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<document id="4F11640BAEDDC9E0781B654BB0FEF81A" ID-DOI="10.11646/zootaxa.5569.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14703528" ID-ZooBank="96194340-054D-49C0-BD89-D47899BCAF14" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1737371250141" checkinUser="plazi" docAuthor="Yamana, Yusuke, Hirashima, Kentarou, Sato, Masanori &amp; Yamamori, Luna" docDate="2025" docId="03CC879FF94D9D68FF6326248822AA22" docLanguage="en" docName="zootaxa.5569.1.1.pdf" docOrigin="Zootaxa 5569 (1)" docSource="https://doi.org/10.11646/zootaxa.5569.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Patinapta parvaspiculus Yamana, Hirashima, Sato &amp; Yamamori, 2025, sp. nov." docType="treatment" docVersion="1" lastPageNumber="37" masterDocId="FFF5FFE7F9529D4CFFF4243A8C2BAC76" masterDocTitle="First revision of the apodid holothurian genus Patinapta Heding, 1928 (Synaptidae) in Japanese waters, with establishment of four new species from Kyushu Island and Ryukyu Islands, southwestern Japan" masterLastPageNumber="54" masterPageNumber="1" pageNumber="32" updateTime="1737473723657" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:title id="4984AE6F946B45EA5E0D182AAD8C9C6A">First revision of the apodid holothurian genus Patinapta Heding, 1928 (Synaptidae) in Japanese waters, with establishment of four new species from Kyushu Island and Ryukyu Islands, southwestern Japan</mods:title>
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<mods:namePart id="7AB593B408D4989DA112377668EDFB32">Yamana, Yusuke</mods:namePart>
<mods:affiliation id="07F16CE9D1A01D0E455C4BB97521403B">Wakayama Prefectural Museum of Natural History (WMNH), Invertebrate Specimen House, Funo 370 - 1, Kainan, Wakayama 642 - 0001, Japan</mods:affiliation>
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<mods:namePart id="04FEF8947739B9A9ACDB15D076AB8639">Hirashima, Kentarou</mods:namePart>
<mods:affiliation id="1013E4E5B7998043DCBC01F86E1F8DAC">Wakayama Prefectural Museum of Natural History (WMNH), 1 Room Curators Head Office, Funo 370 - 1, Kainan, Wakayama 642 - 0001, Japan</mods:affiliation>
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<mods:namePart id="026401540AB0136EE2F4D0460B284CE8">Sato, Masanori</mods:namePart>
<mods:affiliation id="12476C83794C68AE2C9FB932EF6E5F2E">Graduate School of Science and Engineering, Kagoshima University, Kagoshima 890 - 0065, Japan &amp; Present address: 5233 Hiramatsu, Aira, Kagoshima 899 - 5652, Japan</mods:affiliation>
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<mods:namePart id="7CA77E6A750DDDF58CFF3C16AEC37A34">Yamamori, Luna</mods:namePart>
<mods:affiliation id="58048663879B26F4B19127144144753C">Seto Marine biological laboratory (SMBL) of Kyoto University, 459, Shirahama, Wakayama 649 - 2211, Japan</mods:affiliation>
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<mods:date id="274DD8A33C6C37A2967F118FE2C8BD3F">2025</mods:date>
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<mods:number id="CCE02DA7F044814BDC740040A57ACA24">5569</mods:number>
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<treatment id="03CC879FF94D9D68FF6326248822AA22" ID-DOI="http://doi.org/10.5281/zenodo.14704974" ID-Zenodo-Dep="14704974" LSID="urn:lsid:plazi:treatment:03CC879FF94D9D68FF6326248822AA22" httpUri="http://treatment.plazi.org/id/03CC879FF94D9D68FF6326248822AA22" lastPageId="36" lastPageNumber="37" pageId="31" pageNumber="32">
<subSubSection id="C37F6502F94D9D53FF6326248E0AAE4E" box="[151,545,541,568]" pageId="31" pageNumber="32" type="nomenclature">
<paragraph id="8BDA3689F94D9D53FF6326248E0AAE4E" blockId="31.[151,732,541,640]" box="[151,545,541,568]" pageId="31" pageNumber="32">
<heading id="D09281E5F94D9D53FF6326248E0AAE4E" bold="true" box="[151,545,541,568]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
<emphasis id="B911EA9BF94D9D53FF6326248E0AAE4E" bold="true" box="[151,545,541,568]" pageId="31" pageNumber="32">
<taxonomicName id="4C654D0AF94D9D53FF6326248D97AE4E" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[151,444,541,568]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="species" species="parvaspiculus" status="sp. nov.">
<emphasis id="B911EA9BF94D9D53FF6326248D97AE4E" bold="true" box="[151,444,541,568]" italics="true" pageId="31" pageNumber="32">Patinapta parvaspiculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257E0F94D9D53FE3726248E0AAE4E" box="[451,545,542,568]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F94D9D53FF6326788EF7AE2A" box="[151,732,578,604]" pageId="31" pageNumber="32" type="vernacular_names">
<paragraph id="8BDA3689F94D9D53FF6326788EF7AE2A" blockId="31.[151,732,541,640]" box="[151,732,578,604]" pageId="31" pageNumber="32">[New Japanese name: Kokotsu-himo-ikari-namako]</paragraph>
</subSubSection>
<subSubSection id="C37F6502F94D9D53FF63265C8DF1AEF6" box="[151,474,613,640]" pageId="31" pageNumber="32" type="description">
<paragraph id="8BDA3689F94D9D53FF63265C8DF1AEF6" blockId="31.[151,732,541,640]" box="[151,474,613,640]" pageId="31" pageNumber="32">
(
<figureCitation id="135E2A0CF94D9D53FF6B265C8D0AAEF6" box="[159,289,613,640]" captionStart-0="FIGURE 15" captionStart-1="FIGURE 16" captionStart-2="FIGURE 17" captionStartId-0="33.[151,250,1258,1282]" captionStartId-1="34.[151,250,1674,1698]" captionStartId-2="35.[151,250,628,652]" captionTargetBox-0="[151,1436,181,1234]" captionTargetBox-1="[158,1430,181,1643]" captionTargetBox-2="[164,1422,281,600]" captionTargetId-0="figure-20@33.[151,1436,178,1237]" captionTargetId-1="figure-29@34.[158,1430,181,1650]" captionTargetId-2="figure-65@35.[151,1435,273,605]" captionTargetPageId-0="33" captionTargetPageId-1="34" captionTargetPageId-2="35" captionText-0="FIGURE 15. External view of a damaged specimen collected from Yoron-jima Island, Kagoshima, erected as Patinapta parvaspiculus sp. nov., in the present study (holotype specimen, WMNH-2023-INV-157): A, lateral view, anterior left, dorsal top with damage on anterior body wall (from sampling shovel; gonad tubules also damaged); B, oral-side view of the tentacles; C, outer-side view of the tentacles." captionText-1="FIGURE 16. SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, erected as Patinapta parvaspiculus sp. nov., in the present study (holotype specimen, WMNH-2023-INV-157): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." captionText-2="FIGURE 17. Outer-surface view of calcareous ring plates of a specimen collected from Yoron-jima Island, Kagoshima, erected as Patinapta parvaspiculus sp. nov., in the present study (holotype specimen, WMNH-2023-INV-157). The positions of the plates are denoted by Ludwigs [18891892 (1889), p156] system: RIRV and IR1IR5 where R and IR stands for positions for radius and interradius, respectively. Hence RI, mid-ventral; RII, left ventral; RIII, right ventral; RIV, left dorsal; RV, right dorsal; and: IR1, left ventral; IR2, right ventral; IR3-IR3, left dorso-lateral; IR4-IR4, right dorso-lateral; IR5, mid-dorsal. Anterior top." figureDoi-0="http://doi.org/10.5281/zenodo.14703564" figureDoi-1="http://doi.org/10.5281/zenodo.14703566" figureDoi-2="http://doi.org/10.5281/zenodo.14703568" httpUri-0="https://zenodo.org/record/14703564/files/figure.png" httpUri-1="https://zenodo.org/record/14703566/files/figure.png" httpUri-2="https://zenodo.org/record/14703568/files/figure.png" pageId="31" pageNumber="32">Figs 1517</figureCitation>
;
<tableCitation id="C6E70332F94D9D53FED8265C8DA1AEF6" box="[300,394,614,640]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionTargetPageId="6" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="31" pageNumber="32">Tables 1</tableCitation>
,
<tableCitation id="C6E70332F94D9D53FE6C265C8D83AEF6" box="[408,424,614,640]" captionStart="TABLE 9" captionStartId="32.[152,243,152,176]" captionTargetPageId="32" captionText="TABLE 9. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta parvaspiculus sp. nov." pageId="31" pageNumber="32">9</tableCitation>
,
<tableCitation id="C6E70332F94D9D53FE40265C8DFAAEF6" box="[436,465,614,640]" captionStart="TABLE 10" captionStartId="36.[152,244,872,896]" captionTargetPageId="36" captionText="TABLE 10. List of WMNH-INV specimens of Patinapta parvaspiculus sp. nov. as of 2024." pageId="31" pageNumber="32">10</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F94D9D53FF63269189A9AE97" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="8BDA3689F94D9D53FF63269189A9AE97" blockId="31.[151,1436,682,737]" pageId="31" pageNumber="32">
<materialsCitation id="3B0D3CD4F94D9D53FF6326918FD6AEB4" box="[151,1021,682,706]" location="Yoron-jima Island" pageId="31" pageNumber="32" specimenCount="2">
<specimenCount id="9D63FD00F94D9D53FF6326918D1FAEB4" box="[151,308,682,706]" count="2" pageId="31" pageNumber="32" type="generic">Two specimens</specimenCount>
from
<location id="8EBA6052F94D9D53FE8526908E02AEB4" LSID="urn:lsid:plazi:treatment:03CC879FF94D9D68FF6326248822AA22:8EBA6052F94D9D53FE8526908E02AEB4" box="[369,553,682,706]" name="Yoron-jima Island" pageId="31" pageNumber="32">Yoron-jima Island</location>
(WMNH-2015-INV-107; 2023-INV-157) and
</materialsCitation>
<materialsCitation id="3B0D3CD4F94D9D53FBF726918956AE97" location="Imari Bay" municipality="We" pageId="31" pageNumber="32" specimenCount="1">
<specimenCount id="9D63FD00F94D9D53FBF7269188A5AEB4" box="[1027,1166,682,706]" count="1" pageId="31" pageNumber="32" type="generic">one specimen</specimenCount>
from
<location id="8EBA6052F94D9D53FB3826908919AEB5" LSID="urn:lsid:plazi:treatment:03CC879FF94D9D68FF6326248822AA22:8EBA6052F94D9D53FB3826908919AEB5" box="[1228,1330,682,707]" municipality="We" name="Imari Bay" pageId="31" pageNumber="32">Imari Bay</location>
(WMNH-2024-INV-156) were identified as new to science.
<collectingMunicipality id="6BBEACF3F94D9D53FD3226F38ECCAE96" box="[710,743,713,736]" pageId="31" pageNumber="32">We</collectingMunicipality>
herein describe the morphological characters of this new species
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F94D9D53FF6327358EBFAFAB" pageId="31" pageNumber="32" type="diagnosis">
<paragraph id="8BDA3689F94D9D53FF6327358EBFAFAB" blockId="31.[151,1437,783,1602]" pageId="31" pageNumber="32">
<emphasis id="B911EA9BF94D9D53FF6327358D39AF5F" bold="true" box="[151,274,783,809]" pageId="31" pageNumber="32">Diagnosis.</emphasis>
Anchor and anchor plate ossicles small. Anchor plate ossicles smaller than 100 μm, while anchor ossicles up to 150 μm, resulting in unbalanced size combinations of bounded ossicles. Total shapes of anchor plates strongly distorted (because of small total numbers of perforations), basally with a concave shallow or deep, resulting in a two-tipped ovoid shape. Perforations of anchor plate ossicles mostly without teeth, while outer edge of anchor plate ossicles equipped with minute teeth in small numbers. Ciliated funnels attached to inner side body wall along midline of left dorso-lateral interradius (
<emphasis id="B911EA9BF94D9D53FDAF27F98EA8AFAB" box="[603,643,963,989]" italics="true" pageId="31" pageNumber="32">IR3</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F94D9D53FF3327DD8CE3A850" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="8BDA3689F94D9D53FF3327DD8CE3A850" blockId="31.[151,1437,783,1602]" pageId="31" pageNumber="32">
<emphasis id="B911EA9BF94D9D53FF3327DD8D7DA877" bold="true" box="[199,342,999,1025]" pageId="31" pageNumber="32">Type series.</emphasis>
<typeStatus id="54DE882BF94D9D53FE9427DD8DE0A877" box="[352,459,999,1025]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
, WMNH-2023-INV-157;
<specimenCount id="9D63FD00F94D9D53FD0727D28FBEA874" box="[755,917,1000,1026]" count="2" pageId="31" pageNumber="32" type="generic" typeStatus="paratype">two paratypes</specimenCount>
, WMNH-2015-INV-107; WMNH-2024-INV-156.
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F94D9D53FF3320158E22AA6B" pageId="31" pageNumber="32" type="description">
<paragraph id="8BDA3689F94D9D53FF3320158F22A8C3" blockId="31.[151,1437,783,1602]" pageId="31" pageNumber="32">
<emphasis id="B911EA9BF94D9D53FF332015888AA83F" bold="true" box="[199,1185,1071,1097]" pageId="31" pageNumber="32">
Description of
<typeStatus id="54DE882BF94D9D53FE8920158DCFA83F" box="[381,484,1071,1097]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
(WMNH-2023-INV-157: middle-size damaged specimen).
</emphasis>
External and internal morphologies as follows (
<tableCitation id="C6E70332F94D9D53FE3620698E31A81B" box="[450,538,1107,1133]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionTargetPageId="6" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="31" pageNumber="32">Table 1</tableCitation>
): 30 year-old preserved well-matured specimen pale peanut color, body large about
<quantity id="4C9D9B6CF94D9D53FF2A204D8D16A8E4" box="[222,317,1143,1170]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="31" pageNumber="32" unit="mm" value="100.0">100 mm</quantity>
length (
<figureCitation id="135E2A0CF94D9D53FE68204D8DDAA8E4" box="[412,497,1143,1170]" captionStart="FIGURE 15" captionStartId="33.[151,250,1258,1282]" captionTargetBox="[151,1436,181,1234]" captionTargetId="figure-20@33.[151,1436,178,1237]" captionTargetPageId="33" captionText="FIGURE 15. External view of a damaged specimen collected from Yoron-jima Island, Kagoshima, erected as Patinapta parvaspiculus sp. nov., in the present study (holotype specimen, WMNH-2023-INV-157): A, lateral view, anterior left, dorsal top with damage on anterior body wall (from sampling shovel; gonad tubules also damaged); B, oral-side view of the tentacles; C, outer-side view of the tentacles." figureDoi="http://doi.org/10.5281/zenodo.14703564" httpUri="https://zenodo.org/record/14703564/files/figure.png" pageId="31" pageNumber="32">Fig. 15</figureCitation>
). Tentacles 12, each with 10 digits of 5 pairs. On oral-side of tentacle stem, about 1016 sensory cups sporadically arranged in a U-shape.
</paragraph>
<paragraph id="8BDA3689F94D9D53FF3320858DFBA933" blockId="31.[151,1437,783,1602]" pageId="31" pageNumber="32">
Polian vesicle in medio-ventral position, fusiform, triple (
<tableCitation id="C6E70332F94D9D53FCB920858F8AA8AF" box="[845,929,1215,1241]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionTargetPageId="6" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="31" pageNumber="32">Table 1</tableCitation>
), stone canal single and thread-like, attached to narrow space between basal parts of tentacle ampullae at mediodorsal (
<emphasis id="B911EA9BF94D9D53FC3420D98FC2A88B" box="[960,1001,1251,1277]" italics="true" pageId="31" pageNumber="32">IR5</emphasis>
), suddenly turned upward and forward, resulted in it facing forward, distally with moldy surface madreporite. Ciliated funnels in one crowded band along left dorsal interradius (
<emphasis id="B911EA9BF94D9D53FE6321118D94A933" box="[407,447,1323,1349]" italics="true" pageId="31" pageNumber="32">IR3</emphasis>
).
