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<document id="12018CE1560C37C994507716C39D87D1" ID-CLB-Dataset="59554" ID-DOI="10.5852/ejt.2023.885.2189" ID-GBIF-Dataset="f8509ab7-b884-41aa-a5ee-f8c425a46031" ID-ISSN="2118-9773" ID-Zenodo-Dep="8205500" ID-ZooBank="444CE1D2-8098-4CDE-9B97-927CA13FC39F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1690898759111" checkinUser="felipe" docAuthor="Means, Jackson C., Bouzan, Rodrigo S., Iniesta, Luiz Felipe Moretti, Martínez-Torres, Daniela, Vasquez-Valverde, Luisa F., Brescovit, Antonio D. &amp; Ivanov, Kaloyan" docDate="2023" docId="03CC1013FFCB3C7C97D8F93231E7FB4B" docLanguage="en" docName="ejt-885_65-85.pdf" docOrigin="European Journal of Taxonomy 885 (1)" docSource="http://dx.doi.org/10.5852/ejt.2023.885.2189" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dibolostethus inopinatus Means, Bouzan &amp; Ivanov 2023, sp. nov." docType="treatment" docUuid="8C1D05D6-D758-4825-9BDF-69E642BA219D" docUuidSource="ZooBank" docVersion="6" lastPageNumber="76" masterDocId="FFF5686BFFC23C77966FFF973518FFDD" masterDocTitle="A review of the previously monotypic tribe Dibolostethini (Chelodesmidae: Chelodesminae) with description of two new species and a summary of the Chelodesmidae of the Tropical Andes Biodiversity Hotspot" masterLastPageNumber="85" masterPageNumber="65" pageNumber="74" updateTime="1699550199751" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="E37FCE44501114F1793F7F8530FB6C17">A review of the previously monotypic tribe Dibolostethini (Chelodesmidae: Chelodesminae) with description of two new species and a summary of the Chelodesmidae of the Tropical Andes Biodiversity Hotspot</mods:title>
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<mods:namePart id="4C55D5B68718B1D566B181F8A8DDA505">Means, Jackson C.</mods:namePart>
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<mods:namePart id="305694FC87C9766030B8283E40B2573B">Bouzan, Rodrigo S.</mods:namePart>
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<mods:affiliation id="B49B75EACE6F9DEAE8CEDC8234132A33">Pós-graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, Brazil. &amp; Laboratório Especial de Coleções Zoológicas, Instituto Butantan, Avenida Vital Brasil, 1500, 05503 - 090, São Paulo, Brazil.</mods:affiliation>
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<mods:namePart id="D33BA02EB19AFFC9979D06788D57B828">Iniesta, Luiz Felipe Moretti</mods:namePart>
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<mods:namePart id="E087E7D228F6580665632D7DB65D769A">Vasquez-Valverde, Luisa F.</mods:namePart>
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<mods:affiliation id="8650ED3460FCDA5CD00870CEFCEEF5BF">Department of Entomology, Virginia Tech, 216 A Price Hall, Blacksburg, VA 24061, USA.</mods:affiliation>
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<taxonomicName id="4C65DA86FFCB3C7E97D8F9323102F91D" authority="Means, Bouzan &amp; Ivanov" authorityName="Means, Bouzan &amp; Ivanov" authorityYear="2023" box="[439,1050,1701,1728]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="74" phylum="Arthropoda" rank="species" species="inopinatus" status="sp. nov.">
<emphasis id="B9117D17FFCB3C7E97D8F93237FEF962" bold="true" box="[439,742,1701,1728]" italics="true" pageId="9" pageNumber="74">Dibolostethus inopinatus</emphasis>
Means, Bouzan &amp; Ivanov
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<figureCitation id="135EBD80FFCB3C7E94D3F97B3627F8DB" box="[700,831,1772,1798]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[190,1352,1052,1886]" captionTargetId="figure-271@8.[189,1398,1044,1886]" captionTargetPageId="8" captionText="Fig. 4. Body ring 4, sternal projections. A–B. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810). C–D. D. inopinatus Means, Bouzan &amp; Ivanov sp. nov., holotype, ♂ (VMNH110813). A, C. Lateral views. B, D. Posterolateral views. Abbreviation: Cx = coxae." figureDoi="http://doi.org/10.5281/zenodo.8205508" httpUri="https://zenodo.org/record/8205508/files/figure.png" pageId="9" pageNumber="74">Figs 4C–D</figureCitation>
, 
<figureCitation id="135EBD80FFCB3C7E9524F97B3644F8DB" box="[843,860,1772,1798]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[189,1398,1270,1881]" captionTargetId="figure-240@10.[189,1398,1266,1881]" captionTargetPageId="10" captionText="Fig. 6. Dibolostethus inopinatus Means, Bouzan &amp; Ivanov sp. nov., holotype, ♂ (VMNH110813), left gonopod. A. Lateral view, red circle = cingulum. B. Anterior view. C. Mesal view. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." figureDoi="http://doi.org/10.5281/zenodo.8205512" httpUri="https://zenodo.org/record/8205512/files/figure.png" pageId="9" pageNumber="74">6</figureCitation>
, 
<figureCitation id="135EBD80FFCB3C7E9507F97B366FF8DB" box="[872,887,1772,1798]" captionStart="Fig" captionStartId="14.[188,231,1892,1918]" captionTargetBox="[243,1357,497,1838]" captionTargetId="figure-72@14.[212,1380,484,1853]" captionTargetPageId="14" captionText="Fig. 9. Distribution of Chelodesmidae spp. in the Tropical Andes Biodiversity Hotspot, by country. Note: the type locality of Dibolostethus sicarius Hoffman, 2009 is included for completeness although it falls just outside of the boundaries of the Tropical Andes Biodiversity Hotspot. For information on these taxa see Supp. file 1." figureDoi="http://doi.org/10.5281/zenodo.8205518" httpUri="https://zenodo.org/record/8205518/files/figure.png" pageId="9" pageNumber="74">9</figureCitation>
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<treatmentCitationGroup id="AB75862BFFCB3C7E96D2F8B9369CF89A" box="[189,900,1837,1864]" pageId="9" pageNumber="74">
<taxonomicName id="4C65DA86FFCB3C7E96D2F8B93426F89A" ID-CoL="35GSJ" authorityName="Hoffman" authorityYear="2009" box="[189,318,1838,1863]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="74" phylum="Arthropoda" rank="species" species="sicarius">
<emphasis id="B9117D17FFCB3C7E96D2F8B93426F89A" box="[189,318,1838,1863]" italics="true" pageId="9" pageNumber="74">D. sicarius</emphasis>
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(in part) – 
<treatmentCitation id="0AC48714FFCB3C7E97ADF8BA37B3F895" author="Hoffman R. L." box="[450,683,1837,1864]" page="151" pageId="9" pageNumber="74" year="2009">
<bibRefCitation id="EFF4DCF4FFCB3C7E97ADF8BA37B3F895" author="Hoffman" box="[450,683,1837,1864]" firstAuthor="Hoffman" page="151" pageId="9" pageNumber="74" pagination="149 - 154" refId="ref9561" refString="Hoffman R. L. 2009. Chelodesmid studies. XXVI. A new genus and tribe of chelodesmid millipeds from Ecuador (Polydesmida: Chelodesmidae). International Journal of Myriapodology 2: 149 - 154. https: // doi. org / 10.1163 / 187525409 X 12577705044584" type="journal article" year="2009">Hoffman 2009: 151</bibRefCitation>
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, misidentification.
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<emphasis id="B9117D17FFCB3C7E96D2F8F9342EF855" bold="true" box="[189,310,1902,1928]" pageId="9" pageNumber="74">Diagnosis</emphasis>
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Adult males of 
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<emphasis id="B9117D17FFCB3C7E971DF809349FF86A" box="[370,391,1950,1975]" italics="true" pageId="9" pageNumber="74">D</emphasis>
. 
<emphasis id="B9117D17FFCB3C7E97F9F809370AF86A" box="[406,530,1950,1975]" italics="true" pageId="9" pageNumber="74">inopinatus</emphasis>
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differ from those of 
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<emphasis id="B9117D17FFCB3C7E9509F80936C3F86A" box="[870,987,1949,1975]" italics="true" pageId="9" pageNumber="74">D. kattani</emphasis>
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<taxonomicNameLabel id="A222C06CFFCB3C7E958DF8093125F865" box="[994,1085,1950,1976]" pageId="9" pageNumber="74" rank="species">sp. nov.</taxonomicNameLabel>
based on the lack of short, blunt projections on the sternite of the 5 
<superScript id="7C100C4DFFCB3C7E94C4F85737A0F813" attach="left" box="[683,696,1984,1998]" fontSize="6" pageId="9" pageNumber="74">th</superScript>
body ring, and from 
<taxonomicName id="4C65DA86FFCB3C7E95ACF856315FF807" authorityName="Hoffman" authorityYear="2009" box="[963,1095,1985,2010]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="74" phylum="Arthropoda" rank="species" species="sicarius">
<emphasis id="B9117D17FFCB3C7E95ACF856315FF807" box="[963,1095,1985,2010]" italics="true" pageId="9" pageNumber="74">D. sicarius</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C65DA86FFCB3C7E92E7F8563018F807" authorityName="Means &amp; Bouzan &amp; Iniesta &amp; Martínez-Torres &amp; Vasquez-Valverde &amp; Brescovit &amp; Ivanov" authorityYear="2023" box="[1160,1280,1984,2010]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="74" phylum="Arthropoda" rank="species" species="kattani">
<emphasis id="B9117D17FFCB3C7E92E7F8563018F807" box="[1160,1280,1984,2010]" italics="true" pageId="9" pageNumber="74">D. kattani</emphasis>
</taxonomicName>
based on the following combination of gonopodal characteristics: solenomere sinuous before curving dorsally at nearly 90° at apex, outer edge with wide lateral flange just before apex, tip curving laterally (
<figureCitation id="135EBD80FFC83C7D937CFEBB35CCFEB4" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[189,1398,1270,1881]" captionTargetId="figure-240@10.[189,1398,1266,1881]" captionTargetPageId="10" captionText="Fig. 6. Dibolostethus inopinatus Means, Bouzan &amp; Ivanov sp. nov., holotype, ♂ (VMNH110813), left gonopod. A. Lateral view, red circle = cingulum. B. Anterior view. C. Mesal view. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." figureDoi="http://doi.org/10.5281/zenodo.8205512" httpUri="https://zenodo.org/record/8205512/files/figure.png" pageId="10" pageNumber="75">Fig. 6A, C</figureCitation>
); prefemoral process only slightly narrower than solenomere at base, with pronounced 45° angle on dorsal side, expanding at midpoint with lateral and medial edges becoming pronounced ridges widening into lamellate flanges on either side of prefemoral process; apex of prefemoral process acute, directed anteriad (
<figureCitation id="135EBD80FFC83C7D9743FE2E3490FE0E" box="[300,392,441,467]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[189,1398,1270,1881]" captionTargetId="figure-240@10.[189,1398,1266,1881]" captionTargetPageId="10" captionText="Fig. 6. Dibolostethus inopinatus Means, Bouzan &amp; Ivanov sp. nov., holotype, ♂ (VMNH110813), left gonopod. A. Lateral view, red circle = cingulum. B. Anterior view. C. Mesal view. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." figureDoi="http://doi.org/10.5281/zenodo.8205512" httpUri="https://zenodo.org/record/8205512/files/figure.png" pageId="10" pageNumber="75">Fig. 6B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37FF28EFFC83C7D96D2FE6C31FFFDBB" pageId="10" pageNumber="75" type="etymology">
<paragraph id="8BDAA105FFC83C7D96D2FE6C345CFDC8" blockId="10.[189,324,507,533]" box="[189,324,507,533]" pageId="10" pageNumber="75">
<heading id="D0921669FFC83C7D96D2FE6C345CFDC8" bold="true" box="[189,324,507,533]" fontSize="11" level="3" pageId="10" pageNumber="75" reason="3">
<emphasis id="B9117D17FFC83C7D96D2FE6C345CFDC8" bold="true" box="[189,324,507,533]" pageId="10" pageNumber="75">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BDAA105FFC83C7D96D2FDBE31FFFDBB" blockId="10.[189,1398,553,615]" pageId="10" pageNumber="75">
The species epithet ‘ 
<taxonomicName id="4C65DA86FFC83C7D97C8FDBD373BFD9E" authorityName="Means &amp; Bouzan &amp; Iniesta &amp; Martínez-Torres &amp; Vasquez-Valverde &amp; Brescovit &amp; Ivanov" authorityYear="2023" box="[423,547,554,579]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="75" phylum="Arthropoda" rank="species" species="inopinatus">
<emphasis id="B9117D17FFC83C7D97C8FDBD373BFD9E" box="[423,547,554,579]" italics="true" pageId="10" pageNumber="75">inopinatus</emphasis>
</taxonomicName>
’ is Latin for ‘unexpected’, ‘unforeseen’, reflecting its surprising discovery among the 
<typeStatus id="54DE1FA7FFC83C7D9750FDDA346AFDBA" box="[319,370,589,615]" pageId="10" pageNumber="75">type</typeStatus>
series of 
<taxonomicName id="4C65DA86FFC83C7D978CFDDA377CFDBB" authorityName="Hoffman" authorityYear="2009" box="[483,612,589,614]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="75" phylum="Arthropoda" rank="species" species="sicarius">
<emphasis id="B9117D17FFC83C7D978CFDDA377CFDBB" box="[483,612,589,614]" italics="true" pageId="10" pageNumber="75">D. sicarius</emphasis>
</taxonomicName>
. The species epithet is a masculine, singular adjective.
</paragraph>
</subSubSection>
<subSubSection id="C37FF28EFFC83C7D96D2FD193753FC9D" pageId="10" pageNumber="75" type="materials_examined">
<paragraph id="8BDAA105FFC83C7D96D2FD193474FD7A" blockId="10.[189,364,653,680]" box="[189,364,653,680]" pageId="10" pageNumber="75">
<heading id="D0921669FFC83C7D96D2FD193474FD7A" bold="true" box="[189,364,653,680]" fontSize="11" level="3" pageId="10" pageNumber="75" reason="3">
<emphasis id="B9117D17FFC83C7D96D2FD193474FD7A" bold="true" box="[189,364,653,680]" pageId="10" pageNumber="75">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BDAA105FFC83C7D9685FD2B3753FC9D" blockId="10.[189,1399,700,832]" pageId="10" pageNumber="75">
<materialsCitation id="3B0DAB58FFC83C7D9685FD2B3753FC9D" ID-GBIF-Occurrence="4167732302" collectingDate="1976-07-22" collectionCode="VMNH" country="Ecuador" latitude="-3.0666666" location="Cueva de Los Tayos" longLatPrecision="1307" longitude="-78.25" municipality="Los Tayos" pageId="10" pageNumber="75" specimenCode="VMNH110813" specimenCount="1" specimenCount-male="1" stateProvince="Cordillera del Condor" typeStatus="holotype">
<emphasis id="B9117D17FFC83C7D9685FD2B3444FD0B" bold="true" box="[234,348,700,726]" pageId="10" pageNumber="75">
<typeStatus id="54DE1FA7FFC83C7D9685FD2B3444FD0B" box="[234,348,700,726]" pageId="10" pageNumber="75" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F372E195FFC83C7D96D2FD483453FD24" box="[189,331,735,761]" name="Ecuador" pageId="10" pageNumber="75">ECUADOR</collectingCountry>
• 
<specimenCount id="9D636A8CFFC83C7D970FFD77346EFD27" box="[352,374,736,762]" pageId="10" pageNumber="75" type="male">♂</specimenCount>
; 
<collectingRegion id="49A16FE7FFC83C7D97EBFD48379CFD24" box="[388,644,735,761]" pageId="10" pageNumber="75">Cordillera del Cóndor</collectingRegion>
, 
<collectingRegion id="49A16FE7FFC83C7D94E2FD4836D3FD24" box="[653,971,735,761]" country="Ecuador" name="Morona-Santiago" pageId="10" pageNumber="75">Morona-Santiago Province</collectingRegion>
, 
<collectingMunicipality id="6BBE3B7FFFC83C7D95BAFD773155FD27" box="[981,1101,736,762]" pageId="10" pageNumber="75">
Los 
<taxonomicName id="4C65DA86FFC83C7D9268FD773155FD27" box="[1031,1101,736,762]" class="Arachnida" family="Hubbardiidae" genus="Tayos" kingdom="Animalia" order="Schizomida" pageId="10" pageNumber="75" phylum="Arthropoda" rank="genus">Tayos</taxonomicName>
</collectingMunicipality>
[vicinity of 
<location id="8EBAF7DEFFC83C7D92B6FD483411FCC0" LSID="urn:lsid:plazi:treatment:03CC1013FFCB3C7C97D8F93231E7FB4B:8EBAF7DEFFC83C7D92B6FD483411FCC0" country="Ecuador" latitude="-3.0666666" longLatPrecision="1307" longitude="-78.25" municipality="Los Tayos" name="Cueva de Los Tayos" pageId="10" pageNumber="75" stateProvince="Cordillera del Condor">
Cueva de Los 
<taxonomicName id="4C65DA86FFC83C7D96D2FC943411FCC0" box="[189,265,771,797]" class="Arachnida" family="Hubbardiidae" genus="Tayos" kingdom="Animalia" order="Schizomida" pageId="10" pageNumber="75" phylum="Arthropoda" rank="genus">Tayos</taxonomicName>
</location>
]; 
<geoCoordinate id="EE51C7C2FFC83C7D9772FC96346EFCC0" box="[285,374,769,797]" degrees="3" direction="south" minutes="04" orientation="latitude" pageId="10" pageNumber="75" precision="925" value="-3.0666666">3°04′ S</geoCoordinate>
, 
<geoCoordinate id="EE51C7C2FFC83C7D97EBFC9634ECFCC0" box="[388,500,769,797]" degrees="78" direction="west" minutes="15" orientation="longitude" pageId="10" pageNumber="75" precision="925" value="-78.25">78°15′ W</geoCoordinate>
[originally reported incorrectly in decimal degrees]; 
<date id="FFDB87C5FFC83C7D9203FC94301AFCC1" box="[1132,1282,770,797]" pageId="10" pageNumber="75" value="1976-07-22">
<collectingDate id="EF9F7E2DFFC83C7D9203FC94301AFCC1" box="[1132,1282,770,797]" pageId="10" pageNumber="75" value="1976-07-22">22 Jul. 1976</collectingDate>
</date>
; collector unknown; on log; 
<specimenCode id="DBC3097EFFC83C7D97FCFCB13750FC9D" box="[403,584,806,832]" pageId="10" pageNumber="75">VMNH110813</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37FF28EFFC83C7C96D2FCF03420FCF7" lastPageId="11" lastPageNumber="76" pageId="10" pageNumber="75" type="description">
<paragraph id="8BDAA105FFC83C7D96D2FCF03456FC5C" blockId="10.[189,334,871,897]" box="[189,334,871,897]" pageId="10" pageNumber="75">
<heading id="D0921669FFC83C7D96D2FCF03456FC5C" bold="true" box="[189,334,871,897]" fontSize="11" level="3" pageId="10" pageNumber="75" reason="3">
<emphasis id="B9117D17FFC83C7D96D2FCF03456FC5C" bold="true" box="[189,334,871,897]" pageId="10" pageNumber="75">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BDAA105FFC83C7D9685FC013772FC6D" blockId="10.[189,1399,918,1155]" box="[234,618,918,944]" pageId="10" pageNumber="75">
<emphasis id="B9117D17FFC83C7D9685FC013432FC6D" bold="true" box="[234,298,918,944]" pageId="10" pageNumber="75">Male</emphasis>
(
<typeStatus id="54DE1FA7FFC83C7D9756FC0134BCFC6D" box="[313,420,918,944]" pageId="10" pageNumber="75" type="holotype">holotype</typeStatus>
, VMNH110813)
</paragraph>
<paragraph id="8BDAA105FFC83C7D96D2FC2E340FFB5E" blockId="10.[189,1399,918,1155]" pageId="10" pageNumber="75">
With characteristics typical for the genus. Coloration (preserved in 70% isopropanol): body rings, all appendages, and telson uniformly light yellow. Head: epicranial macrosetae 2–2, interantennal macrosetae 1–1, frontal macrosetae 2–2. Sternites: 4 
<superScript id="7C100C4DFFC83C7D9525FC68364FFBD0" attach="left" box="[842,855,1023,1037]" fontSize="6" pageId="10" pageNumber="75">th</superScript>
body ring with two slender, conical, acute projections, appressed along entire length (
<figureCitation id="135EBD80FFC83C7D94ADFBB5365FFBE0" box="[706,839,1058,1085]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[190,1352,1052,1886]" captionTargetId="figure-271@8.[189,1398,1044,1886]" captionTargetPageId="8" captionText="Fig. 4. Body ring 4, sternal projections. A–B. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810). C–D. D. inopinatus Means, Bouzan &amp; Ivanov sp. nov., holotype, ♂ (VMNH110813). A, C. Lateral views. B, D. Posterolateral views. Abbreviation: Cx = coxae." figureDoi="http://doi.org/10.5281/zenodo.8205508" httpUri="https://zenodo.org/record/8205508/files/figure.png" pageId="10" pageNumber="75">Fig. 4C, D</figureCitation>
). Gonopod aperture on 7 
<superScript id="7C100C4DFFC83C7D921AFBB5319AFBED" attach="left" box="[1141,1154,1058,1072]" fontSize="6" pageId="10" pageNumber="75">th</superScript>
body ring oval, rim notably projecting above sternite (
<quantity id="4C9D0CE0FFC83C7D9437FBD137D1FBBD" box="[600,713,1094,1120]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="10" pageNumber="75" unit="mm" value="0.48">0.48 mm</quantity>
), with a deep U-shaped concavity (
<quantity id="4C9D0CE0FFC83C7D921AFBD131CDFBBD" box="[1141,1237,1094,1120]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="75" unit="mm" value="0.2">0.2 mm</quantity>
) on posterior margin.
</paragraph>
<caption id="DF1AF18DFFC83C7D96D2F81F312CF835" ID-DOI="http://doi.org/10.5281/zenodo.8205512" ID-Zenodo-Dep="8205512" httpUri="https://zenodo.org/record/8205512/files/figure.png" pageId="10" pageNumber="75" startId="10.[189,232,1928,1954]" targetBox="[189,1398,1270,1881]" targetPageId="10">
<paragraph id="8BDAA105FFC83C7D96D2F81F312CF835" blockId="10.[189,1399,1928,2025]" pageId="10" pageNumber="75">
<emphasis id="B9117D17FFC83C7D96D2F81F3414F87F" bold="true" box="[189,268,1928,1954]" pageId="10" pageNumber="75">Fig. 6.</emphasis>
<taxonomicName id="4C65DA86FFC83C7D977AF81F366DF87F" authority="Means, Bouzan &amp; Ivanov" authorityName="Means, Bouzan &amp; Ivanov" authorityYear="2023" box="[277,885,1928,1954]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="75" phylum="Arthropoda" rank="species" species="inopinatus" status="sp. nov.">
<emphasis id="B9117D17FFC83C7D977AF81F3724F87F" box="[277,572,1928,1954]" italics="true" pageId="10" pageNumber="75">Dibolostethus inopinatus</emphasis>
Means, Bouzan &amp; Ivanov
</taxonomicName>
<taxonomicNameLabel id="A222C06CFFC83C7D9512F81F36C3F87F" box="[893,987,1928,1954]" pageId="10" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (VMNH110813), left gonopod. 
<emphasis id="B9117D17FFC83C7D975AF83C3452F818" bold="true" box="[309,330,1963,1989]" pageId="10" pageNumber="75">A</emphasis>
. Lateral view, red circle = cingulum. 
<emphasis id="B9117D17FFC83C7D9571F83C362AF818" bold="true" box="[798,818,1963,1989]" pageId="10" pageNumber="75">B</emphasis>
. Anterior view. 
<emphasis id="B9117D17FFC83C7D9593F83C3109F818" bold="true" box="[1020,1041,1963,1989]" pageId="10" pageNumber="75">C</emphasis>
. Mesal view. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere.
</paragraph>
</caption>
<paragraph id="8BDAA105FFC93C7C96D2FE9E3152FE2B" blockId="11.[189,1399,265,503]" pageId="11" pageNumber="76">
<smallCapsWord id="8D3C37D9FFC93C7C96D2FE9E342CFEFC" baselines="284,284" box="[189,308,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Gonopods" pageId="11" pageNumber="76">GONOPODS</smallCapsWord>
(
<figureCitation id="135EBD80FFC93C7C9729FE9E3496FEFE" box="[326,398,265,291]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[189,1398,1270,1881]" captionTargetId="figure-240@10.[189,1398,1266,1881]" captionTargetPageId="10" captionText="Fig. 6. Dibolostethus inopinatus Means, Bouzan &amp; Ivanov sp. nov., holotype, ♂ (VMNH110813), left gonopod. A. Lateral view, red circle = cingulum. B. Anterior view. C. Mesal view. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." figureDoi="http://doi.org/10.5281/zenodo.8205512" httpUri="https://zenodo.org/record/8205512/files/figure.png" pageId="11" pageNumber="76">Fig. 6</figureCitation>
). Gonocoxae rounded, equivalent to about three quarters the length of the telopodite; with two macrosetae on the dorsum. Cannula appearing larger than in 
<taxonomicName id="4C65DA86FFC93C7C927EFEBA318CFE9B" authorityName="Hoffman" authorityYear="2009" box="[1041,1172,301,326]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="76" phylum="Arthropoda" rank="species" species="sicarius">
<emphasis id="B9117D17FFC93C7C927EFEBA318CFE9B" box="[1041,1172,301,326]" italics="true" pageId="11" pageNumber="76">D. sicarius</emphasis>
</taxonomicName>
. Solenomere long, sinuous, unbranched, apex falciform and tip curving laterally (
<figureCitation id="135EBD80FFC93C7C95D0FED83112FEB4" box="[959,1034,335,361]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[189,1398,1270,1881]" captionTargetId="figure-240@10.[189,1398,1266,1881]" captionTargetPageId="10" captionText="Fig. 6. Dibolostethus inopinatus Means, Bouzan &amp; Ivanov sp. nov., holotype, ♂ (VMNH110813), left gonopod. A. Lateral view, red circle = cingulum. B. Anterior view. C. Mesal view. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." figureDoi="http://doi.org/10.5281/zenodo.8205512" httpUri="https://zenodo.org/record/8205512/files/figure.png" pageId="11" pageNumber="76">Fig. 6</figureCitation>
, S). Prefemoral process: long (similar in length to solenomere), narrow at base (width subequal to solenomere), widening at halfway point with lateral and medial flanges resembling a hooded cobra in lateral and medial views (
<figureCitation id="135EBD80FFC93C7C937BFE0135CCFE0E" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[189,1398,1270,1881]" captionTargetId="figure-240@10.[189,1398,1266,1881]" captionTargetPageId="10" captionText="Fig. 6. Dibolostethus inopinatus Means, Bouzan &amp; Ivanov sp. nov., holotype, ♂ (VMNH110813), left gonopod. A. Lateral view, red circle = cingulum. B. Anterior view. C. Mesal view. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." figureDoi="http://doi.org/10.5281/zenodo.8205512" httpUri="https://zenodo.org/record/8205512/files/figure.png" pageId="11" pageNumber="76">Fig. 6A, C</figureCitation>
); medial flange with acute projection directed ventrally; prefemoral process and solenomere together resembling blacksmithing bolt tongs in lateral and medial views (
<figureCitation id="135EBD80FFC93C7C95D3FE4B3125FE2B" box="[956,1085,476,502]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[189,1398,1270,1881]" captionTargetId="figure-240@10.[189,1398,1266,1881]" captionTargetPageId="10" captionText="Fig. 6. Dibolostethus inopinatus Means, Bouzan &amp; Ivanov sp. nov., holotype, ♂ (VMNH110813), left gonopod. A. Lateral view, red circle = cingulum. B. Anterior view. C. Mesal view. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." figureDoi="http://doi.org/10.5281/zenodo.8205512" httpUri="https://zenodo.org/record/8205512/files/figure.png" pageId="11" pageNumber="76">Fig. 6A, C</figureCitation>
).
</paragraph>
<paragraph id="8BDAA105FFC93C7C96D2FD893450FD18" blockId="11.[189,1399,541,709]" pageId="11" pageNumber="76">
<smallCapsWord id="8D3C37D9FFC93C7C96D2FD8935E6FDEB" baselines="561,561" box="[189,254,542,568]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Body" pageId="11" pageNumber="76">BODY</smallCapsWord>
<smallCapsWord id="8D3C37D9FFC93C7C9769FDB434B7FDEB" baselines="561" box="[262,431,547,566]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="measurements" pageId="11" pageNumber="76">MEASUREMENTS</smallCapsWord>
(mm). Total length 29.7, width of 10 
<superScript id="7C100C4DFFC93C7C9505FD8A366FFDF6" attach="left" box="[874,887,541,555]" fontSize="6" pageId="11" pageNumber="76">th</superScript>
segment 1.95. Antennomere lengths (1–7): 0.48, 1.1, 1.1, 1.0, 1.05, 1.01, 0.28. Collum: length 0.8, width 1.55. Podomere lengths of 10 
<superScript id="7C100C4DFFC93C7C929FFDD731E5FD93" attach="left" box="[1264,1277,576,590]" fontSize="6" pageId="11" pageNumber="76">th</superScript>
body ring (1–7): 0.33, 0.55, 1.0, 0.55, 0.65, 1.01, 0.11. Gonopod aperture: length 0.50, width 0.75. Gonopod: length 0.93, width 0.71. Gonocoxae: length 0.7, width 0.38. Telopodite: length 0.89, width 0.35. Telson length 0.95.
</paragraph>
<paragraph id="8BDAA105FFC93C7C9685FD7B345DFCDB" blockId="11.[189,325,748,810]" box="[234,325,748,774]" pageId="11" pageNumber="76">
<emphasis id="B9117D17FFC93C7C9685FD7B345DFCDB" bold="true" box="[234,325,748,774]" pageId="11" pageNumber="76">Female</emphasis>
</paragraph>
<paragraph id="8BDAA105FFC93C7C96D2FC983420FCF7" blockId="11.[189,325,748,810]" box="[189,312,783,810]" pageId="11" pageNumber="76">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C37FF28EFFC93C7C96D2FCC736CDFC44" box="[189,981,848,922]" pageId="11" pageNumber="76" type="distribution">
<paragraph id="8BDAA105FFC93C7C96D2FCC7344EFCB7" blockId="11.[189,342,848,874]" box="[189,342,848,874]" pageId="11" pageNumber="76">
<heading id="D0921669FFC93C7C96D2FCC7344EFCB7" bold="true" box="[189,342,848,874]" fontSize="11" level="3" pageId="11" pageNumber="76" reason="3">
<emphasis id="B9117D17FFC93C7C96D2FCC7344EFCB7" bold="true" box="[189,342,848,874]" pageId="11" pageNumber="76">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BDAA105FFC93C7C96D2FC1736CDFC44" blockId="11.[189,981,895,922]" box="[189,981,895,922]" pageId="11" pageNumber="76">
Known only from the 
<collectingRegion id="49A16FE7FFC93C7C97ADFCE83619FC44" box="[450,769,895,921]" country="Ecuador" name="Morona-Santiago" pageId="11" pageNumber="76">Morona-Santiago Province</collectingRegion>
, 
<collectingCountry id="F372E195FFC93C7C9562FCE83677FC44" box="[781,879,895,921]" name="Ecuador" pageId="11" pageNumber="76">Ecuador</collectingCountry>
(
<figureCitation id="135EBD80FFC93C7C9510FCE836DFFC44" box="[895,967,895,921]" captionStart="Fig" captionStartId="14.[188,231,1892,1918]" captionTargetBox="[243,1357,497,1838]" captionTargetId="figure-72@14.[212,1380,484,1853]" captionTargetPageId="14" captionText="Fig. 9. Distribution of Chelodesmidae spp. in the Tropical Andes Biodiversity Hotspot, by country. Note: the type locality of Dibolostethus sicarius Hoffman, 2009 is included for completeness although it falls just outside of the boundaries of the Tropical Andes Biodiversity Hotspot. For information on these taxa see Supp. file 1." figureDoi="http://doi.org/10.5281/zenodo.8205518" httpUri="https://zenodo.org/record/8205518/files/figure.png" pageId="11" pageNumber="76">Fig. 9</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37FF28EFFC93C7C96D2FC5631E7FB4B" pageId="11" pageNumber="76" type="discussion">
<paragraph id="8BDAA105FFC93C7C96D2FC563437FC06" blockId="11.[189,303,961,987]" box="[189,303,961,987]" pageId="11" pageNumber="76">
<heading id="D0921669FFC93C7C96D2FC563437FC06" bold="true" box="[189,303,961,987]" fontSize="11" level="3" pageId="11" pageNumber="76" reason="3">
<emphasis id="B9117D17FFC93C7C96D2FC563437FC06" bold="true" box="[189,303,961,987]" pageId="11" pageNumber="76">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BDAA105FFC93C7C96D2FC7831E7FB4B" blockId="11.[189,1399,1007,1175]" pageId="11" pageNumber="76">
Although no collector information is given on the specimen labels, review of material housed at VMNH revealed that 
<taxonomicName id="4C65DA86FFC93C7C97D5FB843742FBF1" authorityName="Means, Bouzan &amp; Ivanov" authorityYear="2023" box="[442,602,1043,1068]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="76" phylum="Arthropoda" rank="species" species="inopinatus">
<emphasis id="B9117D17FFC93C7C97D5FB843742FBF1" box="[442,602,1043,1068]" italics="true" pageId="11" pageNumber="76">D. inopinatus</emphasis>
</taxonomicName>
was collected during the “Los 
<taxonomicName id="4C65DA86FFC93C7C95ABFB84318FFBF1" authority="Expedition" authorityName="Expedition" box="[964,1175,1042,1069]" class="Arachnida" family="Hubbardiidae" genus="Tayos" kingdom="Animalia" order="Schizomida" pageId="11" pageNumber="76" phylum="Arthropoda" rank="genus">Tayos Expedition</taxonomicName>
”. The joint British-Ecuadorian expedition, which included over 100 researchers and supporting personnel, focused on exploring the archeology, geology, and biology of Cueva de Los 
<taxonomicName id="4C65DA86FFC93C7C95ADFBCE3113FBAE" box="[962,1035,1113,1139]" class="Arachnida" family="Hubbardiidae" genus="Tayos" kingdom="Animalia" order="Schizomida" pageId="11" pageNumber="76" phylum="Arthropoda" rank="genus">Tayos</taxonomicName>
, situated in the foothills of the Cordillera del Cóndor mountain range, and its vicinity between 
<date id="FFDB87C5FFC93C7C95C3FBEB31E3FB4B" box="[940,1275,1147,1175]" pageId="11" pageNumber="76" value="1976-07-04" valueMax="1976-08-05" valueMin="1976-07-04">
July 4 
<superScript id="7C100C4DFFC93C7C959EFBEC36E6FB54" attach="left" box="[1009,1022,1147,1161]" fontSize="6" pageId="11" pageNumber="76">th</superScript>
and August 5 
<superScript id="7C100C4DFFC93C7C92CEFBEC31B6FB54" attach="none" box="[1185,1198,1147,1161]" fontSize="6" pageId="11" pageNumber="76">th</superScript>
, 1976
</date>
.
</paragraph>
</subSubSection>
</treatment>
</document>