927 lines
106 KiB
XML
927 lines
106 KiB
XML
<document id="D41C6006133D78E38EB2A85DAD9C3C76" ID-DOI="10.11646/zootaxa.5569.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14703528" ID-ZooBank="96194340-054D-49C0-BD89-D47899BCAF14" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1737371250141" checkinUser="plazi" docAuthor="Yamana, Yusuke, Hirashima, Kentarou, Sato, Masanori & Yamamori, Luna" docDate="2025" docId="03CC879FF9769D66FF63229C8E96A923" docLanguage="en" docName="zootaxa.5569.1.1.pdf" docOrigin="Zootaxa 5569 (1)" docSource="https://doi.org/10.11646/zootaxa.5569.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Patinapta neglectus Yamana, Hirashima, Sato & Yamamori, 2025, sp. nov." docType="treatment" docVersion="1" lastPageNumber="43" masterDocId="FFF5FFE7F9529D4CFFF4243A8C2BAC76" masterDocTitle="First revision of the apodid holothurian genus Patinapta Heding, 1928 (Synaptidae) in Japanese waters, with establishment of four new species from Kyushu Island and Ryukyu Islands, southwestern Japan" masterLastPageNumber="54" masterPageNumber="1" pageNumber="37" updateTime="1737473723657" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:titleInfo id="584AA35BAC4866EAA8E33A34485A7855">
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<mods:title id="B37AA38CB961FF2690409FE5543A7380">First revision of the apodid holothurian genus Patinapta Heding, 1928 (Synaptidae) in Japanese waters, with establishment of four new species from Kyushu Island and Ryukyu Islands, southwestern Japan</mods:title>
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<mods:name id="AE37823D9D0B725B5CBF3EFCAEA2BBB3" type="personal">
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<mods:role id="D42F058A856E01FB0962F9741D1718C7">
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<mods:roleTerm id="298233F8A6A2EC75CDE456280BDF4345">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="92F161DC1EBF2EAACDE003CDBDA9F30F">Yamana, Yusuke</mods:namePart>
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<mods:affiliation id="3300DE568BAF023EAE441A13AA9701C0">Wakayama Prefectural Museum of Natural History (WMNH), Invertebrate Specimen House, Funo 370 - 1, Kainan, Wakayama 642 - 0001, Japan</mods:affiliation>
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</mods:name>
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<mods:name id="D1BE9AF7DF8920F3ADB7BCC9479FB2C0" type="personal">
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<mods:role id="C4F0A484FAB0E28900EEB40ECD90D407">
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<mods:roleTerm id="F6B4E6819502BFE9A01623CD7D3EF405">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="98F67E6B09C6A38366D9CC1E4C8B5A2D">Hirashima, Kentarou</mods:namePart>
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<mods:affiliation id="CC44D6C8AC39B42A64912347DE5BC919">Wakayama Prefectural Museum of Natural History (WMNH), 1 Room Curators Head Office, Funo 370 - 1, Kainan, Wakayama 642 - 0001, Japan</mods:affiliation>
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</mods:name>
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<mods:name id="00CFDD8264D7F4DE7EBF297225AF5248" type="personal">
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<mods:role id="20B3EC2A79CA293913D70D5C1A6EACE3">
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<mods:roleTerm id="A2D464138F17EB8E20F0988A7493921A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7DDD93731D78C6DA88EDE0CB78D19AED">Sato, Masanori</mods:namePart>
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<mods:affiliation id="933D33EDD8F67416802EEC157FE7FC90">Graduate School of Science and Engineering, Kagoshima University, Kagoshima 890 - 0065, Japan & Present address: 5233 Hiramatsu, Aira, Kagoshima 899 - 5652, Japan</mods:affiliation>
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</mods:name>
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<mods:name id="9230B62FC2B00F94CBE6CB02082049A7" type="personal">
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<mods:role id="83239995EB70386E788F2D7FCECE1011">
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<mods:roleTerm id="240141EE3EE9913AE72D9C6BA5BAC403">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FE4F18E009C6067A955C523976CCEAA2">Yamamori, Luna</mods:namePart>
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<mods:affiliation id="A80E4B7E4EEC948FD04323E52E6CDF64">Seto Marine biological laboratory (SMBL) of Kyoto University, 459, Shirahama, Wakayama 649 - 2211, Japan</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="F0C7200C6A8F041696FD0AAD1365E0E1">text</mods:typeOfResource>
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<mods:title id="C8226E7FB1EF2957FE9FE84D7D5F34DF">Zootaxa</mods:title>
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<mods:part id="EC993A9590FC5C957F7815126F8E3979">
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<mods:date id="A273DE1EAF1256C5FA907B9E043DC3F1">2025</mods:date>
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<mods:number id="9F89D206F70D66AA9BCD4B3108BAE024">2025-01-13</mods:number>
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<mods:detail id="B7EF9675F1F201519CE3D7C3978CEDC0" type="volume">
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<mods:number id="16CE23F16197C0AB73F594C3848C098B">5569</mods:number>
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<mods:detail id="1508C4E1097D3F8EA10951E3CE9DB0C8" type="issue">
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<mods:number id="F1F90B34DE1A69CF8634C8C42CB73926">1</mods:number>
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<mods:url id="238BD626AC1DD62D418330A6CA6C4FE7">https://doi.org/10.11646/zootaxa.5569.1.1</mods:url>
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</mods:location>
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<mods:classification id="5BEA93DA1DD1A617C3345FC64DCF2546">journal article</mods:classification>
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<mods:identifier id="3814531C4FBBFF106E96A22C995350CC" type="DOI">10.11646/zootaxa.5569.1.1</mods:identifier>
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<mods:identifier id="5AA9952CD2A89E4351DCB50F3563A03A" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="E231A6CF582908D92E03D8A0C12F5419" type="Zenodo-Dep">14703528</mods:identifier>
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<mods:identifier id="20F4C53514EAE534A6367787C393ACDC" type="ZooBank">96194340-054D-49C0-BD89-D47899BCAF14</mods:identifier>
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</mods:mods>
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<treatment id="03CC879FF9769D66FF63229C8E96A923" ID-DOI="http://doi.org/10.5281/zenodo.14704984" ID-Zenodo-Dep="14704984" LSID="urn:lsid:plazi:treatment:03CC879FF9769D66FF63229C8E96A923" httpUri="http://treatment.plazi.org/id/03CC879FF9769D66FF63229C8E96A923" lastPageId="42" lastPageNumber="43" pageId="36" pageNumber="37">
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<subSubSection id="C37F6502F9769D68FF63229C8DC0AAB6" box="[151,491,1702,1729]" pageId="36" pageNumber="37" type="nomenclature">
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<paragraph id="8BDA3689F9769D68FF63229C8DC0AAB6" blockId="36.[151,732,1702,1800]" box="[151,491,1702,1729]" pageId="36" pageNumber="37">
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<heading id="D09281E5F9769D68FF63229C8DC0AAB6" bold="true" box="[151,491,1702,1729]" fontSize="11" level="1" pageId="36" pageNumber="37" reason="1">
|
||
<emphasis id="B911EA9BF9769D68FF63229C8DC0AAB6" bold="true" box="[151,491,1702,1729]" pageId="36" pageNumber="37">
|
||
<taxonomicName id="4C654D0AF9769D68FF63229C8DADAAB7" authorityName="Yamana & Hirashima & Sato & Yamamori" authorityYear="2025" box="[151,390,1702,1729]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="36" pageNumber="37" phylum="Echinodermata" rank="species" species="neglectus" status="sp. nov.">
|
||
<emphasis id="B911EA9BF9769D68FF63229C8DADAAB7" bold="true" box="[151,390,1702,1729]" italics="true" pageId="36" pageNumber="37">Patinapta neglectus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22257E0F9769D68FE79229C8DC0AAB6" box="[397,491,1702,1728]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F6502F9769D68FF6322F18EF7AA92" box="[151,732,1738,1765]" pageId="36" pageNumber="37" type="vernacular_names">
|
||
<paragraph id="8BDA3689F9769D68FF6322F18EF7AA92" blockId="36.[151,732,1702,1800]" box="[151,732,1738,1765]" pageId="36" pageNumber="37">[New Japanese name: Satsuma-himo-ikari-namako]</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F6502F9769D68FF6322D48DCDAB7E" box="[151,486,1774,1800]" pageId="36" pageNumber="37" type="description">
|
||
<paragraph id="8BDA3689F9769D68FF6322D48DCDAB7E" blockId="36.[151,732,1702,1800]" box="[151,486,1774,1800]" pageId="36" pageNumber="37">
|
||
(
|
||
<figureCitation id="135E2A0CF9769D68FF6B22D48D0AAB7E" box="[159,289,1774,1800]" captionStart-0="FIGURE 18" captionStart-1="FIGURE 19" captionStart-2="FIGURE 20" captionStartId-0="37.[151,250,1319,1343]" captionStartId-1="38.[151,250,1613,1637]" captionStartId-2="41.[151,250,585,609]" captionTargetBox-0="[151,1436,181,1295]" captionTargetBox-1="[155,1433,181,1581]" captionTargetBox-2="[163,1422,188,552]" captionTargetId-0="figure-20@37.[151,1436,181,1295]" captionTargetId-1="figure-29@38.[155,1433,181,1589]" captionTargetId-2="figure-20@41.[151,1435,181,561]" captionTargetPageId-0="37" captionTargetPageId-1="38" captionTargetPageId-2="41" captionText-0="FIGURE 18. External view of a depressed specimen collected from Iso beach inner coast of Kagoshima Bay, Kagoshima, erected as Patinapta neglectus sp. nov., in the present study (holotype specimen, WMNH-2022-INV-154): A, lateral view, anterior left; B, oral-side view of the tentacles; C, outer-side view of the tentacles." captionText-1="FIGURE 19. SEM images of the ossicles extracted from a specimen collected from Iso Beach, Kagoshima Bay Kagoshima, erected as Patinapta neglectus sp. nov., in the present study (holotype specimen, WMNH-2022-INV-154): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." captionText-2="FIGURE 20. Outer-surface view of calcareous ring of the calcareous ring plates of a specimen collected from Iso Beach, Kagoshima Bay Kagoshima, erected as Patinapta neglectus sp. nov., in the present study (holotype specimen, WMNH-2022- INV-154): Positions of the plates denoted by Ludwig’s [1889–1892 (1889), p156] system: RI–RV and IR1–IR5 where R and IR stands for positions for radius and interradius, respectively. Hence RI, mid-ventral; RII, left ventral; RIII, right ventral; RIV, left dorsal; RV, right dorsal; and: IR1, left ventral; IR2, right ventral; IR3-IR3’, left dorso-lateral; IR4-IR4’, right dorso-lateral; IR5, mid-dorsal. Anterior top." figureDoi-0="http://doi.org/10.5281/zenodo.14703570" figureDoi-1="http://doi.org/10.5281/zenodo.14703572" figureDoi-2="http://doi.org/10.5281/zenodo.14703574" httpUri-0="https://zenodo.org/record/14703570/files/figure.png" httpUri-1="https://zenodo.org/record/14703572/files/figure.png" httpUri-2="https://zenodo.org/record/14703574/files/figure.png" pageId="36" pageNumber="37">Figs 18–20</figureCitation>
|
||
;
|
||
<tableCitation id="C6E70332F9769D68FED822D48DA1AB7E" box="[300,394,1774,1800]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionTargetPageId="6" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="36" pageNumber="37">Tables 1</tableCitation>
|
||
,
|
||
<tableCitation id="C6E70332F9769D68FE6C22D48D9EAB7E" box="[408,437,1774,1800]" captionStart="TABLE 11" captionStartId="39.[152,244,152,176]" captionTargetPageId="39" captionText="TABLE 11. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta neglectus sp. nov." pageId="36" pageNumber="37">11</tableCitation>
|
||
,
|
||
<tableCitation id="C6E70332F9769D68FE3522D48DF5AB7E" box="[449,478,1774,1800]" captionStart="TABLE 12" captionStartId="42.[152,244,1412,1436]" captionTargetPageId="42" captionText="TABLE 12. List of WMNH-INV specimens of Patinapta neglectus sp. nov., as of 2024." pageId="36" pageNumber="37">12</tableCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F6502F9769D68FF6323098EADAB1F" pageId="36" pageNumber="37" type="materials_examined">
|
||
<paragraph id="8BDA3689F9769D68FF6323098EADAB1F" blockId="36.[151,1437,1843,1897]" pageId="36" pageNumber="37">
|
||
<materialsCitation id="3B0D3CD4F9769D68FF6323098F57AB3D" box="[151,892,1843,1867]" location="Nagaura-higata" pageId="36" pageNumber="37" specimenCount="2">
|
||
<specimenCount id="9D63FD00F9769D68FF6323098D1DAB3D" box="[151,310,1843,1867]" count="2" pageId="36" pageNumber="37" type="generic">Two specimens</specimenCount>
|
||
from
|
||
<location id="8EBA6052F9769D68FE8223098E39AB3D" LSID="urn:lsid:plazi:treatment:03CC879FF9769D66FF63229C8E96A923:8EBA6052F9769D68FE8223098E39AB3D" box="[374,530,1843,1867]" name="Nagaura-higata" pageId="36" pageNumber="37">Nagaura-higata</location>
|
||
(WMNH-2024-INV-193, 194) and
|
||
</materialsCitation>
|
||
<materialsCitation id="3B0D3CD4F9769D68FC7023098EA9AB1F" location="Bay" pageId="36" pageNumber="37" specimenCount="1" stateProvince="Kagoshima">
|
||
<specimenCount id="9D63FD00F9769D68FC702309883BAB3D" box="[900,1040,1843,1867]" count="1" pageId="36" pageNumber="37" type="generic">one specimen</specimenCount>
|
||
from
|
||
<collectingRegion id="49A1F86BF9769D68FBA5230988EFAB3D" box="[1105,1220,1843,1867]" country="Japan" name="Kagoshima" pageId="36" pageNumber="37">Kagoshima</collectingRegion>
|
||
<location id="8EBA6052F9769D68FB38230988DEAB3D" LSID="urn:lsid:plazi:treatment:03CC879FF9769D66FF63229C8E96A923:8EBA6052F9769D68FB38230988DEAB3D" box="[1228,1269,1843,1867]" name="Bay" pageId="36" pageNumber="37" stateProvince="Kagoshima">Bay</location>
|
||
(WMNH-2022- INV-154) were identified as new to science
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF1A6601F9779D69FF63211D8FFCA9F1" ID-DOI="http://doi.org/10.5281/zenodo.14703570" ID-Zenodo-Dep="14703570" httpUri="https://zenodo.org/record/14703570/files/figure.png" pageId="37" pageNumber="38" startId="37.[151,250,1319,1343]" targetBox="[151,1436,181,1295]" targetPageId="37" targetType="figure">
|
||
<paragraph id="8BDA3689F9779D69FF63211D8FFCA9F1" blockId="37.[151,1436,1319,1415]" pageId="37" pageNumber="38">
|
||
<emphasis id="B911EA9BF9779D69FF63211D8D09A949" bold="true" box="[151,290,1319,1343]" pageId="37" pageNumber="38">FIGURE 18.</emphasis>
|
||
External view of a depressed specimen collected from Iso beach inner coast of Kagoshima Bay, Kagoshima, erected as
|
||
<taxonomicName id="4C654D0AF9779D69FEFC21768DFDA915" authorityName="Yamana & Hirashima & Sato & Yamamori" authorityYear="2025" box="[264,470,1355,1379]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="37" pageNumber="38" phylum="Echinodermata" rank="species" species="neglectus" status="sp. nov.">
|
||
<emphasis id="B911EA9BF9779D69FEFC21768DFDA915" box="[264,470,1355,1379]" italics="true" pageId="37" pageNumber="38">Patinapta neglectus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911EA9BF9779D69FE2B21718E19A915" bold="true" box="[479,562,1355,1379]" pageId="37" pageNumber="38">
|
||
<taxonomicNameLabel id="A22257E0F9779D69FE2B21718E19A915" box="[479,562,1355,1379]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, in the present study (holotype specimen, WMNH-2022-INV-154): A, lateral view, anterior left; B, oral-side view of the tentacles; C, outer-side view of the tentacles.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C37F6502F9779D69FF6321E388B5AABD" pageId="37" pageNumber="38" type="diagnosis">
|
||
<paragraph id="8BDA3689F9779D69FF6321E388B5AABD" blockId="37.[151,1437,1497,1920]" pageId="37" pageNumber="38">
|
||
<emphasis id="B911EA9BF9779D69FF6321E38D39A985" bold="true" box="[151,274,1497,1523]" pageId="37" pageNumber="38">Diagnosis.</emphasis>
|
||
Tentacles possessing sensory cups, situated on basal part of stem, not only on oral-side, but also on outer oral-side, numerous (50–80 cups in total). Body wall soft and thick, covered with verrucous surface layer, which sparsely includes ossicles. Anchor and anchor plate ossicles small. Anchor plate ossicles often exceed 110 μm, while anchor ossicles often exceed 140 μm, rarely equipped with teeth on bills (arms) of anchor ossicles, Total shapes of anchor plates strongly distorted, however basally without a concave. Outer edge of anchor plate ossicles lacking minute teeth at all, while one or two perforations of anchor plate ossicles with only one or two minute teeth. Ciliated funnels attached to inner side body wall along midline of left dorso-lateral interradius (
|
||
<emphasis id="B911EA9BF9779D69FB91228B88A5AABD" box="[1125,1166,1713,1739]" italics="true" pageId="37" pageNumber="38">IR3</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F6502F9779D69FF3322EF88F2AA99" box="[199,1241,1749,1776]" pageId="37" pageNumber="38" type="materials_examined">
|
||
<paragraph id="8BDA3689F9779D69FF3322EF88F2AA99" blockId="37.[151,1437,1497,1920]" box="[199,1241,1749,1776]" pageId="37" pageNumber="38">
|
||
<emphasis id="B911EA9BF9779D69FF3322EF8D78AA99" bold="true" box="[199,339,1749,1775]" pageId="37" pageNumber="38">Type series.</emphasis>
|
||
<typeStatus id="54DE882BF9779D69FEAE22EF8DEEAA99" box="[346,453,1749,1775]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
|
||
, WMNH-2022-INV-154;
|
||
<specimenCount id="9D63FD00F9779D69FD1322EC8FADAA86" box="[743,902,1750,1776]" count="2" pageId="37" pageNumber="38" type="generic" typeStatus="paratype">two paratypes</specimenCount>
|
||
, WMNH-2024-INV-193, 194.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F6502F9779D64FF3322C38CE9A89F" lastPageId="40" lastPageNumber="41" pageId="37" pageNumber="38" type="description">
|
||
<paragraph id="8BDA3689F9779D69FF3322C388A2AB09" blockId="37.[151,1437,1497,1920]" pageId="37" pageNumber="38">
|
||
<emphasis id="B911EA9BF9779D69FF3322C3888FAB65" bold="true" box="[199,1188,1785,1811]" pageId="37" pageNumber="38">
|
||
Description of
|
||
<typeStatus id="54DE882BF9779D69FE8F22C38DC9AB65" box="[379,482,1785,1811]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
|
||
(WMNH-2022-INV-154: middle-size depressed specimen).
|
||
</emphasis>
|
||
External and internal morphologies as follows (
|
||
<tableCitation id="C6E70332F9779D69FE4923278E38AB41" box="[445,531,1821,1847]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionTargetPageId="6" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="37" pageNumber="38">Table 1</tableCitation>
|
||
): 5 year-old preserved specimen pale peanut color (
|
||
<figureCitation id="135E2A0CF9779D69FB962327889CAB41" box="[1122,1207,1821,1847]" captionStart="FIGURE 18" captionStartId="37.[151,250,1319,1343]" captionTargetBox="[151,1436,181,1295]" captionTargetId="figure-20@37.[151,1436,181,1295]" captionTargetPageId="37" captionText="FIGURE 18. External view of a depressed specimen collected from Iso beach inner coast of Kagoshima Bay, Kagoshima, erected as Patinapta neglectus sp. nov., in the present study (holotype specimen, WMNH-2022-INV-154): A, lateral view, anterior left; B, oral-side view of the tentacles; C, outer-side view of the tentacles." figureDoi="http://doi.org/10.5281/zenodo.14703570" httpUri="https://zenodo.org/record/14703570/files/figure.png" pageId="37" pageNumber="38">Fig. 18</figureCitation>
|
||
). Tentacles 12, each with 9 digits of 4 pairs and unpaired 1. On oral-side of tentacle stem, about 30–40 sensory cups, also on outer oral-side tentacle stem, about 20–40 sensory cups present densely arranged in a V- shape band.
|
||
</paragraph>
|
||
<caption id="DF1A6601F9749D6AFF6322778ED5AAA7" ID-DOI="http://doi.org/10.5281/zenodo.14703572" ID-Zenodo-Dep="14703572" httpUri="https://zenodo.org/record/14703572/files/figure.png" pageId="38" pageNumber="39" startId="38.[151,250,1613,1637]" targetBox="[155,1433,181,1581]" targetPageId="38" targetType="figure">
|
||
<paragraph id="8BDA3689F9749D6AFF6322778ED5AAA7" blockId="38.[151,1436,1613,1745]" pageId="38" pageNumber="39">
|
||
<emphasis id="B911EA9BF9749D6AFF6322778D0AAA13" bold="true" box="[151,289,1613,1637]" pageId="38" pageNumber="39">FIGURE 19.</emphasis>
|
||
SEM images of the ossicles extracted from a specimen collected from Iso Beach, Kagoshima Bay Kagoshima, erected as
|
||
<taxonomicName id="4C654D0AF9749D6AFEF322488DF8AAFF" authorityName="Yamana & Hirashima & Sato & Yamamori" authorityYear="2025" box="[263,467,1649,1673]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="38" pageNumber="39" phylum="Echinodermata" rank="species" species="neglectus" status="sp. nov.">
|
||
<emphasis id="B911EA9BF9749D6AFEF322488DF8AAFF" box="[263,467,1649,1673]" italics="true" pageId="38" pageNumber="39">Patinapta neglectus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911EA9BF9749D6AFE28224B8E04AAFF" bold="true" box="[476,559,1649,1673]" pageId="38" pageNumber="39">
|
||
<taxonomicNameLabel id="A22257E0F9749D6AFE28224B8E04AAFF" box="[476,559,1649,1673]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, in the present study (holotype specimen, WMNH-2022-INV-154): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDA3689F9749D6AFF3323198DFBABDF" blockId="38.[151,1436,1827,1961]" pageId="38" pageNumber="39">
|
||
Polian vesicle in medio-ventral position, fusiform, single (
|
||
<tableCitation id="C6E70332F9749D6AFCA523198F8FAB4B" box="[849,932,1827,1853]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionTargetPageId="6" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="38" pageNumber="39">Table 1</tableCitation>
|
||
), stone canal single and thread-like, attached to narrow space between basal parts of tentacle ampullae at mediodorsal (
|
||
<emphasis id="B911EA9BF9749D6AFC3423728FC2AB14" box="[960,1001,1864,1890]" italics="true" pageId="38" pageNumber="39">IR5</emphasis>
|
||
), suddenly turned upward and forward, resulted in it facing forward, distally with moldy surface madreporite. Ciliated funnels in one crowded band along left dorsal interradius (
|
||
<emphasis id="B911EA9BF9749D6AFE6323B58D94ABDF" box="[407,447,1935,1961]" italics="true" pageId="38" pageNumber="39">IR3</emphasis>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF1A6601F9759D6BFF6C24A28957ACC7" box="[152,1404,152,177]" pageId="39" pageNumber="40" startId="39.[152,244,152,176]" targetBox="[163,1418,192,1867]" targetIsTable="true" targetPageId="39" targetType="table">
|
||
<paragraph id="8BDA3689F9759D6BFF6C24A28957ACC7" blockId="39.[152,1404,152,177]" box="[152,1404,152,177]" pageId="39" pageNumber="40">
|
||
<emphasis id="B911EA9BF9759D6BFF6C24A28D30ACC6" bold="true" box="[152,283,152,176]" pageId="39" pageNumber="40">TABLE 11.</emphasis>
|
||
Measurements (mean±SD μm) and counts of ossicles of three specimens of
|
||
<taxonomicName id="4C654D0AF9759D6BFBB824A3890AACC6" authorityName="Yamana & Hirashima & Sato & Yamamori" authorityYear="2025" box="[1100,1313,152,176]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="39" pageNumber="40" phylum="Echinodermata" rank="species" species="neglectus" status="sp. nov.">
|
||
<emphasis id="B911EA9BF9759D6BFBB824A3890AACC6" box="[1100,1313,152,176]" italics="true" pageId="39" pageNumber="40">Patinapta neglectus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22257E0F9759D6BFAD324A38957ACC7" box="[1319,1404,153,177]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDA3689F9759D6BFF5724FB89A1AB3D" pageId="39" pageNumber="40">
|
||
<table id="F965C429F97562B3FF5724FA89A1AB3D" box="[163,1418,192,1867]" gridcols="7" gridrows="42" pageId="39" pageNumber="40">
|
||
<tr id="355534CBF97562B3FF5724FA89A1ACAF" box="[163,1418,192,217]" gridrow="0" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF5724FA8D32ACAF" box="[163,281,192,217]" gridcol="0" gridrow="0" pageId="39" pageNumber="40">Body parts</th>
|
||
<th id="76845DB7F97562B3FEBE24FA8DD0ACAF" box="[330,507,192,217]" gridcol="1" gridrow="0" pageId="39" pageNumber="40">(Ossicle-type)</th>
|
||
<th id="76845DB7F97562B3FDD424FA8F31ACAF" box="[544,794,192,217]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="39" pageNumber="40">2024-INV-193</th>
|
||
<th id="76845DB7F97562B3FCC224FA8863ACAF" box="[822,1096,192,217]" colspan="2" colspanRight="1" gridcol="4" gridrow="0" pageId="39" pageNumber="40">2024-INV-194</th>
|
||
<th id="76845DB7F97562B3FB8324FA89A1ACAF" box="[1143,1418,192,217]" gridcol="6" gridrow="0" pageId="39" pageNumber="40">2022-INV-154</th>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF5724D389A1AD77" box="[163,1418,233,257]" gridrow="1" pageId="39" pageNumber="40" rowspan-4="1">
|
||
<th id="76845DB7F97562B3FF5724D38D32AD77" box="[163,281,233,257]" gridcol="0" gridrow="1" pageId="39" pageNumber="40">Tentacle</th>
|
||
<td id="76845DB7F97562B3FEBE24D38DD0AD77" box="[330,507,233,257]" gridcol="1" gridrow="1" pageId="39" pageNumber="40">
|
||
Rods (
|
||
<emphasis id="B911EA9BF9759D6BFE4E24D08DECAD77" box="[442,455,234,257]" italics="true" pageId="39" pageNumber="40">n</emphasis>
|
||
)
|
||
</td>
|
||
<td id="76845DB7F97562B3FDD424D38F31AD77" box="[544,794,233,257]" colspan="2" colspanRight="1" gridcol="2" gridrow="1" pageId="39" pageNumber="40">9</td>
|
||
<td id="76845DB7F97562B3FC4224D38863AD77" box="[950,1096,233,257]" gridcol="5" gridrow="1" pageId="39" pageNumber="40">14</td>
|
||
<td id="76845DB7F97562B3FB8324D389A1AD77" box="[1143,1418,233,257]" gridcol="6" gridrow="1" pageId="39" pageNumber="40">15</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57252B89A1AD5F" box="[163,1418,273,297]" gridrow="2" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF57252B8D32AD5F" box="[163,281,273,297]" gridcol="0" gridrow="2" pageId="39" pageNumber="40">Length</th>
|
||
<td id="76845DB7F97562B3FEBE252B8DD0AD5F" box="[330,507,273,297]" gridcol="1" gridrow="2" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD4252B8EACAD5F" box="[544,647,273,297]" gridcol="2" gridrow="2" pageId="39" pageNumber="40">54.8±7.4</td>
|
||
<td id="76845DB7F97562B3FD50252B8F31AD5F" box="[676,794,273,297]" gridcol="3" gridrow="2" pageId="39" pageNumber="40">(44–63)</td>
|
||
<td id="76845DB7F97562B3FCC2252B8F88AD5F" box="[822,931,273,297]" gridcol="4" gridrow="2" pageId="39" pageNumber="40">73.6±13.0</td>
|
||
<td id="76845DB7F97562B3FC42252B8863AD5F" box="[950,1096,273,297]" gridcol="5" gridrow="2" pageId="39" pageNumber="40">(51–98)</td>
|
||
<td id="76845DB7F97562B3FB83252B89A1AD5F" box="[1143,1418,273,297]" gridcol="6" gridrow="2" pageId="39" pageNumber="40">74.2±6.7 (62–85)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57250389A1AD24" box="[163,1418,313,338]" gridrow="3" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF5725038D32AD24" box="[163,281,313,338]" gridcol="0" gridrow="3" pageId="39" pageNumber="40">Breadth</th>
|
||
<td id="76845DB7F97562B3FEBE25038DD0AD24" box="[330,507,313,338]" gridcol="1" gridrow="3" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD425038EACAD24" box="[544,647,313,338]" gridcol="2" gridrow="3" pageId="39" pageNumber="40">7.3± 0.9</td>
|
||
<td id="76845DB7F97562B3FD5025038F31AD24" box="[676,794,313,338]" gridcol="3" gridrow="3" pageId="39" pageNumber="40">(6–8)</td>
|
||
<td id="76845DB7F97562B3FCC225038F88AD24" box="[822,931,313,338]" gridcol="4" gridrow="3" pageId="39" pageNumber="40">9.4±2.5</td>
|
||
<td id="76845DB7F97562B3FC4225038863AD24" box="[950,1096,313,338]" gridcol="5" gridrow="3" pageId="39" pageNumber="40">(7–17)</td>
|
||
<td id="76845DB7F97562B3FB83250389A1AD24" box="[1143,1418,313,338]" gridcol="6" gridrow="3" pageId="39" pageNumber="40">10.7±1.9 (8–14)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57255B89A1AD0C" box="[163,1418,353,378]" gridrow="4" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF57255B8D32AD0C" box="[163,281,353,378]" gridcol="0" gridrow="4" pageId="39" pageNumber="40">
|
||
<emphasis id="B911EA9BF9759D6BFF4425588CE4AD0F" box="[176,207,354,377]" italics="true" pageId="39" pageNumber="40">RP</emphasis>
|
||
a (%)
|
||
</th>
|
||
<td id="76845DB7F97562B3FEBE255B8DD0AD0C" box="[330,507,353,378]" gridcol="1" gridrow="4" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD4255B8EACAD0C" box="[544,647,353,378]" gridcol="2" gridrow="4" pageId="39" pageNumber="40">13.5±2.1</td>
|
||
<td id="76845DB7F97562B3FD50255B8F31AD0C" box="[676,794,353,378]" gridcol="3" gridrow="4" pageId="39" pageNumber="40">(10.5–16.3)</td>
|
||
<td id="76845DB7F97562B3FCC2255B8F88AD0C" box="[822,931,353,378]" gridcol="4" gridrow="4" pageId="39" pageNumber="40">12.9±3.2</td>
|
||
<td id="76845DB7F97562B3FC42255B8863AD0C" box="[950,1096,353,378]" gridcol="5" gridrow="4" pageId="39" pageNumber="40">(9.6–21.5)</td>
|
||
<td id="76845DB7F97562B3FB83255B89A1AD0C" box="[1143,1418,353,378]" gridcol="6" gridrow="4" pageId="39" pageNumber="40">14.7±3.8 (9.6–21.5)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF5725B089A1ADD4" box="[163,1418,394,418]" gridrow="5" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF5725B08D32ADD4" box="[163,281,394,418]" gridcol="0" gridrow="5" pageId="39" pageNumber="40">
|
||
<emphasis id="B911EA9BF9759D6BFF4425B08CFBADD7" box="[176,208,394,417]" italics="true" pageId="39" pageNumber="40">RC</emphasis>
|
||
b
|
||
</th>
|
||
<td id="76845DB7F97562B3FEBE25B08DD0ADD4" box="[330,507,394,418]" gridcol="1" gridrow="5" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD425B08EACADD4" box="[544,647,394,418]" gridcol="2" gridrow="5" pageId="39" pageNumber="40">9.1±1.4</td>
|
||
<td id="76845DB7F97562B3FD5025B08F31ADD4" box="[676,794,394,418]" gridcol="3" gridrow="5" pageId="39" pageNumber="40">(6.5–11.1)</td>
|
||
<td id="76845DB7F97562B3FCC225B08F88ADD4" box="[822,931,394,418]" gridcol="4" gridrow="5" pageId="39" pageNumber="40">13.9±5.5</td>
|
||
<td id="76845DB7F97562B3FC4225B08863ADD4" box="[950,1096,394,418]" gridcol="5" gridrow="5" pageId="39" pageNumber="40">(7.1–27.0)</td>
|
||
<td id="76845DB7F97562B3FB8325B089A1ADD4" box="[1143,1418,394,418]" gridcol="6" gridrow="5" pageId="39" pageNumber="40">9.3±4.4 (2.7–17.8)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57258889A1ADBC" box="[163,1418,434,458]" gridrow="6" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF57258889A1ADBC" box="[163,1418,434,458]" colspan="7" colspanRight="6" gridcol="0" gridrow="6" pageId="39" pageNumber="40">Ventral side</th>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF5725E089A1AD85" box="[163,1418,474,499]" gridrow="7" pageId="39" pageNumber="40" rowspan-4="1">
|
||
<th id="76845DB7F97562B3FF5725E08D32AD85" box="[163,281,474,499]" gridcol="0" gridrow="7" pageId="39" pageNumber="40">L-Muscle</th>
|
||
<td id="76845DB7F97562B3FEBE25E08DD0AD85" box="[330,507,474,499]" gridcol="1" gridrow="7" pageId="39" pageNumber="40">
|
||
Granules (
|
||
<emphasis id="B911EA9BF9759D6BFE3A25E18DF0AD84" box="[462,475,475,498]" italics="true" pageId="39" pageNumber="40">n</emphasis>
|
||
)
|
||
</td>
|
||
<td id="76845DB7F97562B3FDD425E08F31AD85" box="[544,794,474,499]" colspan="2" colspanRight="1" gridcol="2" gridrow="7" pageId="39" pageNumber="40">12</td>
|
||
<td id="76845DB7F97562B3FC4225E08863AD85" box="[950,1096,474,499]" gridcol="5" gridrow="7" pageId="39" pageNumber="40">12</td>
|
||
<td id="76845DB7F97562B3FB8325E089A1AD85" box="[1143,1418,474,499]" gridcol="6" gridrow="7" pageId="39" pageNumber="40">13</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57263989A1AE6D" box="[163,1418,515,539]" gridrow="8" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF5726398D32AE6D" box="[163,281,515,539]" gridcol="0" gridrow="8" pageId="39" pageNumber="40">Length</th>
|
||
<td id="76845DB7F97562B3FEBE26398DD0AE6D" box="[330,507,515,539]" gridcol="1" gridrow="8" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD426398EACAE6D" box="[544,647,515,539]" gridcol="2" gridrow="8" pageId="39" pageNumber="40">35.6±2.8</td>
|
||
<td id="76845DB7F97562B3FD5026398F31AE6D" box="[676,794,515,539]" gridcol="3" gridrow="8" pageId="39" pageNumber="40">(31–41)</td>
|
||
<td id="76845DB7F97562B3FCC226398F88AE6D" box="[822,931,515,539]" gridcol="4" gridrow="8" pageId="39" pageNumber="40">34.7±4.8</td>
|
||
<td id="76845DB7F97562B3FC4226398863AE6D" box="[950,1096,515,539]" gridcol="5" gridrow="8" pageId="39" pageNumber="40">(28–43)</td>
|
||
<td id="76845DB7F97562B3FB83263989A1AE6D" box="[1143,1418,515,539]" gridcol="6" gridrow="8" pageId="39" pageNumber="40">37.2±6.1 (28–50)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57261189A1AE35" box="[163,1418,555,579]" gridrow="9" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF5726118D32AE35" box="[163,281,555,579]" gridcol="0" gridrow="9" pageId="39" pageNumber="40">Width</th>
|
||
<td id="76845DB7F97562B3FEBE26118DD0AE35" box="[330,507,555,579]" gridcol="1" gridrow="9" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD426118EACAE35" box="[544,647,555,579]" gridcol="2" gridrow="9" pageId="39" pageNumber="40">24.2± 4.9</td>
|
||
<td id="76845DB7F97562B3FD5026118F31AE35" box="[676,794,555,579]" gridcol="3" gridrow="9" pageId="39" pageNumber="40">(18–34)</td>
|
||
<td id="76845DB7F97562B3FCC226118F88AE35" box="[822,931,555,579]" gridcol="4" gridrow="9" pageId="39" pageNumber="40">15.6±1.9</td>
|
||
<td id="76845DB7F97562B3FC4226118863AE35" box="[950,1096,555,579]" gridcol="5" gridrow="9" pageId="39" pageNumber="40">(11–18)</td>
|
||
<td id="76845DB7F97562B3FB83261189A1AE35" box="[1143,1418,555,579]" gridcol="6" gridrow="9" pageId="39" pageNumber="40">20.4±7.7 (11–38)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57266889A1AE1D" box="[163,1418,594,619]" gridrow="10" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF5726688D32AE1D" box="[163,281,594,619]" gridcol="0" gridrow="10" pageId="39" pageNumber="40">
|
||
<emphasis id="B911EA9BF9759D6BFF44266E8CF9AE1D" box="[176,210,596,619]" italics="true" pageId="39" pageNumber="40">GP</emphasis>
|
||
c
|
||
</th>
|
||
<td id="76845DB7F97562B3FEBE26688DD0AE1D" box="[330,507,594,619]" gridcol="1" gridrow="10" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD426688EACAE1D" box="[544,647,594,619]" gridcol="2" gridrow="10" pageId="39" pageNumber="40">67.6±10.3</td>
|
||
<td id="76845DB7F97562B3FD5026688F31AE1D" box="[676,794,594,619]" gridcol="3" gridrow="10" pageId="39" pageNumber="40">(32.4–80.6)</td>
|
||
<td id="76845DB7F97562B3FCC226688F88AE1D" box="[822,931,594,619]" gridcol="4" gridrow="10" pageId="39" pageNumber="40">45.7±8.0</td>
|
||
<td id="76845DB7F97562B3FC4226688863AE1D" box="[950,1096,594,619]" gridcol="5" gridrow="10" pageId="39" pageNumber="40">(28.9–53.5)</td>
|
||
<td id="76845DB7F97562B3FB83266889A1AE1D" box="[1143,1418,594,619]" gridcol="6" gridrow="10" pageId="39" pageNumber="40">54.9±21.3 (29.5–123)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57264189A1AEE2" box="[163,1418,635,660]" gridrow="11" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF5726418D32AEE2" box="[163,281,635,660]" gridcol="0" gridrow="11" pageId="39" pageNumber="40">
|
||
<emphasis id="B911EA9BF9759D6BFF4426468CCEAEE5" box="[176,229,636,659]" italics="true" pageId="39" pageNumber="40">GCO</emphasis>
|
||
d (%)
|
||
</th>
|
||
<td id="76845DB7F97562B3FEBE26418DD0AEE2" box="[330,507,635,660]" gridcol="1" gridrow="11" pageId="39" pageNumber="40">(Ratio)</td>
|
||
<td id="76845DB7F97562B3FDD426418EACAEE2" box="[544,647,635,660]" gridcol="2" gridrow="11" pageId="39" pageNumber="40">66.7</td>
|
||
<td id="76845DB7F97562B3FD5026418F31AEE2" box="[676,794,635,660]" gridcol="3" gridrow="11" pageId="39" pageNumber="40">(O8: C4)</td>
|
||
<td id="76845DB7F97562B3FCC226418F88AEE2" box="[822,931,635,660]" gridcol="4" gridrow="11" pageId="39" pageNumber="40">50.0</td>
|
||
<td id="76845DB7F97562B3FC4226418863AEE2" box="[950,1096,635,660]" gridcol="5" gridrow="11" pageId="39" pageNumber="40">(O6: C6)</td>
|
||
<td id="76845DB7F97562B3FB83264189A1AEE2" box="[1143,1418,635,660]" gridcol="6" gridrow="11" pageId="39" pageNumber="40">46.2 (O6: C7)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57269E89A1AECA" box="[163,1418,676,700]" gridrow="12" pageId="39" pageNumber="40" rowspan-4="1">
|
||
<th id="76845DB7F97562B3FF57269E8D32AECA" box="[163,281,676,700]" gridcol="0" gridrow="12" pageId="39" pageNumber="40">Body wall</th>
|
||
<td id="76845DB7F97562B3FEBE269E8DD0AECA" box="[330,507,676,700]" gridcol="1" gridrow="12" pageId="39" pageNumber="40">
|
||
Anchor plates (
|
||
<emphasis id="B911EA9BF9759D6BFE13269E8DDFAECD" box="[487,500,676,699]" italics="true" pageId="39" pageNumber="40">n</emphasis>
|
||
)
|
||
</td>
|
||
<td id="76845DB7F97562B3FDD4269E8F31AECA" box="[544,794,676,700]" colspan="2" colspanRight="1" gridcol="2" gridrow="12" pageId="39" pageNumber="40">4</td>
|
||
<td id="76845DB7F97562B3FC42269E8863AECA" box="[950,1096,676,700]" gridcol="5" gridrow="12" pageId="39" pageNumber="40">9</td>
|
||
<td id="76845DB7F97562B3FB83269E89A1AECA" box="[1143,1418,676,700]" gridcol="6" gridrow="12" pageId="39" pageNumber="40">9</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF5726F689A1AE92" box="[163,1418,716,740]" gridrow="13" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF5726F68D32AE92" box="[163,281,716,740]" gridcol="0" gridrow="13" pageId="39" pageNumber="40">Length</th>
|
||
<td id="76845DB7F97562B3FEBE26F68DD0AE92" box="[330,507,716,740]" gridcol="1" gridrow="13" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD426F68EACAE92" box="[544,647,716,740]" gridcol="2" gridrow="13" pageId="39" pageNumber="40">95.3±6.8</td>
|
||
<td id="76845DB7F97562B3FD5026F68F31AE92" box="[676,794,716,740]" gridcol="3" gridrow="13" pageId="39" pageNumber="40">(85–99)</td>
|
||
<td id="76845DB7F97562B3FCC226F68F88AE92" box="[822,931,716,740]" gridcol="4" gridrow="13" pageId="39" pageNumber="40">99.1±8.7</td>
|
||
<td id="76845DB7F97562B3FC4226F68863AE92" box="[950,1096,716,740]" gridcol="5" gridrow="13" pageId="39" pageNumber="40">(85–114)</td>
|
||
<td id="76845DB7F97562B3FB8326F689A1AE92" box="[1143,1418,716,740]" gridcol="6" gridrow="13" pageId="39" pageNumber="40">101.1±7.8 (85–113)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF5726CE89A1AF7A" box="[163,1418,756,780]" gridrow="14" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF5726CE8D32AF7A" box="[163,281,756,780]" gridcol="0" gridrow="14" pageId="39" pageNumber="40">Width</th>
|
||
<td id="76845DB7F97562B3FEBE26CE8DD0AF7A" box="[330,507,756,780]" gridcol="1" gridrow="14" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD426CE8EACAF7A" box="[544,647,756,780]" gridcol="2" gridrow="14" pageId="39" pageNumber="40">67.5±2.9</td>
|
||
<td id="76845DB7F97562B3FD5026CE8F31AF7A" box="[676,794,756,780]" gridcol="3" gridrow="14" pageId="39" pageNumber="40">(64–71)</td>
|
||
<td id="76845DB7F97562B3FCC226CE8F88AF7A" box="[822,931,756,780]" gridcol="4" gridrow="14" pageId="39" pageNumber="40">70.6±7.8</td>
|
||
<td id="76845DB7F97562B3FC4226CE8863AF7A" box="[950,1096,756,780]" gridcol="5" gridrow="14" pageId="39" pageNumber="40">(58–85)</td>
|
||
<td id="76845DB7F97562B3FB8326CE89A1AF7A" box="[1143,1418,756,780]" gridcol="6" gridrow="14" pageId="39" pageNumber="40">66.2±5.7 (53–74)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57272689A1AF43" box="[163,1418,796,821]" gridrow="15" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF5727268D32AF43" box="[163,281,796,821]" gridcol="0" gridrow="15" pageId="39" pageNumber="40">
|
||
<emphasis id="B911EA9BF9759D6BFF4427278CF7AF42" box="[176,220,797,820]" italics="true" pageId="39" pageNumber="40">APS</emphasis>
|
||
e
|
||
</th>
|
||
<td id="76845DB7F97562B3FEBE27268DD0AF43" box="[330,507,796,821]" gridcol="1" gridrow="15" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD427268EACAF43" box="[544,647,796,821]" gridcol="2" gridrow="15" pageId="39" pageNumber="40">0.9±1.5</td>
|
||
<td id="76845DB7F97562B3FD5027268F31AF43" box="[676,794,796,821]" gridcol="3" gridrow="15" pageId="39" pageNumber="40">(0–3.2)</td>
|
||
<td id="76845DB7F97562B3FCC227268F88AF43" box="[822,931,796,821]" gridcol="4" gridrow="15" pageId="39" pageNumber="40">0.6±0.9</td>
|
||
<td id="76845DB7F97562B3FC4227268863AF43" box="[950,1096,796,821]" gridcol="5" gridrow="15" pageId="39" pageNumber="40">(0–2.5)</td>
|
||
<td id="76845DB7F97562B3FB83272689A1AF43" box="[1143,1418,796,821]" gridcol="6" gridrow="15" pageId="39" pageNumber="40">0.7±0.8 (0–2.5)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57277F89A1AF2B" box="[163,1418,837,861]" gridrow="16" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF57277F8D32AF2B" box="[163,281,837,861]" gridcol="0" gridrow="16" pageId="39" pageNumber="40">
|
||
<emphasis id="B911EA9BF9759D6BFF44277C8CFEAF2B" box="[176,213,838,861]" italics="true" pageId="39" pageNumber="40">Ntp</emphasis>
|
||
f
|
||
</th>
|
||
<td id="76845DB7F97562B3FEBE277F8DD0AF2B" box="[330,507,837,861]" gridcol="1" gridrow="16" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD4277F8EACAF2B" box="[544,647,837,861]" gridcol="2" gridrow="16" pageId="39" pageNumber="40">0.8±0.5</td>
|
||
<td id="76845DB7F97562B3FD50277F8F31AF2B" box="[676,794,837,861]" gridcol="3" gridrow="16" pageId="39" pageNumber="40">(0–1)</td>
|
||
<td id="76845DB7F97562B3FCC2277F8F88AF2B" box="[822,931,837,861]" gridcol="4" gridrow="16" pageId="39" pageNumber="40">0.7±0.5</td>
|
||
<td id="76845DB7F97562B3FC42277F8863AF2B" box="[950,1096,837,861]" gridcol="5" gridrow="16" pageId="39" pageNumber="40">(0–1)</td>
|
||
<td id="76845DB7F97562B3FB83277F89A1AF2B" box="[1143,1418,837,861]" gridcol="6" gridrow="16" pageId="39" pageNumber="40">0 -</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57275689A1AFF3" box="[163,1418,876,901]" gridrow="17" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF5727568D32AFF3" box="[163,281,876,901]" gridcol="0" gridrow="17" pageId="39" pageNumber="40">
|
||
<emphasis id="B911EA9BF9759D6BFF4427578CF6AFF2" box="[176,221,877,900]" italics="true" pageId="39" pageNumber="40">APT</emphasis>
|
||
g (%)
|
||
</th>
|
||
<td id="76845DB7F97562B3FEBE27568DD0AFF3" box="[330,507,876,901]" gridcol="1" gridrow="17" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD427568EACAFF3" box="[544,647,876,901]" gridcol="2" gridrow="17" pageId="39" pageNumber="40">5.0±3.4</td>
|
||
<td id="76845DB7F97562B3FD5027568F31AFF3" box="[676,794,876,901]" gridcol="3" gridrow="17" pageId="39" pageNumber="40">(0–7.7)</td>
|
||
<td id="76845DB7F97562B3FCC227568F88AFF3" box="[822,931,876,901]" gridcol="4" gridrow="17" pageId="39" pageNumber="40">3.2±3.4</td>
|
||
<td id="76845DB7F97562B3FC4227568863AFF3" box="[950,1096,876,901]" gridcol="5" gridrow="17" pageId="39" pageNumber="40">(0–9.5)</td>
|
||
<td id="76845DB7F97562B3FB83275689A1AFF3" box="[1143,1418,876,901]" gridcol="6" gridrow="17" pageId="39" pageNumber="40">0 -</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF5727AF89A1AFDB" box="[163,1418,917,941]" gridrow="18" pageId="39" pageNumber="40" rowspan-4="1">
|
||
<th id="76845DB7F97562B3FF5727AF8D32AFDB" box="[163,281,917,941]" gridcol="0" gridrow="18" pageId="39" pageNumber="40">Body wall</th>
|
||
<td id="76845DB7F97562B3FEBE27AF8DD0AFDB" box="[330,507,917,941]" gridcol="1" gridrow="18" pageId="39" pageNumber="40">
|
||
Anchor (
|
||
<emphasis id="B911EA9BF9759D6BFE3227AC8DF8AFDB" box="[454,467,918,941]" italics="true" pageId="39" pageNumber="40">n</emphasis>
|
||
)
|
||
</td>
|
||
<td id="76845DB7F97562B3FDD427AF8F31AFDB" box="[544,794,917,941]" colspan="2" colspanRight="1" gridcol="2" gridrow="18" pageId="39" pageNumber="40">7</td>
|
||
<td id="76845DB7F97562B3FC4227AF8863AFDB" box="[950,1096,917,941]" gridcol="5" gridrow="18" pageId="39" pageNumber="40">7</td>
|
||
<td id="76845DB7F97562B3FB8327AF89A1AFDB" box="[1143,1418,917,941]" gridcol="6" gridrow="18" pageId="39" pageNumber="40">12</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57278789A1AFA0" box="[163,1418,957,982]" gridrow="19" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF5727878D32AFA0" box="[163,281,957,982]" gridcol="0" gridrow="19" pageId="39" pageNumber="40">Length</th>
|
||
<td id="76845DB7F97562B3FEBE27878DD0AFA0" box="[330,507,957,982]" gridcol="1" gridrow="19" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD427878EACAFA0" box="[544,647,957,982]" gridcol="2" gridrow="19" pageId="39" pageNumber="40">120.7±6.9</td>
|
||
<td id="76845DB7F97562B3FD5027878F31AFA0" box="[676,794,957,982]" gridcol="3" gridrow="19" pageId="39" pageNumber="40">(108–127)</td>
|
||
<td id="76845DB7F97562B3FCC227878F88AFA0" box="[822,931,957,982]" gridcol="4" gridrow="19" pageId="39" pageNumber="40">136.1±7.8</td>
|
||
<td id="76845DB7F97562B3FC4227878863AFA0" box="[950,1096,957,982]" gridcol="5" gridrow="19" pageId="39" pageNumber="40">(128–149)</td>
|
||
<td id="76845DB7F97562B3FB83278789A1AFA0" box="[1143,1418,957,982]" gridcol="6" gridrow="19" pageId="39" pageNumber="40">160.2±0.9 (149–188)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF5727DC89A1AF88" box="[163,1418,998,1022]" gridrow="20" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF5727DC8D32AF88" box="[163,281,998,1022]" gridcol="0" gridrow="20" pageId="39" pageNumber="40">Width</th>
|
||
<td id="76845DB7F97562B3FEBE27DC8DD0AF88" box="[330,507,998,1022]" gridcol="1" gridrow="20" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD427DC8EACAF88" box="[544,647,998,1022]" gridcol="2" gridrow="20" pageId="39" pageNumber="40">79.4±3.9</td>
|
||
<td id="76845DB7F97562B3FD5027DC8F31AF88" box="[676,794,998,1022]" gridcol="3" gridrow="20" pageId="39" pageNumber="40">(74–85)</td>
|
||
<td id="76845DB7F97562B3FCC227DC8F88AF88" box="[822,931,998,1022]" gridcol="4" gridrow="20" pageId="39" pageNumber="40">73.7±4.2</td>
|
||
<td id="76845DB7F97562B3FC4227DC8863AF88" box="[950,1096,998,1022]" gridcol="5" gridrow="20" pageId="39" pageNumber="40">(66–77)</td>
|
||
<td id="76845DB7F97562B3FB8327DC89A1AF88" box="[1143,1418,998,1022]" gridcol="6" gridrow="20" pageId="39" pageNumber="40">67.7±1.7 (64–73)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57203489A1A850" box="[163,1418,1038,1062]" gridrow="21" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF5720348D32A850" box="[163,281,1038,1062]" gridcol="0" gridrow="21" pageId="39" pageNumber="40">Teeth</th>
|
||
<td id="76845DB7F97562B3FEBE20348DD0A850" box="[330,507,1038,1062]" gridcol="1" gridrow="21" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD420348EACA850" box="[544,647,1038,1062]" gridcol="2" gridrow="21" pageId="39" pageNumber="40">1.1± 1.6</td>
|
||
<td id="76845DB7F97562B3FD5020348F31A850" box="[676,794,1038,1062]" gridcol="3" gridrow="21" pageId="39" pageNumber="40">(0–4)</td>
|
||
<td id="76845DB7F97562B3FCC220348F88A850" box="[822,931,1038,1062]" gridcol="4" gridrow="21" pageId="39" pageNumber="40">0.6±1.1</td>
|
||
<td id="76845DB7F97562B3FC4220348863A850" box="[950,1096,1038,1062]" gridcol="5" gridrow="21" pageId="39" pageNumber="40">(0–3)</td>
|
||
<td id="76845DB7F97562B3FB83203489A1A850" box="[1143,1418,1038,1062]" gridcol="6" gridrow="21" pageId="39" pageNumber="40">0.9±1.4 (0–4)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57200C89A1A838" box="[163,1418,1078,1102]" gridrow="22" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF57200C8D32A838" box="[163,281,1078,1102]" gridcol="0" gridrow="22" pageId="39" pageNumber="40">
|
||
<emphasis id="B911EA9BF9759D6BFF44200D8CCAA838" box="[176,225,1079,1102]" italics="true" pageId="39" pageNumber="40">ASW</emphasis>
|
||
h(%)
|
||
</th>
|
||
<td id="76845DB7F97562B3FEBE200C8DD0A838" box="[330,507,1078,1102]" gridcol="1" gridrow="22" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD4200C8EACA838" box="[544,647,1078,1102]" gridcol="2" gridrow="22" pageId="39" pageNumber="40">19.7±1.4</td>
|
||
<td id="76845DB7F97562B3FD50200C8F31A838" box="[676,794,1078,1102]" gridcol="3" gridrow="22" pageId="39" pageNumber="40">(18.0–21.4)</td>
|
||
<td id="76845DB7F97562B3FCC2200C8F88A838" box="[822,931,1078,1102]" gridcol="4" gridrow="22" pageId="39" pageNumber="40">21.3±1.6</td>
|
||
<td id="76845DB7F97562B3FC42200C8863A838" box="[950,1096,1078,1102]" gridcol="5" gridrow="22" pageId="39" pageNumber="40">(19.4–24.2)</td>
|
||
<td id="76845DB7F97562B3FB83200C89A1A838" box="[1143,1418,1078,1102]" gridcol="6" gridrow="22" pageId="39" pageNumber="40">26.0±1.1 (24.3–28.3)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57206489A1A801" box="[163,1418,1118,1143]" gridrow="23" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF5720648D32A801" box="[163,281,1118,1143]" gridcol="0" gridrow="23" pageId="39" pageNumber="40">
|
||
<emphasis id="B911EA9BF9759D6BFF4420658CCFA800" box="[176,228,1119,1142]" italics="true" pageId="39" pageNumber="40">AEW</emphasis>
|
||
i (%)
|
||
</th>
|
||
<td id="76845DB7F97562B3FEBE20648DD0A801" box="[330,507,1118,1143]" gridcol="1" gridrow="23" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD420648EACA801" box="[544,647,1118,1143]" gridcol="2" gridrow="23" pageId="39" pageNumber="40">36.7±3.6</td>
|
||
<td id="76845DB7F97562B3FD5020648F31A801" box="[676,794,1118,1143]" gridcol="3" gridrow="23" pageId="39" pageNumber="40">(29.6–40.8)</td>
|
||
<td id="76845DB7F97562B3FCC220648F88A801" box="[822,931,1118,1143]" gridcol="4" gridrow="23" pageId="39" pageNumber="40">47.5±4.3</td>
|
||
<td id="76845DB7F97562B3FC4220648863A801" box="[950,1096,1118,1143]" gridcol="5" gridrow="23" pageId="39" pageNumber="40">(40.0–53.0)</td>
|
||
<td id="76845DB7F97562B3FB83206489A1A801" box="[1143,1418,1118,1143]" gridcol="6" gridrow="23" pageId="39" pageNumber="40">47.9±5.3 (38.5–55.4)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF5720BD89A1A8E9" box="[163,1418,1159,1183]" gridrow="24" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF5720BD89A1A8E9" box="[163,1418,1159,1183]" colspan="7" colspanRight="6" gridcol="0" gridrow="24" pageId="39" pageNumber="40">Dorsal side</th>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57209589A1A8B1" box="[163,1418,1199,1223]" gridrow="25" pageId="39" pageNumber="40" rowspan-4="1">
|
||
<th id="76845DB7F97562B3FF5720958D32A8B1" box="[163,281,1199,1223]" gridcol="0" gridrow="25" pageId="39" pageNumber="40">L-Muscle</th>
|
||
<td id="76845DB7F97562B3FEBE20958DD0A8B1" box="[330,507,1199,1223]" gridcol="1" gridrow="25" pageId="39" pageNumber="40">
|
||
Granules (
|
||
<emphasis id="B911EA9BF9759D6BFE3A208A8DF0A8B1" box="[462,475,1200,1223]" italics="true" pageId="39" pageNumber="40">n</emphasis>
|
||
)
|
||
</td>
|
||
<td id="76845DB7F97562B3FDD420958F31A8B1" box="[544,794,1199,1223]" colspan="2" colspanRight="1" gridcol="2" gridrow="25" pageId="39" pageNumber="40">12</td>
|
||
<td id="76845DB7F97562B3FC4220958863A8B1" box="[950,1096,1199,1223]" gridcol="5" gridrow="25" pageId="39" pageNumber="40">12</td>
|
||
<td id="76845DB7F97562B3FB83209589A1A8B1" box="[1143,1418,1199,1223]" gridcol="6" gridrow="25" pageId="39" pageNumber="40">17</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF5720ED89A1A899" box="[163,1418,1239,1263]" gridrow="26" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF5720ED8D32A899" box="[163,281,1239,1263]" gridcol="0" gridrow="26" pageId="39" pageNumber="40">Length</th>
|
||
<td id="76845DB7F97562B3FEBE20ED8DD0A899" box="[330,507,1239,1263]" gridcol="1" gridrow="26" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD420ED8EACA899" box="[544,647,1239,1263]" gridcol="2" gridrow="26" pageId="39" pageNumber="40">35.6±2.8</td>
|
||
<td id="76845DB7F97562B3FD5020ED8F31A899" box="[676,794,1239,1263]" gridcol="3" gridrow="26" pageId="39" pageNumber="40">(31–41)</td>
|
||
<td id="76845DB7F97562B3FCC220ED8F88A899" box="[822,931,1239,1263]" gridcol="4" gridrow="26" pageId="39" pageNumber="40">30.8±5.0</td>
|
||
<td id="76845DB7F97562B3FC4220ED8863A899" box="[950,1096,1239,1263]" gridcol="5" gridrow="26" pageId="39" pageNumber="40">(26–42)</td>
|
||
<td id="76845DB7F97562B3FB8320ED89A1A899" box="[1143,1418,1239,1263]" gridcol="6" gridrow="26" pageId="39" pageNumber="40">37.9±8.4 (21–54)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF5720C589A1A96E" box="[163,1418,1279,1304]" gridrow="27" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF5720C58D32A96E" box="[163,281,1279,1304]" gridcol="0" gridrow="27" pageId="39" pageNumber="40">Width</th>
|
||
<td id="76845DB7F97562B3FEBE20C58DD0A96E" box="[330,507,1279,1304]" gridcol="1" gridrow="27" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD420C58EACA96E" box="[544,647,1279,1304]" gridcol="2" gridrow="27" pageId="39" pageNumber="40">24.2± 4.9</td>
|
||
<td id="76845DB7F97562B3FD5020C58F31A96E" box="[676,794,1279,1304]" gridcol="3" gridrow="27" pageId="39" pageNumber="40">(18–34)</td>
|
||
<td id="76845DB7F97562B3FCC220C58F88A96E" box="[822,931,1279,1304]" gridcol="4" gridrow="27" pageId="39" pageNumber="40">15.4±3.7</td>
|
||
<td id="76845DB7F97562B3FC4220C58863A96E" box="[950,1096,1279,1304]" gridcol="5" gridrow="27" pageId="39" pageNumber="40">(10–24)</td>
|
||
<td id="76845DB7F97562B3FB8320C589A1A96E" box="[1143,1418,1279,1304]" gridcol="6" gridrow="27" pageId="39" pageNumber="40">20.4±7.7 (11–38)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57211D89A1A936" box="[163,1418,1319,1344]" gridrow="28" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF57211D8D32A936" box="[163,281,1319,1344]" gridcol="0" gridrow="28" pageId="39" pageNumber="40">
|
||
<emphasis id="B911EA9BF9759D6BFF4421128CF9A949" box="[176,210,1320,1343]" italics="true" pageId="39" pageNumber="40">GP</emphasis>
|
||
c
|
||
</th>
|
||
<td id="76845DB7F97562B3FEBE211D8DD0A936" box="[330,507,1319,1344]" gridcol="1" gridrow="28" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD4211D8EACA936" box="[544,647,1319,1344]" gridcol="2" gridrow="28" pageId="39" pageNumber="40">67.6±10.3</td>
|
||
<td id="76845DB7F97562B3FD50211D8F31A936" box="[676,794,1319,1344]" gridcol="3" gridrow="28" pageId="39" pageNumber="40">(32.4–80.6)</td>
|
||
<td id="76845DB7F97562B3FCC2211D8F88A936" box="[822,931,1319,1344]" gridcol="4" gridrow="28" pageId="39" pageNumber="40">51.0±14.6</td>
|
||
<td id="76845DB7F97562B3FC42211D8863A936" box="[950,1096,1319,1344]" gridcol="5" gridrow="28" pageId="39" pageNumber="40">(32.3–88.9)</td>
|
||
<td id="76845DB7F97562B3FB83211D89A1A936" box="[1143,1418,1319,1344]" gridcol="6" gridrow="28" pageId="39" pageNumber="40">54.9±21.3 (29.5–123)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57216A89A1A91E" box="[163,1418,1360,1384]" gridrow="29" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF57216A8D32A91E" box="[163,281,1360,1384]" gridcol="0" gridrow="29" pageId="39" pageNumber="40">
|
||
<emphasis id="B911EA9BF9759D6BFF44216B8CCEA91E" box="[176,229,1361,1384]" italics="true" pageId="39" pageNumber="40">GCO</emphasis>
|
||
d (%)
|
||
</th>
|
||
<td id="76845DB7F97562B3FEBE216A8DD0A91E" box="[330,507,1360,1384]" gridcol="1" gridrow="29" pageId="39" pageNumber="40">(Ratio)</td>
|
||
<td id="76845DB7F97562B3FDD4216A8EACA91E" box="[544,647,1360,1384]" gridcol="2" gridrow="29" pageId="39" pageNumber="40">66.7</td>
|
||
<td id="76845DB7F97562B3FD50216A8F31A91E" box="[676,794,1360,1384]" gridcol="3" gridrow="29" pageId="39" pageNumber="40">(O8: C4)</td>
|
||
<td id="76845DB7F97562B3FCC2216A8F88A91E" box="[822,931,1360,1384]" gridcol="4" gridrow="29" pageId="39" pageNumber="40">33.3</td>
|
||
<td id="76845DB7F97562B3FC42216A8863A91E" box="[950,1096,1360,1384]" gridcol="5" gridrow="29" pageId="39" pageNumber="40">(O4: C8)</td>
|
||
<td id="76845DB7F97562B3FB83216A89A1A91E" box="[1143,1418,1360,1384]" gridcol="6" gridrow="29" pageId="39" pageNumber="40">52.9 (O9: C8)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57214289A1A9E7" box="[163,1418,1400,1425]" gridrow="30" pageId="39" pageNumber="40" rowspan-4="1">
|
||
<th id="76845DB7F97562B3FF5721428D32A9E7" box="[163,281,1400,1425]" gridcol="0" gridrow="30" pageId="39" pageNumber="40">Body wall</th>
|
||
<td id="76845DB7F97562B3FEBE21428DD0A9E7" box="[330,507,1400,1425]" gridcol="1" gridrow="30" pageId="39" pageNumber="40">
|
||
Anchor plates (
|
||
<emphasis id="B911EA9BF9759D6BFE1321438DDFA9E6" box="[487,500,1401,1424]" italics="true" pageId="39" pageNumber="40">n</emphasis>
|
||
)
|
||
</td>
|
||
<td id="76845DB7F97562B3FDD421428F31A9E7" box="[544,794,1400,1425]" colspan="2" colspanRight="1" gridcol="2" gridrow="30" pageId="39" pageNumber="40">4</td>
|
||
<td id="76845DB7F97562B3FC4221428863A9E7" box="[950,1096,1400,1425]" gridcol="5" gridrow="30" pageId="39" pageNumber="40">7</td>
|
||
<td id="76845DB7F97562B3FB83214289A1A9E7" box="[1143,1418,1400,1425]" gridcol="6" gridrow="30" pageId="39" pageNumber="40">11</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57219A89A1A9CF" box="[163,1418,1440,1465]" gridrow="31" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF57219A8D32A9CF" box="[163,281,1440,1465]" gridcol="0" gridrow="31" pageId="39" pageNumber="40">Length</th>
|
||
<td id="76845DB7F97562B3FEBE219A8DD0A9CF" box="[330,507,1440,1465]" gridcol="1" gridrow="31" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD4219A8EACA9CF" box="[544,647,1440,1465]" gridcol="2" gridrow="31" pageId="39" pageNumber="40">114.8±3.2</td>
|
||
<td id="76845DB7F97562B3FD50219A8F31A9CF" box="[676,794,1440,1465]" gridcol="3" gridrow="31" pageId="39" pageNumber="40">(110–117)</td>
|
||
<td id="76845DB7F97562B3FCC2219A8F88A9CF" box="[822,931,1440,1465]" gridcol="4" gridrow="31" pageId="39" pageNumber="40">105.9±4.9</td>
|
||
<td id="76845DB7F97562B3FC42219A8863A9CF" box="[950,1096,1440,1465]" gridcol="5" gridrow="31" pageId="39" pageNumber="40">(99–113)</td>
|
||
<td id="76845DB7F97562B3FB83219A89A1A9CF" box="[1143,1418,1440,1465]" gridcol="6" gridrow="31" pageId="39" pageNumber="40">108.0±8.8 (94–123)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF5721F389A1A997" box="[163,1418,1481,1505]" gridrow="32" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF5721F38D32A997" box="[163,281,1481,1505]" gridcol="0" gridrow="32" pageId="39" pageNumber="40">Width</th>
|
||
<td id="76845DB7F97562B3FEBE21F38DD0A997" box="[330,507,1481,1505]" gridcol="1" gridrow="32" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD421F38EACA997" box="[544,647,1481,1505]" gridcol="2" gridrow="32" pageId="39" pageNumber="40">74.5±7.2</td>
|
||
<td id="76845DB7F97562B3FD5021F38F31A997" box="[676,794,1481,1505]" gridcol="3" gridrow="32" pageId="39" pageNumber="40">(68–83)</td>
|
||
<td id="76845DB7F97562B3FCC221F38F88A997" box="[822,931,1481,1505]" gridcol="4" gridrow="32" pageId="39" pageNumber="40">73.1±3.9</td>
|
||
<td id="76845DB7F97562B3FC4221F38863A997" box="[950,1096,1481,1505]" gridcol="5" gridrow="32" pageId="39" pageNumber="40">(68–81)</td>
|
||
<td id="76845DB7F97562B3FB8321F389A1A997" box="[1143,1418,1481,1505]" gridcol="6" gridrow="32" pageId="39" pageNumber="40">69.3±3.3 (65–75)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF5721CA89A1AA7F" box="[163,1418,1520,1545]" gridrow="33" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF5721CA8D32AA7F" box="[163,281,1520,1545]" gridcol="0" gridrow="33" pageId="39" pageNumber="40">
|
||
<emphasis id="B911EA9BF9759D6BFF4421CB8CF7AA7E" box="[176,220,1521,1544]" italics="true" pageId="39" pageNumber="40">APS</emphasis>
|
||
e
|
||
</th>
|
||
<td id="76845DB7F97562B3FEBE21CA8DD0AA7F" box="[330,507,1520,1545]" gridcol="1" gridrow="33" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD421CA8EACAA7F" box="[544,647,1520,1545]" gridcol="2" gridrow="33" pageId="39" pageNumber="40">1.5±1.4</td>
|
||
<td id="76845DB7F97562B3FD5021CA8F31AA7F" box="[676,794,1520,1545]" gridcol="3" gridrow="33" pageId="39" pageNumber="40">(0–2.9)</td>
|
||
<td id="76845DB7F97562B3FCC221CA8F88AA7F" box="[822,931,1520,1545]" gridcol="4" gridrow="33" pageId="39" pageNumber="40">0.6±0.9</td>
|
||
<td id="76845DB7F97562B3FC4221CA8863AA7F" box="[950,1096,1520,1545]" gridcol="5" gridrow="33" pageId="39" pageNumber="40">(0–2.5)</td>
|
||
<td id="76845DB7F97562B3FB8321CA89A1AA7F" box="[1143,1418,1520,1545]" gridcol="6" gridrow="33" pageId="39" pageNumber="40">1.8±3.2 (0–11.3)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57222389A1AA47" box="[163,1418,1561,1585]" gridrow="34" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF5722238D32AA47" box="[163,281,1561,1585]" gridcol="0" gridrow="34" pageId="39" pageNumber="40">
|
||
<emphasis id="B911EA9BF9759D6BFF4422208CFEAA47" box="[176,213,1562,1585]" italics="true" pageId="39" pageNumber="40">Ntp</emphasis>
|
||
f
|
||
</th>
|
||
<td id="76845DB7F97562B3FEBE22238DD0AA47" box="[330,507,1561,1585]" gridcol="1" gridrow="34" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD422238EACAA47" box="[544,647,1561,1585]" gridcol="2" gridrow="34" pageId="39" pageNumber="40">0.75±0.5</td>
|
||
<td id="76845DB7F97562B3FD5022238F31AA47" box="[676,794,1561,1585]" gridcol="3" gridrow="34" pageId="39" pageNumber="40">(0–1)</td>
|
||
<td id="76845DB7F97562B3FCC222238F88AA47" box="[822,931,1561,1585]" gridcol="4" gridrow="34" pageId="39" pageNumber="40">0.7±0.5</td>
|
||
<td id="76845DB7F97562B3FC4222238863AA47" box="[950,1096,1561,1585]" gridcol="5" gridrow="34" pageId="39" pageNumber="40">(0–1)</td>
|
||
<td id="76845DB7F97562B3FB83222389A1AA47" box="[1143,1418,1561,1585]" gridcol="6" gridrow="34" pageId="39" pageNumber="40">0.83±1.0 (0–2)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57227B89A1AA2C" box="[163,1418,1601,1626]" gridrow="35" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF57227B8D32AA2C" box="[163,281,1601,1626]" gridcol="0" gridrow="35" pageId="39" pageNumber="40">
|
||
<emphasis id="B911EA9BF9759D6BFF4422788CF6AA2F" box="[176,221,1602,1625]" italics="true" pageId="39" pageNumber="40">APT</emphasis>
|
||
g (%)
|
||
</th>
|
||
<td id="76845DB7F97562B3FEBE227B8DD0AA2C" box="[330,507,1601,1626]" gridcol="1" gridrow="35" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD4227B8EACAA2C" box="[544,647,1601,1626]" gridcol="2" gridrow="35" pageId="39" pageNumber="40">4.9±3.4</td>
|
||
<td id="76845DB7F97562B3FD50227B8F31AA2C" box="[676,794,1601,1626]" gridcol="3" gridrow="35" pageId="39" pageNumber="40">(0–7.7)</td>
|
||
<td id="76845DB7F97562B3FCC2227B8F88AA2C" box="[822,931,1601,1626]" gridcol="4" gridrow="35" pageId="39" pageNumber="40">3.5±2.5</td>
|
||
<td id="76845DB7F97562B3FC42227B8863AA2C" box="[950,1096,1601,1626]" gridcol="5" gridrow="35" pageId="39" pageNumber="40">(0–5.9)</td>
|
||
<td id="76845DB7F97562B3FB83227B89A1AA2C" box="[1143,1418,1601,1626]" gridcol="6" gridrow="35" pageId="39" pageNumber="40">3.5±4.4 (0–11.8)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57225089A1AAF4" box="[163,1418,1642,1666]" gridrow="36" pageId="39" pageNumber="40" rowspan-4="1">
|
||
<th id="76845DB7F97562B3FF5722508D32AAF4" box="[163,281,1642,1666]" gridcol="0" gridrow="36" pageId="39" pageNumber="40">Body wall</th>
|
||
<td id="76845DB7F97562B3FEBE22508DD0AAF4" box="[330,507,1642,1666]" gridcol="1" gridrow="36" pageId="39" pageNumber="40">
|
||
Anchor (
|
||
<emphasis id="B911EA9BF9759D6BFE3222518DF8AAF4" box="[454,467,1643,1666]" italics="true" pageId="39" pageNumber="40">n</emphasis>
|
||
)
|
||
</td>
|
||
<td id="76845DB7F97562B3FDD422508F31AAF4" box="[544,794,1642,1666]" colspan="2" colspanRight="1" gridcol="2" gridrow="36" pageId="39" pageNumber="40">9</td>
|
||
<td id="76845DB7F97562B3FC4222508863AAF4" box="[950,1096,1642,1666]" gridcol="5" gridrow="36" pageId="39" pageNumber="40">7</td>
|
||
<td id="76845DB7F97562B3FB83225089A1AAF4" box="[1143,1418,1642,1666]" gridcol="6" gridrow="36" pageId="39" pageNumber="40">11</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF5722A889A1AADC" box="[163,1418,1682,1706]" gridrow="37" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF5722A88D32AADC" box="[163,281,1682,1706]" gridcol="0" gridrow="37" pageId="39" pageNumber="40">Length</th>
|
||
<td id="76845DB7F97562B3FEBE22A88DD0AADC" box="[330,507,1682,1706]" gridcol="1" gridrow="37" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD422A88EACAADC" box="[544,647,1682,1706]" gridcol="2" gridrow="37" pageId="39" pageNumber="40">123.3±9.7</td>
|
||
<td id="76845DB7F97562B3FD5022A88F31AADC" box="[676,794,1682,1706]" gridcol="3" gridrow="37" pageId="39" pageNumber="40">(106–136)</td>
|
||
<td id="76845DB7F97562B3FCC222A88863AADC" box="[822,1096,1682,1706]" colspan="2" colspanRight="1" gridcol="4" gridrow="37" pageId="39" pageNumber="40">134.1±11.5 (120–147)</td>
|
||
<td id="76845DB7F97562B3FB8322A889A1AADC" box="[1143,1418,1682,1706]" gridcol="6" gridrow="37" pageId="39" pageNumber="40">158.2±11.8 (140–178)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57228089A1AAA5" box="[163,1418,1722,1747]" gridrow="38" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF5722808D32AAA5" box="[163,281,1722,1747]" gridcol="0" gridrow="38" pageId="39" pageNumber="40">Width</th>
|
||
<td id="76845DB7F97562B3FEBE22808DD0AAA5" box="[330,507,1722,1747]" gridcol="1" gridrow="38" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD422808EACAAA5" box="[544,647,1722,1747]" gridcol="2" gridrow="38" pageId="39" pageNumber="40">76.6±6.4</td>
|
||
<td id="76845DB7F97562B3FD5022808F31AAA5" box="[676,794,1722,1747]" gridcol="3" gridrow="38" pageId="39" pageNumber="40">(66–84)</td>
|
||
<td id="76845DB7F97562B3FCC222808F88AAA5" box="[822,931,1722,1747]" gridcol="4" gridrow="38" pageId="39" pageNumber="40">78.9±4.5</td>
|
||
<td id="76845DB7F97562B3FC4222808863AAA5" box="[950,1096,1722,1747]" gridcol="5" gridrow="38" pageId="39" pageNumber="40">(73–84)</td>
|
||
<td id="76845DB7F97562B3FB83228089A1AAA5" box="[1143,1418,1722,1747]" gridcol="6" gridrow="38" pageId="39" pageNumber="40">72.4±4.1 (66–78)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF5722D989A1AA8D" box="[163,1418,1763,1787]" gridrow="39" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF5722D98D32AA8D" box="[163,281,1763,1787]" gridcol="0" gridrow="39" pageId="39" pageNumber="40">Teeth</th>
|
||
<td id="76845DB7F97562B3FEBE22D98DD0AA8D" box="[330,507,1763,1787]" gridcol="1" gridrow="39" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD422D98EACAA8D" box="[544,647,1763,1787]" gridcol="2" gridrow="39" pageId="39" pageNumber="40">1.4±1.4</td>
|
||
<td id="76845DB7F97562B3FD5022D98F31AA8D" box="[676,794,1763,1787]" gridcol="3" gridrow="39" pageId="39" pageNumber="40">(0–3)</td>
|
||
<td id="76845DB7F97562B3FCC222D98F88AA8D" box="[822,931,1763,1787]" gridcol="4" gridrow="39" pageId="39" pageNumber="40">1.6±1.4</td>
|
||
<td id="76845DB7F97562B3FC4222D98863AA8D" box="[950,1096,1763,1787]" gridcol="5" gridrow="39" pageId="39" pageNumber="40">(0–3)</td>
|
||
<td id="76845DB7F97562B3FB8322D989A1AA8D" box="[1143,1418,1763,1787]" gridcol="6" gridrow="39" pageId="39" pageNumber="40">0.9±1.1 (0–3)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57233189A1AB55" box="[163,1418,1803,1827]" gridrow="40" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF5723318D32AB55" box="[163,281,1803,1827]" gridcol="0" gridrow="40" pageId="39" pageNumber="40">
|
||
<emphasis id="B911EA9BF9759D6BFF4423318CCAAB54" box="[176,225,1803,1826]" italics="true" pageId="39" pageNumber="40">ASW</emphasis>
|
||
h(%)
|
||
</th>
|
||
<td id="76845DB7F97562B3FEBE23318DD0AB55" box="[330,507,1803,1827]" gridcol="1" gridrow="40" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD423318EACAB55" box="[544,647,1803,1827]" gridcol="2" gridrow="40" pageId="39" pageNumber="40">20.8± 1.4</td>
|
||
<td id="76845DB7F97562B3FD5023318F31AB55" box="[676,794,1803,1827]" gridcol="3" gridrow="40" pageId="39" pageNumber="40">(18.9–22.7)</td>
|
||
<td id="76845DB7F97562B3FCC223318F88AB55" box="[822,931,1803,1827]" gridcol="4" gridrow="40" pageId="39" pageNumber="40">18.8±0.8</td>
|
||
<td id="76845DB7F97562B3FC4223318863AB55" box="[950,1096,1803,1827]" gridcol="5" gridrow="40" pageId="39" pageNumber="40">(17.3–19.6)</td>
|
||
<td id="76845DB7F97562B3FB83233189A1AB55" box="[1143,1418,1803,1827]" gridcol="6" gridrow="40" pageId="39" pageNumber="40">24.5±1.2 (22.7–26.8)</td>
|
||
</tr>
|
||
<tr id="355534CBF97562B3FF57230989A1AB3D" box="[163,1418,1843,1867]" gridrow="41" pageId="39" pageNumber="40">
|
||
<th id="76845DB7F97562B3FF5723098D32AB3D" box="[163,281,1843,1867]" gridcol="0" gridrow="41" pageId="39" pageNumber="40">
|
||
<emphasis id="B911EA9BF9759D6BFF44230E8CCFAB3D" box="[176,228,1844,1867]" italics="true" pageId="39" pageNumber="40">AEW</emphasis>
|
||
i (%)
|
||
</th>
|
||
<td id="76845DB7F97562B3FEBE23098DD0AB3D" box="[330,507,1843,1867]" gridcol="1" gridrow="41" pageId="39" pageNumber="40">(Range)</td>
|
||
<td id="76845DB7F97562B3FDD423098EACAB3D" box="[544,647,1843,1867]" gridcol="2" gridrow="41" pageId="39" pageNumber="40">37.9±4.5</td>
|
||
<td id="76845DB7F97562B3FD5023098F31AB3D" box="[676,794,1843,1867]" gridcol="3" gridrow="41" pageId="39" pageNumber="40">(30.5–47.3)</td>
|
||
<td id="76845DB7F97562B3FCC223098F88AB3D" box="[822,931,1843,1867]" gridcol="4" gridrow="41" pageId="39" pageNumber="40">45.2±2.7</td>
|
||
<td id="76845DB7F97562B3FC4223098863AB3D" box="[950,1096,1843,1867]" gridcol="5" gridrow="41" pageId="39" pageNumber="40">(42.5–49.3)</td>
|
||
<td id="76845DB7F97562B3FB83230989A1AB3D" box="[1143,1418,1843,1867]" gridcol="6" gridrow="41" pageId="39" pageNumber="40">48.9±3.2 (42.9–55.1)</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BDA3689F9759D6BFF63236289B8AB06" blockId="39.[151,1427,1880,1905]" box="[151,1427,1880,1905]" pageId="39" pageNumber="40">
|
||
<tableNote id="76833707F9759D6BFF63236289B8AB06" box="[151,1427,1880,1905]" pageId="39" pageNumber="40" targetBox="[163,1418,192,1867]" targetPageId="39">
|
||
a–i, (see, Materials and Methods) Indicators for ossicle shapes, and one parameter (
|
||
<emphasis id="B911EA9BF9759D6BFBFD23638804AB06" box="[1033,1071,1881,1904]" italics="true" pageId="39" pageNumber="40">Ntp</emphasis>
|
||
, number of teethed perforations).
|
||
</tableNote>
|
||
</paragraph>
|
||
<paragraph id="8BDA3689F97A9D64FF3324AD8CF1AC8F" blockId="40.[151,1436,151,393]" pageId="40" pageNumber="41">Inside body, two tufts of gonad tubules attached with both sides of anterior dorsal mesentery. Intestine canal lacking loop, upper and under sides of intestine canal left blank, no whitish tubule organs adhered to intestine canal.</paragraph>
|
||
<paragraph id="8BDA3689F97A9D64FF3325398FC6ADFF" blockId="40.[151,1436,151,393]" pageId="40" pageNumber="41">Body wall smooth surface, soft but thick, covered by a surface layer like "intestinal villi," with ossicle arranging latitudinal layout of all as sets of anchor and plate. Both sides along five longitudinal muscles with distal ends of anchor ossicles facing towards interradial area, however ossicles on radial body walls upon longitudinal muscles and interradial body walls have distal ends randomly facing either direction.</paragraph>
|
||
<paragraph id="8BDA3689F97A9D64FF6325E18E86AD82" blockId="40.[151,685,474,501]" box="[151,685,474,501]" pageId="40" pageNumber="41">
|
||
<heading id="D09281E5F97A9D64FF6325E18E86AD82" bold="true" box="[151,685,474,501]" fontSize="11" level="1" pageId="40" pageNumber="41" reason="1">
|
||
<emphasis id="B911EA9BF97A9D64FF6325E18E86AD82" bold="true" box="[151,685,474,501]" pageId="40" pageNumber="41">The present results of ossicle morphologies</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDA3689F97A9D64FF6326198E76AF63" blockId="40.[151,1437,546,1401]" pageId="40" pageNumber="41">
|
||
Body wall ossicles anchors and anchor plates (
|
||
<figureCitation id="135E2A0CF97A9D64FD4026198F69AE4B" box="[692,834,547,573]" captionStart="FIGURE 19" captionStartId="38.[151,250,1613,1637]" captionTargetBox="[155,1433,181,1581]" captionTargetId="figure-29@38.[155,1433,181,1589]" captionTargetPageId="38" captionText="FIGURE 19. SEM images of the ossicles extracted from a specimen collected from Iso Beach, Kagoshima Bay Kagoshima, erected as Patinapta neglectus sp. nov., in the present study (holotype specimen, WMNH-2022-INV-154): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703572" httpUri="https://zenodo.org/record/14703572/files/figure.png" pageId="40" pageNumber="41">Fig. 19B; D</figureCitation>
|
||
,
|
||
<tableCitation id="C6E70332F97A9D64FCBB26198F9FAE4B" box="[847,948,547,573]" captionStart="TABLE 11" captionStartId="39.[152,244,152,176]" captionTargetPageId="39" captionText="TABLE 11. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta neglectus sp. nov." pageId="40" pageNumber="41">Table 11</tableCitation>
|
||
). Anchor plates ranged within 85–123 μm in length. Size of perforations not noticeably variable between central area and marginal area, and instead almost constant between central area and marginal area, because of only few marginal perforations rarely equipped. Plates showed strong distortion, with approximate hexagonal arrangement perforations. Perforations of anchor plate ossicles mostly teethless, but seldomly possessing 1–2 minute teeth, while outer edge of all anchor plate ossicles lacking minute teeth. Two calculated indicators for anchor plates and a parameter varied;
|
||
<emphasis id="B911EA9BF97A9D64FB8926ED8886AE87" box="[1149,1197,727,753]" italics="true" pageId="40" pageNumber="41">APS</emphasis>
|
||
= 0–11.3;
|
||
<emphasis id="B911EA9BF97A9D64FAD126ED8966AE86" box="[1317,1357,727,752]" italics="true" pageId="40" pageNumber="41">Ntp</emphasis>
|
||
= 0–2;
|
||
<emphasis id="B911EA9BF97A9D64FF6326C18CE2AF63" box="[151,201,763,789]" italics="true" pageId="40" pageNumber="41">APT</emphasis>
|
||
= 0–11.8% (
|
||
<figureCitation id="135E2A0CF97A9D64FEAD26C18DC9AF63" box="[345,482,763,789]" captionStart="FIGURE 19" captionStartId="38.[151,250,1613,1637]" captionTargetBox="[155,1433,181,1581]" captionTargetId="figure-29@38.[155,1433,181,1589]" captionTargetPageId="38" captionText="FIGURE 19. SEM images of the ossicles extracted from a specimen collected from Iso Beach, Kagoshima Bay Kagoshima, erected as Patinapta neglectus sp. nov., in the present study (holotype specimen, WMNH-2022-INV-154): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703572" httpUri="https://zenodo.org/record/14703572/files/figure.png" pageId="40" pageNumber="41">Fig. 19B; D</figureCitation>
|
||
,
|
||
<tableCitation id="C6E70332F97A9D64FE1926C18E64AF63" box="[493,591,763,789]" captionStart="TABLE 11" captionStartId="39.[152,244,152,176]" captionTargetPageId="39" captionText="TABLE 11. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta neglectus sp. nov." pageId="40" pageNumber="41">Table 11</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDA3689F97A9D64FF3327258DA0AFBF" blockId="40.[151,1437,546,1401]" pageId="40" pageNumber="41">
|
||
Anchor ossicles ranged within 140–188 μm in length (
|
||
<figureCitation id="135E2A0CF97A9D64FCDE27258F98AF4F" box="[810,947,799,825]" captionStart="FIGURE 19" captionStartId="38.[151,250,1613,1637]" captionTargetBox="[155,1433,181,1581]" captionTargetId="figure-29@38.[155,1433,181,1589]" captionTargetPageId="38" captionText="FIGURE 19. SEM images of the ossicles extracted from a specimen collected from Iso Beach, Kagoshima Bay Kagoshima, erected as Patinapta neglectus sp. nov., in the present study (holotype specimen, WMNH-2022-INV-154): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703572" httpUri="https://zenodo.org/record/14703572/files/figure.png" pageId="40" pageNumber="41">Fig. 19B; D</figureCitation>
|
||
,
|
||
<tableCitation id="C6E70332F97A9D64FC4927258834AF4F" box="[957,1055,799,825]" captionStart="TABLE 11" captionStartId="39.[152,244,152,176]" captionTargetPageId="39" captionText="TABLE 11. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta neglectus sp. nov." pageId="40" pageNumber="41">Table 11</tableCitation>
|
||
). Most anchor arms (bills) devoid of minute teeth on their outer tips, numbers of teeth very small within 0–4 (mostly
|
||
<quantity id="4C9D9B6CF97A9D64FBBB277988B3AF2B" box="[1103,1176,835,861]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" metricValueMax="5.08" metricValueMin="0.0" pageId="40" pageNumber="41" unit="in" value="1.0" valueMax="2.0" valueMin="0.0">0–2 in</quantity>
|
||
one arm of the anchor ossicle), and anchor ossicles of this specimen possessed narrow stem and also relatively narrow anchor arm width, with
|
||
<emphasis id="B911EA9BF97A9D64FF2627B18D23AFD3" box="[210,264,907,933]" italics="true" pageId="40" pageNumber="41">ASW</emphasis>
|
||
= 22.7–28.3%;
|
||
<emphasis id="B911EA9BF97A9D64FE3627B18DD7AFD3" box="[450,508,907,933]" italics="true" pageId="40" pageNumber="41">AEW</emphasis>
|
||
= 38.5–55.1%. T-shaped basal end of the anchor ossicle of this species crescent-shaped (
|
||
<figureCitation id="135E2A0CF97A9D64FF0027958D55AFBF" box="[244,382,943,969]" captionStart="FIGURE 19" captionStartId="38.[151,250,1613,1637]" captionTargetBox="[155,1433,181,1581]" captionTargetId="figure-29@38.[155,1433,181,1589]" captionTargetPageId="38" captionText="FIGURE 19. SEM images of the ossicles extracted from a specimen collected from Iso Beach, Kagoshima Bay Kagoshima, erected as Patinapta neglectus sp. nov., in the present study (holotype specimen, WMNH-2022-INV-154): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703572" httpUri="https://zenodo.org/record/14703572/files/figure.png" pageId="40" pageNumber="41">Fig. 19B; D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDA3689F97A9D64FF3327E98D4BA80B" blockId="40.[151,1437,546,1401]" pageId="40" pageNumber="41">
|
||
Body longitudinal muscle, granule ossicles of this specimen ranged within 21–54 μm in length (
|
||
<figureCitation id="135E2A0CF97A9D64FAFB27E989BCAF9B" box="[1295,1431,979,1005]" captionStart="FIGURE 19" captionStartId="38.[151,250,1613,1637]" captionTargetBox="[155,1433,181,1581]" captionTargetId="figure-29@38.[155,1433,181,1589]" captionTargetPageId="38" captionText="FIGURE 19. SEM images of the ossicles extracted from a specimen collected from Iso Beach, Kagoshima Bay Kagoshima, erected as Patinapta neglectus sp. nov., in the present study (holotype specimen, WMNH-2022-INV-154): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703572" httpUri="https://zenodo.org/record/14703572/files/figure.png" pageId="40" pageNumber="41">Fig. 19C; E</figureCitation>
|
||
,
|
||
<tableCitation id="C6E70332F97A9D64FF6327CD8CD0A867" box="[151,251,1015,1041]" captionStart="TABLE 11" captionStartId="39.[152,244,152,176]" captionTargetPageId="39" captionText="TABLE 11. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta neglectus sp. nov." pageId="40" pageNumber="41">Table 11</tableCitation>
|
||
). Both C-shaped and O-shaped granule ossicles, mostly flattened, distorted and roundish triangle form, rarely not distorted and usual oval form. The calculated indicator for granule proportions were varied within
|
||
<emphasis id="B911EA9BF97A9D64FA83202189B7A843" box="[1399,1436,1051,1077]" italics="true" pageId="40" pageNumber="41">GP</emphasis>
|
||
= 29.5–123.0%;
|
||
<emphasis id="B911EA9BF97A9D64FEA720058DA5A82F" box="[339,398,1087,1113]" italics="true" pageId="40" pageNumber="41">GCO</emphasis>
|
||
= 46.1–52.9% and did not have noticeable differences in amounts between the O-shape and C-shape granules.
