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<document id="F2F308D1AFA8E6A6F75068E24C829A14" ID-CLB-Dataset="39001" ID-DOI="10.11646/zootaxa.4114.4.3" ID-GBIF-Dataset="c1842fc0-8483-47f2-aa73-697840678713" ID-ISSN="1175-5326" ID-Zenodo-Dep="271605" ID-ZooBank="8A2A98DF-33B4-4B2C-BA3F-88B49EF3E5D2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1464158826947" checkinUser="plazi" docAuthor="Huang, Hao &amp; Zhu, Jian-Qing" docDate="2016" docId="03C587E1FFBBFFD419CB7820FC1117DF" docLanguage="en" docName="zootaxa.4114.4.3.pdf.imf" docOrigin="Zootaxa 4114 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ahlbergia chalybeia Leech" docType="treatment" docVersion="9" lastPageNumber="430" masterDocId="FFFCFF99FFAFFFC1195C7F38FFA1124D" masterDocTitle="Ahlbergia maoweiweii sp. n. from Shaanxi, China with revisional notes on similar species (Lepidoptera: Lycaenidae)" masterLastPageNumber="433" masterPageNumber="409" pageNumber="429" updateTime="1698662613831" updateUser="ExternalLinkService">
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<mods:title id="E53504148D8EA1B2E519A0A6698282FB">Ahlbergia maoweiweii sp. n. from Shaanxi, China with revisional notes on similar species (Lepidoptera: Lycaenidae)</mods:title>
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<mods:namePart id="8BD079023BE52DBCF726CB8F431F0B6E">Huang, Hao</mods:namePart>
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<mods:namePart id="E93683DCE679D1005151FD3D5BA19833">Zhu, Jian-Qing</mods:namePart>
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<mods:title id="513AD94D5D223DF4FC6E2E05B8C585A4">Zootaxa</mods:title>
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<mods:date id="2D8636677E0E6604CFEAC49CD8C3C5D2">2016</mods:date>
<mods:detail id="51F16063C2018EB04B8B76B24F0CD1B9" type="volume">
<mods:number id="D4CB13488FEBB717986D60B6E2A9CE9E">4114</mods:number>
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<mods:identifier id="2ACF7886405388531106B58B782CAE1F" type="DOI">10.11646/zootaxa.4114.4.3</mods:identifier>
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<treatment id="03C587E1FFBBFFD419CB7820FC1117DF" ID-DOI="http://doi.org/10.5281/zenodo.6080429" ID-GBIF-Taxon="120701279" ID-Zenodo-Dep="6080429" LSID="urn:lsid:plazi:treatment:03C587E1FFBBFFD419CB7820FC1117DF" httpUri="http://treatment.plazi.org/id/03C587E1FFBBFFD419CB7820FC1117DF" lastPageId="21" lastPageNumber="430" pageId="20" pageNumber="429">
<subSubSection id="C376657CFFBBFFD519CB7820FE3C151E" pageId="20" pageNumber="429" type="nomenclature">
<paragraph id="8BD336F7FFBBFFD519CB7820FE4E157F" blockId="20.[151,495,1816,1876]" box="[151,495,1816,1842]" pageId="20" pageNumber="429">
<heading id="D09B819BFFBBFFD519CB7820FE4E157F" bold="true" box="[151,495,1816,1842]" fontSize="11" level="1" pageId="20" pageNumber="429" reason="1">
<taxonomicName id="4C6C4D74FFBBFFD519CB7820FE4E157F" authority="Leech" authorityName="Leech" box="[151,495,1816,1842]" class="Insecta" family="Lycaenidae" genus="Ahlbergia" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="429" phylum="Arthropoda" rank="species" species="chalybeia">
<emphasis id="B918EAE5FFBBFFD519CB7820FE4E157F" bold="true" box="[151,495,1816,1842]" pageId="20" pageNumber="429">
<emphasis id="B918EAE5FFBBFFD519CB7820FE2A157F" bold="true" box="[151,395,1816,1842]" italics="true" pageId="20" pageNumber="429">Ahlbergia chalybeia</emphasis>
(Leech)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BD336F7FFBBFFD519CB7803FE3C151E" blockId="20.[151,495,1816,1876]" box="[151,413,1851,1876]" pageId="20" pageNumber="429">
(
<figureCitation id="13572A72FFBBFFD519C37803FE861519" box="[159,295,1851,1876]" captionStart="FIGURES 61 – 75" captionStartId="11.[151,264,1904,1927]" captionTargetBox="[151,1436,193,1881]" captionTargetId="figure@11.[151,1436,193,1883]" captionTargetPageId="11" captionText="FIGURES 61 – 75. Habitus of Ahlbergia females at same scale. 61 – 62 — A. maoweiweii, paratype. 63 – 66, 71 – 72 — A. clarolinea; 63 – 64 — holotype, Yulongxueshan; 65 – 66 — Ludian, Lijiang; 71 – 72 — Muli, Sichuan. 67 – 68 — A. chalcidis, Ludian, Lijiang. 69 – 70 — A. chalybeia, Jiulong. 73 – 75 — A. caesius, holotype. 61, 63, 65, 67, 69, 71, 73 — upperside; 62, 64, 66, 68, 70, 72, 74 — underside." httpUri="https://zenodo.org/record/271609/files/figure.png" pageId="20" pageNumber="429">Figs. 69–70</figureCitation>
,
<figureCitation id="13572A72FFBBFFD5186E7803FEFF151E" box="[306,350,1851,1875]" captionStart="FIGURES 94 – 104" captionStartId="15.[151,264,1462,1485]" captionTargetBox="[151,1436,193,1441]" captionTargetId="figure@15.[151,1436,193,1442]" captionTargetPageId="15" captionText="FIGURES 94 – 104. Female genitalia of Ahlbergia taxa in lateral view at same scale. 94 — A. maoweiweii, paratype (Figs. 31 – 32); 95 — A. leei, Liaoning (Figs. 35 – 36); 96 — A. ferrea, Liaoning (Fig. 43); 97 — A. tricaudata confusa, holotype (Fig. 45); 98 — A. luoliangi, paratype (Figs. 39 – 40); 99 — A. frivaldszkyi, Shaanxi (Fig. 38); 100 — A. frivaldszkyi, Tianjin (Fig. 37); 101 — A. leei ssp., Shaanxi (Figs. 41 – 42); 102 — A. chalcidis, Lijiang; 103 — A. chalybeia, Jiulong (Figs. 69 – 70); 104 — A. clarolinea, holotype (Figs. 63 – 64)." httpUri="https://zenodo.org/record/271613/files/figure.png" pageId="20" pageNumber="429">103</figureCitation>
,
<figureCitation id="13572A72FFBBFFD518367803FE35151E" box="[362,404,1851,1875]" captionStart="FIGURES 105 – 129" captionStartId="17.[151,264,1786,1809]" captionTargetBox="[151,1436,193,1764]" captionTargetId="figure@17.[151,1436,193,1765]" captionTargetPageId="17" captionText="FIGURES 105 – 129. Female genitalia of Ahlbergia taxa in ventral view at same scale (except for 109). 105 — A. maoweiweii, paratype (Figs. 31 – 32); 106 — A. ferrea, Liaoning (Fig. 43); 107 — A. luoliangi, paratype (Figs. 39 – 40); 108 — A. clarolinea, holotype (Figs. 63 – 64); 109 — A. chalybeia, lectotype, after Johnson (1992); 110 — A. chalybeia, Jiulong (Figs. 69 – 70); 111 — A. chalcidis, Lijiang; 112 — A. frivaldszkyi, Shaanxi (Fig. 38); 113 — A. tricaudata confusa, holotype (Fig. 45); 114 — A. tricaudata tricaudata, Liaoning (Fig. 44); 115 — A. tricaudata ssp., Shaanxi; 116 – 121, 126 – 129 — A. frivaldszkyi; 116 – 118, 127 – 129 — Shaanxi; 119 — Kangxian, Gansu (Figs. 48 – 49); 120 — Huixian, Gansu (Figs. 52 – 53); 121 — Wenxian, Gansu (Figs. 50 – 51); 126 — Tianjin (Fig. 37); 122 – 125 — A. leei; 122 — Liaoning (Figs. 35 – 36); 123 — Shaanxi (Figs. 41 – 42); 124 – 125 — Shaanxi." httpUri="https://zenodo.org/record/271614/files/figure.png" pageId="20" pageNumber="429">110</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C376657CFFBBFFD419CB78BBFDBB1281" lastPageId="21" lastPageNumber="430" pageId="20" pageNumber="429" type="reference_group">
<paragraph id="8BD336F7FFBBFFD519CB78BBFCA215D4" blockId="20.[151,1437,1923,2037]" box="[151,771,1923,1945]" pageId="20" pageNumber="429">
<taxonomicName id="4C6C4D74FFBBFFD519CB78BBFDAF15D4" authority="Leech, 1893: 355" authorityName="Leech" authorityPageNumber="355" authorityYear="1893" box="[151,526,1923,1945]" class="Insecta" family="Lycaenidae" genus="Satsuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="429" phylum="Arthropoda" rank="species" species="chalybeia">
<emphasis id="B918EAE5FFBBFFD519CB78BBFEF715D4" box="[151,342,1923,1945]" italics="true" pageId="20" pageNumber="429">Satsuma chalybeia</emphasis>
Leech, 1893: 355
</taxonomicName>
, TL: Changyang, Hubei
</paragraph>
<paragraph id="8BD336F7FFBBFFD519CB7899FDC615FA" blockId="20.[151,1437,1923,2037]" box="[151,615,1953,1975]" pageId="20" pageNumber="429">
<treatmentCitationGroup id="AB7C11D9FFBBFFD519CB7899FDC615FA" box="[151,615,1953,1975]" pageId="20" pageNumber="429">
<taxonomicName id="4C6C4D74FFBBFFD519CB7899FEF315FA" box="[151,338,1953,1975]" class="Insecta" family="Lycaenidae" genus="Incisalia" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="429" phylum="Arthropoda" rank="species" species="chalybea">
<emphasis id="B918EAE5FFBBFFD519CB7899FEF315FA" box="[151,338,1953,1975]" italics="true" pageId="20" pageNumber="429">Incisalia chalybea</emphasis>
</taxonomicName>
(sic!):
<treatmentCitation id="0ACD10E6FFBBFFD518C67899FDC215FA" author="Gillham" box="[410,611,1953,1975]" page="150" pageId="20" pageNumber="429" year="1955">
<bibRefCitation id="EFFD4B06FFBBFFD518C67899FD9115FA" author="Gillham" box="[410,560,1953,1975]" pageId="20" pageNumber="429" refString="Gillham, N. W. (1955) Incisalia Scudder, a Holarctic genus (Lepidoptera: Lycaenidae). Psyche, 62, 145 - 151. http: // dx. doi. org / 10.1155 / 1955 / 86218" type="journal article" year="1955">Gillham, 1955</bibRefCitation>
: 150
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD336F7FFBBFFD519CB78F8FDF315B8" blockId="20.[151,1437,1923,2037]" pageId="20" pageNumber="429">
<treatmentCitationGroup id="AB7C11D9FFBBFFD519CB78F8FDF315B8" pageId="20" pageNumber="429">
<taxonomicName id="4C6C4D74FFBBFFD519CB78F8FD92159B" authority="Johnson, 1992: 16" authorityName="Johnson" authorityPageNumber="16" authorityYear="1992" box="[151,563,1984,2006]" class="Insecta" family="Lycaenidae" genus="Ahlbergia" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="429" phylum="Arthropoda" rank="species" species="chalybeia">
<emphasis id="B918EAE5FFBBFFD519CB78F8FEC4159B" box="[151,357,1984,2006]" italics="true" pageId="20" pageNumber="429">Ahlbergia chalybeia</emphasis>
:
<treatmentCitation id="0ACD10E6FFBBFFD5182978F8FD92159B" author="Johnson" box="[373,563,1984,2006]" page="16" pageId="20" pageNumber="429" year="1992">
<bibRefCitation id="EFFD4B06FFBBFFD5182978F8FDAD159B" author="Johnson" box="[373,524,1984,2006]" pageId="20" pageNumber="429" refString="Johnson, K. (1992) The Palaearctic &quot; Elfin &quot; Butterflies (Lycaenidae, Theclinae). Neue Entomologische Nachrichten, 29, 1 - 141." type="journal article" year="1992">Johnson, 1992</bibRefCitation>
: 16
</treatmentCitation>
</taxonomicName>
,
<typeStatus id="54D78855FFBBFFD51B1C78F8FD3C159B" box="[576,669,1984,2006]" pageId="20" pageNumber="429" type="lectotype">lectotype</typeStatus>
designation, figs. 4, 5, 59 for habitus and genitalia; Huang &amp; Chen 2006: 319, discussion on sexual dimorphism.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD336F7FFBAFFD419CB7FA0FDBB1281" blockId="21.[151,1436,152,204]" pageId="21" pageNumber="430">
<treatmentCitationGroup id="AB7C11D9FFBAFFD419CB7FA0FDBB1281" pageId="21" pageNumber="430">
?
<taxonomicName id="4C6C4D74FFBAFFD419F47FA1FDE512E3" authority="Johnson, 1992: 67" authorityName="Johnson" authorityPageNumber="67" authorityYear="1992" box="[168,580,152,174]" class="Insecta" family="Lycaenidae" genus="Novosatsuma" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="430" phylum="Arthropoda" rank="species" species="cibdela">
<emphasis id="B918EAE5FFBAFFD419F47FA1FE2312E3" box="[168,386,152,174]" italics="true" pageId="21" pageNumber="430">Novosatsuma cibdela</emphasis>
<treatmentCitation id="0ACD10E6FFBAFFD418D47FA0FDE512E3" author="Johnson" box="[392,580,152,174]" page="67" pageId="21" pageNumber="430" year="1992">
<bibRefCitation id="EFFD4B06FFBAFFD418D47FA0FDBF12E3" author="Johnson" box="[392,542,152,174]" pageId="21" pageNumber="430" refString="Johnson, K. (1992) The Palaearctic &quot; Elfin &quot; Butterflies (Lycaenidae, Theclinae). Neue Entomologische Nachrichten, 29, 1 - 141." type="journal article" year="1992">Johnson, 1992</bibRefCitation>
: 67
</treatmentCitation>
</taxonomicName>
, figs. 46, 84 for habitus and male genitalia; Huang &amp; Chen 2006: 319, discussion on status;
<treatmentCitation id="0ACD10E6FFBAFFD418527F8FFDB71281" author="Huang" box="[270,534,182,204]" page="141" pageId="21" pageNumber="430" year="2014">
<bibRefCitation id="EFFD4B06FFBAFFD418527F8FFE451281" author="Huang" box="[270,484,182,204]" pageId="21" pageNumber="430" refString="Huang, H. &amp; Zhou, L. - P. (2014) Discovery of two new species of the &quot; elfin &quot; butterflies from Shaanxi province, China. Atalanta, 45, 139 - 150." type="journal article" year="2014">Huang &amp; Zhou 2014</bibRefCitation>
: 141
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C376657CFFBAFFD419CB7FC1FBFC16F7" pageId="21" pageNumber="430" type="materials_examined">
<paragraph id="8BD336F7FFBAFFD419CB7FC1FD80137B" blockId="21.[151,1437,249,1426]" pageId="21" pageNumber="430">
<emphasis id="B918EAE5FFBAFFD419CB7FC1FF5E135F" bold="true" box="[151,255,249,274]" pageId="21" pageNumber="430">Material</emphasis>
.
<collectingCountry id="F37B7667FFBAFFD418527FC1FECA135C" box="[270,363,249,273]" name="China" pageId="21" pageNumber="430">CHINA</collectingCountry>
: Sichuan Province: 1♀ (CHH, dissected), Ganzi Tibetan Autonomous Region, Jiulong County,
<quantity id="4C949B12FFBAFFD419CB7E26FF49137B" box="[151,232,286,310]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="21" pageNumber="430" unit="m" value="2900.0">2900m</quantity>
,
<date id="FFD21037FFBAFFD419AF7E26FED0137B" box="[243,369,286,310]" pageId="21" pageNumber="430" value="2014-04-27">27.IV.2014</date>
, H. Huang leg..
</paragraph>
<paragraph id="8BD336F7FFBAFFD4199B7E79FDE8108F" blockId="21.[151,1437,249,1426]" pageId="21" pageNumber="430">
<emphasis id="B918EAE5FFBAFFD4199B7E79FED11317" bold="true" box="[199,368,321,346]" pageId="21" pageNumber="430">Identification.</emphasis>
A female
<typeStatus id="54D78855FFBAFFD418AE7E7AFDE81317" box="[498,585,322,346]" pageId="21" pageNumber="430" type="syntype">syntype</typeStatus>
from Changyang, Hubei Province was designated by
<bibRefCitation id="EFFD4B06FFBAFFD41D947E79FAD81314" author="Johnson" box="[1224,1401,321,346]" pageId="21" pageNumber="430" refString="Johnson, K. (1992) The Palaearctic &quot; Elfin &quot; Butterflies (Lycaenidae, Theclinae). Neue Entomologische Nachrichten, 29, 1 - 141." type="journal article" year="1992">Johnson (1992)</bibRefCitation>
as
<typeStatus id="54D78855FFBAFFD419CB7E5DFF5F1333" box="[151,254,357,382]" pageId="21" pageNumber="430" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C6C4D74FFBAFFD418797E5EFE121330" box="[293,435,357,381]" class="Insecta" family="Lycaenidae" genus="Ahlbergia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="430" phylum="Arthropoda" rank="species" species="chalybeia">
<emphasis id="B918EAE5FFBAFFD418797E5EFE121330" box="[293,435,357,381]" italics="true" pageId="21" pageNumber="430">A. chalybeia</emphasis>
</taxonomicName>
(female genitalia reproduced herein as
<figureCitation id="13572A72FFBAFFD41A2A7E5DFC791333" box="[886,984,357,382]" captionStart="FIGURES 105 – 129" captionStartId="17.[151,264,1786,1809]" captionTargetBox="[151,1436,193,1764]" captionTargetId="figure@17.[151,1436,193,1765]" captionTargetPageId="17" captionText="FIGURES 105 – 129. Female genitalia of Ahlbergia taxa in ventral view at same scale (except for 109). 105 — A. maoweiweii, paratype (Figs. 31 – 32); 106 — A. ferrea, Liaoning (Fig. 43); 107 — A. luoliangi, paratype (Figs. 39 – 40); 108 — A. clarolinea, holotype (Figs. 63 – 64); 109 — A. chalybeia, lectotype, after Johnson (1992); 110 — A. chalybeia, Jiulong (Figs. 69 – 70); 111 — A. chalcidis, Lijiang; 112 — A. frivaldszkyi, Shaanxi (Fig. 38); 113 — A. tricaudata confusa, holotype (Fig. 45); 114 — A. tricaudata tricaudata, Liaoning (Fig. 44); 115 — A. tricaudata ssp., Shaanxi; 116 – 121, 126 – 129 — A. frivaldszkyi; 116 – 118, 127 – 129 — Shaanxi; 119 — Kangxian, Gansu (Figs. 48 – 49); 120 — Huixian, Gansu (Figs. 52 – 53); 121 — Wenxian, Gansu (Figs. 50 – 51); 126 — Tianjin (Fig. 37); 122 – 125 — A. leei; 122 — Liaoning (Figs. 35 – 36); 123 — Shaanxi (Figs. 41 – 42); 124 – 125 — Shaanxi." httpUri="https://zenodo.org/record/271614/files/figure.png" pageId="21" pageNumber="430">Fig. 109</figureCitation>
). Two female specimens collected from Changyang kept in
<collectionCode id="ED7DAE32FFBAFFD4182E7EB2FE1213EC" box="[370,435,394,417]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="21" pageNumber="430">NHM</collectionCode>
were figured by D’Abrera (1993). All these can be regarded as basis of identification for
<taxonomicName id="4C6C4D74FFBAFFD419CB7E96FE881388" box="[151,297,429,453]" class="Insecta" family="Lycaenidae" genus="Ahlbergia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="430" phylum="Arthropoda" rank="species" species="chalybeia">
<emphasis id="B918EAE5FFBAFFD419CB7E96FE881388" box="[151,297,429,453]" italics="true" pageId="21" pageNumber="430">A. chalybeia</emphasis>
</taxonomicName>
. A peculiar character in female genitalia described by Johnson needs to be clarified: “lamella antevaginalis curved ventrally around caudal tip of ductus bursae with caudal one-tenth of ductus bursae protruding terminally beyond the juncture of the lamella lips”. After dissecting many specimens of elfin butterflies, the first author realized that such peculiar character was caused by the everted condition of membrane between lodix and lamella antevaginalis. The membrane between lodix and lamella antevaginalis covers mostly lamella postvaginalis in natural shape, it can be everted with the whole lamella postvaginalis exposed when the lodix is drawn forwards or the lamella is drawn backwards. Johnson’s (1992) illustration for female genitalia of
<taxonomicName id="4C6C4D74FFBAFFD41DCD7DBEFA8110D0" box="[1169,1312,645,669]" class="Insecta" family="Lycaenidae" genus="Ahlbergia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="430" phylum="Arthropoda" rank="species" species="chalybeia">
<emphasis id="B918EAE5FFBAFFD41DCD7DBEFA8110D0" box="[1169,1312,645,669]" italics="true" pageId="21" pageNumber="430">A. chalybeia</emphasis>
</taxonomicName>
was made when the membrane was fully everted.
</paragraph>
<paragraph id="8BD336F7FFBAFFD4199B7DF5FA9811F3" blockId="21.[151,1437,249,1426]" pageId="21" pageNumber="430">
This species was also recorded by
<bibRefCitation id="EFFD4B06FFBAFFD41B047DF5FCA910AB" author="Johnson" box="[600,776,717,742]" pageId="21" pageNumber="430" refString="Johnson, K. (1992) The Palaearctic &quot; Elfin &quot; Butterflies (Lycaenidae, Theclinae). Neue Entomologische Nachrichten, 29, 1 - 141." type="journal article" year="1992">Johnson (1992)</bibRefCitation>
from “Tatsienlu” (Kangding), western Sichuan. The only specimen examined in this work is a female specimen (
<figureCitation id="13572A72FFBAFFD41A5C7DC9FC261147" box="[768,903,753,778]" captionStart="FIGURES 61 – 75" captionStartId="11.[151,264,1904,1927]" captionTargetBox="[151,1436,193,1881]" captionTargetId="figure@11.[151,1436,193,1883]" captionTargetPageId="11" captionText="FIGURES 61 – 75. Habitus of Ahlbergia females at same scale. 61 – 62 — A. maoweiweii, paratype. 63 – 66, 71 – 72 — A. clarolinea; 63 – 64 — holotype, Yulongxueshan; 65 – 66 — Ludian, Lijiang; 71 – 72 — Muli, Sichuan. 67 – 68 — A. chalcidis, Ludian, Lijiang. 69 – 70 — A. chalybeia, Jiulong. 73 – 75 — A. caesius, holotype. 61, 63, 65, 67, 69, 71, 73 — upperside; 62, 64, 66, 68, 70, 72, 74 — underside." httpUri="https://zenodo.org/record/271609/files/figure.png" pageId="21" pageNumber="430">Figs. 69–70</figureCitation>
,
<figureCitation id="13572A72FFBAFFD41ACE7DCAFC1D1147" box="[914,956,754,778]" captionStart="FIGURES 105 – 129" captionStartId="17.[151,264,1786,1809]" captionTargetBox="[151,1436,193,1764]" captionTargetId="figure@17.[151,1436,193,1765]" captionTargetPageId="17" captionText="FIGURES 105 – 129. Female genitalia of Ahlbergia taxa in ventral view at same scale (except for 109). 105 — A. maoweiweii, paratype (Figs. 31 – 32); 106 — A. ferrea, Liaoning (Fig. 43); 107 — A. luoliangi, paratype (Figs. 39 – 40); 108 — A. clarolinea, holotype (Figs. 63 – 64); 109 — A. chalybeia, lectotype, after Johnson (1992); 110 — A. chalybeia, Jiulong (Figs. 69 – 70); 111 — A. chalcidis, Lijiang; 112 — A. frivaldszkyi, Shaanxi (Fig. 38); 113 — A. tricaudata confusa, holotype (Fig. 45); 114 — A. tricaudata tricaudata, Liaoning (Fig. 44); 115 — A. tricaudata ssp., Shaanxi; 116 – 121, 126 – 129 — A. frivaldszkyi; 116 – 118, 127 – 129 — Shaanxi; 119 — Kangxian, Gansu (Figs. 48 – 49); 120 — Huixian, Gansu (Figs. 52 – 53); 121 — Wenxian, Gansu (Figs. 50 – 51); 126 — Tianjin (Fig. 37); 122 – 125 — A. leei; 122 — Liaoning (Figs. 35 – 36); 123 — Shaanxi (Figs. 41 – 42); 124 – 125 — Shaanxi." httpUri="https://zenodo.org/record/271614/files/figure.png" pageId="21" pageNumber="430">110</figureCitation>
) collected by the first author from Jiulong, historically regarded as part of “Tatsienlu” area; it shares all the external characters with the female specimens figured by D’Abrera (1993: 437) from Changyang. The female genitalia of this specimen agree with Johnson’s (1992) figure in having a depression at distal margin of lamella postvaginalis, but differ by having a relatively larger lamella postvaginalis bearing stronger convolutions. More specimens from Hubei need to be examined for a better understanding of this species. Thus, identification of the specimen from Sichuan remains uncertain.
</paragraph>
<paragraph id="8BD336F7FFBAFFD4199B7CF1FBFC16F7" blockId="21.[151,1437,249,1426]" pageId="21" pageNumber="430">
<emphasis id="B918EAE5FFBAFFD4199B7CF1FE1011AF" bold="true" box="[199,433,969,994]" pageId="21" pageNumber="430">Sexual dimorphism</emphasis>
.
<taxonomicName id="4C6C4D74FFBAFFD4189C7CF2FDE711AC" box="[448,582,969,993]" class="Insecta" family="Lycaenidae" genus="Ahlbergia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="430" phylum="Arthropoda" rank="species" species="chalcidis">
<emphasis id="B918EAE5FFBAFFD4189C7CF2FDE711AC" box="[448,582,969,993]" italics="true" pageId="21" pageNumber="430">A. chalcidis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D74FFBAFFD41B237CF2FCB311AC" box="[639,786,969,993]" class="Insecta" family="Lycaenidae" genus="Ahlbergia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="430" phylum="Arthropoda" rank="species" species="clarolinea">
<emphasis id="B918EAE5FFBAFFD41B237CF2FCB311AC" box="[639,786,969,993]" italics="true" pageId="21" pageNumber="430">A. clarolinea</emphasis>
</taxonomicName>
are closest to
<taxonomicName id="4C6C4D74FFBAFFD41AE47CF2FBE411AC" box="[952,1093,969,993]" class="Insecta" family="Lycaenidae" genus="Ahlbergia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="430" phylum="Arthropoda" rank="species" species="chalybeia">
<emphasis id="B918EAE5FFBAFFD41AE47CF2FBE411AC" box="[952,1093,969,993]" italics="true" pageId="21" pageNumber="430">A. chalybeia</emphasis>
</taxonomicName>
in morphological distance, so that it is reasonable to deduce that the males of these species are also similar to one another in morphology. Both
<taxonomicName id="4C6C4D74FFBAFFD41CD87CD6FF5D1664" class="Insecta" family="Lycaenidae" genus="Ahlbergia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="430" phylum="Arthropoda" rank="species" species="chalcidis">
<emphasis id="B918EAE5FFBAFFD41CD87CD6FF5D1664" italics="true" pageId="21" pageNumber="430">A. chalcidis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D74FFBAFFD4186B7B2AFE6A1664" box="[311,459,1041,1065]" class="Insecta" family="Lycaenidae" genus="Ahlbergia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="430" phylum="Arthropoda" rank="species" species="clarolinea">
<emphasis id="B918EAE5FFBAFFD4186B7B2AFE6A1664" box="[311,459,1041,1065]" italics="true" pageId="21" pageNumber="430">A. clarolinea</emphasis>
</taxonomicName>
have their male genitalia more similar to those of the bright male
<typeStatus id="54D78855FFBAFFD41D937B2AFA871667" box="[1231,1318,1042,1066]" pageId="21" pageNumber="430" type="syntype">syntype</typeStatus>
(
<bibRefCitation id="EFFD4B06FFBAFFD41C6B7B29FEFE1603" author="Johnson" pageId="21" pageNumber="430" refString="Johnson, K. (1992) The Palaearctic &quot; Elfin &quot; Butterflies (Lycaenidae, Theclinae). Neue Entomologische Nachrichten, 29, 1 - 141." type="journal article" year="1992">Johnson, 1992: 108, fig. 46</bibRefCitation>
) of
<taxonomicName id="4C6C4D74FFBAFFD418D77B0EFDB61600" box="[395,535,1077,1101]" class="Insecta" family="Lycaenidae" genus="Ahlbergia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="430" phylum="Arthropoda" rank="species" species="chalybeia">
<emphasis id="B918EAE5FFBAFFD418D77B0EFDB61600" box="[395,535,1077,1101]" italics="true" pageId="21" pageNumber="430">A. chalybeia</emphasis>
</taxonomicName>
than to the dark male specimen identified by
<bibRefCitation id="EFFD4B06FFBAFFD41D487B0DFAE01603" author="Johnson" box="[1044,1345,1077,1102]" pageId="21" pageNumber="430" refString="Johnson, K. (1992) The Palaearctic &quot; Elfin &quot; Butterflies (Lycaenidae, Theclinae). Neue Entomologische Nachrichten, 29, 1 - 141." type="journal article" year="1992">Johnson (1992: 103, fig. 4)</bibRefCitation>
as male of
<taxonomicName id="4C6C4D74FFBAFFD419EB7B62FEE4163C" box="[183,325,1113,1137]" class="Insecta" family="Lycaenidae" genus="Ahlbergia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="430" phylum="Arthropoda" rank="species" species="chalybeia">
<emphasis id="B918EAE5FFBAFFD419EB7B62FEE4163C" box="[183,325,1113,1137]" italics="true" pageId="21" pageNumber="430">A. chalybeia</emphasis>
</taxonomicName>
. Therefore the male of
<taxonomicName id="4C6C4D74FFBAFFD41B047B62FD47163C" box="[600,742,1113,1137]" class="Insecta" family="Lycaenidae" genus="Ahlbergia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="430" phylum="Arthropoda" rank="species" species="chalybeia">
<emphasis id="B918EAE5FFBAFFD41B047B62FD47163C" box="[600,742,1113,1137]" italics="true" pageId="21" pageNumber="430">A. chalybeia</emphasis>
</taxonomicName>
should have extensive blue suffusion on wing uppersides as that of
<taxonomicName id="4C6C4D74FFBAFFD419B97B46FEC816D8" box="[229,361,1149,1173]" class="Insecta" family="Lycaenidae" genus="Ahlbergia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="430" phylum="Arthropoda" rank="species" species="chalcidis">
<emphasis id="B918EAE5FFBAFFD419B97B46FEC816D8" box="[229,361,1149,1173]" italics="true" pageId="21" pageNumber="430">A. chalcidis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D74FFBAFFD418C27B46FD8E16D8" box="[414,559,1149,1173]" class="Insecta" family="Lycaenidae" genus="Ahlbergia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="430" phylum="Arthropoda" rank="species" species="clarolinea">
<emphasis id="B918EAE5FFBAFFD418C27B46FD8E16D8" box="[414,559,1149,1173]" italics="true" pageId="21" pageNumber="430">A. clarolinea</emphasis>
</taxonomicName>
, and in that case
<taxonomicName id="4C6C4D74FFBAFFD41BB37B46FC4116D8" box="[751,992,1149,1173]" class="Insecta" family="Lycaenidae" genus="Novosatsuma" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="430" phylum="Arthropoda" rank="species" species="cibdela">
<emphasis id="B918EAE5FFBAFFD41BB37B46FC4116D8" box="[751,992,1149,1173]" italics="true" pageId="21" pageNumber="430">Novosatsuma cibdela</emphasis>
</taxonomicName>
is a synonym of
<taxonomicName id="4C6C4D74FFBAFFD41DC07B46FA8916D8" box="[1180,1320,1149,1173]" class="Insecta" family="Lycaenidae" genus="Ahlbergia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="430" phylum="Arthropoda" rank="species" species="chalybeia">
<emphasis id="B918EAE5FFBAFFD41DC07B46FA8916D8" box="[1180,1320,1149,1173]" italics="true" pageId="21" pageNumber="430">A. chalybeia</emphasis>
</taxonomicName>
. We leave the formal revision to the future when more specimens from Changyang are available.
</paragraph>
</subSubSection>
<subSubSection id="C376657CFFBAFFD4199B7BFDFAB81723" pageId="21" pageNumber="430" type="diagnosis">
<paragraph id="8BD336F7FFBAFFD4199B7BFDFAB81723" blockId="21.[151,1437,249,1426]" pageId="21" pageNumber="430">
<emphasis id="B918EAE5FFBAFFD4199B7BFDFE9B1693" bold="true" box="[199,314,1221,1246]" pageId="21" pageNumber="430">Diagnosis</emphasis>
. It is very possible that
<taxonomicName id="4C6C4D74FFBAFFD41B1F7BFDFC401693" authority="Chou &amp; Li, 1994" authorityName="Chou &amp; Li" authorityYear="1994" box="[579,993,1221,1246]" class="Insecta" family="Lycaenidae" genus="Ahlbergia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="430" phylum="Arthropoda" rank="species" species="chalcidis">
<emphasis id="B918EAE5FFBAFFD41B1F7BFDFCBC1690" box="[579,797,1221,1245]" italics="true" pageId="21" pageNumber="430">Ahlbergia chalcidis</emphasis>
<bibRefCitation id="EFFD4B06FFBAFFD41A787BFDFC401693" author="Chou" box="[804,993,1221,1246]" pageId="21" pageNumber="430" refString="Chou, I. &amp; al. (1994) Monographia Rhopalocerorum Sinensium. Henan Science and Technology." type="book" year="1994">Chou &amp; Li, 1994</bibRefCitation>
</taxonomicName>
from Yunnan represents a geographical variation of
<taxonomicName id="4C6C4D74FFBAFFD418437BD2FE0A174C" box="[287,427,1257,1281]" class="Insecta" family="Lycaenidae" genus="Ahlbergia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="430" phylum="Arthropoda" rank="species" species="chalybeia">
<emphasis id="B918EAE5FFBAFFD418437BD2FE0A174C" box="[287,427,1257,1281]" italics="true" pageId="21" pageNumber="430">A. chalybeia</emphasis>
</taxonomicName>
.
<taxonomicName id="4C6C4D74FFBAFFD418E57BD2FDE5174C" box="[441,580,1257,1281]" class="Insecta" family="Lycaenidae" genus="Ahlbergia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="430" phylum="Arthropoda" rank="species" species="chalybeia">
<emphasis id="B918EAE5FFBAFFD418E57BD2FDE5174C" box="[441,580,1257,1281]" italics="true" pageId="21" pageNumber="430">A. chalybeia</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C6C4D74FFBAFFD41A2F7BD1FAF1174F" authority="Huang &amp; Chen, 2006" authorityName="Huang &amp; Chen" authorityYear="2006" box="[883,1360,1257,1282]" class="Insecta" family="Lycaenidae" genus="Ahlbergia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="430" phylum="Arthropoda" rank="species" species="clarolinea">
<emphasis id="B918EAE5FFBAFFD41A2F7BD1FBFA174C" box="[883,1115,1257,1281]" italics="true" pageId="21" pageNumber="430">Ahlbergia clarolinea</emphasis>
Huang &amp; Chen, 2006
</taxonomicName>
by the following combination of characters: 1) hindwing underside ground color more reddish and less greyish; 2) hindwing underside with more clearly defined postdiscal markings; 3) hindwing underside with more ill-defined black dashes or lines in subbasal and discal areas; 4) ductus bursae of female genitalia markedly wider.
</paragraph>
</subSubSection>
<subSubSection id="C376657CFFBAFFD4199B7A41FC1117DF" box="[199,944,1401,1426]" pageId="21" pageNumber="430" type="distribution">
<paragraph id="8BD336F7FFBAFFD4199B7A41FC1117DF" blockId="21.[151,1437,249,1426]" box="[199,944,1401,1426]" pageId="21" pageNumber="430">
<emphasis id="B918EAE5FFBAFFD4199B7A41FEC117DF" bold="true" box="[199,352,1401,1426]" pageId="21" pageNumber="430">Distribution.</emphasis>
Hubei (Changyang),? Sichuan (Kangding, Jiulong).
</paragraph>
</subSubSection>
</treatment>
</document>
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