</paragraph>
<paragraph id="8BDA3689F94D9D53FF332175889CA9FB" blockId="31.[151,1437,783,1602]" pageId="31" pageNumber="32">Inside body, two tufts of gonad tubules attached to both sides of anterior dorsal mesentery, also two whitish tubule organs adhered to upper and under sides of intestine canal. Intestine canal lacking loop.</paragraph>
<paragraph id="8BDA3689F94D9D53FF3321AD8E22AA6B" blockId="31.[151,1437,783,1602]" pageId="31" pageNumber="32">Body wall smooth surface, hard but thin, with ossicle arranging latitudinal layout of all as sets of anchor and plate. Both sides along five longitudinal muscles with distal ends of anchor ossicles facing towards interradial area, however ossicles on radial body walls upon longitudinal muscles and interradial body walls have distal ends randomly facing either direction.</paragraph>
</subSubSection>
<subSubSection id="C37F6502F94D9D68FF33221D8816AD37" lastPageId="36" lastPageNumber="37" pageId="31" pageNumber="32" type="etymology">
<paragraph id="8BDA3689F94D9D53FF33221D8879AA37" blockId="31.[151,1437,783,1602]" box="[199,1106,1575,1602]" pageId="31" pageNumber="32">
<emphasis id="B911EA9BF94D9D53FF33221D8D65AA37" bold="true" box="[199,334,1575,1601]" pageId="31" pageNumber="32">Etymology.</emphasis>
The species name
<taxonomicName id="4C654D0AF94D9D53FDD6221D8E94AA37" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[546,703,1575,1601]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="species" species="parvaspiculus">
<emphasis id="B911EA9BF94D9D53FDD6221D8E94AA37" box="[546,703,1575,1601]" italics="true" pageId="31" pageNumber="32">parvaspiculus</emphasis>
</taxonomicName>
, alluding to the ossicles small size.
</paragraph>
<paragraph id="8BDA3689F94D9D53FF6322A98E86AADB" blockId="31.[151,685,1683,1709]" box="[151,685,1683,1709]" pageId="31" pageNumber="32">
<heading id="D09281E5F94D9D53FF6322A98E86AADB" bold="true" box="[151,685,1683,1709]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
<emphasis id="B911EA9BF94D9D53FF6322A98E86AADB" bold="true" box="[151,685,1683,1709]" pageId="31" pageNumber="32">The present results of ossicle morphologies</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA3689F94D9D53FF6322E18EA0AB87" blockId="31.[151,1437,1755,2034]" pageId="31" pageNumber="32">
Body wall ossicles anchors and anchor plates (
<figureCitation id="135E2A0CF94D9D53FD4622E18F14AA80" box="[690,831,1755,1782]" captionStart="FIGURE 16" captionStartId="34.[151,250,1674,1698]" captionTargetBox="[158,1430,181,1643]" captionTargetId="figure-29@34.[158,1430,181,1650]" captionTargetPageId="34" captionText="FIGURE 16. SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, erected as Patinapta parvaspiculus sp. nov., in the present study (holotype specimen, WMNH-2023-INV-157): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703566" httpUri="https://zenodo.org/record/14703566/files/figure.png" pageId="31" pageNumber="32">Fig. 16B, D</figureCitation>
;
<tableCitation id="C6E70332F94D9D53FCB922E18F8FAA83" box="[845,932,1755,1781]" captionStart="TABLE 9" captionStartId="32.[152,243,152,176]" captionTargetPageId="32" captionText="TABLE 9. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta parvaspiculus sp. nov." pageId="31" pageNumber="32">Table 9</tableCitation>
). Anchor plates ranged within 7598 μm in length. Size of perforations not noticeably variable between central area and marginal area, and instead almost constant between central area and marginal area, because of only few marginal perforations rarely equipped. Plates showed slight distortion, with small total numbers of perforations arranged more or less in two rows as holothuroid button ossicles, and basally with a concave shallow or deep, resulting in a two-tipped ovoid shape. Perforations of anchor plate ossicles mostly toothless, while outer edge of all anchor plate ossicles equipped with small numbers of sharp minute teeth. Two calculated indicators for anchor plates and a parameter were variated within
<emphasis id="B911EA9BF94D9D53FB012389890EABBB" box="[1269,1317,1971,1997]" italics="true" pageId="31" pageNumber="32">APS</emphasis>
=00.7;
<emphasis id="B911EA9BF94D9D53FA76238E89B7ABBB" box="[1410,1436,1972,1997]" italics="true" pageId="31" pageNumber="32">Nt</emphasis>
=03;
<emphasis id="B911EA9BF94D9D53FF1423ED8D39AB87" box="[224,274,2007,2033]" italics="true" pageId="31" pageNumber="32">APT</emphasis>
=027.3% (
<figureCitation id="135E2A0CF94D9D53FE6123ED8E36AB84" box="[405,541,2007,2034]" captionStart="FIGURE 16" captionStartId="34.[151,250,1674,1698]" captionTargetBox="[158,1430,181,1643]" captionTargetId="figure-29@34.[158,1430,181,1650]" captionTargetPageId="34" captionText="FIGURE 16. SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, erected as Patinapta parvaspiculus sp. nov., in the present study (holotype specimen, WMNH-2023-INV-157): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703566" httpUri="https://zenodo.org/record/14703566/files/figure.png" pageId="31" pageNumber="32">Fig. 16B, D</figureCitation>
;
<tableCitation id="C6E70332F94D9D53FDDC23ED8E56AB87" box="[552,637,2007,2033]" captionStart="TABLE 9" captionStartId="32.[152,243,152,176]" captionTargetPageId="32" captionText="TABLE 9. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta parvaspiculus sp. nov." pageId="31" pageNumber="32">Table 9</tableCitation>
).
</paragraph>
<caption id="DF1A6601F9729D6CFF6C24A289BEACC7" box="[152,1429,152,177]" pageId="32" pageNumber="33" startId="32.[152,243,152,176]" targetBox="[163,1414,190,1688]" targetIsTable="true" targetPageId="32" targetType="table">
<paragraph id="8BDA3689F9729D6CFF6C24A289BEACC7" blockId="32.[152,1429,152,177]" box="[152,1429,152,177]" pageId="32" pageNumber="33">
<emphasis id="B911EA9BF9729D6CFF6C24A28D26ACC6" bold="true" box="[152,269,152,176]" pageId="32" pageNumber="33">TABLE 9.</emphasis>
Measurements (mean±SD μm) and counts of ossicles of three specimens of
<taxonomicName id="4C654D0AF9729D6CFBC224A38912ACC6" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[1078,1337,152,176]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="32" pageNumber="33" phylum="Echinodermata" rank="species" species="parvaspiculus" status="sp. nov.">
<emphasis id="B911EA9BF9729D6CFBC224A38912ACC6" box="[1078,1337,152,176]" italics="true" pageId="32" pageNumber="33">Patinapta parvaspiculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257E0F9729D6CFAB424A389BEACC7" box="[1344,1429,153,177]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BDA3689F9729D6CFF57248489ADAAEE" pageId="32" pageNumber="33">
<table id="F965C429F97262B3FF57248489ADAAEE" box="[163,1414,190,1688]" gridcols="8" gridrows="42" pageId="32" pageNumber="33">
<tr id="355534CBF97262B3FF57248489ADACA0" box="[163,1414,190,214]" gridrow="0" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF5724848D32ACA0" box="[163,281,190,214]" gridcol="0" gridrow="0" pageId="32" pageNumber="33">Body parts</th>
<th id="76845DB7F97262B3FEB724848DDFACA0" box="[323,500,190,214]" gridcol="1" gridrow="0" pageId="32" pageNumber="33">(Ossicle-type)</th>
<th id="76845DB7F97262B3FDE124848F0CACA0" box="[533,807,190,214]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="32" pageNumber="33">2015-INV-107</th>
<th id="76845DB7F97262B3FCA624848876ACA0" box="[850,1117,190,214]" colspan="2" colspanRight="1" gridcol="4" gridrow="0" pageId="32" pageNumber="33">2023-INV-157</th>
<th id="76845DB7F97262B3FB75248489ADACA0" box="[1153,1414,190,214]" colspan="2" colspanRight="1" gridcol="6" gridrow="0" pageId="32" pageNumber="33">2024-INV-156</th>
</tr>
<tr id="355534CBF97262B3FF5724D889ADAC8C" box="[163,1414,226,250]" gridrow="1" pageId="32" pageNumber="33" rowspan-3="1" rowspan-5="1" rowspan-7="1">
<th id="76845DB7F97262B3FF5724D88D32AC8C" box="[163,281,226,250]" gridcol="0" gridrow="1" pageId="32" pageNumber="33">Tentacle</th>
<td id="76845DB7F97262B3FEB724D88DDFAC8C" box="[323,500,226,250]" gridcol="1" gridrow="1" pageId="32" pageNumber="33">
Rods (
<emphasis id="B911EA9BF9729D6CFE4624D98D94AC8C" box="[434,447,227,250]" italics="true" pageId="32" pageNumber="33">n</emphasis>
)
</td>
<td id="76845DB7F97262B3FDE124D88EA3AC8C" box="[533,648,226,250]" gridcol="2" gridrow="1" pageId="32" pageNumber="33">9</td>
<td id="76845DB7F97262B3FCA624D88FEEAC8C" box="[850,965,226,250]" gridcol="4" gridrow="1" pageId="32" pageNumber="33">15</td>
<td id="76845DB7F97262B3FB7524D888C3AC8C" box="[1153,1256,226,250]" gridcol="6" gridrow="1" pageId="32" pageNumber="33">10</td>
</tr>
<tr id="355534CBF97262B3FF57253C89ADAD68" box="[163,1414,262,286]" gridrow="2" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF57253C8D32AD68" box="[163,281,262,286]" gridcol="0" gridrow="2" pageId="32" pageNumber="33">Length</th>
<td id="76845DB7F97262B3FEB7253C8DDFAD68" box="[323,500,262,286]" gridcol="1" gridrow="2" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE1253C8EA3AD68" box="[533,648,262,286]" gridcol="2" gridrow="2" pageId="32" pageNumber="33">62.0±5.6</td>
<td id="76845DB7F97262B3FD45253C8F0CAD68" box="[689,807,262,286]" gridcol="3" gridrow="2" pageId="32" pageNumber="33">(5574)</td>
<td id="76845DB7F97262B3FCA6253C8FEEAD68" box="[850,965,262,286]" gridcol="4" gridrow="2" pageId="32" pageNumber="33">75.0±1.4</td>
<td id="76845DB7F97262B3FC13253C8876AD68" box="[999,1117,262,286]" gridcol="5" gridrow="2" pageId="32" pageNumber="33">(53102)</td>
<td id="76845DB7F97262B3FB75253C88C3AD68" box="[1153,1256,262,286]" gridcol="6" gridrow="2" pageId="32" pageNumber="33">56.1±10.5</td>
<td id="76845DB7F97262B3FAE4253C89ADAD68" box="[1296,1414,262,286]" gridcol="7" gridrow="2" pageId="32" pageNumber="33">(4579)</td>
</tr>
<tr id="355534CBF97262B3FF57251389ADAD34" box="[163,1414,297,322]" gridrow="3" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF5725138D32AD34" box="[163,281,297,322]" gridcol="0" gridrow="3" pageId="32" pageNumber="33">Breadth</th>
<td id="76845DB7F97262B3FEB725138DDFAD34" box="[323,500,297,322]" gridcol="1" gridrow="3" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE125138EA3AD34" box="[533,648,297,322]" gridcol="2" gridrow="3" pageId="32" pageNumber="33">8.1±0.9</td>
<td id="76845DB7F97262B3FD4525138F0CAD34" box="[689,807,297,322]" gridcol="3" gridrow="3" pageId="32" pageNumber="33">(710)</td>
<td id="76845DB7F97262B3FCA625138FEEAD34" box="[850,965,297,322]" gridcol="4" gridrow="3" pageId="32" pageNumber="33">7.8±1.4</td>
<td id="76845DB7F97262B3FC1325138876AD34" box="[999,1117,297,322]" gridcol="5" gridrow="3" pageId="32" pageNumber="33">(611)</td>
<td id="76845DB7F97262B3FB75251388C3AD34" box="[1153,1256,297,322]" gridcol="6" gridrow="3" pageId="32" pageNumber="33">7.8±2.3</td>
<td id="76845DB7F97262B3FAE4251389ADAD34" box="[1296,1414,297,322]" gridcol="7" gridrow="3" pageId="32" pageNumber="33">(512)</td>
</tr>
<tr id="355534CBF97262B3FF57257689ADAD13" box="[163,1414,332,357]" gridrow="4" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF5725768D32AD13" box="[163,281,332,357]" gridcol="0" gridrow="4" pageId="32" pageNumber="33">
<emphasis id="B911EA9BF9729D6CFF4425748CE4AD13" box="[176,207,334,357]" italics="true" pageId="32" pageNumber="33">RP</emphasis>
a (%)
</th>
<td id="76845DB7F97262B3FEB725768DDFAD13" box="[323,500,332,357]" gridcol="1" gridrow="4" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE125768EA3AD13" box="[533,648,332,357]" gridcol="2" gridrow="4" pageId="32" pageNumber="33">13.2±2.1</td>
<td id="76845DB7F97262B3FD4525768F0CAD13" box="[689,807,332,357]" gridcol="3" gridrow="4" pageId="32" pageNumber="33">(10.816.9)</td>
<td id="76845DB7F97262B3FCA625768FEEAD13" box="[850,965,332,357]" gridcol="4" gridrow="4" pageId="32" pageNumber="33">10.7±2.5</td>
<td id="76845DB7F97262B3FC1325768876AD13" box="[999,1117,332,357]" gridcol="5" gridrow="4" pageId="32" pageNumber="33">(7.116.9)</td>
<td id="76845DB7F97262B3FB75257688C3AD13" box="[1153,1256,332,357]" gridcol="6" gridrow="4" pageId="32" pageNumber="33">14.1±4.1</td>
<td id="76845DB7F97262B3FAE4257689ADAD13" box="[1296,1414,332,357]" gridcol="7" gridrow="4" pageId="32" pageNumber="33">(8.220.7)</td>
</tr>
<tr id="355534CBF97262B3FF57254B89ADADFF" box="[163,1414,369,393]" gridrow="5" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF57254B8D32ADFF" box="[163,281,369,393]" gridcol="0" gridrow="5" pageId="32" pageNumber="33">
<emphasis id="B911EA9BF9729D6CFF44254B8CFBADFE" box="[176,208,369,392]" italics="true" pageId="32" pageNumber="33">RC</emphasis>
b
</th>
<td id="76845DB7F97262B3FEB7254B8DDFADFF" box="[323,500,369,393]" gridcol="1" gridrow="5" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE1254B8EA3ADFF" box="[533,648,369,393]" gridcol="2" gridrow="5" pageId="32" pageNumber="33">8.0±1.7</td>
<td id="76845DB7F97262B3FD45254B8F0CADFF" box="[689,807,369,393]" gridcol="3" gridrow="5" pageId="32" pageNumber="33">(5.310.2)</td>
<td id="76845DB7F97262B3FCA6254B8FEEADFF" box="[850,965,369,393]" gridcol="4" gridrow="5" pageId="32" pageNumber="33">6.4±2.3</td>
<td id="76845DB7F97262B3FC13254B8876ADFF" box="[999,1117,369,393]" gridcol="5" gridrow="5" pageId="32" pageNumber="33">(2.711.7)</td>
<td id="76845DB7F97262B3FB75254B88C3ADFF" box="[1153,1256,369,393]" gridcol="6" gridrow="5" pageId="32" pageNumber="33">13.4±6.2</td>
<td id="76845DB7F97262B3FAE4254B89ADADFF" box="[1296,1414,369,393]" gridcol="7" gridrow="5" pageId="32" pageNumber="33">(6.625.9)</td>
</tr>
<tr id="355534CBF97262B3FF5725AF89ADADDB" box="[163,1414,405,429]" gridrow="6" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF5725AF89ADADDB" box="[163,1414,405,429]" colspan="8" colspanRight="7" gridcol="0" gridrow="6" pageId="32" pageNumber="33">Ventral side</th>
</tr>
<tr id="355534CBF97262B3FF57258289ADADA7" box="[163,1414,440,465]" gridrow="7" pageId="32" pageNumber="33" rowspan-3="1" rowspan-5="1" rowspan-7="1">
<th id="76845DB7F97262B3FF5725828D32ADA7" box="[163,281,440,465]" gridcol="0" gridrow="7" pageId="32" pageNumber="33">L-Muscle</th>
<td id="76845DB7F97262B3FEB725828DDFADA7" box="[323,500,440,465]" gridcol="1" gridrow="7" pageId="32" pageNumber="33">
Granules (
<emphasis id="B911EA9BF9729D6CFE3225838DF8ADA6" box="[454,467,441,464]" italics="true" pageId="32" pageNumber="33">n</emphasis>
)
</td>
<td id="76845DB7F97262B3FDE125828EA3ADA7" box="[533,648,440,465]" gridcol="2" gridrow="7" pageId="32" pageNumber="33">12</td>
<td id="76845DB7F97262B3FCA625828FEEADA7" box="[850,965,440,465]" gridcol="4" gridrow="7" pageId="32" pageNumber="33">11</td>
<td id="76845DB7F97262B3FB75258288C3ADA7" box="[1153,1256,440,465]" gridcol="6" gridrow="7" pageId="32" pageNumber="33">12</td>
</tr>
<tr id="355534CBF97262B3FF5725E689ADAD82" box="[163,1414,476,500]" gridrow="8" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF5725E68D32AD82" box="[163,281,476,500]" gridcol="0" gridrow="8" pageId="32" pageNumber="33">Length</th>
<td id="76845DB7F97262B3FEB725E68DDFAD82" box="[323,500,476,500]" gridcol="1" gridrow="8" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE125E68EA3AD82" box="[533,648,476,500]" gridcol="2" gridrow="8" pageId="32" pageNumber="33">28.4±4.3</td>
<td id="76845DB7F97262B3FD4525E68F0CAD82" box="[689,807,476,500]" gridcol="3" gridrow="8" pageId="32" pageNumber="33">(2336)</td>
<td id="76845DB7F97262B3FCA625E68FEEAD82" box="[850,965,476,500]" gridcol="4" gridrow="8" pageId="32" pageNumber="33">27.8±2.3</td>
<td id="76845DB7F97262B3FC1325E68876AD82" box="[999,1117,476,500]" gridcol="5" gridrow="8" pageId="32" pageNumber="33">(2432)</td>
<td id="76845DB7F97262B3FB7525E688C3AD82" box="[1153,1256,476,500]" gridcol="6" gridrow="8" pageId="32" pageNumber="33">29.6±4.2</td>
<td id="76845DB7F97262B3FAE425E689ADAD82" box="[1296,1414,476,500]" gridcol="7" gridrow="8" pageId="32" pageNumber="33">(2134)</td>
</tr>
<tr id="355534CBF97262B3FF57263A89ADAE6E" box="[163,1414,512,536]" gridrow="9" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF57263A8D32AE6E" box="[163,281,512,536]" gridcol="0" gridrow="9" pageId="32" pageNumber="33">Width</th>
<td id="76845DB7F97262B3FEB7263A8DDFAE6E" box="[323,500,512,536]" gridcol="1" gridrow="9" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE1263A8EA3AE6E" box="[533,648,512,536]" gridcol="2" gridrow="9" pageId="32" pageNumber="33">16.0± 2.1</td>
<td id="76845DB7F97262B3FD45263A8F0CAE6E" box="[689,807,512,536]" gridcol="3" gridrow="9" pageId="32" pageNumber="33">(1021)</td>
<td id="76845DB7F97262B3FCA6263A8FEEAE6E" box="[850,965,512,536]" gridcol="4" gridrow="9" pageId="32" pageNumber="33">12.7±2.1</td>
<td id="76845DB7F97262B3FC13263A8876AE6E" box="[999,1117,512,536]" gridcol="5" gridrow="9" pageId="32" pageNumber="33">(1018)</td>
<td id="76845DB7F97262B3FB75263A88C3AE6E" box="[1153,1256,512,536]" gridcol="6" gridrow="9" pageId="32" pageNumber="33">16.5±5.9</td>
<td id="76845DB7F97262B3FAE4263A89ADAE6E" box="[1296,1414,512,536]" gridcol="7" gridrow="9" pageId="32" pageNumber="33">(1233)</td>
</tr>
<tr id="355534CBF97262B3FF57261989ADAE4A" box="[163,1414,547,572]" gridrow="10" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF5726198D32AE4A" box="[163,281,547,572]" gridcol="0" gridrow="10" pageId="32" pageNumber="33">
<emphasis id="B911EA9BF9729D6CFF44261E8CF9AE4D" box="[176,210,548,571]" italics="true" pageId="32" pageNumber="33">GP</emphasis>
c
</th>
<td id="76845DB7F97262B3FEB726198DDFAE4A" box="[323,500,547,572]" gridcol="1" gridrow="10" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE126198EA3AE4A" box="[533,648,547,572]" gridcol="2" gridrow="10" pageId="32" pageNumber="33">50.7±15.5</td>
<td id="76845DB7F97262B3FD4526198F0CAE4A" box="[689,807,547,572]" gridcol="3" gridrow="10" pageId="32" pageNumber="33">(30.680.8)</td>
<td id="76845DB7F97262B3FCA626198FEEAE4A" box="[850,965,547,572]" gridcol="4" gridrow="10" pageId="32" pageNumber="33">45.9±7.8</td>
<td id="76845DB7F97262B3FC1326198876AE4A" box="[999,1117,547,572]" gridcol="5" gridrow="10" pageId="32" pageNumber="33">(35.564.3)</td>
<td id="76845DB7F97262B3FB75261988C3AE4A" box="[1153,1256,547,572]" gridcol="6" gridrow="10" pageId="32" pageNumber="33">33.3±0.5</td>
<td id="76845DB7F97262B3FAE4261989ADAE4A" box="[1296,1414,547,572]" gridcol="7" gridrow="10" pageId="32" pageNumber="33">(35.3106)</td>
</tr>
<tr id="355534CBF97262B3FF57267D89ADAE29" box="[163,1414,583,607]" gridrow="11" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF57267D8D32AE29" box="[163,281,583,607]" gridcol="0" gridrow="11" pageId="32" pageNumber="33">
<emphasis id="B911EA9BF9729D6CFF4426728CCEAE29" box="[176,229,584,607]" italics="true" pageId="32" pageNumber="33">GCO</emphasis>
d (%)
</th>
<td id="76845DB7F97262B3FEB7267D8DDFAE29" box="[323,500,583,607]" gridcol="1" gridrow="11" pageId="32" pageNumber="33">(Ratio)</td>
<td id="76845DB7F97262B3FDE1267D8EA3AE29" box="[533,648,583,607]" gridcol="2" gridrow="11" pageId="32" pageNumber="33">50.0</td>
<td id="76845DB7F97262B3FD45267D8F0CAE29" box="[689,807,583,607]" gridcol="3" gridrow="11" pageId="32" pageNumber="33">(O6: C6)</td>
<td id="76845DB7F97262B3FCA6267D8FEEAE29" box="[850,965,583,607]" gridcol="4" gridrow="11" pageId="32" pageNumber="33">36.4</td>
<td id="76845DB7F97262B3FC13267D8876AE29" box="[999,1117,583,607]" gridcol="5" gridrow="11" pageId="32" pageNumber="33">(O4: C7)</td>
<td id="76845DB7F97262B3FB75267D88C3AE29" box="[1153,1256,583,607]" gridcol="6" gridrow="11" pageId="32" pageNumber="33">56.7</td>
<td id="76845DB7F97262B3FAE4267D89ADAE29" box="[1296,1414,583,607]" gridcol="7" gridrow="11" pageId="32" pageNumber="33">(O4: C8)</td>
</tr>
<tr id="355534CBF97262B3FF57265189ADAEF5" box="[163,1414,619,643]" gridrow="12" pageId="32" pageNumber="33" rowspan-3="1" rowspan-5="1" rowspan-7="1">
<th id="76845DB7F97262B3FF5726518D32AEF5" box="[163,281,619,643]" gridcol="0" gridrow="12" pageId="32" pageNumber="33">Body wall</th>
<td id="76845DB7F97262B3FEB726518DDFAEF5" box="[323,500,619,643]" gridcol="1" gridrow="12" pageId="32" pageNumber="33">
Anchor plates (
<emphasis id="B911EA9BF9729D6CFE2B26568DC7AEF5" box="[479,492,620,643]" italics="true" pageId="32" pageNumber="33">n</emphasis>
)
</td>
<td id="76845DB7F97262B3FDE126518EA3AEF5" box="[533,648,619,643]" gridcol="2" gridrow="12" pageId="32" pageNumber="33">5</td>
<td id="76845DB7F97262B3FCA626518FEEAEF5" box="[850,965,619,643]" gridcol="4" gridrow="12" pageId="32" pageNumber="33">7</td>
<td id="76845DB7F97262B3FB75265188C3AEF5" box="[1153,1256,619,643]" gridcol="6" gridrow="12" pageId="32" pageNumber="33">9</td>
</tr>
<tr id="355534CBF97262B3FF5726B589ADAED1" box="[163,1414,655,679]" gridrow="13" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF5726B58D32AED1" box="[163,281,655,679]" gridcol="0" gridrow="13" pageId="32" pageNumber="33">Length</th>
<td id="76845DB7F97262B3FEB726B58DDFAED1" box="[323,500,655,679]" gridcol="1" gridrow="13" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE126B58EA3AED1" box="[533,648,655,679]" gridcol="2" gridrow="13" pageId="32" pageNumber="33">87.6±7.6</td>
<td id="76845DB7F97262B3FD4526B58F0CAED1" box="[689,807,655,679]" gridcol="3" gridrow="13" pageId="32" pageNumber="33">(7594)</td>
<td id="76845DB7F97262B3FCA626B58FEEAED1" box="[850,965,655,679]" gridcol="4" gridrow="13" pageId="32" pageNumber="33">90.9±5.6</td>
<td id="76845DB7F97262B3FC1326B58876AED1" box="[999,1117,655,679]" gridcol="5" gridrow="13" pageId="32" pageNumber="33">(8097)</td>
<td id="76845DB7F97262B3FB7526B588C3AED1" box="[1153,1256,655,679]" gridcol="6" gridrow="13" pageId="32" pageNumber="33">93.0±3.5</td>
<td id="76845DB7F97262B3FAE426B589ADAED1" box="[1296,1414,655,679]" gridcol="7" gridrow="13" pageId="32" pageNumber="33">(8697)</td>
</tr>
<tr id="355534CBF97262B3FF57268889ADAEBD" box="[163,1414,690,715]" gridrow="14" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF5726888D32AEBD" box="[163,281,690,715]" gridcol="0" gridrow="14" pageId="32" pageNumber="33">Width</th>
<td id="76845DB7F97262B3FEB726888DDFAEBD" box="[323,500,690,715]" gridcol="1" gridrow="14" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE126888EA3AEBD" box="[533,648,690,715]" gridcol="2" gridrow="14" pageId="32" pageNumber="33">49.0±5.8</td>
<td id="76845DB7F97262B3FD4526888F0CAEBD" box="[689,807,690,715]" gridcol="3" gridrow="14" pageId="32" pageNumber="33">(3954)</td>
<td id="76845DB7F97262B3FCA626888FEEAEBD" box="[850,965,690,715]" gridcol="4" gridrow="14" pageId="32" pageNumber="33">57.3±4.9</td>
<td id="76845DB7F97262B3FC1326888876AEBD" box="[999,1117,690,715]" gridcol="5" gridrow="14" pageId="32" pageNumber="33">(5364)</td>
<td id="76845DB7F97262B3FB75268888C3AEBD" box="[1153,1256,690,715]" gridcol="6" gridrow="14" pageId="32" pageNumber="33">62.3±8.6</td>
<td id="76845DB7F97262B3FAE4268889ADAEBD" box="[1296,1414,690,715]" gridcol="7" gridrow="14" pageId="32" pageNumber="33">(4979)</td>
</tr>
<tr id="355534CBF97262B3FF5726EF89ADAE98" box="[163,1414,725,750]" gridrow="15" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF5726EF8D32AE98" box="[163,281,725,750]" gridcol="0" gridrow="15" pageId="32" pageNumber="33">
<emphasis id="B911EA9BF9729D6CFF4426ED8CF7AE98" box="[176,220,727,750]" italics="true" pageId="32" pageNumber="33">APS</emphasis>
e
</th>
<td id="76845DB7F97262B3FEB726EF8DDFAE98" box="[323,500,725,750]" gridcol="1" gridrow="15" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE126EF8EA3AE98" box="[533,648,725,750]" gridcol="2" gridrow="15" pageId="32" pageNumber="33">1.2±0.9</td>
<td id="76845DB7F97262B3FD4526EF8F0CAE98" box="[689,807,725,750]" gridcol="3" gridrow="15" pageId="32" pageNumber="33">(0.62.2)</td>
<td id="76845DB7F97262B3FCA626EF8FEEAE98" box="[850,965,725,750]" gridcol="4" gridrow="15" pageId="32" pageNumber="33">0.2±0.3</td>
<td id="76845DB7F97262B3FC1326EF8876AE98" box="[999,1117,725,750]" gridcol="5" gridrow="15" pageId="32" pageNumber="33">(00.6)</td>
<td id="76845DB7F97262B3FB7526EF88C3AE98" box="[1153,1256,725,750]" gridcol="6" gridrow="15" pageId="32" pageNumber="33">3.2±2.2</td>
<td id="76845DB7F97262B3FAE426EF89ADAE98" box="[1296,1414,725,750]" gridcol="7" gridrow="15" pageId="32" pageNumber="33">(2.39.11)</td>
</tr>
<tr id="355534CBF97262B3FF5726C089ADAF64" box="[163,1414,762,786]" gridrow="16" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF5726C08D32AF64" box="[163,281,762,786]" gridcol="0" gridrow="16" pageId="32" pageNumber="33">
<emphasis id="B911EA9BF9729D6CFF4426C18CFEAF64" box="[176,213,763,786]" italics="true" pageId="32" pageNumber="33">Ntp</emphasis>
f
</th>
<td id="76845DB7F97262B3FEB726C08DDFAF64" box="[323,500,762,786]" gridcol="1" gridrow="16" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE126C08EA3AF64" box="[533,648,762,786]" gridcol="2" gridrow="16" pageId="32" pageNumber="33">1.6±0.5</td>
<td id="76845DB7F97262B3FD4526C08F0CAF64" box="[689,807,762,786]" gridcol="3" gridrow="16" pageId="32" pageNumber="33">(12)</td>
<td id="76845DB7F97262B3FCA626C08FEEAF64" box="[850,965,762,786]" gridcol="4" gridrow="16" pageId="32" pageNumber="33">0.4±0.8</td>
<td id="76845DB7F97262B3FC1326C08876AF64" box="[999,1117,762,786]" gridcol="5" gridrow="16" pageId="32" pageNumber="33">(02)</td>
<td id="76845DB7F97262B3FB7526C088C3AF64" box="[1153,1256,762,786]" gridcol="6" gridrow="16" pageId="32" pageNumber="33">1.3±0.7</td>
<td id="76845DB7F97262B3FAE426C089ADAF64" box="[1296,1414,762,786]" gridcol="7" gridrow="16" pageId="32" pageNumber="33">(02)</td>
</tr>
<tr id="355534CBF97262B3FF57272789ADAF40" box="[163,1414,797,822]" gridrow="17" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF5727278D32AF40" box="[163,281,797,822]" gridcol="0" gridrow="17" pageId="32" pageNumber="33">
<emphasis id="B911EA9BF9729D6CFF4427248CF6AF43" box="[176,221,798,821]" italics="true" pageId="32" pageNumber="33">APT</emphasis>
g (%)
</th>
<td id="76845DB7F97262B3FEB727278DDFAF40" box="[323,500,797,822]" gridcol="1" gridrow="17" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE127278EA3AF40" box="[533,648,797,822]" gridcol="2" gridrow="17" pageId="32" pageNumber="33">11.9±4.3</td>
<td id="76845DB7F97262B3FD4527278F0CAF40" box="[689,807,797,822]" gridcol="3" gridrow="17" pageId="32" pageNumber="33">(6.716.7)</td>
<td id="76845DB7F97262B3FCA627278FEEAF40" box="[850,965,797,822]" gridcol="4" gridrow="17" pageId="32" pageNumber="33">3.0±5.2</td>
<td id="76845DB7F97262B3FC1327278876AF40" box="[999,1117,797,822]" gridcol="5" gridrow="17" pageId="32" pageNumber="33">(012.5)</td>
<td id="76845DB7F97262B3FB75272788C3AF40" box="[1153,1256,797,822]" gridcol="6" gridrow="17" pageId="32" pageNumber="33">8.2±5.2</td>
<td id="76845DB7F97262B3FAE4272789ADAF40" box="[1296,1414,797,822]" gridcol="7" gridrow="17" pageId="32" pageNumber="33">(016.7)</td>
</tr>
<tr id="355534CBF97262B3FF57277B89ADAF2C" box="[163,1414,833,858]" gridrow="18" pageId="32" pageNumber="33" rowspan-3="1" rowspan-5="1" rowspan-7="1">
<th id="76845DB7F97262B3FF57277B8D32AF2C" box="[163,281,833,858]" gridcol="0" gridrow="18" pageId="32" pageNumber="33">Body wall</th>
<td id="76845DB7F97262B3FEB7277B8DDFAF2C" box="[323,500,833,858]" gridcol="1" gridrow="18" pageId="32" pageNumber="33">
Anchor (
<emphasis id="B911EA9BF9729D6CFE4A27788DE0AF2F" box="[446,459,834,857]" italics="true" pageId="32" pageNumber="33">n</emphasis>
)
</td>
<td id="76845DB7F97262B3FDE1277B8EA3AF2C" box="[533,648,833,858]" gridcol="2" gridrow="18" pageId="32" pageNumber="33">8</td>
<td id="76845DB7F97262B3FCA6277B8FEEAF2C" box="[850,965,833,858]" gridcol="4" gridrow="18" pageId="32" pageNumber="33">8</td>
<td id="76845DB7F97262B3FB75277B88C3AF2C" box="[1153,1256,833,858]" gridcol="6" gridrow="18" pageId="32" pageNumber="33">12</td>
</tr>
<tr id="355534CBF97262B3FF57275F89ADAF0B" box="[163,1414,869,893]" gridrow="19" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF57275F8D32AF0B" box="[163,281,869,893]" gridcol="0" gridrow="19" pageId="32" pageNumber="33">Length</th>
<td id="76845DB7F97262B3FEB7275F8DDFAF0B" box="[323,500,869,893]" gridcol="1" gridrow="19" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE1275F8EA3AF0B" box="[533,648,869,893]" gridcol="2" gridrow="19" pageId="32" pageNumber="33">165.4±5.6</td>
<td id="76845DB7F97262B3FD45275F8F0CAF0B" box="[689,807,869,893]" gridcol="3" gridrow="19" pageId="32" pageNumber="33">(160174)</td>
<td id="76845DB7F97262B3FCA6275F8FEEAF0B" box="[850,965,869,893]" gridcol="4" gridrow="19" pageId="32" pageNumber="33">160.4±5.9</td>
<td id="76845DB7F97262B3FC13275F8876AF0B" box="[999,1117,869,893]" gridcol="5" gridrow="19" pageId="32" pageNumber="33">(153173)</td>
<td id="76845DB7F97262B3FB75275F88C3AF0B" box="[1153,1256,869,893]" gridcol="6" gridrow="19" pageId="32" pageNumber="33">155.4±6.0</td>
<td id="76845DB7F97262B3FAE4275F89ADAF0B" box="[1296,1414,869,893]" gridcol="7" gridrow="19" pageId="32" pageNumber="33">(143164)</td>
</tr>
<tr id="355534CBF97262B3FF5727B389ADAFD7" box="[163,1414,905,929]" gridrow="20" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF5727B38D32AFD7" box="[163,281,905,929]" gridcol="0" gridrow="20" pageId="32" pageNumber="33">Width</th>
<td id="76845DB7F97262B3FEB727B38DDFAFD7" box="[323,500,905,929]" gridcol="1" gridrow="20" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE127B38EA3AFD7" box="[533,648,905,929]" gridcol="2" gridrow="20" pageId="32" pageNumber="33">64.8± 4.5</td>
<td id="76845DB7F97262B3FD4527B38F0CAFD7" box="[689,807,905,929]" gridcol="3" gridrow="20" pageId="32" pageNumber="33">(5771)</td>
<td id="76845DB7F97262B3FCA627B38FEEAFD7" box="[850,965,905,929]" gridcol="4" gridrow="20" pageId="32" pageNumber="33">68.0±3.2</td>
<td id="76845DB7F97262B3FC1327B38876AFD7" box="[999,1117,905,929]" gridcol="5" gridrow="20" pageId="32" pageNumber="33">(6171)</td>
<td id="76845DB7F97262B3FB7527B388C3AFD7" box="[1153,1256,905,929]" gridcol="6" gridrow="20" pageId="32" pageNumber="33">76.9±4.3</td>
<td id="76845DB7F97262B3FAE427B389ADAFD7" box="[1296,1414,905,929]" gridcol="7" gridrow="20" pageId="32" pageNumber="33">(6984)</td>
</tr>
<tr id="355534CBF97262B3FF57279689ADAFB3" box="[163,1414,940,965]" gridrow="21" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF5727968D32AFB3" box="[163,281,940,965]" gridcol="0" gridrow="21" pageId="32" pageNumber="33">Teeth</th>
<td id="76845DB7F97262B3FEB727968DDFAFB3" box="[323,500,940,965]" gridcol="1" gridrow="21" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE127968EA3AFB3" box="[533,648,940,965]" gridcol="2" gridrow="21" pageId="32" pageNumber="33">6.5±0.9</td>
<td id="76845DB7F97262B3FD4527968F0CAFB3" box="[689,807,940,965]" gridcol="3" gridrow="21" pageId="32" pageNumber="33">(58)</td>
<td id="76845DB7F97262B3FCA627968FEEAFB3" box="[850,965,940,965]" gridcol="4" gridrow="21" pageId="32" pageNumber="33">4.0±1.3</td>
<td id="76845DB7F97262B3FC1327968876AFB3" box="[999,1117,940,965]" gridcol="5" gridrow="21" pageId="32" pageNumber="33">(15)</td>
<td id="76845DB7F97262B3FB75279688C3AFB3" box="[1153,1256,940,965]" gridcol="6" gridrow="21" pageId="32" pageNumber="33">9.3±1.5</td>
<td id="76845DB7F97262B3FAE4279689ADAFB3" box="[1296,1414,940,965]" gridcol="7" gridrow="21" pageId="32" pageNumber="33">(712)</td>
</tr>
<tr id="355534CBF97262B3FF5727EA89ADAF9E" box="[163,1414,976,1000]" gridrow="22" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF5727EA8D32AF9E" box="[163,281,976,1000]" gridcol="0" gridrow="22" pageId="32" pageNumber="33">
<emphasis id="B911EA9BF9729D6CFF4427EB8CCAAF9E" box="[176,225,977,1000]" italics="true" pageId="32" pageNumber="33">ASW</emphasis>
h(%)
</th>
<td id="76845DB7F97262B3FEB727EA8DDFAF9E" box="[323,500,976,1000]" gridcol="1" gridrow="22" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE127EA8EA3AF9E" box="[533,648,976,1000]" gridcol="2" gridrow="22" pageId="32" pageNumber="33">27.8±2.0</td>
<td id="76845DB7F97262B3FD4527EA8F0CAF9E" box="[689,807,976,1000]" gridcol="3" gridrow="22" pageId="32" pageNumber="33">(25.832.5)</td>
<td id="76845DB7F97262B3FCA627EA8FEEAF9E" box="[850,965,976,1000]" gridcol="4" gridrow="22" pageId="32" pageNumber="33">24.7±2.7</td>
<td id="76845DB7F97262B3FC1327EA8876AF9E" box="[999,1117,976,1000]" gridcol="5" gridrow="22" pageId="32" pageNumber="33">(21.829.5)</td>
<td id="76845DB7F97262B3FB7527EA88C3AF9E" box="[1153,1256,976,1000]" gridcol="6" gridrow="22" pageId="32" pageNumber="33">20.3±0.9</td>
<td id="76845DB7F97262B3FAE427EA89ADAF9E" box="[1296,1414,976,1000]" gridcol="7" gridrow="22" pageId="32" pageNumber="33">(19.121.6)</td>
</tr>
<tr id="355534CBF97262B3FF5727CE89ADA87A" box="[163,1414,1012,1036]" gridrow="23" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF5727CE8D32A87A" box="[163,281,1012,1036]" gridcol="0" gridrow="23" pageId="32" pageNumber="33">
<emphasis id="B911EA9BF9729D6CFF4427CE8CCFA87D" box="[176,228,1012,1035]" italics="true" pageId="32" pageNumber="33">AEW</emphasis>
i (%)
</th>
<td id="76845DB7F97262B3FEB727CE8DDFA87A" box="[323,500,1012,1036]" gridcol="1" gridrow="23" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE127CE8EA3A87A" box="[533,648,1012,1036]" gridcol="2" gridrow="23" pageId="32" pageNumber="33">55.1±5.2</td>
<td id="76845DB7F97262B3FD4527CE8F0CA87A" box="[689,807,1012,1036]" gridcol="3" gridrow="23" pageId="32" pageNumber="33">(50.063.2)</td>
<td id="76845DB7F97262B3FCA627CE8FEEA87A" box="[850,965,1012,1036]" gridcol="4" gridrow="23" pageId="32" pageNumber="33">51.8±5.0</td>
<td id="76845DB7F97262B3FC1327CE8876A87A" box="[999,1117,1012,1036]" gridcol="5" gridrow="23" pageId="32" pageNumber="33">(45.160.7)</td>
<td id="76845DB7F97262B3FB7527CE88C3A87A" box="[1153,1256,1012,1036]" gridcol="6" gridrow="23" pageId="32" pageNumber="33">48.9±2.0</td>
<td id="76845DB7F97262B3FAE427CE89ADA87A" box="[1296,1414,1012,1036]" gridcol="7" gridrow="23" pageId="32" pageNumber="33">(45.653.1)</td>
</tr>
<tr id="355534CBF97262B3FF57202289ADA846" box="[163,1414,1048,1072]" gridrow="24" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF57202289ADA846" box="[163,1414,1048,1072]" colspan="8" colspanRight="7" gridcol="0" gridrow="24" pageId="32" pageNumber="33">Dorsal side</th>
</tr>
<tr id="355534CBF97262B3FF57200189ADA822" box="[163,1414,1083,1108]" gridrow="25" pageId="32" pageNumber="33" rowspan-3="1" rowspan-5="1" rowspan-7="1">
<th id="76845DB7F97262B3FF5720018D32A822" box="[163,281,1083,1108]" gridcol="0" gridrow="25" pageId="32" pageNumber="33">L-Muscle</th>
<td id="76845DB7F97262B3FEB720018DDFA822" box="[323,500,1083,1108]" gridcol="1" gridrow="25" pageId="32" pageNumber="33">
Granules (
<emphasis id="B911EA9BF9729D6CFE3220068DF8A825" box="[454,467,1084,1107]" italics="true" pageId="32" pageNumber="33">n</emphasis>
)
</td>
<td id="76845DB7F97262B3FDE120018EA3A822" box="[533,648,1083,1108]" gridcol="2" gridrow="25" pageId="32" pageNumber="33">12</td>
<td id="76845DB7F97262B3FCA620018FEEA822" box="[850,965,1083,1108]" gridcol="4" gridrow="25" pageId="32" pageNumber="33">14</td>
<td id="76845DB7F97262B3FB75200188C3A822" box="[1153,1256,1083,1108]" gridcol="6" gridrow="25" pageId="32" pageNumber="33">11</td>
</tr>
<tr id="355534CBF97262B3FF57206589ADA801" box="[163,1414,1119,1143]" gridrow="26" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF5720658D32A801" box="[163,281,1119,1143]" gridcol="0" gridrow="26" pageId="32" pageNumber="33">Length</th>
<td id="76845DB7F97262B3FEB720658DDFA801" box="[323,500,1119,1143]" gridcol="1" gridrow="26" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE120658EA3A801" box="[533,648,1119,1143]" gridcol="2" gridrow="26" pageId="32" pageNumber="33">32.5±4.7</td>
<td id="76845DB7F97262B3FD4520658F0CA801" box="[689,807,1119,1143]" gridcol="3" gridrow="26" pageId="32" pageNumber="33">(2842)</td>
<td id="76845DB7F97262B3FCA620658FEEA801" box="[850,965,1119,1143]" gridcol="4" gridrow="26" pageId="32" pageNumber="33">28.7±3.9</td>
<td id="76845DB7F97262B3FC1320658876A801" box="[999,1117,1119,1143]" gridcol="5" gridrow="26" pageId="32" pageNumber="33">(2134)</td>
<td id="76845DB7F97262B3FB75206588C3A801" box="[1153,1256,1119,1143]" gridcol="6" gridrow="26" pageId="32" pageNumber="33">27.9±6.3</td>
<td id="76845DB7F97262B3FAE4206589ADA801" box="[1296,1414,1119,1143]" gridcol="7" gridrow="26" pageId="32" pageNumber="33">(2139)</td>
</tr>
<tr id="355534CBF97262B3FF5720B989ADA8ED" box="[163,1414,1155,1179]" gridrow="27" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF5720B98D32A8ED" box="[163,281,1155,1179]" gridcol="0" gridrow="27" pageId="32" pageNumber="33">Width</th>
<td id="76845DB7F97262B3FEB720B98DDFA8ED" box="[323,500,1155,1179]" gridcol="1" gridrow="27" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE120B98EA3A8ED" box="[533,648,1155,1179]" gridcol="2" gridrow="27" pageId="32" pageNumber="33">13.5±2.0</td>
<td id="76845DB7F97262B3FD4520B98F0CA8ED" box="[689,807,1155,1179]" gridcol="3" gridrow="27" pageId="32" pageNumber="33">(917)</td>
<td id="76845DB7F97262B3FCA620B98FEEA8ED" box="[850,965,1155,1179]" gridcol="4" gridrow="27" pageId="32" pageNumber="33">13.8±1.7</td>
<td id="76845DB7F97262B3FC1320B98876A8ED" box="[999,1117,1155,1179]" gridcol="5" gridrow="27" pageId="32" pageNumber="33">(1116)</td>
<td id="76845DB7F97262B3FB7520B988C3A8ED" box="[1153,1256,1155,1179]" gridcol="6" gridrow="27" pageId="32" pageNumber="33">15.0±1.8</td>
<td id="76845DB7F97262B3FAE420B989ADA8ED" box="[1296,1414,1155,1179]" gridcol="7" gridrow="27" pageId="32" pageNumber="33">(1218)</td>
</tr>
<tr id="355534CBF97262B3FF57209C89ADA8C9" box="[163,1414,1190,1215]" gridrow="28" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF57209C8D32A8C9" box="[163,281,1190,1215]" gridcol="0" gridrow="28" pageId="32" pageNumber="33">
<emphasis id="B911EA9BF9729D6CFF44209D8CF9A8C8" box="[176,210,1191,1214]" italics="true" pageId="32" pageNumber="33">GP</emphasis>
c
</th>
<td id="76845DB7F97262B3FEB7209C8DDFA8C9" box="[323,500,1190,1215]" gridcol="1" gridrow="28" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE1209C8EA3A8C9" box="[533,648,1190,1215]" gridcol="2" gridrow="28" pageId="32" pageNumber="33">42.4±9.1</td>
<td id="76845DB7F97262B3FD45209C8F0CA8C9" box="[689,807,1190,1215]" gridcol="3" gridrow="28" pageId="32" pageNumber="33">(28.256.7)</td>
<td id="76845DB7F97262B3FCA6209C8FEEA8C9" box="[850,965,1190,1215]" gridcol="4" gridrow="28" pageId="32" pageNumber="33">48.6±6.6</td>
<td id="76845DB7F97262B3FC13209C8876A8C9" box="[999,1117,1190,1215]" gridcol="5" gridrow="28" pageId="32" pageNumber="33">(35.359.1)</td>
<td id="76845DB7F97262B3FB75209C88C3A8C9" box="[1153,1256,1190,1215]" gridcol="6" gridrow="28" pageId="32" pageNumber="33">56.7±15.9</td>
<td id="76845DB7F97262B3FAE4209C89ADA8C9" box="[1296,1414,1190,1215]" gridcol="7" gridrow="28" pageId="32" pageNumber="33">(35.381.8)</td>
</tr>
<tr id="355534CBF97262B3FF5720F089ADA894" box="[163,1414,1226,1250]" gridrow="29" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF5720F08D32A894" box="[163,281,1226,1250]" gridcol="0" gridrow="29" pageId="32" pageNumber="33">
<emphasis id="B911EA9BF9729D6CFF4420F18CCEA894" box="[176,229,1227,1250]" italics="true" pageId="32" pageNumber="33">GCO</emphasis>
d (%)
</th>
<td id="76845DB7F97262B3FEB720F08DDFA894" box="[323,500,1226,1250]" gridcol="1" gridrow="29" pageId="32" pageNumber="33">(Ratio)</td>
<td id="76845DB7F97262B3FDE120F08EA3A894" box="[533,648,1226,1250]" gridcol="2" gridrow="29" pageId="32" pageNumber="33">33.3</td>
<td id="76845DB7F97262B3FD4520F08F0CA894" box="[689,807,1226,1250]" gridcol="3" gridrow="29" pageId="32" pageNumber="33">(O4: C8)</td>
<td id="76845DB7F97262B3FCA620F08FEEA894" box="[850,965,1226,1250]" gridcol="4" gridrow="29" pageId="32" pageNumber="33">78.6</td>
<td id="76845DB7F97262B3FC1320F08876A894" box="[999,1117,1226,1250]" gridcol="5" gridrow="29" pageId="32" pageNumber="33">(O11: C3)</td>
<td id="76845DB7F97262B3FB7520F088C3A894" box="[1153,1256,1226,1250]" gridcol="6" gridrow="29" pageId="32" pageNumber="33">54.5</td>
<td id="76845DB7F97262B3FAE420F089ADA894" box="[1296,1414,1226,1250]" gridcol="7" gridrow="29" pageId="32" pageNumber="33">(O6: C5)</td>
</tr>
<tr id="355534CBF97262B3FF5720D489ADA970" box="[163,1414,1262,1286]" gridrow="30" pageId="32" pageNumber="33" rowspan-3="1" rowspan-5="1" rowspan-7="1">
<th id="76845DB7F97262B3FF5720D48D32A970" box="[163,281,1262,1286]" gridcol="0" gridrow="30" pageId="32" pageNumber="33">Body wall</th>
<td id="76845DB7F97262B3FEB720D48DDFA970" box="[323,500,1262,1286]" gridcol="1" gridrow="30" pageId="32" pageNumber="33">
Anchor plates (
<emphasis id="B911EA9BF9729D6CFE2B20D58DC7A970" box="[479,492,1263,1286]" italics="true" pageId="32" pageNumber="33">n</emphasis>
)
</td>
<td id="76845DB7F97262B3FDE120D48EA3A970" box="[533,648,1262,1286]" gridcol="2" gridrow="30" pageId="32" pageNumber="33">8</td>
<td id="76845DB7F97262B3FCA620D48FEEA970" box="[850,965,1262,1286]" gridcol="4" gridrow="30" pageId="32" pageNumber="33">9</td>
<td id="76845DB7F97262B3FB7520D488C3A970" box="[1153,1256,1262,1286]" gridcol="6" gridrow="30" pageId="32" pageNumber="33">10</td>
</tr>
<tr id="355534CBF97262B3FF57212889ADA95C" box="[163,1414,1298,1322]" gridrow="31" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF5721288D32A95C" box="[163,281,1298,1322]" gridcol="0" gridrow="31" pageId="32" pageNumber="33">Length</th>
<td id="76845DB7F97262B3FEB721288DDFA95C" box="[323,500,1298,1322]" gridcol="1" gridrow="31" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE121288EA3A95C" box="[533,648,1298,1322]" gridcol="2" gridrow="31" pageId="32" pageNumber="33">93.9±4.2</td>
<td id="76845DB7F97262B3FD4521288F0CA95C" box="[689,807,1298,1322]" gridcol="3" gridrow="31" pageId="32" pageNumber="33">(8899)</td>
<td id="76845DB7F97262B3FCA621288FEEA95C" box="[850,965,1298,1322]" gridcol="4" gridrow="31" pageId="32" pageNumber="33">89.4±8.1</td>
<td id="76845DB7F97262B3FC1321288876A95C" box="[999,1117,1298,1322]" gridcol="5" gridrow="31" pageId="32" pageNumber="33">(7598)</td>
<td id="76845DB7F97262B3FB75212888C3A95C" box="[1153,1256,1298,1322]" gridcol="6" gridrow="31" pageId="32" pageNumber="33">94.5±3.8</td>
<td id="76845DB7F97262B3FAE4212889ADA95C" box="[1296,1414,1298,1322]" gridcol="7" gridrow="31" pageId="32" pageNumber="33">(8899)</td>
</tr>
<tr id="355534CBF97262B3FF57210F89ADA938" box="[163,1414,1333,1358]" gridrow="32" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF57210F8D32A938" box="[163,281,1333,1358]" gridcol="0" gridrow="32" pageId="32" pageNumber="33">Width</th>
<td id="76845DB7F97262B3FEB7210F8DDFA938" box="[323,500,1333,1358]" gridcol="1" gridrow="32" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE1210F8EA3A938" box="[533,648,1333,1358]" gridcol="2" gridrow="32" pageId="32" pageNumber="33">50.4±3.0</td>
<td id="76845DB7F97262B3FD45210F8F0CA938" box="[689,807,1333,1358]" gridcol="3" gridrow="32" pageId="32" pageNumber="33">(4755)</td>
<td id="76845DB7F97262B3FCA6210F8FEEA938" box="[850,965,1333,1358]" gridcol="4" gridrow="32" pageId="32" pageNumber="33">55.6±2.8</td>
<td id="76845DB7F97262B3FC13210F8876A938" box="[999,1117,1333,1358]" gridcol="5" gridrow="32" pageId="32" pageNumber="33">(5362)</td>
<td id="76845DB7F97262B3FB75210F88C3A938" box="[1153,1256,1333,1358]" gridcol="6" gridrow="32" pageId="32" pageNumber="33">59.1±5.7</td>
<td id="76845DB7F97262B3FAE4210F89ADA938" box="[1296,1414,1333,1358]" gridcol="7" gridrow="32" pageId="32" pageNumber="33">(4967)</td>
</tr>
<tr id="355534CBF97262B3FF57216289ADA907" box="[163,1414,1368,1393]" gridrow="33" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF5721628D32A907" box="[163,281,1368,1393]" gridcol="0" gridrow="33" pageId="32" pageNumber="33">
<emphasis id="B911EA9BF9729D6CFF4421608CF7A907" box="[176,220,1370,1393]" italics="true" pageId="32" pageNumber="33">APS</emphasis>
e
</th>
<td id="76845DB7F97262B3FEB721628DDFA907" box="[323,500,1368,1393]" gridcol="1" gridrow="33" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE121628EA3A907" box="[533,648,1368,1393]" gridcol="2" gridrow="33" pageId="32" pageNumber="33">0.7±0.6</td>
<td id="76845DB7F97262B3FD4521628F0CA907" box="[689,807,1368,1393]" gridcol="3" gridrow="33" pageId="32" pageNumber="33">(02.1)</td>
<td id="76845DB7F97262B3FCA621628FEEA907" box="[850,965,1368,1393]" gridcol="4" gridrow="33" pageId="32" pageNumber="33">0.2±0.3</td>
<td id="76845DB7F97262B3FC1321628876A907" box="[999,1117,1368,1393]" gridcol="5" gridrow="33" pageId="32" pageNumber="33">(00.7)</td>
<td id="76845DB7F97262B3FB75216288C3A907" box="[1153,1256,1368,1393]" gridcol="6" gridrow="33" pageId="32" pageNumber="33">1.8 ±1.6</td>
<td id="76845DB7F97262B3FAE4216289ADA907" box="[1296,1414,1368,1393]" gridcol="7" gridrow="33" pageId="32" pageNumber="33">(0.55.4)</td>
</tr>
<tr id="355534CBF97262B3FF57214789ADA9E3" box="[163,1414,1405,1429]" gridrow="34" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF5721478D32A9E3" box="[163,281,1405,1429]" gridcol="0" gridrow="34" pageId="32" pageNumber="33">
<emphasis id="B911EA9BF9729D6CFF4421448CFEA9E3" box="[176,213,1406,1429]" italics="true" pageId="32" pageNumber="33">Ntp</emphasis>
f
</th>
<td id="76845DB7F97262B3FEB721478DDFA9E3" box="[323,500,1405,1429]" gridcol="1" gridrow="34" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE121478EA3A9E3" box="[533,648,1405,1429]" gridcol="2" gridrow="34" pageId="32" pageNumber="33">2.5±0.9</td>
<td id="76845DB7F97262B3FD4521478F0CA9E3" box="[689,807,1405,1429]" gridcol="3" gridrow="34" pageId="32" pageNumber="33">(14)</td>
<td id="76845DB7F97262B3FCA621478FEEA9E3" box="[850,965,1405,1429]" gridcol="4" gridrow="34" pageId="32" pageNumber="33">1.2±1.0</td>
<td id="76845DB7F97262B3FC1321478876A9E3" box="[999,1117,1405,1429]" gridcol="5" gridrow="34" pageId="32" pageNumber="33">(03)</td>
<td id="76845DB7F97262B3FB75214788C3A9E3" box="[1153,1256,1405,1429]" gridcol="6" gridrow="34" pageId="32" pageNumber="33">1.7±0.7</td>
<td id="76845DB7F97262B3FAE4214789ADA9E3" box="[1296,1414,1405,1429]" gridcol="7" gridrow="34" pageId="32" pageNumber="33">(13)</td>
</tr>
<tr id="355534CBF97262B3FF57219A89ADA9CF" box="[163,1414,1440,1465]" gridrow="35" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF57219A8D32A9CF" box="[163,281,1440,1465]" gridcol="0" gridrow="35" pageId="32" pageNumber="33">
<emphasis id="B911EA9BF9729D6CFF44219B8CF6A9CE" box="[176,221,1441,1464]" italics="true" pageId="32" pageNumber="33">APT</emphasis>
g (%)
</th>
<td id="76845DB7F97262B3FEB7219A8DDFA9CF" box="[323,500,1440,1465]" gridcol="1" gridrow="35" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE1219A8EA3A9CF" box="[533,648,1440,1465]" gridcol="2" gridrow="35" pageId="32" pageNumber="33">16.8 ± 4.9</td>
<td id="76845DB7F97262B3FD45219A8F0CA9CF" box="[689,807,1440,1465]" gridcol="3" gridrow="35" pageId="32" pageNumber="33">(8.323.1)</td>
<td id="76845DB7F97262B3FCA6219A8FEEA9CF" box="[850,965,1440,1465]" gridcol="4" gridrow="35" pageId="32" pageNumber="33">10.1 ± 8.8</td>
<td id="76845DB7F97262B3FC13219A8876A9CF" box="[999,1117,1440,1465]" gridcol="5" gridrow="35" pageId="32" pageNumber="33">(027.3)</td>
<td id="76845DB7F97262B3FB75219A88C3A9CF" box="[1153,1256,1440,1465]" gridcol="6" gridrow="35" pageId="32" pageNumber="33">9.8±3.3</td>
<td id="76845DB7F97262B3FAE4219A89ADA9CF" box="[1296,1414,1440,1465]" gridcol="7" gridrow="35" pageId="32" pageNumber="33">(5.313.6)</td>
</tr>
<tr id="355534CBF97262B3FF5721FE89ADA9AB" box="[163,1414,1476,1501]" gridrow="36" pageId="32" pageNumber="33" rowspan-3="1" rowspan-5="1" rowspan-7="1">
<th id="76845DB7F97262B3FF5721FE8D32A9AB" box="[163,281,1476,1501]" gridcol="0" gridrow="36" pageId="32" pageNumber="33">Body wall</th>
<td id="76845DB7F97262B3FEB721FE8DDFA9AB" box="[323,500,1476,1501]" gridcol="1" gridrow="36" pageId="32" pageNumber="33">
Anchor (
<emphasis id="B911EA9BF9729D6CFE4A21FF8DE0A9AA" box="[446,459,1477,1500]" italics="true" pageId="32" pageNumber="33">n</emphasis>
)
</td>
<td id="76845DB7F97262B3FDE121FE8EA3A9AB" box="[533,648,1476,1501]" gridcol="2" gridrow="36" pageId="32" pageNumber="33">8</td>
<td id="76845DB7F97262B3FCA621FE8FEEA9AB" box="[850,965,1476,1501]" gridcol="4" gridrow="36" pageId="32" pageNumber="33">12</td>
<td id="76845DB7F97262B3FB7521FE88C3A9AB" box="[1153,1256,1476,1501]" gridcol="6" gridrow="36" pageId="32" pageNumber="33">10</td>
</tr>
<tr id="355534CBF97262B3FF5721CB89ADAA7F" box="[163,1414,1521,1545]" gridrow="37" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF5721CB8D32AA7F" box="[163,281,1521,1545]" gridcol="0" gridrow="37" pageId="32" pageNumber="33">Length</th>
<td id="76845DB7F97262B3FEB721CB8DDFAA7F" box="[323,500,1521,1545]" gridcol="1" gridrow="37" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE121CB8EA3AA7F" box="[533,648,1521,1545]" gridcol="2" gridrow="37" pageId="32" pageNumber="33">162.3 ± 8.5</td>
<td id="76845DB7F97262B3FD4521CB8F0CAA7F" box="[689,807,1521,1545]" gridcol="3" gridrow="37" pageId="32" pageNumber="33">(145172)</td>
<td id="76845DB7F97262B3FCA621CB8FEEAA7F" box="[850,965,1521,1545]" gridcol="4" gridrow="37" pageId="32" pageNumber="33">153.1 ± 7.9</td>
<td id="76845DB7F97262B3FC1321CB8876AA7F" box="[999,1117,1521,1545]" gridcol="5" gridrow="37" pageId="32" pageNumber="33">(144168)</td>
<td id="76845DB7F97262B3FB7521CB88C3AA7F" box="[1153,1256,1521,1545]" gridcol="6" gridrow="37" pageId="32" pageNumber="33">157.4±6.7</td>
<td id="76845DB7F97262B3FAE421CB89ADAA7F" box="[1296,1414,1521,1545]" gridcol="7" gridrow="37" pageId="32" pageNumber="33">(145168)</td>
</tr>
<tr id="355534CBF97262B3FF57222F89ADAA5B" box="[163,1414,1557,1581]" gridrow="38" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF57222F8D32AA5B" box="[163,281,1557,1581]" gridcol="0" gridrow="38" pageId="32" pageNumber="33">Width</th>
<td id="76845DB7F97262B3FEB7222F8DDFAA5B" box="[323,500,1557,1581]" gridcol="1" gridrow="38" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE1222F8EA3AA5B" box="[533,648,1557,1581]" gridcol="2" gridrow="38" pageId="32" pageNumber="33">61.5 ± 4.7</td>
<td id="76845DB7F97262B3FD45222F8F0CAA5B" box="[689,807,1557,1581]" gridcol="3" gridrow="38" pageId="32" pageNumber="33">(5467)</td>
<td id="76845DB7F97262B3FCA6222F8FEEAA5B" box="[850,965,1557,1581]" gridcol="4" gridrow="38" pageId="32" pageNumber="33">64.8 ± 3.2</td>
<td id="76845DB7F97262B3FC13222F8876AA5B" box="[999,1117,1557,1581]" gridcol="5" gridrow="38" pageId="32" pageNumber="33">(5870)</td>
<td id="76845DB7F97262B3FB75222F88C3AA5B" box="[1153,1256,1557,1581]" gridcol="6" gridrow="38" pageId="32" pageNumber="33">79.7±4.9</td>
<td id="76845DB7F97262B3FAE4222F89ADAA5B" box="[1296,1414,1557,1581]" gridcol="7" gridrow="38" pageId="32" pageNumber="33">(7389)</td>
</tr>
<tr id="355534CBF97262B3FF57220389ADAA27" box="[163,1414,1593,1617]" gridrow="39" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF5722038D32AA27" box="[163,281,1593,1617]" gridcol="0" gridrow="39" pageId="32" pageNumber="33">Teeth</th>
<td id="76845DB7F97262B3FEB722038DDFAA27" box="[323,500,1593,1617]" gridcol="1" gridrow="39" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE122038EA3AA27" box="[533,648,1593,1617]" gridcol="2" gridrow="39" pageId="32" pageNumber="33">6.2 ± 0.6</td>
<td id="76845DB7F97262B3FD4522038F0CAA27" box="[689,807,1593,1617]" gridcol="3" gridrow="39" pageId="32" pageNumber="33">(57)</td>
<td id="76845DB7F97262B3FCA622038FEEAA27" box="[850,965,1593,1617]" gridcol="4" gridrow="39" pageId="32" pageNumber="33">3.2 ± 1.9</td>
<td id="76845DB7F97262B3FC1322038876AA27" box="[999,1117,1593,1617]" gridcol="5" gridrow="39" pageId="32" pageNumber="33">(06)</td>
<td id="76845DB7F97262B3FB75220388C3AA27" box="[1153,1256,1593,1617]" gridcol="6" gridrow="39" pageId="32" pageNumber="33">9.9±1.2</td>
<td id="76845DB7F97262B3FAE4220389ADAA27" box="[1296,1414,1593,1617]" gridcol="7" gridrow="39" pageId="32" pageNumber="33">(812)</td>
</tr>
<tr id="355534CBF97262B3FF57226689ADAA03" box="[163,1414,1628,1653]" gridrow="40" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF5722668D32AA03" box="[163,281,1628,1653]" gridcol="0" gridrow="40" pageId="32" pageNumber="33">
<emphasis id="B911EA9BF9729D6CFF4422678CCAAA02" box="[176,225,1629,1652]" italics="true" pageId="32" pageNumber="33">ASW</emphasis>
h(%)
</th>
<td id="76845DB7F97262B3FEB722668DDFAA03" box="[323,500,1628,1653]" gridcol="1" gridrow="40" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE122668EA3AA03" box="[533,648,1628,1653]" gridcol="2" gridrow="40" pageId="32" pageNumber="33">28.4 ± 2.5</td>
<td id="76845DB7F97262B3FD4522668F0CAA03" box="[689,807,1628,1653]" gridcol="3" gridrow="40" pageId="32" pageNumber="33">(25.832.4)</td>
<td id="76845DB7F97262B3FCA622668FEEAA03" box="[850,965,1628,1653]" gridcol="4" gridrow="40" pageId="32" pageNumber="33">25.0 ± 1.9</td>
<td id="76845DB7F97262B3FC1322668876AA03" box="[999,1117,1628,1653]" gridcol="5" gridrow="40" pageId="32" pageNumber="33">(22.228.1)</td>
<td id="76845DB7F97262B3FB75226688C3AA03" box="[1153,1256,1628,1653]" gridcol="6" gridrow="40" pageId="32" pageNumber="33">19.9±1.1</td>
<td id="76845DB7F97262B3FAE4226689ADAA03" box="[1296,1414,1628,1653]" gridcol="7" gridrow="40" pageId="32" pageNumber="33">(18.522.1)</td>
</tr>
<tr id="355534CBF97262B3FF5722BA89ADAAEE" box="[163,1414,1664,1688]" gridrow="41" pageId="32" pageNumber="33">
<th id="76845DB7F97262B3FF5722BA8D32AAEE" box="[163,281,1664,1688]" gridcol="0" gridrow="41" pageId="32" pageNumber="33">
<emphasis id="B911EA9BF9729D6CFF4422BB8CCFAAEE" box="[176,228,1665,1688]" italics="true" pageId="32" pageNumber="33">AEW</emphasis>
i (%)
</th>
<td id="76845DB7F97262B3FEB722BA8DDFAAEE" box="[323,500,1664,1688]" gridcol="1" gridrow="41" pageId="32" pageNumber="33">(Range)</td>
<td id="76845DB7F97262B3FDE122BA8EA3AAEE" box="[533,648,1664,1688]" gridcol="2" gridrow="41" pageId="32" pageNumber="33">59.2 ± 4.3</td>
<td id="76845DB7F97262B3FD4522BA8F0CAAEE" box="[689,807,1664,1688]" gridcol="3" gridrow="41" pageId="32" pageNumber="33">(52.264.9)</td>
<td id="76845DB7F97262B3FCA622BA8FEEAAEE" box="[850,965,1664,1688]" gridcol="4" gridrow="41" pageId="32" pageNumber="33">50.8 ± 4.7</td>
<td id="76845DB7F97262B3FC1322BA8876AAEE" box="[999,1117,1664,1688]" gridcol="5" gridrow="41" pageId="32" pageNumber="33">(41.255.6)</td>
<td id="76845DB7F97262B3FB7522BA88C3AAEE" box="[1153,1256,1664,1688]" gridcol="6" gridrow="41" pageId="32" pageNumber="33">48.6±2.7</td>
<td id="76845DB7F97262B3FAE422BA89ADAAEE" box="[1296,1414,1664,1688]" gridcol="7" gridrow="41" pageId="32" pageNumber="33">(43.852.0)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BDA3689F9729D6CFF63229E89BFAACD" blockId="32.[151,1428,1699,1723]" box="[151,1428,1699,1723]" pageId="32" pageNumber="33">
<tableNote id="76833707F9729D6CFF63229E89BFAACD" box="[151,1428,1699,1723]" pageId="32" pageNumber="33" targetBox="[163,1414,190,1688]" targetPageId="32">
aI, (see, Materials and Methods) Indicators for ossicle shapes, and one parameter (
<emphasis id="B911EA9BF9729D6CFBFF229E881AAACD" box="[1035,1073,1700,1723]" italics="true" pageId="32" pageNumber="33">Ntp</emphasis>
, number of teethed perforations).
</tableNote>
</paragraph>
<caption id="DF1A6601F9739D6DFF6320D08DD3A918" ID-DOI="http://doi.org/10.5281/zenodo.14703564" ID-Zenodo-Dep="14703564" httpUri="https://zenodo.org/record/14703564/files/figure.png" pageId="33" pageNumber="34" startId="33.[151,250,1258,1282]" targetBox="[151,1436,181,1234]" targetPageId="33" targetType="figure">
<paragraph id="8BDA3689F9739D6DFF6320D08DD3A918" blockId="33.[151,1436,1258,1391]" pageId="33" pageNumber="34">
<emphasis id="B911EA9BF9739D6DFF6320D08D0FA974" bold="true" box="[151,292,1258,1282]" pageId="33" pageNumber="34">FIGURE 15.</emphasis>
External view of a damaged specimen collected from Yoron-jima Island, Kagoshima, erected as
<taxonomicName id="4C654D0AF9739D6DFACC20D18D0EA950" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="33" pageNumber="34" phylum="Echinodermata" rank="species" species="parvaspiculus" status="sp. nov.">
<emphasis id="B911EA9BF9739D6DFACC20D18D0EA950" italics="true" pageId="33" pageNumber="34">Patinapta parvaspiculus</emphasis>
</taxonomicName>
<emphasis id="B911EA9BF9739D6DFED921358D55A951" bold="true" box="[301,382,1295,1319]" pageId="33" pageNumber="34">
<taxonomicNameLabel id="A22257E0F9739D6DFED921358D55A951" box="[301,382,1295,1319]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, in the present study (holotype specimen, WMNH-2023-INV-157): A, lateral view, anterior left, dorsal top with damage on anterior body wall (from sampling shovel; gonad tubules also damaged); B, oral-side view of the tentacles; C, outer-side view of the tentacles.
</paragraph>
</caption>
<paragraph id="8BDA3689F9739D6DFF3321FB8D1DAA1D" blockId="33.[151,1437,1472,1823]" pageId="33" pageNumber="34">
Anchor ossicles ranged within 144173 μm in length (
<figureCitation id="135E2A0CF9739D6DFCDA21FB8F93A9AD" box="[814,952,1472,1499]" captionStart="FIGURE 16" captionStartId="34.[151,250,1674,1698]" captionTargetBox="[158,1430,181,1643]" captionTargetId="figure-29@34.[158,1430,181,1650]" captionTargetPageId="34" captionText="FIGURE 16. SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, erected as Patinapta parvaspiculus sp. nov., in the present study (holotype specimen, WMNH-2023-INV-157): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703566" httpUri="https://zenodo.org/record/14703566/files/figure.png" pageId="33" pageNumber="34">Fig. 16B, D</figureCitation>
;
<tableCitation id="C6E70332F9739D6DFC3021FB8832A9AD" box="[964,1049,1473,1499]" captionStart="TABLE 9" captionStartId="32.[152,243,152,176]" captionTargetPageId="32" captionText="TABLE 9. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta parvaspiculus sp. nov." pageId="33" pageNumber="34">Table 9</tableCitation>
). All anchor arms (bills) equipped with minute teeth on their outer tips, numbers of teeth variable within 06 (mostly
<quantity id="4C9D9B6CF9739D6DFBBB21DF88BCA989" box="[1103,1175,1509,1535]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="5.08" metricValueMin="2.54" pageId="33" pageNumber="34" unit="in" value="1.5" valueMax="2.0" valueMin="1.0">12 in</quantity>
one arm of the anchor ossicle), and anchor ossicles of this specimen possessed narrow stem and also relatively narrow anchor arm width, with
<emphasis id="B911EA9BF9739D6DFF3B22178D2EAA31" box="[207,261,1581,1607]" italics="true" pageId="33" pageNumber="34">ASW</emphasis>
=21.829.5%;
<emphasis id="B911EA9BF9739D6DFE5E22178DCFAA31" box="[426,484,1581,1607]" italics="true" pageId="33" pageNumber="34">AEW</emphasis>
=41.260.7%. T-shaped basal end of the anchor ossicle of this species crescent-shaped (
<figureCitation id="135E2A0CF9739D6DFF6B226B8D02AA1D" box="[159,297,1616,1643]" captionStart="FIGURE 16" captionStartId="34.[151,250,1674,1698]" captionTargetBox="[158,1430,181,1643]" captionTargetId="figure-29@34.[158,1430,181,1650]" captionTargetPageId="34" captionText="FIGURE 16. SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, erected as Patinapta parvaspiculus sp. nov., in the present study (holotype specimen, WMNH-2023-INV-157): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703566" httpUri="https://zenodo.org/record/14703566/files/figure.png" pageId="33" pageNumber="34">Fig. 16B; D</figureCitation>
).
</paragraph>
<paragraph id="8BDA3689F9739D6DFF33224F88B8AAA1" blockId="33.[151,1437,1472,1823]" pageId="33" pageNumber="34">
Body longitudinal muscle, granule ossicles of this specimen ranged within 2134 μm in length (
<figureCitation id="135E2A0CF9739D6DFAFB224F89BCAAF9" box="[1295,1431,1652,1679]" captionStart="FIGURE 16" captionStartId="34.[151,250,1674,1698]" captionTargetBox="[158,1430,181,1643]" captionTargetId="figure-29@34.[158,1430,181,1650]" captionTargetPageId="34" captionText="FIGURE 16. SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, erected as Patinapta parvaspiculus sp. nov., in the present study (holotype specimen, WMNH-2023-INV-157): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703566" httpUri="https://zenodo.org/record/14703566/files/figure.png" pageId="33" pageNumber="34">Fig. 16C, E</figureCitation>
;
<tableCitation id="C6E70332F9739D6DFF6322A38CC0AAC5" box="[151,235,1689,1715]" captionStart="TABLE 9" captionStartId="32.[152,243,152,176]" captionTargetPageId="32" captionText="TABLE 9. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta parvaspiculus sp. nov." pageId="33" pageNumber="34">Table 9</tableCitation>
). The calculated indicator for granule proportions were variated within
<emphasis id="B911EA9BF9739D6DFBF422A3880EAAC5" box="[1024,1061,1689,1715]" italics="true" pageId="33" pageNumber="34">GP</emphasis>
=35.364.3%;
<emphasis id="B911EA9BF9739D6DFB3E22A3892EAAC5" box="[1226,1285,1689,1715]" italics="true" pageId="33" pageNumber="34">GCO</emphasis>
=36.478.6% did not have noticeable differences in amounts between the O-shape and C-shape granules.
</paragraph>
<paragraph id="8BDA3689F9739D6DFF3322DB8883AB69" blockId="33.[151,1437,1472,1823]" pageId="33" pageNumber="34">
Tentacle ossicles rods, ranged within 53102 μm in length (
<figureCitation id="135E2A0CF9739D6DFCAC22DB8F95AA8C" box="[856,958,1760,1787]" captionStart="FIGURE 16" captionStartId="34.[151,250,1674,1698]" captionTargetBox="[158,1430,181,1643]" captionTargetId="figure-29@34.[158,1430,181,1650]" captionTargetPageId="34" captionText="FIGURE 16. SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, erected as Patinapta parvaspiculus sp. nov., in the present study (holotype specimen, WMNH-2023-INV-157): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703566" httpUri="https://zenodo.org/record/14703566/files/figure.png" pageId="33" pageNumber="34">Fig. 16A</figureCitation>
,
<tableCitation id="C6E70332F9739D6DFC3322DB8830AA8D" box="[967,1051,1761,1787]" captionStart="TABLE 9" captionStartId="32.[152,243,152,176]" captionTargetPageId="32" captionText="TABLE 9. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta parvaspiculus sp. nov." pageId="33" pageNumber="34">Table 9</tableCitation>
). Two calculated indicators for rod proportions and complexities were variated within
<emphasis id="B911EA9BF9739D6DFD26233F8EDFAB68" box="[722,756,1797,1822]" italics="true" pageId="33" pageNumber="34">RP</emphasis>
= 7.116.9%;
<emphasis id="B911EA9BF9739D6DFC6E233F8F95AB69" box="[922,958,1797,1823]" italics="true" pageId="33" pageNumber="34">RC</emphasis>
=2.711.7 (
<tableCitation id="C6E70332F9739D6DFBB7233F88B1AB69" box="[1091,1178,1797,1823]" captionStart="TABLE 9" captionStartId="32.[152,243,152,176]" captionTargetPageId="32" captionText="TABLE 9. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta parvaspiculus sp. nov." pageId="33" pageNumber="34">Table 9</tableCitation>
).
</paragraph>
<caption id="DF1A6601F9709D6EFF6322B08EA7AB78" ID-DOI="http://doi.org/10.5281/zenodo.14703566" ID-Zenodo-Dep="14703566" httpUri="https://zenodo.org/record/14703566/files/figure.png" pageId="34" pageNumber="35" startId="34.[151,250,1674,1698]" targetBox="[158,1430,181,1643]" targetPageId="34" targetType="figure">
<paragraph id="8BDA3689F9709D6EFF6322B08EA7AB78" blockId="34.[151,1436,1674,1806]" pageId="34" pageNumber="35">
<emphasis id="B911EA9BF9709D6EFF6322B08D0BAAD4" bold="true" box="[151,288,1674,1698]" pageId="34" pageNumber="35">FIGURE 16.</emphasis>
SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, erected as
<taxonomicName id="4C654D0AF9709D6EFF6322958DBBAAB0" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[151,400,1710,1734]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="34" pageNumber="35" phylum="Echinodermata" rank="species" species="parvaspiculus" status="sp. nov.">
<emphasis id="B911EA9BF9709D6EFF6322958DBBAAB0" box="[151,400,1710,1734]" italics="true" pageId="34" pageNumber="35">Patinapta parvaspiculus</emphasis>
</taxonomicName>
<emphasis id="B911EA9BF9709D6EFE6222948DCCAAB0" bold="true" box="[406,487,1710,1734]" pageId="34" pageNumber="35">
<taxonomicNameLabel id="A22257E0F9709D6EFE6222948DCCAAB0" box="[406,487,1710,1734]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, in the present study (holotype specimen, WMNH-2023-INV-157): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle.
</paragraph>
</caption>
<paragraph id="8BDA3689F9709D6EFF6323638F36AB05" blockId="34.[151,797,1881,1908]" box="[151,797,1881,1908]" pageId="34" pageNumber="35">
<heading id="D09281E5F9709D6EFF6323638F36AB05" bold="true" box="[151,797,1881,1908]" fontSize="11" level="1" pageId="34" pageNumber="35" reason="1">
<emphasis id="B911EA9BF9709D6EFF6323638F36AB05" bold="true" box="[151,797,1881,1908]" pageId="34" pageNumber="35">Observation on the morphologies of calcareous ring</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA3689F9709D6FFF6323988DEDAC8F" blockId="34.[151,1436,1954,2016]" lastBlockId="35.[151,1436,151,249]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
Calcareous plates 12, mostly firmly bound (
<figureCitation id="135E2A0CF9709D6EFD7D23988EF7ABCA" box="[649,732,1954,1980]" captionStart="FIGURE 17" captionStartId="35.[151,250,628,652]" captionTargetBox="[164,1422,281,600]" captionTargetId="figure-65@35.[151,1435,273,605]" captionTargetPageId="35" captionText="FIGURE 17. Outer-surface view of calcareous ring plates of a specimen collected from Yoron-jima Island, Kagoshima, erected as Patinapta parvaspiculus sp. nov., in the present study (holotype specimen, WMNH-2023-INV-157). The positions of the plates are denoted by Ludwigs [18891892 (1889), p156] system: RIRV and IR1IR5 where R and IR stands for positions for radius and interradius, respectively. Hence RI, mid-ventral; RII, left ventral; RIII, right ventral; RIV, left dorsal; RV, right dorsal; and: IR1, left ventral; IR2, right ventral; IR3-IR3, left dorso-lateral; IR4-IR4, right dorso-lateral; IR5, mid-dorsal. Anterior top." figureDoi="http://doi.org/10.5281/zenodo.14703568" httpUri="https://zenodo.org/record/14703568/files/figure.png" pageId="34" pageNumber="35">Fig. 17</figureCitation>
), with slight anterior projection and deep posterior depression in interradial plates, while only posterior depression in radial plates. All radial plates perforated in each anterior half. The connections between 4 plates situated in right and left dorso-lateral positions (
<emphasis id="B911EA9BF9719D6FFB9A24AD896AACC6" box="[1134,1345,151,177]" italics="true" pageId="35" pageNumber="36">IR4; RIII; IR3; RII</emphasis>
), where additional interradial plate (
<emphasis id="B911EA9BF9719D6FFE2D24818E7AACA3" box="[473,593,187,213]" italics="true" pageId="35" pageNumber="36">IR4; IR3</emphasis>
) and adjacent radial plate (
<emphasis id="B911EA9BF9719D6FFC6724818FD3ACA2" box="[915,1016,187,212]" italics="true" pageId="35" pageNumber="36">RIII; RII</emphasis>
) form triangular shapes and weakly bound by their apical ends.
</paragraph>
<caption id="DF1A6601F9719D6FFF63264E8C94AF36" ID-DOI="http://doi.org/10.5281/zenodo.14703568" ID-Zenodo-Dep="14703568" httpUri="https://zenodo.org/record/14703568/files/figure.png" pageId="35" pageNumber="36" startId="35.[151,250,628,652]" targetBox="[164,1422,281,600]" targetPageId="35" targetType="figure">
<paragraph id="8BDA3689F9719D6FFF63264E8C94AF36" blockId="35.[151,1437,628,832]" pageId="35" pageNumber="36">
<emphasis id="B911EA9BF9719D6FFF63264E8D34AEFA" bold="true" box="[151,287,628,652]" pageId="35" pageNumber="36">FIGURE 17.</emphasis>
Outer-surface view of calcareous ring plates of a specimen collected from Yoron-jima Island, Kagoshima, erected as
<taxonomicName id="4C654D0AF9719D6FFF4126A38D84AEC7" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[181,431,665,689]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="35" pageNumber="36" phylum="Echinodermata" rank="species" species="parvaspiculus" status="sp. nov.">
<emphasis id="B911EA9BF9719D6FFF4126A38D84AEC7" box="[181,431,665,689]" italics="true" pageId="35" pageNumber="36">Patinapta parvaspiculus</emphasis>
</taxonomicName>
<emphasis id="B911EA9BF9719D6FFE4C26A38E21AEC7" bold="true" box="[440,522,665,689]" pageId="35" pageNumber="36">
<taxonomicNameLabel id="A22257E0F9719D6FFE4C26A38E21AEC7" box="[440,522,665,689]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, in the present study (holotype specimen, WMNH-2023-INV-157). The positions of the plates are denoted by Ludwigs [18891892 (1889), p156] system:
<emphasis id="B911EA9BF9719D6FFCB426878FA8AEA2" box="[832,899,701,724]" italics="true" pageId="35" pageNumber="36">RIRV</emphasis>
and
<emphasis id="B911EA9BF9719D6FFC4026878822AEA2" box="[948,1033,701,725]" italics="true" pageId="35" pageNumber="36">IR1IR5</emphasis>
where
<emphasis id="B911EA9BF9719D6FFBA026878848AEA2" box="[1108,1123,701,724]" italics="true" pageId="35" pageNumber="36">R</emphasis>
and
<emphasis id="B911EA9BF9719D6FFB6126878886AEA2" box="[1173,1197,701,724]" italics="true" pageId="35" pageNumber="36">IR</emphasis>
stands for positions for radius and interradius, respectively. Hence
<emphasis id="B911EA9BF9719D6FFDBC26DB8E4BAE8E" box="[584,608,737,760]" italics="true" pageId="35" pageNumber="36">RI</emphasis>
, mid-ventral;
<emphasis id="B911EA9BF9719D6FFD1B26DB8F24AE8E" box="[751,783,737,760]" italics="true" pageId="35" pageNumber="36">RII</emphasis>
, left ventral;
<emphasis id="B911EA9BF9719D6FFC6126DB8F95AE8E" box="[917,958,737,760]" italics="true" pageId="35" pageNumber="36">RIII</emphasis>
, right ventral;
<emphasis id="B911EA9BF9719D6FFBA626DB8852AE8E" box="[1106,1145,737,760]" italics="true" pageId="35" pageNumber="36">RIV</emphasis>
, left dorsal;
<emphasis id="B911EA9BF9719D6FFB0326DB893EAE8E" box="[1271,1301,737,760]" italics="true" pageId="35" pageNumber="36">RV</emphasis>
, right dorsal; and:
<emphasis id="B911EA9BF9719D6FFF38273F8CDBAF6B" box="[204,240,773,797]" italics="true" pageId="35" pageNumber="36">IR1</emphasis>
, left ventral;
<emphasis id="B911EA9BF9719D6FFE74273F8D8EAF6B" box="[384,421,773,797]" italics="true" pageId="35" pageNumber="36">IR2</emphasis>
, right ventral;
<emphasis id="B911EA9BF9719D6FFDB6273F8EB7AF6B" box="[578,668,773,797]" italics="true" pageId="35" pageNumber="36">IR3-IR3</emphasis>
, left dorso-lateral;
<emphasis id="B911EA9BF9719D6FFC92273F8FEBAF6B" box="[870,960,773,797]" italics="true" pageId="35" pageNumber="36">IR4-IR4</emphasis>
, right dorso-lateral;
<emphasis id="B911EA9BF9719D6FFB63273F8890AF6A" box="[1175,1211,773,796]" italics="true" pageId="35" pageNumber="36">IR5</emphasis>
, mid-dorsal. Anterior top.
</paragraph>
</caption>
<paragraph id="8BDA3689F9719D6FFF6327BF8EB8AFE9" blockId="35.[151,659,901,928]" box="[151,659,901,928]" pageId="35" pageNumber="36">
<heading id="D09281E5F9719D6FFF6327BF8EB8AFE9" bold="true" box="[151,659,901,928]" fontSize="11" level="1" pageId="35" pageNumber="36" reason="1">
<emphasis id="B911EA9BF9719D6FFF6327BF8EB8AFE9" bold="true" box="[151,659,901,928]" pageId="35" pageNumber="36">
Ossicle variation among
<specimenCount id="9D63FD00F9719D6FFE3827BC8EB8AFE9" box="[460,659,901,928]" count="3" pageId="35" pageNumber="36" type="generic">three specimens</specimenCount>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA3689F9719D6FFF6327F78F92A822" blockId="35.[151,1437,973,2008]" pageId="35" pageNumber="36">
<specimenCount id="9D63FD00F9719D6FFF6327F78D7CAF91" box="[151,343,973,999]" count="3" pageId="35" pageNumber="36" type="generic">Three specimens</specimenCount>
approximately
<quantity id="4C9D9B6CF9719D6FFDFF27F78EBEAF9E" box="[523,661,973,1000]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="35" pageNumber="36" unit="mm" value="75.0" valueMax="100.0" valueMin="50.0">50100 mm</quantity>
in length (
<tableCitation id="C6E70332F9719D6FFCE027F78F40AF9E" box="[788,875,973,1000]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionTargetPageId="6" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="35" pageNumber="36">Table 1</tableCitation>
),
<specimenCount id="9D63FD00F9719D6FFC7227F48805AF91" box="[902,1070,973,1000]" count="2" pageId="35" pageNumber="36" type="generic">two specimens</specimenCount>
(WMNH-2015-INV-107; 2023- INV-157) well-matured and possessing fully equipped gonad tubules in their body cavities, while
<specimenCount id="9D63FD00F9719D6FFAF527C889B6A87D" box="[1281,1437,1009,1036]" count="1" pageId="35" pageNumber="36" type="generic">one specimen</specimenCount>
(WMNH-2024-INV-187) immature and extended by strong contraction of latitudinal muscle. Their external and internal morphologies agreed with each others in many parts (
<tableCitation id="C6E70332F9719D6FFCA020038F83A822" box="[852,936,1081,1108]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionTargetPageId="6" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="35" pageNumber="36">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8BDA3689F9719D6FFF3320678DF8A9A9" blockId="35.[151,1437,973,2008]" pageId="35" pageNumber="36">
Body wall ossicles anchors and anchor plates. Anchor plate ossicle length and calculated indicators of
<specimenCount id="9D63FD00F9719D6FFA9020678D27A8ED" count="3" pageId="35" pageNumber="36" type="generic">three specimens</specimenCount>
ranged within: 75123 μm in length,
<emphasis id="B911EA9BF9719D6FFD5A20B88EF5A8EA" box="[686,734,1154,1180]" italics="true" pageId="35" pageNumber="36">APS</emphasis>
=011.3,
<emphasis id="B911EA9BF9719D6FFCB320B88F44A8ED" box="[839,879,1154,1179]" italics="true" pageId="35" pageNumber="36">Ntp</emphasis>
=04,
<emphasis id="B911EA9BF9719D6FFC4220B88FC3A8EA" box="[950,1000,1154,1180]" italics="true" pageId="35" pageNumber="36">APT</emphasis>
=027.3%. Among these values, length of anchor plate ossicles (both ventrally and dorsally) was significantly different (
<emphasis id="B911EA9BF9719D6FFBDF209C886CA8C9" box="[1067,1095,1190,1215]" italics="true" pageId="35" pageNumber="36">Ps</emphasis>
&lt;0.005, Kruskal-Wallis test), while non-size dependance indicator of plate skewness
<emphasis id="B911EA9BF9719D6FFCD420F08F7BA892" box="[800,848,1226,1252]" italics="true" pageId="35" pageNumber="36">APS</emphasis>
was not significantly different (
<emphasis id="B911EA9BF9719D6FFB3820F088C3A895" box="[1228,1256,1226,1251]" italics="true" pageId="35" pageNumber="36">Ps</emphasis>
&gt;0.05, Kruskal-Wallis test). However, the frequency of teethed perforations
<emphasis id="B911EA9BF9719D6FFCB920D48F54A97E" box="[845,895,1262,1288]" italics="true" pageId="35" pageNumber="36">APT</emphasis>
was significantly different (
<emphasis id="B911EA9BF9719D6FFB3420D488F7A971" box="[1216,1244,1262,1287]" italics="true" pageId="35" pageNumber="36">Ps</emphasis>
&lt;0.005, Kruskal-Wallis test), while contrary to this parameter
<emphasis id="B911EA9BF9719D6FFD5521288EE2A95D" box="[673,713,1298,1323]" italics="true" pageId="35" pageNumber="36">Ntp</emphasis>
was not significantly different (
<emphasis id="B911EA9BF9719D6FFBCC2128887FA95D" box="[1080,1108,1298,1323]" italics="true" pageId="35" pageNumber="36">Ps</emphasis>
&gt;0.05, Kruskal-Wallis test). From these results, the anchor plate ossicles in the body wall of the present
<specimenCount id="9D63FD00F9719D6FFC2D210F88A7A939" box="[985,1164,1333,1359]" count="3" pageId="35" pageNumber="36" type="generic">three specimens</specimenCount>
were different in ossicle length and frequency (
<emphasis id="B911EA9BF9719D6FFE6121608DECA902" box="[405,455,1370,1396]" italics="true" pageId="35" pageNumber="36">APT</emphasis>
) of teethed perforations (
<emphasis id="B911EA9BF9719D6FFD1221608EDCA905" box="[742,759,1370,1395]" italics="true" pageId="35" pageNumber="36">P</emphasis>
s&lt;0.005, Kruskal-Wallis test), while the numbers of teethed perforations (
<emphasis id="B911EA9BF9719D6FFEC421448D73A9E1" box="[304,344,1406,1431]" italics="true" pageId="35" pageNumber="36">Ntp</emphasis>
) were not significantly different. Therefore, other than the number of teethed perforations (04) and noticeable plate skewness (
<emphasis id="B911EA9BF9719D6FFDF621988E19A9CA" box="[514,562,1442,1468]" italics="true" pageId="35" pageNumber="36">APS</emphasis>
=011.3), all the other morphological characters of the anchor plate ossicles cannot be the accurate keys.
</paragraph>
<paragraph id="8BDA3689F9719D6FFF3321D3883CAAC1" blockId="35.[151,1437,973,2008]" pageId="35" pageNumber="36">
Anchor ossicle lengths and calculated indicators ranged within: 140178 μm in length,
<emphasis id="B911EA9BF9719D6FFB3C21D088D5AA72" box="[1224,1278,1514,1540]" italics="true" pageId="35" pageNumber="36">ASW</emphasis>
=21.832.5%,
<emphasis id="B911EA9BF9719D6FFF6322348CFAAA5E" box="[151,209,1550,1576]" italics="true" pageId="35" pageNumber="36">AEW</emphasis>
=38.564.9% (
<tableCitation id="C6E70332F9719D6FFE8D22378DF9AA5E" box="[377,466,1549,1576]" captionStart="TABLE 9" captionStartId="32.[152,243,152,176]" captionTargetPageId="32" captionText="TABLE 9. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta parvaspiculus sp. nov." pageId="35" pageNumber="36">Table 9</tableCitation>
). Among these values, lengths of anchor ossicles (both ventrally and dorsally) were not significantly different (
<emphasis id="B911EA9BF9719D6FFE3C22088DCFAA3D" box="[456,484,1586,1611]" italics="true" pageId="35" pageNumber="36">Ps</emphasis>
&gt;0.05, Kruskal-Wallis test), also the indicator for anchor stem breadth
<emphasis id="B911EA9BF9719D6FFAF922088968AA3A" box="[1293,1347,1586,1612]" italics="true" pageId="35" pageNumber="36">ASW</emphasis>
and for basal / distal ends width
<emphasis id="B911EA9BF9719D6FFE45226C8DC0AA06" box="[433,491,1622,1648]" italics="true" pageId="35" pageNumber="36">AEW</emphasis>
were not significantly different (
<emphasis id="B911EA9BF9719D6FFC96226C8F55AA19" box="[866,894,1622,1647]" italics="true" pageId="35" pageNumber="36">Ps</emphasis>
&gt;0.05, Kruskal-Wallis test). From these results, the anchor ossicles in the body wall of the present
<specimenCount id="9D63FD00F9719D6FFD1422438FBDAAE5" box="[736,918,1657,1683]" count="3" pageId="35" pageNumber="36" type="generic">three specimens</specimenCount>
were not different in ossicle length or shape. Therefore, proportion and shape of the anchor ossicles can be the accurate keys.
</paragraph>
<paragraph id="8BDA3689F9719D6FFF3322FB8876ABF9" blockId="35.[151,1437,973,2008]" pageId="35" pageNumber="36">
Longitudinal muscle had both O-shaped and C-shaped granule ossicles, length and calculated indicators ranged within: 2154 μm in length,
<emphasis id="B911EA9BF9719D6FFE2722DC8DD3AB76" box="[467,504,1766,1792]" italics="true" pageId="35" pageNumber="36">GP</emphasis>
=28.280.8%,
<emphasis id="B911EA9BF9719D6FFD6922DC8EF3AB76" box="[669,728,1766,1792]" italics="true" pageId="35" pageNumber="36">GCO</emphasis>
=33.378.6% (
<tableCitation id="C6E70332F9719D6FFC8822DF8FF9AB76" box="[892,978,1765,1792]" captionStart="TABLE 9" captionStartId="32.[152,243,152,176]" captionTargetPageId="32" captionText="TABLE 9. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta parvaspiculus sp. nov." pageId="35" pageNumber="36">Table 9</tableCitation>
). Among these values, lengths of granule ossicles (both ventrally and dorsally) were significantly different (
<emphasis id="B911EA9BF9719D6FFC7123308F8AAB55" box="[901,929,1802,1827]" italics="true" pageId="35" pageNumber="36">Ps</emphasis>
&lt;0.005, Kruskal-Wallis test), also calculated indicator
<emphasis id="B911EA9BF9719D6FFEF723148D03AB3E" box="[259,296,1838,1864]" italics="true" pageId="35" pageNumber="36">GP</emphasis>
was significantly different (
<emphasis id="B911EA9BF9719D6FFD9D23148EAEAB31" box="[617,645,1838,1863]" italics="true" pageId="35" pageNumber="36">Ps</emphasis>
&lt;0.005, Kruskal-Wallis test), but
<emphasis id="B911EA9BF9719D6FFBF22314886AAB3E" box="[1030,1089,1838,1864]" italics="true" pageId="35" pageNumber="36">GCO</emphasis>
was not different significantly (
<emphasis id="B911EA9BF9719D6FFF5523688C96AB1D" box="[161,189,1874,1899]" italics="true" pageId="35" pageNumber="36">Ps</emphasis>
&gt;0.05, χ
<superScript id="7C109BC1F9719D6FFEE3236B8D34AB29" attach="left" box="[279,287,1873,1887]" fontSize="6" pageId="35" pageNumber="36">2</superScript>
-multiple test). Therefore, the length and proportion of the granule ossicles cannot be the accurate keys, while the ratio C- or O-shaped granule ossicles of this species can be the accurate key.
</paragraph>
<paragraph id="8BDA3689F9719D68FF3323A38816AD37" blockId="35.[151,1437,973,2008]" lastBlockId="36.[151,1437,150,321]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
Tentacle ossicles rods. Lengths and calculated indicators of
<specimenCount id="9D63FD00F9719D6FFC7023A38817ABC5" box="[900,1084,1945,1971]" count="3" pageId="35" pageNumber="36" type="generic">three specimens</specimenCount>
ranged within: 53102 μm in length,
<emphasis id="B911EA9BF9719D6FFF1F23848D26ABA1" box="[235,269,1982,2007]" italics="true" pageId="35" pageNumber="36">RP</emphasis>
=7.121.5%, and
<emphasis id="B911EA9BF9719D6FFE2723848DDCABAE" box="[467,503,1982,2008]" italics="true" pageId="35" pageNumber="36">RC</emphasis>
=2.717.8. Among these values, length and non-size dependence complexities (
<emphasis id="B911EA9BF9719D6FFA9B238489B8ABAE" box="[1391,1427,1982,2008]" italics="true" pageId="35" pageNumber="36">RC</emphasis>
) of rod ossicles were not significantly different (
<emphasis id="B911EA9BF9769D68FD5824AD8E96ACC6" box="[684,701,151,176]" italics="true" pageId="36" pageNumber="37">P</emphasis>
&gt;0.05, Kruskal-Wallis test), while non-size dependence indicator for proportion (
<emphasis id="B911EA9BF9769D68FEB224818D43ACA2" box="[326,360,187,212]" italics="true" pageId="36" pageNumber="37">RP</emphasis>
) was significantly different (
<emphasis id="B911EA9BF9769D68FD5B24818EEBACA2" box="[687,704,187,212]" italics="true" pageId="36" pageNumber="37">P</emphasis>
&lt;0.005, Kruskal-Wallis test). From these results, the rod ossicles in the tentacles of the present
<specimenCount id="9D63FD00F9769D68FE2924E58EBBAC8F" box="[477,656,223,249]" count="3" pageId="36" pageNumber="37" type="generic">three specimens</specimenCount>
have similar length, however their proportion was noticeably variable. Therefore, the length (about 50100 μm) and the high complexity of the rod ossicles shown by
<emphasis id="B911EA9BF9769D68FB4E253988F5AD6B" box="[1210,1246,259,285]" italics="true" pageId="36" pageNumber="37">RC</emphasis>
=2.717.8 of this species can be accurate keys, however their proportions cannot be the accurate key.
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9769D68FF6325A9897AAF4F" pageId="36" pageNumber="37" type="discussion">
<paragraph id="8BDA3689F9769D68FF6325A98D22ADDB" blockId="36.[151,265,403,429]" box="[151,265,403,429]" pageId="36" pageNumber="37">
<emphasis id="B911EA9BF9769D68FF6325A98D22ADDB" bold="true" box="[151,265,403,429]" pageId="36" pageNumber="37">Remarks</emphasis>
</paragraph>
<paragraph id="8BDA3689F9769D68FF6325E1897AAF4F" blockId="36.[151,1437,474,825]" pageId="36" pageNumber="37">
Among the present
<specimenCount id="9D63FD00F9769D68FE8C25E18E1AAD83" box="[376,561,475,501]" count="3" pageId="36" pageNumber="37" type="generic">three specimens</specimenCount>
, the smallest specimen (WMNH-2024-INV-156) had no &quot; two-tipped anchor plate&quot; in the body wall, that was deviation for one diagnostic character noted above, however, that simultaneously had other two diagnostic characters &quot;perforations of anchor plate ossicles mostly without teeth&quot; and &quot;outer edge of anchor plate equipped with minute teeth in small numbers, &quot;therefore, we concluded that the anchor plate of immature animals would be underdeveloped and without basal concave which divide the &quot;two-tips&quot;. Among the previously nominated six species of the genus
<taxonomicName id="4C654D0AF9769D68FD4326B58FE0AEDF" authority="Heding, 1928" authorityName="Heding" authorityYear="1928" box="[695,971,655,681]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="36" pageNumber="37" phylum="Echinodermata" rank="genus">
<emphasis id="B911EA9BF9769D68FD4326B58F0EAEDE" box="[695,805,655,680]" italics="true" pageId="36" pageNumber="37">Patinapta</emphasis>
<bibRefCitation id="EFF44B78F9769D68FCC426B58FE0AEDF" author="Heding, S. G." box="[816,971,655,681]" pageId="36" pageNumber="37" pagination="105 - 323" refId="ref31634" refString="Heding, S. G. (1928) Synaptidae. Papers from Dr. Th. Mortensen's Pacific Expedition 1914 - 1916. No. 66. Videnskabelige Meddeleser fra Dansk naturhistorisk Forening i KObenhavn, 85, 105 - 323." type="journal article" year="1928">Heding, 1928</bibRefCitation>
</taxonomicName>
(WoRMS 2024, listed in Table 7), only one species,
<taxonomicName id="4C654D0AF9769D68FED326898E6FAEBB" authority="(Bedford, 1899)" baseAuthorityName="Bedford" baseAuthorityYear="1899" box="[295,580,691,717]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="36" pageNumber="37" phylum="Echinodermata" rank="species" species="laevis">
<emphasis id="B911EA9BF9769D68FED326898DAFAEBB" box="[295,388,691,717]" italics="true" pageId="36" pageNumber="37">P. laevis</emphasis>
(
<bibRefCitation id="EFF44B78F9769D68FE6026898E17AEBB" author="Bedford, F. P." box="[404,572,691,717]" pageId="36" pageNumber="37" pagination="141 - 150" refId="ref31264" refString="Bedford, F. P. (1899) Holothurians. In: Willey, A. (Ed.), Zoological Results based on material from New Britain, New Guinea, Loyalty Islands and elsewhere, collected during the years 1895, 1896 and 1897. Part II. University Press, Cambridge, pp. 141 - 150." type="book chapter" year="1899">Bedford, 1899</bibRefCitation>
)
</taxonomicName>
, was reported as a species possessing toothless perforations on anchor plate ossicles (Table 7-2), which showed a concave on basal tip resulting in two tips basal end, also resembling the present new species
<taxonomicName id="4C654D0AF9769D68FEDC26C18DEEAF63" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[296,453,763,789]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="36" pageNumber="37" phylum="Echinodermata" rank="species" species="parvaspiculus" status="sp. nov.">
<emphasis id="B911EA9BF9769D68FEDC26C18DEEAF63" box="[296,453,763,789]" italics="true" pageId="36" pageNumber="37">parvaspiculus</emphasis>
</taxonomicName>
<emphasis id="B911EA9BF9769D68FE3926C18E03AF63" bold="true" box="[461,552,763,789]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="A22257E0F9769D68FE3926C18E03AF63" box="[461,552,763,789]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
However,
<taxonomicName id="4C654D0AF9769D68FD5226C18F2FAF63" baseAuthorityName="Bedford" baseAuthorityYear="1899" box="[678,772,763,789]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="36" pageNumber="37" phylum="Echinodermata" rank="species" species="laevis">
<emphasis id="B911EA9BF9769D68FD5226C18F2FAF63" box="[678,772,763,789]" italics="true" pageId="36" pageNumber="37">P. laevis</emphasis>
</taxonomicName>
also possesses unique ossicles in its longitudinal muscles, mostly two perforated oval granule ossicles, and thus is easily distinguishable from the present new species.
</paragraph>
</subSubSection>
<caption id="DF1A6601F9769D68FF6C2752885CAFF7" box="[152,1143,872,897]" pageId="36" pageNumber="37" startId="36.[152,244,872,896]" targetBox="[163,1405,913,1259]" targetIsTable="true" targetPageId="36" targetType="table">
<paragraph id="8BDA3689F9769D68FF6C2752885CAFF7" blockId="36.[152,1143,872,897]" box="[152,1143,872,897]" pageId="36" pageNumber="37">
<emphasis id="B911EA9BF9769D68FF6C27528D37AFF6" bold="true" box="[152,284,872,896]" pageId="36" pageNumber="37">TABLE 10.</emphasis>
List of WMNH-INV specimens of
<taxonomicName id="4C654D0AF9769D68FD6C27538FB5AFF6" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[664,926,872,896]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="36" pageNumber="37" phylum="Echinodermata" rank="species" species="parvaspiculus" status="sp. nov.">
<emphasis id="B911EA9BF9769D68FD6C27538FB5AFF6" box="[664,926,872,896]" italics="true" pageId="36" pageNumber="37">Patinapta parvaspiculus</emphasis>
</taxonomicName>
<emphasis id="B911EA9BF9769D68FC5027538FD1AFF7" bold="true" box="[932,1018,873,897]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="A22257E0F9769D68FC5027538FD1AFF7" box="[932,1018,873,897]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
as of 2024.
</paragraph>
</caption>
<paragraph id="8BDA3689F9769D68FF5727AB8919A89D" pageId="36" pageNumber="37">
<table id="F965C429F97662B3FF5727AB8956A89D" box="[163,1405,913,1259]" colsContinueFrom="31.[163,1406,193,458]" colsContinueIn="42.[163,1385,1453,1638]" gridcols="3" gridrows="7" pageId="36" pageNumber="37">
<tr id="355534CBF97662B3FF5727AB8956AFDF" box="[163,1405,913,937]" gridrow="0" pageId="36" pageNumber="37">
<th id="76845DB7F97662B3FF5727AB8D47AFDF" box="[163,364,913,937]" gridcol="0" gridrow="0" pageId="36" pageNumber="37">WMNH-INV No.</th>
<th id="76845DB7F97662B3FE3227AB8FE4AFDF" box="[454,975,913,937]" gridcol="1" gridrow="0" pageId="36" pageNumber="37">Locality (arranged under north to south)</th>
<th id="76845DB7F97662B3FBFF27AB8956AFDF" box="[1035,1405,913,937]" gridcol="2" gridrow="0" pageId="36" pageNumber="37">Collectors (Date)</th>
</tr>
<tr id="355534CBF97662B3FF5727838956AFA7" box="[163,1405,953,977]" gridrow="1" pageId="36" pageNumber="37">
<th id="76845DB7F97662B3FF5727838956AFA7" box="[163,1405,953,977]" colspan="3" colspanRight="2" gridcol="0" gridrow="1" pageId="36" pageNumber="37">Kyushu Island</th>
</tr>
<tr id="355534CBF97662B3FF5727DB8956A854" box="[163,1405,993,1058]" gridrow="2" pageId="36" pageNumber="37">
<th id="76845DB7F97662B3FF5727DB8D47A854" box="[163,364,993,1058]" gridcol="0" gridrow="2" pageId="36" pageNumber="37">2024-INV-152 2024-INV-156, 157</th>
<td id="76845DB7F97662B3FE3227DB8FE4A854" box="[454,975,993,1058]" gridcol="1" gridrow="2" pageId="36" pageNumber="37">Yasaka-gawa, Kitsuki City, Oita Imari Bay, Imari City, Saga</td>
<td id="76845DB7F97662B3FBFF27DB8956A854" box="[1035,1405,993,1058]" gridcol="2" gridrow="2" pageId="36" pageNumber="37">Nishikawa, T. (July 2, 2004) Nishikawa, T. (April 24, 2002)</td>
</tr>
<tr id="355534CBF97662B3FF5720088956A83C" box="[163,1405,1074,1098]" gridrow="3" pageId="36" pageNumber="37">
<th id="76845DB7F97662B3FF5720088956A83C" box="[163,1405,1074,1098]" colspan="3" colspanRight="2" gridcol="0" gridrow="3" pageId="36" pageNumber="37">Shikoku Island</th>
</tr>
<tr id="355534CBF97662B3FF5720608956A804" box="[163,1405,1114,1138]" gridrow="4" pageId="36" pageNumber="37">
<th id="76845DB7F97662B3FF5720608D47A804" box="[163,364,1114,1138]" gridcol="0" gridrow="4" pageId="36" pageNumber="37">2024-INV-162163</th>
<td id="76845DB7F97662B3FE3220608FE4A804" box="[454,975,1114,1138]" gridcol="1" gridrow="4" pageId="36" pageNumber="37">Uranouchi Bay, Suzaki City, Kochi</td>
<td id="76845DB7F97662B3FBFF20608956A804" box="[1035,1405,1114,1138]" gridcol="2" gridrow="4" pageId="36" pageNumber="37">Nishikawa, T. (September 27, 2003)</td>
</tr>
<tr id="355534CBF97662B3FF5720B88956A8EC" box="[163,1405,1154,1178]" gridrow="5" pageId="36" pageNumber="37">
<th id="76845DB7F97662B3FF5720B88956A8EC" box="[163,1405,1154,1178]" colspan="3" colspanRight="2" gridcol="0" gridrow="5" pageId="36" pageNumber="37">Ryukyu Islands</th>
</tr>
<tr id="355534CBF97662B3FF5720908956A89D" box="[163,1405,1194,1259]" gridrow="6" pageId="36" pageNumber="37">
<th id="76845DB7F97662B3FF5720908D47A89D" box="[163,364,1194,1259]" gridcol="0" gridrow="6" pageId="36" pageNumber="37">2024-INV-181, 182 2015-INV-108</th>
<td id="76845DB7F97662B3FE3220908FE4A89D" box="[454,975,1194,1259]" gridcol="1" gridrow="6" pageId="36" pageNumber="37">Urauchi Bay, Kamikoshiki-jima Island, Kagoshima Akasaki beach, Yoron-jima Island, Kagoshima</td>
<td id="76845DB7F97662B3FBFF20908956A89D" box="[1035,1405,1194,1259]" gridcol="2" gridrow="6" pageId="36" pageNumber="37">Nishikawa, T. (March 15, 2003) S. M. (May 28, 1994)</td>
</tr>
</table>
</paragraph>
<subSubSection id="C37F6502F9769D68FF6321048822AA22" pageId="36" pageNumber="37" type="distribution">
<paragraph id="8BDA3689F9769D68FF6321048D1BA92E" blockId="36.[151,304,1342,1368]" box="[151,304,1342,1368]" pageId="36" pageNumber="37">
<emphasis id="B911EA9BF9769D68FF6321048D1BA92E" bold="true" box="[151,304,1342,1368]" pageId="36" pageNumber="37">Distribution</emphasis>
</paragraph>
<paragraph id="8BDA3689F9769D68FF6321BC8822AA22" blockId="36.[151,1436,1414,1621]" pageId="36" pageNumber="37">
Southern
<collectingCountry id="F3727619F9769D68FEF321BD8D6CA9D7" box="[263,327,1415,1441]" name="Japan" pageId="36" pageNumber="37">Japan</collectingCountry>
(Kyushu Island, Shikoku Island and the Ryukyu Islands). In Yoron-jima Island (
<typeStatus id="54DE882BF9769D68FB0F21BC8906A9D6" box="[1275,1325,1414,1440]" pageId="36" pageNumber="37">type</typeStatus>
locality), the specimens were collected from intertidal sandy beach with boulders originated from dead corals. Among the specimens housed in WMNH-INV, including the present specimens, 5 bottles including
<specimenCount id="9D63FD00F9769D68FB9221F488C4A99E" box="[1126,1263,1486,1512]" count="8" pageId="36" pageNumber="37" type="generic">8 specimens</specimenCount>
were identified as this species (
<tableCitation id="C6E70332F9769D68FEB021C88D81AA7A" box="[324,426,1522,1548]" captionStart="TABLE 10" captionStartId="36.[152,244,872,896]" captionTargetPageId="36" captionText="TABLE 10. List of WMNH-INV specimens of Patinapta parvaspiculus sp. nov. as of 2024." pageId="36" pageNumber="37">Table 10</tableCitation>
). Other than the
<typeStatus id="54DE882BF9769D68FD9121C98EBEAA7B" box="[613,661,1523,1549]" pageId="36" pageNumber="37">type</typeStatus>
locality, 6 young specimens collected from
<collectingRegion id="49A1F86BF9769D68FB7721C88898AA7A" box="[1155,1203,1522,1548]" country="Japan" name="Oita" pageId="36" pageNumber="37">Oita</collectingRegion>
and
<collectingRegion id="49A1F86BF9769D68FB1D21C88934AA7A" box="[1257,1311,1522,1548]" country="Japan" name="Saga" pageId="36" pageNumber="37">Saga</collectingRegion>
of Kyushu Island, from
<collectingRegion id="49A1F86BF9769D68FED2222C8D41AA46" box="[294,362,1558,1584]" country="Japan" name="Kochi" pageId="36" pageNumber="37">Kochi</collectingRegion>
of Shikoku Island, and from two islands of
<collectingRegion id="49A1F86BF9769D68FCA5222C8FFAAA46" box="[849,977,1558,1584]" country="Japan" name="Kagoshima" pageId="36" pageNumber="37">Kagoshima</collectingRegion>
in Ryukyu Islands,
<collectingCountry id="F3727619F9769D68FB5A222D88C5AA47" box="[1198,1262,1559,1585]" name="Japan" pageId="36" pageNumber="37">Japan</collectingCountry>
were identified as this species. All the specimens were collected from estuarine intertidal flats.
</paragraph>
</subSubSection>
</treatment>
</document>