME. OT hairs:&gt;12 (C2) directly posterior to AMEs, 4 to 5 between PME and center of OT (directly behind each AME). 18 (C2) along median anterior OT surface. OT entirely covered in small, weak, pallid setae (C3), point to center of OT, posteriorly and anteriorly. Lateral small pallids sparse with no uniform direction/random arrangment.
</paragraph>
<paragraph id="8BDA36A77646FFB392C0FA4CFE18D2E6" blockId="27.[151,1437,1276,2011]" pageId="27" pageNumber="28">
Chelicerae (
<figureCitation id="135E2A227646FFB39354FA4CFE49D186" box="[336,475,1462,1488]" captionStart="FIGURES 77 80" captionStartId="27.[151,269,1142,1166]" captionTargetBox="[152,1434,194,1119]" captionTargetId="figure@27.[152,1435,194,1121]" captionTargetPageId="27" captionText="FIGURES 77 80. Lyrognathus lessunda sp. nov. paratype female QM S 74066: 77. Left chelicerae, prolateral view. 78. Left intercheliceral pegs, prolateral view. 79. Left cheliceral strikers, ectal view. 80. Left basomedial cheliceral spines, retrolateral view. Scale bar = 2 mm for 77; 1 mm for 78, 79; 0.5 mm for 80. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275745/files/figure.png" pageId="27" pageNumber="28">Figs 7780</figureCitation>
): length 4.0, width at base of each lobe 2.85. Ectal lyrate region a series of strikers (&gt;50) up to 7 horizontal rows (
<figureCitation id="135E2A227646FFB3900BFA26FDF9D1A0" box="[527,619,1500,1526]" captionStart="FIGURES 77 80" captionStartId="27.[151,269,1142,1166]" captionTargetBox="[152,1434,194,1119]" captionTargetId="figure@27.[152,1435,194,1121]" captionTargetPageId="27" captionText="FIGURES 77 80. Lyrognathus lessunda sp. nov. paratype female QM S 74066: 77. Left chelicerae, prolateral view. 78. Left intercheliceral pegs, prolateral view. 79. Left cheliceral strikers, ectal view. 80. Left basomedial cheliceral spines, retrolateral view. Scale bar = 2 mm for 77; 1 mm for 78, 79; 0.5 mm for 80. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275745/files/figure.png" pageId="27" pageNumber="28">Fig. 79</figureCitation>
). Strongest and longest strikers on lowest rows. Each striker blade/ scimitar shape, with some filiform ends. Teeth, 14,&gt;25 basomesal teeth. Intercheliceral pegs (&gt;16 larger,&gt;23 smaller) in tight cluster on basodorsal surface (
<figureCitation id="135E2A227646FFB390D7F9DCFCBDD216" box="[723,815,1574,1600]" captionStart="FIGURES 77 80" captionStartId="27.[151,269,1142,1166]" captionTargetBox="[152,1434,194,1119]" captionTargetId="figure@27.[152,1435,194,1121]" captionTargetPageId="27" captionText="FIGURES 77 80. Lyrognathus lessunda sp. nov. paratype female QM S 74066: 77. Left chelicerae, prolateral view. 78. Left intercheliceral pegs, prolateral view. 79. Left cheliceral strikers, ectal view. 80. Left basomedial cheliceral spines, retrolateral view. Scale bar = 2 mm for 77; 1 mm for 78, 79; 0.5 mm for 80. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275745/files/figure.png" pageId="27" pageNumber="28">Fig. 78</figureCitation>
). Retrolaterally basomedially, with small cluster of short strong spines (&gt;3), point distad (
<figureCitation id="135E2A227646FFB39057F9B6FD38D230" box="[595,682,1612,1638]" captionStart="FIGURES 77 80" captionStartId="27.[151,269,1142,1166]" captionTargetBox="[152,1434,194,1119]" captionTargetId="figure@27.[152,1435,194,1121]" captionTargetPageId="27" captionText="FIGURES 77 80. Lyrognathus lessunda sp. nov. paratype female QM S 74066: 77. Left chelicerae, prolateral view. 78. Left intercheliceral pegs, prolateral view. 79. Left cheliceral strikers, ectal view. 80. Left basomedial cheliceral spines, retrolateral view. Scale bar = 2 mm for 77; 1 mm for 78, 79; 0.5 mm for 80. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275745/files/figure.png" pageId="27" pageNumber="28">Fig. 80</figureCitation>
). Antero-dorsally to retrolateral spines: patch of fine setae, dark, of medium length, blends into typical thicker pallid cheliceral setae. Setae filiform, seem to emerge from hemispherical bases.
</paragraph>
<paragraph id="8BDA36A77646FFB492C0F946FE0CD4AA" blockId="27.[151,1437,1276,2011]" lastBlockId="28.[151,1437,152,2007]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
Maxillae (
<figureCitation id="135E2A227646FFB39338F946FE0BD280" box="[316,409,1724,1750]" captionStart="FIGURES 64 70" captionStartId="25.[151,269,1653,1677]" captionTargetBox="[163,1423,343,1632]" captionTargetId="figure@25.[163,1424,343,1632]" captionTargetPageId="25" captionText="FIGURES 64 70. Lyrognathus lessunda sp. nov. holotype female: 64. Carapace, dorsal view. 65. Eyes, dorsal view. 66. Sternum, labium, maxillae and coxa, ventral view. 67. Labium, ventral view. 68. Left palp, legs I, II, ventral view. 69. Abdomen, legs IV, ventral view. 70. Spermathecae, dorsal view. Scale bar = 10 mm for 64, 66, 69; 5 mm for 68; 2 mm for 65, 67, 70. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275743/files/figure.png" pageId="27" pageNumber="28">Figs 66</figureCitation>
,
<figureCitation id="135E2A227646FFB393A1F946FE55D280" box="[421,455,1724,1750]" captionStart="FIGURES 71 76" captionStartId="26.[151,270,1906,1930]" captionTargetBox="[165,1409,415,1873]" captionTargetId="figure@26.[165,1409,415,1874]" captionTargetPageId="26" captionText="FIGURES 71 76. Lyrognathus lessunda sp. nov., holotype female (except: paratype female QM S 74066; 75, 76): 71. Abdomen and legs IV, dorsal view. 72. Left leg IV, retrolateral view. 73. Left leg IV, prolateral view. 74. Left met. and tar. IV, ventral view. 75. Left maxillae, prolateral view. 76. Left maxillary lyra, prolateral view. Scale bar = 10 mm for 71 73; 5 mm for 74; 2 mm for 75; 1 mm for 76. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275744/files/figure.png" pageId="27" pageNumber="28">75</figureCitation>
): prolaterally planoconvex, anterior lobe well pronounced, many cuspules (
<emphasis id="B911EAB57646FFB39756F946FAF4D283" box="[1362,1382,1724,1749]" italics="true" pageId="27" pageNumber="28">&gt;</emphasis>
120) on inner basoventral surface. Ventrally, diagonal to anterior lobe, basal projection evident, overly large, setose.
<taxonomicName id="4C654D247646FFB392F6F8FCFEB8D376" box="[242,298,1798,1824]" class="Arachnida" family="Theraphosidae" genus="Selenocosmia" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="42" phylum="Arthropoda" rank="species" species="lyra">Lyra</taxonomicName>
(
<figureCitation id="135E2A227646FFB3933FF8FCFE5FD376" box="[315,461,1798,1824]" captionStart="FIGURES 71 76" captionStartId="26.[151,270,1906,1930]" captionTargetBox="[165,1409,415,1873]" captionTargetId="figure@26.[165,1409,415,1874]" captionTargetPageId="26" captionText="FIGURES 71 76. Lyrognathus lessunda sp. nov., holotype female (except: paratype female QM S 74066; 75, 76): 71. Abdomen and legs IV, dorsal view. 72. Left leg IV, retrolateral view. 73. Left leg IV, prolateral view. 74. Left met. and tar. IV, ventral view. 75. Left maxillae, prolateral view. 76. Left maxillary lyra, prolateral view. Scale bar = 10 mm for 71 73; 5 mm for 74; 2 mm for 75; 1 mm for 76. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275744/files/figure.png" pageId="27" pageNumber="28">Figs 75, 76</figureCitation>
): many bacilliform rods (&gt;180) form dense, ovoid patch medially on prolateral surface, lowest row up to&gt;20 bacillae longest rods in center of lowest row. Rods distally paddle-shaped with short to medium length shafts, largest with distal blades. At highest point, lyrate patch is&gt;11 rows deep with smallest rods dorsally. Posterior end of patch truncate, anterior end slightly pointed distad/rounded. Immediately above the suture 10 to 15 small spines on anterior margin in disordered rows. Labium (
<figureCitation id="135E2A227646FFB39732F866FA1DD3E0" box="[1334,1423,1948,1974]" captionStart="FIGURES 64 70" captionStartId="25.[151,269,1653,1677]" captionTargetBox="[163,1423,343,1632]" captionTargetId="figure@25.[163,1424,343,1632]" captionTargetPageId="25" captionText="FIGURES 64 70. Lyrognathus lessunda sp. nov. holotype female: 64. Carapace, dorsal view. 65. Eyes, dorsal view. 66. Sternum, labium, maxillae and coxa, ventral view. 67. Labium, ventral view. 68. Left palp, legs I, II, ventral view. 69. Abdomen, legs IV, ventral view. 70. Spermathecae, dorsal view. Scale bar = 10 mm for 64, 66, 69; 5 mm for 68; 2 mm for 65, 67, 70. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275743/files/figure.png" pageId="27" pageNumber="28">Fig. 67</figureCitation>
): length 1.75, width 2.4. Many small cuspules (&gt;350) along anterior 1/3 surface. Labium with 2 discernible hair
<typeStatus id="54DE88057641FFB49293FF62FF47D4E4" box="[151,213,152,178]" pageId="28" pageNumber="29">types</typeStatus>
(L1, L2);
<typeStatus id="54DE88057641FFB49355FF62FE11D4E4" box="[337,387,152,178]" pageId="28" pageNumber="29">type</typeStatus>
L1: long, reasonably dark spiniform, curving distally toward anterior, most concentrated along lateral borders. Setae longest along anterior edge (&gt;30);
<typeStatus id="54DE88057641FFB49170FF47FC35D481" box="[884,935,189,215]" pageId="28" pageNumber="29">type</typeStatus>
L2, 1/2 length of L1, interspersed with L1 but more needle-form.
</paragraph>
<paragraph id="8BDA36A77641FFB492C0FEF2FE4FD61A" blockId="28.[151,1437,152,2007]" pageId="28" pageNumber="29">
Sternum (
<figureCitation id="135E2A227641FFB49332FEF2FE1DD574" box="[310,399,264,290]" captionStart="FIGURES 64 70" captionStartId="25.[151,269,1653,1677]" captionTargetBox="[163,1423,343,1632]" captionTargetId="figure@25.[163,1424,343,1632]" captionTargetPageId="25" captionText="FIGURES 64 70. Lyrognathus lessunda sp. nov. holotype female: 64. Carapace, dorsal view. 65. Eyes, dorsal view. 66. Sternum, labium, maxillae and coxa, ventral view. 67. Labium, ventral view. 68. Left palp, legs I, II, ventral view. 69. Abdomen, legs IV, ventral view. 70. Spermathecae, dorsal view. Scale bar = 10 mm for 64, 66, 69; 5 mm for 68; 2 mm for 65, 67, 70. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275743/files/figure.png" pageId="28" pageNumber="29">Fig. 66</figureCitation>
): length 7.25, width 6.2. Slightly wider posteriorly than anteriorly with 5 discernible hair
<typeStatus id="54DE88057641FFB49293FED7FF47D511" box="[151,213,301,327]" pageId="28" pageNumber="29">types</typeStatus>
(S1 to S5):
<typeStatus id="54DE88057641FFB49362FED7FE0BD511" box="[358,409,301,327]" pageId="28" pageNumber="29">type</typeStatus>
S1, longest, thick blunt-tipped, darker basally, become pallid distally, entire cover, but sparse:
<typeStatus id="54DE88057641FFB492F5FEA8FEB6D53A" box="[241,292,338,364]" pageId="28" pageNumber="29">type</typeStatus>
S2, elongate spiniforms, dark basally, distally pallid, on borders, most concentrated posteriorly:
<typeStatus id="54DE88057641FFB49293FE82FF58D5C4" box="[151,202,376,402]" pageId="28" pageNumber="29">type</typeStatus>
S3, short spiniforms, overall sparse:
<typeStatus id="54DE88057641FFB49080FE82FD25D5C4" box="[644,695,376,402]" pageId="28" pageNumber="29">type</typeStatus>
S4 short, thin pallids, form basis of mat:
<typeStatus id="54DE88057641FFB496A1FE82FB4AD5C4" box="[1189,1240,376,402]" pageId="28" pageNumber="29">type</typeStatus>
S5 dark basally, but apically pallid, short blunt tipped, entirecover in sparse unorganised arrangement. Posteriorly between left/right cox. IV, border greatly acuminate, lateral sternal points also acuminate. Sigilla: 3 pairs, posterior, very large, ovoid, 1.7 lengths apart, 0.6 to 0.95 of their length from margin. Median pair 1/2 size of posterior, similar, 0.20.1 of their length from margin. Anterior pair somewhat obscured, on margin. Labio-sternal sigilla small, thin, but wide.
</paragraph>
<paragraph id="8BDA36A77641FFB492C0FDA2FD89D7CA" blockId="28.[151,1437,152,2007]" pageId="28" pageNumber="29">
Legs. Setation: retrolateral surfaces of pat., tib., met. and tar. IV with distinct thick brushes of hair-like setae (
<figureCitation id="135E2A227641FFB492E1FD87FED2D6C1" box="[229,320,637,663]" captionStart="FIGURES 71 76" captionStartId="26.[151,270,1906,1930]" captionTargetBox="[165,1409,415,1873]" captionTargetId="figure@26.[165,1409,415,1874]" captionTargetPageId="26" captionText="FIGURES 71 76. Lyrognathus lessunda sp. nov., holotype female (except: paratype female QM S 74066; 75, 76): 71. Abdomen and legs IV, dorsal view. 72. Left leg IV, retrolateral view. 73. Left leg IV, prolateral view. 74. Left met. and tar. IV, ventral view. 75. Left maxillae, prolateral view. 76. Left maxillary lyra, prolateral view. Scale bar = 10 mm for 71 73; 5 mm for 74; 2 mm for 75; 1 mm for 76. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275744/files/figure.png" pageId="28" pageNumber="29">Fig. 71</figureCitation>
), penicillate. Legs: formula; (length) IV, I, II, III, (width) IV, III, I, II. Leg
<collectingCountry id="F37276377641FFB496DFFD87FA93D6C1" box="[1243,1281,637,663]" name="Russia" pageId="28" pageNumber="29">RF</collectingCountry>
~ 87.92. Leg lengths (fem., pat., tib., met., tar., total):
<typeStatus id="54DE88057641FFB49073FD58FD4FD6EA" box="[631,733,674,700]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
S74067: palp: 7.2, 4.3, 4.65, 0.00, 5.15, 21.3. I: 10.4, 6.15, 7.25, 6.3, 4.1, 34.2. II: 8.1, 5.2, 5.2, 5.25, 4.15, 27.9. III: 8.9, 4.8, 4.8, 5.6, 4.05, 28.15. IV: 10.65, 5.85, 8.25, 9.3, 4.85, 38.9.
<typeStatus id="54DE88057641FFB49350FD17FE2ED751" box="[340,444,749,775]" pageId="28" pageNumber="29" type="paratype">Paratype</typeStatus>
<accessionNumber id="9436AB447641FFB493C0FD17FDB0D751" box="[452,546,749,775]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S74070" pageId="28" pageNumber="29" type="EnaNcbi">S74070</accessionNumber>
: palp: 7.3, 5.15, 5.2, 0.00, 5.25, 22.9. I: 10.25, 6.7, 8.0, 7.3, 4.65, 36.9. II: 8.55, 5.9, 6.2, 6.3, 4.4, 31.35. III: 7.75, 5.25, 6.7, 6.45, 4.15, 30.3. IV: 10.3, 5.95, 9.05, 9.80, 5.35, 40.35. Leg mid-widths (fem., pat., tib., met., tar., total.): (
<typeStatus id="54DE88057641FFB490B2FCC2FC8CD704" box="[694,798,824,850]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
only): palp: 1.85, 2.13, 1.71, 0.00, 2.05, 7.74. I: 2.45, 2.51, 2.63, 2.1, 2.15, 11.84. II: 2.35, 2.35, 2.12, 1.95, 1.75, 10.52. III: 3.15, 2.55, 2.35, 1.95, 1.85, 11.85. IV: 2.45, 2.83, 3.02, 2.35, 2.15, 12.8.
</paragraph>
<paragraph id="8BDA36A77641FFB492C0FC52FCF4D794" blockId="28.[151,1437,152,2007]" box="[196,870,936,962]" pageId="28" pageNumber="29">Scopula: undivided, for full length of met./tar. on all legs.</paragraph>
<paragraph id="8BDA36A77641FFB492C0FC37FD60D0BA" blockId="28.[151,1437,152,2007]" pageId="28" pageNumber="29">
Coxae: some small black thorns prolatero-dorsally, no thorns retrolaterally on IIV; easily seen dorsally; I longest, about 1.6 times length of II. IV widest, as long as III, basally rectangular with rounded corners; with ventral thorns prolaterally on IIV. IIII ventrally with many long thick blunt setae proximally, pallid. No short black setae. IV with mixture of long thick blunt setae proximally, pallid intermixed with shorter thin pallid setae. Ventral surface of IIV gently sloping anteriorly. Retrolateral setation: IIII with median narrow light brush, IV completely cover of light setae. IIV retrolaterally lack ventral ledge. Ventral measurements for coxae (
<typeStatus id="54DE88057641FFB49310FB57FEEED091" box="[276,380,1197,1223]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
only): Ilength 6.8, width 3.1; II5.8, 2.9; III5.1, 2.8; IV6.1, 3.4. Trochantera: Ilength 2.6, width 2.7; II2.6, 2.1; III2.6, 2.8; IV3.3, 3.1.
</paragraph>
<paragraph id="8BDA36A77641FFB492C0FB02FAF6D1D4" blockId="28.[151,1437,152,2007]" pageId="28" pageNumber="29">
Trichobothria: on all tar., basal filiform field slightly wider than clavate field, merging evenly. Clavates on tar. I only in distal 1/2, long filiforms only in basal 1/2, shorter filiforms intermixed with clavates distally. Clavate extent on IIIV
<emphasis id="B911EAB57641FFB493C7FAB9FE73D10A" box="[451,481,1347,1372]" italics="true" pageId="28" pageNumber="29">cf.</emphasis>
I, only in distal 1/2. Shorter filiforms for length. Short epitrichobothrial field (
<bibRefCitation id="EFF44B567641FFB4929BFA92FEBDD1D4" author="Raven" box="[159,303,1384,1410]" pageId="28" pageNumber="43" refString="Raven, R. J. (2005) A new tarantula species from northern Australia (Araneae, Theraphosidae). Zootaxa, 1004, 15 - 28." type="journal article" year="2005">Raven 2005</bibRefCitation>
) on I shorter than clavates, uniform height for length. Tarsal organ not evident on legs IIV.
</paragraph>
<paragraph id="8BDA36A77641FFB492C0FA77FD48D19A" blockId="28.[151,1437,152,2007]" pageId="28" pageNumber="29">Spines: met. I with 1 DV, met. II with 1 DV, 1 DPV, 1 DRV, met. III with 2 DV, 1 DPV, 1 DRV, 1 DD and met. IV with 2 DV, 1 DPV, 1 DRV, 1 DD, 1 DPL.</paragraph>
<paragraph id="8BDA36A77641FFB492C0FA22FBB9D1A4" blockId="28.[151,1437,152,2007]" box="[196,1067,1496,1522]" pageId="28" pageNumber="29">Claws: unarmed paired claws on all legs. No reduced third claw on leg IV.</paragraph>
<paragraph id="8BDA36A77641FFB492C0FA07FB4ED34A" blockId="28.[151,1437,152,2007]" pageId="28" pageNumber="29">
Abdomen (
<figureCitation id="135E2A227641FFB4934BFA07FE23D241" box="[335,433,1533,1559]" captionStart="FIGURES 64 70" captionStartId="25.[151,269,1653,1677]" captionTargetBox="[163,1423,343,1632]" captionTargetId="figure@25.[163,1424,343,1632]" captionTargetPageId="25" captionText="FIGURES 64 70. Lyrognathus lessunda sp. nov. holotype female: 64. Carapace, dorsal view. 65. Eyes, dorsal view. 66. Sternum, labium, maxillae and coxa, ventral view. 67. Labium, ventral view. 68. Left palp, legs I, II, ventral view. 69. Abdomen, legs IV, ventral view. 70. Spermathecae, dorsal view. Scale bar = 10 mm for 64, 66, 69; 5 mm for 68; 2 mm for 65, 67, 70. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275743/files/figure.png" pageId="28" pageNumber="29">Figs 69</figureCitation>
,
<figureCitation id="135E2A227641FFB493BBFA07FE71D241" box="[447,483,1533,1559]" captionStart="FIGURES 71 76" captionStartId="26.[151,270,1906,1930]" captionTargetBox="[165,1409,415,1873]" captionTargetId="figure@26.[165,1409,415,1874]" captionTargetPageId="26" captionText="FIGURES 71 76. Lyrognathus lessunda sp. nov., holotype female (except: paratype female QM S 74066; 75, 76): 71. Abdomen and legs IV, dorsal view. 72. Left leg IV, retrolateral view. 73. Left leg IV, prolateral view. 74. Left met. and tar. IV, ventral view. 75. Left maxillae, prolateral view. 76. Left maxillary lyra, prolateral view. Scale bar = 10 mm for 71 73; 5 mm for 74; 2 mm for 75; 1 mm for 76. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275744/files/figure.png" pageId="28" pageNumber="29">71</figureCitation>
): ovular, elongate, reddish brown; with 4 discernible hair
<typeStatus id="54DE88057641FFB496BBFA07FA93D241" box="[1215,1281,1533,1559]" pageId="28" pageNumber="29">types</typeStatus>
(A1 to A4): dorsally with 3 hair
<typeStatus id="54DE88057641FFB49385F9D8FE51D26A" box="[385,451,1570,1596]" pageId="28" pageNumber="29">types</typeStatus>
:
<typeStatus id="54DE88057641FFB493CAF9D8FD93D26A" box="[462,513,1570,1596]" pageId="28" pageNumber="29">type</typeStatus>
A1, long, dark blunt-tipped, uniform color:
<typeStatus id="54DE88057641FFB49600F9D8FBA4D26A" box="[1028,1078,1570,1596]" pageId="28" pageNumber="29">type</typeStatus>
A2, mid-length, also dark, but distally spiniform, unlike
<typeStatus id="54DE88057641FFB493CCF9B2FE68D234" box="[456,506,1608,1634]" pageId="28" pageNumber="29">type</typeStatus>
A1:
<typeStatus id="54DE88057641FFB49033F9B2FDF8D234" box="[567,618,1608,1634]" pageId="28" pageNumber="29">type</typeStatus>
A3, form dense mat, mid-length, uniformly dark, needle-form (some stand erect, some run posteriorly but with similar morphology). Longest hairs (
<typeStatus id="54DE88057641FFB4964FF997FB12D2D1" box="[1099,1152,1645,1671]" pageId="28" pageNumber="29">type</typeStatus>
A1) more concentrated posteriorly toward spinnerets, point distad, but proximally, hairs stand erect, medially curved (toward posterior). Ventral hair
<typeStatus id="54DE88057641FFB493ACF942FE74D284" box="[424,486,1720,1746]" pageId="28" pageNumber="29">types</typeStatus>
almost same as dorsal, but
<typeStatus id="54DE88057641FFB4912CF942FCC8D284" box="[808,858,1720,1746]" pageId="28" pageNumber="29">type</typeStatus>
A2 and A3 more dense entirely.
<typeStatus id="54DE88057641FFB496D9F942FA85D284" box="[1245,1303,1720,1746]" pageId="28" pageNumber="29">Type</typeStatus>
A4 hair, in very dense patch, trapezoidal, posterior of pedicel between median regions of cox. IV. Hairs of medium length, straight, pallid. Between anterior pair of booklungs, setal mat similar to rest of venter.
</paragraph>
<paragraph id="8BDA36A77641FFB492C0F8D2FF4ED3DA" blockId="28.[151,1437,152,2007]" pageId="28" pageNumber="29">
Genitalia (
<figureCitation id="135E2A227641FFB4933BF8D2FE0AD314" box="[319,408,1832,1858]" captionStart="FIGURES 64 70" captionStartId="25.[151,269,1653,1677]" captionTargetBox="[163,1423,343,1632]" captionTargetId="figure@25.[163,1424,343,1632]" captionTargetPageId="25" captionText="FIGURES 64 70. Lyrognathus lessunda sp. nov. holotype female: 64. Carapace, dorsal view. 65. Eyes, dorsal view. 66. Sternum, labium, maxillae and coxa, ventral view. 67. Labium, ventral view. 68. Left palp, legs I, II, ventral view. 69. Abdomen, legs IV, ventral view. 70. Spermathecae, dorsal view. Scale bar = 10 mm for 64, 66, 69; 5 mm for 68; 2 mm for 65, 67, 70. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275743/files/figure.png" pageId="28" pageNumber="29">Fig. 70</figureCitation>
): epigastric fold 4.95. Spermathecae bilobular. Lobes apically swollen with lateral lobes 2/3 length of medials. Medial lobes very long. Epigastric fold extends
<emphasis id="B911EAB57641FFB491D9F8B7FB92D330" box="[989,1024,1869,1894]" italics="true" pageId="28" pageNumber="29">ca.</emphasis>
2/3 length of medial spermathecal lobes.
</paragraph>
<paragraph id="8BDA36A77641FFB592C0F862FD34D481" blockId="28.[151,1437,152,2007]" lastBlockId="29.[151,1437,152,364]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
Spinnerets: PMS: length 1.45, width (medially) 0.65. PLS: section lengths; basal 2.85, medial 1.30, apical 2.10. PLS section widths (medially); basal 1.32, medial 0.98, apical 0.67; with 2 discernible hair
<typeStatus id="54DE88057641FFB49711F847FAC1D381" box="[1301,1363,1981,2007]" pageId="28" pageNumber="29">types</typeStatus>
(SP1, SP2): SP1 very short and distally blunt pallid, covers both pairs ventrally; second form (SP2), very similar but up to twice as long, dorsally on all segments.
</paragraph>
<paragraph id="8BDA36A77640FFB592C0FF18FE87D53A" blockId="29.[151,1437,152,364]" pageId="29" pageNumber="30">
Male
<typeStatus id="54DE88057640FFB59303FF18FEFED4AA" box="[263,364,226,252]" pageId="29" pageNumber="30" type="paratype">paratype</typeStatus>
(
<figureCitation id="135E2A227640FFB59378FF18FE41D4AA" box="[380,467,226,252]" captionStart="FIGURES 81 90" captionStartId="29.[151,271,1676,1700]" captionTargetBox="[151,1434,412,1651]" captionTargetId="figure@29.[151,1436,412,1652]" captionTargetPageId="29" captionText="FIGURES 81 90. Lyrognathus lessunda sp. nov. paratype male QM S 74069: 81. Habitus, dorsal view. 82. Carapace, dorsal view. 83. Eyes, dorsal view. 84. Sternum, labium, maxillae and coxa, ventral view. 85. Right intercheliceral pegs, prolateral view. 86. Right cheliceral strikers, ectal view. 87. Right maxillae showing lyra, prolateral view. 88. Left bulb, retrolateral view. 89. Left bulb, prolateral view. 90. Left bulb, anterior view. Scale bar = 10 mm for 81; 5 mm for 82, 84; 2 mm for 83, 86, 87; 1 mm for 85, 88 90. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275746/files/figure.png" pageId="29" pageNumber="30">Fig. 81</figureCitation>
, as in female except) with body length: 30.28. Color (in life): reddish brown with slight purple sheen in natural light. Carapace (
<figureCitation id="135E2A227640FFB59343FED7FE37D511" box="[327,421,301,327]" captionStart="FIGURES 81 90" captionStartId="29.[151,271,1676,1700]" captionTargetBox="[151,1434,412,1651]" captionTargetId="figure@29.[151,1436,412,1652]" captionTargetPageId="29" captionText="FIGURES 81 90. Lyrognathus lessunda sp. nov. paratype male QM S 74069: 81. Habitus, dorsal view. 82. Carapace, dorsal view. 83. Eyes, dorsal view. 84. Sternum, labium, maxillae and coxa, ventral view. 85. Right intercheliceral pegs, prolateral view. 86. Right cheliceral strikers, ectal view. 87. Right maxillae showing lyra, prolateral view. 88. Left bulb, retrolateral view. 89. Left bulb, prolateral view. 90. Left bulb, anterior view. Scale bar = 10 mm for 81; 5 mm for 82, 84; 2 mm for 83, 86, 87; 1 mm for 85, 88 90. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275746/files/figure.png" pageId="29" pageNumber="30">Fig. 84</figureCitation>
): length 11.9, width 9.35 (width across anterior edge 6.9). Fovea 1.75 wide, deep, procurved.
</paragraph>
<caption id="DF1A662F7640FFB59293F976FCB0D375" httpUri="https://zenodo.org/record/275746/files/figure.png" pageId="29" pageNumber="30" targetBox="[151,1434,412,1651]" targetPageId="29">
<paragraph id="8BDA36A77640FFB59293F976FCB0D375" blockId="29.[151,1436,1676,1827]" pageId="29" pageNumber="30">
<emphasis id="B911EAB57640FFB59293F976FEF2D2F2" bold="true" box="[151,352,1676,1700]" pageId="29" pageNumber="30">FIGURES 8190.</emphasis>
<taxonomicName id="4C654D247640FFB5936CF976FDC7D2F5" box="[360,597,1676,1699]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="lessunda" status="sp. nov.">
<emphasis id="B911EAB57640FFB5936CF976FDC7D2F5" box="[360,597,1676,1699]" italics="true" pageId="29" pageNumber="30">Lyrognathus lessunda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257CE7640FFB59058F976FD26D2F2" box="[604,692,1676,1700]" pageId="29" pageNumber="30" rank="species">
<emphasis id="B911EAB57640FFB59058F976FD26D2F2" bold="true" box="[604,692,1676,1700]" pageId="29" pageNumber="30">sp. nov.</emphasis>
</taxonomicNameLabel>
paratype male QM
<accessionNumber id="9436AB447640FFB5918BF976FC74D2F5" box="[911,998,1676,1699]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S74069" pageId="29" pageNumber="30" type="EnaNcbi">S74069</accessionNumber>
: 81. Habitus, dorsal view. 82. Carapace, dorsal view. 83. Eyes, dorsal view. 84. Sternum, labium, maxillae and coxa, ventral view. 85. Right intercheliceral pegs, prolateral view. 86. Right cheliceral strikers, ectal view. 87. Right maxillae showing
<taxonomicName id="4C654D247640FFB59620F936FBC2D2B5" box="[1060,1104,1740,1763]" class="Arachnida" family="Theraphosidae" genus="Selenocosmia" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="lyra">lyra</taxonomicName>
, prolateral view. 88. Left bulb, retrolateral view. 89. Left bulb, prolateral view. 90. Left bulb, anterior view. Scale bar = 10mm for 81; 5mm for 82, 84; 2mm for 83, 86, 87; 1mm for 85, 8890. Photos: S. C. Nunn.
</paragraph>
</caption>
<paragraph id="8BDA36A77640FFB592C0F8A8FCC6D3C7" blockId="29.[151,1437,1874,2012]" pageId="29" pageNumber="30">
Eyes (
<figureCitation id="135E2A227640FFB5930BF8A8FEF8D33A" box="[271,362,1874,1900]" captionStart="FIGURES 81 90" captionStartId="29.[151,271,1676,1700]" captionTargetBox="[151,1434,412,1651]" captionTargetId="figure@29.[151,1436,412,1652]" captionTargetPageId="29" captionText="FIGURES 81 90. Lyrognathus lessunda sp. nov. paratype male QM S 74069: 81. Habitus, dorsal view. 82. Carapace, dorsal view. 83. Eyes, dorsal view. 84. Sternum, labium, maxillae and coxa, ventral view. 85. Right intercheliceral pegs, prolateral view. 86. Right cheliceral strikers, ectal view. 87. Right maxillae showing lyra, prolateral view. 88. Left bulb, retrolateral view. 89. Left bulb, prolateral view. 90. Left bulb, anterior view. Scale bar = 10 mm for 81; 5 mm for 82, 84; 2 mm for 83, 86, 87; 1 mm for 85, 88 90. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275746/files/figure.png" pageId="29" pageNumber="30">Fig. 83</figureCitation>
): ocular tubercle; length 1.86, width 2.17. Anterior row transverse. Posterior row slightly recurved. Eyes, largest to smallest: AME, ALE, PLE, PME.
</paragraph>
<paragraph id="8BDA36A77640FFB592C0F866FB54D38A" blockId="29.[151,1437,1874,2012]" pageId="29" pageNumber="30">
Chelicerae (
<figureCitation id="135E2A227640FFB5935FF866FE29D3E0" box="[347,443,1948,1974]" captionStart="FIGURES 81 90" captionStartId="29.[151,271,1676,1700]" captionTargetBox="[151,1434,412,1651]" captionTargetId="figure@29.[151,1436,412,1652]" captionTargetPageId="29" captionText="FIGURES 81 90. Lyrognathus lessunda sp. nov. paratype male QM S 74069: 81. Habitus, dorsal view. 82. Carapace, dorsal view. 83. Eyes, dorsal view. 84. Sternum, labium, maxillae and coxa, ventral view. 85. Right intercheliceral pegs, prolateral view. 86. Right cheliceral strikers, ectal view. 87. Right maxillae showing lyra, prolateral view. 88. Left bulb, retrolateral view. 89. Left bulb, prolateral view. 90. Left bulb, anterior view. Scale bar = 10 mm for 81; 5 mm for 82, 84; 2 mm for 83, 86, 87; 1 mm for 85, 88 90. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275746/files/figure.png" pageId="29" pageNumber="30">Fig. 86</figureCitation>
): length 3.9, width at base of each lobe 2.5. Teeth,12,&gt;15 smaller basomesals. Intercheliceral pegs (&gt;9 larger,&gt;15 smaller) in tight cluster on basodorsal surface (
<figureCitation id="135E2A227640FFB59664F838FB2AD38A" box="[1120,1208,1986,2012]" captionStart="FIGURES 81 90" captionStartId="29.[151,271,1676,1700]" captionTargetBox="[151,1434,412,1651]" captionTargetId="figure@29.[151,1436,412,1652]" captionTargetPageId="29" captionText="FIGURES 81 90. Lyrognathus lessunda sp. nov. paratype male QM S 74069: 81. Habitus, dorsal view. 82. Carapace, dorsal view. 83. Eyes, dorsal view. 84. Sternum, labium, maxillae and coxa, ventral view. 85. Right intercheliceral pegs, prolateral view. 86. Right cheliceral strikers, ectal view. 87. Right maxillae showing lyra, prolateral view. 88. Left bulb, retrolateral view. 89. Left bulb, prolateral view. 90. Left bulb, anterior view. Scale bar = 10 mm for 81; 5 mm for 82, 84; 2 mm for 83, 86, 87; 1 mm for 85, 88 90. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275746/files/figure.png" pageId="29" pageNumber="30">Fig. 85</figureCitation>
).
</paragraph>
<paragraph id="8BDA36A77643FFB692C0FF62FD1CD574" blockId="30.[151,1437,152,1372]" pageId="30" pageNumber="31">
Maxillae (
<figureCitation id="135E2A227643FFB69340FF62FE36D4E4" box="[324,420,152,178]" captionStart="FIGURES 81 90" captionStartId="29.[151,271,1676,1700]" captionTargetBox="[151,1434,412,1651]" captionTargetId="figure@29.[151,1436,412,1652]" captionTargetPageId="29" captionText="FIGURES 81 90. Lyrognathus lessunda sp. nov. paratype male QM S 74069: 81. Habitus, dorsal view. 82. Carapace, dorsal view. 83. Eyes, dorsal view. 84. Sternum, labium, maxillae and coxa, ventral view. 85. Right intercheliceral pegs, prolateral view. 86. Right cheliceral strikers, ectal view. 87. Right maxillae showing lyra, prolateral view. 88. Left bulb, retrolateral view. 89. Left bulb, prolateral view. 90. Left bulb, anterior view. Scale bar = 10 mm for 81; 5 mm for 82, 84; 2 mm for 83, 86, 87; 1 mm for 85, 88 90. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275746/files/figure.png" pageId="30" pageNumber="31">Fig. 87</figureCitation>
): anterior lobe well pronounced with many cuspules (&gt;140) on inner basoventral surface.
<taxonomicName id="4C654D247643FFB69305FF47FEA8D481" box="[257,314,189,215]" class="Arachnida" family="Theraphosidae" genus="Selenocosmia" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="42" phylum="Arthropoda" rank="species" species="lyra">Lyra</taxonomicName>
(
<figureCitation id="135E2A227643FFB69348FF47FE3ED481" box="[332,428,189,215]" captionStart="FIGURES 81 90" captionStartId="29.[151,271,1676,1700]" captionTargetBox="[151,1434,412,1651]" captionTargetId="figure@29.[151,1436,412,1652]" captionTargetPageId="29" captionText="FIGURES 81 90. Lyrognathus lessunda sp. nov. paratype male QM S 74069: 81. Habitus, dorsal view. 82. Carapace, dorsal view. 83. Eyes, dorsal view. 84. Sternum, labium, maxillae and coxa, ventral view. 85. Right intercheliceral pegs, prolateral view. 86. Right cheliceral strikers, ectal view. 87. Right maxillae showing lyra, prolateral view. 88. Left bulb, retrolateral view. 89. Left bulb, prolateral view. 90. Left bulb, anterior view. Scale bar = 10 mm for 81; 5 mm for 82, 84; 2 mm for 83, 86, 87; 1 mm for 85, 88 90. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275746/files/figure.png" pageId="30" pageNumber="31">Fig. 87</figureCitation>
): immediately above suture 1015 elongated spines medially to anterior margin. Bacillate rods relatively slightly shorter than in female. Labium (
<figureCitation id="135E2A227643FFB6918DFF18FC70D4AA" box="[905,994,226,252]" captionStart="FIGURES 81 90" captionStartId="29.[151,271,1676,1700]" captionTargetBox="[151,1434,412,1651]" captionTargetId="figure@29.[151,1436,412,1652]" captionTargetPageId="29" captionText="FIGURES 81 90. Lyrognathus lessunda sp. nov. paratype male QM S 74069: 81. Habitus, dorsal view. 82. Carapace, dorsal view. 83. Eyes, dorsal view. 84. Sternum, labium, maxillae and coxa, ventral view. 85. Right intercheliceral pegs, prolateral view. 86. Right cheliceral strikers, ectal view. 87. Right maxillae showing lyra, prolateral view. 88. Left bulb, retrolateral view. 89. Left bulb, prolateral view. 90. Left bulb, anterior view. Scale bar = 10 mm for 81; 5 mm for 82, 84; 2 mm for 83, 86, 87; 1 mm for 85, 88 90. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275746/files/figure.png" pageId="30" pageNumber="31">Fig. 84</figureCitation>
): length 1.95, width 2.65. Many small cuspules (&gt;300) along anterior 1/4 surface.
</paragraph>
<paragraph id="8BDA36A77643FFB692C0FED7FEE8D5C4" blockId="30.[151,1437,152,1372]" pageId="30" pageNumber="31">
Sternum (
<figureCitation id="135E2A227643FFB6933CFED7FE03D511" box="[312,401,301,327]" captionStart="FIGURES 81 90" captionStartId="29.[151,271,1676,1700]" captionTargetBox="[151,1434,412,1651]" captionTargetId="figure@29.[151,1436,412,1652]" captionTargetPageId="29" captionText="FIGURES 81 90. Lyrognathus lessunda sp. nov. paratype male QM S 74069: 81. Habitus, dorsal view. 82. Carapace, dorsal view. 83. Eyes, dorsal view. 84. Sternum, labium, maxillae and coxa, ventral view. 85. Right intercheliceral pegs, prolateral view. 86. Right cheliceral strikers, ectal view. 87. Right maxillae showing lyra, prolateral view. 88. Left bulb, retrolateral view. 89. Left bulb, prolateral view. 90. Left bulb, anterior view. Scale bar = 10 mm for 81; 5 mm for 82, 84; 2 mm for 83, 86, 87; 1 mm for 85, 88 90. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275746/files/figure.png" pageId="30" pageNumber="31">Fig. 84</figureCitation>
): length 5.85, width 5.05. Wider posteriorly than anteriorly, with thin but even cover of short setae, interspersed with many longer darker setae. No spines around edges. Lateral points acuminate, but not posterior angle.
</paragraph>
<paragraph id="8BDA36A77643FFB692C0FE67FC9DD721" blockId="30.[151,1437,152,1372]" pageId="30" pageNumber="31">
Legs: formula; (length) IV, I, II, III; (width) III, IV, I, II. Leg
<collectingCountry id="F37276377643FFB6918FFE67FC22D5E1" box="[907,944,413,439]" name="Russia" pageId="30" pageNumber="31">RF</collectingCountry>
~97.07 (
<accessionNumber id="9436AB447643FFB69614FE67FBFDD5E1" box="[1040,1135,413,439]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S74069" pageId="30" pageNumber="31" type="EnaNcbi">S74069</accessionNumber>
). Lengths (fem., pat., tib., met., tar., total):
<typeStatus id="54DE88057643FFB6935AFE38FE56D58A" box="[350,452,450,476]" pageId="30" pageNumber="31" type="paratype">paratype</typeStatus>
<accessionNumber id="9436AB447643FFB693C8FE38FDB8D58A" box="[460,554,450,476]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S74069" pageId="30" pageNumber="31" type="EnaNcbi">S74069</accessionNumber>
: palp: 7.25, 4.7, 6.9, 0.00, 2.4, 21.25. I: 11.4, 6.7, 9.4, 8.45, 5.45, 41.4. II: 10.4, 5.5, 8.15, 7.45, 4.6, 36.1. III: 8.55, 5.3, 5.5, 6.75, 4.15, 30.25. IV: 11.85, 5.8, 9.35, 10.85, 4.8, 42.65.
<typeStatus id="54DE88057643FFB69293FDF7FF6FD671" box="[151,253,525,551]" pageId="30" pageNumber="31" type="paratype">paratype</typeStatus>
S74068: palp: 6.9, 4.75, 6.35, 0.00, 2.35, 20.35. I: 11.1, 6.15, 9.45, 8.1, 5.15, 39.95. II: 9.05, 5.35, 7.2, 7.25, 5.05, 33.9. III: 8.0, 4.85, 5.35, 6.85, 3.95, 29.0. IV: 11.25, 5.55, 9.25, 11.45, 4.75, 42.25.
<typeStatus id="54DE88057643FFB69732FDC8FA09D61A" box="[1334,1435,562,588]" pageId="30" pageNumber="31" type="paratype">paratype</typeStatus>
S74066: palp: 8.6, 5.0, 7.1, 0.00, 2.45, 23.15. I: 12.15, 6.75, 10.6, 9.15, 5.9, 44.55. II: 10.0, 5.75, 8.05, 8.15, 5.25, 37.2. III: 8.45, 5.3, 5.65, 7.85, 4.25, 31.5. IV: 12.25, 5.65, 10.25, 12.65, 5.15, 49.15. Leg mid-widths (fem., pat., tib., met., tar., total.):
<typeStatus id="54DE88057643FFB69022FD58FD1ED6EA" box="[550,652,674,700]" pageId="30" pageNumber="31" type="paratype">paratype</typeStatus>
<accessionNumber id="9436AB447643FFB69090FD58FD61D6EA" box="[660,755,674,700]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S74069" pageId="30" pageNumber="31" type="EnaNcbi">S74069</accessionNumber>
: palp: 1.85,1.95, 1.9, 0.00, 2.03, 7.73. I: 2.26, 2.45, 1.98, 1.47, 1.35, 9.51. II: 2.25, 2.17, 1.95, 1.37, 1.27, 9.01. III: 3.15, 2.3, 2.1, 1.37, 1.51, 10.43. IV: 2.35, 2.37, 2.27, 1.52, 1.36, 9.87. Leg IV with reduced retrolateral setal brush, most concentrated on retrolateral metatarsal surface, proximally, but on tib. (
<figureCitation id="135E2A227643FFB69024FCE8FDE9D77A" box="[544,635,786,812]" captionStart="FIGURES 81 90" captionStartId="29.[151,271,1676,1700]" captionTargetBox="[151,1434,412,1651]" captionTargetId="figure@29.[151,1436,412,1652]" captionTargetPageId="29" captionText="FIGURES 81 90. Lyrognathus lessunda sp. nov. paratype male QM S 74069: 81. Habitus, dorsal view. 82. Carapace, dorsal view. 83. Eyes, dorsal view. 84. Sternum, labium, maxillae and coxa, ventral view. 85. Right intercheliceral pegs, prolateral view. 86. Right cheliceral strikers, ectal view. 87. Right maxillae showing lyra, prolateral view. 88. Left bulb, retrolateral view. 89. Left bulb, prolateral view. 90. Left bulb, anterior view. Scale bar = 10 mm for 81; 5 mm for 82, 84; 2 mm for 83, 86, 87; 1 mm for 85, 88 90. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275746/files/figure.png" pageId="30" pageNumber="31">Fig. 81</figureCitation>
). Coxae ventral measurements (
<accessionNumber id="9436AB447643FFB69606FCE8FBF4D77A" box="[1026,1126,786,812]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S74069" pageId="30" pageNumber="31" type="EnaNcbi">S74069</accessionNumber>
): palplength 5.97, width 2.66; I5.52, 2.87; II4.69, 2.67; III4.2, 2.73; IV5.44, 3.2. Trochantera: palplength 2.31, width 1.87; I 3.36, 2.78; II3.04, 2.69; III2.34, 2.7; IV2.96, 2.86.
</paragraph>
<paragraph id="8BDA36A77643FFB692C0FC78FB75D7CA" blockId="30.[151,1437,152,1372]" box="[196,1255,898,924]" pageId="30" pageNumber="31">Claws: as in female, except on apical ventral region of paired claws of IV, 23 small teeth.</paragraph>
<paragraph id="8BDA36A77643FFB692C0FC52FBB8D064" blockId="30.[151,1437,152,1372]" pageId="30" pageNumber="31">
Genitalia. Palp (
<figureCitation id="135E2A227643FFB6938EFC52FD8BD794" box="[394,537,936,962]" captionStart="FIGURES 81 90" captionStartId="29.[151,271,1676,1700]" captionTargetBox="[151,1434,412,1651]" captionTargetId="figure@29.[151,1436,412,1652]" captionTargetPageId="29" captionText="FIGURES 81 90. Lyrognathus lessunda sp. nov. paratype male QM S 74069: 81. Habitus, dorsal view. 82. Carapace, dorsal view. 83. Eyes, dorsal view. 84. Sternum, labium, maxillae and coxa, ventral view. 85. Right intercheliceral pegs, prolateral view. 86. Right cheliceral strikers, ectal view. 87. Right maxillae showing lyra, prolateral view. 88. Left bulb, retrolateral view. 89. Left bulb, prolateral view. 90. Left bulb, anterior view. Scale bar = 10 mm for 81; 5 mm for 82, 84; 2 mm for 83, 86, 87; 1 mm for 85, 88 90. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275746/files/figure.png" pageId="30" pageNumber="31">Figs 8890</figureCitation>
): tib. mildly incrassate, tapers distally. Cymbium bipartite, tegulum large. Lanceolate embolus slightly tapers distally with very little flaring, proximally emerges anterior to tegulum. Relative length of palpal tibia
<emphasis id="B911EAB57643FFB69002FC08FDB8D05D" box="[518,554,1010,1035]" italics="true" pageId="30" pageNumber="31">ca.</emphasis>
twice length of embolus. Longitudinal keel distinct (embolus not terete), runs entire length of embolus, twisted through 90° prolaterally at tip (
<figureCitation id="135E2A227643FFB691C1FBE2FB8CD064" box="[965,1054,1048,1074]" captionStart="FIGURES 81 90" captionStartId="29.[151,271,1676,1700]" captionTargetBox="[151,1434,412,1651]" captionTargetId="figure@29.[151,1436,412,1652]" captionTargetPageId="29" captionText="FIGURES 81 90. Lyrognathus lessunda sp. nov. paratype male QM S 74069: 81. Habitus, dorsal view. 82. Carapace, dorsal view. 83. Eyes, dorsal view. 84. Sternum, labium, maxillae and coxa, ventral view. 85. Right intercheliceral pegs, prolateral view. 86. Right cheliceral strikers, ectal view. 87. Right maxillae showing lyra, prolateral view. 88. Left bulb, retrolateral view. 89. Left bulb, prolateral view. 90. Left bulb, anterior view. Scale bar = 10 mm for 81; 5 mm for 82, 84; 2 mm for 83, 86, 87; 1 mm for 85, 88 90. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275746/files/figure.png" pageId="30" pageNumber="31">Fig. 90</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37F652C7643FFB692C0FBC7FC39D0F4" pageId="30" pageNumber="31" type="distribution">
<paragraph id="8BDA36A77643FFB692C0FBC7FC39D0F4" blockId="30.[151,1437,152,1372]" pageId="30" pageNumber="31">
<emphasis id="B911EAB57643FFB692C0FBC7FDF0D001" bold="true" box="[196,610,1085,1111]" pageId="30" pageNumber="31">Distribution and natural history</emphasis>
(
<figureCitation id="135E2A227643FFB69077FBC7FD4FD001" box="[627,733,1085,1111]" captionStart="FIGURE 114" captionStartId="38.[151,255,1297,1321]" captionTargetBox="[151,1436,194,1276]" captionTargetId="figure@38.[151,1436,194,1276]" captionTargetPageId="38" captionText="FIGURE 114. Records of Lyrognathus species in Southeast Asia. C. Senescall" httpUri="https://zenodo.org/record/275751/files/figure.png" pageId="30" pageNumber="31">Fig. 114</figureCitation>
): The natural history of this spider remains unknown. It is claimed to live in ground burrows on the higher slopes of Gunung Rinjani, Lombok Island, Lesser Sunda Islands,
<collectingCountry id="F37276377643FFB692FCFB72FEF8D0F4" box="[248,362,1160,1186]" name="Indonesia" pageId="30" pageNumber="31">Indonesia</collectingCountry>
(G. Albanjari and I. Wayan Suana, pers. comm.).
</paragraph>
</subSubSection>
<subSubSection id="C37F652C7643FFB692C0FB57FF78D10A" pageId="30" pageNumber="31" type="discussion">
<paragraph id="8BDA36A77643FFB692C0FB57FF78D10A" blockId="30.[151,1437,152,1372]" pageId="30" pageNumber="31">
<emphasis id="B911EAB57643FFB692C0FB57FEADD091" bold="true" box="[196,319,1197,1223]" pageId="30" pageNumber="31">Remarks:</emphasis>
This is the first confirmed record of any theraphosid species on Lombok Island.
<bibRefCitation id="EFF44B567643FFB696EAFB57FF7DD0BA" author="Roewer" pageId="30" pageNumber="43" refString="Roewer, C. F. (1942) Katalog der Araneae von 1758 bis 1940. Bremen 1, 1040 pp." type="book" year="1942" yearSuffix="p">Roewer (1942, p. 265)</bibRefCitation>
listed
<taxonomicName id="4C654D247643FFB69346FB29FC93D0BA" authority="Ausserer 1871" authorityName="Ausserer" authorityYear="1871" box="[322,769,1234,1260]" class="Arachnida" family="Theraphosidae" genus="Phlogiellus" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="B911EAB57643FFB69346FB29FDA0D0BA" box="[322,562,1235,1260]" italics="true" pageId="30" pageNumber="31">Phlogiellus inermis</emphasis>
(
<bibRefCitation id="EFF44B567643FFB69046FB28FD6AD0BA" author="Ausserer" box="[578,760,1234,1260]" pageId="30" pageNumber="42" refString="Ausserer, A. (1871) Beitrage zur Kenntniss der Arachniden - Familie der Territelariae Thorell (Mygalidae Autor). Verhandlungen der k. k. zoologisch - botanischen Gesellschaft in Wien, 21, 17 - 224." type="journal article" year="1871">Ausserer 1871</bibRefCitation>
)
</taxonomicName>
from Lombok Island, however, the listing is probably erroneous as no description or record of
<emphasis id="B911EAB57643FFB6907CFB02FD79D147" box="[632,747,1272,1297]" italics="true" pageId="30" pageNumber="31">P. i n e r m i s</emphasis>
, previous or since, cites this spider from Lombok and the original
<typeStatus id="54DE88057643FFB692FDFAE7FEBED161" box="[249,300,1309,1335]" pageId="30" pageNumber="31">type</typeStatus>
locality for the specimens Ausserer examined were from Java and Sumatra only (
<bibRefCitation id="EFF44B567643FFB696EDFAE7FF4FD10A" author="Ausserer" pageId="30" pageNumber="42" refString="Ausserer, A. (1871) Beitrage zur Kenntniss der Arachniden - Familie der Territelariae Thorell (Mygalidae Autor). Verhandlungen der k. k. zoologisch - botanischen Gesellschaft in Wien, 21, 17 - 224." type="journal article" year="1871" yearSuffix="p">Ausserer 1871, p.189</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>