|
||
</paragraph>
|
||
<paragraph id="8BDA3689F97A9D64FF3320BD8CE9A89F" blockId="40.[151,1437,546,1401]" pageId="40" pageNumber="41">
|
||
Tentacle ossicles rods, often with complex shape, ranging within 62–85 μm in length (
|
||
<figureCitation id="135E2A0CF97A9D64FB7B20BD88DDA8D7" box="[1167,1270,1159,1185]" captionStart="FIGURE 19" captionStartId="38.[151,250,1613,1637]" captionTargetBox="[155,1433,181,1581]" captionTargetId="figure-29@38.[155,1433,181,1589]" captionTargetPageId="38" captionText="FIGURE 19. SEM images of the ossicles extracted from a specimen collected from Iso Beach, Kagoshima Bay Kagoshima, erected as Patinapta neglectus sp. nov., in the present study (holotype specimen, WMNH-2022-INV-154): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703572" httpUri="https://zenodo.org/record/14703572/files/figure.png" pageId="40" pageNumber="41">Fig. 19A</figureCitation>
|
||
,
|
||
<tableCitation id="C6E70332F97A9D64FAF520BD897DA8D7" box="[1281,1366,1159,1185]" captionStart="TABLE 9" captionStartId="32.[152,243,152,176]" captionTargetPageId="32" captionText="TABLE 9. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta parvaspiculus sp. nov." pageId="40" pageNumber="41">Table 9</tableCitation>
|
||
). Two calculated indicators for rod proportions and complexities varied within
|
||
<emphasis id="B911EA9BF97A9D64FC2820918FD5A8B2" box="[988,1022,1195,1220]" italics="true" pageId="40" pageNumber="41">RP</emphasis>
|
||
= 9.6–21.5%;
|
||
<emphasis id="B911EA9BF97A9D64FB58209188FBA8B3" box="[1196,1232,1195,1221]" italics="true" pageId="40" pageNumber="41">RC</emphasis>
|
||
=2.7–17.8 (
|
||
<tableCitation id="C6E70332F97A9D64FAA820918C9FA89F" captionStart="TABLE 11" captionStartId="39.[152,244,152,176]" captionTargetPageId="39" captionText="TABLE 11. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta neglectus sp. nov." pageId="40" pageNumber="41">Table 11</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F6502F97A9D66FF3320C98DA0ADA7" lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41" type="etymology">
|
||
<paragraph id="8BDA3689F97A9D64FF3320C989A0A90F" blockId="40.[151,1437,546,1401]" pageId="40" pageNumber="41">
|
||
<emphasis id="B911EA9BF97A9D64FF3320C98D65A97B" bold="true" box="[199,334,1267,1293]" pageId="40" pageNumber="41">Etymology.</emphasis>
|
||
The species name
|
||
<taxonomicName id="4C654D0AF97A9D64FDB520C98E80A97B" authorityName="Yamana & Hirashima & Sato & Yamamori" authorityYear="2025" box="[577,683,1267,1293]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="40" pageNumber="41" phylum="Echinodermata" rank="species" species="neglectus">
|
||
<emphasis id="B911EA9BF97A9D64FDB520C98E80A97B" box="[577,683,1267,1293]" italics="true" pageId="40" pageNumber="41">neglectus</emphasis>
|
||
</taxonomicName>
|
||
, alluding to Y.Y.’s misidentification of the
|
||
<typeStatus id="54DE882BF97A9D64FB3620C9890FA97B" box="[1218,1316,1267,1293]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
|
||
specimen (WMNH-2022-INV-154). Previously, this specimen was reported as
|
||
<taxonomicName id="4C654D0AF97A9D64FC67212D8FD1A947" box="[915,1018,1303,1329]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="40" pageNumber="41" phylum="Echinodermata" rank="species" species="ooplax">
|
||
<emphasis id="B911EA9BF97A9D64FC67212D8FD1A947" box="[915,1018,1303,1329]" italics="true" pageId="40" pageNumber="41">P. ooplax</emphasis>
|
||
</taxonomicName>
|
||
with young and small ossicles, to Mr. Soichiro Tashima of Kusunoki-Shizenkan (N.P.O. Kusunoki Nature Study Museum), Aira City,
|
||
<collectingRegion id="49A1F86BF97A9D64FB2821018974A923" box="[1244,1375,1339,1365]" country="Japan" name="Kagoshima" pageId="40" pageNumber="41">Kagoshima</collectingRegion>
|
||
. This misjudgment was made due to Y.Y.’s unsympathetic view on ossicle morphologies at that time (
|
||
<date id="FFDB1049F97A9D64FB3221658954A90F" box="[1222,1407,1375,1401]" pageId="40" pageNumber="41" value="2022-12">December 2022</date>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDA3689F97A9D64FF6321F08F36A992" blockId="40.[151,797,1482,1509]" box="[151,797,1482,1509]" pageId="40" pageNumber="41">
|
||
<heading id="D09281E5F97A9D64FF6321F08F36A992" bold="true" box="[151,797,1482,1509]" fontSize="11" level="1" pageId="40" pageNumber="41" reason="1">
|
||
<emphasis id="B911EA9BF97A9D64FF6321F08F36A992" bold="true" box="[151,797,1482,1509]" pageId="40" pageNumber="41">Observation on the morphologies of calcareous ring</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDA3689F97A9D64FF632229884DAA97" blockId="40.[151,1436,1555,1761]" pageId="40" pageNumber="41">
|
||
Calcareous plates 12, mostly firmly bound (
|
||
<figureCitation id="135E2A0CF97A9D64FD8A22298EFBAA5B" box="[638,720,1555,1581]" captionStart="FIGURE 20" captionStartId="41.[151,250,585,609]" captionTargetBox="[163,1422,188,552]" captionTargetId="figure-20@41.[151,1435,181,561]" captionTargetPageId="41" captionText="FIGURE 20. Outer-surface view of calcareous ring of the calcareous ring plates of a specimen collected from Iso Beach, Kagoshima Bay Kagoshima, erected as Patinapta neglectus sp. nov., in the present study (holotype specimen, WMNH-2022- INV-154): Positions of the plates denoted by Ludwig’s [1889–1892 (1889), p156] system: RI–RV and IR1–IR5 where R and IR stands for positions for radius and interradius, respectively. Hence RI, mid-ventral; RII, left ventral; RIII, right ventral; RIV, left dorsal; RV, right dorsal; and: IR1, left ventral; IR2, right ventral; IR3-IR3’, left dorso-lateral; IR4-IR4’, right dorso-lateral; IR5, mid-dorsal. Anterior top." figureDoi="http://doi.org/10.5281/zenodo.14703574" httpUri="https://zenodo.org/record/14703574/files/figure.png" pageId="40" pageNumber="41">Fig. 20</figureCitation>
|
||
), with slight anterior projection and deep posterior depression in interradial plates, while only posterior depression in radial plates. Only medio-ventral radial plate perforated in its anterior half, relatively large perforation. Other four radial plates devoid of perforation, however, there were narrow “interradial perforations” situated in the connections between 4 plates situated in right and left dorso-lateral positions (
|
||
<emphasis id="B911EA9BF97A9D64FF5522998DACAACA" box="[161,391,1699,1725]" italics="true" pageId="40" pageNumber="41">IR4’; RIII; IR3’; RII</emphasis>
|
||
), where additional interradial plate (
|
||
<emphasis id="B911EA9BF97A9D64FCD022998FB2AACB" box="[804,921,1699,1725]" italics="true" pageId="40" pageNumber="41">IR4’; IR3’</emphasis>
|
||
) formed triangular shapes and adjacent radial plate (
|
||
<emphasis id="B911EA9BF97A9D64FF2A22FD8D14AA96" box="[222,319,1735,1760]" italics="true" pageId="40" pageNumber="41">RIII; RII</emphasis>
|
||
) form creviced squarish shapes, and weakly bound by their apical ends.
|
||
</paragraph>
|
||
<caption id="DF1A6601F97B9D65FF6326738DBFAF63" ID-DOI="http://doi.org/10.5281/zenodo.14703574" ID-Zenodo-Dep="14703574" httpUri="https://zenodo.org/record/14703574/files/figure.png" pageId="41" pageNumber="42" startId="41.[151,250,585,609]" targetBox="[163,1422,188,552]" targetPageId="41" targetType="figure">
|
||
<paragraph id="8BDA3689F97B9D65FF6326738DBFAF63" blockId="41.[151,1436,585,789]" pageId="41" pageNumber="42">
|
||
<emphasis id="B911EA9BF97B9D65FF6326738D08AE17" bold="true" box="[151,291,585,609]" pageId="41" pageNumber="42">FIGURE 20.</emphasis>
|
||
Outer-surface view of calcareous ring of the calcareous ring plates of a specimen collected from Iso Beach, Kagoshima Bay Kagoshima, erected as
|
||
<taxonomicName id="4C654D0AF97B9D65FDDB26548ED0AEF3" authorityName="Yamana & Hirashima & Sato & Yamamori" authorityYear="2025" box="[559,763,621,645]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="41" pageNumber="42" phylum="Echinodermata" rank="species" species="neglectus" status="sp. nov.">
|
||
<emphasis id="B911EA9BF97B9D65FDDB26548ED0AEF3" box="[559,763,621,645]" italics="true" pageId="41" pageNumber="42">Patinapta neglectus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911EA9BF97B9D65FCF626578F78AEF3" bold="true" box="[770,851,621,645]" pageId="41" pageNumber="42">
|
||
<taxonomicNameLabel id="A22257E0F97B9D65FCF626578F78AEF3" box="[770,851,621,645]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, in the present study (holotype specimen, WMNH-2022- INV-154): Positions of the plates denoted by Ludwig’s [1889–1892 (1889), p156] system:
|
||
<emphasis id="B911EA9BF97B9D65FBDB26A8885AAEDF" box="[1071,1137,658,681]" italics="true" pageId="41" pageNumber="42">RI–RV</emphasis>
|
||
and
|
||
<emphasis id="B911EA9BF97B9D65FB5726AB88D3AEDF" box="[1187,1272,657,681]" italics="true" pageId="41" pageNumber="42">IR1–IR5</emphasis>
|
||
where
|
||
<emphasis id="B911EA9BF97B9D65FAB726A88979AEDF" box="[1347,1362,658,681]" italics="true" pageId="41" pageNumber="42">R</emphasis>
|
||
and
|
||
<emphasis id="B911EA9BF97B9D65FA7026A889B7AEDF" box="[1412,1436,658,681]" italics="true" pageId="41" pageNumber="42">IR</emphasis>
|
||
stands for positions for radius and interradius, respectively. Hence
|
||
<emphasis id="B911EA9BF97B9D65FCCD268C8F7AAEBB" box="[825,849,694,717]" italics="true" pageId="41" pageNumber="42">RI</emphasis>
|
||
, mid-ventral;
|
||
<emphasis id="B911EA9BF97B9D65FC14268C882BAEBB" box="[992,1024,694,717]" italics="true" pageId="41" pageNumber="42">RII</emphasis>
|
||
, left ventral;
|
||
<emphasis id="B911EA9BF97B9D65FB7C268C889AAEBB" box="[1160,1201,694,717]" italics="true" pageId="41" pageNumber="42">RIII</emphasis>
|
||
, right ventral;
|
||
<emphasis id="B911EA9BF97B9D65FAB3268C8945AEBB" box="[1351,1390,694,717]" italics="true" pageId="41" pageNumber="42">RIV</emphasis>
|
||
, left dorsal;
|
||
<emphasis id="B911EA9BF97B9D65FF1726E08D2AAE87" box="[227,257,730,753]" italics="true" pageId="41" pageNumber="42">RV</emphasis>
|
||
, right dorsal; and:
|
||
<emphasis id="B911EA9BF97B9D65FE3626E38DCDAE87" box="[450,486,729,753]" italics="true" pageId="41" pageNumber="42">IR1</emphasis>
|
||
, left ventral;
|
||
<emphasis id="B911EA9BF97B9D65FD9B26E38EBFAE87" box="[623,660,729,753]" italics="true" pageId="41" pageNumber="42">IR2</emphasis>
|
||
, right ventral;
|
||
<emphasis id="B911EA9BF97B9D65FCDE26E38FAFAE87" box="[810,900,729,753]" italics="true" pageId="41" pageNumber="42">IR3-IR3’</emphasis>
|
||
, left dorso-lateral;
|
||
<emphasis id="B911EA9BF97B9D65FBB326E3888AAE87" box="[1095,1185,729,753]" italics="true" pageId="41" pageNumber="42">IR4-IR4’</emphasis>
|
||
, right dorso-lateral;
|
||
<emphasis id="B911EA9BF97B9D65FA8526E089BEAE87" box="[1393,1429,730,753]" italics="true" pageId="41" pageNumber="42">IR5</emphasis>
|
||
, mid-dorsal. Anterior top.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDA3689F97B9D65FF63275D8E9CAFF7" blockId="41.[151,695,871,898]" box="[151,695,871,898]" pageId="41" pageNumber="42">
|
||
<heading id="D09281E5F97B9D65FF63275D8E9CAFF7" bold="true" box="[151,695,871,898]" fontSize="11" level="1" pageId="41" pageNumber="42" reason="1">
|
||
<emphasis id="B911EA9BF97B9D65FF63275D8E9CAFF7" bold="true" box="[151,695,871,898]" pageId="41" pageNumber="42">
|
||
Ossicle variation among the
|
||
<typeStatus id="54DE882BF97B9D65FE0E27528E1BAFF4" box="[506,560,872,898]" pageId="41" pageNumber="42">type</typeStatus>
|
||
specimens
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDA3689F97B9D65FF63278A8E3AAA7F" blockId="41.[151,1437,943,2014]" pageId="41" pageNumber="42">
|
||
Among
|
||
<specimenCount id="9D63FD00F97B9D65FF0C27958D9FAFBF" box="[248,436,943,969]" count="3" pageId="41" pageNumber="42" type="generic">three specimens</specimenCount>
|
||
,
|
||
<specimenCount id="9D63FD00F97B9D65FE30278A8E5BAFBF" box="[452,624,943,970]" count="2" pageId="41" pageNumber="42" type="generic">two specimens</specimenCount>
|
||
(WMNH-2022-INV-154; 2024-INV-194) were mature and possessed gonad tubules, while
|
||
<specimenCount id="9D63FD00F97B9D65FE7C27EE8E0AAF9B" box="[392,545,979,1006]" count="1" pageId="41" pageNumber="42" type="generic">one specimen</specimenCount>
|
||
(WMNH-2024-INV-193) was immature and was the smallest specimen, which also possessed under-developed ossicles as below: short rod ossicles in tentacles, stout anchor and small anchor plate ossicles in body wall. In tentacles, simple short rod ossicles possess up to only two perforations and only four minute branches, anchor stem short and stout; the single basal tip of anchor plate ossicles with besides noticeable defections, resulting in the “table-tennis racket-form” those of congeners of the apodid genus
|
||
<taxonomicName id="4C654D0AF97B9D65FB56205989B3A8F6" authority="Ostergren" authorityName="Ostergren" authorityYear="1898" box="[1186,1432,1123,1152]" class="Holothuroidea" family="Synaptidae" genus="Labidoplax" kingdom="Animalia" order="Apodida" pageId="41" pageNumber="42" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B911EA9BF97B9D65FB5620598909A80B" box="[1186,1314,1123,1149]" italics="true" pageId="41" pageNumber="42">Labidoplax</emphasis>
|
||
Ōstergren
|
||
</taxonomicName>
|
||
, as such feature was not observed from two mature specimens, which possessed normal ovoid anchor plates. Body wall ossicles anchors and anchor plates. Anchor plate ossicle length and calculated indicators of
|
||
<specimenCount id="9D63FD00F97B9D65FB13209189B6A8B3" box="[1255,1437,1195,1221]" count="3" pageId="41" pageNumber="42" type="generic">three specimens</specimenCount>
|
||
ranged within: 85–123 μm in length,
|
||
<emphasis id="B911EA9BF97B9D65FDA320EA8EACA89C" box="[599,647,1232,1258]" italics="true" pageId="41" pageNumber="42">APS</emphasis>
|
||
=0–11.3,
|
||
<emphasis id="B911EA9BF97B9D65FD0220EA8F35A89F" box="[758,798,1232,1257]" italics="true" pageId="41" pageNumber="42">Ntp</emphasis>
|
||
=0–2,
|
||
<emphasis id="B911EA9BF97B9D65FC9F20EA8FB6A89C" box="[875,925,1232,1258]" italics="true" pageId="41" pageNumber="42">APT</emphasis>
|
||
=0–11.8%. Among these values, length and non—size dependance indicator of plate skewness
|
||
<emphasis id="B911EA9BF97B9D65FD2520CE8F2AA978" box="[721,769,1268,1294]" italics="true" pageId="41" pageNumber="42">APS</emphasis>
|
||
of anchor plate ossicles were not significantly different (
|
||
<emphasis id="B911EA9BF97B9D65FA7420CE89B7A97B" box="[1408,1436,1268,1293]" italics="true" pageId="41" pageNumber="42">Ps</emphasis>
|
||
>0.05, Kruskal-Wallis’ test), while frequency of teethed perforations
|
||
<emphasis id="B911EA9BF97B9D65FC6621228FEFA944" box="[914,964,1304,1330]" italics="true" pageId="41" pageNumber="42">APT</emphasis>
|
||
and their parameter
|
||
<emphasis id="B911EA9BF97B9D65FB52212288E5A947" box="[1190,1230,1304,1329]" italics="true" pageId="41" pageNumber="42">Ntp</emphasis>
|
||
were significantly different on the ventral side (
|
||
<emphasis id="B911EA9BF97B9D65FE2221068DD9A923" box="[470,498,1340,1365]" italics="true" pageId="41" pageNumber="42">Ps</emphasis>
|
||
<0.005, Kruskal-Wallis’ test) but not significantly different on dorsal side (
|
||
<emphasis id="B911EA9BF97B9D65FAC721068964A923" box="[1331,1359,1340,1365]" italics="true" pageId="41" pageNumber="42">Ps</emphasis>
|
||
>0.05, Kruskal-Wallis’ test). From these results, the anchor plate ossicles in the body wall of the present
|
||
<specimenCount id="9D63FD00F97B9D65FB1C216589B6A90F" box="[1256,1437,1375,1401]" count="3" pageId="41" pageNumber="42" type="generic">three specimens</specimenCount>
|
||
were not different in ossicle length or the skewness of the layout for perforations However, number of teethed perforation and their occupancies were different on the ventral body wall. Therefore, the length (about 85–120 μm), numbers of teethed perforations (0–2) on dorsal side, and slightly distorted (up to 11.3) anchor plate ossicles can be appropriate diagnostic characters.
