126 lines
14 KiB
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126 lines
14 KiB
XML
<document id="0D7B23D07F5F8CA5F040813207498767" ID-DOI="10.5281/zenodo.5400520" ID-ISSN="1638-9387" ID-Zenodo-Dep="5400520" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630333700421" checkinUser="marcus" docAuthor="Pisera, Andrzej" docDate="2000" docId="03C6E07B1324FFC2BED56B596F42F938" docLanguage="en" docName="Zoosystema.2000.22.2.285-298.pdf" docOrigin="Zoosystema 22 (2)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Theonella ukrainica Pisera 2000, n. sp." docType="treatment" docVersion="4" lastPageNumber="288" masterDocId="FFFF98031327FFC1BE386E206C5BFF90" masterDocTitle="New species of lithistid sponges from the Paleogene of the Ukraine" masterLastPageNumber="298" masterPageNumber="285" pageNumber="288" updateTime="1699232094796" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="B9BCCD95EB97CBA3ADB722CBF58DB0B1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="D3DF375218685B01D6903364BA42F3F7">
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<mods:title id="357D8E34F763BC3F8584FD41843FEE6D">New species of lithistid sponges from the Paleogene of the Ukraine</mods:title>
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</mods:titleInfo>
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<mods:name id="E02FD71A65276ADBA5EC544EAB8D9504" type="personal">
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<mods:role id="1BEF2686DF1F79E9AC1313AF0A250CF3">
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<mods:roleTerm id="F895DB186BDC013A090231D93DC4EEC0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6F993BC06A1931B0E90483C876145C06">Pisera, Andrzej</mods:namePart>
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<mods:affiliation id="09AD23EFAABB73BB96A3E578E68C4F87">Institute of Paleobiology, Polish Academy of Sciences, ul. Twarda 51 / 55, 00 - 818 Warszawa (Poland) apis @ twarda. pan. pl</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="19EF996EB263A619ADEA82B1AFC0365C">text</mods:typeOfResource>
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<mods:relatedItem id="56A640AF3F5B6E4E261A68BE181A303E" type="host">
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<mods:titleInfo id="D5E7A5B538FF8F771A305EC1381E4C3D">
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<mods:title id="3A8938E8FDE2676822E03F8BD2BBB66E">Zoosystema</mods:title>
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</mods:titleInfo>
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<mods:part id="2C37B0739E50053806E4B55BEE521253">
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<mods:date id="0BA8380AF0F7C7AD80C044CBE7612D10">2000</mods:date>
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<mods:detail id="322E7A2497CD1529DB6709D52219C9B1" type="volume">
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<mods:number id="F22E171AFB7C8A0B315366C7342F890D">22</mods:number>
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</mods:detail>
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<mods:detail id="FC398FA7FBFAEDA72C002A765CBED60F" type="issue">
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<mods:number id="C2D001C1950459463BE5601564104E72">2</mods:number>
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</mods:detail>
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<mods:extent id="F50957D0FDA73FCC63511088898D5736" unit="page">
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<mods:start id="2EA8B3A8DA156FFFD8C3BA0AFA4A6B59">285</mods:start>
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<mods:end id="8A5CEFD7EF6CBE8AF7B5287F66CE3AA3">298</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="D9310B2C6929C02C30E3A5D2B320DD86">journal article</mods:classification>
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<mods:identifier id="E06C338C2871DFBFED7E2F31683F3A61" type="DOI">10.5281/zenodo.5400520</mods:identifier>
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<mods:identifier id="2563EE64C57426FEA2F5790D826E37B4" type="ISSN">1638-9387</mods:identifier>
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<mods:identifier id="7D008E6EE3784435A7171124741C7E0B" type="Zenodo-Dep">5400520</mods:identifier>
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</mods:mods>
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<treatment id="03C6E07B1324FFC2BED56B596F42F938" LSID="urn:lsid:plazi:treatment:03C6E07B1324FFC2BED56B596F42F938" httpUri="http://treatment.plazi.org/id/03C6E07B1324FFC2BED56B596F42F938" lastPageNumber="288" pageId="3" pageNumber="288">
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<subSubSection id="C37502E61324FFC2BED56B596E50FA02" box="[237,523,1400,1427]" pageId="3" pageNumber="288" type="nomenclature">
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<paragraph id="8BD0516D1324FFC2BED56B596E50FA02" blockId="3.[237,523,1400,1457]" box="[237,523,1400,1427]" pageId="3" pageNumber="288">
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<heading id="D098E6011324FFC2BED56B596E50FA02" box="[237,523,1400,1427]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="288" reason="2">
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<taxonomicName id="4C6F2AEE1324FFC2BED56B596D93FA03" authority="Pisera, 2000" authorityName="Pisera" authorityYear="2000" box="[237,456,1401,1427]" class="Demospongiae" family="Theonellidae" genus="Theonella" kingdom="Animalia" order="Tetractinellida" pageId="3" pageNumber="288" phylum="Porifera" rank="species" species="ukrainica" status="sp. nov.">
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<emphasis id="B91B8D7F1324FFC2BED56B596D93FA03" bold="true" box="[237,456,1401,1427]" italics="true" pageId="3" pageNumber="288">Theonella ukrainica</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22830041324FFC2BFE86B586E50FA02" box="[464,523,1400,1426]" pageId="3" pageNumber="288" rank="species">n. sp.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37502E61324FFC2BF156BB76D90FA21" box="[301,459,1431,1457]" pageId="3" pageNumber="288" type="description">
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<paragraph id="8BD0516D1324FFC2BF156BB76D90FA21" blockId="3.[237,523,1400,1457]" box="[301,459,1431,1457]" pageId="3" pageNumber="288">
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(
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<figureCitation id="13544DE81324FFC2BF0C6BB76D2EFA21" box="[308,373,1431,1457]" captionStart="FIG" captionStartId="4.[114,125,1616,1633]" captionTargetBox="[144,1184,208,1579]" captionTargetId="figure-3@4.[144,1184,208,1582]" captionTargetPageId="4" captionText="FIG. 2. — Habit of the investigated sponges; A-D, Theonella ukrainica n. sp.; A, NHM ZN10D; B, holotype NHM ZN5A; C, NHM ZN5E; D, NHM ZN5F; E-H, Chenendopora piaskovskii n. sp.; E, F, paratype NHM ZN11 in lower side (E) and upper side (F) views; G, H, holotype NHM ZN1 in upper (G) and lower side (H) views; I-K, Lerouxia digitata n. sp.; I, NHM ZN4; J, holotype, NHM ZN7; K, paratype, NHM ZN8; L, Plinthosella magna n. sp., holotype NHM ZN3, oblique view. Scale bars: A-D, G, H, L, 1 cm; E, F, I-K, 2 cm." figureDoi="http://doi.org/10.5281/zenodo.5400524" httpUri="https://zenodo.org/record/5400524/files/figure.png" pageId="3" pageNumber="288">Figs 2</figureCitation>
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A-D; 3)
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</paragraph>
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</subSubSection>
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<subSubSection id="C37502E61324FFC2BE496BF26D37F98E" pageId="3" pageNumber="288" type="materials_examined">
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<paragraph id="8BD0516D1324FFC2BE496BF26D37F98E" blockId="3.[113,647,1490,1705]" pageId="3" pageNumber="288">
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MATERIAL. — 24 fragmentary specimens (NHM ZN5, ZN10).
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<typeStatus id="54D4EFCF1324FFC2BF376BCC6D2CF994" box="[271,375,1516,1540]" pageId="3" pageNumber="288" type="holotype">Holotype</typeStatus>
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, specimen NHM ZN5A;
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<typeStatus id="54D4EFCF1324FFC2BE4968256C9CF98D" box="[113,199,1541,1565]" pageId="3" pageNumber="288" type="paratype">paratype</typeStatus>
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, NHM ZN10A.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37502E61324FFC2BE4968116E41F9F3" pageId="3" pageNumber="288" type="etymology">
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<paragraph id="8BD0516D1324FFC2BE4968116E41F9F3" blockId="3.[113,647,1490,1705]" pageId="3" pageNumber="288">
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ETYMOLOGY. — The species name refers to the
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<collectingCountry id="F37811FD1324FFC2BE49686B6C9AF9F3" box="[113,193,1611,1635]" name="Ukraine" pageId="3" pageNumber="288">Ukraine</collectingCountry>
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where the material was discovered.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37502E61324FFC2BE4968576803FE88" pageId="3" pageNumber="288" type="diagnosis">
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<paragraph id="8BD0516D1324FFC2BE4968576EDCF937" blockId="3.[113,647,1490,1705]" pageId="3" pageNumber="288">
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DIAGNOSIS. — These are small branching
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<taxonomicName id="4C6F2AEE1324FFC2BC1F68576EDDF91F" authorityName="Gray" authorityYear="1867" box="[551,646,1655,1679]" class="Demospongiae" family="Theonellidae" genus="Theonella" kingdom="Animalia" order="Tetractinellida" pageId="3" pageNumber="288" phylum="Porifera" rank="genus">
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<emphasis id="B91B8D7F1324FFC2BC1F68576EDDF91F" box="[551,646,1655,1679]" italics="true" pageId="3" pageNumber="288">Theonella</emphasis>
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</taxonomicName>
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with one to several longitudinal canals in the centre.
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</paragraph>
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<paragraph id="8BD0516D1324FFC2BC9D6EEE6803FE88" blockId="3.[677,1210,205,280]" pageId="3" pageNumber="288">Choanosomal desmas as smooth regular tetraclones. Dermalia as phyllotriaenes with slender clads and pointed tips, or small irregular discotriaenes.</paragraph>
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</subSubSection>
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<subSubSection id="C37502E61324FFC2BC9D6F686EABFB2D" pageId="3" pageNumber="288" type="description">
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<paragraph id="8BD0516D1324FFC2BC9D6F686F6EFEF0" blockId="3.[677,1210,328,1213]" box="[677,821,328,354]" pageId="3" pageNumber="288">DESCRIPTION</paragraph>
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<paragraph id="8BD0516D1324FFC2BC9D6F466EABFB2D" blockId="3.[677,1210,328,1213]" pageId="3" pageNumber="288">
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Numerous fragments of small, up to
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<quantity id="4C97FC881324FFC2BA7A6F476821FE10" box="[1090,1146,358,385]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="288" unit="cm" value="5.0">5 cm</quantity>
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long, branching sponges with branches
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<quantity id="4C97FC881324FFC2BA226FA66829FE0F" box="[1050,1138,389,416]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="3" pageNumber="288" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6-8 mm</quantity>
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thick. One to several vertical, 0.5 to 1.0 mm wide, round canals run along the sponge in the centre. Sometimes these canals approach the lateral surface and then may be partly open. In the main trunk several canals always occur, in branches less or only one. These canals usually do not open at the tips of the branches. Branches usually rounded more rarely laterally flattened with rounded tips, when preserved. Skeleton consists of smooth, regular tetraclones, with moderately branched terminal articulations; they are 255- 430
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<emphasis id="B91B8D7F1324FFC2BCE56CD56EB0FC9E" box="[733,747,757,782]" italics="true" pageId="3" pageNumber="288">µ</emphasis>
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m in size. Dermal skeleton developed mostly as phyllotriaenes with slender pointed clads, which are 250-580
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<emphasis id="B91B8D7F1324FFC2BDEB6D136FBAFCDC" box="[979,993,819,844]" italics="true" pageId="3" pageNumber="288">µ</emphasis>
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m in size. In some areas, however, small discotriaenes that are irregular in outline also occur, and are 220-280
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<emphasis id="B91B8D7F1324FFC2BAAD6D5068F8FC19" box="[1173,1187,880,905]" italics="true" pageId="3" pageNumber="288">µ</emphasis>
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m in size. Choanosomal tetraclones are very regular in the upper and inner part of branches and measure 255-430
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<emphasis id="B91B8D7F1324FFC2BD476DEC6FD6FC75" box="[895,909,972,997]" italics="true" pageId="3" pageNumber="288">µ</emphasis>
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m in size, while in lower (older) parts, close to the surface, and around canals they are very densely packed and irregular. The space between dermal phyllotriaenes is often infilled in the older part of branches with very densely packed and tightly interlocking small, flat, irregular spicules resembling rhizoclones.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37502E61324FFC2BC9D6AC06F42F938" pageId="3" pageNumber="288" type="discussion">
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<paragraph id="8BD0516D1324FFC2BC9D6AC06F52FB68" blockId="3.[677,1210,1248,1704]" box="[677,777,1248,1274]" pageId="3" pageNumber="288">REMARKS</paragraph>
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<paragraph id="8BD0516D1324FFC2BC9D6ADF6F42F938" blockId="3.[677,1210,1248,1704]" pageId="3" pageNumber="288">
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The specimens included by me into
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<taxonomicName id="4C6F2AEE1324FFC2BA0D6ADF68E2FA89" authorityName="Pisera" authorityYear="2000" box="[1077,1209,1278,1305]" class="Demospongiae" family="Theonellidae" genus="Theonella" kingdom="Animalia" order="Tetractinellida" pageId="3" pageNumber="288" phylum="Porifera" rank="species" species="ukrainica">
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<emphasis id="B91B8D7F1324FFC2BA0D6ADF681CFA88" box="[1077,1095,1279,1304]" italics="true" pageId="3" pageNumber="288">T</emphasis>
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.
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<emphasis id="B91B8D7F1324FFC2BA6E6ADF68E2FA89" box="[1110,1209,1279,1305]" italics="true" pageId="3" pageNumber="288">ukrainica</emphasis>
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</taxonomicName>
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were most probably one of those included into
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<taxonomicName id="4C6F2AEE1324FFC2BC9D6B1D6F60FAC7" box="[677,827,1341,1367]" class="Demospongiae" family="Theonellidae" genus="Discodermia" kingdom="Animalia" order="Tetractinellida" pageId="3" pageNumber="288" phylum="Porifera" rank="genus">
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<emphasis id="B91B8D7F1324FFC2BC9D6B1D6F60FAC7" box="[677,827,1341,1367]" italics="true" pageId="3" pageNumber="288">Discodermia</emphasis>
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</taxonomicName>
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by Oakley (1942). The genus
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<taxonomicName id="4C6F2AEE1324FFC2BC9D6B7B6F4DFAE5" authorityName="Gray" authorityYear="1867" box="[677,790,1371,1397]" class="Demospongiae" family="Theonellidae" genus="Theonella" kingdom="Animalia" order="Tetractinellida" pageId="3" pageNumber="288" phylum="Porifera" rank="genus">
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<emphasis id="B91B8D7F1324FFC2BC9D6B7B6F4DFAE5" box="[677,790,1371,1397]" italics="true" pageId="3" pageNumber="288">Theonella</emphasis>
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</taxonomicName>
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, to my knowledge, has never been found as fossil before, except for an indication by Laubenfels (1955) which I was unable to resolve. Assignment of my material to
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<taxonomicName id="4C6F2AEE1324FFC2BDCE6B976807FA41" authorityName="Gray" authorityYear="1867" box="[1014,1116,1463,1489]" class="Demospongiae" family="Theonellidae" genus="Theonella" kingdom="Animalia" order="Tetractinellida" pageId="3" pageNumber="288" phylum="Porifera" rank="genus">
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<emphasis id="B91B8D7F1324FFC2BDCE6B976807FA41" box="[1014,1116,1463,1489]" italics="true" pageId="3" pageNumber="288">Theonella</emphasis>
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</taxonomicName>
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is based on the
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<typeStatus id="54D4EFCF1324FFC2BCCD6BF56F78FA7F" box="[757,803,1493,1519]" pageId="3" pageNumber="288">type</typeStatus>
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of desmas and their articulation and accompanying dermal phyllotriaenes. No microscleres have been found. Most theonellas have a spongocoel developed, but my specimens are tiny-branching in habit, and thus lack of spongocoel, instead several vertical canals are developed.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |