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<document id="1EF44ADC3DBC37FDACA5865E69623357" ID-CLB-Dataset="29039" ID-DOI="10.11646/zootaxa.4459.1.1" ID-GBIF-Dataset="9acf71bc-6414-4878-8af0-a64d3ec1fea6" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458302" ID-ZooBank="633A9498-FCC6-4439-BE33-A45B9E51B550" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539287574730" checkinUser="plazi" docAuthor="Grandinete, Yuri Campanholo, Noll, Fernando Barbosa &amp; Carpenter, James" docDate="2018" docId="03C5878FFFD0F324FF309089D2C2F8DA" docLanguage="en" docName="zootaxa.4459.1.1.pdf" docOrigin="Zootaxa 4459 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eumenes rufomaculatus Fox 1899" docType="treatment" docVersion="9" lastPageNumber="37" masterDocId="FFFCFFF7FFF3F300FFA79155D339FFDF" masterDocTitle="Taxonomic Review of Eumenes Latreille, 1802 (Hymenoptera, Vespidae, Eumeninae) from the New World" masterLastPageNumber="52" masterPageNumber="1" pageNumber="36" updateTime="1698577660185" updateUser="ExternalLinkService">
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<mods:title id="897D0420AB3F55B8908958D0727A5394">Taxonomic Review of Eumenes Latreille, 1802 (Hymenoptera, Vespidae, Eumeninae) from the New World</mods:title>
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<mods:namePart id="510DB887EE7554EDF3E8C3615B20AF4A">Grandinete, Yuri Campanholo</mods:namePart>
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<mods:namePart id="86963368CE6FF4841C35C3A67B351F17">Carpenter, James</mods:namePart>
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<mods:title id="BB6DC7D3C27246FD6AD795B52DEAAF89">Zootaxa</mods:title>
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<mods:date id="51FE321E3919328F4915C7D9A852C6DB">2018</mods:date>
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<mods:number id="29C3E3CD379DC4FEE50815F9B75EB4BF">2018-08-14</mods:number>
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<mods:identifier id="DFE19F8CAEBF3C43F441DEAA1BFBC71A" type="DOI">10.11646/zootaxa.4459.1.1</mods:identifier>
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<treatment id="03C5878FFFD0F324FF309089D2C2F8DA" ID-DOI="http://doi.org/10.5281/zenodo.5986399" ID-GBIF-Taxon="148692493" ID-Zenodo-Dep="5986399" LSID="urn:lsid:plazi:treatment:03C5878FFFD0F324FF309089D2C2F8DA" httpUri="http://treatment.plazi.org/id/03C5878FFFD0F324FF309089D2C2F8DA" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
<subSubSection id="C3766512FFD0F323FF309089D19FFDE7" pageId="35" pageNumber="36" type="nomenclature">
<paragraph id="8BD33699FFD0F323FF309089D179FE29" blockId="35.[151,576,476,502]" box="[151,576,476,502]" pageId="35" pageNumber="36">
<heading id="D09B81F5FFD0F323FF309089D179FE29" bold="true" box="[151,576,476,502]" fontSize="11" level="1" pageId="35" pageNumber="36" reason="1">
<taxonomicName id="4C6C4D1AFFD0F323FF309089D179FE29" ID-CoL="3CDNN" authority="Fox, 1899" authorityName="Fox" authorityYear="1899" box="[151,576,476,502]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="rufomaculatus">
<emphasis id="B918EA8BFFD0F323FF309089D179FE29" bold="true" box="[151,576,476,502]" pageId="35" pageNumber="36">
<emphasis id="B918EA8BFFD0F323FF309089D286FE29" bold="true" box="[151,447,476,502]" italics="true" pageId="35" pageNumber="36">Eumenes rufomaculatus</emphasis>
Fox, 1899
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BD33699FFD0F323FF309376D19FFDE7" blockId="35.[151,1436,546,1366]" box="[151,678,546,568]" pageId="35" pageNumber="36">
<taxonomicName id="4C6C4D1AFFD0F323FF309376D149FDE7" authority="Fox, 1899" authorityName="Fox" authorityYear="1899" box="[151,624,546,568]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="rufomaculata">
<emphasis id="B918EA8BFFD0F323FF309376D2C3FDE7" bold="true" box="[151,506,546,568]" pageId="35" pageNumber="36">
<emphasis id="B918EA8BFFD0F323FF309376D2B5FDE7" bold="true" box="[151,396,546,568]" italics="true" pageId="35" pageNumber="36">Eumenes rufomaculata</emphasis>
Fox, 1899
</emphasis>
, 1899: 457
</taxonomicName>
, 461.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFD0F323FF609314D1BFFCF1" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="8BD33699FFD0F323FF609314D0A2FD88" blockId="35.[151,1436,546,1366]" box="[199,923,577,599]" pageId="35" pageNumber="36">
<emphasis id="B918EA8BFFD0F323FF609314D203FD88" bold="true" box="[199,314,577,599]" pageId="35" pageNumber="36">Type data:</emphasis>
Lectotype and paralectotype female at CMNH
<emphasis id="B918EA8BFFD0F323FCBB9314D0ACFD88" bold="true" box="[796,917,577,599]" pageId="35" pageNumber="36">(examined)</emphasis>
.
</paragraph>
<paragraph id="8BD33699FFD0F323FF609335D129FDA9" blockId="35.[151,1436,546,1366]" box="[199,528,608,630]" pageId="35" pageNumber="36">
<materialsCitation id="3B043CC4FFD0F323FF609335D129FDA9" ID-GBIF-Occurrence="1917355176" box="[199,528,608,630]" country="Brazil" location="Corumba" pageId="35" pageNumber="36" specimenCount="1" typeStatus="holotype">
<emphasis id="B918EA8BFFD0F323FF609335D263FDA9" bold="true" box="[199,346,608,630]" pageId="35" pageNumber="36">
<typeStatus id="54D7883BFFD0F323FF609335D3C5FDA9" box="[199,252,608,630]" pageId="35" pageNumber="36">Type</typeStatus>
locality:
</emphasis>
<location id="8EB36042FFD0F323FEC79335D2F8FDA9" LSID="urn:lsid:plazi:treatment:03C5878FFFD0F324FF309089D2C2F8DA:8EB36042FFD0F323FEC79335D2F8FDA9" box="[352,449,608,630]" country="Brazil" name="Corumba" pageId="35" pageNumber="36">Corumbá</location>
,
<collectingCountry id="F37B7609FFD0F323FE6C9335D135FDA9" box="[459,524,608,630]" name="Brazil" pageId="35" pageNumber="36">Brazil</collectingCountry>
.
</materialsCitation>
</paragraph>
<paragraph id="8BD33699FFD0F323FF60932BD2F4FD4B" blockId="35.[151,1436,546,1366]" box="[199,461,638,660]" pageId="35" pageNumber="36">Dalla Torre, 1904, 19: 24.</paragraph>
<paragraph id="8BD33699FFD0F323FF6093C8D11FFD6C" blockId="35.[151,1436,546,1366]" box="[199,550,669,691]" pageId="35" pageNumber="36">Brèthes, 1906, 6 (13): 336, fig. 18.</paragraph>
<paragraph id="8BD33699FFD0F323FF6093E9D2F3FD0D" blockId="35.[151,1436,546,1366]" box="[199,458,700,722]" pageId="35" pageNumber="36">Bertoni, 1918, (2) 3: 206.</paragraph>
<paragraph id="8BD33699FFD0F323FF30938ED1FDFD2F" blockId="35.[151,1436,546,1366]" box="[151,708,730,752]" pageId="35" pageNumber="36">
<taxonomicName id="4C6C4D1AFFD0F323FF30938ED2B2FD2F" authorityName="Fox" authorityYear="1899" box="[151,395,730,752]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="rufomaculatus">
<emphasis id="B918EA8BFFD0F323FF30938ED2B2FD2F" box="[151,395,730,752]" italics="true" pageId="35" pageNumber="36">Eumenes rufomaculatus</emphasis>
</taxonomicName>
; Giordani Soika, 1961, 1: 241.
</paragraph>
<paragraph id="8BD33699FFD0F323FF6093ACD139FCD0" blockId="35.[151,1436,546,1366]" box="[199,512,761,783]" pageId="35" pageNumber="36">Rodríguez-Palafox, 1996: 479.</paragraph>
<paragraph id="8BD33699FFD0F323FF60924CD1BFFCF1" blockId="35.[151,1436,546,1366]" box="[199,646,792,814]" pageId="35" pageNumber="36">
Hermes
<emphasis id="B918EA8BFFD0F323FEBB924CD271FCF1" box="[284,328,792,814]" italics="true" pageId="35" pageNumber="36">et al</emphasis>
., 2014, 30: 455, 459, 462, 469.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFD0F323FF30926DD188FCB4" pageId="35" pageNumber="36" type="reference_group">
<paragraph id="8BD33699FFD0F323FF30926DD188FCB4" blockId="35.[151,1436,546,1366]" pageId="35" pageNumber="36">
<taxonomicName id="4C6C4D1AFFD0F323FF30926DD100FC93" ID-CoL="3CDNN" authorityName="Fox" authorityYear="1899" box="[151,569,822,845]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="rufomaculatus" subGenus="Zeteumenoides">
<emphasis id="B918EA8BFFD0F323FF30926DD3CAFC92" box="[151,243,824,845]" italics="true" pageId="35" pageNumber="36">Eumenes</emphasis>
(
<emphasis id="B918EA8BFFD0F323FEA69263D2A0FC93" box="[257,409,822,844]" italics="true" pageId="35" pageNumber="36">Zeteumenoides</emphasis>
)
<emphasis id="B918EA8BFFD0F323FE009263D100FC93" box="[423,569,822,844]" italics="true" pageId="35" pageNumber="36">rufomaculatus</emphasis>
</taxonomicName>
; Carpenter and van der
<bibRefCitation id="EFFD4B68FFD0F323FC899263D090FC93" box="[814,937,822,844]" pageId="35" pageNumber="36" refId="ref42766" refString="Carpenter, J. M. &amp; van der Vecht, J. (1991) A study of the Vespidae described by William J. Fox (Insecta: Hymenoptera), with assessment of taxonomic implications. Annals of the Carnegie Museum of Natural History, 60 (3), 211 - 241." type="journal article">Vecht, 1991</bibRefCitation>
, 60: 211, 232 (designation of
<typeStatus id="54D7883BFFD0F323FB709263D600FC93" box="[1239,1337,822,844]" pageId="35" pageNumber="36" type="lectotype">lectotype</typeStatus>
).
<taxonomicName id="4C6C4D1AFFD0F323FF309203D124FCB4" authority="Zavattari, 1912" authorityName="Zavattari" authorityYear="1912" box="[151,541,853,875]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="colombianus">
<emphasis id="B918EA8BFFD0F323FF309203D240FCB4" box="[151,377,853,875]" italics="true" pageId="35" pageNumber="36">Eumenes colombianus</emphasis>
<bibRefCitation id="EFFD4B68FFD0F323FE279200D124FCB4" box="[384,541,853,875]" pageId="35" pageNumber="36" refId="ref44216" refString="Zavattari, E. (1912) Materialien fur eine Monographie der neotropischen Eumeniden. Archiv fur Naturgeschichte, 78 A (4), 1 - 272." type="journal article">Zavattari, 1912</bibRefCitation>
</taxonomicName>
, 78A (4): 130.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFD0F323FF609221D142FC39" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="8BD33699FFD0F323FF609221D123FC55" blockId="35.[151,1436,546,1366]" box="[199,538,884,906]" pageId="35" pageNumber="36">
<emphasis id="B918EA8BFFD0F323FF609221D203FC55" bold="true" box="[199,314,884,906]" pageId="35" pageNumber="36">Type data:</emphasis>
Type male at ZMHB.
</paragraph>
<paragraph id="8BD33699FFD0F323FF6092C7D2F2FC77" blockId="35.[151,1436,546,1366]" box="[199,459,914,936]" pageId="35" pageNumber="36">
<materialsCitation id="3B043CC4FFD0F323FF6092C7D2F2FC77" ID-GBIF-Occurrence="1917355140" box="[199,459,914,936]" country="Colombia" location="Colombia" pageId="35" pageNumber="36" specimenCount="1" typeStatus="holotype">
<emphasis id="B918EA8BFFD0F323FF6092C7D263FC77" bold="true" box="[199,346,914,936]" pageId="35" pageNumber="36">
<typeStatus id="54D7883BFFD0F323FF6092C7D3C5FC77" box="[199,252,914,936]" pageId="35" pageNumber="36">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="F37B7609FFD0F323FEC792C7D2FEFC77" box="[352,455,914,936]" name="Colombia" pageId="35" pageNumber="36">Colombia</collectingCountry>
.
</materialsCitation>
</paragraph>
<paragraph id="8BD33699FFD0F323FF6092E4D2CDFC18" blockId="35.[151,1436,546,1366]" box="[199,500,945,967]" pageId="35" pageNumber="36">Giordani Soika, 1961, 1: 241.</paragraph>
<paragraph id="8BD33699FFD0F323FF309284D142FC39" blockId="35.[151,1436,546,1366]" box="[151,635,976,998]" pageId="35" pageNumber="36">
<taxonomicName id="4C6C4D1AFFD0F323FF309284D2A5FC39" box="[151,412,976,998]" class="Insecta" family="Vespidae" genus="Zeteumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="rufomaculata">
<emphasis id="B918EA8BFFD0F323FF309284D2A5FC39" box="[151,412,976,998]" italics="true" pageId="35" pageNumber="36">Zeteumenes rufomaculata</emphasis>
</taxonomicName>
; Bertoni, 1921, 1: 11.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFD0F323FF3092A5D029FBDB" box="[151,784,1006,1029]" pageId="35" pageNumber="36" type="description">
<paragraph id="8BD33699FFD0F323FF3092A5D029FBDB" blockId="35.[151,1436,546,1366]" box="[151,784,1006,1029]" pageId="35" pageNumber="36">
<taxonomicName id="4C6C4D1AFFD0F323FF3092A5D23AFBDA" authorityName="Bertoni" authorityYear="1925" box="[151,259,1008,1029]" class="Insecta" family="Vespidae" genus="Zetamenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA8BFFD0F323FF3092A5D23AFBDA" box="[151,259,1008,1029]" italics="true" pageId="35" pageNumber="36">Zetamenes</emphasis>
</taxonomicName>
[!]
<taxonomicName id="4C6C4D1AFFD0F323FE8E92BBD1F5FBDB" authority="Bertoni, 1925" authorityName="Bertoni" authorityYear="1925" box="[297,716,1006,1028]" class="Insecta" family="Vespidae" genus="Zeteumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="subSpecies" species="rufomaculata" subSpecies="meridionalis">
<emphasis id="B918EA8BFFD0F323FE8E92BBD101FBDB" box="[297,568,1006,1028]" italics="true" pageId="35" pageNumber="36">rufomaculata meridionalis</emphasis>
Bertoni, 1925
</taxonomicName>
, 2: 75.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFD0F323FF609558D070FBA0" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="8BD33699FFD0F323FF609558D1B6FBFC" blockId="35.[151,1436,546,1366]" box="[199,655,1037,1059]" pageId="35" pageNumber="36">
<emphasis id="B918EA8BFFD0F323FF609558D203FBFC" bold="true" box="[199,314,1037,1059]" pageId="35" pageNumber="36">Type data:</emphasis>
Type female and male at IBNPY.
</paragraph>
<paragraph id="8BD33699FFD0F323FF609579D149FB9D" blockId="35.[151,1436,546,1366]" box="[199,624,1068,1090]" pageId="35" pageNumber="36">
<materialsCitation id="3B043CC4FFD0F323FF609579D149FB9D" ID-GBIF-Occurrence="1917355138" box="[199,624,1068,1090]" country="Paraguay" location="Puerto Bertoni" pageId="35" pageNumber="36" specimenCount="1" typeStatus="holotype">
<emphasis id="B918EA8BFFD0F323FF609579D263FB9D" bold="true" box="[199,346,1068,1090]" pageId="35" pageNumber="36">
<typeStatus id="54D7883BFFD0F323FF609579D3C5FB9D" box="[199,252,1068,1090]" pageId="35" pageNumber="36">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="F37B7609FFD0F323FEC79578D287FB9D" box="[352,446,1069,1090]" name="Paraguay" pageId="35" pageNumber="36">Paraguay</collectingCountry>
(
<location id="8EB36042FFD0F323FE6C9579D15FFB9D" LSID="urn:lsid:plazi:treatment:03C5878FFFD0F324FF309089D2C2F8DA:8EB36042FFD0F323FE6C9579D15FFB9D" box="[459,614,1068,1090]" country="Paraguay" name="Puerto Bertoni" pageId="35" pageNumber="36">Puerto Bertoni</location>
).
</materialsCitation>
</paragraph>
<paragraph id="8BD33699FFD0F323FF60951FD2CDFBBF" blockId="35.[151,1436,546,1366]" box="[199,500,1098,1120]" pageId="35" pageNumber="36">
<bibRefCitation id="EFFD4B68FFD0F323FF60951FD29BFBBF" box="[199,418,1098,1120]" pageId="35" pageNumber="36" refId="ref43151" refString="Giordani Soika, A. (1978) Revisione degli Eumenidi neotropicali appartenenti ai generi Eumenes Latr., Omicron (Sauss.), Pararhaphidoglossa Schulth. ed afinni. Bolletino della Museo Civico di Storia Naturale di Uenezia, 29, 1 - 420." type="journal article">Giordani Soika, 1978</bibRefCitation>
, 29: 41.
</paragraph>
<paragraph id="8BD33699FFD0F323FF30953FD070FBA0" blockId="35.[151,1436,546,1366]" box="[151,841,1129,1151]" pageId="35" pageNumber="36">
<taxonomicName id="4C6C4D1AFFD0F323FF30953FD115FBA0" box="[151,556,1129,1151]" class="Insecta" family="Vespidae" genus="Zeteumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="rufomaculata" subGenus="Beteumenes">
<emphasis id="B918EA8BFFD0F323FF30953FD236FBA0" box="[151,271,1130,1151]" italics="true" pageId="35" pageNumber="36">Zeteumenes</emphasis>
(
<emphasis id="B918EA8BFFD0F323FEBA953FD2AFFBA0" box="[285,406,1130,1151]" italics="true" pageId="35" pageNumber="36">Beteumenes</emphasis>
)
<emphasis id="B918EA8BFFD0F323FE03953CD115FBA0" box="[420,556,1129,1151]" italics="true" pageId="35" pageNumber="36">rufomaculata</emphasis>
</taxonomicName>
; Bertoni, 1934, 3: 111, 115.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFD0F323FF3095DCD0AAFB04" pageId="35" pageNumber="36" type="description">
<paragraph id="8BD33699FFD0F323FF3095DCD7A7FB41" blockId="35.[151,1436,546,1366]" box="[151,1182,1160,1182]" pageId="35" pageNumber="36">
<taxonomicName id="4C6C4D1AFFD0F323FF3095DCD18AFB41" authorityName="Bertoni" authorityYear="1925" box="[151,691,1160,1182]" class="Insecta" family="Vespidae" genus="Zeteumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="subSpecies" species="rufomaculata" subGenus="Beteumenes" subSpecies="meridionalis">
<emphasis id="B918EA8BFFD0F323FF3095DCD236FB41" box="[151,271,1161,1182]" italics="true" pageId="35" pageNumber="36">Zeteumenes</emphasis>
(
<emphasis id="B918EA8BFFD0F323FEBA95DCD2AFFB41" box="[285,406,1161,1182]" italics="true" pageId="35" pageNumber="36">Beteumenes</emphasis>
)
<emphasis id="B918EA8BFFD0F323FE0395DDD18AFB41" box="[420,691,1160,1182]" italics="true" pageId="35" pageNumber="36">rufomaculata meridionalis</emphasis>
</taxonomicName>
; Bertoni, 1934, 3: 111, (
<emphasis id="B918EA8BFFD0F323FC0B95DDD77DFB41" box="[940,1092,1160,1182]" italics="true" pageId="35" pageNumber="36">ruformaculada</emphasis>
[!]) 115.
</paragraph>
<paragraph id="8BD33699FFD0F323FF3095F3D023FB63" blockId="35.[151,1436,546,1366]" box="[151,794,1190,1212]" pageId="35" pageNumber="36">
<taxonomicName id="4C6C4D1AFFD0F323FF3095F3D2FEFB63" box="[151,455,1190,1212]" class="Insecta" genus="Zeteumenoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="rufomaculatus">
<emphasis id="B918EA8BFFD0F323FF3095F3D2FEFB63" box="[151,455,1190,1212]" italics="true" pageId="35" pageNumber="36">Zeteumenoides rufomaculatus</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFFD4B68FFD0F323FE7395F3D189FB63" box="[468,688,1190,1212]" pageId="35" pageNumber="36" refId="ref43124" refString="Giordani Soika, A. (1972) Notulae vespidologicae XXXII Nuovi Eumenidi Indomalesi. Bolletino dela Societa Entomologica Italiana, 104, 99 - 110." type="journal article">Giordani Soika, 1972</bibRefCitation>
, 104: 110.
</paragraph>
<paragraph id="8BD33699FFD0F323FF309593D0AAFB04" blockId="35.[151,1436,546,1366]" box="[151,915,1221,1243]" pageId="35" pageNumber="36">
<taxonomicName id="4C6C4D1AFFD0F323FF309593D2CEFB04" box="[151,503,1221,1243]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="subSpecies" species="versicolor" subSpecies="rufomaculatus">
<emphasis id="B918EA8BFFD0F323FF309593D2CEFB04" box="[151,503,1221,1243]" italics="true" pageId="35" pageNumber="36">Eumenes versicolor rufomaculatus</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFFD4B68FFD0F323FDA29590D1D9FB04" box="[517,736,1221,1243]" pageId="35" pageNumber="36" refId="ref43151" refString="Giordani Soika, A. (1978) Revisione degli Eumenidi neotropicali appartenenti ai generi Eumenes Latr., Omicron (Sauss.), Pararhaphidoglossa Schulth. ed afinni. Bolletino della Museo Civico di Storia Naturale di Uenezia, 29, 1 - 420." type="journal article">Giordani Soika, 1978</bibRefCitation>
, 29: 17 (key), 41.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFD0F323FF6095B1D2CAFA89" pageId="35" pageNumber="36" type="distribution">
<paragraph id="8BD33699FFD0F323FF6095B1D148FAE8" blockId="35.[151,1436,546,1366]" pageId="35" pageNumber="36">
<emphasis id="B918EA8BFFD0F323FF6095B1D26AFB25" bold="true" box="[199,339,1252,1274]" pageId="35" pageNumber="36">Distribution:</emphasis>
<collectingCountry id="F37B7609FFD0F323FEFC95B1D2F9FB25" box="[347,448,1252,1274]" name="Argentina" pageId="35" pageNumber="36">Argentina</collectingCountry>
(
<collectingRegion id="49A8F87BFFD0F323FE6995B1D102FB25" box="[462,571,1252,1274]" country="Argentina" name="Corrientes" pageId="35" pageNumber="36">Corrientes</collectingRegion>
,
<collectingRegion id="49A8F87BFFD0F323FDEF95B0D18EFB25" box="[584,695,1252,1274]" country="Argentina" name="Entre Rios" pageId="35" pageNumber="36">Entre Rios</collectingRegion>
,
<collectingRegion id="49A8F87BFFD0F323FD6495B0D027FB25" box="[707,798,1253,1274]" country="Argentina" name="Formosa" pageId="35" pageNumber="36">
<collectingCountry id="F37B7609FFD0F323FD6495B0D027FB25" box="[707,798,1253,1274]" name="Taiwan" pageId="35" pageNumber="36">Formosa</collectingCountry>
</collectingRegion>
,
<collectingRegion id="49A8F87BFFD0F323FC8E95B1D0B3FB25" box="[809,906,1252,1274]" country="Paraguay" name="Misiones" pageId="35" pageNumber="36">Misiones</collectingRegion>
);
<collectingCountry id="F37B7609FFD0F323FC3995B1D0E5FB25" box="[926,988,1252,1274]" name="Brazil" pageId="35" pageNumber="36">Brazil</collectingCountry>
(DF, MS*; RJ*; SC*, SP*);
<collectingCountry id="F37B7609FFD0F323FAAF95B1D66AFB25" box="[1288,1363,1252,1274]" name="Bolivia" pageId="35" pageNumber="36">Bolivia</collectingCountry>
(
<collectingRegion id="49A8F87BFFD0F323FAC595B1D3C5FAC7" country="Bolivia" name="Santa Cruz" pageId="35" pageNumber="36">Santa Cruz</collectingRegion>
);
<collectingCountry id="F37B7609FFD0F323FEB79457D24EFAC7" box="[272,375,1282,1304]" name="Colombia" pageId="35" pageNumber="36">Colombia</collectingCountry>
* (
<collectingRegion id="49A8F87BFFD0F323FE379457D2E4FAC7" box="[400,477,1282,1304]" country="Colombia" name="Arauca" pageId="35" pageNumber="36">Arauca</collectingRegion>
,
<collectingRegion id="49A8F87BFFD0F323FE4E9457D10EFAC7" box="[489,567,1282,1304]" country="Colombia" name="Bogota" pageId="35" pageNumber="36">Bogotá</collectingRegion>
);
<collectingCountry id="F37B7609FFD0F323FDED9457D183FAC7" box="[586,698,1282,1304]" name="Costa Rica" pageId="35" pageNumber="36">Costa Rica</collectingCountry>
(
<collectingRegion id="49A8F87BFFD0F323FD6E9457D010FAC7" box="[713,809,1282,1304]" country="Costa Rica" name="San Jose" pageId="35" pageNumber="36">San José</collectingRegion>
);
<collectingRegion id="49A8F87BFFD0F323FC9A9457D0B4FAC7" box="[829,909,1282,1304]" country="Mexico" name="Mexico" pageId="35" pageNumber="36">
<collectingCountry id="F37B7609FFD0F323FC9A9457D0B4FAC7" box="[829,909,1282,1304]" name="Mexico" pageId="35" pageNumber="36">Mexico</collectingCountry>
</collectingRegion>
* (
<collectingRegion id="49A8F87BFFD0F323FC019457D73CFAC7" box="[934,1029,1282,1304]" country="Mexico" name="Guerrero" pageId="35" pageNumber="36">Guerrero</collectingRegion>
,
<collectingRegion id="49A8F87BFFD0F323FBB69456D75EFAC7" box="[1041,1127,1283,1304]" country="Mexico" name="Yucatan" pageId="35" pageNumber="36">Yucatan</collectingRegion>
);
<collectingRegion id="49A8F87BFFD0F323FBDC9456D7E0FAC7" box="[1147,1241,1283,1304]" country="Paraguay" name="Asuncion" pageId="35" pageNumber="36">
<collectingCountry id="F37B7609FFD0F323FBDC9456D7E0FAC7" box="[1147,1241,1283,1304]" name="Paraguay" pageId="35" pageNumber="36">Paraguay</collectingCountry>
</collectingRegion>
(Distrito Capital*,
<collectingRegion id="49A8F87BFFD0F323FF609474D20BFAE8" box="[199,306,1313,1335]" country="Paraguay" name="San Pedro" pageId="35" pageNumber="36">
<collectingCountry id="F37B7609FFD0F323FF609474D20BFAE8" box="[199,306,1313,1335]" name="Saint Pierre and Miquelon" pageId="35" pageNumber="36">San Pedro</collectingCountry>
</collectingRegion>
);
<collectingCountry id="F37B7609FFD0F323FEE39474D2A4FAE8" box="[324,413,1313,1335]" name="Trinidad and Tobago" pageId="35" pageNumber="36">Trinidad</collectingCountry>
* (Tanapuna-Piarco).
</paragraph>
<paragraph id="8BD33699FFD0F323FF609415D2CAFA89" blockId="35.[151,1436,546,1366]" box="[199,499,1344,1366]" pageId="35" pageNumber="36">
<emphasis id="B918EA8BFFD0F323FF609415D3EDFA89" bold="true" box="[199,212,1344,1366]" pageId="35" pageNumber="36">*</emphasis>
Records from the literature.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFD0F323FF3094D7D776F990" pageId="35" pageNumber="36" type="discussion">
<paragraph id="8BD33699FFD0F323FF3094D7D776F990" blockId="35.[151,1437,1410,2011]" pageId="35" pageNumber="36">
<emphasis id="B918EA8BFFD0F323FF3094D7D2DBFA44" bold="true" box="[151,482,1410,1435]" pageId="35" pageNumber="36">Comments and diagnosis.</emphasis>
<taxonomicName id="4C6C4D1AFFD0F323FE5294D6D029FA44" authorityName="Fox" authorityYear="1899" box="[501,784,1411,1435]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="rufomaculatus">
<emphasis id="B918EA8BFFD0F323FE5294D6D029FA44" box="[501,784,1411,1435]" italics="true" pageId="35" pageNumber="36">Eumenes rufomaculatus</emphasis>
</taxonomicName>
can be identified by the following features: body predominantly ferrugineous; clypeus slightly wider and shorter than
<taxonomicName id="4C6C4D1AFFD0F323FC3E94F2D726FA61" authorityName="de Saussure" authorityYear="1855" box="[921,1055,1446,1470]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="B918EA8BFFD0F323FC3E94F2D726FA61" box="[921,1055,1446,1470]" italics="true" pageId="35" pageNumber="36">E. filiformis</emphasis>
</taxonomicName>
with weak but evident punctation homogeneously distributed (fig. 75); humeral region more projecting than
<taxonomicName id="4C6C4D1AFFD0F323FC43949ED753FA3C" authorityName="de Saussure" authorityYear="1855" box="[996,1130,1483,1507]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="B918EA8BFFD0F323FC43949ED753FA3C" box="[996,1130,1483,1507]" italics="true" pageId="35" pageNumber="36">E. filiformis</emphasis>
</taxonomicName>
. T1 very long and narrow; apical region from the ventral surface of T1 ventrally projecting. Digitus broad on basal half, becoming narrow towards apex (fig. 124), with the dorsal edge weakly angled (not strongly as in
<taxonomicName id="4C6C4D1AFFD0F323FB8B9746D78AF9F4" authorityName="de Saussure" authorityYear="1855" box="[1068,1203,1555,1579]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="B918EA8BFFD0F323FB8B9746D78AF9F4" box="[1068,1203,1555,1579]" italics="true" pageId="35" pageNumber="36">E. filiformis</emphasis>
</taxonomicName>
). Digitus with short bristles along the dorsal edge, while on the broad region there are no evident bristles.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFD0F324FF60970FD094F98F" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="description">
<paragraph id="8BD33699FFD0F323FF60970FD04AF9AC" blockId="35.[151,1437,1410,2011]" box="[199,883,1626,1651]" pageId="35" pageNumber="36">
<emphasis id="B918EA8BFFD0F323FF60970FD033F9AC" bold="true" box="[199,778,1626,1651]" pageId="35" pageNumber="36">
Redescription
<taxonomicName id="4C6C4D1AFFD0F323FED3970ED033F9AC" authority="Fox, 1899" authorityName="Fox" authorityYear="1899" box="[372,778,1626,1651]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="rufomaculatus">
<emphasis id="B918EA8BFFD0F323FED3970ED1A9F9AC" bold="true" box="[372,656,1626,1651]" italics="true" pageId="35" pageNumber="36">Eumenes rufomaculatus</emphasis>
Fox, 1899
</taxonomicName>
</emphasis>
(
<figureCitation id="13572A1CFFD0F323FCBD970FD052F9AC" box="[794,875,1626,1651]" captionStart="FIGURES 13–26" captionStartId="42.[151,264,1649,1671]" captionTargetBox="[152,1436,193,1628]" captionTargetId="figure@42.[151,1436,193,1628]" captionTargetPageId="42" captionText="FIGURES 13–26. Eumenes species, lateral view. 13) E. cubensis Cresson, 1865 (female); 14) E. flavitinctus Bohart, 1950 (Holotype, female); 15) E. fervens Say, 1824 (Lectotype, female); 16) E. pictus (Smith, 1857) (female); 17) E. sculleni Bohart, 1950 (Holotype, male); 18) E. smithii de Saussure, 1852 (female); 19) E. belfragei Cresson, 1872 (Lectotype, female); 20) E. verticalis Say, 1824 (female); 21) Eumenes verticalis coloradensis Cresson, 1875 (lectotype, female); 22) E. verticalis neoboreus Bequaert, 1944 (Holotype, male); 23) E. verticalis tricinctus Isely, 1917 (Holotype, female); 24) E. filiformis de Saussure, 1855 (famele); 25) E. rufomaculata Fox, 1899 (Lectotype, female); 26) Eumenes versicolor de Saussure, 1852 (Lectotype, female). Scale bar = 1mm." httpUri="https://zenodo.org/record/1458306/files/figure.png" pageId="35" pageNumber="36">Fig. 25</figureCitation>
)
</paragraph>
<paragraph id="8BD33699FFD0F324FF60972BD2E0FF27" blockId="35.[151,1437,1410,2011]" lastBlockId="36.[151,1437,151,1797]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
<emphasis id="B918EA8BFFD0F323FF60972BD2FEF948" bold="true" box="[199,455,1662,1687]" pageId="35" pageNumber="36">Female. Color, head:</emphasis>
Black, except for yellow-ferrugineous marks as follows: two-third apical region of mandibles; labrum; clypeus (except for a dark sport on center); scape, pedicel and mostly F8–F10. A narrow band on upper half of gena, adjacent to the compound eye.
<emphasis id="B918EA8BFFD0F323FCB39793D0A5F900" bold="true" box="[788,924,1734,1759]" pageId="35" pageNumber="36">Mesosoma:</emphasis>
Ferrugineous, with black marks as follows: mesoscutum (with two wide ferrugineous spots on each side of it); narrow transverse stripe on apical edge of metanotum; broad longitudinal stripe on posterior face of the propodeum; entirely mesepimeron and anterior region from the lateral face of the propodeum. Parategula and metanotum with yellow marks. Legs yellowferrugineous, with black marks as follows: coxa, trochanter and basal half of fore leg; trochanter and inner surface of mid legs; inner half of coxa, trochanter and entirely femur of hind legs. The external surface of the mid and hind tarsi is also slightly blackish.
<emphasis id="B918EA8BFFD0F323FE4196CBD155F868" bold="true" box="[486,620,1950,1975]" pageId="35" pageNumber="36">Metasoma:</emphasis>
dorsal and ventral surface of T1 black with lateral surface ferrugineous. Dorsal surface of T2 with a blackish broad longitudinal stripe while lateral surface is reddish-ferrugineous with some blackish stripes. Apex border, including lamella, yellow. This color pattern of T2 also occurs for S2. Apical half of T3–T6 and S3–S6 yellow.
<emphasis id="B918EA8BFFD7F324FD9B91E9D1A8FF0A" bold="true" box="[572,657,188,213]" pageId="36" pageNumber="37">Wings:</emphasis>
hyaline, except for costal medial and submedial cells darkened. Pterostigma and veins black.
</paragraph>
<paragraph id="8BD33699FFD7F324FF609051D0EEFDC7" blockId="36.[151,1437,151,1797]" pageId="36" pageNumber="37">
<emphasis id="B918EA8BFFD7F324FF609051D29AFEC2" bold="true" box="[199,419,260,285]" pageId="36" pageNumber="37">Pubescence, head:</emphasis>
Clypeus with short whitish (better observed in oblique view) pubescence. Frons, vertex and occipital region with long and golden pubescence, while on gena it is short (shorter than on clypeus). Scape with golden pubescence as short as on clypeus.
<emphasis id="B918EA8BFFD7F324FD609019D077FEBA" bold="true" box="[711,846,332,357]" pageId="36" pageNumber="37">Mesosoma:</emphasis>
pronotum, mesoscutum, scutellum and metanotum with long golden pubescence (slightly shorter on pronotum and mesoscutum). Mesepisternum and posterior and lateral face of propodeum with whitish pubescence, longer on the latter region. Trochanter and basal half of femur of the fore leg with erect golden bristles. External surface of tibiae also with golden and erect bristles.
<emphasis id="B918EA8BFFD7F324FAB190E2D6A5FE0F" bold="true" box="[1302,1436,439,464]" pageId="36" pageNumber="37">Metasoma:</emphasis>
Covered with golden pubescence. Short pubescence on dorsal face of T1 and other segments. Ventral surface of T1 with scattered long bristles. Apex of the other segments with long bristles.
</paragraph>
<paragraph id="8BD33699FFD7F324FF609371D6A4FD5B" blockId="36.[151,1437,151,1797]" pageId="36" pageNumber="37">
<emphasis id="B918EA8BFFD7F324FF609371D127FDE2" bold="true" box="[199,542,548,573]" pageId="36" pageNumber="37">Surface of integument, head:</emphasis>
Weak but evident punctation homogeneously distributed. Frons with coarse and moderately dense punctation; reduced punctation behind the ocelli and on gena.
<emphasis id="B918EA8BFFD7F324FB829312D795FDBF" bold="true" box="[1061,1196,583,608]" pageId="36" pageNumber="37">Mesosoma:</emphasis>
Covered with coarse and moderately dense punctation as on frons, except on basal half of mesepisternum where it is sparser.
</paragraph>
<paragraph id="8BD33699FFD7F324FF3093DAD2F2FCCA" blockId="36.[151,1437,151,1797]" pageId="36" pageNumber="37">
<emphasis id="B918EA8BFFD7F324FF3093DAD224FD77" bold="true" box="[151,285,655,680]" pageId="36" pageNumber="37">Metasoma:</emphasis>
Dorsal surface of T1 with coarser and moderately dense punctation, becoming denser on apex. Lateral surface of T1 with scattered punctures, while ventral surface without evident punctation. Dorsal surface of T2 with coarse and dense punctation, becoming slightly weaker and sparser towards lateral surface. T3–T6 and S2–S6 without evident punctation.
</paragraph>
<paragraph id="8BD33699FFD7F324FF60924AD038FB7F" blockId="36.[151,1437,151,1797]" pageId="36" pageNumber="37">
<emphasis id="B918EA8BFFD7F324FF60924AD2B3FCE7" bold="true" box="[199,394,799,824]" pageId="36" pageNumber="37">Structure, head:</emphasis>
Apex of labrum straight; clypeus convex, longer than wide, with the apex concave forming two rounded projections with weak carinae on apex (
<figureCitation id="13572A1CFFD7F324FD539211D073FC82" box="[756,842,836,861]" captionStart="FIGURES 66–76" captionStartId="47.[151,264,1772,1794]" captionTargetBox="[151,1436,193,1750]" captionTargetId="figure@47.[151,1436,193,1750]" captionTargetPageId="47" captionText="FIGURES 66–76. Clypeus. 66) Eumenes americanus (female); 67) E. smithii (female); 68) E. crucifera (female); 69) E. crucifera (male); 70) E. verticalis (female); 71) E. fraternus (female); 72) E. fraternus (male); 73) E. sculleni (female); 74) E. filiformis (female); 75) E. rufomaculatus (female); 76) E. versicolor (female). Scale bar = 1mm." httpUri="https://zenodo.org/record/1458316/files/figure.png" pageId="36" pageNumber="37">Fig. 75</figureCitation>
); inter-antennal area wider than the antennal socket, with a central tubercle; lateral ocelli closer to the compound eye than to each other; occipital carina angled on middle region of gena and close to the compound eyes next to the mandible.
<emphasis id="B918EA8BFFD7F324FBA092D9D7B7FC7A" bold="true" box="[1031,1166,908,933]" pageId="36" pageNumber="37">Mesosoma:</emphasis>
pronotal fovea present; pronotal present dorsally and and more developed laterally; humeral region weakly projecting; mesepimeron slightly elevated regarding the mesepisternum; posterior projection of tegula weakle developed and rounded; parategula lamelliform; posterior face of propodeum strongly concave on basal half. Basal region of the external surface of fore coxa with a short and weak carina.
<emphasis id="B918EA8BFFD7F324FD6E9549D076FBEA" bold="true" box="[713,847,1052,1077]" pageId="36" pageNumber="37">Metasoma:</emphasis>
T1 very long, more than five times longer than wide (
<figureCitation id="13572A1CFFD7F324FF38956AD3CDFB87" box="[159,244,1087,1112]" captionStart="FIGURES 27–42" captionStartId="43.[151,264,1902,1924]" captionTargetBox="[151,1435,193,1881]" captionTargetId="figure@43.[151,1436,193,1881]" captionTargetPageId="43" captionText="FIGURES 27–42. First tergum, dorsal view. 27) Eumenes americanus; 28) E. bequaerti; 29) E. bollii; 30) E. consobrinus; 31) E. coyotae; 32) E. crucifera; 33) E. cubensis; 34) E. flavitinctus; 35) E. fraternus; 36) E. pictus; 37) E. sculleni; 38) E. smithii; 39) E. verticalis; 40) E. filiformis; 41) E. rufomaculactus; 42) E. versicolor. Scale bar = 1mm." httpUri="https://zenodo.org/record/1458308/files/figure.png" pageId="36" pageNumber="37">Fig. 41</figureCitation>
); T2 longer than wide, with the lateral margins weakly concave (
<figureCitation id="13572A1CFFD7F324FC47956AD70EFB87" box="[992,1079,1087,1112]" captionStart="FIGURES 43–58" captionStartId="44.[151,264,1946,1968]" captionTargetBox="[202,1388,195,1934]" captionTargetId="figure@44.[201,1388,193,1934]" captionTargetPageId="44" captionText="FIGURES 43–58. Second tergum, dorsal view. 43) Eumenes americanus; 44) E. bequaerti; 45) E. bollii; 46) E. consobrinus; 47) E. coyotae; 48) E. crucifera; 49) E. cubensis; 50) E. flavitinctus; 51) E. fraternus; 52) E. pictus; 53) E. sculleni; 54) E. smithii; 55) E. verticalis; 56) E. filiformis; 57) E. rufomaculactus; 58) E. versicolor. Scale bar = 1mm." httpUri="https://zenodo.org/record/1458310/files/figure.png" pageId="36" pageNumber="37">Fig. 57</figureCitation>
); apical lamella on apex of T2, becoming reduced towards lateral margins. Apical region of T2 (in lateral view) slightly elevated; S2 with a weak longitudinal elevation, better observed in oblique view.
</paragraph>
<paragraph id="8BD33699FFD7F324FF6095F9D7E5FAD2" blockId="36.[151,1437,151,1797]" pageId="36" pageNumber="37">
<emphasis id="B918EA8BFFD7F324FF6095F9D234FB1A" bold="true" box="[199,269,1196,1221]" pageId="36" pageNumber="37">Male:</emphasis>
Clypeus narrower than females; F11 long, without longitudinal carina, apex pointed and surpassing the apical edge of F8, and without microscopic erect bristles on ventral surface (
<figureCitation id="13572A1CFFD7F324FBAD959AD759FB37" box="[1034,1120,1231,1256]" captionStart="FIGURES 77–92" captionStartId="48.[151,264,1946,1968]" captionTargetBox="[151,1436,193,1925]" captionTargetId="figure@48.[151,1436,193,1925]" captionTargetPageId="48" captionText="FIGURES 77–92. First tergum, dorsal view. 77) Eumenes americanus; 78) E. bequaerti; 79) E. bollii; 80) E. consobrinus; 81) E. coyotae; 82) E. crucifera; 83) E. cubensis; 84) E. flavitinctus; 85) E. fraternus; 86) E. pictus; 87) E. sculleni; 88) E. smithii; 89) E. verticalis; 90) E. filiformis; 91) E. rufomaculactus; 92) E. versicolor. Scale bar = 1mm." httpUri="https://zenodo.org/record/1458318/files/figure.png" pageId="36" pageNumber="37">Fig. 91</figureCitation>
). S7 flattened apically with erect bristles on apex. Color, pubescence, surface of integument and other structure as in females.
</paragraph>
<paragraph id="8BD33699FFD7F324FF609442D776FA42" blockId="36.[151,1437,151,1797]" pageId="36" pageNumber="37">
<emphasis id="B918EA8BFFD7F324FF609442D245FAEF" bold="true" box="[199,380,1303,1328]" pageId="36" pageNumber="37">Male genitalia:</emphasis>
Aedeagus as in figure 107a, b. Paramere (
<figureCitation id="13572A1CFFD7F324FCCE9442D0EFFAEF" box="[873,982,1303,1328]" captionStart="FIGURES 102–109" captionStartId="50.[151,264,1926,1948]" captionTargetBox="[151,1436,193,1903]" captionTargetId="figure@50.[151,1436,193,1904]" captionTargetPageId="50" captionText="FIGURES 102–109. Male genitalia: a, b) aedeagus (lateral and ventral view, respectively); c) paramere, internal view. 102) E. pictus; 103) E. sculleni; 104) E. smithii; 105) E. verticalis; 106) E. filiformis; 107) E. rufomaculatus; 108) E. versicolor; 109) General scheme of the apical projection of the aedeagus: a) weakly concave/truncate; b) strongly concave. Scale bar = 1mm." httpUri="https://zenodo.org/record/1458322/files/figure.png" pageId="36" pageNumber="37">Fig. 107c</figureCitation>
) with long bristles on middle region of gonostyle; digitus flat dorso-ventrally, broad on basal half, becoming narrow towards apex (fig. 124), with the dorsal edge weakly angled. Short bristles along the dorsal edge of digitus, while on the broad region there is no evident bristles. Cuspis with erect short bristles; distal lobe truncate, without bristles.
</paragraph>
<paragraph id="8BD33699FFD7F324FF6094F2D094F98F" blockId="36.[151,1437,151,1797]" pageId="36" pageNumber="37">
<emphasis id="B918EA8BFFD7F324FF6094F2D278FA1F" bold="true" box="[199,321,1447,1472]" pageId="36" pageNumber="37">Variation:</emphasis>
Clypeus yellowish, sometimes with wide black spot on center. Marks on metasoma and mesoscutum may vary in size and may present some yellow marks mostly on the pronotum and propodeum. Some specimens, mostly from
<collectingCountry id="F37B7609FFD7F324FE1194BAD137F9D7" box="[438,526,1519,1544]" name="Mexico" pageId="36" pageNumber="37">
<collectingRegion id="49A8F87BFFD7F324FE1194BAD137F9D7" box="[438,526,1519,1544]" country="Mexico" name="Mexico" pageId="36" pageNumber="37">Mexico</collectingRegion>
</collectingCountry>
, showed a strong sinuosity on the dorsal surface of T2, although in the same locality we examined specimens with weak sinuosity. Males of these specimens had their genitalia examined and no significant differences were observed from those in other localities.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFD7F324FF609709D2C2F8DA" pageId="36" pageNumber="37" type="materials_examined">
<paragraph id="8BD33699FFD7F324FF609709D2C2F8DA" blockId="36.[151,1437,151,1797]" pageId="36" pageNumber="37">
<emphasis id="B918EA8BFFD7F324FF609709D244F9AA" bold="true" box="[199,381,1628,1653]" pageId="36" pageNumber="37">
<typeStatus id="54D7883BFFD7F324FF609709D23BF9AA" box="[199,258,1628,1653]" pageId="36" pageNumber="37">Type</typeStatus>
material:
</emphasis>
<typeStatus id="54D7883BFFD7F324FE2C9708D2C2F9AA" box="[395,507,1629,1653]" pageId="36" pageNumber="37" type="lectotype">Lectotype</typeStatus>
<specimenCount id="9D6AFD10FFD7F324FDAF9708D126F9AB" box="[520,543,1629,1652]" pageId="36" pageNumber="37" type="female">♀</specimenCount>
:
<taxonomicName id="4C6C4D1AFFD7F324FD949708D0FBF9AB" authority="Fox, 1899" authorityName="Fox" authorityYear="1899" box="[563,962,1628,1652]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="rufomaculata">
<emphasis id="B918EA8BFFD7F324FD949708D005F9AB" box="[563,828,1628,1652]" italics="true" pageId="36" pageNumber="37">Eumenes rufomaculata</emphasis>
Fox, 1899
</taxonomicName>
(
<collectionCode id="ED7DAE5CFFD7F324FC709709D70BF9AB" box="[983,1074,1628,1652]" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="36" pageNumber="37">CMNH</collectionCode>
): ‘April’ ‘Corumbá’ ‘lowland’ ‘
<taxonomicName id="4C6C4D1AFFD7F324FF0797D4D23CF947" box="[160,261,1665,1688]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA8BFFD7F324FF0797D4D23CF947" box="[160,261,1665,1688]" italics="true" pageId="36" pageNumber="37">Eumenes</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6C4D1AFFD7F324FEBA97D5D28DF947" authorityName="Fox" authorityYear="1899" box="[285,436,1664,1688]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="rufomaculata">
<emphasis id="B918EA8BFFD7F324FEBA97D5D28DF947" box="[285,436,1664,1688]" italics="true" pageId="36" pageNumber="37">rufomaculata</emphasis>
</taxonomicName>
\
<typeStatus id="54D7883BFFD7F324FE6A97D4D136F946" box="[461,527,1665,1689]" pageId="36" pageNumber="37">Types</typeStatus>
Fox’ ‘
<typeStatus id="54D7883BFFD7F324FDFE97D5D1C7F947" box="[601,766,1664,1688]" pageId="36" pageNumber="37" type="lectotype">LECTOTYPE</typeStatus>
<specimenCount id="9D6AFD10FFD7F324FCA197D4D022F947" box="[774,795,1665,1688]" pageId="36" pageNumber="37" type="female">♀</specimenCount>
\
<taxonomicName id="4C6C4D1AFFD7F324FC9497D4D0A1F947" box="[819,920,1665,1688]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA8BFFD7F324FC9497D4D0A1F947" box="[819,920,1665,1688]" italics="true" pageId="36" pageNumber="37">Eumenes</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6C4D1AFFD7F324FC1697D5D7FAF947" authority="Fox, 1899" authorityName="Fox" authorityYear="1899" box="[945,1219,1664,1688]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="rufomaculata">
<emphasis id="B918EA8BFFD7F324FC1697D5D771F947" box="[945,1096,1664,1688]" italics="true" pageId="36" pageNumber="37">rufomaculata</emphasis>
Fox, 1899
</taxonomicName>
\ des. Carpenter &amp; Vecht’.
<typeStatus id="54D7883BFFD7F324FF4997F1D2BFF962" box="[238,390,1700,1725]" pageId="36" pageNumber="37" type="paralectotype">Paralectotype</typeStatus>
<specimenCount id="9D6AFD10FFD7F324FE2997F0D29CF963" box="[398,421,1701,1724]" pageId="36" pageNumber="37" type="female">♀</specimenCount>
:
<taxonomicName id="4C6C4D1AFFD7F324FE1597F0D017F963" authority="Fox, 1899" authorityName="Fox" authorityYear="1899" box="[434,814,1700,1724]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="rufomaculata">
<emphasis id="B918EA8BFFD7F324FE1597F0D18CF963" box="[434,693,1700,1724]" italics="true" pageId="36" pageNumber="37">Eumenes rufomaculata</emphasis>
Fox, 1899
</taxonomicName>
(
<collectionCode id="ED7DAE5CFFD7F324FC9A97F1D0A1F963" box="[829,920,1700,1724]" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="36" pageNumber="37">CMNH</collectionCode>
): ‘April’ ‘Pedra Branca’ ‘
<taxonomicName id="4C6C4D1AFFD7F324FB1D97F0D626F963" box="[1210,1311,1701,1724]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA8BFFD7F324FB1D97F0D626F963" box="[1210,1311,1701,1724]" italics="true" pageId="36" pageNumber="37">Eumenes</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6C4D1AFFD7F324FA9297F1D6A5F963" authorityName="de Saussure" authorityYear="1855" box="[1333,1436,1700,1724]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="B918EA8BFFD7F324FA9297F1D6A5F963" box="[1333,1436,1700,1724]" italics="true" pageId="36" pageNumber="37">filiformis</emphasis>
</taxonomicName>
var. \
<taxonomicName id="4C6C4D1AFFD7F324FF70979DD240F93F" authorityName="Fox" authorityYear="1899" box="[215,377,1736,1760]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="rufomaculatus">
<emphasis id="B918EA8BFFD7F324FF70979DD240F93F" box="[215,377,1736,1760]" italics="true" pageId="36" pageNumber="37">rufomaculatus</emphasis>
</taxonomicName>
\ cotype
<specimenCount id="9D6AFD10FFD7F324FE45979CD2CEF93F" box="[482,503,1737,1760]" pageId="36" pageNumber="37" type="female">♀</specimenCount>
Fox’
<emphasis id="B918EA8BFFD7F324FD9D979DD1DCF93F" box="[570,741,1736,1760]" italics="true" pageId="36" pageNumber="37">
‘
<taxonomicName id="4C6C4D1AFFD7F324FD99979DD1DCF93F" authorityName="Fox" authorityYear="1899" box="[574,741,1736,1760]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="rufomaculatus">rufomaculatus</taxonomicName>
</emphasis>
\ Fx’ ‘
<typeStatus id="54D7883BFFD7F324FC89979DD722F93F" box="[814,1051,1736,1760]" pageId="36" pageNumber="37" type="paralectotype">PARALECTOTYPE</typeStatus>
<specimenCount id="9D6AFD10FFD7F324FB84979CD701F93F" box="[1059,1080,1737,1760]" pageId="36" pageNumber="37" type="female">♀</specimenCount>
\
<taxonomicName id="4C6C4D1AFFD7F324FBE8979CD78DF93F" box="[1103,1204,1737,1760]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA8BFFD7F324FBE8979CD78DF93F" box="[1103,1204,1737,1760]" italics="true" pageId="36" pageNumber="37">Eumenes</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6C4D1AFFD7F324FB6C979DD3F6F8DB" authority="Fox, 1899" authorityName="Fox" authorityYear="1899" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="rufomaculata">
<emphasis id="B918EA8BFFD7F324FB6C979DD65BF93F" box="[1227,1378,1736,1760]" italics="true" pageId="36" pageNumber="37">rufomaculata</emphasis>
Fox, 1899
</taxonomicName>
\ des. Carpenter &amp; Vecht’.
</paragraph>
</subSubSection>
</treatment>
</document>
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