|
||
</paragraph>
|
||
<paragraph id="8BDA3689F97B9D65FF3322298CEEAB73" blockId="41.[151,1437,943,2014]" pageId="41" pageNumber="42">
|
||
Anchor ossicle length and calculated indicators ranged within: 108–188 μm in length,
|
||
<emphasis id="B911EA9BF97B9D65FB3C222E88D5AA58" box="[1224,1278,1556,1582]" italics="true" pageId="41" pageNumber="42">ASW</emphasis>
|
||
=17.3–28.3%,
|
||
<emphasis id="B911EA9BF97B9D65FF6322028CFAAA24" box="[151,209,1592,1618]" italics="true" pageId="41" pageNumber="42">AEW</emphasis>
|
||
=29.6–55.4% (
|
||
<tableCitation id="C6E70332F97B9D65FE83220D8DF7AA24" box="[375,476,1591,1618]" captionStart="TABLE 11" captionStartId="39.[152,244,152,176]" captionTargetPageId="39" captionText="TABLE 11. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta neglectus sp. nov." pageId="41" pageNumber="42">Table 11</tableCitation>
|
||
). Among these values, lengths, the indicator for anchor stem breadth
|
||
<emphasis id="B911EA9BF97B9D65FB0622028903AA24" box="[1266,1320,1592,1618]" italics="true" pageId="41" pageNumber="42">ASW</emphasis>
|
||
of anchor ossicles and for basal / distal ends width
|
||
<emphasis id="B911EA9BF97B9D65FD9F22668E8EAA00" box="[619,677,1628,1654]" italics="true" pageId="41" pageNumber="42">AEW</emphasis>
|
||
were significantly different (
|
||
<emphasis id="B911EA9BF97B9D65FC0522668826AA03" box="[1009,1037,1628,1653]" italics="true" pageId="41" pageNumber="42">Ps</emphasis>
|
||
<0.005, Kruskal-Wallis’ test, both in ventrally and dorsally), while number of the teeth was not significantly different (
|
||
<emphasis id="B911EA9BF97B9D65FBB222BA8849AAEF" box="[1094,1122,1664,1689]" italics="true" pageId="41" pageNumber="42">Ps</emphasis>
|
||
>0.05, Kruskal-Wallis’ test, both ventrally and dorsally). From these results, total shapes of anchor ossicles in the body wall of the present
|
||
<specimenCount id="9D63FD00F97B9D65FA9022998D27AA97" count="3" pageId="41" pageNumber="42" type="generic">three specimens</specimenCount>
|
||
cannot be appropriate keys, but the excessively low number of the counts for teeth can be the accurate key.
|
||
</paragraph>
|
||
<paragraph id="8BDA3689F97B9D66FF3323358859ACA3" blockId="41.[151,1437,943,2014]" lastBlockId="42.[151,1437,151,465]" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
|
||
Longitudinal muscle involved both O-shaped and C-shaped granule ossicles, length and calculated indicators ranged within: 28–54 μm length,
|
||
<emphasis id="B911EA9BF97B9D65FDEA230E8E68AB38" box="[542,579,1844,1870]" italics="true" pageId="41" pageNumber="42">GP</emphasis>
|
||
=28.9–123.0%,
|
||
<emphasis id="B911EA9BF97B9D65FD0E230E8F1EAB38" box="[762,821,1844,1870]" italics="true" pageId="41" pageNumber="42">GCO</emphasis>
|
||
=33.3–66.7% (
|
||
<tableCitation id="C6E70332F97B9D65FC292309886FAB38" box="[989,1092,1843,1870]" captionStart="TABLE 11" captionStartId="39.[152,244,152,176]" captionTargetPageId="39" captionText="TABLE 11. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta neglectus sp. nov." pageId="41" pageNumber="42">Table 11</tableCitation>
|
||
). Among these values, length of granule ossicles and calculated indicator for proportions
|
||
<emphasis id="B911EA9BF97B9D65FCB323628F47AB04" box="[839,876,1880,1906]" italics="true" pageId="41" pageNumber="42">GP</emphasis>
|
||
were different significantly (
|
||
<emphasis id="B911EA9BF97B9D65FB4B236288F0AB07" box="[1215,1243,1880,1905]" italics="true" pageId="41" pageNumber="42">Ps</emphasis>
|
||
<0.005, Kruskal-Wallis’ test, both in ventrally and dorsally), while the ratio C- or O-shaped granule ossicles
|
||
<emphasis id="B911EA9BF97B9D65FB67234688E5ABE0" box="[1171,1230,1916,1942]" italics="true" pageId="41" pageNumber="42">GCO</emphasis>
|
||
were not different significantly (
|
||
<emphasis id="B911EA9BF97B9D65FECE239A8D7DABCF" box="[314,342,1952,1977]" italics="true" pageId="41" pageNumber="42">Ps</emphasis>
|
||
>0.05, Kruskal-Wallis’ test, both ventrally and dorsally), but different significantly in
|
||
<emphasis id="B911EA9BF97B9D65FA95239A89B7ABCC" box="[1377,1436,1952,1978]" italics="true" pageId="41" pageNumber="42">GCO</emphasis>
|
||
(
|
||
<emphasis id="B911EA9BF97B9D65FF5523FE8C96ABAB" box="[161,189,1988,2013]" italics="true" pageId="41" pageNumber="42">Ps</emphasis>
|
||
<0.05, χ
|
||
<superScript id="7C109BC1F97B9D65FEE823F98D0FABA7" attach="right" box="[284,292,1987,2001]" fontSize="6" pageId="41" pageNumber="42">2</superScript>
|
||
-multiple test, both in ventrally and dorsally). Therefore, the length and proportion of the granule ossicles cannot be appropriate keys, while ratio C- or O-shaped granule and the feature of granule ossicles being “flattened, distorted and roundish triangle form” of this species can be appropriate keys.
|
||
</paragraph>
|
||
<paragraph id="8BDA3689F9789D66FF3324E58DA0ADA7" blockId="42.[151,1437,151,465]" pageId="42" pageNumber="43">
|
||
Tentacle ossicles rods, length and calculated indicators of
|
||
<specimenCount id="9D63FD00F9789D66FCA024E58823AC8F" box="[852,1032,223,249]" count="3" pageId="42" pageNumber="43" type="generic">three specimens</specimenCount>
|
||
ranged within: 44–98 μm in length,
|
||
<emphasis id="B911EA9BF9789D66FF6325398C92AD6A" box="[151,185,259,284]" italics="true" pageId="42" pageNumber="43">RP</emphasis>
|
||
=9.6–21.5%, and
|
||
<emphasis id="B911EA9BF9789D66FE7025398D83AD6B" box="[388,424,259,285]" italics="true" pageId="42" pageNumber="43">RC</emphasis>
|
||
=2.7–17.8. Among these values, length of rod ossicles, nonsize dependence complexities (
|
||
<emphasis id="B911EA9BF9789D66FF55251D8CEEAD37" box="[161,197,295,321]" italics="true" pageId="42" pageNumber="43">RC</emphasis>
|
||
) were significantly different (
|
||
<emphasis id="B911EA9BF9789D66FDF8251D8E03AD36" box="[524,552,295,320]" italics="true" pageId="42" pageNumber="43">Ps</emphasis>
|
||
<0.005, Kruskal-Wallis’ test), while nonsize dependence indicator for proportion (
|
||
<emphasis id="B911EA9BF9789D66FF5525718CE8AD12" box="[161,195,331,356]" italics="true" pageId="42" pageNumber="43">RP</emphasis>
|
||
) was not different significantly (
|
||
<emphasis id="B911EA9BF9789D66FDB325718E48AD12" box="[583,611,331,356]" italics="true" pageId="42" pageNumber="43">Ps</emphasis>
|
||
>0.05, Kruskal-Wallis’ test). From these results, the rod ossicles in the tentacles of the present
|
||
<specimenCount id="9D63FD00F9789D66FE5425558E7FADFF" box="[416,596,367,393]" count="3" pageId="42" pageNumber="43" type="generic">three specimens</specimenCount>
|
||
had similar proportions, simultaneously complexities and length increased by body growth (
|
||
<tableCitation id="C6E70332F9789D66FEA825A98DEAADDB" box="[348,449,403,429]" captionStart="TABLE 11" captionStartId="39.[152,244,152,176]" captionTargetPageId="39" captionText="TABLE 11. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta neglectus sp. nov." pageId="42" pageNumber="43">Table 11</tableCitation>
|
||
). Therefore, the morphologies of the rod ossicles of this species cannot be appropriate diagnostic characters.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F6502F9789D66FF6326198DD6AFBF" pageId="42" pageNumber="43" type="discussion">
|
||
<paragraph id="8BDA3689F9789D66FF6326198D22AE4B" blockId="42.[151,265,547,573]" box="[151,265,547,573]" pageId="42" pageNumber="43">
|
||
<emphasis id="B911EA9BF9789D66FF6326198D22AE4B" bold="true" box="[151,265,547,573]" pageId="42" pageNumber="43">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDA3689F9789D66FF6326518DD6AFBF" blockId="42.[151,1437,619,969]" pageId="42" pageNumber="43">
|
||
In the present
|
||
<specimenCount id="9D63FD00F9789D66FECE26518D81AEF3" box="[314,426,619,645]" count="3" pageId="42" pageNumber="43" type="generic">three type</specimenCount>
|
||
materials, ossicle morphologies were noticeably different between matured
|
||
<typeStatus id="54DE882BF9789D66FAFE26518947AEF3" box="[1290,1388,619,645]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
|
||
and two small
|
||
<typeStatus id="54DE882BF9789D66FEF826B58D51AEDF" box="[268,378,655,681]" pageId="42" pageNumber="43" type="paratype">paratypes</typeStatus>
|
||
,
|
||
<specimenCount id="9D63FD00F9789D66FE7126B58E34AEDF" box="[389,543,655,681]" count="1" pageId="42" pageNumber="43" type="immature">one immature</specimenCount>
|
||
and one in the development-stage of tubules (
|
||
<tableCitation id="C6E70332F9789D66FBD726B5885DAEDF" box="[1059,1142,655,681]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionTargetPageId="6" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="42" pageNumber="43">Table 1</tableCitation>
|
||
). From the present results of measurements and counts on ossicles (
|
||
<tableCitation id="C6E70332F9789D66FD9C26898EE7AEBB" box="[616,716,691,717]" captionStart="TABLE 11" captionStartId="39.[152,244,152,176]" captionTargetPageId="39" captionText="TABLE 11. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta neglectus sp. nov." pageId="42" pageNumber="43">Table 11</tableCitation>
|
||
), it can be assumed that several morphological changes relating with individual growth: i) in anchor ossicles, minute teeth on outer edge of anchor-bills take numbers larger in small animals than in large animals; ii) in anchor plate ossicles, teethed perforationstake larger number in small animals than in large animals; however, iii) in tentacle rods, complexity of rod-shape (
|
||
<emphasis id="B911EA9BF9789D66FB8C272588B7AF4F" box="[1144,1180,799,825]" italics="true" pageId="42" pageNumber="43">RC</emphasis>
|
||
) take values higher in large animals than in small animals, and complexity may not be increasing or decreasing by individual growth. In the previous nominal species, none of the species possessed calcareous ring only perforated on medio-ventral plate (Table 7-3), and this must be largest difference of the present new species among all the congeners of genus
|
||
<taxonomicName id="4C654D0AF9789D66FF6327958D2EAFBE" authorityName="Heding" authorityYear="1928" box="[151,261,943,968]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="42" pageNumber="43" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B911EA9BF9789D66FF6327958D2EAFBE" box="[151,261,943,968]" italics="true" pageId="42" pageNumber="43">Patinapta</emphasis>
|
||
</taxonomicName>
|
||
previously described.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F6502F9789D66FF6320208E96A923" pageId="42" pageNumber="43" type="distribution">
|
||
<paragraph id="8BDA3689F9789D66FF6320208D1BA842" blockId="42.[151,304,1050,1076]" box="[151,304,1050,1076]" pageId="42" pageNumber="43">
|
||
<emphasis id="B911EA9BF9789D66FF6320208D1BA842" bold="true" box="[151,304,1050,1076]" pageId="42" pageNumber="43">Distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDA3689F9789D66FF6320598E96A923" blockId="42.[151,1437,1123,1365]" pageId="42" pageNumber="43">
|
||
Only known from Kyushu Island and adjacent islands, Mature specimen has been collected only from Iso Beach (the type locality) on the inner coast of
|
||
<collectingRegion id="49A1F86BF9789D66FDA220BD8EFDA8D7" box="[598,726,1159,1185]" country="Japan" name="Kagoshima" pageId="42" pageNumber="43">Kagoshima</collectingRegion>
|
||
Bay (
|
||
<tableCitation id="C6E70332F9789D66FCEE20BD8FABA8D7" box="[794,896,1159,1185]" captionStart="TABLE 12" captionStartId="42.[152,244,1412,1436]" captionTargetPageId="42" captionText="TABLE 12. List of WMNH-INV specimens of Patinapta neglectus sp. nov., as of 2024." pageId="42" pageNumber="43">Table 12</tableCitation>
|
||
). At the type locality, the
|
||
<typeStatus id="54DE882BF9789D66FB5020BD892DA8D7" box="[1188,1286,1159,1185]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
|
||
material was collected from intertidal sandy beach with boulders. Among the specimens housed in WMNH-INV, including the present specimens, 3 bottles including
|
||
<specimenCount id="9D63FD00F9789D66FDBB20F48EF1A89F" box="[591,730,1230,1257]" count="5" pageId="42" pageNumber="43" type="generic">5 specimens</specimenCount>
|
||
were identified as this species (
|
||
<tableCitation id="C6E70332F9789D66FBB120F58887A89F" box="[1093,1196,1231,1257]" captionStart="TABLE 12" captionStartId="42.[152,244,1412,1436]" captionTargetPageId="42" captionText="TABLE 12. List of WMNH-INV specimens of Patinapta neglectus sp. nov., as of 2024." pageId="42" pageNumber="43">Table 12</tableCitation>
|
||
). Other than the type locality,
|
||
<specimenCount id="9D63FD00F9789D66FF0E20C98D56A97B" box="[250,381,1267,1293]" count="2" pageId="42" pageNumber="43" type="immature">2 immature</specimenCount>
|
||
and mature specimens were collected from Nagaura-higata flat, Amakusa City,
|
||
<collectingRegion id="49A1F86BF9789D66FAEA20C989B7A97B" box="[1310,1436,1267,1293]" country="Japan" name="Kumamoto" pageId="42" pageNumber="43">Kumamoto</collectingRegion>
|
||
and
|
||
<specimenCount id="9D63FD00F9789D66FF33212D8D6CA947" box="[199,327,1303,1329]" count="2" pageId="42" pageNumber="43" type="immature">2 immature</specimenCount>
|
||
specimens collected from Uchiura Bay, Kamikoshiki-jima Island, adjacent to
|
||
<collectingRegion id="49A1F86BF9789D66FB43212D8912A947" box="[1207,1337,1303,1329]" country="Japan" name="Kagoshima" pageId="42" pageNumber="43">Kagoshima</collectingRegion>
|
||
, Kyushu Island,
|
||
<collectingCountry id="F3727619F9789D66FF1E21018D06A923" box="[234,301,1339,1365]" name="Japan" pageId="42" pageNumber="43">Japan</collectingCountry>
|
||
, and were identified as this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |