452 lines
86 KiB
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452 lines
86 KiB
XML
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<mods:title id="9498B5C154F9138E6AE7EF0C6B79E7D6">A review of the previously monotypic tribe Dibolostethini (Chelodesmidae: Chelodesminae) with description of two new species and a summary of the Chelodesmidae of the Tropical Andes Biodiversity Hotspot</mods:title>
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<mods:namePart id="93850E1C604A4011A268847C1297D952">Means, Jackson C.</mods:namePart>
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<mods:namePart id="25F17858DE2FA68B62E96FC65EC30391">Bouzan, Rodrigo S.</mods:namePart>
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<mods:affiliation id="AB3D5020A57B77C253F579C504F362E5">Pós-graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, Brazil. & Laboratório Especial de Coleções Zoológicas, Instituto Butantan, Avenida Vital Brasil, 1500, 05503 - 090, São Paulo, Brazil.</mods:affiliation>
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<mods:namePart id="C120EBE93DB86F399EC4C0D261F2C50F">Iniesta, Luiz Felipe Moretti</mods:namePart>
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<mods:nameIdentifier id="93F49A420AC8DF6FFE73D1033760AF07" type="email">luisafvv@vt.edu</mods:nameIdentifier>
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<mods:affiliation id="18732DD2156085EC5D53FF848D5F4DF7">Virginia Museum of Natural History, 21 Starling Ave., Martinsville, Virginia 24112, USA.</mods:affiliation>
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<mods:title id="65C229C07226FE8514F33E4BB526BA6E">European Journal of Taxonomy</mods:title>
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Genus
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<emphasis id="B9117D17FFC13C7494FCFD5A3621FD35" bold="true" box="[659,825,717,744]" italics="true" pageId="3" pageNumber="68">Dibolostethus</emphasis>
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<treatmentCitation id="0AC48714FFC13C7496D2FC983740FCF4" author="Hoffman R. L." box="[189,600,782,809]" page="151" pageId="3" pageNumber="68" year="2009">
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<taxonomicName id="4C65DA86FFC13C7496D2FC983740FCF4" ID-CoL="7P99B" authority="Hoffman, 2009: 151" authorityName="Hoffman" authorityPageNumber="151" authorityYear="2009" box="[189,600,782,809]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="68" phylum="Arthropoda" rank="genus">
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<emphasis id="B9117D17FFC13C7496D2FC983478FCF4" box="[189,352,783,809]" italics="true" pageId="3" pageNumber="68">Dibolostethus</emphasis>
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<bibRefCitation id="EFF4DCF4FFC13C749708FC983740FCF4" author="Hoffman" box="[359,600,782,809]" firstAuthor="Hoffman" page="151" pageId="3" pageNumber="68" pagination="149 - 154" refId="ref9561" refString="Hoffman R. L. 2009. Chelodesmid studies. XXVI. A new genus and tribe of chelodesmid millipeds from Ecuador (Polydesmida: Chelodesmidae). International Journal of Myriapodology 2: 149 - 154. https: // doi. org / 10.1163 / 187525409 X 12577705044584" type="journal article" year="2009">Hoffman, 2009: 151</bibRefCitation>
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.
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<typeStatus id="54DE1FA7FFC13C7496D2FCC735E2FCB7" box="[189,250,848,874]" pageId="3" pageNumber="68">Type</typeStatus>
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species
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<taxonomicName id="4C65DA86FFC13C7496D2FCE8379AFC44" ID-CoL="35GSJ" authority="Hoffman, 2009" authorityName="Hoffman" authorityYear="2009" box="[189,642,895,921]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="sicarius">
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<emphasis id="B9117D17FFC13C7496D2FCE834DCFC45" box="[189,452,895,921]" italics="true" pageId="3" pageNumber="68">Dibolostethus sicarius</emphasis>
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, by monotypy.
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<emphasis id="B9117D17FFC13C7496D2FC57342EFC07" bold="true" box="[189,310,960,986]" pageId="3" pageNumber="68">Diagnosis</emphasis>
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<paragraph id="8BDAA105FFC13C7496D2FC7831AFFADD" blockId="3.[189,1400,1007,1280]" pageId="3" pageNumber="68">
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The genus differs from all other chelodesmid genera by the following combination of characters: body rings constricted between pro- and metazonites and paranota reduced, resulting in a moniliform body outline (
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<figureCitation id="135EBD80FFC13C74974EFBA23474FB92" box="[289,364,1077,1103]" captionStart="Fig" captionStartId="4.[189,232,1963,1989]" captionTargetBox="[378,1032,415,1893]" captionTargetId="figure-14@4.[310,1277,265,1924]" captionTargetPageId="4" captionText="Fig. 1. Dibolostethus kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., ♂ (ICN-MD-1367), habitus. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8205502" httpUri="https://zenodo.org/record/8205502/files/figure.png" pageId="3" pageNumber="68">Figs 1</figureCitation>
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,
|
||
<figureCitation id="135EBD80FFC13C749714FBA234BBFB92" box="[379,419,1077,1103]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[189,1398,642,1776]" captionTargetId="figure-157@5.[189,1398,642,1776]" captionTargetPageId="5" captionText="Fig. 2. Scanning electron microscope images of Dibolostethus Hoffman, 2009 showing somatic characters. Dibolostethus kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., paratype, ♂ (ICN- MD-1317B). A. Body rings 9th and 10th, red rectangle = median metazonal sulcus. B. Venter of 4th body ring, showing paired sternal projections, red oval = stigma. C. Venter of head, showing narrow gnathochilarium. D. Tip of seventh antennomere, red arrow = narrow slit connecting pocket containing sensilla basiconica and sensory cone area." figureDoi="http://doi.org/10.5281/zenodo.8205504" httpUri="https://zenodo.org/record/8205504/files/figure.png" pageId="3" pageNumber="68">2A</figureCitation>
|
||
); spiracles reduced as compared to other members of the family, oval (
|
||
<figureCitation id="135EBD80FFC13C749286FBA2305EFB92" box="[1257,1350,1077,1103]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[189,1398,642,1776]" captionTargetId="figure-157@5.[189,1398,642,1776]" captionTargetPageId="5" captionText="Fig. 2. Scanning electron microscope images of Dibolostethus Hoffman, 2009 showing somatic characters. Dibolostethus kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., paratype, ♂ (ICN- MD-1317B). A. Body rings 9th and 10th, red rectangle = median metazonal sulcus. B. Venter of 4th body ring, showing paired sternal projections, red oval = stigma. C. Venter of head, showing narrow gnathochilarium. D. Tip of seventh antennomere, red arrow = narrow slit connecting pocket containing sensilla basiconica and sensory cone area." figureDoi="http://doi.org/10.5281/zenodo.8205504" httpUri="https://zenodo.org/record/8205504/files/figure.png" pageId="3" pageNumber="68">Fig. 2B</figureCitation>
|
||
, red circle); gonopores located on subterminal convexity on the mesal side of 2
|
||
<superScript id="7C100C4DFFC13C749245FBCF3123FBBB" attach="left" box="[1066,1083,1112,1126]" fontSize="6" pageId="3" pageNumber="68">nd</superScript>
|
||
coxae (
|
||
<figureCitation id="135EBD80FFC13C7492F7FBCF31EEFBAF" box="[1176,1270,1112,1138]" captionStart="Fig" captionStartId="7.[189,232,1782,1808]" captionTargetBox="[189,1398,812,1742]" captionTargetId="figure-103@7.[189,1398,806,1742]" captionTargetPageId="7" captionText="Fig. 3. Dibolostethus Hoffman, 2009 somatic characters continued. A, E. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810). B–D. D. kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., holotype, ♂ (ICN-MD-1317-1). A. Coxa 2, mesal view, red arrow = gonopore. B. Left leg 4, posterior view, red arrow = tibial ventro-apical projection. C. Left leg of 5th leg pair showing incrassate femur, red arrow = tibial ventro-apical projection. D. Body ring 5, lateral view, red circle = anterior pair of sternal projections. E. Prefemur 5, red arrow = basal pore. Abbreviations: Cx = coxa; Pf = prefemur." figureDoi="http://doi.org/10.5281/zenodo.8205506" httpUri="https://zenodo.org/record/8205506/files/figure.png" pageId="3" pageNumber="68">Fig. 3A</figureCitation>
|
||
); in males, sternite of 4
|
||
<superScript id="7C100C4DFFC13C749721FBEC3443FB54" attach="left" box="[334,347,1147,1161]" fontSize="6" pageId="3" pageNumber="68">th</superScript>
|
||
body ring prominent, with a pair of acute projections curving anteriad (
|
||
<figureCitation id="135EBD80FFC13C7492ABFBEC303FFB4B" box="[1220,1319,1147,1174]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[189,1398,642,1776]" captionTargetId="figure-157@5.[189,1398,642,1776]" captionTargetPageId="5" captionText="Fig. 2. Scanning electron microscope images of Dibolostethus Hoffman, 2009 showing somatic characters. Dibolostethus kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., paratype, ♂ (ICN- MD-1317B). A. Body rings 9th and 10th, red rectangle = median metazonal sulcus. B. Venter of 4th body ring, showing paired sternal projections, red oval = stigma. C. Venter of head, showing narrow gnathochilarium. D. Tip of seventh antennomere, red arrow = narrow slit connecting pocket containing sensilla basiconica and sensory cone area." figureDoi="http://doi.org/10.5281/zenodo.8205504" httpUri="https://zenodo.org/record/8205504/files/figure.png" pageId="3" pageNumber="68">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="135EBD80FFC13C74935AFBEB305FFB4B" box="[1333,1351,1148,1174]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[190,1352,1052,1886]" captionTargetId="figure-271@8.[189,1398,1044,1886]" captionTargetPageId="8" captionText="Fig. 4. Body ring 4, sternal projections. A–B. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810). C–D. D. inopinatus Means, Bouzan & Ivanov sp. nov., holotype, ♂ (VMNH110813). A, C. Lateral views. B, D. Posterolateral views. Abbreviation: Cx = coxae." figureDoi="http://doi.org/10.5281/zenodo.8205508" httpUri="https://zenodo.org/record/8205508/files/figure.png" pageId="3" pageNumber="68">4</figureCitation>
|
||
); in males, femora of 4
|
||
<superScript id="7C100C4DFFC13C7497F3FB0934B1FB71" attach="left" box="[412,425,1182,1196]" fontSize="6" pageId="3" pageNumber="68">th</superScript>
|
||
pair of legs with large acute process (
|
||
<figureCitation id="135EBD80FFC13C749506FB0836DEFB64" box="[873,966,1183,1209]" captionStart="Fig" captionStartId="7.[189,232,1782,1808]" captionTargetBox="[189,1398,812,1742]" captionTargetId="figure-103@7.[189,1398,806,1742]" captionTargetPageId="7" captionText="Fig. 3. Dibolostethus Hoffman, 2009 somatic characters continued. A, E. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810). B–D. D. kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., holotype, ♂ (ICN-MD-1317-1). A. Coxa 2, mesal view, red arrow = gonopore. B. Left leg 4, posterior view, red arrow = tibial ventro-apical projection. C. Left leg of 5th leg pair showing incrassate femur, red arrow = tibial ventro-apical projection. D. Body ring 5, lateral view, red circle = anterior pair of sternal projections. E. Prefemur 5, red arrow = basal pore. Abbreviations: Cx = coxa; Pf = prefemur." figureDoi="http://doi.org/10.5281/zenodo.8205506" httpUri="https://zenodo.org/record/8205506/files/figure.png" pageId="3" pageNumber="68">Fig. 3B</figureCitation>
|
||
); in males, femora of 5
|
||
<superScript id="7C100C4DFFC13C7492B4FB0931F0FB71" attach="left" box="[1243,1256,1182,1196]" fontSize="6" pageId="3" pageNumber="68">th</superScript>
|
||
pair of legs incrassate (
|
||
<figureCitation id="135EBD80FFC13C74972BFB5534BBFB01" box="[324,419,1218,1244]" captionStart="Fig" captionStartId="7.[189,232,1782,1808]" captionTargetBox="[189,1398,812,1742]" captionTargetId="figure-103@7.[189,1398,806,1742]" captionTargetPageId="7" captionText="Fig. 3. Dibolostethus Hoffman, 2009 somatic characters continued. A, E. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810). B–D. D. kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., holotype, ♂ (ICN-MD-1317-1). A. Coxa 2, mesal view, red arrow = gonopore. B. Left leg 4, posterior view, red arrow = tibial ventro-apical projection. C. Left leg of 5th leg pair showing incrassate femur, red arrow = tibial ventro-apical projection. D. Body ring 5, lateral view, red circle = anterior pair of sternal projections. E. Prefemur 5, red arrow = basal pore. Abbreviations: Cx = coxa; Pf = prefemur." figureDoi="http://doi.org/10.5281/zenodo.8205506" httpUri="https://zenodo.org/record/8205506/files/figure.png" pageId="3" pageNumber="68">Fig. 3C</figureCitation>
|
||
); gonopods large, straight; solenomere long, slender, and unbranched; prefemoral process widening at midpoint and terminating in several well-defined lobes (
|
||
<figureCitation id="135EBD80FFC13C749250FB7231B2FADD" box="[1087,1194,1253,1280]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="9.[189,232,970,996]" captionStartId-1="10.[189,232,1928,1954]" captionStartId-2="12.[188,231,1892,1918]" captionTargetBox-0="[189,1397,293,928]" captionTargetBox-1="[189,1398,1270,1881]" captionTargetBox-2="[190,1398,1234,1849]" captionTargetId-0="figure-255@9.[189,1398,287,928]" captionTargetId-1="figure-240@10.[189,1398,1266,1881]" captionTargetId-2="figure-296@12.[189,1398,1228,1849]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="12" captionText-0="Fig. 5. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810), left gonopod. A. Lateral view. B. Anterior view. C. Mesal view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." captionText-1="Fig. 6. Dibolostethus inopinatus Means, Bouzan & Ivanov sp. nov., holotype, ♂ (VMNH110813), left gonopod. A. Lateral view, red circle = cingulum. B. Anterior view. C. Mesal view. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." captionText-2="Fig. 7. Dibolostethus kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., holotype, ♂ (ICN- MD-1317A), right gonopod. A. Mesal view. B. Anterior view. C. Lateral view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: Lb = lobe; PfP = prefemoral process; S = solenomere; Sh = shelf." figureDoi-0="http://doi.org/10.5281/zenodo.8205510" figureDoi-1="http://doi.org/10.5281/zenodo.8205512" figureDoi-2="http://doi.org/10.5281/zenodo.8205514" httpUri-0="https://zenodo.org/record/8205510/files/figure.png" httpUri-1="https://zenodo.org/record/8205512/files/figure.png" httpUri-2="https://zenodo.org/record/8205514/files/figure.png" pageId="3" pageNumber="68">Figs 5–7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37FF28EFFC13C7196D2FAB13157FCF3" lastPageId="6" lastPageNumber="71" pageId="3" pageNumber="68" type="description">
|
||
<paragraph id="8BDAA105FFC13C7496D2FAB13456FA9D" blockId="3.[189,334,1318,1344]" box="[189,334,1318,1344]" pageId="3" pageNumber="68">
|
||
<heading id="D0921669FFC13C7496D2FAB13456FA9D" bold="true" box="[189,334,1318,1344]" fontSize="11" level="3" pageId="3" pageNumber="68" reason="3">
|
||
<emphasis id="B9117D17FFC13C7496D2FAB13456FA9D" bold="true" box="[189,334,1318,1344]" pageId="3" pageNumber="68">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDAA105FFC13C7296D2FAC2364BFDE1" blockId="3.[187,1401,1365,1990]" lastBlockId="5.[188,1400,265,573]" lastPageId="5" lastPageNumber="70" pageId="3" pageNumber="68">
|
||
Body composed of 20 rings. Length of males between
|
||
<quantity id="4C9D0CE0FFC13C749551FAC23688FAAD" box="[830,912,1365,1392]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="68" unit="mm" value="25.0">25 mm</quantity>
|
||
(
|
||
<taxonomicName id="4C65DA86FFC13C7495CEFAC13139FAB2" authorityName="Hoffman" authorityYear="2009" box="[929,1057,1366,1391]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="sicarius">
|
||
<emphasis id="B9117D17FFC13C7495CEFAC13139FAB2" box="[929,1057,1366,1391]" italics="true" pageId="3" pageNumber="68">D. sicarius</emphasis>
|
||
</taxonomicName>
|
||
) and ~
|
||
<quantity id="4C9D0CE0FFC13C74921CFAC231DEFAAD" box="[1139,1222,1365,1392]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="68" unit="mm" value="30.0">30 mm</quantity>
|
||
(
|
||
<taxonomicName id="4C65DA86FFC13C7492B8FAC1306FFAB2" authority="Means & Bouzan & Iniesta & Martínez-Torres & Vasquez-Valverde & Brescovit & Ivanov, 2023" authorityName="Means & Bouzan & Iniesta & Martínez-Torres & Vasquez-Valverde & Brescovit & Ivanov" authorityYear="2023" box="[1239,1399,1366,1391]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="inopinatus" status="sp. nov.">
|
||
<emphasis id="B9117D17FFC13C7492B8FAC1306FFAB2" box="[1239,1399,1366,1391]" italics="true" pageId="3" pageNumber="68">D. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A222C06CFFC13C7496D2FAEE3405FA4F" box="[189,285,1400,1427]" pageId="3" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
|
||
). Coloration (preserved in 70% isopropanol/ethanol): body rings yellowish (
|
||
<taxonomicName id="4C65DA86FFC13C7492C3FAEE3055FA4F" authorityName="Means & Bouzan & Iniesta & Martínez-Torres & Vasquez-Valverde & Brescovit & Ivanov" authorityYear="2023" box="[1196,1357,1401,1426]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="B9117D17FFC13C7492C3FAEE3055FA4F" box="[1196,1357,1401,1426]" italics="true" pageId="3" pageNumber="68">D. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
) to dark reddish brown (nearly purple;
|
||
<taxonomicName id="4C65DA86FFC13C749431FA0B37CAFA68" authority="Means & Bouzan & Iniesta & Martínez-Torres & Vasquez-Valverde & Brescovit & Ivanov, 2023" authorityName="Means & Bouzan & Iniesta & Martínez-Torres & Vasquez-Valverde & Brescovit & Ivanov" authorityYear="2023" box="[606,722,1435,1461]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="kattani" status="sp. nov.">
|
||
<emphasis id="B9117D17FFC13C749431FA0B37CAFA68" box="[606,722,1435,1461]" italics="true" pageId="3" pageNumber="68">D. kattani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A222C06CFFC13C7494B5FA0B362FFA6B" box="[730,823,1436,1462]" pageId="3" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<figureCitation id="135EBD80FFC13C74952CFA0C369FFA6B" box="[835,903,1435,1462]" captionStart="Fig" captionStartId="4.[189,232,1963,1989]" captionTargetBox="[378,1032,415,1893]" captionTargetId="figure-14@4.[310,1277,265,1924]" captionTargetPageId="4" captionText="Fig. 1. Dibolostethus kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., ♂ (ICN-MD-1367), habitus. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8205502" httpUri="https://zenodo.org/record/8205502/files/figure.png" pageId="3" pageNumber="68">Fig. 1</figureCitation>
|
||
); antennae, head, legs, and paranota same color as the body (
|
||
<taxonomicName id="4C65DA86FFC13C7497FFFA283736FA05" authorityName="Means & Bouzan & Iniesta & Martínez-Torres & Vasquez-Valverde & Brescovit & Ivanov" authorityYear="2023" box="[400,558,1471,1496]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="B9117D17FFC13C7497FFFA283736FA05" box="[400,558,1471,1496]" italics="true" pageId="3" pageNumber="68">D. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
) or light yellow, contrasting with the body; telson same color as the body (
|
||
<taxonomicName id="4C65DA86FFC13C7496A8FA743471FA21" authorityName="Means & Bouzan & Iniesta & Martínez-Torres & Vasquez-Valverde & Brescovit & Ivanov" authorityYear="2023" box="[199,361,1507,1532]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="B9117D17FFC13C7496A8FA743471FA21" box="[199,361,1507,1532]" italics="true" pageId="3" pageNumber="68">D. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
) or reddish with posterior margin yellowish. Head: smooth, with epicranial setae 2–2, interantennal 1–1, and frontal 2–2; other facial setae sparse and irregular, becoming more dense closer to the mouthparts. Gnathochilarium without modifications, but narrower in relation to other members of
|
||
<taxonomicName id="4C65DA86FFC13C7496D2F9DC346DF9B8" authorityName="Cook" authorityYear="1895" box="[189,373,1611,1637]" class="Diplopoda" family="Chelodesmidae" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="68" phylum="Arthropoda" rank="family">Chelodesmidae</taxonomicName>
|
||
, covered by few setae (
|
||
<figureCitation id="135EBD80FFC13C7494EFF9DC37C3F9BB" box="[640,731,1611,1638]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[189,1398,642,1776]" captionTargetId="figure-157@5.[189,1398,642,1776]" captionTargetPageId="5" captionText="Fig. 2. Scanning electron microscope images of Dibolostethus Hoffman, 2009 showing somatic characters. Dibolostethus kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., paratype, ♂ (ICN- MD-1317B). A. Body rings 9th and 10th, red rectangle = median metazonal sulcus. B. Venter of 4th body ring, showing paired sternal projections, red oval = stigma. C. Venter of head, showing narrow gnathochilarium. D. Tip of seventh antennomere, red arrow = narrow slit connecting pocket containing sensilla basiconica and sensory cone area." figureDoi="http://doi.org/10.5281/zenodo.8205504" httpUri="https://zenodo.org/record/8205504/files/figure.png" pageId="3" pageNumber="68">Fig. 2C</figureCitation>
|
||
). Mandibular bases prominent. Incisura lateralis rounded, not completely closed. Antennae: sixth antennomere with an invagination possibly containing sensilla basiconica, obscured by debris in examined specimen; seventh antennomere having two well-demarcated invaginations and one slight invagination between the sensory cones; sensilla basiconica arranged into a small pocket, connected to the cones by a slit (
|
||
<figureCitation id="135EBD80FFC13C7494B5F94F362FF92F" box="[730,823,1752,1778]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[189,1398,642,1776]" captionTargetId="figure-157@5.[189,1398,642,1776]" captionTargetPageId="5" captionText="Fig. 2. Scanning electron microscope images of Dibolostethus Hoffman, 2009 showing somatic characters. Dibolostethus kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., paratype, ♂ (ICN- MD-1317B). A. Body rings 9th and 10th, red rectangle = median metazonal sulcus. B. Venter of 4th body ring, showing paired sternal projections, red oval = stigma. C. Venter of head, showing narrow gnathochilarium. D. Tip of seventh antennomere, red arrow = narrow slit connecting pocket containing sensilla basiconica and sensory cone area." figureDoi="http://doi.org/10.5281/zenodo.8205504" httpUri="https://zenodo.org/record/8205504/files/figure.png" pageId="3" pageNumber="68">Fig. 2D</figureCitation>
|
||
, red arrow). Body ring cuticle smooth, appearing slightly rugose at high magnification, with a median sulcus on the metazonites (
|
||
<figureCitation id="135EBD80FFC13C74920CF96C31D9F8CB" box="[1123,1217,1787,1814]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[189,1398,642,1776]" captionTargetId="figure-157@5.[189,1398,642,1776]" captionTargetPageId="5" captionText="Fig. 2. Scanning electron microscope images of Dibolostethus Hoffman, 2009 showing somatic characters. Dibolostethus kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., paratype, ♂ (ICN- MD-1317B). A. Body rings 9th and 10th, red rectangle = median metazonal sulcus. B. Venter of 4th body ring, showing paired sternal projections, red oval = stigma. C. Venter of head, showing narrow gnathochilarium. D. Tip of seventh antennomere, red arrow = narrow slit connecting pocket containing sensilla basiconica and sensory cone area." figureDoi="http://doi.org/10.5281/zenodo.8205504" httpUri="https://zenodo.org/record/8205504/files/figure.png" pageId="3" pageNumber="68">Fig. 2A</figureCitation>
|
||
, red rectangle). Collum: small, more slender than the head; posterior margin concave, anterior margin convex; corners rounded. Paranota reduced, distantly separated, rounded, and without projections, not developed on 19
|
||
<superScript id="7C100C4DFFC13C7496B5F8F335FFF8AF" attach="left" box="[218,231,1892,1906]" fontSize="6" pageId="3" pageNumber="68">th</superScript>
|
||
body ring, and body rings constricted between pro- and metazonites, all resulting in a moniliform appearance (
|
||
<figureCitation id="135EBD80FFC13C74973CF81F34B8F87F" box="[339,416,1928,1954]" captionStart="Fig" captionStartId="4.[189,232,1963,1989]" captionTargetBox="[378,1032,415,1893]" captionTargetId="figure-14@4.[310,1277,265,1924]" captionTargetPageId="4" captionText="Fig. 1. Dibolostethus kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., ♂ (ICN-MD-1367), habitus. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8205502" httpUri="https://zenodo.org/record/8205502/files/figure.png" pageId="3" pageNumber="68">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="135EBD80FFC13C7497DFF81F34C0F87F" box="[432,472,1928,1954]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[189,1398,642,1776]" captionTargetId="figure-157@5.[189,1398,642,1776]" captionTargetPageId="5" captionText="Fig. 2. Scanning electron microscope images of Dibolostethus Hoffman, 2009 showing somatic characters. Dibolostethus kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., paratype, ♂ (ICN- MD-1317B). A. Body rings 9th and 10th, red rectangle = median metazonal sulcus. B. Venter of 4th body ring, showing paired sternal projections, red oval = stigma. C. Venter of head, showing narrow gnathochilarium. D. Tip of seventh antennomere, red arrow = narrow slit connecting pocket containing sensilla basiconica and sensory cone area." figureDoi="http://doi.org/10.5281/zenodo.8205504" httpUri="https://zenodo.org/record/8205504/files/figure.png" pageId="3" pageNumber="68">2A</figureCitation>
|
||
). Stigma oval (
|
||
<figureCitation id="135EBD80FFC13C7494E0F81F37F5F87F" box="[655,749,1928,1954]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[189,1398,642,1776]" captionTargetId="figure-157@5.[189,1398,642,1776]" captionTargetPageId="5" captionText="Fig. 2. Scanning electron microscope images of Dibolostethus Hoffman, 2009 showing somatic characters. Dibolostethus kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., paratype, ♂ (ICN- MD-1317B). A. Body rings 9th and 10th, red rectangle = median metazonal sulcus. B. Venter of 4th body ring, showing paired sternal projections, red oval = stigma. C. Venter of head, showing narrow gnathochilarium. D. Tip of seventh antennomere, red arrow = narrow slit connecting pocket containing sensilla basiconica and sensory cone area." figureDoi="http://doi.org/10.5281/zenodo.8205504" httpUri="https://zenodo.org/record/8205504/files/figure.png" pageId="3" pageNumber="68">Fig. 2B</figureCitation>
|
||
, red circle). Sternites of males: 4
|
||
<superScript id="7C100C4DFFC13C749215F810319FF848" attach="left" box="[1146,1159,1927,1941]" fontSize="6" pageId="3" pageNumber="68">th</superScript>
|
||
body ring with two slender acute projections (
|
||
<figureCitation id="135EBD80FFC13C749799F83C374FF81B" box="[502,599,1963,1990]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[189,1398,642,1776]" captionTargetId="figure-157@5.[189,1398,642,1776]" captionTargetPageId="5" captionText="Fig. 2. Scanning electron microscope images of Dibolostethus Hoffman, 2009 showing somatic characters. Dibolostethus kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., paratype, ♂ (ICN- MD-1317B). A. Body rings 9th and 10th, red rectangle = median metazonal sulcus. B. Venter of 4th body ring, showing paired sternal projections, red oval = stigma. C. Venter of head, showing narrow gnathochilarium. D. Tip of seventh antennomere, red arrow = narrow slit connecting pocket containing sensilla basiconica and sensory cone area." figureDoi="http://doi.org/10.5281/zenodo.8205504" httpUri="https://zenodo.org/record/8205504/files/figure.png" pageId="3" pageNumber="68">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="135EBD80FFC13C74940BF83B376EF81B" box="[612,630,1964,1990]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[190,1352,1052,1886]" captionTargetId="figure-271@8.[189,1398,1044,1886]" captionTargetPageId="8" captionText="Fig. 4. Body ring 4, sternal projections. A–B. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810). C–D. D. inopinatus Means, Bouzan & Ivanov sp. nov., holotype, ♂ (VMNH110813). A, C. Lateral views. B, D. Posterolateral views. Abbreviation: Cx = coxae." figureDoi="http://doi.org/10.5281/zenodo.8205508" httpUri="https://zenodo.org/record/8205508/files/figure.png" pageId="3" pageNumber="68">4</figureCitation>
|
||
); 5
|
||
<superScript id="7C100C4DFFC13C7494F5F83C37BFF864" attach="left" box="[666,679,1963,1977]" fontSize="6" pageId="3" pageNumber="68">th</superScript>
|
||
body ring with two pairs of short, blunt projections (only in
|
||
<taxonomicName id="4C65DA86FFC73C7296D2FE9D342BFEFE" authorityName="Means & Bouzan & Iniesta & Martínez-Torres & Vasquez-Valverde & Brescovit & Ivanov" authorityYear="2023" box="[189,307,265,291]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="70" phylum="Arthropoda" rank="species" species="kattani">
|
||
<emphasis id="B9117D17FFC73C7296D2FE9D342BFEFE" box="[189,307,265,291]" italics="true" pageId="5" pageNumber="70">D. kattani</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="135EBD80FFC73C72972AFE9E34BAFEFE" box="[325,418,265,291]" captionStart="Fig" captionStartId="7.[189,232,1782,1808]" captionTargetBox="[189,1398,812,1742]" captionTargetId="figure-103@7.[189,1398,806,1742]" captionTargetPageId="7" captionText="Fig. 3. Dibolostethus Hoffman, 2009 somatic characters continued. A, E. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810). B–D. D. kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., holotype, ♂ (ICN-MD-1317-1). A. Coxa 2, mesal view, red arrow = gonopore. B. Left leg 4, posterior view, red arrow = tibial ventro-apical projection. C. Left leg of 5th leg pair showing incrassate femur, red arrow = tibial ventro-apical projection. D. Body ring 5, lateral view, red circle = anterior pair of sternal projections. E. Prefemur 5, red arrow = basal pore. Abbreviations: Cx = coxa; Pf = prefemur." figureDoi="http://doi.org/10.5281/zenodo.8205506" httpUri="https://zenodo.org/record/8205506/files/figure.png" pageId="5" pageNumber="70">Fig. 3D</figureCitation>
|
||
, red oval); post-gonopodal sternites unmodified. Ozopores: posteriorly situated on paranota (
|
||
<figureCitation id="135EBD80FFC73C729759FEBB3480FE9B" box="[310,408,300,326]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[189,1398,642,1776]" captionTargetId="figure-157@5.[189,1398,642,1776]" captionTargetPageId="5" captionText="Fig. 2. Scanning electron microscope images of Dibolostethus Hoffman, 2009 showing somatic characters. Dibolostethus kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., paratype, ♂ (ICN- MD-1317B). A. Body rings 9th and 10th, red rectangle = median metazonal sulcus. B. Venter of 4th body ring, showing paired sternal projections, red oval = stigma. C. Venter of head, showing narrow gnathochilarium. D. Tip of seventh antennomere, red arrow = narrow slit connecting pocket containing sensilla basiconica and sensory cone area." figureDoi="http://doi.org/10.5281/zenodo.8205504" httpUri="https://zenodo.org/record/8205504/files/figure.png" pageId="5" pageNumber="70">Fig. 2A</figureCitation>
|
||
); peritremata indistinguishable; arrangement typical for
|
||
<taxonomicName id="4C65DA86FFC73C72922CFEBB31C3FE9B" authorityName="Pocock" authorityYear="1887" box="[1091,1243,300,326]" class="Diplopoda" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="70" phylum="Arthropoda" rank="order">Polydesmida</taxonomicName>
|
||
(5, 7, 9, 10, 12, 13, 15–19); without a well-demarcated rim. Legs of males: femur of 4
|
||
<superScript id="7C100C4DFFC73C729241FED83123FE80" attach="left" box="[1070,1083,335,349]" fontSize="6" pageId="5" pageNumber="70">th</superScript>
|
||
pair produced into a large acute process (
|
||
<figureCitation id="135EBD80FFC73C729701FEE534D4FE50" box="[366,460,370,397]" captionStart="Fig" captionStartId="7.[189,232,1782,1808]" captionTargetBox="[189,1398,812,1742]" captionTargetId="figure-103@7.[189,1398,806,1742]" captionTargetPageId="7" captionText="Fig. 3. Dibolostethus Hoffman, 2009 somatic characters continued. A, E. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810). B–D. D. kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., holotype, ♂ (ICN-MD-1317-1). A. Coxa 2, mesal view, red arrow = gonopore. B. Left leg 4, posterior view, red arrow = tibial ventro-apical projection. C. Left leg of 5th leg pair showing incrassate femur, red arrow = tibial ventro-apical projection. D. Body ring 5, lateral view, red circle = anterior pair of sternal projections. E. Prefemur 5, red arrow = basal pore. Abbreviations: Cx = coxa; Pf = prefemur." figureDoi="http://doi.org/10.5281/zenodo.8205506" httpUri="https://zenodo.org/record/8205506/files/figure.png" pageId="5" pageNumber="70">Fig. 3B</figureCitation>
|
||
); femur of 5
|
||
<superScript id="7C100C4DFFC73C72940EFEE53776FE5D" attach="left" box="[609,622,370,384]" fontSize="6" pageId="5" pageNumber="70">th</superScript>
|
||
pair of legs incrassate and glandular; prefemur of 5
|
||
<superScript id="7C100C4DFFC73C7292B5FEE531FFFE5D" attach="left" box="[1242,1255,370,384]" fontSize="6" pageId="5" pageNumber="70">th</superScript>
|
||
pair of legs with a small basal pore (
|
||
<figureCitation id="135EBD80FFC73C72978BFE01377BFE6D" box="[484,611,406,432]" captionStart="Fig" captionStartId="7.[189,232,1782,1808]" captionTargetBox="[189,1398,812,1742]" captionTargetId="figure-103@7.[189,1398,806,1742]" captionTargetPageId="7" captionText="Fig. 3. Dibolostethus Hoffman, 2009 somatic characters continued. A, E. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810). B–D. D. kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., holotype, ♂ (ICN-MD-1317-1). A. Coxa 2, mesal view, red arrow = gonopore. B. Left leg 4, posterior view, red arrow = tibial ventro-apical projection. C. Left leg of 5th leg pair showing incrassate femur, red arrow = tibial ventro-apical projection. D. Body ring 5, lateral view, red circle = anterior pair of sternal projections. E. Prefemur 5, red arrow = basal pore. Abbreviations: Cx = coxa; Pf = prefemur." figureDoi="http://doi.org/10.5281/zenodo.8205506" httpUri="https://zenodo.org/record/8205506/files/figure.png" pageId="5" pageNumber="70">Fig. 3C, E</figureCitation>
|
||
, red arrow); tibia of all legs with a ventro-apical translucent acute projection, reducing in size posteriorly (
|
||
<figureCitation id="135EBD80FFC73C7294E0FE2E3614FE0E" box="[655,780,441,467]" captionStart="Fig" captionStartId="7.[189,232,1782,1808]" captionTargetBox="[189,1398,812,1742]" captionTargetId="figure-103@7.[189,1398,806,1742]" captionTargetPageId="7" captionText="Fig. 3. Dibolostethus Hoffman, 2009 somatic characters continued. A, E. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810). B–D. D. kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., holotype, ♂ (ICN-MD-1317-1). A. Coxa 2, mesal view, red arrow = gonopore. B. Left leg 4, posterior view, red arrow = tibial ventro-apical projection. C. Left leg of 5th leg pair showing incrassate femur, red arrow = tibial ventro-apical projection. D. Body ring 5, lateral view, red circle = anterior pair of sternal projections. E. Prefemur 5, red arrow = basal pore. Abbreviations: Cx = coxa; Pf = prefemur." figureDoi="http://doi.org/10.5281/zenodo.8205506" httpUri="https://zenodo.org/record/8205506/files/figure.png" pageId="5" pageNumber="70">Fig. 3B–C</figureCitation>
|
||
, red arrows). Podosterna of 4
|
||
<superScript id="7C100C4DFFC73C729231FE2F3173FE1B" attach="none" box="[1118,1131,440,454]" fontSize="6" pageId="5" pageNumber="70">th</superScript>
|
||
–6
|
||
<superScript id="7C100C4DFFC73C7292E6FE2F318EFE1B" attach="left" box="[1161,1174,440,454]" fontSize="6" pageId="5" pageNumber="70">th</superScript>
|
||
body rings notably elevated (
|
||
<figureCitation id="135EBD80FFC73C72975EFE4B348CFE2B" box="[305,404,476,502]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[189,1398,642,1776]" captionTargetId="figure-157@5.[189,1398,642,1776]" captionTargetPageId="5" captionText="Fig. 2. Scanning electron microscope images of Dibolostethus Hoffman, 2009 showing somatic characters. Dibolostethus kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., paratype, ♂ (ICN- MD-1317B). A. Body rings 9th and 10th, red rectangle = median metazonal sulcus. B. Venter of 4th body ring, showing paired sternal projections, red oval = stigma. C. Venter of head, showing narrow gnathochilarium. D. Tip of seventh antennomere, red arrow = narrow slit connecting pocket containing sensilla basiconica and sensory cone area." figureDoi="http://doi.org/10.5281/zenodo.8205504" httpUri="https://zenodo.org/record/8205504/files/figure.png" pageId="5" pageNumber="70">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="135EBD80FFC73C7297CBFE4B34D3FE2B" box="[420,459,476,502]" captionStart="Fig" captionStartId="7.[189,232,1782,1808]" captionTargetBox="[189,1398,812,1742]" captionTargetId="figure-103@7.[189,1398,806,1742]" captionTargetPageId="7" captionText="Fig. 3. Dibolostethus Hoffman, 2009 somatic characters continued. A, E. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810). B–D. D. kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., holotype, ♂ (ICN-MD-1317-1). A. Coxa 2, mesal view, red arrow = gonopore. B. Left leg 4, posterior view, red arrow = tibial ventro-apical projection. C. Left leg of 5th leg pair showing incrassate femur, red arrow = tibial ventro-apical projection. D. Body ring 5, lateral view, red circle = anterior pair of sternal projections. E. Prefemur 5, red arrow = basal pore. Abbreviations: Cx = coxa; Pf = prefemur." figureDoi="http://doi.org/10.5281/zenodo.8205506" httpUri="https://zenodo.org/record/8205506/files/figure.png" pageId="5" pageNumber="70">3D</figureCitation>
|
||
). Telson: subtriangular, with three pairs of macrosetae dorsolaterally and two pairs in the apical region. The apical pairs of macrosetae separate from and equidistant to the mesal lips of the paraproct, forming the four corners of a rectangle.
|
||
</paragraph>
|
||
<caption id="DF1AF18DFFC63C7396D2F83C34EFF834" ID-DOI="http://doi.org/10.5281/zenodo.8205502" ID-Zenodo-Dep="8205502" httpUri="https://zenodo.org/record/8205502/files/figure.png" pageId="4" pageNumber="69" startId="4.[189,232,1963,1989]" targetBox="[378,1032,415,1893]" targetPageId="4">
|
||
<paragraph id="8BDAA105FFC63C7396D2F83C34EFF834" blockId="4.[189,1399,1963,2025]" pageId="4" pageNumber="69">
|
||
<emphasis id="B9117D17FFC63C7396D2F83C3415F818" bold="true" box="[189,269,1963,1989]" pageId="4" pageNumber="69">Fig. 1.</emphasis>
|
||
<taxonomicName id="4C65DA86FFC63C739779F83C3101F81B" authority="Means, Bouzan, Martinez-Torres & Ivanov" authorityName="Means, Bouzan, Martinez-Torres & Ivanov" authorityYear="2023" box="[278,1049,1963,1990]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="69" phylum="Arthropoda" rank="species" species="kattani" status="sp. nov.">
|
||
<emphasis id="B9117D17FFC63C739779F83C370BF818" box="[278,531,1963,1989]" italics="true" pageId="4" pageNumber="69">Dibolostethus kattani</emphasis>
|
||
Means, Bouzan, Martínez-Torres & Ivanov
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A222C06CFFC63C73924CF83B3199F81B" box="[1059,1153,1964,1990]" pageId="4" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, ♂ (ICN-MD-1367), habitus. Scale bar = 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1AF18DFFC73C7296D2F88937B2F834" ID-DOI="http://doi.org/10.5281/zenodo.8205504" ID-Zenodo-Dep="8205504" httpUri="https://zenodo.org/record/8205504/files/figure.png" pageId="5" pageNumber="70" startId="5.[189,232,1822,1848]" targetBox="[189,1398,642,1776]" targetPageId="5">
|
||
<paragraph id="8BDAA105FFC73C7296D2F88937B2F834" blockId="5.[189,1399,1822,2025]" pageId="5" pageNumber="70">
|
||
<emphasis id="B9117D17FFC73C7296D2F889340BF8E5" bold="true" box="[189,275,1822,1848]" pageId="5" pageNumber="70">Fig. 2.</emphasis>
|
||
Scanning electron microscope images of
|
||
<taxonomicName id="4C65DA86FFC73C729542F8893182F8E5" authority="Hoffman, 2009" authorityName="Hoffman" authorityYear="2009" box="[813,1178,1822,1848]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="70" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9117D17FFC73C729542F88936C8F8E5" box="[813,976,1822,1848]" italics="true" pageId="5" pageNumber="70">Dibolostethus</emphasis>
|
||
<bibRefCitation id="EFF4DCF4FFC73C7295B0F8893182F8E5" author="Hoffman" box="[991,1178,1822,1848]" firstAuthor="Hoffman" pageId="5" pageNumber="70" pagination="149 - 154" refId="ref9561" refString="Hoffman R. L. 2009. Chelodesmid studies. XXVI. A new genus and tribe of chelodesmid millipeds from Ecuador (Polydesmida: Chelodesmidae). International Journal of Myriapodology 2: 149 - 154. https: // doi. org / 10.1163 / 187525409 X 12577705044584" type="journal article" year="2009">Hoffman, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
showing somatic characters.
|
||
<taxonomicName id="4C65DA86FFC73C72972EF8D6312CF881" authority="Means, Bouzan, Martinez-Torres & Ivanov" authorityName="Means, Bouzan, Martinez-Torres & Ivanov" authorityYear="2023" box="[321,1076,1857,1884]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="70" phylum="Arthropoda" rank="species" species="kattani" status="sp. nov.">
|
||
<emphasis id="B9117D17FFC73C72972EF8D63723F886" box="[321,571,1857,1883]" italics="true" pageId="5" pageNumber="70">Dibolostethus kattani</emphasis>
|
||
Means, Bouzan, Martínez-Torres & Ivanov
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A222C06CFFC73C729254F8D5318EF881" box="[1083,1174,1858,1884]" pageId="5" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype, ♂ (ICN- MD-1317B).
|
||
<emphasis id="B9117D17FFC73C72970EF8F3346EF8A3" bold="true" box="[353,374,1892,1918]" pageId="5" pageNumber="70">A</emphasis>
|
||
. Body rings 9
|
||
<superScript id="7C100C4DFFC73C729447F8F3372DF8AF" attach="left" box="[552,565,1892,1906]" fontSize="6" pageId="5" pageNumber="70">th</superScript>
|
||
and 10
|
||
<superScript id="7C100C4DFFC73C7294FBF8F337B9F8AF" attach="right" box="[660,673,1892,1906]" fontSize="6" pageId="5" pageNumber="70">th</superScript>
|
||
, red rectangle = median metazonal sulcus.
|
||
<emphasis id="B9117D17FFC73C7292D6F8F231D5F8A2" bold="true" box="[1209,1229,1893,1919]" pageId="5" pageNumber="70">B</emphasis>
|
||
. Venter of 4
|
||
<superScript id="7C100C4DFFC73C729306F8F3306EF8AF" attach="none" box="[1385,1398,1892,1906]" fontSize="6" pageId="5" pageNumber="70">th</superScript>
|
||
body ring, showing paired sternal projections, red oval = stigma.
|
||
<emphasis id="B9117D17FFC73C7295BDF81F36FDF87F" bold="true" box="[978,997,1928,1954]" pageId="5" pageNumber="70">C</emphasis>
|
||
. Venter of head, showing narrow gnathochilarium.
|
||
<emphasis id="B9117D17FFC73C7297E3F83C34B9F818" bold="true" box="[396,417,1963,1989]" pageId="5" pageNumber="70">D</emphasis>
|
||
. Tip of seventh antennomere, red arrow = narrow slit connecting pocket containing sensilla basiconica and sensory cone area.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDAA105FFC43C719685FE9E34ADFEFE" blockId="6.[187,1399,265,643]" box="[234,437,265,291]" pageId="6" pageNumber="71">
|
||
<emphasis id="B9117D17FFC43C719685FE9E34ADFEFE" bold="true" box="[234,437,265,291]" pageId="6" pageNumber="71">Male characters</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDAA105FFC43C7196D2FEBB31D2FD5E" blockId="6.[187,1399,265,643]" pageId="6" pageNumber="71">
|
||
Gonopores without modifications, located on the coxae of the 2
|
||
<superScript id="7C100C4DFFC43C7195A9FEBC36CFFEE4" attach="left" box="[966,983,299,313]" fontSize="6" pageId="6" pageNumber="71">nd</superScript>
|
||
leg-pair, with a subterminal pore. Gonopod aperture: on 7
|
||
<superScript id="7C100C4DFFC43C71978BFED934E9FE81" attach="left" box="[484,497,334,348]" fontSize="6" pageId="6" pageNumber="71">th</superScript>
|
||
body ring; oval; rim notably projecting above sternite, with a U-shaped concavity on posterior margin, posterior edge without folds. Gonopods: gonocoxae length variable relative to telopodite length; largely obscured by telopodite in ectal view; without a spiniform process (
|
||
<figureCitation id="135EBD80FFC43C7196A9FE2F3429FE0E" box="[198,305,440,467]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="9.[189,232,970,996]" captionStartId-1="10.[189,232,1928,1954]" captionStartId-2="12.[188,231,1892,1918]" captionTargetBox-0="[189,1397,293,928]" captionTargetBox-1="[189,1398,1270,1881]" captionTargetBox-2="[190,1398,1234,1849]" captionTargetId-0="figure-255@9.[189,1398,287,928]" captionTargetId-1="figure-240@10.[189,1398,1266,1881]" captionTargetId-2="figure-296@12.[189,1398,1228,1849]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="12" captionText-0="Fig. 5. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810), left gonopod. A. Lateral view. B. Anterior view. C. Mesal view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." captionText-1="Fig. 6. Dibolostethus inopinatus Means, Bouzan & Ivanov sp. nov., holotype, ♂ (VMNH110813), left gonopod. A. Lateral view, red circle = cingulum. B. Anterior view. C. Mesal view. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." captionText-2="Fig. 7. Dibolostethus kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., holotype, ♂ (ICN- MD-1317A), right gonopod. A. Mesal view. B. Anterior view. C. Lateral view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: Lb = lobe; PfP = prefemoral process; S = solenomere; Sh = shelf." figureDoi-0="http://doi.org/10.5281/zenodo.8205510" figureDoi-1="http://doi.org/10.5281/zenodo.8205512" figureDoi-2="http://doi.org/10.5281/zenodo.8205514" httpUri-0="https://zenodo.org/record/8205510/files/figure.png" httpUri-1="https://zenodo.org/record/8205512/files/figure.png" httpUri-2="https://zenodo.org/record/8205514/files/figure.png" pageId="6" pageNumber="71">Figs 5–7</figureCitation>
|
||
). Cannula hook-shaped, without modifications (
|
||
<figureCitation id="135EBD80FFC43C719508FE2F36D0FE0E" box="[871,968,440,467]" captionStart="Fig" captionStartId="9.[189,232,970,996]" captionTargetBox="[189,1397,293,928]" captionTargetId="figure-255@9.[189,1398,287,928]" captionTargetPageId="9" captionText="Fig. 5. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810), left gonopod. A. Lateral view. B. Anterior view. C. Mesal view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." figureDoi="http://doi.org/10.5281/zenodo.8205510" httpUri="https://zenodo.org/record/8205510/files/figure.png" pageId="6" pageNumber="71">Figs 5C</figureCitation>
|
||
,
|
||
<figureCitation id="135EBD80FFC43C7195BAFE2F36E1FE0F" box="[981,1017,440,466]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[189,1398,1270,1881]" captionTargetId="figure-240@10.[189,1398,1266,1881]" captionTargetPageId="10" captionText="Fig. 6. Dibolostethus inopinatus Means, Bouzan & Ivanov sp. nov., holotype, ♂ (VMNH110813), left gonopod. A. Lateral view, red circle = cingulum. B. Anterior view. C. Mesal view. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." figureDoi="http://doi.org/10.5281/zenodo.8205512" httpUri="https://zenodo.org/record/8205512/files/figure.png" pageId="6" pageNumber="71">6C</figureCitation>
|
||
,
|
||
<figureCitation id="135EBD80FFC43C719268FE2E3137FE0E" box="[1031,1071,441,467]" captionStart="Fig" captionStartId="12.[188,231,1892,1918]" captionTargetBox="[190,1398,1234,1849]" captionTargetId="figure-296@12.[189,1398,1228,1849]" captionTargetPageId="12" captionText="Fig. 7. Dibolostethus kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., holotype, ♂ (ICN- MD-1317A), right gonopod. A. Mesal view. B. Anterior view. C. Lateral view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: Lb = lobe; PfP = prefemoral process; S = solenomere; Sh = shelf." figureDoi="http://doi.org/10.5281/zenodo.8205514" httpUri="https://zenodo.org/record/8205514/files/figure.png" pageId="6" pageNumber="71">7A</figureCitation>
|
||
). Solenomere long, slender, unbranched, apex falciform (
|
||
<figureCitation id="135EBD80FFC43C719464FE4B376CFE2B" box="[523,628,475,502]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="9.[189,232,970,996]" captionStartId-1="10.[189,232,1928,1954]" captionStartId-2="12.[188,231,1892,1918]" captionTargetBox-0="[189,1397,293,928]" captionTargetBox-1="[189,1398,1270,1881]" captionTargetBox-2="[190,1398,1234,1849]" captionTargetId-0="figure-255@9.[189,1398,287,928]" captionTargetId-1="figure-240@10.[189,1398,1266,1881]" captionTargetId-2="figure-296@12.[189,1398,1228,1849]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="12" captionText-0="Fig. 5. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810), left gonopod. A. Lateral view. B. Anterior view. C. Mesal view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." captionText-1="Fig. 6. Dibolostethus inopinatus Means, Bouzan & Ivanov sp. nov., holotype, ♂ (VMNH110813), left gonopod. A. Lateral view, red circle = cingulum. B. Anterior view. C. Mesal view. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." captionText-2="Fig. 7. Dibolostethus kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., holotype, ♂ (ICN- MD-1317A), right gonopod. A. Mesal view. B. Anterior view. C. Lateral view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: Lb = lobe; PfP = prefemoral process; S = solenomere; Sh = shelf." figureDoi-0="http://doi.org/10.5281/zenodo.8205510" figureDoi-1="http://doi.org/10.5281/zenodo.8205512" figureDoi-2="http://doi.org/10.5281/zenodo.8205514" httpUri-0="https://zenodo.org/record/8205510/files/figure.png" httpUri-1="https://zenodo.org/record/8205512/files/figure.png" httpUri-2="https://zenodo.org/record/8205514/files/figure.png" pageId="6" pageNumber="71">Figs 5–7</figureCitation>
|
||
). Prostatic groove visible along the entire length of the solenomere in mesal view, without a trace of torsion (
|
||
<figureCitation id="135EBD80FFC43C7194CFFE683618FDC7" box="[672,768,511,538]" captionStart="Fig" captionStartId="9.[189,232,970,996]" captionTargetBox="[189,1397,293,928]" captionTargetId="figure-255@9.[189,1398,287,928]" captionTargetPageId="9" captionText="Fig. 5. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810), left gonopod. A. Lateral view. B. Anterior view. C. Mesal view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." figureDoi="http://doi.org/10.5281/zenodo.8205510" httpUri="https://zenodo.org/record/8205510/files/figure.png" pageId="6" pageNumber="71">Figs 5C</figureCitation>
|
||
,
|
||
<figureCitation id="135EBD80FFC43C719563FE683629FDC4" box="[780,817,511,537]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[189,1398,1270,1881]" captionTargetId="figure-240@10.[189,1398,1266,1881]" captionTargetPageId="10" captionText="Fig. 6. Dibolostethus inopinatus Means, Bouzan & Ivanov sp. nov., holotype, ♂ (VMNH110813), left gonopod. A. Lateral view, red circle = cingulum. B. Anterior view. C. Mesal view. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." figureDoi="http://doi.org/10.5281/zenodo.8205512" httpUri="https://zenodo.org/record/8205512/files/figure.png" pageId="6" pageNumber="71">6C</figureCitation>
|
||
,
|
||
<figureCitation id="135EBD80FFC43C719552FE68367CFDC4" box="[829,868,511,537]" captionStart="Fig" captionStartId="12.[188,231,1892,1918]" captionTargetBox="[190,1398,1234,1849]" captionTargetId="figure-296@12.[189,1398,1228,1849]" captionTargetPageId="12" captionText="Fig. 7. Dibolostethus kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., holotype, ♂ (ICN- MD-1317A), right gonopod. A. Mesal view. B. Anterior view. C. Lateral view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: Lb = lobe; PfP = prefemoral process; S = solenomere; Sh = shelf." figureDoi="http://doi.org/10.5281/zenodo.8205514" httpUri="https://zenodo.org/record/8205514/files/figure.png" pageId="6" pageNumber="71">7A</figureCitation>
|
||
, dashed lines). Prefemoral region (
|
||
<figureCitation id="135EBD80FFC43C719293FE683070FDC7" box="[1276,1384,511,538]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="9.[189,232,970,996]" captionStartId-1="10.[189,232,1928,1954]" captionStartId-2="12.[188,231,1892,1918]" captionTargetBox-0="[189,1397,293,928]" captionTargetBox-1="[189,1398,1270,1881]" captionTargetBox-2="[190,1398,1234,1849]" captionTargetId-0="figure-255@9.[189,1398,287,928]" captionTargetId-1="figure-240@10.[189,1398,1266,1881]" captionTargetId-2="figure-296@12.[189,1398,1228,1849]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="12" captionText-0="Fig. 5. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810), left gonopod. A. Lateral view. B. Anterior view. C. Mesal view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." captionText-1="Fig. 6. Dibolostethus inopinatus Means, Bouzan & Ivanov sp. nov., holotype, ♂ (VMNH110813), left gonopod. A. Lateral view, red circle = cingulum. B. Anterior view. C. Mesal view. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." captionText-2="Fig. 7. Dibolostethus kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., holotype, ♂ (ICN- MD-1317A), right gonopod. A. Mesal view. B. Anterior view. C. Lateral view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: Lb = lobe; PfP = prefemoral process; S = solenomere; Sh = shelf." figureDoi-0="http://doi.org/10.5281/zenodo.8205510" figureDoi-1="http://doi.org/10.5281/zenodo.8205512" figureDoi-2="http://doi.org/10.5281/zenodo.8205514" httpUri-0="https://zenodo.org/record/8205510/files/figure.png" httpUri-1="https://zenodo.org/record/8205512/files/figure.png" httpUri-2="https://zenodo.org/record/8205514/files/figure.png" pageId="6" pageNumber="71">Figs 5–7</figureCitation>
|
||
): small, about ⅓ the length of the telopodite; a slight cingulum demarcates the transition zone between the prefemoral and solenomere regions (
|
||
<figureCitation id="135EBD80FFC43C719405FDD237CCFDBD" box="[618,724,581,608]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="9.[189,232,970,996]" captionStartId-1="10.[189,232,1928,1954]" captionStartId-2="12.[188,231,1892,1918]" captionTargetBox-0="[189,1397,293,928]" captionTargetBox-1="[189,1398,1270,1881]" captionTargetBox-2="[190,1398,1234,1849]" captionTargetId-0="figure-255@9.[189,1398,287,928]" captionTargetId-1="figure-240@10.[189,1398,1266,1881]" captionTargetId-2="figure-296@12.[189,1398,1228,1849]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="12" captionText-0="Fig. 5. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810), left gonopod. A. Lateral view. B. Anterior view. C. Mesal view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." captionText-1="Fig. 6. Dibolostethus inopinatus Means, Bouzan & Ivanov sp. nov., holotype, ♂ (VMNH110813), left gonopod. A. Lateral view, red circle = cingulum. B. Anterior view. C. Mesal view. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." captionText-2="Fig. 7. Dibolostethus kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., holotype, ♂ (ICN- MD-1317A), right gonopod. A. Mesal view. B. Anterior view. C. Lateral view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: Lb = lobe; PfP = prefemoral process; S = solenomere; Sh = shelf." figureDoi-0="http://doi.org/10.5281/zenodo.8205510" figureDoi-1="http://doi.org/10.5281/zenodo.8205512" figureDoi-2="http://doi.org/10.5281/zenodo.8205514" httpUri-0="https://zenodo.org/record/8205510/files/figure.png" httpUri-1="https://zenodo.org/record/8205512/files/figure.png" httpUri-2="https://zenodo.org/record/8205514/files/figure.png" pageId="6" pageNumber="71">Figs 5–7</figureCitation>
|
||
, red ovals). Prefemoral process: long (similar in length to solenomere); apex broad, with a series of distinct lobes and lamellate flanges (
|
||
<figureCitation id="135EBD80FFC43C71923EFDFF31A4FD5E" box="[1105,1212,616,643]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="9.[189,232,970,996]" captionStartId-1="10.[189,232,1928,1954]" captionStartId-2="12.[188,231,1892,1918]" captionTargetBox-0="[189,1397,293,928]" captionTargetBox-1="[189,1398,1270,1881]" captionTargetBox-2="[190,1398,1234,1849]" captionTargetId-0="figure-255@9.[189,1398,287,928]" captionTargetId-1="figure-240@10.[189,1398,1266,1881]" captionTargetId-2="figure-296@12.[189,1398,1228,1849]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="12" captionText-0="Fig. 5. Dibolostethus sicarius Hoffman, 2009, holotype, ♂ (VMNH110810), left gonopod. A. Lateral view. B. Anterior view. C. Mesal view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." captionText-1="Fig. 6. Dibolostethus inopinatus Means, Bouzan & Ivanov sp. nov., holotype, ♂ (VMNH110813), left gonopod. A. Lateral view, red circle = cingulum. B. Anterior view. C. Mesal view. Dashed lines = prostatic groove. Abbreviations: PfP = prefemoral process; S = solenomere." captionText-2="Fig. 7. Dibolostethus kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., holotype, ♂ (ICN- MD-1317A), right gonopod. A. Mesal view. B. Anterior view. C. Lateral view. Red circles = cingulum. Dashed lines = prostatic groove. Abbreviations: Lb = lobe; PfP = prefemoral process; S = solenomere; Sh = shelf." figureDoi-0="http://doi.org/10.5281/zenodo.8205510" figureDoi-1="http://doi.org/10.5281/zenodo.8205512" figureDoi-2="http://doi.org/10.5281/zenodo.8205514" httpUri-0="https://zenodo.org/record/8205510/files/figure.png" httpUri-1="https://zenodo.org/record/8205512/files/figure.png" httpUri-2="https://zenodo.org/record/8205514/files/figure.png" pageId="6" pageNumber="71">Figs 5–7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDAA105FFC43C719685FD3D34C9FD19" blockId="6.[188,1398,682,814]" box="[234,465,682,708]" pageId="6" pageNumber="71">
|
||
<emphasis id="B9117D17FFC43C719685FD3D34C9FD19" bold="true" box="[234,465,682,708]" pageId="6" pageNumber="71">Female characters</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDAA105FFC43C7196D2FD593157FCF3" blockId="6.[188,1398,682,814]" pageId="6" pageNumber="71">
|
||
Known only for
|
||
<taxonomicName id="4C65DA86FFC43C719713FD5934E4FD3A" authorityName="Hoffman" authorityYear="2009" box="[380,508,718,743]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="71" phylum="Arthropoda" rank="species" species="sicarius">
|
||
<emphasis id="B9117D17FFC43C719713FD5934E4FD3A" box="[380,508,718,743]" italics="true" pageId="6" pageNumber="71">D. sicarius</emphasis>
|
||
</taxonomicName>
|
||
. Vulvae: small, oval-shaped, unmodified (
|
||
<figureCitation id="135EBD80FFC43C719586FD5A315EFD3A" box="[1001,1094,717,743]" captionStart="Fig" captionStartId="13.[189,232,1928,1954]" captionTargetBox="[190,1397,1511,1885]" captionTargetId="figure-343@13.[189,1398,1505,1885]" captionTargetPageId="13" captionText="Fig. 8. Dibolostethus sicarius Hoffman, 2009, paratype, ♀ (VMNH110811), vulvae. A. Left vulva, lateroventral view, with operculum at base. B. Vulvae in situ, held appressed to 2nd leg pair coxae with valve openings facing each other. C. Left vulva, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8205516" httpUri="https://zenodo.org/record/8205516/files/figure.png" pageId="6" pageNumber="71">Fig. 8A</figureCitation>
|
||
); held against coxae of 2
|
||
<superScript id="7C100C4DFFC43C71930AFD5B306EFD07" attach="none" box="[1381,1398,716,730]" fontSize="6" pageId="6" pageNumber="71">nd</superScript>
|
||
leg pair within deep emargination of 3
|
||
<superScript id="7C100C4DFFC43C719415FD673790FD23" attach="left" box="[634,648,752,766]" fontSize="6" pageId="6" pageNumber="71">rd</superScript>
|
||
segment, not protruding beyond U-shaped vulvar rim (
|
||
<figureCitation id="135EBD80FFC43C719363FD67307FFCD7" box="[1292,1383,752,778]" captionStart="Fig" captionStartId="13.[189,232,1928,1954]" captionTargetBox="[190,1397,1511,1885]" captionTargetId="figure-343@13.[189,1398,1505,1885]" captionTargetPageId="13" captionText="Fig. 8. Dibolostethus sicarius Hoffman, 2009, paratype, ♀ (VMNH110811), vulvae. A. Left vulva, lateroventral view, with operculum at base. B. Vulvae in situ, held appressed to 2nd leg pair coxae with valve openings facing each other. C. Left vulva, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8205516" httpUri="https://zenodo.org/record/8205516/files/figure.png" pageId="6" pageNumber="71">Fig. 8B</figureCitation>
|
||
); composed of two subequal valves and a small operculum; setose (
|
||
<figureCitation id="135EBD80FFC43C7195AEFC84315AFCF3" box="[961,1090,787,814]" captionStart="Fig" captionStartId="13.[189,232,1928,1954]" captionTargetBox="[190,1397,1511,1885]" captionTargetId="figure-343@13.[189,1398,1505,1885]" captionTargetPageId="13" captionText="Fig. 8. Dibolostethus sicarius Hoffman, 2009, paratype, ♀ (VMNH110811), vulvae. A. Left vulva, lateroventral view, with operculum at base. B. Vulvae in situ, held appressed to 2nd leg pair coxae with valve openings facing each other. C. Left vulva, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8205516" httpUri="https://zenodo.org/record/8205516/files/figure.png" pageId="6" pageNumber="71">Fig. 8A, C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37FF28EFFC43C7196D2FCC23777FA70" pageId="6" pageNumber="71" type="discussion">
|
||
<paragraph id="8BDAA105FFC43C7196D2FCC23437FCB2" blockId="6.[189,303,853,879]" box="[189,303,853,879]" pageId="6" pageNumber="71">
|
||
<heading id="D0921669FFC43C7196D2FCC23437FCB2" bold="true" box="[189,303,853,879]" fontSize="11" level="3" pageId="6" pageNumber="71" reason="3">
|
||
<emphasis id="B9117D17FFC43C7196D2FCC23437FCB2" bold="true" box="[189,303,853,879]" pageId="6" pageNumber="71">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDAA105FFC43C7196D2FC143777FA70" blockId="6.[188,1401,899,1454]" pageId="6" pageNumber="71">
|
||
Although the structure of
|
||
<taxonomicName id="4C65DA86FFC43C71979BFC14378FFC40" authorityName="Hoffman" authorityYear="2009" box="[500,663,899,925]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="71" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9117D17FFC43C71979BFC14378FFC40" box="[500,663,899,925]" italics="true" pageId="6" pageNumber="71">Dibolostethus</emphasis>
|
||
</taxonomicName>
|
||
gonopods (i.e., presence of a prefemoral process and a hook-shaped canula) shows definitive placement within the
|
||
<taxonomicName id="4C65DA86FFC43C719558FC3036F7FC1C" authorityName="Cook" authorityYear="1895" box="[823,1007,935,961]" class="Diplopoda" family="Chelodesmidae" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="71" phylum="Arthropoda" rank="family">Chelodesmidae</taxonomicName>
|
||
, all known
|
||
<taxonomicName id="4C65DA86FFC43C719219FC303006FC1C" authorityName="Hoffman" authorityYear="2009" box="[1142,1310,935,961]" class="Diplopoda" family="Chelodesmidae" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="68" phylum="Arthropoda" rank="tribe" tribe="Dibolostethini">Dibolostethini</taxonomicName>
|
||
exhibit striking similarities with members of
|
||
<taxonomicName id="4C65DA86FFC43C7194EDFC5D3674FC39" authorityName="Daday" authorityYear="1889" box="[642,876,970,996]" class="Diplopoda" family="Paradoxosomatidae" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="71" phylum="Arthropoda" rank="family">Paradoxosomatidae</taxonomicName>
|
||
. These include the presence of a transverse metatergal sulcus, reduction of paranota and constriction between the pro- and metazonites giving the body a moniliform appearance (shared with several other groups of
|
||
<taxonomicName id="4C65DA86FFC43C719594FB8731ADFBF7" authorityName="Cook" authorityYear="1895" box="[1019,1205,1040,1066]" class="Diplopoda" family="Chelodesmidae" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="71" phylum="Arthropoda" rank="family">Chelodesmidae</taxonomicName>
|
||
), elongated legs and antennae, and the positioning of the dorsal pair of the macrosetae in relation to the mesal lips of the paraproct (
|
||
<figureCitation id="135EBD80FFC43C719701FBC034A1FBAC" box="[366,441,1111,1137]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[189,1398,642,1776]" captionTargetId="figure-157@5.[189,1398,642,1776]" captionTargetPageId="5" captionText="Fig. 2. Scanning electron microscope images of Dibolostethus Hoffman, 2009 showing somatic characters. Dibolostethus kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov., paratype, ♂ (ICN- MD-1317B). A. Body rings 9th and 10th, red rectangle = median metazonal sulcus. B. Venter of 4th body ring, showing paired sternal projections, red oval = stigma. C. Venter of head, showing narrow gnathochilarium. D. Tip of seventh antennomere, red arrow = narrow slit connecting pocket containing sensilla basiconica and sensory cone area." figureDoi="http://doi.org/10.5281/zenodo.8205504" httpUri="https://zenodo.org/record/8205504/files/figure.png" pageId="6" pageNumber="71">Fig. 2</figureCitation>
|
||
). In the
|
||
<taxonomicName id="4C65DA86FFC43C71944DFBC03613FBAC" authorityName="Daday" authorityYear="1889" box="[546,779,1111,1137]" class="Diplopoda" family="Paradoxosomatidae" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="71" phylum="Arthropoda" rank="family">Paradoxosomatidae</taxonomicName>
|
||
, these macrosetae are positioned far away from the mesal lips of the paraproct, and located directly above the ventral pair of macrosetae, forming the four corners of a rectangle (
|
||
<bibRefCitation id="EFF4DCF4FFC43C7197A5FB0A365DFB6A" author="Nguyen & Sierwald" box="[458,837,1181,1207]" firstAuthor="Nguyen" page="1147" pageId="6" pageNumber="71" pagination="1132 - 1353" refId="ref9850" refString="Nguyen A. D. & Sierwald P. 2013. A worldwide catalog of the family Paradoxosomatidae Daday, 1889 (Diplopoda: Polydesmida). Check List 9 (6): 1132 - 1353. https: // doi. org / 10.15560 / 9.6.1132" type="journal article" year="2013">Nguyen & Sierwald 2013: 1147</bibRefCitation>
|
||
). In the vast majority of the
|
||
<taxonomicName id="4C65DA86FFC43C7192FFFB0A3050FB6A" authorityName="Cook" authorityYear="1895" box="[1168,1352,1181,1207]" class="Diplopoda" family="Chelodesmidae" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="71" phylum="Arthropoda" rank="family">Chelodesmidae</taxonomicName>
|
||
, the dorsal pair of macrosetae is located directly on the mesal lips of the paraproct, although this character is somewhat variable. For example, in the Lepturodesmini
|
||
<bibRefCitation id="EFF4DCF4FFC43C719533FB74310AFB23" author="Hoffman" box="[860,1042,1251,1278]" firstAuthor="Hoffman" pageId="6" pageNumber="71" pagination="127 - 144" refId="ref9265" refString="Hoffman R. L. 1975. Chelodesmid studies VI. A synopsis of the tribe Trachelodesmini (Diplopoda: Polydesmida). Studies on Neotropical Fauna 10 (2): 127 - 144. https: // doi. org / 10.1080 / 01650527509360488" type="journal article" year="1975">Hoffman, 1975</bibRefCitation>
|
||
, the dorsal pair of macrosetae is immediately adjacent to the mesal lips of the paraproct but not mounted on them. In species of
|
||
<taxonomicName id="4C65DA86FFC43C7196D2FABD3478FA99" authorityName="Hoffman" authorityYear="2009" box="[189,352,1322,1348]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="71" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9117D17FFC43C7196D2FABD3478FA99" box="[189,352,1322,1348]" italics="true" pageId="6" pageNumber="71">Dibolostethus</emphasis>
|
||
</taxonomicName>
|
||
, the two pairs of macrosetae are positioned far away and equidistant from the mesal lips of the paraproct as in the
|
||
<taxonomicName id="4C65DA86FFC43C7197ADFADA37B4FABA" authorityName="Daday" authorityYear="1889" box="[450,684,1357,1383]" class="Diplopoda" family="Paradoxosomatidae" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="71" phylum="Arthropoda" rank="family">Paradoxosomatidae</taxonomicName>
|
||
. The elongated antennae and legs (similar to the chelodesmid
|
||
<taxonomicName id="4C65DA86FFC43C7196D3FAE73462FA57" authorityName="Hoffman" authorityYear="1979" box="[188,378,1392,1418]" class="Diplopoda" family="Chelodesmidae" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="84" phylum="Arthropoda" rank="tribe" tribe="Trichomorphini">Trichomorphini</taxonomicName>
|
||
), and the presence of an adenostyle-like femoral process on leg pair 4 also resemble many species of
|
||
<taxonomicName id="4C65DA86FFC43C7197EEFA043773FA70" authorityName="Daday" authorityYear="1889" box="[385,619,1427,1453]" class="Diplopoda" family="Paradoxosomatidae" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="71" phylum="Arthropoda" rank="family">Paradoxosomatidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37FF28EFFC43C7196D2FA4236C1F9C3" box="[189,985,1493,1566]" pageId="6" pageNumber="71" type="distribution">
|
||
<paragraph id="8BDAA105FFC43C7196D2FA42344EFA32" blockId="6.[189,342,1493,1519]" box="[189,342,1493,1519]" pageId="6" pageNumber="71">
|
||
<heading id="D0921669FFC43C7196D2FA42344EFA32" bold="true" box="[189,342,1493,1519]" fontSize="11" level="3" pageId="6" pageNumber="71" reason="3">
|
||
<emphasis id="B9117D17FFC43C7196D2FA42344EFA32" bold="true" box="[189,342,1493,1519]" pageId="6" pageNumber="71">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDAA105FFC43C7196D2F99336C1F9C3" blockId="6.[189,985,1539,1566]" box="[189,985,1539,1566]" pageId="6" pageNumber="71">
|
||
Known from the Andean regions of
|
||
<collectingCountry id="F372E195FFC43C71940AF99437C1F9C0" box="[613,729,1539,1565]" name="Colombia" pageId="6" pageNumber="71">Colombia</collectingCountry>
|
||
and
|
||
<collectingCountry id="F372E195FFC43C71957EF994366BF9C0" box="[785,883,1539,1565]" name="Ecuador" pageId="6" pageNumber="71">Ecuador</collectingCountry>
|
||
(
|
||
<figureCitation id="135EBD80FFC43C7195ECF99436D3F9C3" box="[899,971,1539,1566]" captionStart="Fig" captionStartId="14.[188,231,1892,1918]" captionTargetBox="[243,1357,497,1838]" captionTargetId="figure-72@14.[212,1380,484,1853]" captionTargetPageId="14" captionText="Fig. 9. Distribution of Chelodesmidae spp. in the Tropical Andes Biodiversity Hotspot, by country. Note: the type locality of Dibolostethus sicarius Hoffman, 2009 is included for completeness although it falls just outside of the boundaries of the Tropical Andes Biodiversity Hotspot. For information on these taxa see Supp. file 1." figureDoi="http://doi.org/10.5281/zenodo.8205518" httpUri="https://zenodo.org/record/8205518/files/figure.png" pageId="6" pageNumber="71">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37FF28EFFC43C7196D2F9D231BEF953" box="[189,1190,1605,1678]" pageId="6" pageNumber="71" type="taxon_list">
|
||
<paragraph id="8BDAA105FFC43C7196D2F9D23445F982" blockId="6.[189,349,1605,1631]" box="[189,349,1605,1631]" pageId="6" pageNumber="71">
|
||
<heading id="D0921669FFC43C7196D2F9D23445F982" bold="true" box="[189,349,1605,1631]" fontSize="11" level="3" pageId="6" pageNumber="71" reason="3">
|
||
<emphasis id="B9117D17FFC43C7196D2F9D23445F982" bold="true" box="[189,349,1605,1631]" pageId="6" pageNumber="71">Composition</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDAA105FFC43C7196D2F9E431BEF953" blockId="6.[189,1190,1651,1678]" box="[189,1190,1651,1678]" pageId="6" pageNumber="71">
|
||
<taxonomicName id="4C65DA86FFC43C7196D2F9E4379AF953" authority="Hoffman, 2009" authorityName="Hoffman" authorityYear="2009" box="[189,642,1651,1678]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="71" phylum="Arthropoda" rank="species" species="sicarius">
|
||
<emphasis id="B9117D17FFC43C7196D2F9E434DCF950" box="[189,452,1651,1677]" italics="true" pageId="6" pageNumber="71">Dibolostethus sicarius</emphasis>
|
||
<bibRefCitation id="EFF4DCF4FFC43C7197A3F9E4379AF953" author="Hoffman" box="[460,642,1651,1678]" firstAuthor="Hoffman" pageId="6" pageNumber="71" pagination="149 - 154" refId="ref9561" refString="Hoffman R. L. 2009. Chelodesmid studies. XXVI. A new genus and tribe of chelodesmid millipeds from Ecuador (Polydesmida: Chelodesmidae). International Journal of Myriapodology 2: 149 - 154. https: // doi. org / 10.1163 / 187525409 X 12577705044584" type="journal article" year="2009">Hoffman, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C65DA86FFC43C7194E2F9E33635F950" authority="Means & Bouzan & Iniesta & Martínez-Torres & Vasquez-Valverde & Brescovit & Ivanov, 2023" authorityName="Means & Bouzan & Iniesta & Martínez-Torres & Vasquez-Valverde & Brescovit & Ivanov" authorityYear="2023" box="[653,813,1652,1677]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="71" phylum="Arthropoda" rank="species" species="inopinatus" status="sp. nov.">
|
||
<emphasis id="B9117D17FFC43C7194E2F9E33635F950" box="[653,813,1652,1677]" italics="true" pageId="6" pageNumber="71">D. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A222C06CFFC43C71955BF9E33689F953" box="[820,913,1652,1678]" pageId="6" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, and
|
||
<taxonomicName id="4C65DA86FFC43C7195A1F9E3315BF950" authority="Means & Bouzan & Iniesta & Martínez-Torres & Vasquez-Valverde & Brescovit & Ivanov, 2023" authorityName="Means & Bouzan & Iniesta & Martínez-Torres & Vasquez-Valverde & Brescovit & Ivanov" authorityYear="2023" box="[974,1091,1651,1677]" class="Diplopoda" family="Chelodesmidae" genus="Dibolostethus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="71" phylum="Arthropoda" rank="species" species="kattani" status="sp. nov.">
|
||
<emphasis id="B9117D17FFC43C7195A1F9E3315BF950" box="[974,1091,1651,1677]" italics="true" pageId="6" pageNumber="71">D. kattani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A222C06CFFC43C719224F9E331BEF953" box="[1099,1190,1652,1678]" pageId="6" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |